Mackay's Horsetail.Pdf

Total Page:16

File Type:pdf, Size:1020Kb

Mackay's Horsetail.Pdf MACKAYS HORSETAIL (EQUISETUM TRACHYODON) Coastal Sand Dune Stems grow up to 100 cm high and be 2-6 mm diameter. Mackay's horsetail (Equisetum x variegatum) is a hybrid of Variegated horsetail and Dutch horsetail and is only found on a couple of sites in the UK. The stems of the plant are usually unbranched although often have an odd branch which are found very low down and so look like a separate stem. The stems have rough ribs each with 2 rows of tiny tubercles. The sheaths have both teeth and hair-points, which during winter fall together. The number of stem ribs or sheath teeth is usually between 7 and 13. The cone of the plant and looks like that of Variegated horsetail but is sterile and so is less developed and often remains partly included in the top sheath. The sheath is pale green or grey with a black band at the top, eventually turning entirely black. The teeth have a black centre with pale edges and may also turn wholly black. MacKay's horsetail prefers coastal areas. This species, a sterile hybrid of rough horsetail and variegated horsetail, occurs in three sites in the Cheshire region, Red Rocks Marsh Nature Reserve and Royal Liverpool Golf Course at Hoylake, and Wallasey Golf Course, all on the Wirral peninsula. The species has been lost from a site in Meols but a substantial population was found a couple of years ago along a footpath south of Hoylake station. Nationally, the species occurs locally in two dozen locations in Ireland and five in Scotland but no other sites in England. It thrives particularly in sites where competition from other species is low and especially where this state is maintained by surface erosion or continual addition of wind blown sand. * Habitat loss to coastal accretion - it is not clear whether this natural change will provide new good habitat or not. * Habitat loss to cycleway * Open Golf Championship at Royal Liverpool Golf Course, 2006. Objectives, targets and actions to help conserve mackays horsetail in the Cheshire region can be found on the Biodiversity Action Reporting System (BARS) along with full details of our progress so far. LBAP Chair Hilary Ash Page, C. N.: The Ferns of Britain and Ireland, 2nd edition. Page, C. N.: A Natural History of Britains Ferns, New Naturalist Series http://www.plant-identification.co.uk/skye/equisetaceae/equisetum-x-trachyodon.htm .
Recommended publications
  • Equisetaceae – Horsetail Family
    EQUISETACEAE – HORSETAIL FAMILY Plant: Stem: jointed, with nodes Root: Leaves: small, whorled, reduced and fused into sheaths with free tips (usually termed teeth) Flowers: no true flowers; spores (all alike) from sporphylls on sporangia located on cones (strobilus), spores usually green except in hybrids; male and female gametophytes green, male smaller than female Fruit: spores Other: worldwide; Division Equisetophyta, Horsetail Group Genera: 1 genus – Equisetum (horsetails or scouring rush), 15+ species WARNING – family descriptions are only a layman’s guide and should not be used as definitive EQUISETACEAE – HORSETAIL FAMILY Field Horsetail; Equisetum arvense L. [Common] Scouring Rush Horsetail; Equisetum hyemale L. var. affine (Engelm.) A.A. Eaton Field Horsetail USDA Equisetum arvense L. Equisetaceae (Horsetail Family) Oak Openings Metropark, Lucas County, Ohio Notes: Dimorphic (fertile and vegetative stems); fertile stems non-green, usually brownish, lacking stomata (pores), non-branching, shorter than veg. stems, sheath teeth dark, usually 14 or less, dies back after spores released; vegetative stems hollow and green, branched in whorls, branches solid with 3-4 ridges; spring [V Max Brown, 2008] [Common] Scouring Rush USDA Horsetail Equisetum hyemale L. var. affine (Engelm.) A.A. Eaton Equisetaceae (Horsetail Family) Alley Springs, Shannon County, Missouri Notes: medium to tall plant, up to 220 cm, unbranched stem (or with a few scattered branches), usually persists more than one year (perennial), with 14 to 50 ridges, stomatal lines single, often rough to the touch; sheaths dark at most nodes (often 2 dark bands separated by a white band), 14 or more teeth; apex of cone fairly sharp or pointed; spores green and spherical; often found on banks of streams, ponds, and margins of lakes as well as along ditches, roadsides, etc.; spring to summer [V Max Brown, 2008].
