The New El Yunque National Forest Vegetation Map Methodology Note: the Following Document Was Inserted As Originally Written and Formatted

Home , Alsophila (plant), Pouteria multiflora

Forest Plan Assessment

Appendix A: The New El Yunque National Forest Vegetation Map Methodology Note: The following document was inserted as originally written and formatted.

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Appendix B: National Wetland Plant List, Caribbean Region US Army Corp. of Engineer, Puerto Rico 2013 (updated) Final Regional Wetland Plant List: http://rsgisias.crrel.usace.army.mil/NWPL/

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Appendix C: Map Unit Description for El Yunque National Forest Note: Please refer to Map 2-9 for location of soil units. Minor map unit components are not included from this report.

Map Unit/Component Discussion CaD - Caguabo gravelly clay loam, 12 to 20 percent slopes Caguabo (90%) The Caguabo component makes up 90 percent of the map unit. Slopes are 20 to 40 percent. This component is on ridgetops, ledges, and side slopes of strongly dissected uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, lithic, is 10 to 20 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 4 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. CzE - Cristal-Zarzal complex, 5 to 40 percent slopes Cristal (55%) The Cristal component makes up 55 percent of the map unit. Slopes are 5 to 40 percent. This component is on in concave positions in mountain coves and on lower side slopes of volcanic hills and mountains. The parent material consists of clayey colluvium derived from volcanic rock over silty and clayey residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is high. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 15 inches during April, May, June, July, August, September, October, and November. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. Zarzal (40%) The Zarzal component makes up 40 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain ranges on uplands, mountain slopes on uplands. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. DwF - Dwarf-El Duque complex, 5 to 60 percent slopes, windswept Dwarf, windswept (70%) The Dwarf, windswept component makes up 70 percent of the map unit. Slopes are 5 to 60 percent. This component is on mountains, mountain ranges. The parent material consists of residuum weathered from sandstone. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is high. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A

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Map Unit/Component Discussion seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 38 percent. Nonirrigated land capability classification is 3w. This soil meets hydric criteria. El Duque, windswept The El Duque, windswept component makes up 20 percent of the map unit. Slopes are 5 to 60 percent. This component is on (20%) mountains, mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer, bedrock, paralithic, is 10 to 40 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 38 percent. Nonirrigated land capability classification is 6w. This soil meets hydric criteria. GuF - Guayabota-Yunque complex, 20 to 60 percent slopes Guayabota (70%) The Guayabota component makes up 70 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain ranges, mountains. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer, bedrock, paralithic, is 40 to 64 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 17 percent. Nonirrigated land capability classification is 7w. This soil meets hydric criteria. Yunque (25%) The Yunque component makes up 25 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer inches, plinthite. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is high. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 7w. This soil does not meet hydric criteria. HmE - Humatas-Zarzal complex, 5 to 40 percent slopes Humatas (50%) The Humatas component makes up 50 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountain slopes, ridges, uplands. The parent material consists of residuum weathered from igneous rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is high. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 4e. This soil does not meet hydric criteria. Zarzal (45%) The Zarzal component makes up 45 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain ranges on uplands, mountain slopes on uplands. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content

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Map Unit/Component Discussion in the surface horizon is about 80 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria.

HtF2 - Humatas clay, 40 to 60 percent slopes, eroded Humatas (100%) The Humatas component makes up 100 percent of the map unit. Slopes are 40 to 60 percent. This component is on mountain slopes on mountain ranges, ridges on mountain ranges. The parent material consists of residuum clayey and loamy materials. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately high. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 5 percent. Nonirrigated land capability classification is 6e. This soil does not meet hydric criteria. IcA - Icacos loam, occasionally flooded Icacos, occasionally The Icacos, occasionally flooded component makes up 90 percent of the map unit. Slopes are 0 to 5 percent. This component is on flooded (90%) mountain valleys, mountain ranges. The parent material consists of alluvium derived from diorite. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is high. Shrink-swell potential is low. This soil is frequently flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 8 percent. Nonirrigated land capability classification is 3w. This soil meets hydric criteria. LuB - Luquillo-El Verde complex, 0 to 5 percent slopes, occasionally flooded

Luquillo (50%) The Luquillo component makes up 60 percent of the map unit. Slopes are 0 to 5 percent. This component is on terraces and flood plains along large streams and rivers. The parent material is unconsolidated Quaternary-terrace- and bouldery alluvial deposits that weathered from material of sandstone origin. Depth to a root restrictive layer, abrupt textural change, is 40 to 59 inches. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is occasionally flooded. It is not ponded. A seasonal zone of water saturation is at 8 inches during May, June, July, August, September, October, and November. Organic matter content in the surface horizon is about 8 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. El Verde (40%) The El Verde component makes up 40 percent of the map unit. Slopes are 0 to 4 percent. The parent material is unconsolidated Quaternary-terrace and bouldery alluvial deposits that weathered from material of sandstone origin. Depth to a root restrictive layer, abrupt textural change, is 11 to 20 inches. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is occasionally flooded. It is not ponded. A seasonal zone of water saturation is at 18 inches during May, June, July, August, September, October, and November. Organic matter content in the surface horizon is about 8 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. PaG - Palm-Yunque complex, 40 to 90 percent slopes, extremely stony

