World Catalog of Fossil Cantharidae
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See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/317166287 World catalog of fossil Cantharidae Article · May 2017 CITATIONS READS 2 362 1 author: Fabrizio Fanti 36 PUBLICATIONS 86 CITATIONS SEE PROFILE Some of the authors of this publication are also working on these related projects: Studies of the world fossil Soldier Beetles (Cantharidae) View project All content following this page was uploaded by Fabrizio Fanti on 26 May 2017. The user has requested enhancement of the downloaded file. Gruppo Umbro Mineralogico Paleontogico Special Issue World catalog of fossil Cantharidae N° 2/2017 Fossils & minerals index World Catalog of foSSIl CantharIdaE Fabrizio FANTI Pag 1 Cover: Cantharidae. Amber. Photo courtesy of Artur R. Michalski Editorial Boarding Chief: Boccali Roldano Balocchi Pamela Calzolari Prospero Famiani Federico Giorgi Maria Pia Nessi Enrico Partenzi Chiara Sensi Claudio Tassi Luca Scientific commitee: Bogni Giorgio Venturi Federico Biagini Publio Carlini Rossano Tipography : Tipolito Properzio Adresse : Via dei Carrettieri, 12 - 06081 Località: Santa Maria degli Angeli - ASSISI - PERUGIA Italia Authorization No. 17 of 2 November 2016 - Court of Perugia Fossils & minerals World Catalog of foSSIl CantharIdaE Fabrizio FANTI [email protected] abstract acknowledgements A catalogue and bibliography of the world fossil Cantharidae are I am grateful to: Susana Adamonis and Cecilia Soledad given. Rhagonycha germari (Heer, 1847) n. comb., Rhagonycha Cataldo (Buenos Aires University), Vitalii I. Alekseev, Clive tertiaria (Heer, 1847) n. comb. and Rhagonycha hesperus A. Auton, Gabriel Biffi (Brazil), Albertas Bitinas (Klaipėda (Wickham, 1914) n. comb. are transferred from Telephorus University - Lithuania), Guido Bonamie (Merendree - Belgium), Schaeffer, 1766 = Cantharis Linnaeus, 1758 to Rhagonycha Jan Bosselaers, Christopher J. Duffin (Natural History Museum, Eschscholtz, 1830. Atalantycha humata (Wickham, 1913) n. London), Scott A. Elias, Michael S. Engel, Yun Hsiao (Taiwan); comb. is transferred from Telephorus Schaeffer, 1766 =Cantharis Jin Zhenyu (Sun Yat-Sen University, Guangzhou - China), Linnaeus, 1758 to Atalantycha Kazantsev, 2005. Lycocerus guttula Sergey V. Kazantsev; Wieslaw Krzemiński (Museum PAN, (J. Zhang, 1989) n. comb. is transferred from Cantharis Linnaeus, Krakow), Ulrich Kotthoff (CeNak - Hamburg University), 1758 to Lycocerus Gorham, 1889. Curticantharis J. Zhang, 1989 Jan-Michael Lange (Senckenberg Research Institute), Carlo = Themus Motschulsky, 1858 n. syn. Themus capacis (J. Zhang, Maria Legittimo, Cosmin-Ovidiu Manci (Romania), Yûichi 1989) n. comb., Themus thermophilus (J. Zhang, 1989) n. comb., Okushima (Kurashiki Museum of Natural History), Weston Themus trapezialis (J. Zhang, Sun & X. Zhang, 1994) n. comb. Opitz, Maurizio Pavesi, Evgeny E. Perkovsky, George O. and Themus sp. (J. Zhang, 1989) n. comb. are transferred from Poinar Jr., Ren Dong and Wang Yongjie (College of Life Curticantharis J. Zhang, 1989 to Themus Motschulsky, 1858. Sciences, Capital Normal University - Beijing, China), Mónica Telephorus tertiarius oeningensis Heer, 1847 n. syn. and Telephorus Morayma Solórzano Kraemer, Francesco Vitali (Luxembourg), tertiarius radobojanus Heer, 1847 n. syn. are synonymized with the Biodiversity Heritage Library (archive.org), the George Rhagonycha tertiaria (Heer, 1847). Lectotype and paralectotype Peabody Library (Paul Espinosa), the Naturalis Biodiversity of Rhagonycha tertiaria (Heer, 1847) are designated. Malthacus Center - Netherlands Entomological Society, the Sutro Library deceptus (W. J. Brown, 1940) n. comb. is the correct combination - California State Library, and the Biblioteca Civica Gianfranco of Podabrus deceptus Brown = Dichelotarsus deceptus. A Contini of Domodossola for help in finding some papers. preliminary list of subfossil Cantharidae is also provided. And gratefully extend my thanks to: Roberto Antonio Furthermore the Coleoptera Lampyridae Lamprohiza fossilis Pantaleoni and Francesco Vitali for the councils in some (Beier, 1952) n. comb. is transferred from Phausis LeConte, 1851 systematic problems, and Anders L. Damgaard, Sieghard to Lamprohiza Motschulsky, 1853. Ellenberger, Artur R. Michalski, George O. Poinar for the courtesy of the photographs. Key words: Catalog, worldwide, Coleoptera Cantharidae, I wish to dedicate this article to the memory of Milton paleontology Sanderson (entomologist at Illinois Natural History Survey) and express my hearty thanks to his son Steven Sanderson for the English revision. 