A New Genus of the Family Nitidulidae (Coleoptera: Polyphaga) from Australia Новый Род Семейства Nitidulidae (Coleoptera: Polyphaga) Из Австралии

ZOOSYSTEMATICA ROSSICA, 19(2): 54–61 28 JULY 2011

A new genus of the family Nitidulidae (Coleoptera: Polyphaga) from Australia Новый род семейства Nitidulidae (Coleoptera: Polyphaga) из Австралии

A.G. KIREJTSHUK

А.Г. КИРЕЙЧУК

A.G. Kirejtshuk, Zoological Institute of the Russian Academy of Sciences, 1 Universitetskaya Emb., St. Petersburg 199034, Russia; Muséum National d’Histoire Naturelle, CP 50, Entomologie, 45 Rue Buffon, F-75005 Paris, France. E-mail: [email protected], [email protected]

Brittonema gen. nov. of the subfamily Cillaeinae is described. This genus includes two new species from Australia (Queensland), B. mandibulare sp. nov. and B. pygidiatum sp. nov. The new genus has a unique combination of characters, and also peculiar shape of pronotum and structure of genitalia making possible its recognition among other genera of the subfamily. В статье описан Brittonema gen. nov. из подсемейства Cillaeinae, включающий два новых вида из Австралии (Квинсленд): B. mandibulare gen. et sp. nov. и B. pygidiatum gen. et sp. nov. Новый род характеризуется уникальной комбинацией признаков, а также свое- образными формой переднегруди и строением гениталий, которые позволяют легко диа- гностировать его среди остальных родов подсемейства. Key words: Australia, Coleoptera, Nitidulidae, Cillaeinae, new genus, new species Ключевые слова: Австралия, Coleoptera, Nitidulidae, Cillaeinae, жесткокрылые, блестян- ки, новый род, новые виды

INTRODUCTION (2010) with the name ‘Cillaeinae gen. sp.’. Unfortunately the subfamily Cillaeinae in The recently elaborated generic sys- general still remains poorly known in com- tem of the family Nitidulidae (Kirejtshuk, parison with other groups of Nitidulidae. 2008) last two years was added by some The most comprehensive review of the gen- contributions. There have been published era of this subfamily was published only for descriptions of two genera in the tribe Niti- the Hawaiian fauna (Ewing, 2007), how- dulini: Neohebascus Cline, 2009 and Bolivi- ever the rest recent faunas of Cillaeinae (es- toxus Kirejtshuk, 2009, and also one genus from Meligethinae: Sebastiangethes Audi- pecially in the areas with tropical climate) sio, Kirk-Spriggs and Cline, 2008. Besides, need to be studied. Audisio et al. (2009) revised the classifica- The following acronyms are used for col- tion of the subfamily Meligethinae and pro- lections: ANIC, Australian National Insect posed 22 new genera. In the latter version of Collection, Department of Entomology, the subfamily classification all taxa treated Commonwealth Scientific and Industrial before as subgenera were elevated to the Research Organisation, Canberra; BMNH, generic level and some synonyms were re- Natural History Museum (formerly British established as valid genera. The new genus Museum of Natural History), London; QM, described herein was studied by the author Queensland Museum, Brisbane; ZIN, Zoo- many years ago. Last year the picture of one logical Institute of the Russian Academy of species of it was published in Jelìnek et al. Sciences, St. Petersburg.