    [Show full text]
  • Ferns of the National Forests in Alaska
    Ferns of the National Forests in Alaska United States Forest Service R10-RG-182 Department of Alaska Region June 2010 Agriculture Ferns abound in Alaska’s two national forests, the Chugach and the Tongass, which are situated on the southcentral and southeastern coast respectively. These forests contain myriad habitats where ferns thrive. Most showy are the ferns occupying the forest floor of temperate rainforest habitats. However, ferns grow in nearly all non-forested habitats such as beach meadows, wet meadows, alpine meadows, high alpine, and talus slopes. The cool, wet climate highly influenced by the Pacific Ocean creates ideal growing conditions for ferns. In the past, ferns had been loosely grouped with other spore-bearing vascular plants, often called “fern allies.” Recent genetic studies reveal surprises about the relationships among ferns and fern allies. First, ferns appear to be closely related to horsetails; in fact these plants are now grouped as ferns. Second, plants commonly called fern allies (club-mosses, spike-mosses and quillworts) are not at all related to the ferns. General relationships among members of the plant kingdom are shown in the diagram below. Ferns & Horsetails Flowering Plants Conifers Club-mosses, Spike-mosses & Quillworts Mosses & Liverworts Thirty of the fifty-four ferns and horsetails known to grow in Alaska’s national forests are described and pictured in this brochure. They are arranged in the same order as listed in the fern checklist presented on pages 26 and 27. 2 Midrib Blade Pinnule(s) Frond (leaf) Pinna Petiole (leaf stalk) Parts of a fern frond, northern wood fern (p.
    [Show full text]
  • Sobre Los Equisetum × Trachyodon Y Equisetum Variegatum De La Val D´Aiguamòg (Valle De Arán)
    Flora Montiberica 62: 37-42 (I-2016). ISSN: 1138-5952, edic. digital: 1988-799X SOBRE LOS EQUISETUM × TRACHYODON Y EQUISETUM VARIEGATUM DE LA VAL D´AIGUAMÒG (VALLE DE ARÁN) Juan A. ALEJANDRE SÁENZ1, María Josefa ESCALANTE RUIZ1 2 & José Vicente FERRÁNDEZ PALACIO 1C/ Txalaparta, 3, 1º izda. 01006-VITORIA 2C/ Segura, 73. Monzón. 22400-HUESCA RESUMEN: Se aportan testimonios de herbario que contribuyen a con- firmar la presencia actual de Equisetum variegatum Schleich. ex F. Weber & D. Mohr y de una población clonal de Equisetum × trachyodon A. Braun en la zo- na media-alta de la Val d´Aiguamòg (Valle de Arán, Pirineo catalán). Palabras clave: Flora, Equisetum, Pirineo catalán, España. ABSTRACT: Some voucher specimens which contribute to confirm the current presence of Equisetum variegatum Schleich. ex F. Weber & D. Mohr and a clonal population of Equisetum × trachyodon A. Braun in the middle- upper zone of Val d´Aiguamòg (Valle de Arán, Lérida, N Spain) are provided. Keywords: Flora, Equisetum, Catalan Pyrenees, Spain. INTRODUCCIÓN ros en los que observar el contenido esporangial –elemento probatorio de pri- Según lo que se afirma en AYME- mera magnitud en el género Equisetum–. RICH & SÁEZ (2013: 181), los datos com- En una venturosa excursión a esa zona probados con los que se cuenta hasta el que nosotros hicimos el 11 de julio de presente sobre la presencia actual de 2011, con la intención de comparar nues- Equisetum variegatum A. Braun y de tras recolecciones de Equisetum, princi- E. × trachyodon Schleich., en la Val palmente del E. × trachyodon, proce- d´Aiguamòg no llegan a despejar algu- dentes del macizo del Castro Valnera nas dudas sobre la existencia del prime- (Cordillera Cantábrica entre Burgos y ro de ellos en esa zona ni totalmente Cantabria), tuvimos la fortuna de locali- sobre la identidad de los testimonios que zar una población dispersa de E.
