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Carboniferous Rugosa and Heterocorallia In ACT A PAL A EON T 0 LOG I CAP 0 LON• ICA Vol. 22 1977 Nr. 4 NGUYEN DUC KHOA CARBONIFEROUS RUGOSA AND HETEROCORALLIA FROM BOREHOLES IN THE LUBLIN REGION (POLAND) Abstract. - Fourty two species referred to 21 genera and 2 subgenera are described; one genus and 5 species are new: Diphyphyllum rarevesiculosum sp.n., Lithostro­ tion (Siphonodendron) dobrolyubovae sp.n., L. (S.) rossicum strzelcense subsp.n., L. (S.) rossicum parvum subsp.n., Lublinophyllum fedorowskii gen. et sp.n. Genera Orionastraea and Turbinatocaninia are for the first time recorded in Poland. Hete­ rophyllidae with 5 septa and pentagonal cross-section are recorded. Blastogeny has been studied in 13 species and subspecies. Axial increase has been studied in detail in Lithostrotion (Siphonodendron) dobrolyubovae sp.n. Taxonomic division of Ru­ gosa based on blastogeny, is demonstrated to be questionable in some cases. Fusion of corallites has been observed in colonies of Lithostrotion and Lonsdaleia colonies. Existence of holotheca in Orionastraea aff. magna Kato & Mitchell has been proved. INTRODUCTION Carboniferous Rugosa and Heterocorallia in Polapd are known from the outcorps and boreholes in Sudetes, Silesia-Cracow Upland, Holy Cross Mts, Lublin region and Western Pomerania (Kunth 1869; Schindewolf 1941, 1942, 1952; Fedorowski 1968, 1970, 1971, 1975). Their stratigraphic and geographic distribution is shown in Table 1. In the Lublin region the Carboniferous system is represented by Visean, . Namurian, and Westfalian deposits. History of geological investigations in this region as well as descriptive palaeontology, lithology, and tectonics are presented in the studies by Bojkowski (1966a,b), Cebulak & Porzycki (1966), Porzycki (1976), and Zelichowski (1972c) among others. Some bo­ reholes investigated in this paper have already been described by Mila- . czewski and Zelichowski (1976: Strzeloe IG 1 and 2) and Zelichowski (1961: Tyszowce IG 1; 1968: Kaplonosy IG 1; 1972a: Lublin IG 1). Rugosa occur in the Upper Visean and Namurian deposits, while Heterocorallia ar,e restricted to the former ones. Fedorowski (1968) investigated same cor­ als derived from four-boreholes situated in the region (fig. 1). That fauna consists of 10 species assigned to the following genera: Bradyphyllum, Claviphyllum, Cyathaxonia, Dibunophyllum, Hapsiphyllum, Lithostrotion, 302 NGUYEN Due KHOA Table 1 Silesia Holy Lublin Western ~'""'"' Cracow Cross region Pomerania Uplands Mts 1111111 1111111I Namurian A 1/11111 111111/1 1111/1111 1111111I '/111//1 1111/11 1111III1 I111111I IIII f Upper 1111111I1/1111/11 111/1111 11111/11 IIIII Visean 1111111I 11111111 11111111 1111I11I IIIII 1111111I 11111111 11111111 1111111 IIIII 11/11111 --- -- 11111111I 111/1111/1 Middle 11111111I 11111/1/11 Visean IIIIIII II 11111111/1 11/11111 1111111I11 11111/11 11111/1111 Lower 1111111/ 1111111111 Vis~an 11111111 1111111111 11111111 I~I~/!/!