Notes on W. H. Edwards Specimens in Two Midwestern Collections
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Phylogenetic Relationships of Phyciodes Butterfly Species (Lepidoptera: Nymphalidae): Complex Mtdna Variation and Species Delimitations
Systematic Entomology (2003) 28, 257±273 Phylogenetic relationships of Phyciodes butterfly species (Lepidoptera: Nymphalidae): complex mtDNA variation and species delimitations NIKLAS WAHLBERG*, RITA OLIVEIRA* andJAMES A. SCOTTy *Metapopulation Research Group, Department of Ecology and Systematics, University of Helsinki, Finland, and yLakewood, Colorado, U.S.A. Abstract. Mitochondrial DNA variation wasstudied in the butterfly genus Phyciodes (Lepidoptera: Nymphalidae) by sequencing 1450 bp of the COI gene from 140 individuals of all eleven currently recognized species. The study focused on four species in particular that have been taxonomically difficult for the past century, P. tharos, P. cocyta, P. batesii and P. pulchella. A cladistic analysis of ninety-eight unique haplotypesshowedthat Phyciodes formsa monophyletic group with P. graphica as the most basal species. Of the three informal species groupsdescribed for Phyciodes, only one (the mylitta-group) isunambiguously monophyletic. Within the tharos-group, seven well supported clades were found that correspond to three taxa, P. tharos, P. pulchella and a grade consisting of P. cocyta and P. batesii haplotypesinterdigitated with each other. None of the clades is formed exclusively by one species. The patterns of haplotype variation are the result of both retained ancient polymorphism and introgression. Introgression appearsto be mostcommon between P. cocyta and P. batesii; however, these two species occur sympatrically and are morphologically and ecologically distinct, suggesting that the level of current introgression does not seem to be enough to threaten their genetic integrity. The results indicate that mitochondrial DNA sequences must be used with great caution in delimiting species, especially when infraspecific samples are few, or introgression seems to be rampant. -
NATURESCAPE BRITISH COLUMBIA Caring for Wildlife Habitat at Home
The Stewardship Series NATURESCAPE BRITISH COLUMBIA Caring for Wildlife Habitat at Home Native Plant and Animal Booklet, Coast and Mountains Naturescape British Columbia IS FUNDED BY For More information, call 1-800-387-9853 National Library of Canada Cataloguing in Publication Data Main entry under title: Naturescape British Columbia. Native plant and animal booklet, coast & mountains. – [Stewardship series] Consists of 2 v. in 1. Title on v. [2]: Naturescape British Columbia. Resource booklet, coast & mountains. Vol. [2] compiled by Saila Hull. Also available on the Internet. ISBN 0-7726-5179-5 1. Animals - British Columbia. 2. Plants - British Columbia. 3. Biotic communities - British Columbia. 4. Coastal ecology - British Columbia. 5. Nature conservation - Bibliography. 6. Nature conservation - British Columbia – Information resources. 7. Pacific Coast (B.C.) – Environmental conditions. I. MacDonald, Andrew, 1965- II. Hull, Saila. III. Naturescape British Columbia. IV. Title: Naturescape British Columbia. Resource booklet, coast & mountains. QH77.C3 N37 2004 578’.097111 C2004-960069-9 Notice: The information contained in the Naturescape British Columbia Guide and booklets is true and complete to the best of our knowledge. All recommendations are made without any guarantees on the part of the authors and Naturescape British Columbia sponsors. Because the means, materials and procedures followed by home- owners are beyond our control, the authors and Naturescape British Columbia disclaim all liability in connection with the use of this -
Phylogenetic Relationships and Historical Biogeography of Tribes and Genera in the Subfamily Nymphalinae (Lepidoptera: Nymphalidae)