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Map Unit/Component Discussion Palm (70%) The Palm component makes up 70 percent of the map unit. Slopes are 40 to 90 percent. This component is on mountain ranges, mountain slopes. The parent material consists of colluvium derived from volcanic rock and/or residuum weathered from volcanic rock. Depth to a root restrictive layer, bedrock, lithic, is 38 to 39 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 23 percent. Nonirrigated land capability classification is 7w. This soil meets hydric criteria. Yunque, extremely stony The Yunque component makes up 25 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, (20%) mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer inches, plinthite. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is high. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 7w. This soil does not meet hydric criteria. PcE - Picacho-Ciales complex, 5 to 40 percent slopes

Picacho (60%) To be added.

Ciales (30%) The Ciales component makes up 30 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountain slopes, mountain ranges. The parent material consists of colluvium derived from diorite and/or residuum weathered from diorite. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, and December. Organic matter content in the surface horizon is about 28 percent. Nonirrigated land capability classification is 6w. This soil meets hydric criteria. PdF - Pandura-Very stony land complex, 40 to 60 percent slopes Pandura (70%) The Pandura component makes up 70 percent of the map unit. Slopes are 40 to 60 percent. This component is on mountain slopes on mountain ranges. The parent material consists of weathered materials. Depth to a root restrictive layer, bedrock, paralithic, is 12 to 20 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is high. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 2 percent. Nonirrigated land capability classification is 7e. This soil does not meet hydric criteria. Very stony land (30%) Generated brief soil descriptions are created for major soil components. The Very stony land is a miscellaneous area.

PiE - Picacho-Utuado complex, 5 to 40 percent slopes

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Map Unit/Component Discussion Picacho (50%) The Picacho component makes up 60 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountains, mountain ranges. The parent material consists of colluvium derived from diorite and/or residuum weathered from diorite. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 6w. This soil meets hydric criteria. Utuado (35%) The Utuado component makes up 35 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountains, mountain ranges. The parent material consists of colluvium derived from diorite and/or residuum weathered from diorite. Depth to a root restrictive layer inches, bedrock, paralithic. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is moderately high. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. PiG - Picacho-Utuado complex, 40 to 90 percent slope Picacho (60%) The Picacho component makes up 60 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountains, mountain ranges. The parent material consists of colluvium derived from diorite and/or residuum weathered from diorite. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during January, February, March, April, May, June, July, August, September, October, November, and December. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 6w. This soil meets hydric criteria. Utuado (35%) The Utuado component makes up 35 percent of the map unit. Slopes are 5 to 40 percent. This component is on mountains, mountain ranges. The parent material consists of colluvium derived from diorite and/or residuum weathered from diorite. Depth to a root restrictive layer inches, bedrock, paralithic. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is moderately high. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. PrF - Prieto very cobbly clay loam, 20 to 60 percent slopes

Prieto (90%) The Prieto component makes up 90 percent of the map unit. Slopes are 20 to 60 percent. This component is on concave areas of mountain coves, side slopes, and drainageways. These soils formed in residuum and colluvium that weathered from andesitic to basaltic, marine-deposited, metamorphosed volcanic mudstone of the Fajardo Formation. Depth to a root restrictive layer, bedrock, paralithic, is 30 to 39 inches. The natural drainage class is poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is low. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A

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Map Unit/Component Discussion seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 14 percent. Nonirrigated land capability classification is 7w. This soil meets hydric criteria. SoE - Sonadora-Caguabo complex, 20 to 40 percent slopes

Sonadora (70%) The Sonadora component makes up 60 percent of the map unit. Slopes are 20 to 40 percent. This component is on side slopes of lower hills and on footslopes of strongly dissected uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, paralithic, is 20 to 40 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink- swell potential is moderate. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 5 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. Caguabo (15%) The Caguabo component makes up 90 percent of the map unit. Slopes are 20 to 40 percent. This component is on ridgetops, ledges, and side slopes of strongly dissected uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, lithic, is 10 to 20 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 4 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. SoG - Sonadora-Caguabo complex, 40 to 90 percent slopes Sonadora (70%) The Sonadora component makes up 70 percent of the map unit. Slopes are 20 to 40 percent. This component is on side slopes of lower hills and on footslopes of strongly dissected uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, paralithic, is 20 to 40 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink- swell potential is moderate. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 5 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. Caguabo (15%) The Caguabo component makes up 90 percent of the map unit. Slopes are 20 to 40 percent. This component is on ridgetops, ledges, and side slopes of strongly dissected uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, lithic, is 10 to 20 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is very low. Shrink-swell potential is low. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 4 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. YnG - Yunque cobbly clay,40 to 90 percent slopes, extremely stony

