J. Jpn. Bot. 91(5): 302–304 (2016)

a, b c d Hiroyoshi Ohashi *, Tetsuo Ohi-Toma , Toshio Katsuki and Nobuyuki Tanaka : Lectotype of japonica () aHerbarium TUS, Botanical Garden, Tohoku University, Sendai, 980-0862 JAPAN; bBotanical Gardens, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, 112-0001 JAPAN; cTama Forest Science Garden, Forestry and Forest Products Research Institute, National Research and Development Agency, Japan, 1833-81, Todori-machi, Hachioji, Tokyo, 193-0843 JAPAN; dDepartment of Botany, National Museum of Nature and Science, 4-1-1, Amakubo, Tsukuba, 305-0005 JAPAN *Corresponding author: [email protected]

Summary: Tsuga japonica Shiras. is lectotypified and Wakayama Prefectures) in central Honshu by Plate III in the protologue published in Bot. and southeastern Kochi Pref. in Shikoku. The Mag. (Tokyo) 9(no. 96) in 1895. basionym of the name is Tsuga japonica Shiras. japonica (Shiras.) Beissn. It was published as “Tsuga (Pseudo-tsuga) is one of the remarkable endemic japonica Shirasawa” in Bot. Mag. (Tokyo) 9(no. characterizing the flora of Japan. It occurs only 96): 41 (in Japanese) and 84 (in German) with in the southcentral Kii peninsula (Mie, Nara, Plate III in 1895 in the protologue. Pseudotsuga

Fig. 1. Lectotype of Tsuga japonica Shiras.

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(as Pseudo-tsuga) in parenthesis is regarded as Shiras.’; Ohwi, Fl. Jap.: 42 (1953), & Fl. Jap. an infrageneric taxon of the genus Tsuga (ICN ed. Engl.: 112 (1965) as ‘Pseudotsuga japonica Art. 20.1, Rec. 21A.1 and Art. 23.1. McNeill et (Shiras.) Beissn.’; Hayashi, Taxon. Phytogeogr. al. 2012, Ohashi et al. 2014), although Shirasawa Stud. Jap. Conif.: 27 (1960); Kurata, Illust. (1895) stated it belongs to Pseudotsuga in the Import. Forest Tr. Jap. 1: 36, t. 18 (1964); Farjon protologue in Japanese. Shirasawa collected in Regnum Veg. 121 [Pinac.]: 189 (1990); T. the with branchlets and leaves near Yamaz. in K. Iwats. & al., Fl. Jap. 1: 270 (1995); Ryuunotani in Yoshino-gun, Yamato Province, H. Ohashi in H. Ohashi & al., Wild Flow. Japan in July 1893. Further, he got cones and wood 1: 32, pl. 21: 1–4 (2015). from a native resident, Mikio Doi. He described Tsuga japonica Shiras. in Bot. Mag. (Tokyo) the species based on these materials. However, 9: 41 (in Japanese) & 9: 86 (in German), pl. neither herbarium specimens nor cones of the 3 (1895), as Tsuga (Pseudo-tsuga) japonica species can be found in TF (The Herbarium of Shirasawa. Forestry and Forest Products Research Institute, Lectotype (designated here): Plate 3 in Bot. National Research and Development Agency, Mag. (Tokyo) 9(no. 96). (1895). Japan), TI, TNS, and TOFO. No other herbaria have any possibility to keep Shirasawa’s We thank Prof. J. Murata (TI), Dr. H. Ikeda collection in Japan. Also, no previous research (TI), Dr. H. Hoshi of The Forestry and Forest on the species has designated its type. Under Products Research Institute, National Research these circumstances, therefore, we select Plate and Development Agency, and Dr. K. Yonekura III in the protologue of Tsuga japonica as the (TUS) for their help in searching Shirasawa’s lectotype of the name (Fig. 1) in accordance specimens. with ICN Art. 9.2, 9.11 and 9.12 (McNeill et al. 2012, Ohashi et al. 2014). Each illustration in the References plate is explained in Japanese on page 43 and in McNeill J., Barrie F. R., Buck W. R., Demoulin V., Gtriyrt German on page 84 in the protologue. W., Hawksworth D. L., Herendeen P. S., Knapp S., Marhold K., Prado J., Prud’homme van Reine W. F., Smith G. F., Wiersema J. H. and Turland N. J. Pseudotsuga japonica (Shiras.) Beissn., Mitt. (eds.). 2012. International Code of Nomenclature for Deutsch. Dendrol. Ges. 5: 62 (1896); Shiras., algae, fungi, and (Melbourne Code). Regnum Icon. Ess. For. Tr. Jap. 1: 21, t. 7 (1900), as Vegetabile 154. Koeltz Scientific Books, Königstein. ‘Pseudotsuga japonica Shiras.’; Beissn., Handb. Ohashi H., Nagamasu H., Ebihara A., Hayasaka E., Hosoya T., Iketani H., Kurosawa T., Murata J., Nakada T., Nadelh. ed. 2: 111 (1909), as ‘Pseudotsuga Nakai H., Nemoto T., Ohashi K. and Yonekura K. japonica Shiras.’; Jackson in Gard. Chron. ser. 3, 2014. International Code of Nomenclature for Algae, 45: 307, f. 132 (1909), as ‘Pseudotsuga japonica Fungi, and Plants (Melbourne Code). Japanese Edition. Sargent’; E. H. Wilson, Conif. Taxad. Jap.: 52, Hokuryuukan, Tokyo. Shirasawa H. 1895. Shinshu no Nipponsan Shohaku-ka t. 33 & 34 (1916) as ‘Pseudotsuga japonica Shokubutsu. Bot. Mag. (Tokyo) 9: 41–43 (in Japanese); (Shiras.) Beissn.’; Makino, Ill. Fl. Nippon: Eine neue Coniferenart in Japan. Bot. Mag. (Tokyo) 9: 906, f. 2717 (1940), as ‘Pseudotsuga japonica 84–85 (in German), pl. III. 304 植物研究雑誌 第 91 巻 第 5 号 2016 年 10 月

大橋広好 a，大井・東馬哲雄 b，勝木俊雄 c，田中伸幸 d： マツ科トガサワラの学名のレクトタイプ ト ガ サ ワ ラ Pseudotsuga japonica (Shiras.) Beissn. の これまでにトガサワラの学名のタイプは指定されてい 基礎異名 Tsuga japonica Shiras. のタイプ標本を TF（森 ない．このためメルボルン規約に従って，原発表で発表 林総合研究所），TI，TNS，TOFO（さく葉と液浸標本） されたプレート 3 をレクトタイプに指定した． で探索した．原発表によると，大和国吉野郡大字龍の谷 （a東北大学植物園津田記念館， 近傍で白沢保美が枝葉を採集し，後に紀州尾鷲町の土井 b東京大学大学院理学系研究科附属植物園， 幹夫から球果と材幹とを送られ，これらの資料に基づい c森林総合研究所多摩森林科学園， て，白沢が Tsuga japonica Shiras. を発表したという．残 d国立科学博物館植物研究部） 念ながら，その資料のいずれも発見できなかった．また，