Blackwell Science, LtdOxford, UKBIJBiological Journal of the Linnean Society 0024-4066The Linnean Society of London, 2005? 2005 862 227251 Original Article PHYLOGENY OF NYMPHALINAE N. WAHLBERG ET AL Biological Journal of the Linnean Society, 2005, 86, 227–251. With 5 figures . Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) NIKLAS WAHLBERG1*, ANDREW V. Z. BROWER2 and SÖREN NYLIN1 1Department of Zoology, Stockholm University, S-106 91 Stockholm, Sweden 2Department of Zoology, Oregon State University, Corvallis, Oregon 97331–2907, USA Received 10 January 2004; accepted for publication 12 November 2004 We infer for the first time the phylogenetic relationships of genera and tribes in the ecologically and evolutionarily well-studied subfamily Nymphalinae using DNA sequence data from three genes: 1450 bp of cytochrome oxidase subunit I (COI) (in the mitochondrial genome), 1077 bp of elongation factor 1-alpha (EF1-a) and 400–403 bp of wing- less (both in the nuclear genome). We explore the influence of each gene region on the support given to each node of the most parsimonious tree derived from a combined analysis of all three genes using Partitioned Bremer Support. We also explore the influence of assuming equal weights for all characters in the combined analysis by investigating the stability of clades to different transition/transversion weighting schemes. We find many strongly supported and stable clades in the Nymphalinae. We are also able to identify ‘rogue’ -
UC Davis UC Davis Previously Published Works
UC Davis UC Davis Previously Published Works Title The utility of repeated presence data as a surrogate for counts: A case study using butterflies Permalink https://escholarship.org/uc/item/13r5464h Journal Journal of Insect Conservation, 18(1) ISSN 1366-638X Authors Casner, KL Forister, ML Ram, K et al. Publication Date 2014-02-01 DOI 10.1007/s10841-013-9610-8 Peer reviewed eScholarship.org Powered by the California Digital Library University of California The utility of repeated presence data as a surrogate for counts: a case study using butterflies Kayce L. Casner, Matthew L. Forister, Karthik Ram & Arthur M. Shapiro Journal of Insect Conservation An international journal devoted to the conservation of insects and related invertebrates ISSN 1366-638X Volume 18 Number 1 J Insect Conserv (2014) 18:13-27 DOI 10.1007/s10841-013-9610-8 1 23 Your article is protected by copyright and all rights are held exclusively by Springer Science +Business Media Dordrecht. This e-offprint is for personal use only and shall not be self- archived in electronic repositories. If you wish to self-archive your article, please use the accepted manuscript version for posting on your own website. You may further deposit the accepted manuscript version in any repository, provided it is only made publicly available 12 months after official publication or later and provided acknowledgement is given to the original source of publication and a link is inserted to the published article on Springer's website. The link must be accompanied by the following text: "The final publication is available at link.springer.com”. -
Biology and Systematics of Phyciodes (Phyciodes)
cf#']i:z 10 Feb. 20, 1998 PAPILIO ~ $4.00 - ..- ·-- .. -· - ~- - PHYCIODES (PHYCIOD£S): N£W DISCOVERIES, NEW SUBSPECIES, AND comf£R6ENCE by DR. JAMES A. SCOTT 60 Estes Street, Lakewood, Colorado 80226 Abstract. The paper presents new discoveries in Phyciodes (Phyciodes ). New life history information is given for P. mylitta arizonensis, P. pallida pallida, P. pallida barnesi, P. tharos tharos, P. cocyta selenis, P. cocyta dimin'utor, P. batesii lakota, P. b. apsaalooke, P. b. anasazi, P. pulchella camillus, and P. picta picta. P. batesii anasazi was found to significantly converge toward P. cocyta in western Colorado in appearance of some adults larvae and pupae, and P. cocyta selenis from the same area was found to converge a little toward anasazi in adult wing pattern and larval head pattern, so some individuals of these two tax.a are difficult to distinguish. The nomenclatural identity of P. tharos form marcia is clarified. Five new subspecies are named: P. tharos orantain has orange antenna nudum and is somewhat intermediate between P. tharos tharos and P. cocyta. P. cocyta diminutor is multivoltine with late-flying summer generations that are small like P. tharos, and might possibly even be a separate species from the sympatric univoltine larger P. cocyta selenis. P. pulchella deltarufa has oranger ups than ssp. pulchella. P. pulchella owimba differs from ssp. pulchella by having an orange antenna nudum and some different wing pattern details. P. phaon jalapeno has paler median ups bands. INTRODUCTION Scott (1994) reported on the systematics and biology of Phyciodes (Phyciodes), described numerous new life histories, named five new tax.a, and corrected prior nomenclature. -