/I- 1/111111 IIIII Tournaisian 11111111 III / I I 11I11111 IIIII lmill 2C] 1 coral bearing deposits, 2 deposits without corals or lacking. ' Trachylasma, Antiphyllum (=Pentaphyllum), and Caninophyllum (=?Bothrophyllum). The Rugosa and Heterocorallia investigated in the present paper are derived from twenty one boreholes, made by the Geological Institute in Warsaw and situated in the Lublin region up to the state border (fig. 1). The collection described is housed in the Museum of the Geological Insti­ tute in Warsaw, abbreviated as IG. In general, the material is not nume­ rous and poorly preserved. Some species are represented by single or few specimens. Acknowledgments Sincere thanks are due to the authorities of the Geological Institute in Warsaw and to Dr. A. Zelichowski from the same Institute for making the core material available to me for study. The present paper has been completed during my stay in Poland, as a post-graduate student of the Faculty of Geology, University of Warsaw CARBONIFEROUS RUGOSA AND HETEROCORALLIA 303 o o 30km Tr.4 • Ks.l" Tr. 2 " I 3~ \ 2. •• \, Tyl. i \ .1 I Warszawa,-' . If 2 Poznar'l \ \tr.3) o 3 U.l~'l~/ !.J_-\U .y '~ ~6 ~8 "J', KrakOw , -, / \ / / "........., .... -_1'"-...... : 7 ~9 / ....... '_:I [ill] ,. Fig. 1. Geological map of Permian subcrops (after A. Zelichowski 1973, simplified) and location of boreholes: B.1 Bialopole IG 1; By.1 Bychawa IG 1; H.1 Husynne IG 1; Ho.1 Holesz6w IG 1; 1.1 Izbica IG 1; J.4 Jarcz6w IG 1; K.1 Krasnystaw IG 1; Ka.1 Kaplonosy IG 1; Ko.2 Kock IG 2; Kr.1, 2 Koczmin IG 1, IG 2; Ks.1 Kosm6w IG 1; L.1 Lublin IG 1; N.1 Niedrzwica IG 1; R.1'. Rachanie IG 1; Ra.1 RadzyiJ. IG 1; Ru.1 Rudnik IG 1; S.l, 2 Strzelce IG 1, IG 2; T.1 Teptiuk6w IG 1; Tr.1, 2, 3, 4 Terebin IG 1, IG 2, IG 3, IG 4; Ty. 1, 2 Tyszowce IG 1, IG 2; V.1 Ulch6wek IG 1; 2.5 2ubowice IG 5. 1 boreholes with coral-bearing core material studied by the present author, 2 boreholes with coral -bearing core material studied by Fedorowski (1968), 3 other boreholes referred in the text, 4 extent of the Carboni- ferous, 5 faults, 6 Upper Visean, 8 Namurian, 9 Devonian. in 1975-1977. The entire project was conduced under guidance of Dr. J. Fedorowski, formerly of the Palaeowological Institute of the Polish Academy of Sciences, Poznan Branch, and now of the Adam Mickiewicz University in Poznan. My appreciation of his professional assistance and personal interest is profound. I hav,e also benefited from informal con­ versation and discussions with many people, prime among them being: Prof. Maria R6zkowska (Poznan), Dr. Ewa Roniewicz (Institute of Pale­ obiology, Polish Academy of Sciences, Warsaw) and to Dr. Antoni Zeli- 304 NGUYEN Due KHOA chowski (Geological Institute, Warsaw). I am also grateful to Prof. Adam Urbanek (Institute of Geology, Warsaw University), and Dr. Magdalena Borsuk-Bialynicka (Institute of Paleobiology, Polish Academy of Sciences, Warsaw) for their thoughtful care during my post-graduate studies. Mr A. Pietura is acknowledged for help in taking the photos. In the text the following abbreviations are used: av - average dimensions c-c - distance between axes of calices d.ax - diameter of axial structure d.c - diameter of corallite d.col - diameter of columella ew - external wall ext - extremal dimensions 1.s - length of septa 1.smj - length of major septa 1.smn - length of minor septa max - maximum dimensions n.lam - number of septal lamellae n.c. - number of corallites in colony n.colis - number