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Map Unit/Component Discussion Yunque, extremely stony The Yunque component makes up 25 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, (75%) mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer inches, plinthite. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is high. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 7w. This soil does not meet hydric criteria. YqE - Yunque-Los Guineos-Moteado complex, 5 to 40 percent slopes Yunque, extremely stony The Yunque component makes up 25 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, (50%) mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer inches, plinthite. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is high. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 7w. This soil does not meet hydric criteria. Los Guineos (25%) The Los Guineos component makes up 25 percent of the map unit. Slopes are 5 to 40 percent. This component is on ridges, hillslopes, mountain slopes. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 21 percent. Nonirrigated land capability classification is 7e. This soil does not meet hydric criteria. Moteado (20%) Generated brief soil descriptions are created for major components. The Moteado soil is a minor component. YuF - Yunque-Moteado complex, 20 to 60 percent slopes Yunque (50%) The Yunque component makes up 25 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, mountain ranges. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer inches, plinthite. The natural drainage class is moderately well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is high. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 0 inches during June, July, August, September, and October. Organic matter content in the surface horizon is about 22 percent. Nonirrigated land capability classification is 7w. This soil does not meet hydric criteria. Moteado (30%) Generated brief soil descriptions are created for major components. The Moteado soil is a minor component. ZaG - Zarzal very cobbly clay, 40 to 90 percent slopes Zarzal, very cobbly clay The Zarzal, very cobbly clay component makes up 80 percent of the map unit. Slopes are 40 to 90 percent. This component is on (80% mountain ranges on uplands, mountain slopes on uplands. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 7 percent. Nonirrigated land capability classification is 7s. This soil

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Map Unit/Component Discussion does not meet hydric criteria. ZcF - Zarzal-Cristal complex, 20 to 60 percent slopes Zarzal (50%) The Zarzal component makes up 50 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain ranges on uplands, mountain slopes on uplands. The parent material consists of residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. Cristal (25%) The Cristal component makes up 25 percent of the map unit. Slopes are 5 to 40 percent. This component is on in concave positions in mountain coves and on lower side slopes of volcanic hills and mountains. The parent material consists of clayey colluvium derived from volcanic rock over silty and clayey residuum weathered from volcanic rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is high. Shrink-swell potential is low. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 15 inches during April, May, June, July, August, September, October, and November. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria. Sonadora (10%) The Sonadora component makes up 10 percent of the map unit. Slopes are 20 to 60 percent. This component is on low hills, mountain slopes on uplands. The parent material consists of Hato Puerco Formation residuum weathered from mudstone. Depth to a root restrictive layer, bedrock, paralithic, is 20 to 40 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is moderately low. Available water to a depth of 60 inches is low. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. There is no zone of water saturation within a depth of 72 inches. Organic matter content in the surface horizon is about 5 percent. Nonirrigated land capability classification is 7s. This soil does not meet hydric criteria. Humatas (8%) The Humatas component makes up 8 percent of the map unit. Slopes are 20 to 60 percent. This component is on mountain slopes, ridges, uplands. The parent material consists of residuum weathered from igneous rock. Depth to a root restrictive layer is greater than 60 inches. The natural drainage class is well drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is high. Shrink-swell potential is moderate. This soil is not flooded. It is not ponded. A seasonal zone of water saturation is at 65 inches during May, June, July, August, September, and October. Organic matter content in the surface horizon is about 80 percent. Nonirrigated land capability classification is 4e. This soil does not meet hydric criteria. Luquillo (7%) The Luquillo component makes up 7 percent of the map unit. Slopes are 0 to 5 percent. This component is on flood plains, alluvial plains. The parent material consists of Unconsolidated quaternary terrace and/or Bouldery alluvium derived from volcanic sandstone. Depth to a root restrictive layer, abrupt textural change, is 40 to 59 inches. The natural drainage class is somewhat poorly drained. Water movement in the most restrictive layer is low. Available water to a depth of 60 inches is moderate. Shrink- swell potential is moderate. This soil is occasionally flooded. It is not ponded. A seasonal zone of water saturation is at 8 inches during May, June, July, August, September, October, and November. Organic matter content in the surface horizon is about 8 percent. Nonirrigated land capability classification is 6w. This soil does not meet hydric criteria.

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Appendix D. Ecosystems Description (Characterization)

Mature Tabonuco Montane Wet Forest Average annual rainfall: 2,000−4,000 mm (78–156 inches) (Ewell and Whitmore 1973) Average annual temperature: 23.9 °C (Ewell and Whitmore 1973) Altitude range: 150–600 m (492–1,968 feet) (Weaver 2012) Canopy description (structure): Epiphytic ferns, bromeliads and orchids form a dark complete canopy 20 m (66 feet) (Ewell and Whitmore 1973). Four level canopy (Miller and Lugo 2009). Abundance of lianas (i.e., woody vines) dominates. Most abundant lianas are Marcgravia spp., Rourea surinamensis, Schlegia brachyata and Paullinia pinnata (Harris et al. 2012). Number of species present (species diversity): More than 150 species of trees (Ewell and Whitmore 1973) 10 most common species (in order of abundance): Dacryodes excelsa, Prestoea montana (Sierra Palm), Sloanea berteroana, Cordia borinquensis, Manilkara bidentata, Schefflera morototoni, Cecropia schreberiana, Micropholis garciinifolia, Henriettea squamulosa, Quararibea turbinate (Weaver 2012) Species density per hectare: 50 (Weaver 1994) Soils: Non hydric soils Hydrology: Rio Mameyes; Rio Espiritu Santo, Rio Sabana

