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Bulletin 63 New Mexico Museum of Natural History & Science A Bulletin 63 New Mexico Museum of Natural History & Science A Division of the DEPARTMENT OF CULTURAL AFFAIRS ANALYSIS OF INTRASPECIFIC AND ONTOGENETIC VARIATION IN THE DENTITION OF COELOPHYSIS BAURI (LATE TRIASSIC), AND IMPLICATIONS FOR THE SYSTEMATICS OF ISOLATED THEROPOD TEETH by LISA G. BUCKLEY and PHILIP J. CURRIE Albuquerque, 2014 Bulletin 63 New Mexico Museum of Natural History & Science A Division of the DEPARTMENT OF CULTURAL AFFAIRS ANALYSIS OF INTRASPECIFIC AND ONTOGENETIC VARIATION IN THE DENTITION OF COELOPHYSIS BAURI (LATE TRIASSIC), AND IMPLICATIONS FOR THE SYSTEMATICS OF ISOLATED THEROPOD TEETH by LISA G. BUCKLEY and PHILIP J. CURRIE New Mexico Museum of Natural History & Science Albuquerque, 2014 STATE OF NEW MEXICO Department of Cultural Affairs Veronica Gonzales, Secretary NEW MEXICO MUSEUM OF NATURAL HISTORY AND SCIENCE Charles Walter, Executive Director BOARD OF TRUSTEES Susanna Martinez, Governor, State of New Mexico, ex officio Charles Walter, Executive Director, ex officio Gary Friedman, President Deborah Dixon Maya Elrick, Ph.D. Peter F. Gerity, Ph.D. Laurence Lattman, Ph.D. Morton Lieberman, Ph. D. Imogene Lindsay, Emerita Viola Martinez Marvin Moss John Montgomery, Ph.D. Jennifer Riordan Laura Smigielski-Garcia David Smoak Steve West Cover illustration: NMMNH P-42200, skull of Coelophysis bauri in left lateral view. Original Printing ISSN: 1524-4156 Available from the New Mexico Museum of Natural History and Science, 1801 Mountain Road NW, Albuquerque, NM 87104; Telephone (505) 841-2800; Fax (505) 841-2866; www.nmnaturalhistory.org NMMNH Bulletins online at: http://nmnaturalhistory.org/bulletins BULLETIN OF THE NEW MEXICO MUSEUM OF NATURAL HISTORY AND SCIENCE EDITORS Spencer G. Lucas New Mexico Museum of Natural History and Science, Albuquerque, NM, USA (NMMNHS) Robert Sullivan NMMNHS Lawrence H. Tanner Le Moyne College, Syracuse, NY, USA PRODUCTION EDITOR Asher J. Lichtig NMMNHS ASSOCIATE EDITORS Guillermo Alvarado Asociación Costarricense de Geotecnica, San José, Costa Rica Marco Avanzini Museo Tridentino di Scienze Naturali, Trento, Italy David Berman Carnegie Museum of Natural History, Pittsburgh, PA, USA Brent Breithaupt Laramie, WY, USA William DiMichele National Museum of Natural History, Washington, D.C., USA John R. Foster Museum of Western Colorado, Grand Junction, CO, USA Gerard Gierlinski Polish Geological Institute, Warsaw, Poland Jean Guex University of Lausanne, Lausanne, Switzerland Jerald D. Harris Dixie State College, St. George, UT, USA Andrew B. Heckert Appalachian State University, Boone, NC, USA Adrian P. Hunt Flying Heritage Collection, Everett, WA, USA Hendrik Klein Neumarkt, Germany Karl Krainer University of Innsbruck, Innsbruck, Austria Martin G. Lockley University of Colorado at Denver, Denver, CO, USA Claudia Marsicano Universidad de Buenos Aires, Buenos Aires, Argentina Gary S. Morgan NMMNHS Donald R. Prothero Occidental College, Los Angeles, CA, USA Silvio Renesto Università degli Studi dell’Insubria, Varese, Italy Joerg W. Schneider Technical University BergAkademie of Freiberg, Freiberg, Germany Jingeng Sha Nanjing Institute of