RESULTS convex and widely rounded to subtruncate at apex in both sexes. Prosternal process Order COLEOPTERA flattened, not curved along procoxae and Suborder POLYPHAGA strongly widened at rounded apex. All pairs Superfamily CUCUJOIDEA of coxae narrowly separated. Mesoventrite medially flattened and almost not exca- Family NITIDULIDAE Latreille, 1802 vate, but at sides with distinct depressions Subfamily CILLAEINAE Kirejtshuk & for mesofemora. Metaventrite with a dis- Audisio in Kirejtshuk, 1986 tinct median suture (discrimen) along the Brittonema gen. nov. whole length but without submesocoxal lines. Hypopygidium very widely rounded Type species: Brittonema mandibulare to subtruncate at apex in both sexes. Legs gen. et sp. nov. rather narrow and of usual shape. Pro- Diagnosis. Body comparatively medi- tibiae very narrow and with rather short um-sized (6.0–15.5 mm), narrow and elon- spurs. Tarsi with rather widely lobed tar- gate, subflattened dorsally and ventrally. someres 1–3. Male anal sclerite rather long Dorsum rather coarsely punctured, smooth and with subacute apex. Aedeagus heavily to finely microreticulate; elytral punctura- sclerotised, moderately long, with a small tion more or less clearly arranged in longi- median excision at acute apex of penis tudinal rows; underside finely and densely trunk and with sharply acute apex of teg- punctured or with strongly reduced punc- men. Ovipositor heavily sclerotised, inner turation; body with very short pubescence. and outer lobes of gonocoxites well isolat- Head subflattened (either somewhat in- ed, short styli located far from acuminate clined anteriorly or convex medially) and apex of ovipositor. rather strongly narrowed at base, mod- Composition. The new genus includes erately projecting anteriorly, clearly de- the type species and B. pygidiatum gen. et pressed between antennal insertions and sp. nov. with anterior edge of frons truncate; anten- Comparison. This new genus can be nal grooves strongly convergent posteri- compared with the genera characterised by orly and with distinct inner edge; mentum the dorsally flattened body with pronotum very wide and subquadrangular; pregenal subquadrate or widened anteriorly (Cil- process at hypostomal sinus comparatively laeopeplus Sharp, 1908; Cillaeus Laporte, wide. Ultimate labial palpomere short and 1835; Ithyphenes Murray, 1864; Platynema transverse; ultimate maxillary palpomere Ritsema, 1885 and Tokocillaeus Kirejtshuk, long and subcylindrical. Antennae 11-seg- 2001). Indeed the pronotum clearly wid- mented bearing 3-segmented club without ened anteriorly is known only in Brittone- any evident trace of sexual dimorphism. ma gen. nov., however it can be supposed Pronotum somewhat widened anteriorly, that other species of this new genus still subflattened at disk and slightly sloping at remaining undescribed could have a sub- narrowly bordered sides, its anterior and quadrate pronotum or undescribed species posterior angles with somewhat rounded of other mentioned genera could be with vertex; its anterior edge shallowly emar- somewhat transformed pronotum. This new ginate or nearly straight and posterior one genus differs from Cillaeopeplus also in the nearly straight. Elytra with truncate apices shorter head, shorter elytra (less than twice remaining at least last three abdominal seg- as long as wide), widely dilated tarsomeres ments uncovered, with narrowly explanate 1–3 and more or less smoothed integument and bordered sides. Laterosternites of two of dorsum (at least head and pronotum), last abdominal segments moderately wide and in the elytra with longitudinal rows and widened posteriorly. Pygidium slightly of punctures but without distinct striae.

