A Provisional Atlas of the Coleoptera of Yorkshire (Vice Counties 61 –65)

Part 1 – Suborder Adephaga: Family Carabidae

Introduction

There have been 180 years of Coleoptera recording in the historical county of Yorkshire, which includes the vice-counties 61 to 65. The records comprise the YNU Coleoptera database. The purpose of this series of papers is to reassess all our Coleoptera records in the light of modern knowledge of distribution and habitat requirements. The Coleoptera will be covered, it is hoped, in a number of parts. This part will deal with the Carabidae, one of the largest families in terms of species numbers represented in the British fauna.

The first authoritative Coleoptera list for the county was begun in 1884 (Nat 1884:??) by Rev WC Hey and continued by ML Thompson in parts. The project to produce a definitive Yorkshire list was continued with the publication of the Victoria County History, within which was a section on the Coleoptera of the County of Yorkshire. There has been nothing since.

There have been attempts in the past to review the Yorkshire records, but these have usually tended to deal with the more uncommon taxa, or significant records, or small groups. So-called “common” species have often been neglected in the past, and often not included in the records. I think the main reason for this is that records were added by hand to individual species sheets. Entering a long list of, say, Tachyporus hypnorum, into such a medium must have proven extremely tedious and time-consuming. Therefore it is hardly surprising that recorders only entered significant records. This present work, however, is an attempt to treat all species on an equal footing.

Previous reviews and reports include the following – Flint (1988), (Denton 1990, 2003, 2008), Marsh (1991, 2002).

Each species in the database is considered and in each case a distribution map representing records on the database (at the end of 2007) is presented. The number of records on the database for each species is given in the account in the form (a, b, c, d, e) where a to e are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the Vice- county recording system, as the county is thereby divided up into manageable areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas. Thus Yorkshire was divided into “vice-counties”, numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice- county system remains a convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.

For some species there are many records, and for others only one or two. In cases where there are six species or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older recorders tended not to record species they regarded as common quite as assiduously as we do today.

“Recent” records are those considered to be later than 1 January 1990 – this is also the break point used in the production of the distribution maps.

Habitat A brief statement on habitat is included, and for this I have drawn heavily on two publications, Luff (1998) and Luff (2007), wherein is much species habitat and frequency information.

Taxon status Also given is the accepted national status of each taxon, this being the status as defined in JNCC‟s Recorder 3 data program. Status is included if Notable B or above. Please bear in mind that many of these statuses are now in need of revision. The following status criteria are derived from Shirt (1987) and Hyman and Parsons (1994):

RED DATA BOOK CATEGORY 1. RDB1 – ENDANGERED. Species which are known or believed to occur only as a single population within one 10km square of the OS National Grid.

RED DATA BOOK CATEGORY 2. RDB2 – VULNERABLE. Species which are declining throughout their range.

RED DATA BOOK CATEGORY 3. RDB3 – RARE. Species which are estimated to exist in only 15 or fewer 10km squares; this criterion may be relaxed where populations are likely to exist in more than 15 squares but occupy small areas of especially vulnerable habitat.

RED DATA BOOK CATEGORY I. RDB I – INDETERMINATE. Taxa considered to be endangered, Vulnerable or Rare, but where there is not enough information to say which of the three categories (RDB1 to 3) is appropriate.

1

RED DATA BOOK CATEGORY K. RDBK – INSUFFICIENTLY KNOWN. Taxa recently discovered or recognised in Great Britain, which may prove to be more widespread in the future (although some recent discoveries may be placed in other categories if the group to which they belong is thought to be not under-recorded).

PROVISIONAL RED DATA BOOK. Prdb. The prefix „p‟ before any red Data Book category implies that the grading is provisional.

Notable A – species which do not fall within the RDB categories but are thought to occur in 30 or fewer 10km squares of the National Grid, or for less well-recorded groups, within seven or fewer vice-counties. Notable B – species which do not fall within the RDB categories but thought to occur in between 31 and 100 10km squares of the National Grid.

Phenology Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number of records is sufficient to give a meaningful conclusion.

Maps Distribution maps have been produced using Dr Alan Morton‟s DMAP software (version 7.0b). It may be worthwhile to note that as DMAP is asked to map pre- and post-1990 records in this application, any undated records will not be mapped.

Nomenclature The YNU database is presently held in a copy of Recorder 2002, which uses the checklist of Recorder 3, based on Pope (1977). Very recently, a new Coleoptera checklist has emerged (Duff, 2008) and this replaces that of Pope (loc. cit.); the new list incorporates a very large number of taxon name changes. By May 2009, the database will have been upgraded to Recorder 6 and all recording will be done using the new checklist. This report uses the latest checklist as a framework. Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatise on this subject), e.g. “Snail Hunter”, “Crucifix Beetle” and “Heath-glory ground beetle” (Cychrus caraboides, Panagaeus crux-major and Carabus nitens respectively). Such inventions were designed for use in political documents, and seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, vague, confusing and unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.

Quality of the records A regrettable aspect of some of the records within the YNU database in particular (and I suspect sometimes within entomological datasets in general) is the lack of attention to detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/animals, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records onto Recorder requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference.

Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of insect are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is „new‟ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of „difficult‟ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented.

Although some may disagree, I believe any record of a species afforded Notable B or above, especially if a member of a „difficult‟ beetle group, should be confirmed by one specialising in that group before the record is entered onto the database. This of course would be difficult to police with large numbers of records and/or large Orders.

2 Contributors With approaching 180 years of beetle recording in Yorkshire many persons‟ names, or simply initials, appear in the records. The initials of persons used in the species accounts are listed below, alphabetically.

AAF – (identity unclear) KP – Kenneth Payne AEW – AE Winter KS - identity unclear ASL – Alan Lazenby ME – Mick Eyre AMR – AM Robertson MF – M Fishpool BC – Barry Constantine MH – Martin Hammond BE – Brian Eversham MLD - Mike Denton BK – Basil Kitchen MLL – ML Luff BSL – BS Lucas MLT - ML Thompson CDA – CD Ash MT - Mark Telfer CES - CE Stott MU – M Usher CS – C Simms NW – Nikki Whitehouse CHW – C Hyde West PB – P Buckland CJ – Colin Johnson PF – P Flint CWR – CW Russell PK – Peter Kendall DAL – Derek A Lott PS - Peter Skidmore DC – D. Connell RA – R Adams DD – D Dickenson RBA – Robert B Angus DGH – David Hemingway RC - Roy Crossley DHS – DH Smith RCW – R Colin Welch DL – Dick Loxton RGB – Roger Booth ECH – EC Horrell RJM - RJ Marsh EFG – EF Gilmour RK - Roger Key EGB – EG Bayford RLa – R Lawson EJS – Eric Smith RLe – R Leyland EWA – EW (Ted) Aubrook RM – Robert Merritt FE – F Ellis RRUK – RR Uhthoff-Kaufmann FEK – Frank Kenington SLM – SL Mosely GBW - GB Walsh SLS – Stephen L Sutton GM – Graham Maynard SS – S Shaw GTR - GT Rudd ST – Steve Toher HB – Harry Britten jnr TH – Thomas Hick HEJ – HE Johnson TJB – TJ Bird HHC – HH Corbett TS - T Stainforth JBJ – Jim Jobe WAE – Wiliam A Ely JC – John Coulson WCH – WC Hey JDC - John Coldwell WDH – WD Hincks JDF – JD Firth WFB – WF Baker JHF - John Flint WJF – WJ Fordham JS – Jack Spittle WKB – WK Bissill JWC – JW Carter WRD – WR Dolling JWE – JW Ellis WWF – W W Fowler

3 Family CARABIDAE Latreille, 1802 Subfamily CICINDELINAE Latreille, 1802 CICINDELA Linnaeus, 1758 campestris Linnaeus, 1758

Cicindela campestris Linnaeus, 1758 4 WATSONIAN (5, 36, 62, 10, 7) Widely distributed in open areas 3 YORKSHIRE of heath and moorland. A spring breeder and very 2 active on the wing. Well established in all VCs. Many 1 records: (iii-ix, peak vi). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily CARABINAE Latreille, 1802 Tribe CARABINI Latreille, 1802

CALOSOMA Weber, 1801 inquisitor (Linnaeus, 1758)

Calosoma inquisitor (Linnaeus, 1758) 4 WATSONIAN 3 (0,0,1,0,2) Notable A. Associated with old oak YORKSHIRE 2 woodland with adults feeding on Lepidoptera 1 larvae.(Luff, 2007) (Luff, 1998) . There are three records on the YNU database, West Tanfield 0 SE2779, iii 1968 (CS) and Frost Row Fell SD6890 vi 9 1977(PF), both in VC65. The VC63 record is of a 8 subfossil from Norton Common Farm SE5615 in 7 VC63. See Skidmore, 1971. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/2/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

4 Subgenus EUCARABUS Géhin, 1885 arvensis Herbst, 1784 ssp. sylvaticus Dejean, 1826 silvaticus auctt. (misspelling) anglicanus Motschulsky, 1865/6

Carabus arvensis Herbst, 1784 4 WATSONIAN 3 (2, 30, 2, 7, 3) A species of moorland and heaths, YORKSHIRE 2 recorded from all VCs, the majority being in VC62 on the North York Moors and the higher Pennines 1 in VC64 and VC65. Also Allerthorpe district, 1920- 0 1950, WJF, BK). A well-established population 9 exists at Strensall Common. (iv-xii, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus CARABUS Linnaeus, 1758 granulatus Linnaeus, 1758 ssp. granulatus Linnaeus, 1758 ssp. hibernicus Lindroth, 1956

Carabus granulatus Linnaeus, 1758 4 WATSONIAN (24, 3, 23, 4, 0). A species of marshes and fens. 3 YORKSHIRE Often in pitfall traps and under bark of logs in wet 2 places. In grass tussocks and sedge litter in 1 winter. Main centres of population are Doncaster 0 district, Thorne and Hatfield Moors and Lower 9 Derwent Ings with outliers from Bishop Monkton 8 Ings (SE3166) and a 1935 Whitby record (HB). 7 Many records. (i-xii, peak ix). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

5 Subgenus MORPHOCARABUS Géhin, 1885 monilis Fabricius, 1792 consitus sensu auctt. non Panzer, 1809 insularis Born, 1908

Carabus monilis Fabricius, 1792 4 WATSONIAN 3 (4, 8, 7, 14, 0). Notable B. A mainly southern and YORKSHIRE 2 eastern species in Britain, at northern limit of its range in Yorkshire. Few modern records. Before 1 1980 there were two centres of population, in the 0 Leeds/Bradford and Scarborough/Filey districts. 9 There are four post-1980 records, Holmfield (SE02) 8 (MLD), Wheldrake (SE6944) (RC), Low Bramley 7 (SE2074) (RA) and Swinefleet (SE7722) (PK). 6 Recorded from a variety of habitats including carrion 5 traps, in a peat bog, on cultivated lowland and in 4 buildings. (iv-xi, peak vi). 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus ARCHICARABUS Seidlitz, 1887 nemoralis Müller, O.F., 1764

Carabus nemoralis Mueller, 1764 4 WATSONIAN (4, 18, 28, 12, 6). Widely distributed with many 3 YORKSHIRE records, particularly from pitfalls in woodland and 2 heathland, under stones on farmland and in 1 gardens. (i-xii, peak v-vii). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

6 Subgenus AUTOCARABUS Seidlitz, 1887 auratus Linnaeus, 1761

Carabus auratus Linnaeus, 1761 4 WATSONIAN 3 (0, 0, 0, 1, 0) Occasionally introduced (Lindroth, YORKSHIRE 1974), and recently established (Hodge and Jones, 2 1995) in SE England. One record only in VC64, 1 Allerton Bywater iii 1988 (det SLS, specimen in 0 Leeds City Museum). Imported in foreign 9 vegetable produce previously in Britain (Edelsten, 8 1941) (Price, 1968), which may have been the 7 vector in the case of our record. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cancellatus Illiger, 1798 granulatus sensu Stephens, 1827 non Linnaeus, 1758

Carabus cancellatus Illiger, 1798 4 WATSONIAN 3 (0, 0, 0, 0, 1) There is a single 1977 record of this YORKSHIRE species (Coulson and Butterfield, 1979) from Newby 2 Head Moss NZ7983. This record should be viewed 1 with caution unless a voucher specimen can be 0 provided. This species is not included in Luff (2007) 9 on the basis that it is a rare introduction. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

7 Subgenus HEMICARABUS Géhin, 1885 nitens Linnaeus, 1758

Carabus nitens Linnaeus, 1758 4 WATSONIAN 3 (2, 38, 7, 6, 13). Notable B. A flightless species YORKSHIRE 2 generally of upland heaths with Sphagnum. Mainly 1 recorded from the Pennines and North York 0 Moors, with a lowland centre of population at Strensall Common. There is a 1976 record for 9 Thorne Moor (SE71) (Skidmore et al, 1985:) 8 Yorkshire records indicate mainly taken in pitfall 7 traps, but also very numerous at Keld (NY8901) 6 running on road surface in hot sunshine in v1993 5 (RJM). The VC61 records are from Allerthorpe 4 Common (SE7547) (WJF) and Skipwith Common 3 (SE6637) (CDA) on unknown dates. Species 2 apparently declining .(iii-vi, peak v). 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus OREOCARABUS Géhin, 1885 glabratus Paykull, 1790 ssp. lapponicus Born, 1909

Carabus glabratus Paykull, 1790 4 WATSONIAN 3 (0.10, 2, 0, 3) The main centre of population of this YORKSHIRE 2 species of high moorland in the county is in VC62 1 with four post-1980 records. At the southern British 0 limit of its distribution in Yorkshire. Other records 9 emanate from the higher Pennines in VC 63 and 65. Last record Bridestones vi 1985 (DHS). 8 (v-vii, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

8 Subgenus MESOCARABUS Thomson, C.G., 1875 problematicus Herbst, 1786 ssp. feroensis Lapouge, 1910 ssp. harcyniae Sturm, 1815 catenulatus sensu auctt. non Scopoli, 1763 gallicus Géhin, 1885

Carabus problematicus Herbst, 1786 4 WATSONIAN 3 (5, 32, 38, 16,16) In grassland, woodland and YORKSHIRE moorland. Many records widely distributed (i-xii) 2 Most post-1980 records are from VC63, and most 1 of these resulting from pitfall trapping. 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus MEGODONTUS Solier, 1848 violaceus Linnaeus, 1758 ssp. purpurascens Fabricius, 1787 exasperatus sensu auctt. non Duftschmid, 1812 ssp. sollicitans Hartert, 1907 britannicus Born, 1908 browni Deuve, 1999

Carabus violaceus Linnaeus, 1758 4 WATSONIAN 3 (6, 16, 35, 16, 13) Widely distributed in moorland, YORKSHIRE 2 woodland and gardens, with many Yorkshire records 1 (iii-xii). Again, most recent records are from VC63, 0 probably reflecting recording effort. Our records will be referable to ssp. sollicitans . 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

9 Tribe CYCHRINI Laporte, 1834

CYCHRUS Fabricius, 1794 caraboides (Linnaeus, 1758) ssp. rostratus (Linnaeus, 1761) caraboides (Linnaeus, 1758) ssp. rostratus (Linnaeus, 1761)

Cychrus caraboides (Linnaeus, 1758) 4 WATSONIAN 3 (2, 13, 63, 20,9) Generally distributed and YORKSHIRE 2 widespread in woodlands in all VCs, as well as on high moorlands and grasslands at high and low 1 altitudes. Specialised to feed on molluscs. Often in 0 large numbers in pitfall traps. Many records. 9 (i-xii, peak ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe NEBRIINI Laporte, 1834

LEISTUS Frölich, 1799

Subgenus POGONOPHORUS Latreille, 1802 montanus Stephens, 1827 NO RECORDS rufomarginatus (Duftschmid, 1812)