BUTTERFLIES in Thewest Indies of the Caribbean
PO Box 9021, Wilmington, DE 19809, USA E-mail: [email protected]@focusonnature.com Phone: Toll-free in USA 1-888-721-3555 oror 302/529-1876302/529-1876 BUTTERFLIES and MOTHS in the West Indies of the Caribbean in Antigua and Barbuda the Bahamas Barbados the Cayman Islands Cuba Dominica the Dominican Republic Guadeloupe Jamaica Montserrat Puerto Rico Saint Lucia Saint Vincent the Virgin Islands and the ABC islands of Aruba, Bonaire, and Curacao Butterflies in the Caribbean exclusively in Trinidad & Tobago are not in this list. Focus On Nature Tours in the Caribbean have been in: January, February, March, April, May, July, and December. Upper right photo: a HISPANIOLAN KING, Anetia jaegeri, photographed during the FONT tour in the Dominican Republic in February 2012. The genus is nearly entirely in West Indian islands, the species is nearly restricted to Hispaniola. This list of Butterflies of the West Indies compiled by Armas Hill Among the butterfly groupings in this list, links to: Swallowtails: family PAPILIONIDAE with the genera: Battus, Papilio, Parides Whites, Yellows, Sulphurs: family PIERIDAE Mimic-whites: subfamily DISMORPHIINAE with the genus: Dismorphia Subfamily PIERINAE withwith thethe genera:genera: Ascia,Ascia, Ganyra,Ganyra, Glutophrissa,Glutophrissa, MeleteMelete Subfamily COLIADINAE with the genera: Abaeis, Anteos, Aphrissa, Eurema, Kricogonia, Nathalis, Phoebis, Pyrisitia, Zerene Gossamer Wings: family LYCAENIDAE Hairstreaks: subfamily THECLINAE with the genera: Allosmaitia, Calycopis, Chlorostrymon, Cyanophrys, -
POCKET GUIDE to Common Kansas Butterflies ■ ■ ■ ■ ■
A POCKET GUIDE TO Common Kansas Butterflies ■ ■ ■ ■ ■ By Jim Mason Funded by Westar Energy Green Team, Glenn Springs Holdings, Inc., Occidental Chemical Corporation and the Chickadee Checkoff Published by the Friends of the Great Plains Nature Center Table of Contents • Introduction • 2 • Butterflies vs. Moths • 4 • Observing Butterflies • 4 Family Papilionidae - Swallowtails ■ Pipevine Swallowtail • 6 ■ Zebra Swallowtail • 7 ■ Black Swallowtail • 8 ■ Giant Swallowtail • 9 ■ Eastern Tiger Swallowtail • 10 Family Pieridae – Whites & Sulphurs ■ Checkered White • 11 ■ Cabbage White • 12 ■ Clouded Sulphur • 13 ■ Orange Sulphur • 14 ■ Cloudless Sulphur • 15 ■ Sleepy Orange • 16 ■ Little Yellow • 17 ■ Dainty Sulphur • 18 ■ Southern Dogface • 19 Family Lycaenidae – Gossamer-Wings ■ Gray Copper • 20 ■ Bronze Copper • 21 ■ Coral Hairstreak • 22 ■ Gray Hairstreak • 23 ■ Juniper Hairstreak • 24 ■ Reakirts' Blue • 25 ■ Eastern Tailed-Blue • 26 ■ Spring Azure and Summer Azure • 27 Family Nymphalidae – Brushfoots ■ American Snout • 28 ■ Variegated Fritillary • 29 ■ Great Spangled Fritillary • 30 ■ Regal Fritillary • 31 ■ Gorgone Checkerspot • 32 ■ Silvery Checkerspot • 33 ■ Phaon Crescent • 34 ■ Pearl Crescent • 35 ■ Question Mark • 36 ■ Eastern Comma • 37 ■ Mourning Cloak • 38 ■ American Lady • 39 ©Greg Sievert ■ Painted Lady • 40 ■ Red Admiral • 41 ■ Common Buckeye • 42 ■ Red-spotted Purple • 43 ■ Viceroy • 44 ■ Goatweed Leafwing • 45 ■ Hackberry Emperor • 46 ■ Tawny Emperor • 47 ■ Little Wood Satyr • 48 ■ Common Wood Nymph • 49 ■ Monarch • 50 Family -
2010 Butterfly Inventories in Boulder County Open Space Properties