Besides, Brittonema gen. nov. differs also Diagnosis. This new species differs from from Cillaeus and Tokocillaeus in the much its another congener in the larger and much narrower base of head, median excision wider body, transverse head with truncate of labrum, markedly shorter elytra with (not emarginate) anterior edge, gentle out- seriate puncturation, pubescent dorsum; line of temples behind eyes, transverse (not and also from Ithyphenes and Platynema in V-shaped) depression between antennal in- the head narrowed at base, shorter elytra, sertions, more projecting and not subquad- convex or subflattened (not depressed) py- rangular male labrum, strong sexual di- gidium, antennal grooves distinctly conver- morphism in structure of mandibles, much gent posteriorly, narrowly separated coxae shorter tergites of abdomen uncovered by in all pairs, moderately lobed tarsomeres elytra, tergites uncovered by elytra with 1–3, pubescent dorsum (antennal grooves paramedian depressions, truncate apex of of Ithyphenes and Platynema are rather pygidium in both sexes, subacute apex of reduced and (sub) divergent and following anal sclerite far projecting behind pygidial closely the inner edge of eyes). Among the apex, wider mentum, longer ultimate labial mentioned genera, the genus Cillaeus looks palpomeres, subparallel tibiae, wider tarsi, like more archaic and maintaining the most strongly dentate tarsal claws, finer punctur- number of plesiomorphic characters. The ation on dorsum (except elytra) and double latter genus could be in any sense an initial rows of punctures between longitudinal el- one to other mentioned groups with the evated lines on elytra, and also in the struc- more restricted distribution and demons- ture of male genitalia. trating a greater or lesser specialization in Description of holotype (male). Length apomorphic characters. The genera Cillaeus with mandibles 15.3, width 4.0, height and Brittonema gen. nov. are characterised 1.3 mm. Elongate, subflattened dorsally and by the mesoventrite with the quite distinct slightly convex ventrally; yellowish red- lateral depressions for receiption of meso- dish, but base of head, apices of mandibles, femora, while these depressions are weak- scutellum, lateral and apical parts of elytra, ened in Ithyphenes and Platynema. unclear paramedian spots on tergites un- Etymology. The name of this genus is covered by elytra and femoro-tibial articu- formed from the name of outstanding Aus- lations more or less infuscate to dark brown; tralian coleopterist E.B. Britton and Greek body with a faint shine; covered with short ‘nema’ (thread), referring to the long and and subrecumbent brown greyish or yel- flexible body. Gender neuter. lowish hairs, which are about as long as Note. Bionomy of the species of this new distance between insertions; on elytra these genus remains unknown, however, their as- hairs intermixed with longitudinal rows of sociations with interstices on palm trunks somewhat longer hairs disposed between and at base of leaves can be supposed. double rows of punctures. Dorsum, except elytra, and abdominal Brittonema mandibulare gen. et sp. nov. ventrites with very small punctures (sub- (Figs 1–4, 8–11) equal or slightly larger than eye facets), interspaces between them on head about Holotype. Male; Australia, Queensland , ’6 m. a puncture diameter and on rest sclerites up Mt. Lewis Rd., N Q, 27.12.72, A. Walford- markedly greater, with extremely fine and Huggins‘, ’from palm‘, ’Brachypeplus sp. not in dense alutation. Elytra with double regu- B.M., E.B. Britton det. 74‘; ANIC. Paratypes. Ten males and females; same data lar longitudinal rows of large punctures as for holotype; BMNH, ANIC, ZIN. Males and (about three times as great as those on females; Australia, Queensland , ’Mt.Lewis, N other dorsal sclerites), interspaces between Qld, 8 Dec. 1974, A.Walford-Huggins‘, ’E. Gow- punctures less than half of puncture diam- ing-Scopes coll.’; BMNH, ZIN. eter and smoothed; between double rows of

rable in length and each about four fifth as long as scape, their elongate oval club about two ninths of total antennal length, almost twice as long as wide, antennomere 9 about as long as antennomeres 10 and 11 combined, antennomere 11 subacute at apex. Pronotum subquadrangular and rather widened anteriorly, subflattened at disk and gently sloping at extremely narrowly bordered sides, all corners rounded, sides and base finely bordered, anterior edge emarginate, sides and posterior edge more or less straight, lateral edge with nearly regular and very distinct but small notches. Scutellum subpentagonal to subhemicircular, about one and one third as wide as long. Elytra flattened at disk and steeply sloped at extremely narrowly explanate sides, apical outer angle widely rounded, posterior Figs 1–4. Brittonema mandibulare gen. et sp. nov. Teg- edge obliquely truncate and form- men, ventral view (1); penis trunk, dorsal view (2); anal ing a very stump sutural angle, leav- sclerite and spiculum gastrale, ventral view (3); ovipositor, ing three last abdominal segments ventral view (4). Scale bar: 1.0 mm. and half of previous one uncovered. Three last tergites slightly con- punctures a narrow elevated line disposed. vex in the middle and slightly depressed at Thoracic sterna with more or less obsolete sides; pygidium (ultimate tergite) subtri- puncturation and mostly finely alutaceous angular, about as long as wide and rounded to smoothed. to subtruncate at apex. Narrowly rounded Head (Fig. 10) transverse (markedly apex of anal sclerite well exposed dorsally shorter than distance between eyes) and and far projecting posteriorly. strongly narrowing behind extremely fine- Mentum (Fig. 9) rather large and trans- ly faceted eyes, convex at base and sloping verse, about 2.5 times as wide as long, sub- anteriorly, with a transverse depression be- quadrangular, widened anteriorly and with tween rather elevated antennal insertions, widely rounded anterior and posterior an- its anterior edge truncate, unbordered and gles; plane of mentum with two paramedian with rounded lateral angles, base without oval depressions. Lateral parts (lobes) of laterally projecting temples behind eyes. mentum widely cover comparatively wide Labrum entire (not divided into lobes, but hypostomal sinuses. Ligula with very wide with a short blackish line), medially some- lobes. Ultimate labial palpomere transverse, what projecting to truncate apex and ar- slightly wider than long and truncate at cuately widening to anterior edge of frons. apex. Ultimate maxillary palpomere sub- Mandibles with tridentate apices and with cylindrical and about four times as long as bidentate laterodorsal processes. Anten- thick. Antennal grooves S-shaped, strongly nae about 1.5 times as long as head width, convergent posteriorly and turned exter- antennomere 3 and antennomere 5 compa- nally behind eyes. Prosternum slightly con-