    [Show full text]
  • Scouring-Rush Horsetail Scientific Name: Equisetum Hyemale Order
    Common Name: Scouring-rush Horsetail Scientific Name: Equisetum hyemale Order: Equisetales Family: Equisetaceae Wetland Plant Status: Facultative Ecology & Description Scouring-rush horsetail is an evergreen, perennial plant that completes a growing season in two years. At maturity, scouring-rush horsetail usually averages 3 feet in height but can be range anywhere from 2 to 5 feet. It can survive in a variety of environments. One single plant can spread 6 feet in diameter. It has cylindrical stems that averages a third of an inch in diameter. Noticeably spotted are the jointed unions that are located down the plant. The stems are hollow and don’t branch off into additional stems. Also, scouring- rush horsetail has rough ridges that run longitudinal along the stem. Although not covered in leaves, tiny leaves are joined together around the stem which then forms a black or green band, or sheath at each individual joint on the stem. This plant has an enormous root system that can reach 6 feet deep and propagates in two ways: rhizomes and spores. Incredibly, due to the fact that this plant is not full of leaves, it is forced to photosynthesize through the stem rather than leaves. Habitat Scouring-rush horsetail is highly tolerant of tough conditions. It can survive and thrive in full sun or part shade and can successfully grow in a variety of soil types. It can also grow in moderate to wet soils, and can survive in up to 4 inches of water. Distribution Scouring-rush horsetail can be found throughout the United States, Eurasia, and Canada.
    [Show full text]
  • Subclase Equisetidae ¿Tienes Alguna Duda, Sugerencia O Corrección Acerca De Este Taxón? Envíanosla Y Con Gusto La Atenderemos
    subclase Equisetidae ¿Tienes alguna duda, sugerencia o corrección acerca de este taxón? Envíanosla y con gusto la atenderemos. Ver todas las fotos etiquetadas con Equisetidae en Banco de Imagénes » Descripción de WIKIPEDIAES Ver en Wikipedia (español) → Ver Pteridophyta para una introducción a las plantas Equisetos vasculares sin semilla Rango temporal: Devónico-Holoceno PreЄ Є O S D C P T J K Pg N Los equisetos , llamados Equisetidae en la moderna clasificación de Christenhusz et al. 2011,[1] [2] [3] o también Equisetopsida o Equisetophyta, y en paleobotánica es más común Sphenopsida, son plantas vasculares afines a los helechos que aparecieron en el Devónico, pero que actualmente sobrevive únicamente el género Equisetum, si bien hay representantes de órdenes extintos que se verán en este artículo. Este grupo es monofilético, aun con sus representantes extintos, debido a su morfología distintiva. Son plantas pequeñas, aunque en el pasado una variedad de calamitácea alcanzó los 15 metros durante el pérmico.[4] Índice 1 Filogenia 1.1 Ecología y evolución 2 Taxonomía 2.1 Sinonimia Variedades de Equisetum 2.2 Sistema de Christenhusz et al. 2011 Taxonomía 2.3 Clasificación sensu Smith et al. 2006 2.4 Otras clasificaciones Reino: Plantae 3 Caracteres Viridiplantae 4 Véase también Streptophyta 5 Referencias Streptophytina 6 Bibliografía Embryophyta (sin rango) 7 Enlaces externos Tracheophyta Euphyllophyta Monilophyta Filogenia[editar] Equisetopsida o Sphenopsida Introducción teórica en Filogenia Clase: C.Agardh 1825 / Engler 1924 Equisetidae Los análisis moleculares y genéticos de filogenia solo Subclase: se pueden hacer sobre representantes vivientes, Warm. 1883 como circunscripto según Smith et al. (2006) (ver la Órdenes ficha), al menos Equisetales es monofilético (Pryer et Equisetales (DC.