Leistus rufomarginatus (Duftschmid, 1812) 4 WATSONIAN 3 (1, 6, 24, 1, 1). A recent immigrant (Crowson, YORKSHIRE 2 1942), first recorded in Yorkshire in 1980 from 1 Wickersley Wood SK4891 (WAE). The rate at which 0 the species has been recorded in the county has 9 declined markedly since 2000. This decline has been noted nationally. (ii-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

10 spinibarbis (Fabricius, 1775)

Leistus spinibarbis (Fabricius, 1775) 4 WATSONIAN 3 (5, 4, 15, 2, 2) Most of our records are from VC63 YORKSHIRE and post-1980. Widely distributed, particularly from 2 leaf litter and pitfalling in woodland. (iii-x, peak ix). 1 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus LEISTOPHORUS Reitter, 1905 fulvibarbis Dejean, 1826

Leistus fulvibarbis Dejean, 1826 4 WATSONIAN 3 (12, 6, 59, 26, 5) Many records mainly from damp YORKSHIRE 2 woodland, but also in pitfalls in gardens. Few 1 records from the eastern half of the county and 0 probably under-recorded in VC65. (ii-xiii, peak v-vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

11 Subgenus LEISTUS Frölich, 1799 ferrugineus (Linnaeus, 1758) rufescens sensu auctt. partim non (Fabricius, 1775)

Leistus ferrugineus (Linnaeus, 1758) 4 WATSONIAN 3 (16, 9, 58, 17, 0) Many records mainly from the YORKSHIRE centre and south of the county. Mainly from drier 2 situations than the last species. No records yet for 1 VC65. (i-xii, peak vi). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

terminatus (Hellwig in Panzer, 1793) rufescens (Fabricius, 1775) non (Ström, 1768) praeustus (Fabricius, 1792)

Leistus rufescens (Fabricius, 1775) 4 WATSONIAN 3 (11, 24, 55, 37, 7) The most widely distributed YORKSHIRE Leistus in Yorkshire, a species of a wide variety 2 of habitats including woodland, grassland and 1 bogs. Many records. (iv-x, peak v). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

12 NEBRIA Latreille, 1802

Subgenus PARANEBRIA Jeannel, 1937 livida (Linnaeus, 1758) lateralis (Fabricius, 1787) sabulosa (Fabricius, 1787)

Nebria livida (Linnaeus, 1758) 4 WATSONIAN 3 (12, 6, 0, 0, 0) Notable A. Our 18 records are YORKSHIRE 2 entirely coastal, recorded from Withernsea to 1 Redcar, except for a single inland record from a sand pit at Gransmoor TA1159, 1993 (BC). The 0 species is most active during darkness but is 9 usually found in cracks around seepages in clay 8 cliffs in daylight. (v-ix, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus NEBRIA Latreille, 1802 brevicollis (Fabricius, 1792)

Nebria brevicollis (Fabricius, 1792) 4 WATSONIAN 3 (19, 40, 176, 66, 18) One of our commonest YORKSHIRE 2 carabids, generally and widely distributed in all habitat types. Predatory on diptera larvae, the 1 species is very numerous in pitfalls particularly in 0 woodland. The phenology is interesting, with our 9 records showing a marked peak in v, almost no 8 records from vii (summer adult diapause) and 7 another peak in ix (breeding). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

13 salina Fairmaire & Laboulbène, 1854 degenerata Schaufuss, 1862 iberica Oliveira, 1876 klinckowstroemi Mjöberg, 1915

Nebria salina Fairmaire et Laboulbene, 1854 4 WATSONIAN 3 (1, 12, 34, 36, 11) Less frequently recorded than YORKSHIRE brevicollis, this is a species of poorer habitats, 2 particularly upland areas of VCs 62-65. Only one 1 record from VC61 on the YNU database, Houghton 0 Wood SE8837, xi 1974 (RC). There are other 9 records on NBN which require investigation. 8 (ii-xii, peaks v,ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus BOREONEBRIA Jeannel, 1937 nivalis (Paykull, 1790) NO RECORDS

rufescens (Ström, 1768) gyllenhali (Schönherr, 1806) balbii Bonelli, 1810

Nebria rufescens (Strom, 1768) 4 WATSONIAN 3 (1, 18, 31, 26, 11) A distribution map of our YORKSHIRE 2 records agrees with the general habitat of

1 moorland and stream sides in uplands, and stony river margins near the coast in the north of the 0 county, most reports coming from the Pennines 9 and North York Moors. An Askern SE5613 record 8 of vi 1893 (E G Bayford) (Naturalist 1893:288) 7 and a Little Weighton SE9833 (vi 1963, CHW) 6 record are doubtful as both are out of the 5 generally accepted habitat for this species and

4 have not been mapped. (iv-xii, peak vi).

3

2 1 0 9

8 post-1990 records pre-1990 records 7 Map produced by DMAP 14/4/2009 for Yorkshire Naturalists' Union 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

14 EURYNEBRIA Ganglbauer, 1891 complanata (Linnaeus, 1767) NO RECORDS

PELOPHILA Dejean, 1821 borealis (Paykull, 1790) NO RECORDS

Tribe NOTIOPHILINI Motschulsky, 1850

NOTIOPHILUS Duméril, 1806 aesthuans Motschulsky, 1864 aestuans auctt. (misspelling) pusillus Waterhouse, G.R., 1833 non (von Schreber, 1759)

Notiophilus aesthuans Motschulsky, 1864 4 WATSONIAN 3 (1, 0, 0, 1, 6) Notable B. Restricted by its habitat YORKSHIRE 2 to dry, well drained areas. There are eight records, 1 including one from Spurn (Chalk Bank) TA4112, ix 0 1982 (MLD). All the other records are from moorland and under stones and gravels on old 9 mine spoil heaps on the higher Pennines (1972-6, 8 JHF). (iv-viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

aquaticus (Linnaeus, 1758) pusillus (von Schreber, 1759) strigifrons (Baudi, 1864) blacki Edwards, J., 1913

Notiophilus aquaticus (Linnaeus, 1758) 4 WATSONIAN 3 (2, 25, 30, 11, 13) Widely and generally YORKSHIRE 2 distributed. Most post-1980 records are from the south of the county. Recorded from a wide variety 1 of habitats from moorland in the higher Pennines 0 in the west to the dunes of Spurn in the east. 9 (ii-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

15 biguttatus (Fabricius, 1779)

Notiophilus biguttatus (Fabricius, 1779) 4 WATSONIAN 3 (53, 47, 213, 74, 23) Widely distributed and one of YORKSHIRE 2 the most commonly recorded of the Carabidae. 1 Many records particularly from the centre and south 0 of the county. Recorded in Yorkshire from 9 grassland, woodland, gardens and open fields. (ii-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

germinyi Fauvel, 1863 hypocrita sensu auctt. non Putzeys, 1866

Notiophilus germinyi Fauvel, 1863 4 WATSONIAN 3 (2, 22, 6, 18, 12) Although nationally this species YORKSHIRE is generally and evenly distributed, most of our 2 records are in the NE and W of the county, mainly 1 on higher ground. A species of moors, heaths and 0 grasslands, there is a concentration of recent 9 records in the Thorne/Hatfield district of VC63. 8 (iv-viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

16 palustris (Duftschmid, 1812)

Notiophilus palustris (Duftschmid, 1812) 4 WATSONIAN 3 (12, 10, 40, 8, 0) Preferring damp, shaded YORKSHIRE 2 habitats, this species has been recorded in 1 Yorkshire from damp woodland, pond margins, damp roadside mowings and in stable refuse. The 0 few records from VC62 and 65 probably reflects 9 lack of recording effort in these areas. Many 8 recent (post-1995) records are from the south so 7 may be increasing there. (i-xi, peaks iv and vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

quadripunctatus Dejean, 1826 quadriguttatus Fowler, 1886 (error)

Notiophilus quadripunctatus Dejean, 1826 4 WATSONIAN 3 (4, 3, 4, 0, 0) Notable B. An inhabitant of sandy YORKSHIRE 2 areas and thinly vegetated ground, the species has 1 always been very scarce in Yorkshire, and is nearing the northern limit of its national range 0 here. Noted in the county from habitats as diverse 9 as ex-seaweed at Withernsea (GBW) on an 8 unknown date, to Elland Gravel Pits, 1980s (MLD). 7 Recorded from Easington (PK) in 2004 after an 6 interval of 20 years. (vi-ix). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

17 rufipes Curtis, 1829

Notiophilus rufipes Curtis, 1829 4 WATSONIAN 3 (1, 2, 9, 3, 0) A deciduous woodland species YORKSHIRE 2 particularly in ground litter. At the northern limit of 1 its range in our region. Very scarce in Yorkshire. 0 Records peaking in October. (i-x). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

substriatus Waterhouse, G.R., 1833

Notiophilus substriatus Waterhouse, G.R., 1833 4 WATSONIAN 3 (10, 3, 18, 13, 1). Never abundant, but recorded YORKSHIRE 2 from a wide range of habitat types including grass heaps, reservoir and pond edges, in dunes, in 1 ground litter in woodland and in Glyceria beds. 0 (iv-x peaks vi and ix). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

18 Tribe ELAPHRINI Latreille, 1802

BLETHISA Bonelli, 1810 HELOBIUM Leach, 1815 multipunctata (Linnaeus, 1758)

Blethisa multipunctata (Linnaeus, 1758) 4 WATSONIAN 3 (10, 0, 6, 5, 0). Notable B. Widely distributed in YORKSHIRE central Yorkshire but uncommon, in wet sites such 2 as reed-beds and pond edges. Recorded from only 1 eleven Yorkshire sites.A spring breeder. 0 (v-x, peak viii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ELAPHRUS Fabricius, 1775

Subgenus ELAPHRUS Fabricius, 1775 cupreus Duftschmid, 1812

Elaphrus cupreus Duftschmid, 1812 4 WATSONIAN 3 (21, 11, 82, 38, 4). Widespread and common in all YORKSHIRE 2 types of wet situations, particularly on bare mud.. 1 The predominance of VC63 records may reflect collector effort. Probably under-recorded in the 0 north and east. (ii-xi, peaking v-vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

19 uliginosus Fabricius, 1792

Elaphrus uliginosus Fabricius, 1792 4 WATSONIAN 3 (1, 0, 0, 0, 0). Notable B. A single 1990 record from YORKSHIRE 2 Wheldrake Ings SE7044 (MLD). A species of 1 oligotrophic fens and bogs (Luff, 1998). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus TRICHELAPHRUS Semenov, 1926 riparius (Linnaeus, 1758)

Elaphrus riparius (Linnaeus, 1758) 4 WATSONIAN 3 (30, 12, 35, 32, 6). Widespread and common in YORKSHIRE 2 similar habitats to E. cupreus, very often on bare 1 mud by standing water. Many records, the paucity of VC65 entries surely reflecting under-recording. 0 Yorkshire distribution of E. cupreus and E. riparius 9 almost identical, though in Yorkshire riparius is 8 much less commonly recorded in the winter months 7 than cupreus. (v-ix, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 7/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

20 Tribe LORICERINI Bonelli, 1810

LORICERA Latreille, 1802 LOROCERA auctt. (misspelling) pilicornis (Fabricius, 1775) coerulescens sensu auctt. non (Linnaeus, 1758)

Loricera pilicornis (Fabricius, 1775) 4 WATSONIAN 3 (37, 27, 175, 79, 21). Widespread and common YORKSHIRE everywhere. Seems to tolerate all but very dry 2 habitats. Recorded from gardens, woodland, under 1 bark, in compost heaps, on moorland and on coastal 0 dunes. Often very numerous in pitfall traps. 9 (i-xii, peak v-vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe SCARITINI Bonelli, 1810

CLIVINA Latreille, 1802 collaris (Herbst, 1784) contracta (Geoffroy in Fourcroy, 1785)

Clivina collaris (Herbst, 1784) 4 WATSONIAN 3 (6, 7, 5, 16, 2) Records suggest this species tends YORKSHIRE 2 to favour damper situations than C. fossor, often 1 in mud, fine shingle or sand near rivers and ponds, and in Yorkshire tending to the higher 0 ground of the north and west. The two species 9 are sometimes confused in collections. Records 8 also suggest a shorter adult period than fossor. 7 (iv-vii, peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 8/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

21 fossor (Linnaeus, 1758)

Clivina fossor (Linnaeus, 1758) 4 WATSONIAN (35, 16, 55, 50,9). Much more frequent than 3 YORKSHIRE collaris. More widely and generally distributed, in a 2 wide range of habitats including open fields and 1 gardens. Many records. (i-x, peak v). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 8/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

DYSCHIRIUS Bonelli, 1810

Subgenus DYSCHIRIUS Bonelli, 1810 angustatus (Ahrens, 1830)

Dyschirius angustatus (Ahrens, 1830) 4 WATSONIAN (1, 0, 0, 0, 0). RDB3. Very rare nationally. The 3 YORKSHIRE single record on the YNU database is that of W.W. 2 Fowler who reported the species from Bridlington 1 in 1882 (Fowler, 1882). See also Parry, 1975. 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

obscurus (Gyllenhal, 1827) NO RECORDS

22 thoracicus (Rossi, 1790) arenosus Stephens, 1827

Dyschirius thoracicus (Rossi, 1790) 4 WATSONIAN 3 (11, 1, 0, 0, 0) Rarely recorded in Yorkshire and YORKSHIRE 2 only from three sites, Saltburn (record undated 1 and therefore not mapped), Spurn, and repeatedly 0 at Barmston Cliffs from the early 1970s to the present. All records from coastal sand or bare 9 clay. (v-viii, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 8/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus DYSCHIRIODES Jeannel, 1941 aeneus (Dejean, 1825)

Dyschirius aeneus (Dejean, 1825) 4 WATSONIAN 3 (5, 1, 4, 4, 0). Scarce in Yorkshire. Mainly a YORKSHIRE southern insect in Britain and has been confused 2 with D. luedersi in earlier times (before 1915, when 1 that species was described). All the specimens 0 standing as this species in Leed City Museum 9 proved, on closer inspection, to be D. luedersi (MLD, 8 DGH). Near water, often in mud. (v-x, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

extensus Putzeys, 1846 NO RECORDS elongatulus Dawson, 1856

23 globosus (Herbst, 1784) gibbus (Fabricius, 1792)

Dyschirius globosus (Herbst, 1784) 4 WATSONIAN 3 (21, 23, 65, 9, 6). The commonest member of the YORKSHIRE genus, its favoured habitat is bare, rather damp 2 ground. Most of our post-1990 records seem to be 1 from sites at low altitudes, possibly becoming 0 more scarce on higher ground, at least in 9 Yorkshire. (i-ix, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

impunctipennis Dawson, 1854 NO RECORDS

luedersi Wagner, 1915 aeneus sensu auctt. non (Dejean, 1825) ?tristis Stephens, 1827 unicolor sensu auctt. non Motschulsky, 1844

Dyschirius luedersi Wagner, 1915 4 WATSONIAN 3 (14, 3, 15, 2, 0). Much less widespread than D. YORKSHIRE 2 globosus. Favours marshes and damp habitats on 1 heavy soil. The great majority of Yorkshire records 0 appear to be contained in a Lower Derwent/Thorne/Hatfield/Doncaster corridor with 9 a few records from Spurn and the Humber 8 Estuary. (ii-xii, peak vii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

24 nitidus (Dejean, 1825)

Dyschirius nitidus (Dejean, 1825) 4 WATSONIAN 3 (3, 1, 0, 0, 0). Notable A. Yorkshire records are YORKSHIRE 2 entirely coastal. Nationally, mainly in coastal saltmarshes on sand with Bledius (Luff, 1998). 1 Redcar, 1829 (GTR), the 1926 records for 0 “Bridlington” and Barmston Cliffs (CES) may refer 9 to the same event. Spurn TA41, vi 1950 (KS) (Nat 8 1952:172). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 2/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

politus (Dejean, 1825)

Dyschirius politus (Dejean, 1825) 4 WATSONIAN 3 (7, 1, 1, 3, 1). Widely distributed in the east of the YORKSHIRE 2 county. On bare sand or mud, with one population recently established in a suburban garden near 1 Doncaster probably imported with building sand. 0 (iv-viii, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 2/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