© 2011 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 20(1): 54–61 58 A.G. KIREJTSHUK. NEW NITIDULID GENUS FROM AUSTRALIA vex and with somewhat smoothly roof-liked tised, with the distal parts of inner lobes of process, not curved along coxae, strongly gonocoxites clearly isolated from the proxi- widened before widely rounded apex (which mal ones and very short styli located close is about 1.5 times as wide as antennal club). the middle of sides of these inner parts. The distance between mesocoxae 1.5 times Variations. Length with mandibles 11.5– and that between metacoxae nearly as great 15.5 mm. The colouration of paratypes vari- as that between procoxae. Mesoventrite es from nearly unicolorous yellowish body slightly medially depressed in distal half with only head base somewhat darker, scu- and with lateral distinct depressions for tellum and elytral apices to nearly unicolo- receipt of mesofemora. Metaventrite flat- rous brown body with lighter prescutellar tened, slightly longer than prosternum with parts of elytra and tarsi. Some variation is process, its anterior edge almost straight observed in puncturation and sculpture of and posterior edge between coxae angularly the integument and most variation has the excised. Submesocoxal and submetacoxal structure of male mandibles (particularly in lines unexpressed. Abdominal ventrite 1 the shape of apices and laterodorsal process about 1.5 times as long as ventrite 2, but of male mandibles). somewhat shorter than ventrite 3; ventrites Etymology. The new species is named for 4 and 5 longest and comparable in length; the large mandibles in males. hypopygidium truncate at apex. Epipleura gradually narrowed distally, strongly ele- Brittonema pygidiatum gen. et sp. nov. vated laterally and somewhat narrower (Figs 5–7, 12–14) than antennal club. Holotype. Male; Australia, Queensland, ’Mt. Legs moderately long and narrow, of Lewis Rd., N. Q, 3000’, 30 Oct. 1966, E. Britton‘; usual structure. Tibiae about as long as ANIC. antennal flagella and somewhat narrower Paratypes. Male; same data as for holotype; than antennal club, slightly curved; protibia ANIC. Australia, Queensland: male, ’Mt. Lewis, rounded at its apical outer angle; meso- and N. Q., 27.XII.1922, A. & M. Walford-Huggins, metatibiae slightly narrowing to rounded 7121‘ (ANIC); male, ’Russell R., at Bellenden, apex; spurs very small. Femora with more or Ker Landing, N.Q., 5 m., 24 Oct. – 9 Nov.1981, less usual outline and about twice as wide EARTHWATCH/QLD. MUSEUM, Malaise trap, rainforest‘, ’on loan from Queensland Muse- as tibiae. Tarsi about half as long as tibiae, um, Brisbane, 90.14‘ (QM); male, ’Laceys Creek, tarsomere 5 about as long as tarsomeres 1–3 QLD, 10 km SE, El Arish, 40 m, 23 June – 5 Aug. combined, protarsi very widely lobed and 1982, S. & J. Peck, SBP47‘, ’flight intercept trap, about 1.5 times as wide as protibiae, meso- rainforest’ (ZIN). and metatarsi somewhat narrower, claws Diagnosis. See the diagnosis of the previ- strongly toothed at base. ous species. Aedeagus (Figs 1–2) well sclerotised. Description of holotype (male). Length 6.7, Apex of tegmen moderately projecting and width 1.6, height 0.7 mm. Elongate, subflat- angularly acute. tened dorsally and slightly convex ventral- Female. Differs from the male in the sub- ly; yellowish reddish, antennal flagella and flattened head but with a distinct shallow particularly clubs, scutellum, distal third of depression between antennal insertions, elytra and median stripes on tergites uncov- and also with simple and transversely trun- ered by elytra more or less infuscate to dark cate labrum, simple mandibles with biden- brown; body shining; covered with short sub- tate apices, lack of exposed anal sclerite. erect brown greyish hairs, which are about as The apices of the female pygidium and hy- long as distance between insertions; on elytra popygidium, and antennae and tarsi are as these hairs intermixed with yellowish hairs subtruncate as the male ones. Ovipositor and also with longitudinal rows of hairs dis- (Fig. 4) is rather long and heavily sclero- posed between simple rows of punctures.