    [Show full text]
  • Equisetum Ramosissimum Desf. Var. Flagelliferum Milde
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Wulfenia Jahr/Year: 2004 Band/Volume: 11 Autor(en)/Author(s): Skvortsov Vladimir Engelsovich Artikel/Article: An unexpected record of an African horsetail, Equisetum ramosissimum Desf. var. flagelliferum Milde (Equisetaceae), fromCaucasus 69-83 © Landesmuseum für Kärnten; download www.landesmuseum.ktn.gv.at/wulfenia; www.biologiezentrum.at Wulfenia 11 (2004): 69–83 Mitteilungen des Kärntner Botanikzentrums Klagenfurt An unexpected record of an African horsetail, Equisetum ramosissimum Desf. var. flagelliferum Milde (Equisetaceae), from Caucasus Vladimir Engelsovich Skvortsov Summary: A giant evergreen variety of Equisetum ramossisimum Desf. var. flagelliferum Milde (subgenus Hippochaete), was found and identified by the author in the suburbs of Sochi town, Black Sea shore, Caucasus, Russia, in February 2004. In its gross morphology this variety is characterized by very tall (up to 2–2.5m) perennial aerial shoots, climbing growth form, regular unilateral branching and by very long (up to 45cm) whorled branches which are usually branched too (even to the third extent). It looks very similar to E. debile Roxb. (E. ramosissimum Desf. subsp. debile (Roxb.) Hauke), from the tropics of South-Eastern Asia but differs quite distinctly by particular (individual) endo- dermis and by multiserial stomata on the main axis. In their micromorphology and anatomy these Caucasian plants have many resemblances to the tropical South-American species E. giganteum, in regular disposition of stomata in 2–4 rows (this is unique and cannot be found among the other species belonging to the subgenus Hippochaete) and in quite variable vallecular collenchyma patterns.
    [Show full text]
  • The Yellowstone Paleontological Survey
    E PALEONT ON O T LO S G W I O C L A L L E National Y Park The Yellowstone Service Department of the Interior Paleontological Survey SURVEY Vincent L. Santucci Yellowstone Center for Resources National Park Service Yellowstone National Park, Wyoming YCR-NR-98-1 1998 How to cite this document: Santucci, V. L. 1998. The Yellowstone Paleontological Survey. Yellowstone Center for Resources, National Park Service, Yellowstone National Park, Wyoming,YCR-NR-98-1. Current address for Vincent L. Santucci is National Park Service, P.O. Box 592, Kemmerer, WY 83101. The Yellowstone Paleontological Survey To Lt. Col. Luke J. Barnett, III “Uncle by blood, brother in spirit!” Vincent L. Santucci Yellowstone Center for Resources National Park Service Yellowstone National Park, Wyoming YCR-NR-98-1 1998 Table of Contents Introduction .................................................................................................... 1 Stratigraphy .................................................................................................... 4 Fossil Chronology........................................................................................... 6 Taxonomy ..................................................................................................... 12 Localities ...................................................................................................... 15 Interpretation ................................................................................................ 19 Paleontological Resource Management.......................................................
    [Show full text]
  • EQUISETACEAE (Horsetail Family) Equisetum Arvense - Common Horsetail E
    VASCULAR FLORA OF J. CLARK SALYER NATIONAL WILDLIFE REFUGE (2001) EQUISETACEAE (Horsetail Family) Equisetum arvense - common horsetail E. laevigatum - smooth horsetail SELAGINELLACEAE (Spikemoss Family) Selaginella densa - little clubmoss CUPRESSACEAE (Cypress Family) Juniperus scopulorum - rocky mountain juniper ALISMATACEAE (Waterplantain Family) Alisma gramineum - grass waterplantain A. plantago-aquatica - waterplantain Sagittaria cuneata - arrowhead JUNCAGINACEAE (Arrowgrass Family) Triglochin maritima - arrowgrass POTAMOGETONACEAE (Pondweed Family) Potamogeton pectinatus - sago pondweed P. richardsonii - claspingleaf pondweed ZANNICHELLIACEAE (Horned Pondweed Family) Zannichellia palustris - horned pondweed JUNCACEAE (Rush Family) Juncus balticus - Baltic rush J. interior - inland rush CYPERACEAE (Sedge Family) Carex atherodesd - slough sedge C. brevior - short-beaked sedge C. duriuscula (+C. eleocharis) C. eleocharis - needle-leaved sedge C. filifolia - threadleaf sedge C. gravida - heavy sedge C. inops subsp. Heliophila (+C. heliophilia) - sun sedge C. lacustris C. laeviconica - glabrous sedge C. lanuginosa - woolly sedge C. obtusata C. praegracilis - clusterfield sedge C. rosea C. sartwellii - Sartwell’s sedge C. siccata C. sprengwlii - long-beaked sedge C. tetanica Cyperus schweinitzii - Schweinitz’s flatsedge Eleocharis erythropoda - marsh spikesedge Schoenoplectus acutus (+Scirpus acutus) - hardstem bulrush Scirpus fluviatilis - river bulrush S. maritimus - prairie bulrush S. tabernaemontani - softstem bulrush POACEAE (Grass