Geology and Palaeontology, Nanjing, China Sebastian Voigt Urweltmuseum GEOSKOP/Burg Lichtenburg, Thallichtenberg, Germany Ralf Werneburg Naturhistorisches Museum Schloss Bertholdsburg, Schleusingen, Germany Richard S. White, Jr. International Wildlife Museum, Tucson, AZ, USA NEW MEXICO MUSEUM OF NATURAL HISTORY AND SCIENCE BULLETINS 26. Paleogene Mammals, 2004. edited by Spencer G. Lucas, Kate E. Zeigler, and Peter E. Kondrashov, 230 pp. 27. Late Triassic microvertebrates from the lower Chinle Group (Otischalkian-Adamanian: Carnian), 2004. by Andrew B. Heckert, 170 pp. 28. New Mexico’s Ice Ages, 2005. edited by Spencer G. Lucas, Gary S. Morgan, and Kate E. Zeigler, 300+ pp. 29. Vertebrate Paleontology in Arizona, 2005. edited by Andrew B. Heckert & Spencer G. Lucas, 210 pp. 30. The Nonmarine Permian, 2005. edited by Spencer G. Lucas & Kate E. Zeigler, 362 pp. 31. Permian of Central New Mexico, 2005. edited by Spencer G. Lucas, Kate E. Zeigler & Justin A Spielmann, 176 pp. 32. Genética y Mamíferos Mexicanos: Presente y Futuro, 2006. edited by Ella Vázquez-Domínguez and David J. Hafner, 73 pp. 33. Skeletal Impact of Disease, 2006. by Bruce M. Rothschild and Larry D. Martin, 226 pp. 34. America’s Antiquities: 100 Years of Managing Fossils on Federal Lands, 2006. edited by Spencer G. Lucas, Justin A. Spielmann, Patricia M. Hester, Jason P. Kenworthy and Vincent L. Santucci, 185 pp. 35. Late Cretaceous Vertebrates from the Western Interior, 2006. edited by Spencer G. Lucas and Robert M. Sullivan, 410 pp. 36. Paleontology and Geology of the Upper Jurassic Morrison Formation, 2006. edited by John R. Foster and Spencer G. Lucas, 249 pp. 37. The Triassic-Jurassic Terrestrial Transition, 2006. edited by Jerry D. Harris, Spencer G. Lucas, Justin A. Spielmann, Martin G. Lockley, Andrew R.C. Milner and James I. Kirkland, 607 pp. 38. Pennsylvanian-Permian Fusulinaceans of the Big Hatchet Mountains, New Mexico, 2006. by Garner L. Wilde, 331 pp. 39. Upper Aptian-Albian Bivalves of Texas and Sonora: Biostratigraphic, Paleoecologic and Biogeographic Implications, 2007. edited by Robert W. Scott, 39 pp. 40. Triassic of the American West, 2007. edited by Spencer G. Lucas and Justin A. Spielmann, 247 pp. 41. The Global Triassic, 2007. edited by Spencer G. Lucas and Justin A. Spielmann, 415 pp. 42. Cenozoic Vertebrate Tracks and Traces, 2007. edited by Spencer G. Lucas, Justin A. Spielmann and Martin G. Lockley, 330 pp. 43. The Late Triassic archosauromorph Trilophosaurus, 2008. by Justin A. Spielmann, Spencer G. Lucas, Larry F. Rinehart and Andrew B. Heckert, 177 pp. 44. Neogene Mammals, 2008. edited by Spencer G. Lucas, Gary S. Morgan, Justin A. Spielmann and Donald R. Prothero, 442 pp. 45. The Paleobiology of Coelophysis bauri (Cope) from the Upper Triassic (Apachean) Whitaker quarry, New Mexico, with detailed analysis of a single quarry block, 2009. by Larry F. Rinehart, Spencer G. Lucas, Andrew B. Heckert, Justin A. Spielmann and Matthew D. Celeskey, 260 pp. 46. The taxonomy and paleobiology of the Late Triassic (Carnian-Norian: Adamanian-Apachean) drepanosaurs (Diapsida: Archosauromorpha: Drepanosauromorpha, 2010. by Silvio Renesto, Justin A. Spielmann, Spencer G. Lucas and Giorgio Tarditi Spagnoli, 81 pp. 47. Ichnology