25 salinus Schaum, 1843

Dyschirius salinus Schaum, 1843 4 WATSONIAN 3 (9, 0, 1, 0, 0). Well established in Spurn and the YORKSHIRE 2 Stone Creek/Welwick areas of the Humber Estuary. In saltmarsh and muddy banks of watercourses. 1 (iii-viii, peak vi). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe BROSCINI Hope, 1838

BROSCUS Panzer, 1813 cephalotes (Linnaeus, 1758)

Broscus cephalotes (Linnaeus, 1758) 4 WATSONIAN 3 (14, 1, 10, 0, 0). Coastal in dunes in Spurn and YORKSHIRE Bridlington areas, and in sand pits inland, 2 particularly in the Blaxton and Hatfield areas 1 (SE61) and at Pollington SE6120. Also Elland 0 Gravel Pits SE1121. (v-x, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

26 MISCODERA Eschscholtz, 1830 arctica (Paykull, 1798)

Miscodera arctica (Paykull, 1798) 4 WATSONIAN 3 (2, 10, 7, 9, 7). Notable B. Nearly all records are YORKSHIRE pre-1985 from high ground in the county, 2 particularly from moorland and river shingle bars, 1 but also from lowland sites such as Allerthorpe 0 Common SE7547, 19 vi 1931, (WJF); and Strensall 9 SE6559, 1970 (JHF) and 1975 (DGH). Reported 8 twice from the Hatfield Moor area in the 1990s 7 (PS). (v-ix, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 3/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe TRECHINI Bonelli, 1810

PERILEPTUS Schaum, 1860 BLEMUS sensu Laporte, 1840 non Dejean, 1821 areolatus (Creutzer, 1799) NO RECORDS

AEPUS Leach in Samouelle, 1819 AEPOPSIS Jeannel, 1922 marinus (Ström, 1783) fulvescens Samouelle, 1819

Aepus marinus (Stroem, 1783) 4 WATSONIAN 3 (0, 7, 0, 0, 0) All Yorkshire records are from the YORKSHIRE 2 Saltburn/Ravenscar coastal region. Scarce, by 1 virtue of its specialised habitat, i.e. rock crevices in the inter-tidal zone. Recorded first in 1902 0 (MLT) with the latest record 1982 (RK) (iv-xi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 8/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

27 robinii (Laboulbène, 1849) robini auctt. (misspelling)

Aepus robini (Laboulbene, 1849) 4 WATSONIAN 3 (0, 6, 0, 0, 0) Both habitat and area of distribution YORKSHIRE 2 very similar to A. marinus. There are records from Robin Hood‟s Bay NZ5906, 1951 (GBW), 1939 1 (TS), 1986 and 1991 (DGH), Cayton Bay TA0685, 0 1981 (JHF), Cornelian Bay TA0686, 1982 (RSK). 9 (viii-x). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 8/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

TRECHUS Clairville, 1806 BLEMUS sensu Stephens, 1827 non Dejean, 1821

Subgenus EPAPHIUS Leach in Samouelle, 1819 rivularis (Gyllenhal, 1810)

Trechus rivularis (Gyllenhal, 1810) 4 WATSONIAN 3 (0, 0, 0, 5, 0) RDB3 – Rare. Nationally this species YORKSHIRE 2 is recorded from fens and bogs. In Yorkshire only 1 recorded from Askham Bog SE5748 between 1965 and 1996 (EWA, RC, JAO) 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

28 secalis (Paykull, 1790)

Trechus secalis (Paykull, 1790) 4 WATSONIAN 3 (7, 8, 9, 23, 3) Frequently recorded from damp YORKSHIRE 2 grasslands, woods, river banks and ings, both at 1 altitude (e.g. Pockstones Moor SE1160 at 1350 feet) and in lowland (e.g., Fulford Ings SE6049 and 0 Blacktoft SE8524). (v-ix, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus TRECHUS Clairville, 1806 fulvus Dejean, 1831 lapidosus (Dawson, 1849)

Trechus fulvus Dejean, 1831 4 WATSONIAN 3 (0, 1, 0, 0, 0) Notable B. The only record on the YORKSHIRE 2 YNU database is that from Scalby Beck (TA0290) 1 dated August 1909 (TS). Nationally very scarce in freshwater seepages around the coast and under 0 stones on beaches (Luff, 2007). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

29 obtusus Erichson, 1837

Trechus obtusus Erichson, 1837 4 WATSONIAN 3 (39, 29, 77, 33, 14) Many records from all five YORKSHIRE VCs. Ubiquitous and abundant. Recorded habitats 2 on the YNU database include flood refuse, in peat 1 on moorland, in reed-beds, garden compost 0 heaps, grassland, leaf litter in woodland, stable 9 refuse and even in archaelogical deposits of 8 medieval age (Conisbrough SK5198). Can be 7 confused with T. quadristriatus (see Luff 2007:67) 6 and some of our records may be erroneous in this 5 respect. Dissection of males is the best means of determination. Reliable records show a June peak 4 in phenology. (i-xi, peak vi). 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

quadristriatus (Schrank, 1781) minutus (Fabricius, 1781) rubens sensu Clairville, 1806 non (Fabricius, 1792)

Trechus quadristriatus (Schrank, 1781) 4 WATSONIAN 3 (53, 11, 131, 26, 13) Ubiquitous, with many YORKSHIRE records. Recorded from a similar range of habitats 2 to T. obtusus, especially in gardens and post- 1 industrial sites, and in disturbed, dry and open 0 locations. Often recorded at light when conditions 9 are warm and humid. Beware confusion with T. 8 obtusus (see above). The aedeagus is the best 7 character for separation. Reliable records show a 6 later peak in phenology than does obtusus. 5 (i-xii, peak viii). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

30 rubens (Fabricius, 1792) paludosus (Gyllenhal, 1810)

Trechus rubens (F., 1792) 4 WATSONIAN 3 (0, 5, 14, 2, 2) Notable B. Recorded from a wide YORKSHIRE range of habitats including in litter in woods and in 2 pitfalls in moorland; taken in flight in various 1 situations. Our records do not indicate an affinity 0 with coniferous woodland (as suggested in various 9 literature), but regrettably the earlier records are 8 sparse on appended comments. Few recent 7 records with most being from the 1960s and 6 1970s. The species may be declining. (iv-xi, peak 5 vi). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

subnotatus Dejean, 1831

Trechus subnotatus Dejean, 1831 4 WATSONIAN 3 (0, 0, 6, 0, 0) RDB1 - Endangered. A fascinating YORKSHIRE 2 species, and possibly an introduction. Believed to 1 occur otherwise only in Devon and Dublin (Luff, 1998), the species was first noted in Yorkshire at 0 Almondbury SE1615, iii1938 (EFG) (specimens in 9 Doncaster Museum) and then at Dalton 8 Huddersfield SE11, ii 1948 (EWA) (specimen in 7 Tolson Museum). The species was not seen again 6 until found repeatedly in the Huddersfield district 5 from 1983-1986: Dean Wood SE1213, 1983 4 (MLD), Netherton SE2817, xii 1983 (MLD), Spring 3 Wood SE130123, iii 1984 (MLD), and Almondbury SE1615, iii 1986 (MLD) (specimens of the latter in 2 Manchester Museum). A.A.Allen, in his note on the 1 species (Allen, 1950a), was apparently unaware of 0 the EFG and EWA records. Reported from ground 9 litter in woodland situations. (ii-xii). 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

THALASSOPHILUS Wollaston, 1854 TRECHUS sensu Fowler, 1886 partim non Clairville, 1806 longicornis (Sturm, 1825) NO RECORDS

31 BLEMUS Dejean, 1821 LASIOTRECHUS Ganglbauer, 1892 discus (Fabricius, 1792)

Blemus discus (Fabricius, 1792) 4 WATSONIAN 3 (9, 5, 8, 10, 0) Notable B. Widespread but very YORKSHIRE infrequent. Near water, often on mud, also in 2 pitfall traps in a rural garden. Flies strongly 1 especially in hot weather. No reports since 1998, 0 with most records from the 1970s and 1980s. 9 (vi-x, peak ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

TRECHOBLEMUS Ganglbauer, 1891 micros (Herbst, 1784)

Trechoblemus micros (Herbst, 1784) 4 WATSONIAN 3 (2, 6, 6, 8, 4) Widespread and usually recorded YORKSHIRE 2 near water; Yorkshire records reported from flood 1 refuse, in debris below a weir, and in pitfalls in a rural garden in very damp ground. Only five recent 0 (post-1990) records. (ii-xii, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

32 Tribe BEMBIDIINI Stephens, 1827

TACHYS Dejean, 1821

Subgenus PARATACHYS Casey, 1918 EOTACHYS Jeannel, 1941 bistriatus (Duftschmid, 1812) ?pallidulus (Antoine, 1943) non (Ménétriés, 1846)

Tachys bistriatus (Duftschmid, 1812) 4 WATSONIAN 3 (1, 0, 0, 0, 0) Notable B. There is one record only YORKSHIRE 2 on the YNU database: Bubwith SE7237, 1930 1 (CES), one specimen taken from flood refuse (see Nat 1930:392). Nationally the habitat is damp 0 ground, clay or sand, by freshwater (Luff, 2007). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 10/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

micros (Fischer von Waldheim, 1828) NO RECORDS gregarius Chaudoir, 1846

obtusiusculus (Jeannel, 1941) NO RECORDS piceus Edmonds, 1934 non Dalla Torre, 1877 edmondsi Moore, 1956

Subgenus TACHYS Dejean, 1821 scutellaris Stephens, 1828 SEE APPENDIX A

33 ELAPHROPUS Motschulsky, 1839 TACHYS sensu auctt. partim non Dejean, 1821 parvulus (Dejean, 1831)

Elaphropus parvulus (Dejean, 1831) 4 WATSONIAN 3 (0, 0, 10, 0, 0) Notable B. A carabid of sandy soils, YORKSHIRE 2 there are now two reported Yorkshire sites. Found 1 regularly in a suburban garden at Sprotbrough 0 SE553033 between 1994 and 2007 (RJM), in the 9 joints between concrete paving blocks, and almost certainly introduced in building sand, source 8 unknown. Also in a suburban garden in Kettlethorpe 7 SE328166, v 2008 (DGH). (v-vii). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

walkerianus (Sharp, 1913) NO RECORDS

34 ASAPHIDION des Gozis, 1886 TACHYPUS sensu auctt. non Weber, 1801

This genus has given the YNU recorder many problems in interpreting the records. The definitive paper on the revision of the British members of this much confused genus has been published in the entomological literature (Speight et al, 1986). Records for „flavipes‟, „flavipes agg.‟ and „pallipes‟ exist in the YNU database. Many records for this genus previous to 1986 have to be discounted unless vouchers are forthcoming. In the notes which follow the accuracy of records must placed on the expertise of the recording coleopterists (ES, RJM, DGH, MLD, JC, WRD)

curtum (Heyden, 1870) flavipes sensu auctt. Brit. partim non (Linnaeus,1761)

Asaphidion curtum (Heyden, 1870) 4 WATSONIAN (10, 2, 18, 12, 0) Widely distributed in ground 3 YORKSHIRE litter in woodland, farmland field edges, and 2 sometimes around pond edges. Some records for 1 „curtum‟ predate 1986. The records do not make 0 clear whether these records are of specimens that 9 have been determined as curtum after 1986. All 8 records pre-1986 should therefore be considered 7 invalid unless vouchers are forthcoming. To date, 6 this is the most commonly recorded Asaphidion. Recent reliable records include Nostell Fishponds 5 SE393157, 4 v 2002, (DGH), Sharow Mires 4 SE330709 6 viii 2002 (RJM), Abbeydale SK325818, 3 19 vi 2006 (EJS), Elstronwick TA2332, 13 x 2006 2 (WRD) and Withernsea TA3428, 5 iv 2007 (FEK) 1 (i-xii, peak v). 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

flavipes (Linnaeus, 1761)

Asaphidion flavipes (Linnaeus, 1761) 4 WATSONIAN 3 (1, 1, 1, 2, 0) Widely distributed with few records YORKSHIRE that can be confirmed: Cawood SE576374, 1989 2 (KP det MLD); Barlow SE6428, 19 v 1952 (BK det 1 RJM); High Batts SE3076, vi 1992 (MLD) and 1993 0 (JBJ); Duncombe Park SE6082, vi 1993 (MLD); 9 Eggborough Power Station SE5724, xi 1994 8 (RJM); and North Ferriby SE9826, viii 1999 (RGB). 7 Habitats recorded include pitfalls in a garden, 6 ground litter in woodland, roadside grass piles and 5 in seepages at the base of low clay cliffs on the Humber estuary. 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

[‘flavipes agg.’ – these records (of which there are 15) cannot be assigned to a particular species, because they were recorded pre-1980 or are based on indeterminate females. Therefore the records have not been mapped]

35 pallipes (Duftschmid, 1812)

Asaphidion pallipes (Duftschmid, 1812) 4 WATSONIAN (0, 2, 0, 0, 0) Notable B. There are two records in 3 YORKSHIRE the YNU database: Mulgrave Woods NZ8513, 1980 2 (Invertebrate Site register ed. RK), and Cropton 1 Banks Wood SE7487, 1985 (KP). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

stierlini (Heyden, 1880) flavipes sensu auctt. Brit. partim non (Linnaeus, 1761)

Asaphidion stierlini (Heyden, 1880) 4 WATSONIAN 3 (9, 0, 6, 3, 0) Widely distributed in the YORKSHIRE 2 southeastern part of the county. Reported habitats 1 on the database include urban brownfield/demolition sites, farmland, in oak leaf 0 litter on peat moors, in woodland, in piles of 9 woodchips, and in sweepings from roadside 8 verges (i.e. generally more open situations). 7 Recent reliable records include the following: 6 Stainforth SE653125, 4 viii 2008 (RJM); Loversall 5 SK5799, 28 viii 2004 (RJM); Burton Agnes 4 TA1063, 11 x 2003 (WRD); Fountains Abbey 3 SE2868, 24 viii 2000 (RJM). (ii-xi, peak ix). 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

36 OCYS Stephens, 1828 BEMBIDION sensu auctt. partim non Latreille, 1802 harpaloides (Audinet-Serville, 1821) rufescens (Guérin-Méneville, 1823)

Ocys harpaloides (Audinet-Serville, 1821) 4 WATSONIAN 3 (20, 16, 76, 30, 13) Widely distributed and YORKSHIRE 2 frequently recorded from woodland. Under bark of 1 deciduous trees, and occasionally from leaf litter around logs, beating faggots and in fungi in tree 0 roots. (i-xii, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

quinquestriatus (Gyllenhal, 1810)

Ocys quinquestriatus (Gyllenhal, 1810) 4 WATSONIAN (0, 1, 10, 8, 0) Always very scarce in Yorkshire, 3 YORKSHIRE this enigmatic species is widely distributed in the 2 SW quarter of the county, apart from a Whitby 1 NZ8199, 1935 record (HB). Regarding habitat, 0 there seems to be a predeliction for walls, both 9 brick and of the dry stone type. A.A.Allen (pers 8 comm to RJM) suggested that the connection 7 could well be the presence of the runs of small 6 mammals in or under the walls. On both occasions when RJM has found the species the habitat was 5 at the base of dry stone walls in woodland. 4 Another recent record came from a wall by a 3 reservoir (MLD). A record from Thorne Moor came 2 from rubble under a demolished building (BCE). 1 (iii-viii, peak vii). 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

37 CILLENUS Leach in Samouelle, 1819 BEMBIDION sensu auctt. partim non Latreille, 1802 lateralis Samouelle, 1819 NO RECORDS

BRACTEON Bedel, 1879 BEMBIDION sensu auctt. partim non Latreille, 1802 CHRYSOBRACTEON Netolitzky, 1914 argenteolum (Ahrens, 1812) NO RECORDS

litorale (Olivier, 1790) littorale (Olivier, 1791) paludosum (Panzer, 1794)

