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Cellular and Molecular Medicine > > > The BSc Degree Programmes Cancer Biology (B190)
Cancer Biology & Immunology (B131)
Cancer Biology & Virology (BCC5)
Cellular & Molecular Medicine (B130)
Immunology (C551)
Medical Microbiology (C521)
Microbiology (C500)
Pathology & Microbiology (BC15)
Virology and Immunology (C540)
2 > > 2 > > Contents > > >
Introduction 5 The City of Bristol 7 The University of Bristol 8 Cancer Biology, Immunology, 11 Bacteriology, Virology, Stem Cell Biology Teaching and Learning 12 Admissions 13 Year 1 & 2 Units 14 Year in Industry 15 Final Year Research Project & Final Year Topics 16 Postgraduate Opportunities 17 Research 18 Careers 25 Further Information 26 “The Department aims to provide an exciting and challenging academic environment where students and staff can achieve their learning, teaching and research goals in cellular and molecular medicine.” Anthony Hollander, Professor of Rheumatology & Tissue Engineering and Head of Department Introduction > > >
The Department of Cellular and Molecular Medicine is an We are located in the School of Medical Sciences and internationally recognised centre of excellence for our staff include basic scientists as well as clinicians Cancer Biology, Infection and Immunology and Stem working in the major Bristol hospitals. As a result, the Cell Biology. The Department was recognised for its scientific research and teaching of the Department are National and International excellence in the last national conducted with a clear focus on human disease. We Research Assessment Exercise, and was awarded a also have several University spin-out companies maximum 24 out of 24 in the national Teaching Quality embedded within the Department and this allows us to Assurance assessment. These results demonstrate how focus on translating our fundamental discoveries into the Department successfully balances its twin aims of therapies that can be exploited for the benefit of large providing first class teaching whilst carrying out the numbers of patients. This translational research highest quality biomedical research. environment provides an exciting backdrop to the learning experience for our undergraduates who are The Department teaches all aspects of cancer biology, taught both by fundamental scientists and clinicians, as immunology, bacteriology, virology and stem cell biology well as those engaged in commercialisation of medical to its own science undergraduates, to those from other research. departments and to medical and dental students. Our science courses are designed to provide students with a Bristol is an exciting city in which to live and study, and wide variety of choices, allowing them to specialise as the University of Bristol is one of the top universities in their interests develop. the UK. Set against this very attractive background, the Department aims to provide a stimulating and enjoyable All our members of staff draw upon their research environment where students and staff can achieve their expertise to ensure that the teaching enjoyed by our learning, teaching and research goals in the fullest students reflects the cutting edge of the subject matter. possible sense. Indeed, a key element of our undergraduate courses is a major final year research project carried out within one of This brochure will give you a flavour of what the the internationally recognised research teams in the Department has to offer in terms of its teaching and Department. This gives our students experience of the research, and we hope you find it informative and research process and provides them with a competitive interesting. edge when applying for higher degrees or future employment. The projects allow our students to Anthony Hollander contribute to the solving of important problems in Professor of Rheumatology & Tissue contemporary cellular and molecular medicine. Engineering and Head of Department
5 > > www.swns.com The City of Bristol
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Bristol is a multicultural city of balloons and kites, clubs John Cabot sailed from here in 1497 on and DJs, festivals and carnivals, architecture and his historic voyage to Newfoundland. parkland, business and new technology, theatres and museums, artists and animators, music and film. It has Isambard Kingdom Brunel built the been officially designated a ‘Centre of Culture’ and a Great Western Railway from London to ‘Science City’ by the Government. In 2008 Bristol was Bristol in the 1830s. given the ‘European City of the Year’ award by the The foundations of solid-state physics Academy for Urbanism. In the same year it was also were laid here in the 1930s by Nevill selected as Provincial City of the Year and Britain’s most Mott and his team. sustainable city. Bristol is also the only UK city to be short-listed for the European Green Capital Award 2010 In Bristol on 9 April 1969, the British- and is England’s first ‘Cycling City’. Bristol is a wonderful built Concorde prototype made its first place to live and is certainly a great place to be a student. flight. Just ask any past or present Bristol students. The Hollywood Oscar winners, The name Bristol comes from the Old English Brycgstow, Aardman Animations, producers of meaning ‘the place at the bridge’. That’s still a good “Wallace and Gromit”, are based in description, and not just because of Brunel’s renowned Bristol. Clifton Suspension Bridge. Bristol is a bridge to culture, experiences, landscapes, careers – and the future. The band Portishead recorded the groundbreaking Dummy album here Bristol has a history of innovation, creativity, exploration (parts of it were recorded in the Music and risk-taking as illustrated by just a few examples Department’s Victoria Rooms) in 1993. opposite. Banksy, the Bristol-born renowned street artist and ‘trickster’, began his career here in the 1990s.