Mature Tabonuco Montane Rain Forest Average annual rainfall: 4,650 mm (183 inches) (Weaver 2012) Average annual temperature: 21.2 °C (Weaver 2012) Altitude range: 400–700 m (1,312–2,296 feet) (Weaver 2012) Canopy description (structure): Species reaching 30 m height with occasional diameters approaching 1.5 m. Four level canopy (Miller and Lugo 2009) Number of species present (species diversity): More than 150 species (Ewell and Whitmore 1973) 10 most common species (in order of abundance): Prestoea montana, Dacryodes excelsa, Sloanea berteroana, Cordia borinquensis, Manilkara bidentata, Schefflera morototoni, Cecropia schreberiana, Micropholis garciinifolia, Henriettea squamulosa, Quararibea turbinate (Weaver 2012) Species density per hectare: 50 (Weaver 1994) Soils: Both hydric soils: Picacho-Utuado complex; Picacho-Ciales complex Hydrology: Rio Fajardo; Rio Mameyes, Rio Sabana

383 El Yunque National Forest

Mature Sierra Palm Montane Wet Forest Average annual rainfall: 2,000–4,000 mm (78–156 inches) (Ewell and Whitmore 1973) Average annual temperature: 23.9 °C (Ewell and Whitmore 1973) Altitude range: 150–600 m (492–1,968 feet) (Weaver 2012) Canopy description (structure): d.b.h. = 4–30 cm with a height of 17 m. Palms are also found in riparian forest (palm floodplain forest) and on very steep slopes at low elevation. Single canopy (Miller and Lugo 2009) Number of species present (species diversity): 63 (Weaver 2012) 10 most common species (in order of abundance): Prestoea Montana, Cecropia schreberiana, Henriettea squamulosa, Tabebuia rigida, Cordia borinquensis, Eugenia borinquensis, Psychotria berteroana, Croton poecilanthus, Sapium laurocerasus, Clibadeum erosum Species density per hectare: 25–30 (Weaver 2012) Soils: Hydric: Palm Yunque complex Hydrology: Rio Icacos, Rio Espiritu Santo, Quebrada Soñadora

Planted/Secondary Montane Wet Forest Average annual rainfall: 3,670mm (144 inches) (Weaver 2012) Average annual temperature: 24.6 °C (Weaver 2012) Altitude range: 180–600 m (492–1,968 feet) (data based on the permanent and temporary tree plots) (Weaver 2012) Canopy description (structure): Various timber species and secondary native species that have entered as secondary succession species; plantations of mainly mahogany (Ewell and Whitmore 1973) Number of species present (species diversity): Tree permanent and temporary monitoring plots show 39 species in Rio Grande, Sabana 4 and Sabana 8 stations that are within this life zone (Weaver 2012) 10 most common species (in order of abundance): Swietenia macrophylla, Swietenia mahagoni, Cedreal odorata, Cordia alliodora, Casuarina equisetifolia, Montezuma speciossissima, Petitia dominguensis, Tabebula heteryophillia, Tectona grandis, Dalbergia sissoo (Weaver 2012) Species density per hectare: Tree permanent and temporary monitoring plots show 14 species in 8.32 ha within Rio Grande station, 8 species in a 8.48 ha within Sabana 4 station and 17 species in a 8.56 ha within Sabana 8 station. All of these stations are within this life zone (Weaver 2012) Soils: Most Non hydric; Small part of Prieto very cobbly clay loam Hydrology: Rios: Icacos, Quebrada Soñadora, Sabana, Mameyes, Espíritu Santo

Mature Palo Colorado Montane Rain Cloud Forest Average annual rainfall: 4,533 mm (178 feet) (Weaver 2012) Average annual temperature: 18.6 °C (Weaver 2012)

384 Forest Plan Assessment

Altitude range: 600–900 m (1,968–2,952 feet) (Weaver 2012) Canopy description (structure): Similar to the Mature Palo Colorado montane wet cloud forest, but with abundance of epiphytes, epiphyllae, palms, tree ferns (Ewell and Whitmore 1973). Three level canopy: canopy, sub canopy and understory (Miller and Lugo 2009) Number of species present (species diversity): 88 (Weaver 2012) 10 most common species (in order of abundance): Cyrilla racemiflora, Ocotea spathulata Mez. (nemocli), Microphoiis chrysophylloides Pierre (caimitillo), and M. garciniifolia Pierre (caimitillo Verde) (Ewell and Whitmore 1973). Also, Prestoea montana, Tabebuia rigida, Micropholis guyanensis, Cordia borinquensis, Croton poecilanthus. Species density per hectare: 35–40 (Weaver 2012) Soils: Hydric soils: Guayabota Yunque complex; Yunque-Los Guineos moteado; Yunque moteado complex. Hydrology: Rio Mameyes, Rio Fajardo