1.Introduction 2.Methods The family Cantharidae is quite frequently found in fossil record, but currently there are only old or All named fossil taxa of the family Cantharidae incomplete catalogs (Handlirsch 1906-1908; Spahr are listed according to their current classification, 1981a [bibliography], 1981b; Carpenter 1992), and and this is based on the articles of Ramsdale (2002), only recently, on some lists and websites, are cited Kazantsev & Brancucci (2007) and Fanti (2014). more species (Handlirsch 1906-1908; Spahr 1981a The classification for the species is in the systematic- [bibliography], 1981b; Carpenter 1992), or we find alphabetical order, and for the undescribed specimens various data for all fossil species (The Paleobiology also for the date of citation. For each species are Database). Also the world catalog (Delkeskamp indicated: 1- valid name and author(s) and date of 1939, 1977) does not mention these species, therefore description; 2- the original name combination and with this article updated on 31 December 2016, I am page(s) of description (excluded names in index); finding and listing all the fossil species, including 3- eventually incorrect spelling or unjustified all citations that I know (some may be missing), and emendations; 4- all bibliographic references provide a preliminary list of the subfossil (Holocenic) (although it is only mentioned the author or at least species (Table 2.); furthermore furnished an appendix is evident the reference to one or more species) with (Appendix 1.) with the etymology and type species for number of pages, but for these are excluded species the genera. 1 Fossils & minerals names in index; 5- type horizon; 6- type locality; 7- adpression. Lacustrine - large shale, diatomaceous preservation and lithology; 8- collection where is and tuffaceous shale, volcaniclastic. deposited the holotype and, eventually, the paratypes or COLLECTIONS. Museum of Princeton University, other specimens, with number accession; 9- comments No. 5984. and notes, particularly focused on sex, body length, COMMENTS AND NOTES. Body length: 7.50 coloration, relationship with extant or fossil species, mm. Very similar in form and size to the recent syninclusions and others deemed interesting news. Telephorus bilineatus (= Atalantycha bilineata (Say) (Wickham 1913b: 362), and for this, and for the original drawning with pronotum rounded 3.Systematic at sides and angles, second antennomer short and particularly the habitus short but stocky, family Cantharidae Imhoff, 1856 (1815) I tentatively transferred this species to genus Atalantycha Kazantsev. Also for Carpenter (1992: Subfamily Cantharinae Imhoff, 1856 (1815) 331), the assignment to genus Cantharis was doubt. The picture of the holotype is visible in Florissant tribe Cantharini Imhoff, 1856 (1815) Fossil Beds National Monument website at http:// planning.nps.gov/flfo/photos. genus Atalantycha Kazantsev, 2005 genus Cantharis Linnaeus, 1758 †Atalantycha humata (Wickham, 1913) [n. comb.] Telephorus humatus Wickham, 1913b: 360, 362, 366 (explanation Subgenus (Cantharis) Linnaeus, 1758 of plates), Plate XXXVIII. Fig. 2. (but indicated as Plate I. Fig. 2. in the text and explanation) [under family Lampyridae] 1 Lucas 1915: 135; International Catalogue of Scientific Literature †Cantharis (Cantharis) attavina (Heer, 1847)* 1916a: 95; Meunier 1920b: 159; Wickham 1920: 353 [as Cantharis Telephorus attavinus Heer, 1847: 149-150, 225, Taf. IV. Fig. humatus]; Handlirsch 1920-1921: 232; Wilson 1978: 21 (Table 1.); 15., Fig. 15. b. Carpenter 1992: 331 [as Cantharis]; Kirejtshuk & Ponomarenko Telephorus atavinus Pictet, 1854: 334 (incorrect spelling) 2009-2015 [as Cantharis humata]; Mitchell 2013 - EDNA The Telephorus atavinus Scudder, 1891: 587 (unjustified Fossil Insect Database; The Paleobiology Database [as Cantharis emendation) humatus]. Telephorus atavinus Piton & Théobald, 1936: 207 Telephorus atavinus Mitchell 2013 - EDNA The Fossil Insect TYPE HORIZON. Eocene: Chadronian (37.2-33.9 Database Ma), Florissant Formation. Cantharis atavina Kirejtshuk & Ponomarenko, 2009-2015 TYPE LOCALITY. U.S.A.: Colorado, Teller County, Giebel 1852: 650; Pictet 1854: 334; Giebel 1856a: 102, Florissant. 400 (in the “Tabellarische”); Giebel, 1856b: 186; Scudder PRESERVATION/LITHOLOGY. Compression/ 1876: 81; Goss 1878: 50 (reprint 1878: 328); Scudder 1885: 796; Scudder 1886: 76; Scudder 1887: 796 [under family Lampyridae]; Scudder 1891: 587 [under family Lampyridae]; Scudder 1900: 101 [under family Lampyridae]; Handlirsch 1906-1908: 740 [as Cantharis atavina]; Handlirsch 1920- 1921: 232; Piton & Théobald 1936: 207; Carpenter 1992: 331 [as Cantharis]; Kirejtshuk & Ponomarenko 2009-2015 [as Cantharis atavina]; Mitchell 2013 - EDNA The Fossil Insect Database; The Paleobiology Database. TYPE HORIZON. Early Miocene: Burdigalian (16-20 Ma). TYPE LOCALITY. Croatia: Radoboj. PRESERVATION/LITHOLOGY. Mold / impression/compression (adpression). Lagoonal/ restricted