short blackish line), medially somewhat projecting to truncate apex and straight at sides, base with laterally projecting angular temples behind eyes. Mandibles with bidentate apices. Anten- nae about one and one third as long as head width, antennomere 3 markedly longer than antennomere 5 and almost as long as scape, their elongate oval club about two sevenths of total antennal length, somewhat more than twice as long as wide, with antennomeres 9–11 compared Figs 5–7. Brittonema pygidiatum gen. et sp. nov. Tegmen, in length and antennomere 11 ventral view (5); penis trunk, dorsal view (6); anal sclerite subacute at apex. Pronotum sub- and spiculum gastrale, ventral view (7). Scale bar: 1.0 mm. quadrangular and rather widened anteriorly, subflattened at disk Head, pronotum and scutellum with and gently sloping at extremely narrowly more or less distinct punctures about twice bordered sides, all corners rounded, sides as large as eye facets in diameter, interspac- and base finely bordered, anterior edge es between them about a puncture diameter emarginate, sides and posterior edge more and smooth to somewhat alutaceous. Elyt- or less straight, lateral edges with very dis- ra with single regular longitudinal rows of tinct but small notches. Elytra flattened at large transverse punctures (with transverse disk and not steeply sloped at moderately diameter about twice as great as those on explanate sides, apical outer angle widely head and pronotum), interspaces between rounded, posterior edge obliquely truncate punctures less than half of puncture di- and forming a very stump sutural angle, ameter and smoothed; between rows of leaving three last abdominal segments and punctures a narrow elevated line disposed. fourth of previous one uncovered. Three Dorsal and ventral abdominal sclerites with last tergites slightly and gently convex; punctures, subequal or slightly smaller than pygidium (ultimate tergite) subtriangular, those on head and pronotum and slightly about one and two thirds as long as wide shallower, interspaces between them some- and narrowly rounded to subacute at apex. what greater, with extremely fine and dense Widely rounded apex of anal sclerite slight- alutation. Prosternum with shallow and in- ly to scarcely exposed dorsally. distinct punctures with interspaces greater Mentum (Fig. 14) rather large and than puncture diameter and smoothly alu- transverse, about 2.5 times as wide as long, taceous. Metaventrite with fine punctura- subquadrangular slightly widened anteri- tion and mostly finely alutaceous. orly and with rounded anterior and distinct Head (Fig. 13) somewhat longer than posterior angles; plane of mentum without wide at eyes and strongly narrowing be- two paramedical depressions. Lateral parts hind extremely finely faceted eyes, some- (lobes) of mentum narrowly cover com- what convex along the middle, with V- paratively wide hypostomal sinuses. Ligula shaped depression between elevated anten- with moderately narrow lobes. Ultimate nal insertions, its anterior edge emarginate labial palpomere more than twice wider and with rounded lateral angles. Labrum than long and truncate at apex. Ultimate entire (not divided into lobes, but with a maxillary palpomere subcylindrical, about