Mature Palo Colorado Montane Wet Cloud Forest Average annual rainfall: 4,190 mm (164 inches) (Weaver 2012) Average annual temperature: 18.4 °C (Weaver 2012) Altitude range: 700–1,000 m (2,296–3,280 feet) (Ewel and Whitmore 1973) Canopy description (structure): reaches 15 m; characterized by open-crowned trees, many of which have their leaves grouped toward the ends of the branches, and whose crowns, when seen in profile, resemble somewhat-flattened bouquets. The leaves tend to be coriaceous and dark, giving the canopy a brownish or even reddish cast (Ewell and Whitmore 1973). Three level canopy: canopy, sub canopy and understory (Miller and Lugo 2009). Number of species present (species diversity): 53 species found (Ewell and Whitmore 1973) 10 most common species (in order of abundance): Cyrilla racemiflora, Ocotea spathulata Mez. (nemocli), Microphoiis chrysophylloides Pierre (caimitillo), and M. garcinicrefolia Pierre (caimitillo Verde) (Ewell and Whitmore 1973). Also, Prestoea montana, Tabebuia rigida, Micropholis guyanensis, Cordia borinquensis, Croton poecilanthus. Species density per hectare: 35–40 (Weaver 2012) Soils: Hydric soils: Picacho-Ciales complex; Picacho-Utuado complex; Yunque-Los Guineos moteado; Yunque moteado; Palm Yunque complex Hydrology: Rio Cubuy, Espiritu Santo, Icacos y Quebrada Soñadora

Mature Sierra Palm Montane Rain Cloud Forest Average annual rainfall: 4,533 mm (178 inches) (Miller and Lugo 2009) Average annual temperature: 18.6 °C (Miller and Lugo 2009) Altitude range: 600–900 m (1,968–2,952) (Weaver 2012)

385 El Yunque National Forest

Canopy description (structure): Similar to the Mature Palo Colorado montane wet cloud forest, but with abundance of epiphytes, epiphyllae, palms, tree ferns (Ewell and Whitmore 1973). Single canopy (Miller and Lugo 2009). Number of species present (species diversity): 88 (Weaver 2012) 10 most common species (in order of abundance): Prestoea montana, Cyrilla racemiflora, Ocotea spathulata Mez. (nemocli), Microphoiis chrysophylloides Pierre (caimitillo), and M. garcinicrefolia Pierre (caimitillo Verde), Tabebuia rigida, Micropholis guyanensis, Cordia borinquensis, Croton poecilanthus. Species density per hectare: 35–40 (Weaver 2012) Soils: Hydric soils: Picacho-Utuado complex; Picacho-Ciales complex; Palm yunque complex Hydrology: Rios Mameyes y Fajardo

Mature Sierra Palm Montane Wet Cloud Forest Average annual rainfall: 4,190 mm (164 inches) (Weaver 2012) Average annual temperature: 18.4 °C (Weaver 2012) Altitude range: 700–1000 m (2,296–3,280 feet) (Weaver 2012) Canopy description (structure): reaches 15 m; large stands of sierra palm extend up to the mountain peaks on steep slopes and ravines adjacent to elfin cloud forests. Single canopy (Miller and Lugo 2009) Number of species present (species diversity): 53 species found (Ewell and Whitmore 1973) 10 most common species (in order of abundance): Prestoea montana, Cyrilla racemiflora, Ocotea spathulata Mez. (nemocli), Microphoiis chrysophylloides Pierre (caimitillo), and M. garcinicrefolia Pierre (caimitillo Verde) (Ewell and Whitmore 1973) Tabebuia rigida, Micropholis guyanensis, Cordia borinquensis, Croton poecilanthus. Species density per hectare: 25–30 (Weaver 2012) Soils: Both hydric soils: Palm Yunque complex; Yunque-Los guineas moteado Hydrology: Ríos Icacos, Cubuy Espíritu Santo y Quebrada Soñadora

Mature Elfin Woodland Montane Rain Cloud Forest Average annual rainfall: 4,345 mm (171 inches) (Weaver 2012) Average annual temperature: 18.6 °C (Weaver 2012) Altitude range: 900–1,025 m (2,952–3,362 feet) (Weaver 2012) Canopy description (structure): Abundance of Epiphytes. Typical d.b.h. 4–15 cm. Canopy height 3–5 m; 80 t/ha of woody biomass; leaf biomass of 2.9 t/ha; soil organic matter of 550 t/ha; loose litter of 4.34 t/ha; total aboveground net primary productivity 3.70 r/ha/yr (Weaver 1994). Single canopy (Miller and Lugo 2009) Number of species present (species diversity): 54 (Weaver 2012) 10 most common species (in order of abundance): Tabebuia rigida, Ocotea spathulata, Eugenia borinquensis, Cyanthea bryophila, Micropholis garciniifolia,

386 Forest Plan Assessment

Henrietta squamulosa, Prestoea Montana, Clusia clusioides, Calyptranthes krugi, Haenianthus salicifolius (Weaver 2012) Species density per hectare: 15–20 (Weaver 2012) Soils: Hydric: Dwarf-El Duque complex Hydrology: Quebrada Soñadora, Rio Fajardo, Rio Mameyes