7 > > The University of Bristol > > >
The University College of Bristol opened in 1876 in a bid The University continues to invest to maintain its position to bring university culture to the provinces. It was the first at the forefront of research and teaching and has college in the country to admit men and women on an recently broken ground for £80 million-worth of new equal footing. After much lobbying, government grants accommodation for the departments of Mathematics for the University College began in 1889, and in 1893 the and Biological Sciences. Medical and Engineering Schools were opened. An enormous variety of extramural activities on and off In 1908, gifts from the Wills and Fry families enabled the the sports field is available as you might expect from a College to apply for a charter, and in 1909 the King and major university. These take place at the University’s £5 Privy Seal approved the creation of the University of million Centre for Sport, Exercise and Health as well as at Bristol. George and Harry Wills funded the magnificent the 38 acre Coombe Dingle Sports Complex near the Wills Memorial Building, opened by King George V and Stoke Bishop Halls of Residence. Queen Mary in 1925. This is still the centrepiece of the University precinct. The University has been rated as one of the top ten universities in the UK and in 2009 it was ranked within The University’s reputation grew steadily throughout the the top 35 in the World. The University is a member of the 1930s, helped in no small way by the appointment of Worldwide Universities Network, a grouping of 18 Winston Churchill as its third Chancellor in 1929. Four research-led institutions of international standing as well Nobel Laureates worked at Bristol: Paul Dirac (1933), as a member of the Russell Group of universities, an Cecil Frank Powell (1950), Hans Albrecht Bethe (1967) association of 20 major research-intensive universities in and Neville Mott (1977). the UK.
Bristol undergraduates study in an environment of international-class research
8 > > 9> > 10 > > What are cancer biology, immunology, bacteriology, virology and stem cell biology?
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Cancer Biology > > > However, some bacterial infections are dangerous and as time goes by new antibiotic resistant bacteria are Cancer can be simply described as the multiplication of emerging. The mechanisms by which bacteria acquire a cell in the absence of normal controls to produce a antibiotic resistance and how certain bacteria infect tumour. Needless to say the control of cell growth is humans and cause disease are clearly very important enormously complex and cancer is really a general term issues and are a focus of interest in the Department. for a multitude of different diseases where cell growth control has been lost or compromised. It is expected that an understanding of the cellular and molecular Virology > > > mechanisms of growth control of normal cells and its Another group of equally important microbes is the absence in cancer cells will lead to effective treatments, viruses. It was once said by Nobel Laureate, Joshua prevention and ultimately cures for most, if not all, Lederberg, that viruses are man’s only true competitors human cancer. for domination of the planet. This is almost certainly not true but viruses such as HIV and influenza are clearly Immunology > > > having a massive global impact. The study of viruses is of great importance in world health terms. Immunology is the study of immunity to infection and also encompasses autoimmunity where destructive immune responses are made against our own bodies. A Stem Cell Biology > > > healthy immune system can distinguish between self All the specialised cells in our body are derived from and non-self or foreign biological material such as stem cells. In the early embryo there are pluripotent bacteria and viruses. Ultimately it is hoped that we can stem cells that can turn into any other cell type. In the manipulate the immune system to eliminate foetus or in adults there are tissue specific stem cells autoimmune diseases, to make better vaccines against that can turn into a more limited repertoire of specialised infections and even to cause the immune destruction of cells. All stem cells, when grown in the laboratory, can cancers. replicate without ageing. This makes it possible to grow up a large number of cells that can be used to treat a Bacteriology > > > wide range of diseases. By studying these cells we will be able to use them to treat diseases that were once Microbes include bacteria, and bacteriology is, of considered incurable. course, the study of bacteria. They are found almost everywhere and in the main they do not cause harm. In fact they are essential to our lives in many ways.
11 > > Teaching and Learning > > >
The BSc Honours Programmes are broadly based, and allow students flexibility to tailor their studies to their own developing interests.