    [Show full text]
  • Typifications of the Linnaean Name Equisetum Hyemale and E. ×Moorei (Equisetaceae)
    See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/316460415 Typifications of the Linnaean name Equisetum hyemale and E. ×moorei (Equisetaceae) Article in Phytotaxa · April 2017 DOI: 10.11646/phytotaxa.305.2.4 CITATIONS READS 0 115 3 authors: P. Pablo Ferrer-Gallego Emilio Laguna Generalitat Valenciana University of Valencia 366 PUBLICATIONS 297 CITATIONS 604 PUBLICATIONS 1,099 CITATIONS SEE PROFILE SEE PROFILE Josep A. Rossello University of Valencia 164 PUBLICATIONS 2,046 CITATIONS SEE PROFILE Some of the authors of this publication are also working on these related projects: Centaurea Typification Project View project Knowledge of the Valencian Flora View project All content following this page was uploaded by Josep A. Rossello on 26 April 2017. The user has requested enhancement of the downloaded file. Phytotaxa 305 (2): 104–110 ISSN 1179-3155 (print edition) http://www.mapress.com/j/pt/ PHYTOTAXA Copyright © 2017 Magnolia Press Article ISSN 1179-3163 (online edition) https://doi.org/10.11646/phytotaxa.305.2.4 Typifications of the Linnaean name Equisetum hyemale and E. ×moorei (Equisetaceae) P. PABLO FERRER-GALLEGO1,2*, EMILIO LAGUNA1 & JOSEP A. ROSSELLÓ3,4 1Servicio de Vida Silvestre, Centro para la Investigación y Experimentación Forestal (CIEF), Generalitat Valenciana, Avda. Comarques del País Valencià 114, 46930 Quart de Poblet, Valencia, Spain; e-mail: [email protected] 2VAERSA, Avda. Cortes Valencianas, nº 20, 46015, Valencia, Spain 3Jardín Botánico–ICBiBE–Unidad Asociada CSIC, Universitat de València, c./ Quart 80, E46008, Valencia, Spain 4Carl Faust Fdn., PO Box 112, E17300, Blanes, Spain *author for correspondence Abstract The lectotypes of the names Equisetum hyemale Linnaeus (1753: 1062) and E.
    [Show full text]
  • Fern Gazette V19 P2 V8 Contents
    FERN GAZ. 19(2):37-46. 2012 37 IS EQUISETUM RAMOSISSIMUM (EQUISETACEAE: EQUISETOPHYTA) NATIVE TO THE BRITISH ISLES? F.J.RUMSEY & M.SPENCER Angela Marmont Centre for UK Biodiversity, Natural History Museum, Cromwell Road, London, SW7 5BD, UK (Email: [email protected]) Keywords: Equisetum × meridionale, Equisetum × moorei, herbarium, hybrid ABSTRACT A review of historic collections at BM has revealed a specimen of Equisetum ramosissimum collected by Buddle on Hounslow Heath in c.1705. This thermophilous ruderal species has generally been considered a recent introduction in the British Isles, first reported in 1949, its neophyte status having implications for its continuing protection under Schedule 8 of the Wildlife and Countryside Act and other conservation actions. The hybrid with E. hyemale (E. × moorei) has long been known from the Irish east coast, in the absence of E. ramosissimum. Recently its hybrid with E. variegatum (E. × meridionale) has been recorded from Anglesey. Its recognition prompted a review of specimens of Equisetum subgenus Hippochaete from the Cheshire/Lancashire coast which showed obvious similarities, most previously regarded as E. variegatum or E. × trachyodon. The majority show distinctive micro-morphological characters associated with E. ramosissimum and are considered to be E. × meridionale, or possibly an as yet un-described triploid backcross to E. variegatum. Subsequently another overlooked BM herbarium specimen of E. ramosissimum from the Liverpool area, collected in the 19th century, was detected. We present a summary of the known occurrences of the species and, in the light of these new discoveries, re- assess the species status as “Native or Alien”. INTRODUCTION As part of the London Natural History Society’s London Flora Project, that aims to update Rodney Burton’s (1983) Flora of the London Area, Nick Bertrand, John Swindells and MS have been reviewing historic records based upon herbarium specimens in the Sloane Herbarium at the Natural History Museum.