2010 Butterfly Inventories In Boulder County Open Space Properties By Janet Chu October 2, 2010 1 Table of Contents I. Acknowledgments …………………… 3 II. Abstract …………………………… 4 II. Introduction……………………………… 5 IV. Objectives ………………………….. 6 V. Research Methods ………………….. 7 VI. Results and Discussion ………………... 8 VII. Weather ………………………………… 12 VIII. Conclusions …………………………….. 13 VIII. Recommendations …………………….. 15 IX. References …………………………. 16 X. Butterfly Survey Data Tables …………. 17 Table I. Survey Dates and Locations ……………. 17 Table II. Southeast Buffer …………………. 18 Table III. Anne U. White – Fourmile Trail …… 21 Table IV. Heil Valley Open Space –Geer Watershed... 24 Table V. Heil Valley Open Space –Plumely Canyon 27 Table VI. Heil Valley Open Space – North ………… 30 Table VII. Walker Ranch - Meyer’s Gulch ………… 34 Table VIII. Caribou Ranch Open Space ……………… 37 Table IX. Compilation of Species and Locations …… 38 2 I. Acknowledgments Our research team has conducted butterfly surveys for nine consecutive years, from 2002 through 2010, with 2002-2004 introductory to the lands and species, and 2005-2010 in more depth. My valuable field team this year was composed of friends with sharp eyes and ready binoculars Larry Crowley who recorded not only the butterflies but blossoming plants and wildlife joined by Jean Morgan and Amy Chu both joined enthusiastic butterfly chasers. Venice Kelley and John Barr, professional photographers, joined us on many surveys. With their digital photos we are often able to classify the hard-to-identify butterflies later on at the desk. The surveys have been within Boulder County Parks and Open Space (BCPOS) lands. Therese Glowacki, Manager-Resource Manager, issued a Special Collection Permit for access into the Open Spaces; Susan Spaulding, Wildlife Specialist, oversaw research, maintained records of our monographs and organized seminars for presentation of data. -
Papilio (New Series) # 25 2016 Issn 2372-9449
PAPILIO (NEW SERIES) # 25 2016 ISSN 2372-9449 ERNEST J. OSLAR, 1858-1944: HIS TRAVEL AND COLLECTION ITINERARY, AND HIS BUTTERFLIES by James A. Scott, Ph.D. in entomology University of California Berkeley, 1972 (e-mail: [email protected]) Abstract. Ernest John Oslar collected more than 50,000 butterflies and moths and other insects and sold them to many taxonomists and museums throughout the world. This paper attempts to determine his travels in America to collect those specimens, by using data from labeled specimens (most in his remaining collection but some from published papers) plus information from correspondence etc. and a few small field diaries preserved by his descendants. The butterfly specimens and their localities/dates in his collection in the C. P. Gillette Museum (Colorado State University, Fort Collins, Colorado) are detailed. This information will help determine the possible collection locations of Oslar specimens that lack accurate collection data. Many more biographical details of Oslar are revealed, and the 26 insects named for Oslar are detailed. Introduction The last collection of Ernest J. Oslar, ~2159 papered butterfly specimens and several moths, was found in the C. P. Gillette Museum, Colorado State University, Fort Collins, Colorado by Paul A. Opler, providing the opportunity to study his travels and collections. Scott & Fisher (2014) documented specimens sent by Ernest J. Oslar of about 100 Argynnis (Speyeria) nokomis nokomis Edwards labeled from the San Juan Mts. and Hall Valley of Colorado, which were collected by Wilmatte Cockerell at Beulah New Mexico, and documented Oslar’s specimens of Oeneis alberta oslari Skinner labeled from Deer Creek Canyon, [Jefferson County] Colorado, September 25, 1909, which were collected in South Park, Park Co. -
Nevada Butterflies and Their Biology to Forward Such for Inclusion in the Larger Study
Journal of the Lepidopterists' Society 39(2). 1985. 95-118 NEV ADA BUTTERFLIES: PRELIMINARY CHECKLIST AND DISTRIBUTION GEORGE T. AUSTIN Nevada State Museum and Historical Society, 700 Twin Lakes Drive, Las Vegas, Nevada 89107 ABSTRACT. The distribution by county of the 189 species (over 300 taxa) of but terflies occurring in Nevada is presented along with a list of species incorrectly recorded for the state. There are still large areas which are poorly or not collected. Nevada continues as one of the remaining unknown areas in our knowledge of butterfly distribution in North America. Although a com prehensive work on the state's butterflies is in preparation, there is sufficient demand for a preliminary