• Cancer Biology • Cancer Biology and Immunology • Cancer Biology and Virology • Cellular and Molecular Medicine • Immunology • Medical Microbiology • Microbiology • Pathology and Microbiology • Virology and Immunology
During the first and second years, at any one time, students will be taking 3 courses. In general, each will consist of three hours of lectures and a three-hour laboratory class each week. Courses include a substantial element of continual assessment in the form Undergraduates will be assigned different specialist of essays and tutorials that contribute about 20% to the academic tutors in different parts of the degree final sessional mark. During the final year, students programme but will be assigned a Personal Tutor when attend lectures in their chosen topics (on average 5 they arrive in Bristol who will be available for advice and lectures/week), work in the library following up the consultation on any issue, personal or otherwise, original literature relevant to the topics, and usually work throughout the course. at the bench in a research laboratory, to carry out their project. During this time, students are very much their The teaching staff actively seeks student feedback on own masters and they organize their own work every aspect of teaching. Feedback may take the form of accordingly. questionnaires or representation at the Staff/Student Liaison Committee. All marks after the first year contribute towards the final degree grading and marks obtained in year 2 contribute 20% of this. The final year research project accounts for 20% of the final degree grading in accordance with the importance attached to the project work.
12 > > > > > Admissions
A-Levels: Three A levels, with A2 in Chemistry and at least one other A2 in science, are required in all cases.
General Studies: General Studies and Critical Thinking are not included in the terms of conditional offers.
Typical Offers: A-levels: ABB. Scottish Qualifications Authority: AABBB.
International Baccalaureate: 33 points with 6 points (in exceptional cases five) normally required in at least two science subjects (including Chemistry) at Higher Level.
The English Language requirement is IELTS 6.5 in all parts.
Selection: Initial assessment is based on the UCAS form. Shortlisted candidates (except for overseas Candidates taking Access or vocational courses should applicants) are usually invited to attend a UCAS Day be able to demonstrate the relevance of their which includes a one-to-one meeting with a member of qualifications to their chosen course of study. staff. In particular, Chemistry to the equivalent standard of Deferred Entry: Applications are welcomed. A2-level is required.
Broadening Access: The Department welcomes International Students: The department welcomes applications from mature candidates and from applications from international students for both candidates from non-traditional backgrounds. undergraduate and postgraduate study.
13 > > Year 1 Units
Introduction to Microbiology Normal and Tumour Cells Biochemistry I Microorganisms and their impact The structure of mammalian cells Structure and function of proteins, on society and the environment. and tissues; membrane trafficking, nucleic acids, polysaccharides; growth control, developmental enzymes, enzyme kinetics; genetic biology and differentiation; tumour replication and expression; basic Neil Williams cells and the haemopoietic system. metabolic pathways. Duis autem vel eum iriure dolor in
Weeks 1-12 Weeks Membrane structure and function; hendrerit in vulputate velit esse oxidativemolestie consequat,phosphorylation, vel illum dolore photosynthesis.eu feugiat nulla facilisis Control at vero of eros metabolicet accumsan cycles; et iusto hormones odio dignissi and secondary messengers as
Microbes and Disease Pathological Responses signalling mechanisms. Protein The causal relationship between of Cells synthesis. Biochemistry of infection and disease; intervention The basic principles of development. strategies. pathogenesis; development of specific human diseases; how experimental studies have defined the mechanisms underlying
Weeks 13-24 Weeks pathological processes.
Year 2 Units
Infection and Recombinant DNA Membranes and Immunity Technology Signalling Immune defences Genetic structure; Properties of the cell against infection; principles of gene surface; cell immune evasion by cloning; cloning in plants communication; action viruses and bacteria; and animals; DNA of hormones, growth newly emergent sequencing; PCR; factors; secretion Weeks 1-12 Weeks infections; vaccination protein engineering. pathway.
Cellular and Gene Expression Cellular & Molecular Molecular Pathology and Rearrangement Systems Cellular and molecular DNA replication, repair Protein synthesis and basis of diseases not and recombination; gene degradation; the directly related to expression; antibody immune system; infectious variation; molecular basis phagocytic blood cells;
Weeks 13-24 Weeks microorganisms. of disease; animal and chemotaxis; muscle plant molecular genetics. contraction.
14 > > Students who do well in their first two years may, if selected, spend a year in industry between their second and final years. Successful students will be funded by the company (normally a major pharmaceutical firm) that they are placed with during the year.
15 > > Final Year Research Project
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At the beginning of the final year students will become active members of one of the research groups in the department for about four months. They will choose a research project and will investigate a real life research problem (nobody knows what the answers will be) under supervision. We consider this project to be one of the most important parts of the degree programme. As a result, our students have quite an advantage when applying for PhD research studentships because they will have real research experience.