Mature Elfin Woodland Montane Wet Cloud Forest Average annual rainfall: 4,345mm (171 feet) (Weaver 2012) Average annual temperature: 18.6 °C (Weaver 2012) Altitude range: 900–1,025 m (2,952–3,362 feet) (Weaver 2012) Canopy description (structure): gnarled trees less than seven meters tall, high basal area, small diameters, a large number of stems per unit area, and extremely slow growth rates. All trees are evergreen and sclerophyllous here, and the leaves tend to be grouped near the ends of the branches (Ewel and Whitmore 1973). Single canopy (Miller and Lugo 2009) Number of species present (species diversity): 54 (Weaver 2012) 10 most common species (in order of abundance): Tabebuia rigida, Ocotea spathulata, Eugenia borinquensis, Cyanthea bryophila, Micropholis garciniifolia, Henrietta squamulosa, Prestoea montana, Clusia clusioides, Calyptranthes krugi, Haenianthus salicifolius (Weaver 2012) Species density per hectare: 15–20 (Weaver 2012) Soils: Hydric: Dwarf-El Duque complex Hydrology: Rios Cubuy, Icacos, Espíritu Santo

Planted/Secondary Submontane Moist Forest Average annual rainfall: 2,000 to 2,200 millimeters (78 to 86 inches) (Miller and Lugo 2009) Average annual temperature: 18–24 ºC (Miller and Lugo 2009) Altitude range: 350–550 m (1,148–1,804 feet) (Weaver 2012) Canopy description (structure): Various timber species and secondary native species that have entered as secondary succession species. This and other scattered remnants of this zonal association are characterized by trees up to 20 meters (66 feet) tall, with rounded crowns, like those of the mango tree. Number of species present (species diversity): Cubuy tree station within this life zone shows 29 species (Weaver 2012) 10 most common species (in order of abundance): Swietenia macrophylla, Swietenia mahagoni, Cedreal odorata, Cordia alliodora, Casuarina equisetifolia, Montezuma speciossissima, Petitia dominguensis, Tabebula heteryophillia, Tectona grandis, Dalbergia sissoo (Weaver 2012) Species density per hectare: Cubuy tree station within this lifezone shows 29 species on 8.00 ha (Weaver 2012)

387 El Yunque National Forest

Soils: Non hydric Hydrology: Río Canovanillas

Riparian Montane Rain and Wet Forest Average annual rainfall: No data available Average annual temperature: 18–24 °C (Harris et al. 2012) Altitude range: <600 m (<1,968 feet) Canopy description (structure): Upper canopy is 25–30 m in height and a second canopy is found at 15 m with abundant epiphytes and woody vines (Harris et al. 2012). Number of species present (species diversity): Approximately 29 species (Harris et al. 2012) 10 most common species (in order of abundance): Pterocarpus officinalis, Casearia arborea, Cordia borinquensis, Inga laurina, Manilkara bidentata, Nephelea portoricensis, Prestoea montana (Harris et al. 2012) Species density per hectare: No data available Soils: Non hydric soils Hydrology: Rio Mameyes; Rio Espiritu Santo, Rio Sabana

Riparian Montane Rain and Wet Cloud Forest Average annual rainfall: no data available Average annual temperature: 18–24 °C (Harris et al. 2012) Altitude range: >600 m (>1,968 feet) Canopy description (structure): no data available Number of species present (species diversity): Approximately 27 species based on Riparian palm brake tree monitoring station (Weaver 2012) 10 most common species (in order of abundance): no data available Species density per hectare: Approximately 27 species in 0.25 ha based on Riparian palm brake tree monitoring station (Weaver 2012) Soils: Hydric soils: Guayabota Yunque complex; Yunque-Los Guineos moteado; Yunque moteado complex, Picacho-Ciales complex; Picacho-Utuado complex; Yunque-Los Guineos moteado; Yunque moteado; Palm Yunque complex Hydrology: Ríos Icacos, Cubuy, Espíritu Santo y Quebrada Soñadora, Rio Mameyes, Rio Fajardo

Riparian Submontane Moist Forest Average annual rainfall: no data available Average annual temperature: 18–24 °C (Harris et al. 2012) Altitude range: No data available Canopy description (structure): No data available

388 Forest Plan Assessment

Number of species present (species diversity): No data available 10 most common species (in order of abundance): No data available Species density per hectare: No data available Soils: Non hydric soils Hydrology: Río Canovanillas

Literature Cited in Appendix D 1. Ewel and Whitmore. 1973. Puerto Rico Ecological Lifezones.