of corallites with dissepiments present n.s - number of septa n. t - number of tabulae n.vdis - number of verticils of dissepiments s - septum vdis - verticil of dissepiments w.dism - width of dissepimentarium w.tm - width of tabularium C - cardinal septum K - counter septum All dimensions are in milimeters. In text-figures bar 10 mm long, otherwise stated. STRATIGRAPHIC SIGNIFICANCE OF THE INVESTIGATED CORALS In the Lublin region the Carboniferous is most completely developed in the southwestern part, whereas the thickness of all the members of Carboniferous is reduced northeastwards. It also varies in a very wide range (fig. 2). For example, in the borehole Holeszow IG 1 the thickness of Upper Visean deposits does not exceeds 10 m, while in the borehole Niedrzwica IG 1 it is over 250 m (up to 450 m according to Dr. A. Zelichowski's per­ sonal communication). Occurrence of Tournaisian and Lower to Middle Visean deposits in this area is disputable (e.g. Zakowa in: Osika 1970). The Table 2 Distribution of the desc,ribed species in the Carboniferous of Poland .. '"..c: -a. Distribution ~ on :J 0 N - N ..... .. - V :I: CJ - CJ ... N ..... - - ... - '" CJ CJ - CJ CJ CJ CJ - CJ CJ - .. - - ..., CJ ~ on ... CJ CJ CJ CJ - -- CJ - - - - -- :J ., - - CJ CJ CJ N on U >- -- - - - ., - 0 I Species on ., ., '0 - ., ., c: c: ., - ..'" - - CJ on 0 V V ., .>t -0 c: c: c: c: VV .- .- .>t :J v on ., ., - .>t '":J >- c: u ., c: c: ::J :J a. .- .- .- :J :J EE '0 c: - - -'" 0 -., -., c: .- '0 0 .D -.D .D .D 0 0 NN :J N :J '"v .- .. c: .- .. .. V ~ ~ on .>t >- ., ., NN >- E ... ., ., ., ., N N VV '0 '" 0 c: .. ~ V .D -.D "0 V .D V VI VI VI - -a. ... ... VI a. ... ... ... ... ... ... ... ::J N ::J ... ::J 0 -0 - ::J ::J ., 0 ., ., ., ., >- >- 0 0 .. >- '" '" .. .. .. -'" ::> -, c<: N c<: CD :r -VI CD I- l- I- - -' "" VI VI :r I- "" l- I- l- "" "" '" - "" "" '" Arachnolasma biseptatum Fedorowsk i + +[ + A. cylindricum Yli + + Aulophyllum fungites ( Fleming ) + +1+ + + + + + + + Bothrophyllum juddi ( Thomson ) i? B. pater ( Ivanovsky ) 1+ + I + Bothrophyllum .sp. + ? Caninia cornucopiae brockleyensis ( Thomson ) + ? , I? i Cli .. ioph!lll". de-1 i.G-41 t Will ( ·Smyth ) + + + + + + + j+ Cyathaxonia cornu Michelin + + I + + Dibunophyllum bipartitum ( McCoy ) + + + + + + + + + + + + +1+1+ + D. lonsdaleoides Vasslljuk • D. percrassum Gorsky i I - - + + D. pseudoturbinatum Stuckenberg I - + Dibunophyllum sp.l + + Dibunophyllum sp.2 + Diphyphyllum inge·ns Hill - - D. lateseptatum McCoy I + + + D. rarevesiculosum sp. n. + I + D. simplex ( Thomson ) + + + + + + Koninckophyllum ·interruptum ( Thomson & Ni ch. ) + + + K. meathopense ( Garwood ) + + Lithostrotion (Lithostrotion) portlocki ( Bronn ) - L. (5iphonodendron) afEine ( Fleming ) + + L. (5. ) dobrolyubovae sp. n. + + + L. (5. ) junceum ( Fleming ) + + + + + + + + + + + + + + + + L. (5. ) martini M.Edwards S. Haime + - - - - - - - - - + ? + L. (5. ) rossicum parvum subsp. n. + L. (5. ) rossicum rossicum Stuckenberg + + + + + + L. (5. ) rossicum strzelcense subsp. n. + + + + + + + + + + + + + + L. (5. ) volkovae Dobrolyubova + + + + Lonsdaleia EloriEDrmis floriformis ( Martin ) + - Lublinophyllum Eedorowskii sp.
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