Bracteon litorale (Olivier, 1790) 4 WATSONIAN 3 (2, 12, 1, 6, 7) Notable B. A Bembidion of beds of YORKSHIRE 2 fine shingle and sand on river banks. Widely 1 distributed mainly in the north of the county. 0 Sometimes abundant when found. As B. 9 tetracolum Say was once known as littorale (Olivier) some early records of the latter may be 8 erroneous. (v-xi, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

38 BEMBIDION Latreille, 1802 BEMBIDIUM auctt. (misspelling) Subgenus NEJA Motschulsky, 1864 nigricorne Gyllenhal, 1827

Bembidion nigricorne Gyllenhal, 1827 4 WATSONIAN 3 (2, 23, 2, 5, 0) Notable B. An insect mainly of YORKSHIRE 2 upland moors and heaths where Calluna abounds. Most records are from the North York Moors with 1 a few from the Pennines south to Sheffield. There 0 are also records for lowland heaths (Skipwith 9 SE6437 and Strensall SE6559). Very local and 8 usually scarce, but can be abundant where it 7 occurs (especially in pitfall traps). (iii-x, peaks in iv 6 and ix). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 16/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus METALLINA Motschulsky, 1850 lampros (Herbst, 1784) celere (Fabricius, 1792)

Bembidion lampros (Herbst, 1784) 4 WATSONIAN 3 (60, 44, 221, 66, 15) Common and widespread. YORKSHIRE Our most commonly recorded Bembidion. Easily 2 confused with B. properans, the habitat 1 requirements of both species apparently 0 indistinguishable. Some of our records for lampros 9 will be referable to properans and vice versa. Care 8 should be taken with identification. (i-xii, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

39 properans (Stephens, 1828) velox Erichson, 1837 non (Linnaeus, 1761) coeruleotinctum Reitter, 1908 cyaneotinctum Sharp, 1913 caeruleipenne Saunders, 1936

Bembidion properans (Stephens, 1828) 4 WATSONIAN (12, 23, 42, 12, 2) Far less frequently reported 3 YORKSHIRE and apparently not as widespread as B. lampros, 2 with fewer records from the far east and far west 1 of our recording area. The comments above (B. 0 lampros) apply here also. (i-xii, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PRINCIDIUM Motschulsky, 1864 punctulatum Drapiez, 1821

Bembidion punctulatum Drapiez, 1821 4 WATSONIAN 3 (0, 15, 3, 18, 10) Widespread in the northern and YORKSHIRE 2 western areas of the recording area. Often 1 abundant when found. Associated with beds of 0 fine shingle by (mainly upland) rivers and streams, and with gravelly sides of non-flowing bodies of 9 freshwater. There are lowland records (see Luff, 8 1998:71), for example Hatfield Moor SE6804, 7 v2005, (RJM) (DAL). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

40 Subgenus ACTEDIUM Motschulsky, 1864 pallidipenne (Illiger, 1802) NO RECORDS ruficolle (Illiger, 1801) non (Panzer, 1796)

Subgenus TESTEDIUM Motschulsky, 1864 bipunctatum (Linnaeus, 1761)

Bembidion bipunctatum (Linnaeus, 1761) 4 WATSONIAN 3 (1, 1, 1, 2, 1) Notable B. Six records on the YNU YORKSHIRE 2 database for this species of sand and gravel 1 beside still and flowing freshwater. Hull TA02 0 (estimated grid ref) pre-1900, (CWR), Eston NZ5518, vi 1911 (MLT), Aysgarth SE0188, v 1919, 9 (AEW), Ryhill SE3914, v 1919, (WJF), Pen-y- 8 ghent, v 1942, (WDH), Ribblehead Quarry 7 SD766787, v 2006(TJB teste MLD). (v-vi, peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 27/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus EUPETEDROMUS Netolitzky, 1911 dentellum (Thunberg, 1787) flammulatum (Clairville, 1806)

Bembidion dentellum (Thunberg, 1787) 4 WATSONIAN 3 (31, 5, 67, 31, 9) Widespread and frequent except YORKSHIRE in the far west and on the east coast. Recorded 2 from flood refuse, by land drains, marshes, pond 1 margins, reed-beds, river banks and wet pastures. 0 Many records. (i-x, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

41 Subgenus NOTAPHUS Dejean, 1821 obliquum Sturm, 1825

Bembidion obliquum Sturm, 1825 4 WATSONIAN 3 (6, 6, 47, 11, 5) Notable B. Widespread and YORKSHIRE 2 sometimes abundant at sites in the south and centre of the county. Recorded from margins of 1 drains, ponds and lagoons particularly on acid 0 soils in lowland areas. (ii-xi, peak vii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

semipunctatum (Donovan, 1806) NO RECORDS adustum Schaum, 1860

varium (Olivier, 1795) ustulatum sensu Sturm, 1825 non (Linnaeus, 1758) nebulosum (Stephens, 1828)

Bembidion varium (Olivier, 1795) 4 WATSONIAN (40, 1, 19, 0, 1) Restricted mainly to the south- 3 YORKSHIRE east and south of the county, and found near 2 water, on mud at edges of rivers, lakes, reservoirs 1 and dikes, in reedbeds and in saltmarshes. Often 0 reported as very numerous when located. (i-xi, 9 peaks vi and ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

42 Subgenus NOTAPHEMPHANES Netolitzky, 1920 NOTHAPHEMPHANES auctt. (misspelling) ephippium (Marsham, 1802) NO RECORDS

Subgenus PLATAPHUS Motschulsky, 1864 prasinum (Duftschmid, 1812)

Bembidion prasinum (Duftschmid, 1812) 4 WATSONIAN 3 (0, 3, 2, 11, 9) A species of northern and western YORKSHIRE 2 Britain. Our records show an entirely upland 1 distribution, the only exception to this being a specimen, or fragments, of this species recorded 0 (in 1976) from archaeological moat deposits of 9 medieval age at Cowick (SE6520). As the species 8 is one of shingle by usually swiftly flowing rivers, 7 the Cowick situation is most odd. The 6 identification here may be erroneous, especially if 5 the identifiers were unaware of the habitat of the 4 insect and did not have a complete specimen to 3 work with. (i-vii, peak v). 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus TRICHOPLATAPHUS Netolitzky, 1914 BLEPHAROPLATAPHUS Netolitzky, 1920 virens Gyllenhal, 1827 NO RECORDS

Subgenus BEMBIDIONETOLITZKYA Strand, E., 1929 DANIELA Netolitzky, 1910 non Koch, 1891 atrocaeruleum (Stephens, 1828) atrocoeruleum auctt. (misspelling)

Bembidion atrocaeruleum (Stephens, 1828) 4 WATSONIAN 3 (1, 18, 1, 26, 21) The distribution of this species is YORKSHIRE 2 similar to the last, the habitat being, again, 1 exposed shingle by, usually, upland rivers and 0 streams. There are recent records for Barmston Cliffs TA1758, where the beetle was found in 9 coarse sand by seepages on clay cliffs. This 8 species may be confused with B. tibiale and B. 7 geniculatum. Plenty of recent records. 6 (iii-xii, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

43 caeruleum Audinet-Serville, 1826 33 NO RECORDS coeruleum auctt. (misspelling)

geniculatum Heer, 1837/8 redtenbacheri Daniel, K., 1902

Bembidion geniculatum Heer, 1838 4 WATSONIAN (6, 7, 37, 11, 6) Sometimes confused with B. 3 YORKSHIRE tibiale and B. geniculatum, this, as in other 2 members of the subgenus, is a species of sand 1 and gravel by rivers. Our records are 0 predominantly upland, with some older records 9 from coastal sites such as Whitby NZ8911 and 8 Robin Hood‟s Bay NZ9506, and recent ones from 7 Holmpton TA3823. Few recent records. 6 (ii-x, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

tibiale (Duftschmid, 1812)

Bembidion tibiale (Duftschmid, 1812) 4 WATSONIAN 3 (0, 18, 8, 29, 16) An inhabitant of riverside gravels YORKSHIRE and shingles, all our records are from the high 2 ground in the county, being widely distributed 1 especially in the northwest. Luff (1998) shows a 0 clutch of records from the northeast coast of VC62 9 which do not appear on our records. Field workers 8 should beware of confusion with the other two 7 members of the subgenus, viz. atrocaeruleum and 6 geniculatum. Often abundant where its habitat 5 occurs. (iv-viii, peak v). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

44 Subgenus OCYDROMUS Clairville, 1806 PERYPHUS Dejean, 1821 bruxellense Wesmael, 1835 rupestre sensu auctt. non (Linnaeus, 1767)

Bembidion bruxellense Wesmael, 1835 4 WATSONIAN 3 (4, 13, 25, 10, 1) Rather similar to to the common YORKSHIRE 2 tetracolum, this is a species of, generally sandy 1 margins of rivers and lakes, but also recorded from bare peat on Thorne and Hatfield Moors SE61/71, 0 from gravel pits and by water in damp woodland 9 and grassland. Very few recent records. (iii-x). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 24/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

bualei Jacquelin du Val, 1852 andreae sensu auctt. non (Fabricius, 1787) cruciatum sensu auctt. non Dejean, 1831 ssp. anglicanum Sharp, 1869 ssp. polonicum Müller, J., 1930

Bembidion bualei Jacquelin du Val, 1852 4 WATSONIAN 3 (14, 9, 2, 0, 2) The majority of our records are YORKSHIRE 2 coastal and come from sites where there are seepages or areas of wet sand at the bases of clay 1 cliffs. This species was called andreae in earlier 0 checklists and in Luff (1998), but see comments in 9 Luff (2007) on this species. There are, apparently 8 two subspecies, anglicanum and polonicum, in 7 Britain. In the absence of specimens the position 6 of our records in relation to these subspecies 5 cannot be clarified, but there are two distinct 4 areas of population in the county, i.e. records from the coastal strip and those well inland on 3 upland sites. (iv-ix, peak vi). 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

45 decorum (Zenker in Panzer, 1800)

Bembidion decorum (Zenker in Panzer, 1800) 4 WATSONIAN 3 (0, 17, 2, 22, 10) With a distribution which is YORKSHIRE northern and western in Britain, our records follow 2 this pattern, with most records being from the 1 high ground in the north and west. A species of 0 shingle by, usually, fairly swift water courses. 9 There is a Fishlake SE6513, 1976 record from the 8 bank of the Stainforth and Keadby Canal which 7 would require confirmation. This record for the 6 time being remains on the database. (iv-ix, peak 5 vi). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

deletum Audinet-Serville, 1821 nitidulum (Marsham, 1802) non (Schrank, 1781) dalmatinum var. latinum sensu MacKechnie-Jarvis, 1932 non Netolitzky, 1911

Bembidion deletum Audinet-Serville, 1821 4 WATSONIAN 3 (12, 6, 46, 19, 2) This species was the B. nitidulum YORKSHIRE 2 (Marsham) of older checklists. It is widespread in 1 Yorkshire, particularly in the south and west. Apart 0 from strictly coastal sites, there is a marked absence of records from the higher and drier eastern vice 9 counties. The large clutch of records from the 8 northeast in Luff (1998) does not appear in occur in 7 our database. Our records include habitats such as 6 by seepages at the base of clay cliffs, in flood 5 refuse, in decaying vegetation at edges of ponds, 4 lakes, rivers and dikes, and one or two adventitious 3 examples found in buildings. (ii-x, peak v-vi). 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

46 femoratum Sturm, 1825

Bembidion femoratum Sturm, 1825 4 WATSONIAN 3 (6, 7, 14, 10, 1) Widely and rather thinly YORKSHIRE distributed throughout the county, and from a 2 wide variety of damp habitats including on sand at 1 base of clay cliffs, in leaf litter by rivers, on 0 brownfield sites, in gravel pits and by rivers and 9 ponds. Few recent records. (i-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

fluviatile Dejean, 1831

Bembidion fluviatile Dejean, 1831 4 WATSONIAN 3 (0, 0, 0, 2, 0) Very rare in Yorkshire, with only two YORKSHIRE 2 records on the database: Ben Rhydding SE1347, 2 vi 1973 (RC) on sandy river bank, Farnham 1 Gravel Pits SE3560, 1 vii 1976 (RC) no other data. 0 Luff (1998) indicates two further records for SE75 9 and SE36 which require investigation via NBN. 8 Typical habitat is stated to be on river banks (Luff, 7 1998 and Luff, 2007). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

47 lunatum (Duftschmid, 1812)

Bembidion lunatum (Duftschmid, 1812) 4 WATSONIAN 3 (11, 4, 12, 6, 0) With a rather localised YORKSHIRE distribution in Yorkshire, this species is rather 2 uncommon with few recent records. Typical 1 habitat is on lowland river banks which is borne 0 out by our records. Most are from the lower 9 reaches of the Ouse/Aire/Derwent and from the 8 Humber estuary. Some records are from further 7 inland, among them Helmsley area SE6480, 6 13 vi 2000 (DGH). Very few recent records. (v-xi, 5 peak vi). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

maritimum (Stephens, 1835) concinnum sensu auctt. non (Stephens, 1828) dorsuarium Bedel, 1879

Bembidion maritimum (Stephens, 1835) 4 WATSONIAN 3 (7, 2, 4, 0, 0) Very scarce in Yorkshire, with only YORKSHIRE 13 records, none of them recent. Nationally a 2 coastal and estuarine species, our records are of 1 single specimens occurring rarely, in fine debris 0 and sediment at edges of, usually moving, water. 9 There is an odd record for Allerthorpe Common 8 SE7547, 16 iv 1927 (WDH) which is perhaps a 7 little out of normal habitat. The specimen, if it 6 exists, has not yet been traced. (iv-ix,peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

48 monticola Sturm, 1825

Bembidion monticola Sturm, 1825 4 WATSONIAN 3 (0, 4, 2, 8, 4) Notable B. Our records are mainly YORKSHIRE upland ones, with records from the stony banks of 2 rivers and from riverside shingle. The Askern 1 SE5613, 15 vi 1893 (EGB) record is probably an 0 error as the generally accepted habitat does not 9 occur there. Scarce with very few records, with 8 only two being recent. (iv-ix, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

saxatile Gyllenhal, 1827 vectense Fowler, 1886

Bembidion saxatile Gyllenhal, 1827 4 WATSONIAN (2, 9, 2, 1, 1) Notable B. Widely distributed but 3 YORKSHIRE rather scarce. Coastal records from Cornelian Bay 2 TA0686, 1982 (RK) and Cayton Bay TA063850, viii 1 1999 (RGB) came from seepages at the bases of 0 soft cliffs. Other habitats recorded include shingle 9 bars by rivers. There is a record for Spurn (Chalk 8 Bank) TA412124, ix 1982 (MLD) and another for 7 Middleton Wood SE1248, viii 2002 (PK) in shingle 6 by R. Wharfe. (v-ix, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

49 stephensii Crotch, 1866 stephensi auctt. (misspelling) affine (Stephens, 1835) non Say, 1825

Bembidion stephensi Crotch, 1866 4 WATSONIAN 3 (10, 9, 5, 4, 0) Most of our 28 records are post- YORKSHIRE 2 1980. First recorded at Forge Valley SE98, 1913 1 (ECH), our recent records are mainly coastal. Well established in coastal sites particularly in seepages 0 at the bases of clay cliffs. Records from inland 9 sites also indicate a preference for bare clayey 8 ground near water. (iii-ix, peak vii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

testaceum (Duftschmid, 1812)

Bembidion testaceum (Duftschmid, 1812) 4 WATSONIAN 3 (0, 1, 0, 2, 2) Notable B. Extremely scarce in the YORKSHIRE 2 county. The records we have are: Sedburgh 1 SD6693, v 1894 (WFB), (see Nat:1894:225); 0 Forge Valley SE98, 1913 (ECH), (see Nat 9 1914:90); Farnham Gravel Pits SE3560, vii 1976, ix 1976 (RC); Langton Bridge SE295961, 21 v 8 2002 (RJM teste MLL). In shingle by rivers and 7 standing water. (v-ix). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