checklist to justify the following. It is hoped this will stimulate those who have any data on Nevada butterflies and their biology to forward such for inclusion in the larger study. Studies of Nevada butterflies are hampered by a paucity of resident collectors, a large number of mountain and valley systems and vast areas with little or no access. Non-resident collectors usually funnel into known and well worked areas, and, although their data are valu able, large areas of the state remain uncollected. Intensive collecting, with emphasis on poorly known areas, over the past seven years by Nevada State Museum personnel and associates has gone far to clarify butterfly distribution within the state. The gaps in knowledge are now more narrowly identifiable and will be filled during the next few sea sons. There is no all encompassing treatment of Nevada's butterfly fauna. The only state list is an informal recent checklist of species (Harjes, 1980). -
Santa Monica Mountains Butterfly List
Santa Monica Mountains Butterfly List Probable in Griffith Park Abundance Codes: A- Abundant C- Common Possible in Griffith Park U- Uncommon R- Rare ?- Not Yet Found Food Plant Flight Time Swallowtails Papilionidae Anise Swallowtail Papilio zelicaon Fennel or other carrot family All year Giant Swallowtail Papilio cresphones Citrus Mar-Oct Western Tiger Swallowtail Papilio rutulus Mostly sycamore; cottonwood, willow, alder All year Pale Swallowtail Papilio eurymedon Ceanothus, other buckthorn family Feb-Oct White and Sulphurs Pieridae Whites Becker's White Pontia beckerii Bladderpod Feb-Aug Checkered White Pontia protodice Mustards, native and non-native All year Cabbage White Pieris rapae Black mustard, wild radish, nasturtium All year Sara Orangetip Anthocharis sara Native mustards Feb-June Sulphurs Orange Sulphur Colias eurytheme Various pea family: lotus, clover, vetch All year Harford's Sulphur Colias harfordii Astragalus (locoweed or rattleweed) Apr-Sept California Dogface Colias eurydice False indigo (Amorpha californica) Mar-Sept Cloudless Sulphur Phoebis sennae Cassia (Senna) Mar-Oct Sleepy Orange Eurema nicippe Cassia (Senna) Mar-Oct Dainty Sulphur Nathalis iole Various sunflower family Mar-Nov Gossamer-Wings Lycaenidae Coppers Tailed Copper Lycaena arota Wild currant or goooseberry (Ribes) May-July Gorgon Copper Lycaena gorgon Wand Buckwheat (Eriogonum elongatum) Apr-July Hairstreaks Great Purple Hairstreak Atlides halesus Mistletoe in sycamores or cottonwood Mar-Oct California Hairstreak Satyrium californica Ceanothus (wild -
Common Kansas Butterflies ■ ■ ■ ■ ■
A POCKET GUIDE TO Common Kansas Butterflies ■ ■ ■ ■ ■ By Jim Mason Funded by Westar Energy Green Team, Glenn Springs Holdings, Inc., Occidental Chemical Corporation and the Chickadee Checkoff Published by the Friends of the Great Plains Nature Center Table of Contents • Introduction • 2 • Butterflies vs. Moths • 4 • Observing Butterflies • 4 Family Papilionidae - Swallowtails ■ Pipevine Swallowtail • 6 ■ Zebra Swallowtail • 7 ■ Black Swallowtail • 8 ■ Giant Swallowtail • 9 ■ Eastern Tiger Swallowtail • 10 Family Pieridae – Whites & Sulphurs ■ Checkered White • 11 ■ Cabbage White • 12 ■ Clouded Sulphur • 13 ■ Orange Sulphur • 14 ■ Cloudless Sulphur • 15 ■ Sleepy Orange • 16 ■ Little Yellow • 17 ■ Dainty Sulphur • 18 ■ Southern Dogface • 19 Family Lycaenidae – Gossamer-Wings ■ Gray Copper • 20 ■ Bronze Copper • 21 ■ Coral Hairstreak • 22 ■ Gray Hairstreak • 23 ■ Juniper Hairstreak • 24 ■ Reakirts' Blue • 25 ■ Eastern Tailed-Blue • 26 ■ Spring Azure and Summer Azure • 27 Family Nymphalidae – Brushfoots ■ American Snout • 28 ■ Variegated Fritillary • 29 ■ Great Spangled Fritillary • 30 ■ Regal Fritillary • 31 ■ Gorgone Checkerspot • 32 ■ Silvery Checkerspot • 33 ■ Phaon Crescent • 34 ■ Pearl Crescent • 35 ■ Question Mark • 36 ■ Eastern Comma • 37 ■ Mourning Cloak • 38 ■ American Lady • 39 ©Greg Sievert ■ Painted Lady • 40 ■ Red Admiral • 41 ■ Common Buckeye • 42 ■ Red-spotted Purple • 43 ■ Viceroy • 44 ■ Goatweed Leafwing • 45 ■ Hackberry Emperor • 46 ■ Tawny Emperor • 47 ■ Little Wood Satyr • 48 ■ Common Wood Nymph • 49 ■ Monarch • 50 Family