of the Upper Triassic (Apachean) Redonda Formation, east-central New Mexico, 2010. by Spencer G. Lucas, Justin A. Spielmann, Hendrik Klein and Allan J Lerner, 75 pp. 48. New Smithian (Early Triassic) ammonoids from Crittenden Springs, Elko County, Nevada: Implications for taxonomy, biostratigraphy and biogeography, 2010. by James F. Jenks, Arnaud Brayard, Thomas Brühwiler and Hugo Bucher, 41 pp. 49. Carboniferous-Permian transition in Cañon del Cobre, northern New Mexico, 2010. edited by Spencer G. Lucas, Jörg W. Schneider and Justin A. Spielmann, 229 pp. 50. Review of the tetrapod ichnofauna of the Moenkopi Formation/Group (Early-Middle Triassic) of the American Southwest, 2010. by Hendrik Klein and Spencer G. Lucas, 67 pp. 51. Crocodyle tracks and traces, 2010. edited by Jesper Milàn, Spencer G. Lucas, Martin G. Lockley and Justin A. Spielmann, 244 pp. 52. Selachians from the Upper Cretaceous (Santonian) Hosta Tongue of the Point Lookout Sandstone, central New Mexico, 2011. by Jim Bourdon, Keith Wright, Spencer G. Lucas, Justin A. Spielmann and Randy Pence, 54 pp. 53. Fossil Record 3, 2011. edited by Robert M. Sullivan, Spencer G. Lucas and Justin A. Spielmann, 736 pp. 54. Ichnology of the Mississippian Mauch Chunk Formation, eastern Pennsylvania, 2012. by David L. Fillmore, Spencer G. Lucas and Edward L. Simpson, 136 pp. 55. Tetrapod fauna of the Upper Triassic Redonda Formation, east-central New Mexico: The characteristic assemblage of the Apachean land-vertebrate faunachron, 2012. by Justin A. Spielmann and Spencer G. Lucas, 119 pp. 56. Revision of the Lower Triassic tetrapod ichnofauna from Wióry, Holy Cross Mountains, Poland, 2012. by Hendrik Klein and Grzegorz Niedzwiedzki, 62 pp. 57. Vertebrate Coprolites, 2012. edited by Adrian P. Hunt, Jesper Milàn, Spencer G. Lucas and Justin A. Spielmann, 387 pp. 58. A new archaic basking shark (Lamniformes: Cetorhinidae) from the late Eocene of western Oregon, U.S.A., and description of the dentition, gill rakers and vertebrae of the recent basking shark Cetorhinus maximus (Gunnerus), 2013. by Bruce J. Welton, 48 pp. 59. The Carboniferous-Permian transition in central New Mexico, 2013. edited by Spencer G. Lucas, W. John Nelson, William A. DiMichele, Justin A. Spielmann, Karl Krainer, James E. Barrick, Scott Elrick and Sebastian Voigt, 389 pp. 60. The Carboniferous-Permian transition, 2013. edited by Spencer G. Lucas, William A. DiMichele, James E. Barrick, Joerg W. Schneider and Justin A. Spielmann, 465 pp. 61. The Triassic System: New Developments in Stratigraphy and Paleontology, 2013. edited by Lawrence H. Tanner, Justin A. Spielmann and Spencer G. Lucas, 612 pp. 62. Fossil Footprints of Western North America, 2014. edited by Martin G. Lockey and Spencer G. Lucas, 508 pp. Buckley and Currie, 2014, Variation in the Dentition of Coelophysis bauri. New Mexico Museum of Natural History and Science Bulletin 63. v TABLE OF CONTENTS Abstract 1 INTRODUCTION 1 Documentation of Theropod Tooth Morphology 1 Individual Variation in Theropod Teeth 2 Ontogenetic Variation in Theropod Teeth 2 Wrinkles, Ridges, and Tooth Crowns 2 RATIONALE FOR STUDY 2 BACKGROUND TO COELOPHYSIS BAURI 2 Taxonomy of the Ghost
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