50 tetracolum Say, 1825 ustulatum sensu auctt. non (Linnaeus, 1758) littorale sensu auctt. non (Olivier, 1791)

Bembidion tetracolum Say, 1823 4 WATSONIAN 3 (26, 26, 90, 47, 33) Widespread and very YORKSHIRE 2 common. The commonest Bembidion nationally 1 and in Yorkshire. Recorded from many habitat types including gardens, arable land, in leaf litter 0 in damp woodland, by rivers and streams and in 9 reed-beds,in light traps, and in strand line refuse 8 on the shore. At all elevations from Pen-y-ghent to 7 Spurn. Many records (i-xi, peak v-vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus NEPHA Motschulsky, 1864 illigeri Netolitzky, 1914 quadriguttatum sensu auctt. Brit. non (Illiger, 1798) nec (Fabricius, 1775) tetragrammum sensu auctt. non Chaudoir, 1846 genei sensu auctt. Brit. non Küster, 1847

Bembidion illigeri Netolitz ky, 1914 4 WATSONIAN (27, 9, 24, 10, 2) A species of southern and eastern 3 YORKSHIRE Britain, the Yorkshire records are widely distributed in 2 the lowlands and on the coast. Its typical habitat is 1 open situations by water, on bare clay or sand, and 0 especially active in warm conditions. This is the B. 9 genei ssp. illigeri of older checklists and Lindroth 8 (1974). There are more post-1990 records than 7 before that date, so the species may be more 6 frequent than in earlier times, possibly as a result of warmer winters in our latitudes. (ii-xii, peaks vi and 5 viii). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

51 Subgenus SINECHOSTICTUS Motschulsky, 1864 SYNECHOSTICTUS auctt. (misspelling) stomoides Dejean, 1831 atroviolaceum sensu auctt. non Dufour, 1820

Bembidion stomoides Dejean, 1831 4 WATSONIAN (1, 2, 0, 12, 4) Notable B. Rather scarce and not 3 YORKSHIRE recorded since 1988: Semerwater SD911862, viii 2 1988 (RJM and PK). Exclusively by rivers and 1 streams (usually with shingle margins) in mainly 0 upland locations. (iii-xii, peak iv). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PSEUDOLIMNAEUM Kraatz, 1888 inustum Jacquelin du Val, 1857 Subgenus LYMNAEUM Stephens, 1828 NO RECORDS nigropiceum (Marsham, 1802) NO RECORDS

Subgenus SEMICAMPA Netolitzky, 1910 gilvipes Sturm, 1825

Bembidion gilvipes Sturm, 1825 4 WATSONIAN 3 (41, 16, 18, 14, 1) Notable B. Most records are YORKSHIRE 2 from the centre of the county, and is recorded from reed-beds, in ground litter by drains, in damp 1 woodland, riverside ings, fens and gravel pits. The 0 Yorkshire records indicate occurrence away from 9 the high ground and chalk/limestone areas. (i-xi, 8 peak iv). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 17/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

52 schuppelii Dejean, 1831 SEE APPENDIX A schueppeli auctt. (misspelling)

Subgenus DIPLOCAMPA Bedel, 1896 assimile Gyllenhal, 1810

Bembidion assimile Gyllenhal, 1810 4 WATSONIAN 3 (30, 2, 69, 1, 0) Many records, all but a very few YORKSHIRE 2 being from the far south and south-east of the 1 county. Recorded in Yorkshire from lowland marshes, fens and reed-beds, and saltmarshes in 0 the Humber estuary. This species has a south- 9 easterly distribution in England so in Yorkshire is 8 at the north edge of its range. (ii-xi, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

clarkii (Dawson, 1849) clarki auctt. (misspelling)

Bembidion clarkii (Dawson, 1849) 4 WATSONIAN 3 (14, 10, 39, 20, 0) Notable B. Many records. YORKSHIRE 2 Reported from wet sites with plenty of shade, pond edges, wet woodlands, willow carr and 1 shaded fens, nearly always inland with only three 0 records either coastal or estuarine. At the northern 9 edge of its range here. Our records are mainly 8 from the centre and south of the county. (i-xi, 7 peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

53 fumigatum (Duftschmid, 1812)

Bembidion fumigatum (Duftschmid, 1812) 4 WATSONIAN 3 (23, 1, 47, 2, 0) Notable B. Many records from YORKSHIRE 2 lowland marshes, pond edges, river ings, reed- beds and estuarine saltmarshes. Mainly coastal in 1 southern England but occurring inland at the north 0 end of its geographical limit, as in Yorkshire (Luff, 9 1998). (i-xii, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus EMPHANES Motschulsky, 1850 minimum (Fabricius, 1792)

Bembidion minimum (Fabricius, 1792) 4 WATSONIAN 3 (38, 1, 1, 0, 0) Almost exclusively a VC61 species, YORKSHIRE 2 and most records are either estuarine or coastal. 1 Recorded habitats include saltmarshes, strandline debris, in ground litter in riverside reed-beds, and 0 in wet woodland sites near rivers. (i-xii, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 4/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

54 normannum Dejean, 1831

Bembidion normannum Dejean, 1831 4 WATSONIAN 3 (6, 0, 0, 0, 0) Very scarce in Yorkshire. Exclusively YORKSHIRE 2 coastal in England, and mainly in the east and 1 south. Our six records are Welwick TA3421, 9 vi 0 1946 and 18 viii 1949 (WDH , SS); Spurn TA4115, 9 1950 (2 records), 1951 (1 record) (all SS); Spurn TA4115, 23 vii 1963 (RCW). Habitats recorded 8 were pond and drain margins. (vi-viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus BEMBIDION Latreille, 1802 LOPHA Dejean, 1821 non Bolten, 1798 humerale Sturm, 1825

Bembidion humerale Sturm, 1825 4 WATSONIAN (0, 0, 23, 0, 0) RDB - Endangered. In Britain this 3 YORKSHIRE species is only known from Thorne and Hatfield 2 Moors SE60, SE70, SE71. First discovered in 1975 1 (Crossley, Norris, 1975). Located frequently since 0 that date, and widely distributed across Thorne 9 Moor with a few records from Hatfield Moor. 8 Usually on bare peat by water, and may be quite 7 numerous where it occurs. (iv-x, peak viii). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

55 quadrimaculatum (Linnaeus, 1761) quadriguttatum (Fabricius, 1775)

Bembidion quadrimaculatum (Linnaeus, 1761) 4 WATSONIAN 3 (28, 7, 48, 31, 1) Many records. Widely distributed. YORKSHIRE 2 There are no records on the YNU database from the 1 far north and north-east of the county. Recorded 0 habitats include open and rather dry localities such as peatmoors, heaths, chalk pits, arable land, sunny 9 situations at woodland edges and in pitfalls at 8 sparsely vegetated pond margins. (i-xii, peak iv-v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

quadripustulatum Audinet-Serville, 1821 quadriguttatum sensu (Olivier, 1795) non (Fabricius, 1775) antiquorum Crotch, 1871

Bembidion quadripustulatum Audinet-Serville, 1821 4 WATSONIAN 3 (0, 0, 1, 0, 0) A single confirmed record exists on YORKSHIRE the YNU database: Templeborough SK411920, 22 2 vii 2006 (RJM teste MLL), specimen sieved from 1 ground litter in Phragmites beds. Another record 0 from Potteric Carr from the 1960s cannot be 9 verified. The supposed collector (PS) has no 8 knowledge of it and no specimen exists in 7 Doncaster Museum. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

56 Subgenus TREPANEDORIS Netolitzky, 1918 doris (Panzer, 1796)

Bembidion doris (Panz er, 1796) 4 WATSONIAN 3 (16, 3, 16, 20, 1) Widely distributed with many YORKSHIRE 2 records, mainly from the central part of our 1 recording region. This very distinctive Bembidion 0 is recorded from constantly wet areas such as pond and lake edges and in marshes, in reed-beds 9 and in the stems of Typha, in ground litter on 8 riverside ings, and in willow and alder carrs. (i-x, 7 peak v-vii). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus TREPANES Motschulsky, 1864 LEJA Dejean, 1821 28 articulatum (Panzer, 1795)

Bembidion articulatum (Panz er, 1795) 4 WATSONIAN 3 (6, 0, 8, 1, 0) Very scarce in Yorkshire. Restricted YORKSHIRE to SE England, this species is at the northern limit 2 of its geographical range in southern Yorkshire. 1 Bare mud near water is typical of its habitat, with 0 Yorkshire records from peat moors, pond margins, 9 Phragmites marshes and in flood debris at the 8 edge of a saltmarsh. (v-x, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 4/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

octomaculatum (Goeze, 1777) NO RECORDS sturmii (Panzer, 1804) sturmi auctt. (misspelling)

57 Subgenus PHYLA Motschulsky, 1844 PHILA Motschulsky, 1846 36 obtusum Audinet-Serville, 1821

Bembidion obtusum Audinet-Serville, 1821 4 WATSONIAN 3 (43, 14, 61, 24, 2) Widely distributed with many YORKSHIRE 2 records, except for the (under-recorded) far north- west of our recording area. Recorded habitats 1 include open ground, cultivated arable land and 0 pastures, in seepages at the base of soft cliffs, in 9 flood refuse, in leaf litter, in pitfalls in limestone 8 woodlands, in sand dunes, haystack bottoms and 7 in suction samples from ruderal vegetation on 6 disturbed ground. (i-xii, peaks v and viii). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PHILOCHTHUS Stephens, 1828 PHILOCTHUS auctt. (misspelling) aeneum Germar, 1824

Bembidion aeneum Germar, 1824 4 WATSONIAN 3 (50, 19, 94, 46, 4) Widely distributed with many YORKSHIRE records except for the dry chalk areas of VC61 and 2 the high land of VC64 and VC65. Recorded 1 habitats include estuarine and riverside mud flats, 0 in grass tussocks in riverside ings, in leaf litter in 9 damp woodland, in reed-bed litter and in rotting 8 grass heaps in damp farmland. (i-xii, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

58 biguttatum (Fabricius, 1779)

Bembidion biguttatum (Fabricius, 1779) 4 WATSONIAN 3 (56, 19, 66, 37, 5) Many records, mainly from the YORKSHIRE 2 cemtre of the county, and reported from river and 1 pond margins, flood refuse on riverside ings, in mud 0 in rather open reed-beds, in muddy pastureland, by land drains and in willow carr. (i-xi, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

guttula (Fabricius, 1792)

Bembidion guttula (Fabricius, 1792) 4 WATSONIAN 3 (48, 26, 164, 68, 7) Widely distributed with many YORKSHIRE records. The commonest member of the 2 subgenus, particularly in the south of the region. 1 Found near water in a wide variety of habitats, 0 including flood refuse, under stones in gardens, in 9 leaf litter in damp woodland, in grass tussocks, on 8 river banks, in Glyceria swamps, and in estuarine 7 strandline refuse. (i-xii, peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

59 iricolor Bedel, 1879 riparium sensu Fowler, 1886 partim non (Olivier, 1795)

Bembidion iricolor Bedel, 1879 4 WATSONIAN 3 (20, 0, 4, 0, 0) Restricted to the Humber estuary YORKSHIRE 2 and inland only to Blacktoft Sands SE8524, and rather scarce. Reported once or twice a year since 1 the mid-1970s, and the range of the species is 0 restricted by its estuarine mud and saltmarsh 9 habitat requirement. Beware of confusion with 8 lunulatum and biguttatum. (iv-x, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

lunulatum (Geoffroy in Fourcroy, 1785) riparium (Olivier, 1795)

Bembidion lunulatum (Geoffroy in Fourcroy, 1785) 4 WATSONIAN 3 (10, 2, 43, 22, 0) Widely distributed in the south YORKSHIRE 2 and central lowlands with a few coastal records. In mud at pond edges, in seepages at the bases of 1 coastal clay cliffs, in mud byland drains, in reed- 0 beds, in pitfalls in damp gardens and muddy 9 pastures, at stream sides, and in a borrow pit with 8 dense Glyceria. Beware of confusion with 7 biguttatum. (i-x, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

60 mannerheimii Sahlberg, C.R., 1827 mannerheimi auctt. (misspelling) haemorrhoum sensu auctt. non (Stephens, 1828) unicolor Chaudoir, 1850

Bembidion mannerheimi Sahlberg, 1827 4 WATSONIAN 3 (7, 20, 20, 17, 6) [This is the unicolor of Lindroth, YORKSHIRE 2 1974] Widely distributed with many records. Usually recorded from shaded areas which are 1 very damp, including woodland floors, in decaying 0 piles of mowings, flood debris, also in grass 9 tussocks and under bark in winter. Judging from 8 the records, its range seems to be declining in our 7 area with no recent records from many of its 6 previous localities. (i-ix, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe POGONINI Laporte, 1834

POGONUS Dejean, 1821 chalceus (Marsham, 1802)

Pogonus chalceus (Marsham, 1802) 4 WATSONIAN 3 (9, 0, 0, 0, 0) There are nine Yorkshire records all YORKSHIRE 2 from the Humber estuary in VC61: Saltend TA1627 (TS); Spurn TA4-1- from 1949 (see Nat 1 1952:172) (SS and WDH) to 1996 (MLD). Also at 0 Welwick TA3421, vi 1946 (WDH), and most 9 recently from Beacon Field, Kilnsea TA4116, 8 iv 2003 (FEK). All records are coastal, from 7 strandline debris or saltmarsh litter on the Humber 6 shore. Rarely recorded considering the amount of 5 work that has been done at Spurn and on the 4 lower Humber shore. (iv-ix). 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

61 littoralis (Duftschmid, 1812) litoralis auctt. (misspelling)

Pogonus littoralis (Duftschmid, 1812) 4 WATSONIAN 3 (1, 0, 0, 0, 0) Notable B. A single record only: YORKSHIRE 2 Beacon Field, Kilnsea TA4116, 27 viii 2003 (FEK 1 teste WRD and MLD). A small extension to the north of the range shown in (Luff, 1998) and not 0 unexpected. Populations may exist elsewhere in the 9 Humber estuary and North Lincolnshire. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

luridipennis (Germar, 1822) RDB3 - NO RECORDS but could be expected in the Humber estuary. See Luff (1998) for the distribution which includes sites in North Lincolnshire adjacent Spurn peninsula.