Final Year Topics
Microbiology Topics Pathology Topics
Cancer Biology Immunology
Genetics & Advanced Medical Microbiology Development Immunology
Molecular & Cellular Neoplasia 1 Immunopathology Bacteriology
Frontiers in Virology Neoplasia 2 Applied Immunology
Medical Virology Stem Cell Transplantation: From Science To Application
16 > > > > > Postgraduate Opportunities
The majority of our postgraduate students in the Department undertake full-time research at Ph.D. level. However students may also undertake a M.Sc. or M.D. by research or study part-time. These individual programmes of study may start at any time of the year, as is convenient to the student and the research adviser(s), although most students commence their study period at the beginning of October.
Students who enrol in the Department are automatically members of the Graduate School of the Faculty of Medical and Veterinary Sciences. The Graduate School offers an induction course to all new postgraduate students. At this induction new students are given information about the University and the services it offers. They are told what is expected of them throughout their programme of study and what they can expect from the Department and the University by way of support.
Research training is primarily undertaken within the This course is provided jointly by the Department of research group that the student joins, but access to a Cellular and Molecular Medicine and the Bristol wide range of experimental expertise and training Institute for Transfusion Sciences. Further information outside the particular research group is readily available on this course is available on the internet at: in the Department, in other University departments or in http://www.bristol.ac.uk/cellmolmed/postgrads/post an appropriate hospital laboratory. grad-taught/ The Department is well equipped to undertake basic In addition, collectively all the Departments of the School research in many areas, particularly in the areas of of Medical Sciences offer a separate taught/research immunology, molecular biology, molecular genetics, ‘M.Sc. in Biomedical Sciences Research’ that aims to cancer, virology, stem cell biology and aspects of deliver a thorough training in key aspects of biomedical medical microbiology. experimental research techniques. Further information A taught programme is offered in Transfusion and about this course can be found at Transplantation Sciences leading to an MSc degree. http://www.bris.ac.uk/biochemistry/postgrad/msc.html
17 > > Research
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The aim of our research programme is to turn science growth of spin-out companies whenever this is into medicine. We have provided an ideal environment in appropriate. This strategy runs across our entire which to both undertake world class fundamental research portfolio encompassing the scientific fields of research and translate this research into therapeutic immunology, bacteriology, virology, cancer biology and strategies that can be tested clinically and exploited stem cell biology. In this way we hope eventually to commercially. This has been achieved by fostering close provide new or improved treatments for diseases such links between laboratory-based scientists and clinical as multiple sclerosis, colon cancer, meningitis and investigators and by supporting the formation and osteoarthritis.
The Department has completed a major refurbishment of its research facilities. This has created an outstanding working environment with state-of-the-art facilities. The Department also has access to the Wolfson Bioimaging Facility, providing imaging, confocal and electron microscopy facilities within the Faculty of Medical and Veterinary Sciences.
18 > > Cancer Biology
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Cancer is a major cause of death in the UK and the lifetime risk of developing cancer is about one in three. Over 50% of cancers are preventable.
Cancer can be caused by poor diet, tumour viruses, radiation and inherited mutations. The overall research aim is to increase our understanding of the cellular and molecular basis of cancer and to bring advances in these areas to the clinic, in terms of prevention strategies, early diagnosis and targeted novel treatments. Examples of some research areas are as follows: Research in the Cancer Research UK (CRUK) Colorectal Tumour Biology Group investigates how dietary factors and aspirin-like drugs force tumour cells to commit suicide (Chris Paraskeva and Ann Williams). childhood cancers for diagnostic and therapeutic The CRUK group investigates other potentially novel purposes. Other areas of interest include: E- natural agents, such as plant-derived cannabinoids, for cadherin/catenin complexes in colorectal neoplasia; cancer prevention and therapy. Other research involves molecular cross talk between E-cadherin and the EGF the use of high throughput technologies to enable the receptor; dietary fibre and colorectal cancer; and examination of many different types of childhood and epithelial regeneration and mucosal repair (Massimo adult cancers for genome-wide genetic and epigenetic Pignatelli and David Qualtrough); vaccines against changes (Keith Brown and Karim Malik). This research, it tumour viruses (Andy Morgan); and exploiting the is hoped, will identify new genes involved in adult and immune system for novel therapies (David Morgan).
The Department of Cellular and Molecular Medicine is an internationally recognised centre of excellence for the study of Cancer, Infection and Immunology.