2. Peter L. Weaver. 2012. The Luquillo Mountains: Forest Resources and Their History.

3. Peter L. Weaver. 1994. Baño de Oro Natural Area, Luquillo Mouintains.

4. Harris et al. 2012. Luquillo Experimental Forest: Research History and Opportunities.

5. Gary Miller and Ariel Lugo. 2009. Guide to the Ecological Systems of Puerto Rico.

6. Julian Evans John Turnbull. 2004. Plantation Forestry in the Tropics, Third Edition.

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Appendix E: EYNF Forest Type Structural Characteristics and Species Composition Total or Mean Total Standard Standard Number Mean Standard Minimum Maximum Number Basal Deviation Mean Total Deviation of Tree Tree Deviation Tree Tree of Plots Area for Basal Density for Species Height for Tree Height Height Forest Type Sampled (m2/ha) Area (stems/ha) Density Recorded (m) Height (m) (m) Mature Tabonuco Montane 1 49.82 N/A 5,239.45 N/A 17 9.70 5.19 2.13 21.95 Rain Forest Mature Tabonuco Montane 1 32.80 N/A 1,364.86 N/A 15 11.04 4.23 3.35 21.34 Wet Forest Mature Sierra Palm Montane 1 29.24 N/A 1,075.67 N/A 13 10.25 3.41 3.05 15.55 Wet Forest Plantation/Secondary 2 57.26 5.97 4,639.40 439.61 26 10.26 4.42 2.44 25.00 Montane Wet Forest Mature Palo Colorado 1 31.16 N/A 3,535.29 N/A 10 7.65 4.47 2.44 23.17 Montane Rain Cloud Forest Mature Palo Colorado 3 38.37 6.42 2,349.08 476.29 23 7.60 3.31 2.13 19.82 Montane Wet Cloud Forest Mature Sierra Palm Montane 2 24.90 21.74 2,821.59 102.03 11 6.87 2.72 1.52 13.11 Rain Cloud Forest Mature Elfin Woodland 2 26.86 19.12 2,261.53 851.30 9 7.11 2.98 1.52 15.24 Montane Rain Cloud Forest Mature Elfin Woodland 1 16.73 N/A 2,859.65 N/A 8 6.69 2.90 3.05 14.63 Montane Wet Cloud Forest Source: Humfredo Marcano Vega, Research Biologist, USDA Forest Service, Southern Research Station Forest Inventory and Analysis (FIA) Unit, International Institute of Tropical Forestry (IITF). Member of the FIA field crew: Luis Ortiz Lopez, Ivan Vicens Jimenez, Humfredo Marcano Vega. Collaborative effort for the Installation of EFR inventory plots in El Yunque National Forest: Eileen Helmer and Skip Van Bloem. Note: The FIA program revisits study plots in a 5-year cycle.

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Appendix F: Species Present in the Different Vegetation Types at EYNF Mature Tabonuco Montane Rain Forest

Relative Importance Genus Species Relative Density Dominance Value Alchornea latifolia 29.41 10.92 20.16 Cecropia schreberiana 6.24 20.80 13.52 Prestoea acuminata 10.06 15.91 12.98 Dacryodes excelsa 0.85 20.92 10.89 Psychotria berteriana 10.60 1.76 6.18 Ocotea leucoxylon 10.60 1.18 5.89 Inga laurina 7.35 3.64 5.50 Cordia borinquensis 7.07 3.52 5.29 Eugenia borinquensis 4.10 5.68 4.89 Guarea glabra 7.07 2.10 4.59 Hura crepitans 0.28 5.69 2.99 Henriettea squamulosum 4.10 1.56 2.83 Schefflera morototonii 1.13 3.19 2.16 Tetrazygia urbanii 0.28 1.17 0.73 Sloanea berteriana 0.28 0.76 0.52 Micropholis garciniifolia 0.28 0.73 0.51 Sapium laurocerasus 0.28 0.46 0.37

Mature Tabonuco Montane Wet Forest

Relative Importance Genus Species Relative Density Dominance Value Tetragastris balsamifera 41.79 8.46 25.12 Prestoea acuminata 29.90 19.92 24.91 Guarea guidonia 5.44 23.93 14.69 Cecropia schreberiana 4.36 12.90 8.63 Sloanea berteriana 2.18 11.63 6.90 Alchornea latifolia 2.18 6.51 4.35 Swietenia macrophylla 2.18 3.79 2.98 Haenianthus salicifolius 2.18 3.12 2.65 Andira inermis 2.18 2.21 2.19 Schefflera morototonii 2.18 1.27 1.72 Sapium laurocerasus 1.09 2.07 1.58 Ormosia krugii 1.09 1.58 1.34 Ocotea leucoxylon 1.09 1.19 1.14 Cordia sulcata 1.09 0.83 0.96 Cyathea arborea 1.09 0.60 0.84

393 El Yunque National Forest

Mature Elfin Woodland Montane Wet Cloud Forest

Relative Importance Genus Species Relative Density Dominance Value Micropholis garciniifolia 24.62 44.95 34.79 Henriettea squamulosum 32.37 5.23 18.80 Croton poecilanthus 13.99 12.73 13.36 Sloanea berteriana 19.42 6.51 12.97 Clusia rosea 1.56 17.98 9.77 Tree unknown 6.47 6.87 6.67 Cyrilla racemiflora 1.04 4.47 2.75 Prestoea acuminata 0.52 1.26 0.89

Mature Sierra Palm Montane Wet Forest

Relative Importance Genus Species Relative Density Dominance Value Prestoea acuminata 35.18 21.66 28.42 Schefflera morototonii 21.36 8.96 15.16 Inga vera 6.91 12.54 9.72 Guarea guidonia 4.15 14.93 9.54 Ocotea leucoxylon 17.21 1.28 9.25 Dacryodes excelsa 1.38 16.88 9.13 Homalium racemosum 2.76 8.43 5.60 Swietenia macrophylla 1.38 5.12 3.25 Cecropia schreberiana 2.76 2.42 2.59 Ficus citrifolia 1.38 3.77 2.58 Tetragastris balsamifera 2.76 1.60 2.18 Andira inermis 1.38 1.69 1.54 Cyathea arborea 1.38 0.72 1.05