    [Show full text]
  • REMARKS on EQUISETUM ARVENSE (EQUISETACEAE) in TEXAS Department of Biology Baylor University Waco, Texas 76798-7388 J Mink@Bay
    Mink, J.N., J.R. Singhurst, and W.C. Holmes. Remarks on Equisetum arvense (Equisetaceae) in Texas. Phytoneuron 2011-21: 1–3. REMARKS ON EQUISETUM ARVENSE (EQUISETACEAE) IN TEXAS JEFFREY N. MINK Department of Biology Baylor University Waco, Texas 76798-7388 [email protected] JASON R. SINGHURST Wildlife Diversity Program Texas Parks and Wildlife Department 4200 Smith School Road Austin, Texas 78704 WALTER C. HOLMES Department of Biology Baylor University Waco, Texas 76798-7388 ABSTRACT Equisetum arvense , previously recorded in Texas from one collection in Lubbock County in 1932, is documented here from a recent collection in the headwaters of Murtaugh Creek in Wheeler County. KEY WORDS: Equisetum arvense , Equisetaceae, Texas, Great Plains Documentation of the presence of Equisetum arvense L. in Texas is based upon E.L. Reed 3616 , collected August 1932 from Buffalo Spring in Lubbock County (Correll 1956). In Ferns and Fern Allies of Texas , Correll (1956) did not include herbaria as part of the exsiccatae citations but only listed (on p. 16) herbaria consulted. Reed 3616 is presumed to have been deposited at TTC (Reed Herbarium of Texas Tech University, Lubbock). This is supported by Greuter’s (1990) inclusion of all of Reed’s collections catalogued to TTC in Index Herbariorum . Several inquiries by the authors to TTC about the specimen remain without reply. It is not clear if Correll actually saw the specimen of Equisetum arvense or if the information was obtained from another source, possibly casting doubt as to the true identity of the specimen. That some uncertainty existed may be indicated in Correll’s (1956) statement “… I made a brief stop at Buffalo Lakes (Springs) in the hope of relocating this species.
    [Show full text]
  • The Marattiales and Vegetative Features of the Polypodiids We Now
    VI. Ferns I: The Marattiales and Vegetative Features of the Polypodiids We now take up the ferns, order Marattiales - a group of large tropical ferns with primitive features - and subclass Polypodiidae, the leptosporangiate ferns. (See the PPG phylogeny on page 48a: Susan, Dave, and Michael, are authors.) Members of these two groups are spore-dispersed vascular plants with siphonosteles and megaphylls. A. Marattiales, an Order of Eusporangiate Ferns The Marattiales have a well-documented history. They first appear as tree ferns in the coal swamps right in there with Lepidodendron and Calamites. (They will feature in your second critical reading and writing assignment in this capacity!) The living species are prominent in some hot forests, both in tropical America and tropical Asia. They are very like the leptosporangiate ferns (Polypodiids), but they differ in having the common, primitive, thick-walled sporangium, the eusporangium, and in having a distinctive stele and root structure. 1. Living Plants Go with your TA to the greenhouse to view the potted Angiopteris. The largest of the Marattiales, mature Angiopteris plants bear fronds up to 30 feet in length! a.These plants, like all ferns, have megaphylls. These megaphylls are divided into leaflets called pinnae, which are often divided even further. The feather-like design of these leaves is common among the ferns, suggesting that ferns have some sort of narrow definition to the kinds of leaf design they can evolve. b. The leaflets are borne on stem-like axes called rachises, which, as you can see, have swollen bases on some of the plants in the lab.
    [Show full text]