19 > > Immunology
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Our immunology research focuses on two major themes of inflammatory diseases and mucosal immunology.
Research groups within the department work on both basic and applied aspects of immunology. The immune system has evolved in order to protect us from infectious diseases. However, under some circumstances it can be the cause of diseases such as autoimmunity and allergy.
Work on autoimmune diseases includes studies into multiple sclerosis (David Wraith, Lindsay Nicholson) and type 1 diabetes (Susan Wong). These diseases arise when the immune system is active when it should not be. Certain bacterial products can modulate immune Discoveries include mapping cytotoxic T-cell epitopes responses, and work is using one such molecule as a and early immune events in diabetes and the role of T potential asthma therapy (Neil Williams). Other work regulatory cells and cellular tuning in controlling the seeks to devise better approaches to vaccination against immune response. Conversely, in cancer there is a need bacteria that cause meningitis through understanding to make the immune system respond more vigorously. the development of the natural immune response to Our work investigates ways in which antigen presenting these common bacteria in the back of the throat (Neil cells handle tumour antigens, and how tumours may Williams, Adam Finn). suppress immunity (David Morgan). All of our work seeks to balance providing a basic Interests in immune responses at mucosal surfaces understanding of the processes that underlie disease include the studies on how tissues protect themselves and seeking new ways of treating disease. In addition, from immune attack (Gabriele Braun) and how these clinical trials in diabetes, multiple sclerosis and asthma processes can fail leading to ocular diseases (Andrew are being carried out based on the work of members of Dick, Lindsay Nicholson). We also investigate the the department, and new approaches for the relationship between pathogens and the mucosal development of mucosal vaccines for infectious immune system. diseases and for cancer are being developed.
20 > > > > > Bacteriology
Our main bacteriological research interests broadly cover two areas: Pathogenicity and Anti-microbial Resistance.
We are aiming to identify and understand molecular structures that aid the bacterial organism to breach the immune surveillance system (Mumtaz Virji and Darryl Hill) or adversely stimulate the immune system and thereby cause tissue damage. This area of study includes Neisseria meningitidis (Virji) and Haemophilus influenzae (Virji and Hill). Such studies are vital for the design of vaccines that are currently being developed.
Two members of the department (Ariel Blocker and Sarah Daniell) are performing structural studies on the mechanism by which pathogenic bacteria interact with human cells.
Unravelling the complexity of antibiotic resistance is the goal of Matthew Avison, Jim Spencer and Vicky Enne. This research group works together with clinical microbiologists within BCARE (Bristol Centre for resistance gene silencing (Enne); and structure/function Antimicrobial Resistance and Evaluation). Their work studies on proteins that facilitate pathogenicity and/or includes studies on gene regulation and expression of antimicrobial resistance (Spencer). The breadth of resistance determinants from their normal quiescent interest and depth of knowledge of this group have state (Avison); biological cost of antibiotic resistance and contributed to their international reputation.
21 > > Virology
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Virologists in the Department study a number of important viruses that cause diseases in humans.
Both RNA and DNA viruses are being studied including the coronavirus that causes severe acute respiratory syndrome (SARS) (Stuart Siddell and Andrew Davidson), the arthropod-borne flavivirus that causes dengue fever (Andrew Davidson), and Epstein-Barr virus, which is associated with cancers in humans (Andy Morgan). Work is also going on to understand how adenoviruses interfere with cellular systems in the nucleus of the infected cell (David Matthews). The molecular and cellular biology of these viruses are being investigated and, in particular, the way that they interact with the host immune response. This information is being used to devise new ways to control infection. This includes the development of vaccines that prevent virus infection in the first place, antiviral drugs that prevent replication of the virus after We aim to provide a challenging and comprehensive infection has taken place, and therapies that aim to training to both undergraduate and graduate students contain the consequences of virus infection, for example and a stimulating atmosphere in which to conduct first- autoimmune reactions or cancer cells that proliferate class, internationally competitive research that will have uncontrollably. an impact upon both human and animal health.
One of the main aims of the Department is to translate basic research findings directly into clinical applications.
22 > > > > > Stem Cell Biology
Stem cells can replicate without ageing and can be turned into many kinds of specialised cell for use in treating chronic destructive diseases.
Embryonic stem cells have the unique capacity to make every cell type in the body and to grow indefinitely without ageing. This means they could theoretically be used to treat any disease once we understand how to control them and turn them into the cells that we need. Adult stem cells are found in most tissues and organs of the body, where they reside in “niches” allowing them to reproduce without ageing. Their ultimate role is to leave their niche, and then turn into all of the specialised cells needed to maintain tissue or to repair injuries.