Tribe PATROBINI Kirby, 1837

PATROBUS Dejean, 1821 assimilis Chaudoir, 1844 clavipes Thomson, C.G., 1857

Patrobus assimilis Chaudoir, 1844 4 WATSONIAN 3 (1, 13, 3, 14, 12) All our records are from upland YORKSHIRE 2 sites particularly on open moorland and in bogs. There is only one post-1990 record: Rushy Flat 1 Dike SK213924, 2 v 2005 (RM det EJS). The 0 species may be declining, or perhaps collector 9 effort is minimal in such habitats, but was found 8 to be widely distributed and numerous in pitfall 7 traps on the North York Moors in 1977 (moorland 6 entomological survey (see Coulson and Butterfield, 5 1979) (iv-x, peak viii). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

62 atrorufus (Ström, 1768) excavatus (Paykull, 1790) rufipes sensu (Duftschmid, 1812) non (Fabricius, 1792)

Patrobus atrorufus (Stroem, 1768) 4 WATSONIAN 3 (7, 10, 27, 38, 9) Many records. Widely distributed YORKSHIRE 2 with concentrations of records in the west and south, notably absent from the drier parts of 1 VC61. Recorded in the county from damp 0 woodland floors, under hay bales where conditions 9 are warm and damp, in both unbaited and baited 8 pitfall traps, and damp grassland sites. There are 7 plenty of post-1990 records. (i-x, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

septentrionis Dejean, 1828

Patrobus septentrionis Dejean, 1828 4 WATSONIAN 3 (2, 0, 2, 0, 0) Notable B. Very rare in Yorkshire. YORKSHIRE There are four records on the YNU database: 2 Frizinghall SE1436, 1889 (JWC); Brough SE9326, 1 1903 (TS); Bishop Wilton SE8055, 6 vii 1957 0 (DHS); Blackmoorfoot reservoir SE0913, 1980 9 (MLD). The species is montane/northern upland. 8 The Blackmoorfoot record has recently been 7 checked and is confirmed; the Frizinghall record 6 would require a voucher specimen for 5 confirmation, whilst the other two records are questionable. The habitat is in moss, under 4 stones, etc. 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

63 Tribe PTEROSTICHINI Bonelli, 1810

STOMIS Clairville, 1806 pumicatus (Panzer, 1795)

Stomis pumicatus (Panz er, 1795) 4 WATSONIAN 3 (13, 12, 30, 19, 1) Widespread in the centre and YORKSHIRE the south of the county. The species favours damp 2 habitats and has been recorded from pond, river 1 and reservoir margins, damp roadside debris, damp 0 woodland rides, and in flood refuse. Many of our 9 records are recent (post-1990). (i-xi, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

POECILUS Bonelli, 1810 PTEROSTICHUS sensu auctt. partim non Bonelli, 1810 FERONIA Latreille, 1817 cupreus (Linnaeus, 1758) affinis (Sturm, 1824) erythropus (Dejean, 1828) dinniki (Lutshnik, 1912) caesica Donisthorpe, 1931

Poecilus cupreus (Linnaeus, 1758) 4 WATSONIAN 3 (11, 9, 41, 2, 1) There are many records from a YORKSHIRE 2 rather restricted central band within the recording 1 area. This species may be confused easily with P. 0 versicolor, as the characters in (Lindroth, 1974) are rather comparative and may be difficult to 9 appreciate in the absence of authenticated 8 specimens. The treatment in (Luff, 2007) seems 7 simpler and more straightforward. Some of our 6 records from wet areas may be erroneous because 5 of this possible confusion. P. cupreus seems to 4 favour habitats a little drier than versicolor, and is 3 recorded from peat moors, dry sand/gravel 2 locations and the margins of arable land. (ii-ix, peak v-vi). 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

64 kugelanni (Panzer, 1797) NO RECORDS dimidiatus (Olivier, 1795) non (Rossi, 1790)

lepidus (Leske, 1785)

Poecilus lepidus (Leske, 1785) 4 WATSONIAN 3 (3, 15, 0, 0, 0) Notable B. Very few records with YORKSHIRE only one since1985: Strensall Common SE652608, 2 29 v 2002 (MLD). Recorded from dry heaths, 1 usually in exposed upland situations, with Calluna, 0 and so mainly reported from the North York Moors 9 area. There are also records from Allerthorpe 8 Common SE74, 1919 (TS), 1952 (BK), SE7547, 7 1984 (PK). (iv-viii, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

versicolor (Sturm, 1824) caerulescens sensu auctt. non (Linnaeus, 1758)

Poecilus versicolor (Sturm, 1824) 4 WATSONIAN The distribution of this species is similar to P. 3 YORKSHIRE cupreus, tending to occupy a central band in the 2 recording area, with many records especially in 1 the south. Confusion with cupreus (see note 0 above) may mean that some records are 9 erroneous. Our records, where such information is 8 given, indicate habitats including damp areas on 7 peat moors, in damp meadows with Juncus, in 6 damp ground litter on lowland heaths, on riverside ings, and under stones in damp hay meadows. 5 The phenology of this species shows a marked 4 peak of records in August/September, whereas in 3 cupreus, the peak is clearly much earlier, in 2 May/June(iv-x, peak viii-ix). 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

65 PTEROSTICHUS Bonelli, 1810

Subgenus PTEROSTICHUS Bonelli, 1810 cristatus (Dufour, 1820) ssp. parumpunctatus Germar, 1824

Pterostichus cristatus (Dufour, 1820) 4 WATSONIAN (0, 0, 2, 7, 7) Notable B. All our records except four 3 YORKSHIRE of this rarely reported Pterostichus are post-1990, 2 and all are from the far west and north-west of the 1 region, and habitats where recorded are woodland 0 sites. The earlier generations of YNU coleopterists 9 rarely recorded this species, so it may be enjoying 8 an increase. (iv-ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus STEROPUS Stephens, 1828 aethiops (Panzer, 1796)

Pterostichus aethiops (Panzer, 1796) 4 WATSONIAN 3 (0, 3, 0, 4, 3) Notable B. Rare in Yorkshire and YORKSHIRE 2 only recorded from the far north and west on high 1 moorland. There is only one recent record: Cocket 0 Moss SD786619, 21 viii 2005 (TJB teste MLD). (i-viii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

66 madidus (Fabricius, 1775) concinnus (Sturm, 1818)

Pterostichus madidus (Fabricius, 1775) 4 WATSONIAN 3 (26, 45, 331, 95, 34) Ubiquitous with many YORKSHIRE 2 records. Habitats recorded include woodland of all 1 types, particularly in pitfall traps, gardens urban and 0 rural, grasslands, moorland, arable land, under loose bark (especially in winter) flood refuse, in 9 light traps (although wingless), and in estuarine and 8 coastal strandline debris. (i-xii, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 4/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PEDIUS Motschulsky, 1850 longicollis (Duftschmid, 1812) 39 inaequalis (Marsham, 1802) non (Panzer, 1795) ochraceus (Sturm, 1824)

Pterostichus longicollis (Duftschmid, 1812) 4 WATSONIAN (4, 0, 0, 1, 0) Notable B. Very rare in Yorkshire with 3 YORKSHIRE only five records. Located on several occasions in 2 the Spurn area: Kilnsea TA4115, 17 vii 1948 (JS); 1 Easington TA4019, 2 vi 1950 (WDH); Sea Cliffs 0 TA4116, vi 1951 (JHF); Kilnsea Warren TA419150, 9 vi 1951 (SS) (see Nat 1952:173). The species was 8 not reported again until recorded from Askham Bog 7 (Near Wood) SE576486, 28 viii 1998 (MH det MLD). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus LYPEROSOMUS Motschulsky, 1850 aterrimus (Herbst, 1784) NO RECORDS

67 Subgenus ADELOSIA Stephens, 1835 macer (Marsham, 1802) picimanus (Duftschmid, 1812)

Pterostichus macer (Marsham, 1802) 4 WATSONIAN 3 (12, 8, 6, 4, 1) Widely distributed but very YORKSHIRE 2 infrequent. Generally recorded from the heavier 1 soil areas of the Vale of York, and other central localities, with a few coastal records. Habitats 0 recorded vary from moles‟ nests, in slumping soft 9 clay cliffs, and riverside ings. (iii-xi, peak v-vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PLATYSMA Bonelli, 1810 niger (Schaller, 1783) scotus (Jeannel, 1942)

Pterostichus niger (Schaller, 1783) 4 WATSONIAN 3 (36, 20, 187, 45, 18) Widely distributed with many YORKSHIRE 2 records. Recorded from woodland ground litter and pitfall traps, moors, grasslands, in reed-beds, in hay 1 bales, and on arable land. (ii-xii, peak ix). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

68 Subgenus BOTHRIOPTERUS Chaudoir, 1838 adstrictus Eschscholtz, 1823 orinomum (Stephens, 1828) vitreus (Dejean, 1828)

Pterostichus adstrictus Eschscholtz, 1823 4 WATSONIAN 3 (3, 15, 11, 18, 12) Mainly restricted to the higher YORKSHIRE 2 ground, on moors and heaths, with a few records 1 from lowland heaths and moors (Allerhorpe Common, Skipwith Common and Crowle Moor). 0 There are very few recent records – is this a species 9 in decline? (iii-ix, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

oblongopunctatus (Fabricius, 1787)

Pterostichus oblongopunctatus (Fabricius, 1787) 4 WATSONIAN 3 (9, 22, 21, 3, 0) Notable B. A healthy number of YORKSHIRE recent records exist, mainly from the south and 2 centre of the county, and not recorded from the 1 higher land. Apparently exclusively a woodland 0 insect, it can be very numerous in pitfall traps 9 when found (indicating large populations may be 8 present), in leaf litter, and under the loose bark of 7 logs, especially in winter. (i-xii, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

69 quadrifoveolatus Letzner, 1852 angustatus (Duftchmid, 1812) non (Fabricius, 1787)

Pterostichus quadrifoveolatusus Letzner, 1852 4 WATSONIAN (6, 1, 34, 10, 0) Notable B . Recorded mainly from 3 YORKSHIRE the centre of the county, and in a wide variety of 2 habitats including woodland, wetlands and peat 1 moors (Thorne and Hatfield Moors where the 0 species seems particularly well established). An 9 association with burnt ground is mentioned in 8 (Lindroth, 1974).This is not borne out specifically 7 by our records, but Thorne and Hatfield Moors 6 have been much prone to fire in the past which may account for the species‟ presence there. A 5 fairly recent arrival in Britain (Luff, 2007), the first 4 Yorkshire record was Middleton SE3029, 30 ix 3 1939 (JHF) (see Nat. 1940:51). A spring breeder 2 with an emergence in autumn (Luff, 1998), the 1 phenology of the Yorkshire records shows a very 0 marked peak in September. (ii-xii, peak ix). 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus OMASEUS Stephens, 1828 melanarius (Illiger, 1798) vulgaris sensu auctt. non (Linnaeus, 1758)

Pterostichus melanarius (Illiger, 1798) 4 WATSONIAN 3 (37, 14, 85, 53, 9) Ubiquitous, with many YORKSHIRE records. Recorded from moorland, haystack 2 bottoms, dunes, gardens, damp woodland, river 1 banks, arable and pasture land. Few records from 0 the uplands of the north and northeast. (ii-xii, 9 peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

70 Subgenus PSEUDOMASEUS Chaudoir, 1838 anthracinus (Panzer, 1795)

Pterostichus anthracinus (Panzer, 1795) 4 WATSONIAN 3 (15, 0, 5, 0, 0) Notable B. A scarce species in YORKSHIRE 2 Yorkshire. The main centre of population is the 1 lower Derwent valley (locations in the NNR), and most records post-1986 are from this area. In 0 marshes and fens near to water. (iv-ix, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

gracilis (Dejean, 1828)

Pterostichus gracilis (Dejean, 1828) 4 WATSONIAN 3 (6, 0, 3, 6, 0) Notable B. There are very few YORKSHIRE 2 records from Yorkshire. Recorded from the south of the county in grass tussocks, in pitfalls in 1 marshy areas, at pond edges and by seepage 0 pools. At the northern limit of its British range 9 here. (i-ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

71 minor (Gyllenhal, 1827)

Pterostichus minor (Gyllenhal, 1827) 4 WATSONIAN (24, 17, 85, 27, 0) Many records and widespread 3 YORKSHIRE except on the high ground of the north of the 2 county. A marshland insect, it is recorded from 1 situations in carr land and damp woodland, in 0 riverside ings, on peat moors, and on pond and 9 lagoon edges. (i-xii, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nigrita (Paykull, 1790)

Pterostichus nigrita (Paykull, 1790) 4 WATSONIAN 3 (15, 32, 54, 42, 6) This and the following two YORKSHIRE 2 entries have provided your recorder with problems of interpretation. Pterostichus nigrita sensu 1 (Lindroth, 1974) was found to comprise two 0 species, nigrita (Paykull, 1790) and rhaeticus Heer, 9 1837 (Luff, 1990). The two species are best 8 separable on genitalic characters. Records entered 7 under nigrita s.str. are those considered by this 6 recorder to be from bona-fide sources, and are 5 mapped here. Records on the database that predate 4 1990 have to be considered invalid unless vouchers are forthcoming or have been examined, and this 3 has rarely been the case. Such records have not 2 been mapped but have been recorded as nigrita 1 agg.. Habitats include damp woodland, reservoir 0 and pond edges, riverside ings, reed-beds, willow 9 carr and marshes, but also on dry heaths, peat 8 moors and gardens urban and rural. Widely post-1990 records 7 Map produced by DMAP 24/1/2009 for Yorkshire Naturali sts' Union pre-1990 records distributed except for the high west and north-west. 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 (iii-xi, peak viii).

72 rhaeticus Heer, 1837/8 41 nigrita sensu auctt. partim non (Paykull, 1790)

Pterostichus rhaeticus Heer 4 WATSONIAN 3 (2, 9, 89, 9, 5) These records are the result of YORKSHIRE 2 reports from those considered to be bona-fide 1 sources by this recorder. Habitats noted include 0 most of those listed for nigrita (see the previous entry) except for the very wettest of habitats. 9 Again widely distributed but with more records 8 from higher altitudes than nigrita, and more 7 sparsely recorded from the central lowlands of 6 Yorkshire. (iii-xi, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 4/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus LAGARUS Chaudoir, 1838 vernalis (Panzer, 1795) crenatus (Duftschmid, 1812) non (Gmelin in Linnaeus, 1790)

Pterostichus vernalis (Panzer, 1795) 4 WATSONIAN 3 (12, 8, 22, 22, 0) Recorded mainly from the YORKSHIRE southern half of the county, and recorded from 2 damp situations in woodland, in riverside ings, at 1 pond edges, in damp gardens, a Glyceria swamp, 0 and in decaying grass piles in a wet pasture. (i-xii, 9 peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

73 Subgenus ARGUTOR Stephens, 1828 diligens (Sturm, 1824) strenuus sensu Dawson, 1854 non (Panzer, 1796)

Pterostichus diligens (Sturm, 1824) 4 WATSONIAN 3 (29, 42, 214, 69, 17) Widely distributed with YORKSHIRE many records from wet shaded habitats 2 throughout the five vice-counties. This species is 1 easily and often confused with strenuus and some 0 of our diligens records therefore may be 9 erroneous. (i-xii, peak iv). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

strenuus (Panzer, 1796) erythropus (Marsham, 1802)

Pterostichus strenuus (Panzer, 1796) 4 WATSONIAN 3 (7, 38, 270, 106, 21) Very widespread throughout YORKSHIRE 2 Yorkshire, with many records from somewhat 1 drier situations than diligens, and easily confused 0 with that species. Some of our records therefore may be erroneous, particularly those from high 9 altitudes. (i-xii, peak vi – somewhat and notably 8 later than diligens). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

74 ABAX Bonelli, 1810 parallelepipedus (Piller & Mitterpacher, 1783) parallelopipedus auctt. (misspelling) ater (Villers, 1790) striola (Fabricius, 1792)

Abax parallelepipedus (Piller & Mitterpacher, 1783) 4 WATSONIAN 3 (9, 10, 126, 40, 9) Many records and widely YORKSHIRE 2 distributed across all the five VCs, with a hint of a 1 westerly bias. Usually in deciduous woodland, particularly in pitfall traps where the species 0 occurs sometimes in very large numbers. 9 Occasionally under stones in open fields and in 8 gardens. (ii-xii, peak vii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 12/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

parallelus (Duftschmid, 1812) NO RECORDS

Tribe SPHODRINI Laporte, 1834

PLATYDERUS Stephens, 1828 PLATYDERES Stephens, 1827 depressus (Audinet-Serville, 1821) ruficollis (Marsham, 1802) non (Fabricius, 1787)

Platyderus depressus Audinet-Serville, 1821 4 WATSONIAN 3 (7, 1, 1, 0, 0) Notable B. Very scarce, with only YORKSHIRE 2 nine records and all from the east of the county. 1 The only recent records are repeated occurrence 0 at Elstronwick TA2332 in 1993 and 1994 (WRD) and Burshill TA0948, 17 iv 1994 (FEK) . Habitats 9 for Yorkshire records include under chalk pebbles 8 at base of cliffs and in moss. 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

75 SYNUCHUS Gyllenhal, 1810 TAPHRIA Latreille, 1819 ODONTONYX Stephens, 1827 vivalis (Illiger, 1798) nivalis (Panzer, 1797) non (Paykull, 1790)

Synuchus vivalis (Illiger, 1798) 4 WATSONIAN (6, 6, 23, 13, 3) Widely distributed. Before 1980 3 YORKSHIRE finds appeared mainly to be casual and infrequent. 2 Since that date more intensive site surveying, 1 particularly using pitfalls, has yielded many more 0 records. Populations may be larger and more 9 numerous than earlier recording suggests. 8 Recorded habitats include gardens, woodland and 7 waste ground, generally in situations that are 6 neither very wet or very dry. (v-x, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