394 Forest Plan Assessment

Planted/Secondary Montane Wet Forest

Relative Importance Genus Species Relative Density Dominance Value Tabebuia heterophylla 7.60 25.03 16.32 Swietenia macrophylla 5.04 25.47 15.25 Syzygium jambos 17.08 6.60 11.84 Prestoea acuminata 10.72 12.25 11.48 Faramea occidentalis 15.96 3.31 9.63 Guettarda valenzuelana 10.14 4.60 7.37 Andira inermis 6.79 6.68 6.73 Miconia prasina 5.99 1.24 3.61 Manilkara bidentata 2.32 3.52 2.92 Myrcia splendens 3.99 0.41 2.20 Roystonea borinquena 0.32 2.81 1.56 Myrcia deflexa 2.16 0.67 1.41 Sloanea berteriana 2.00 0.43 1.21 Homalium racemosum 0.16 2.20 1.18 Trichilia pallida 2.00 0.36 1.18 Casearia arborea 2.00 0.16 1.08 Cordia borinquensis 2.00 0.10 1.05 Meliosma herbertii 2.00 0.08 1.04 Alchornea latifolia 0.48 0.98 0.73 Schefflera morototonii 0.32 0.78 0.55 Ficus citrifolia 0.16 0.84 0.50 Cinnamomum elongatum 0.16 0.38 0.27 Inga laurina 0.16 0.35 0.26 Buchenavia tetraphylla 0.16 0.33 0.25 Ocotea leucoxylon 0.16 0.24 0.20 Pimenta racemosa 0.16 0.17 0.17

395 El Yunque National Forest

Mature Palo Colorado Montane Rain Cloud Forest

Relative Importance Genus Species Relative Density Dominance Value Cecropia schreberiana 39.41 36.32 37.86 Prestoea acuminata 11.35 26.00 18.68 Alchornea latifolia 15.71 5.95 10.83 Sloanea berteriana 0.42 17.36 8.89 Cordia borinquensis 10.89 2.52 6.71 Croton poecilanthus 5.24 5.06 5.15 Henriettea squamulosum 6.08 3.28 4.68 Byrsonima wadsworthii 5.24 0.68 2.96 Psychotria berteriana 5.24 0.47 2.85 Micropholis garciniifolia 1.39 1.39 1.39

Mature Palo Colorado Montane Wet Cloud Forest

Relative Importance Genus Species Relative Density Dominance Value Prestoea acuminata 43.81 50.12 46.97 Micropholis guyanensis 5.80 15.56 10.68 Cecropia schreberiana 3.48 5.58 4.53 Micropholis garciniifolia 2.11 6.04 4.08 Croton poecilanthus 3.48 3.62 3.55 Alchornea latifolia 5.26 1.70 3.48 Ditta myricoides 5.26 0.53 2.90 Henriettea macfadyenii 5.26 0.45 2.86 Miconia sp. 5.26 0.32 2.79 Unknown Tree unknown 3.69 1.89 2.79 Henriettea squamulosum 1.27 2.65 1.96 Eugenia borinquensis 2.84 0.77 1.81 Magnolia splendens 0.21 3.17 1.69 Ocotea spathulata 2.63 0.47 1.55 Citharexylum caudatum 2.63 0.39 1.51 Daphnopsis philippiana 2.63 0.29 1.46 Nectandra turbacensis 2.63 0.26 1.45 Tabebuia rigida 0.04 2.72 1.38 Haenianthus salicifolius 0.42 1.47 0.95 Byrsonima wadsworthii 0.42 0.80 0.61 Cordia borinquensis 0.42 0.46 0.44 Miconia tetrandra 0.21 0.48 0.35 Cyrilla racemiflora 0.21 0.24 0.22

396 Forest Plan Assessment

Mature Sierra Palm Montane Rain Cloud Forest

Relative Importance Genus Species Relative Density Dominance Value Prestoea acuminata 15.10 42.37 28.73 Henriettea squamulosum 19.96 16.30 18.13 Micropholis garciniifolia 14.18 7.52 10.85 Alsophila bryophila 13.12 4.35 8.74 Eugenia borinquensis 9.45 4.11 6.78 Tabebuia rigida 2.11 11.16 6.63 Cecropia schreberiana 3.16 9.44 6.30 Ocotea spathulata 8.75 1.21 4.98 Psychotria berteriana 8.75 0.61 4.68 Cordia borinquensis 4.73 1.29 3.01 Citharexylum caudatum 0.70 1.64 1.17

Mature Elfin Woodland Montane Rain Cloud Forest

Relative Importance Genus Species Relative Density Dominance Value Prestoea acuminata 28.08 49.19 38.63 Eugenia borinquensis 13.44 20.12 16.78 Alsophila bryophila 20.80 6.95 13.87 Micropholis garciniifolia 9.83 8.52 9.18 Tabebuia rigida 6.39 7.96 7.18 Ocotea spathulata 5.08 5.10 5.09 Psychotria berteriana 8.19 0.86 4.52 Henriettea squamulosum 4.09 1.09 2.59 Cordia borinquensis 4.09 0.21 2.15

Mature Sierra Palm Montane Wet Cloud Forest