Research is underway exploring how stem cells grow and mature into different cell types. The ultimate aim is to be able to develop methods that can be used to create a (ALL) to therapy, as well as improved ways of diagnosing range of cells for the treatment of different diseases (Wael viral infection after stem cell transplantation. There is Kafienah). For example, stem cells are grown to produce also interest in rare diseases of the blood system and new cartilage for treatment of patients with arthritis and metabolic disorders, as well as in translational research related diseases (Anthony Hollander). and improving diagnosis of diseases (Colin Steward). Other research focuses on identifying the stem cells in Research at Bristol Children’s Hospital has resulted in the the bone marrow which may explain relapse of children development of a test, now used by the NHS to monitor undergoing treatment for ALL. The aim is to develop the response of children’s acute lymphoblastic leukaemia better anti-leukaemia drugs (Allison Blair).
23 > > James Barke
The quality of our teaching has been recognised by external assessors. In the ‘Molecular Biosciences’ category we were awarded the maximum 24 points by the Quality Assurance Agency.
24 > > Careers
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Our graduates will have acquired practical research skills Our other graduates go directly into employment in a and an ability to understand complex scientific data. wide variety of careers including education and They will have excellent written and oral communication biomedical research, finance and law, health and skills, as well as numerical, analytical and problem- community work, information technology, management solving abilities. Graduates will also have gained consultancy, manufacturing and many others. experience in several important aspects of information technology during their studies. Graduating with any one of the degrees offered by the Department not only opens the way into professional Many of our graduates go on to further study. A and research careers in biomedical science but also into significant number take up M.Sc. or Ph.D. studentships almost any other area where a scientific approach to in a wide range of biomedical sciences including solving problems is valued. immunology, microbiology, cancer biology, stem cell biology, transfusion & transplantation sciences, toxicology, parasitology and dietetics. Others go on to medicine, dentistry, veterinary science and teaching.
25 > > Further Information
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Departmental Address Vacancies Department of Cellular and Molecular Medicine, All applications should be addressed to School of Medical Sciences, The Recruitment Team, Personnel Services, University of Bristol, University Walk, University of Bristol, Senate House, Bristol, BS8 1TH; Bristol, BS8 1TD, UK. telephone +44 (0)117 954 6947; Email: [email protected] minicom +44 (0)117 928 8894 or Telephone (General Office): email [email protected] quoting the job +44 (0)117 33 12050 reference. Applications can also be submitted on-line Fax: +44 (0)117 33 12091. by following instructions accompanying the advertised job details at http://www.bris.ac.uk/ Undergraduate Admissions cellmolmed/vacancies/index.html All applications must be made through the Universities and Colleges Admissions Service For more general enquiries, or if you (UCAS). Requests for further information should be can’t find what you are looking for in addressed to the Tutor for Undergraduate these pages, please send an email to Admissions (Dr Ann Pullen). She can be contacted [email protected] by email: [email protected] This address should not be used for by telephone: +44 (0)117 33 12025 Undergraduate or Postgraduate or fax: +44 (0)117 33 12091. applications.
Postgraduate Admissions Further information on all areas covered All applications for postgraduate research training or in this brochure can be found at: enquiries for further information should be addressed http://www.bristol.ac.uk/cellmolmed/ to the Tutor for Graduate Admissions (Dr Andrew Davidson). He can be contacted by email: [email protected] by telephone: +44 (0)117 33 12024 or fax: +44 (0)117 33 12091.
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Location Disclaimer Acknowledgements Road, rail and air connections Every care has been taken to Andy Stevens Photography Ltd. from Bristol are excellent, ensure the contents of this [email protected] there being two main line rail brochure are correct at the stations, the M4 and M5 time of going to press. motorways and an However, the University and international airport with the Department of Cellular direct flights to other parts of and Molecular Medicine are the UK, to the USA, to continually striving to improve Europe and connections the quality of their research, worldwide. teaching, and facilities. The University therefore reserves the right to make changes from time to time both in relation to individual programmes of study and the facilities and services available.
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Edinburgh
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Cardiff London BRISTOL > > >
Department of Cellular and Molecular Medicine
School of Medical Sciences University of Bristol University Walk, Bristol, BS8 1TD, UK Tel: +44 (0)117 33 12050 Fax: +44 (0)117 33 12091 www.bristol.ac.uk/cellmolmed/