CALATHUS Bonelli, 1810

Subgenus AMPHIGYNUS Haliday, 1841 rotundicollis Dejean, 1828 piceus sensu (Marsham, 1802) non (Linnaeus, 1758)

Calathus rotundicollis Dejean, 1828 4 WATSONIAN (6, 2, 34, 9, 1) Generally distributed in the 3 YORKSHIRE southern half of the recording area, although the 2 national distribution shows a general extent to 1 northern Scotland (Luff, 1998). Our records show 0 the species to be almost exclusively woodland, in 9 leaf litter, under logs and in pitfalls, but with a few 8 records from rural gardens and in flood refuse. (i- 7 xii, peaks vi and ix). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

76 Subgenus CALATHUS Bonelli, 1810 ambiguus (Paykull, 1790) fuscus (Fabricius, 1792)

Calathus ambiguus (Paykull, 1790) 4 WATSONIAN (4, 3, 6, 0, 0) Notable B. Very scarce, with only 13 3 YORKSHIRE records. Our records are from areas of dunes on the 2 coast, and from sand pits inland, except one of 1 which I am rather suspicious, i.e., that reported 0 from a woodland situation. The species is 9 arenicolous. (i-ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cinctus Motschulsky, 1850 NO RECORDS mollis sensu auctt. partim non Marsham, 1802 erythroderus Gemminger & Harold, 1868

erratus (Sahlberg, C.R., 1827) fulvipes sensu (Gyllenhal, 1810) non (Fabricius, 1792) flavipes sensu (Duftschmid, 1812) non (Geoffroy in Fourcroy, 1785)

Calathus erratus (Sahlberg, 1827) 4 WATSONIAN 3 (4, 9, 6, 2, 0) Recorded from upland and lowland YORKSHIRE heaths and from coastal dunes. Widespread but 2 very infrequent with no recent occurrences, 1 despite many visits by coleopterists to suitable 0 sites since 1990. (iii-ix, peak viii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

77 fuscipes (Goeze, 1777) cisteloides (Panzer, 1793)

Calathus fuscipes (Goeze, 1777) 4 WATSONIAN 3 (19, 7, 55, 18, 8) Common and widespread except YORKSHIRE 2 on the highest ground in the west and northwest. 1 Habitats recorded in Yorkshire include gardens, 0 pitfalls on moorland, on post-industrial sites, in leaf litter in woodland, under stones in fields and 9 on lowland heaths and dunes. (iii-x, peak ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

melanocephalus (Linnaeus, 1758) ochropterus (Duftschmid, 1812) nubigena Haliday, 1838

Calathus melanocephalus sens.str. (Linnaeus, 1758) 4 WATSONIAN 3 (15, 0, 35, 12, 3) Many records, mainly from the YORKSHIRE 2 south of the county. Some records on the YNU 1 database are noted as melanocephalus s. lat and 0 others as s. str. Some records may be referable to cinctus but in absence of vouchers the situation is 9 unknown. C. melanocephalus s. str is mapped here. 8 C. cinctus is more likely to turn up on the coast – all 7 dissected specimens from Spurn, however, have 6 proved to be melanocephalus. (MLD pers. comm.) 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

78 micropterus (Duftschmid, 1812)

Calathus micropterus (Duftschmid, 1812) 4 WATSONIAN 3 (1, 13, 5, 14, 14) Recorded mainly from upland YORKSHIRE 2 localities, particularly higher forests and moors. A 1 species that seems to like plenty of shade. 0 Recorded from pitfall traps, in ground litter and under stones. An Easington Lagoon record of 2004 9 will have to be discounted unless a voucher 8 specimen is forthcoming (and thus has not been 7 mapped). (iii-ix). 6 5 4 3 2 1 0 9

8 post-1990 records pre-1990 records 7 Map produced by DMAP 23/4/2009 for Yorkshire Naturalists' Union 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

mollis (Marsham, 1802) ochropterus sensu auctt. non (Duftschmid, 1812)

Calathus mollis (Marsham, 1802) 4 WATSONIAN 3 (23, 6, 1, 0, 0) All our records except one: Hatfield YORKSHIRE Moor SE70, 1980 (BE), are coastal. The species is 2 almost exclusively coastal as the national 1 distribution indicates (Luff, 1998), and is recorded 0 from sandy places. The two main Yorkshire 9 population centres are Spurn (modern records) 8 and Teesside (older records from 1900 to 1970). 7 (ii-xii, peak vi - the June peak in phenology may 6 reflects collector effort in this case). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

79 SPHODRUS Clairville, 1806 leucophthalmus (Linnaeus, 1758) leucopthalmus Fowler, 1886 (misspelling) planus (Fabricius, 1792)

Sphodrus leucophthalmus (Linnaeus, 1758) 4 WATSONIAN 3 (0, 2, 3, 0, 0) All our records are very old, and YORKSHIRE from synanthropic situations such as buildings, 2 workshops and cellars: Halifax SE02, 1822 (RLe); 1 Lockwood SE1513, 1883 (FE); Huddersfield SE11, 0 1884 (SLM); Scarborough TA08, date unknown 9 (RLa); Scarborough TA08, 1921 (GBW). Said to 8 prey on Blaps species. Introduced probably in 7 stored products. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

LAEMOSTENUS Bonelli, 1810 LAEMOSTHENES Schaufuss, 1865

Subgenus LAEMOSTENUS Bonelli, 1810 complanatus (Dejean, 1828)

Laemostenus complanatus (Dejean, 1828) 4 WATSONIAN 3 (1, 1, 1, 1, 0) A very rare introduction. All four of YORKSHIRE 2 our records are old: Knottingley SE42, 6 ix 1897 (GBW) (a specimen in a basket maker‟s shop); 1 Hull TA12, 1900 (TS) (under rubbish near the 0 docks); Middlesbrough NZ52, 1912 (AAF); Bolton 9 Abbey SE0753, 22 ii 1959 (JHF) (2 specimens in 8 an old nest of Sand Martin). Confusion with its 7 similar and more frequent and widespread 6 congener L. terricola is possible. 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

80 Subgenus PRISTONYCHUS Dejean, 1828 terricola (Herbst, 1784) subcyaneus (Illiger, 1801)

Laemostenus terricola (Herbst, 1784) 4 WATSONIAN 3 (7, 8, 8, 4, 0) Widespread and a little more YORKSHIRE 2 frequent than L. complanatus. A dribble of modern records taken with the older ones 1 suggests a stable situation for this species in 0 Yorkshire. Reported from a variety of habitats 9 including under logs on heathland, in old 8 buildings with mammal runs, in a coal bunker, in 7 carrion traps, birch stumps, industrial slag piles 6 and around rabbit burrows. (iv-x). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe PLATYNINI Bonelli, 1810

OLISTHOPUS Dejean, 1828 ODONTONYX sensu auctt. non Stephens, 1827 rotundatus (Paykull, 1790) rotundicollis (Marsham, 1802)

Olisthopus rotundatus (Paykull, 1790) 4 WATSONIAN 3 (4, 19, 45, 14, 5) Many records for this YORKSHIRE 2 widespread species, and recorded from a variety 1 of rather dry habitats. Especially numerous in 0 pitfall traps over a wide area of the North York Moors during a Calluna moor survey in 1977 (JC), 9 the species is also noted from gravel pits, lowland 8 peat moors, under straw bales on arable land, and 7 at pond margins and seepages. The only recent 6 records are from the east and south of the county. 5 (ii-x, peak ix). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

81 OXYPSELAPHUS Chaudoir, 1843 AGONUM sensu auctt. partim non Bonelli, 1810 ANCHUS LeConte, 1854 obscurus (Herbst, 1784) ?obscurus (Müller, O.F., 1776) oblongus (Fabricius, 1792)

Oxypselaphus obscurus (Herbst, 1784) 4 WATSONIAN 3 (19, 5, 122, 44, 4) Many records. Our distribution YORKSHIRE 2 map shows a tendency to a rather narrow central corridor in the county, avoiding altitude and the 1 drier areas of the east. The species is at its 0 northern British limit in Yorkshire, and our records 9 give marshes and other shaded damp habitats 8 such as reed-beds, damp woodland, willow carr, 7 riverside ings and banks, and flood plain ground 6 litter as typical habitat. Most records are post- 5 1980 and so may be more frequent than in earlier 4 times, or it may have been ignored by earlier recorders because of its commonness. 3 (i-xii, peak v). 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PARANCHUS Lindroth, 1974 AGONUM sensu auctt. partim non Bonelli, 1810 albipes (Fabricius, 1796) ruficornis (Goeze, 1777) non (De Geer, 1774) oblongus (Fabricius, 1792) non (Fabricius, 1792) pallipes (Fabricius, 1801) non (Fabricius, 1787)

Paranchus albipes (Fabricius, 1796) 4 WATSONIAN 3 (28, 38, 171, 65, 32) Widespread and often very YORKSHIRE 2 numerous in mud and ground litter in freshwater wetland situations, such as pond and lake 1 margins, in reed-beds, river banks and ings, and in 0 carr land. Most records are post-1980. 9 (i-xii, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

82 ANCHOMENUS Bonelli, 1810 AGONUM sensu auctt. partim non Bonelli, 1810 dorsalis (Pontoppidan, 1763) prasinus (Thunberg, 1784)

Anchomenus dorsalis (Pontoppidan, 1763) 4 WATSONIAN 3 (42, 9, 70, 20, 1) There are many records of this YORKSHIRE 2 common and widespread species, but not 1 recorded from the far west and northwest. 0 Recorded habitats range from rather dry situations, such as under stones at field margins, 9 in gardens, to wetland situations that are not 8 inundated. The species occurs from an altitude of 7 400m in upland valleys down to strand line refuse 6 on the east coast. (i-xii, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PLATYNUS Bonelli, 1810 AGONUM sensu auctt. partim non Bonelli, 1810 assimilis (Paykull, 1790) ?junceus (Scopoli, 1763) angusticollis (Fabricius, 1801)

Platynus assimilis (Paykull, 1790) 4 WATSONIAN (6, 15, 135, 37, 8) Many records mainly from 3 YORKSHIRE woodland. Recorded from damp situations, in 2 willow carr, in grass tussocks in riverside ings, 1 under the bark of damp rotting logs, in ground 0 litter and often very numerous in pitfall traps, the 9 species having been found in both coniferous and 8 deciduous woodland. Notably absent from most of 7 VC61. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

83 BATENUS Motschulsky, 1864 AGONUM sensu auctt. partim non Bonelli, 1810 livens (Gyllenhal, 1810)

Batenus livens (Gyllenhal, 1810) 4 WATSONIAN 3 (6, 0, 11, 0, 0) Notable B. Rather scarce. YORKSHIRE 2 Recorded from wetland situations such as the riverside ings in the Lower Derwent Valley NNR, 1 also by pools and reservoirs, in Phragmites beds, 0 under the loose bark of rotting Crack Willow, the 9 species also appears in the subfossil record in the 8 peat of Thorne Moor. It reaches its northern limit 7 of its British range at Wheldrake SE6845. (ii-viii). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

SERICODA Kirby, 1837 AGONUM sensu auctt. partim non Bonelli, 1810 quadripunctata (De Geer, 1774)

Sericoda quadripunctata (Degeer, 1774) 4 WATSONIAN 3 (0, 1, 0, 0, 0) RDB – Endangered. There is only YORKSHIRE 2 one record on the YNU database: Seamer TA0183, 1 ix 1951 (EFG, det PS teste MLL) See Nat. 0 1984:119. This species of forests and heaths may 9 be found under bark. Nationally, records suggest 8 an association with burnt ground. 7 6

5

4

3 2 1 0 9

8 post-1990 records pre-1990 records 7 Map produced by DMAP 14/4/2009 for Yorkshire Naturalists' Union 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

84 AGONUM Bonelli, 1810

Subgenus EUROPHILUS Chaudoir, 1859 fuliginosum (Panzer, 1809)

Agonum fuliginosum (Panz er, 1809) 4 WATSONIAN (58, 22, 193, 68, 23) There are many records of 3 YORKSHIRE this widespread and generally distributed species, 2 which is recorded from marshy areas, reed-beds, 1 pond margins and damp woodland, at all altitudes 0 from the high moors down to the coast. It can be 9 very numerous especially in pitfall traps. 8 (i-xii, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

gracile Sturm, 1824

Agonum gracile Sturm, 1824 4 WATSONIAN (12, 10, 80, 23, 7) Widespread with many records 3 YORKSHIRE from damp situations including marshes, 2 woodland, upland moors to lowland pond margins 1 and peatlands. (i-xii, peak vi). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

85 micans Nicolai, 1822

Agonum micans Nicolai, 1822 4 WATSONIAN 3 (8, 2, 18, 12, 3) Recorded from damp, lowland YORKSHIRE situations, mainly from the centre of our recording 2 area, and noted from pond and lake margins, carr 1 land, bogs and damp woodland. (i-xii, peak v). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

piceum (Linnaeus, 1758)

Agonum piceum (Linnaeus, 1758) 4 WATSONIAN (13, 5, 25, 5, 0) Like the preceding species, 3 YORKSHIRE piceum is recorded from wet situations but with 2 an emphasis on more shaded habitats, such as 1 damp woods and dense reed-beds, but rather less 0 widespread than micans. (ii-xi, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

scitulum Dejean, 1828. Records of this species for Yorkshire appear in (Luff, 1998:109). These records, originally in the YNU database and now deleted, were based on an earlier misidentification (pers comm. MLD, CJ and DGH). There are very few British records (pers comm. MT).