© 2011 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 20(1): 54–61 60 A.G. KIREJTSHUK. NEW NITIDULID GENUS FROM AUSTRALIA four times as long as thick and narrow- ACKNOWLEDGEMENTS ing apically. Antennal grooves S-shaped, strongly convergent posteriorly and turned The study of the author would be impossible without assistance of many colleagues and scien- externally behind eyes. Prosternum slightly tific organizations supported him. Particularly, convex and with somewhat smoothly roof- the author greatly appreciates to M.V.L. Barclay liked process, somewhat curved along cox- and H. Mendel (BMNH), and also J.F. Lawrence ae, strongly widened before widely rounded (ANIC), A. Nel and T. Deuve (Muséum Natio- to subtruncate apex (which is slightly wider nal d’Histoire Naturelle in Paris) for providing than antennal club). The distance between specimens and many other things for study. Be- mesocoxae and that between metacoxae sides, the study was supported by the program nearly as great as that between procoxae. of the Presidium of the Russian Academy of Sci- Mesoventrite slightly medially depressed in ences ’Origin of the biosphere and evolution of distal half and with lateral distinct depres- geo-biological systems’ and Russian Foundation of Basic Research (grant 09-04-00789-а). The sions for receipt of mesofemora. Metaven- author sincerely thanks the Muséum National trite flattened, slightly longer than proster- d’Histoire Naturelle for providing him a possi- num with process, its anterior edge almost bility to work in this museum according to the straight and posterior edge between coxae programme of visiting professors. He is also very angularly excised. Submesocoxal and sub- grateful to J. Jelinek (Národní Muzeum, Prague) metacoxal lines unexpressed. Abdominal and S. Hisamatsu (Ehime University, Mat- ventrite 1 slightly longer than ventrite 2, suyama) for their valuable remarks on an earlier but twice shorter than each of ventrites 3 version of the manuscript. and 4; hypopygidium (ventrite 5) truncate at apex. Epipleura gradually narrowed dis- REFERENCES tally, strongly elevated laterally and somewhat narrower than antennal club. Audisio, P., Cline, A.R., De Biase A., Anto- nini, G., Mancini, E., Trizzino, M., Con- Legs moderately long and narrow, of stantini, L., Strika, S., Lamanna, F. & Cer- usual structure. Tibiae markedly shorter retti, P. 2009. Preliminary re-examination than antennal flagella and much narrower of genus-level taxonomy of the pollen beetle than antennal club (about two thirds as subfamily Meligethinae (Coleoptera: Niti- wide as antennal club), with almost straight dulidae). Acta Entomologica Musei Nationalis inner edge; protibiae somewhat rounded at Pragae. 49(2): 341–504. its apical outer angle; meso- and metatibiae Ewing, C. 2007. Phylogenetic analysis of the slightly narrowing to rounded apex; spur genera of endemic Hawaiian sap beetles (Co- very small. Femora with more or less usual leoptera: Nitidulidae) based on morphology outline and about twice as wide as tibiae. with redescription and key to the genera of endemic Hawaiian Nitidulidae. Zootaxa. Tarsi about two thirds as long as tibiae, 1427: 1–36. tarsomere 5 about as long as tarsomeres Jelìnek, J., Carlton, C., Cline, A. & Leschen, R. 1–3 combined, protarsi very widely lobed 2010. 10.26. Nitidulidae Latreille, 390–407. and about somewhat wider than protibiae, In: Beutel, R.G., Leschen, R.A.B. & Law- meso- and metatarsi somewhat narrower, rence, J.F. (Eds) Handbook of Zoology Volume claws simple. IV. Arthropoda: Insecta. Coleoptera, Volume 2: Aedeagus (Fig. 14) well sclerotised. Morphology and systematics (Elateroidea, Bos- Apex of tegmen strongly projecting and trichiformia, Cucujiformia partim): 390–407. very narrowly acute. Walter De Gruyter, Berlin. Female. Unknown. Kirejtshuk, A.G. 2008. A current generic classi- Etymology. The epithet of this new spe- fication of sap beetles (Coleoptera, Nitiduli- dae). Zoosystematica Rossica. 17(1): 107–122. cies refers to the long male pygidium of this new species. Received April 4, 2011 / Accepted June 5, 2011

Figs 8–14. Brittonema gen. n. 8–11, B. mandibulare gen. et sp. nov.; paratype, male of ‘8 Dec. 1974’ with length 15.3 mm (8, 9); paratype, male of ‘27.12.72’ with length 12.3 mm (10); paratype, female of ‘8 Dec. 1974’ with length 13.0 mm (11); 12–14, B. pygidiatum gen. et sp. nov.; paratype, male of ‘27.XII.1922’ with length 8.3 m, (12); paratype, male of ‘23 June – 5 Aug. 1982’ with length 7.6 mm (13, 14). Body, dorsal view (8, 12), mouthparts, ventral view (9), head, dorsal view (10, 13), head, ventral view (11, 14).

Figs 4–7. Libanochrus calvus gen. et sp. nov., holotype. Head and thorax, ventral view (4); head and pronotum, dorsal view (5); head, dorsal view (6): antennal club, dorsal view (7). Head width 2.6 mm.