86 thoreyi Dejean, 1828 pelidnum sensu (Paykull, 1792) non (Herbst, 1784) puellum Dejean, 1828

Agonum thoreyi Dejean, 1828 4 WATSONIAN 3 (40, 6, 101, 24, 5) Widely distributed with many YORKSHIRE 2 records, from marshes, reed-beds, pond, tarn and reservoir margins and from a wide range of 1 altitudes. (i-xi, peak iv-v). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus AGONUM Bonelli, 1810 chalconotum Ménétriés, 1832 NO RECORDS sahlbergii (Chaudoir, 1850) sahlbergi auctt. (misspelling) archangelicum Sahlberg, J., 1874

emarginatum (Gyllenhal, 1827) afrum (Duftschmid, 1812) non (Thunberg, 1784) moestum sensu auctt. non (Duftschmid, 1812) nec (Gmelin in Linnaeus, 1790)

Agonum emarginatum Gyllenhal, 1827 4 WATSONIAN 3 (22, 5, 27, 12, 5) Widely distributed with records YORKSHIRE 2 from shaded sites near fresh water, including 1 marshes, pond and drain edges, and reed-beds. There is the possibility of confusion with A. viduum 0 so some of our (earlier) records may be erroneous. 9 Examination of the male genitalia in the two species 8 is recommended. (i-x, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

87 ericeti (Panzer, 1809)

Agonum ericeti (Panzer, 1809) 4 WATSONIAN 3 (0, 7, 8, 6, 1) Notable B. Widely distributed but YORKSHIRE 2 very scarce in Yorkshire. Recorded from bogs with Sphagnum and bare mud, in pitfall traps and 1 water traps.There are very few recent records. (iii- 0 vii, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

gracilipes (Duftschmid, 1812) elongatum Dejean, 1828

Agonum gracilipes (Duftschmid, 1812) 4 WATSONIAN (2, 0, 0, 0, 0) Very rare with two records only: 3 YORKSHIRE Hornsea TA24, 1860 (WKB) (see Ent. Ann. 2 1860:96) [A note in the Fordham records, now 1 archived with the YNU, says that “Mr Bissell found 0 the species under nettles at cliff base, but 9 specimen was lost in the post” (not much change 8 there in 160 years!). Presumably Mr Bissell was 7 sending the specimen to someone for 6 confirmation. The record was therefore never, to my knowledge, confirmed.]; Spurn TA4114, 22 vii 5 1951 (JHF), one specimen under a stone. This 4 species is considered to be a migrant from 3 continental Europe (Luff, 1998; Luff, 2007) which 2 might explain the two east coastal sites. 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

lugens (Duftschmid, 1812) NO RECORDS

88 marginatum (Linnaeus, 1758)

Agonum marginatum (Linnaeus, 1758) 4 WATSONIAN 3 (23, 8, 79, 13, 4) Many records, mainly from the YORKSHIRE centre of the region. Found at margins of still or 2 slow-flowing fresh water where the vegetation is 1 fairly lush. There is a recent record from a 0 suburban garden: Sprotbrough SE553033, 17 viii 9 2001 (RJM), at the edge of a well established 8 garden pond. (i-xii, peak viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

muelleri (Herbst, 1784) parumpunctatum (Fabricius, 1792) chalybeum Sturm, 1824

Agonum muelleri (Herbst, 1784) 4 WATSONIAN 3 (10, 14, 36, 27, 7) Widely distributed and YORKSHIRE 2 recorded from damp grassland, gardens and 1 arable land, pond margins, ground litter in reed- 0 beds, and in damp woodland. (i-xi, peaks vi and viii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

89 nigrum Dejean, 1828 atratum sensu auctt. non (Duftschmid, 1812) dahli (Preudhomme de Borre, 1879)

Agonum nigrum Dejean, 1828 4 WATSONIAN 3 (0, 0, 1, 0, 0) Notable B There is only a single YORKSHIRE 2 record on the YNU database: Halifax SE02. There

1 are two old specimens in the JR Hardy collection in the Manchester Museum (TH teste WDH) [But 0 see Colin Johnson‟s remarks on locality data in 9 Hardy‟s collection. (Johnson, 2004)] Luff, 1998 8 shows dots at SE74 and SK69. 7 6 5 4 3 2 1 0 9

8 post-1990 records pre-1990 records 7 Map produced by DMAP 23/4/2009 for Yorkshire Naturalists' Union 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sexpunctatum (Linnaeus, 1758) NO RECORDS

versutum Sturm, 1824

Agonum versutum Sturm, 1824 4 WATSONIAN 3 (7, 0, 2, 0, 0) Notable B Very scarce, with both YORKSHIRE 2 VC63 records being pre-1917: Ryhill SE3914, 1909 1 (JWC) (see Nat 1909:341), and 1917 (JDF). More 0 recently the species has occurred repeatedly at various sites within Wheldrake Ings SE74 from 1991 9 to 2000 (MLD), in flood refuse and in pitfall traps. 8 There is also a dot in VC62 at about SE79 – see Luff 7 (1998):111. Its typical habitat is on poorly drained 6 vegetated ground near standing water (Luff, 1998 5 and Luff, 2007). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

90 viduum (Panzer, 1796) moestum sensu Fowler & Donisthorpe, 1913 non (Duftschmid, 1812)

Agonum viduum (Panzer, 1796) 4 WATSONIAN (6, 0, 7, 6, 1) Rather a scarce species. Recorded 3 YORKSHIRE from marshes and at the margins of fresh water 2 bodies. Some early records may be referable to 1 moestum because of confusion with nomenclature. 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe ZABRINI Bonelli, 1810

ZABRUS Clairville, 1806 tenebrioides (Goeze, 1777) NO RECORDS gibbus (Fabricius, 1794)

AMARA Bonelli, 1810

Subgenus ZEZEA Csiki, 1929 TRIAENA LeConte, 1847 non Hübner, 1818 plebeja (Gyllenhal, 1810) plebeia auctt. (misspelling)

Amara plebeja (Gyllenhal, 1810) 4 WATSONIAN 3 (28, 21, 133, 47, 6) Many records, and widely YORKSHIRE 2 distributed though not recorded from the high ground of the north-west and far west. Noted from 1 many moist habitats including grassland and arable 0 field margins, woodland, damp meadows, and 9 margins of ponds and reservoirs. (i-xi, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

91 strenua Zimmermann, 1832 NO RECORDS vectensis Dawson, 1849

Subgenus AMARA Bonelli, 1810 aenea (De Geer, 1774) trivialis (Gyllenhal, 1810)

Amara aenea (Degeer, 1774) 4 WATSONIAN 3 (18, 30, 135, 16, 8) There are many records from YORKSHIRE a variety of habitats including dry grassland, 2 arable fields, meadows and dunes, to somewhat 1 damper (but not saturated) situations, such as 0 pond edges, heaths, and woodland rides and 9 clearings. (i-x, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

anthobia Villa & Villa, 1833 SEE APPENDIX A

communis (Panzer, 1797) vulgaris sensu Dawson, 1854 non (Linnaeus, 1758)

Amara communis (Panz er, 1797) 4 WATSONIAN (4, 8, 24, 9, 4) Widely distributed and reported 3 YORKSHIRE from both damp and dry situations including 2 coastal dunes, reservoir and river sides, 1 woodland margins and in quarries. (i-ix, peak vi). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

92 convexior Stephens, 1828 continua Thomson, C.G., 1873

Amara convexior Stephens, 1828 4 WATSONIAN 3 (0, 3, 3, 1, 0) Very scarce in Yorkshire and rare in YORKSHIRE 2 northern England. All records except one are post- 1 1980: Esholt SE1840, 1890 (Anon). The species is otherwise recorded from the following: Timble Ings 0 SE1653, 10 v 1980 (RC det MLL) in plantation 9 woodland at 300 metres altitude; Orange Wood 8 SE1012, 30 vi 1987 (MLD) in upland deciduous 7 woodland; Sheffield SK3484, 17 vi 1992 (AL), on a 6 post-industrial site; Rawcliffe Meadows SE5755, 5 1997 (MLD) in riverside ings (3 records). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 4/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

curta Dejean, 1828

Amara curta Dejean, 1828 4 WATSONIAN 3 (0, 3, 1, 0, 0) Notable B. Very rare with only one YORKSHIRE recent record: Abbey Brook SK203927, 1 vi 1991 2 (EJS). The other three records are all pre-1951: 1 Eston NZ5518, iv 1910 (MLT) and v 1911 (GBW); 0 Yedingham SE8979, ii 1951 (EFG), in flood refuse. 9 The species‟ habitat requirement is stated to be 8 dry situations, e.g. limestone/chalk grassland and 7 in gravel pits. 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

93 eurynota (Panzer, 1796) acuminata (Paykull, 1798)

Amara eurynota (Panzer, 1796) 4 WATSONIAN 3 (6, 5, 22, 3, 1) Infrequently recorded, with YORKSHIRE 2 records mainly from the south of the county, and 1 typically from dry sites, e.g. dunes, old railway 0 tracks, ex-colliery sites and dry grassland. Some of our recent records from wet sites would need 9 vouchers or confirmation of identification. 8 (i-xi, peak ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

famelica Zimmermann, 1832

Amara famelica Zimmerman, 1832 4 WATSONIAN 3 (1, 3, 0, 0, 0) RDB – Rare. Very rare in Yorkshire, YORKSHIRE 2 this is a species of damp areas on sandy heaths. There are four records on the YNU database: 1 Hutton Buscel SE9784, 1895 (WCH), in moss 0 among heather; Allerthorpe Common SE7547, vii 9 1927 (BSL); Strensall Common SE6559, 11 iv 8 1971 (JHF); Strensall Common SE6559, 6 v 1978 7 (RC det MLD teste MLL). We have no recent 6 records. 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

94 familiaris (Duftschmid, 1812)

Amara familiaris (Duftschmid, 1812) 4 WATSONIAN (38, 20, 115, 35, 6) A very widespread and 3 YORKSHIRE common species with many records. Reported 2 mainly from dry, sunny, open habitats from 430 1 metres down to sea level, and one of the most 0 commonly encountered Amara species. 9 (i-xii, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

lucida (Duftschmid, 1812)

Amara lucida (Duftschmid, 1812) 4 WATSONIAN (7, 1, 2, 1, 0) Notable B. With very few Yorkshire 3 YORKSHIRE records, the species has been reported from dunes 2 and dry grassland. As well as sites such as Spurn 1 and Coatham Sands, there is also a record from 0 the dry dry peat areas of Thorne Moor. There is an 9 old record for Wheatley Wood SE6005, 1902 8 (HHC). There are extra dots in Luff (1998) at 7 about TA09, SE36 and SK49, indicating records 6 from other sources. 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

95 lunicollis Schiødte, 1837 vulgaris sensu auctt. non (Linnaeus, 1758)

Amara lunicollis Schioedte, 1837 4 WATSONIAN 3 (3, 12, 60, 9, 1) Widespread, mainly in the south YORKSHIRE 2 and centre of the county. Recorded from high and low level moors, pond margins on ex-colliery sites, 1 on heaths, at woodland margins, and in hay bales 0 in arable land. (i-ix, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

montivaga Sturm, 1825 NO RECORDS nitida Sturm, 1825 NO RECORDS

ovata (Fabricius, 1792) obsoleta sensu Dejean, 1828 non (Duftschmid, 1812) adamantina Kolenati, 1845

Amara ovata (F., 1792) 4 WATSONIAN 3 (9, 4, 12, 11, 2) Widespread but infrequently YORKSHIRE 2 encountered in all the vice counties. From a variety of habitats including woodland, grassland, 1 gardens, coastal dunes and peat moors, and 0 generally in rather dry situations. (v-xi, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

96 similata (Gyllenhal, 1810) obsoleta (Duftschmid, 1812)

Amara similata (Gyllenhal, 1810) 4 WATSONIAN 3 (20, 10, 61, 19, 5) Widespread and evenly YORKSHIRE 2 distributed throughout the county, with most of 1 our records being from dry, open situations, e.g. 0 coastal dunes, upland limestone areas, dry heaths, gravel pits, and post-industrial sites, but also from 9 some woodlands, lagoon edges and pasture land. 8 (i-xi, peaks v and ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

spreta Dejean, 1831

Amara spreta Dejean, 1831 4 WATSONIAN (0, 2, 0, 0, 0) Notable B. We have only two 3 YORKSHIRE records on our database: Hutton Buscel SE9784, 2 1895 (WCH), in moss among heather; Coatham 1 Sands NZ5725, 1995 (ME), in sand dunes. The 0 latter record is confirmed and in typical habitat, 9 but the former, coupled with the extremely thin 8 known distibution would require a voucher 7 specimen for confirmation, and as such is questionable. The species remains on the 6 database for the time being. A. spreta could be 5 expected to occur at Spurn but has not been found 4 there so far. 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

97 tibialis (Paykull, 1798)

Amara tibialis (Paykull, 1798) 4 WATSONIAN (15, 2, 24, 0, 0) Recorded from the south and 3 YORKSHIRE east of the county from open, dry, often sandy, 2 situations on heaths, in grassland and dry 1 woodland margins, and also on waste ground on 0 post-industrial sites. In Britain generally, the 9 species is widespread but tending to be more 8 coastal in the far north. (iii-xi, peaks vi and ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus CELIA Zimmermann, 1832 bifrons (Gyllenhal, 1810) livida sensu Schiødte, 1841 non (Fabricius, 1792)

Amara bifrons (Gyllenhal, 1810) 4 WATSONIAN 3 (8, 0, 12, 8, 1) A rather infrequent Amara, with YORKSHIRE 2 few recent records. A species of mainly of dry, sandy and well drained situations, it has been 1 noted from dunes, quarries, gravel pits and post- 0 industrial sites. (v-ix, peak viii). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cursitans (Zimmermann, 1832) NO RECORDS fuscicornis (Zimmermann, 1832)

fusca Dejean, 1828 complanata Dejean, 1828 (0, 0, 1, 0, 0) RDB1 There is a single record on the YNU database: Doncaser SE50, 1884 (JWE). Because of its age and lack of voucher this record cannot be confirmed and has therefore not been mapped. infima (Duftschmid, 1812) NO RECORDS but might be expected to occur on the Spurn peninsula. There is an old record for coastal dunes in nearby North Lincolnshire.

98 praetermissa (Sahlberg, C.R., 1827) rufocincta (Sahlberg, C.R., 1827)

Amara praetermissa (Sahlberg, 1827) 4 WATSONIAN (2, 1, 10, 0, 1) Notable B. A rare species in 3 YORKSHIRE Yorkshire, with very few records. Recorded from 2 dry habitats including heaths, peat moors and 1 post-industrial sites. (vi-ix). 0 9 8 7 6

5

4

3 2 1 0 9

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Subgenus PARACELIA Bedel, 1899 quenseli (Schönherr, 1806) NO RECORDS quenselii auctt. (misspelling)

Subgenus BRADYTUS Stephens, 1827 apricaria (Paykull, 1790)

Amara apricaria (Paykull, 1790) 4 WATSONIAN 3 (8, 5, 42, 9, 9) Widely distributed with many YORKSHIRE 2 records from a variety of fairly dry situations, including sand/gravel pits, coastal dunes, roadside 1 waste places, haystack bottoms, and also in light 0 traps, on post-industrial sites and peat moors, but 9 also noted from riverside shingle beds. 8 (iv-x, peak viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

99 consularis (Duftschmid, 1812)

Amara consularis (Duftschmid, 1812) 4 WATSONIAN 3 (4, 2, 2, 4, 0) Notable B. Very scarce, with a county YORKSHIRE 2 tally of 12 records. There are only two recent 1 records: Manor Farm, Eddlethorpe SE772661, 2000 0 (DL teste MLD); Beningbrough SE513579, 11 vii 2005 (AG det MLD).These two were the first since 9 1929. There are dots in Luff, 1998:116 for the 8 Thorne area. These records do not figure in 7 Skidmore (2006). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

fulva (Müller, O.F., 1776)

Amara fulva (Muller, O.F., 1776) 4 WATSONIAN 3 (11, 0, 9, 0, 2) Notable B. A scarce species, with YORKSHIRE 2 records from sand pits, dunes and in sand below 1 coastal soft cliffs, and also from dry heaths. (iv-ix). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

100 Subgenus PERCOSIA Zimmermann, 1832 equestris (Duftschmid, 1812) patricia (Duftschmid, 1812)

Amara equestris (Duftschmid, 1812) 4 WATSONIAN 3 (0, 1, 0, 0, 0) Notable B. There is a single record YORKSHIRE 2 on the YNU database: Danby Low Moor NZ7010, 1 1987 (MU/MF det MLL). This record does not 0 appear in Luff (1998). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

CURTONOTUS Stephens, 1827 CYRTONOTUS auctt. (misspelling) AMARA sensu auctt. partim non Bonelli, 1810 alpinus (Paykull, 1790) NO RECORDS

aulicus (Panzer, 1796) spinipes sensu Schiødte, 1841 non (Linnaeus, 1758)

Curtonotus aulicus (Panzer, 1796) 4 WATSONIAN 3 (24, 17, 64, 22, 3) Widely distributed with many YORKSHIRE 2 records, and noted from a wide variety of habitats 1 including coastal dunes, woodland margins, 0 quarries, gravel pits and arable fields. There are plenty of recent records of this very distinctive 9 Amara. (iii-x, peak vii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

101 convexiusculus (Marsham, 1802)

Curtonotus convexiusculus (Marsham, 1802) 4 WATSONIAN 3 (25, 3, 11, 5, 0) This species is recorded mainly YORKSHIRE 2 from the south of the county from a variety of 1 habitats including coastal dunes, gravel pits, in 0 light traps, in leaf litter at woodland margins, on colliery spoil heaps and in strand refuse by 9 saltmarshes. (ii-x, peak viii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe HARPALINI Bonelli, 1810

HARPALUS Latreille, 1802

Subgenus PSEUDOOPHONUS Motschulsky, 1844 PSEUDOPHONUS auctt. (misspelling) PARDILEUS des Gozis, 1882 calceatus (Duftschmid, 1812)

Harpalus calceatus (Duftschmid, 1812) 4 WATSONIAN 3 (1, 0, 0, 0, 0) A single record exists on the YNU YORKSHIRE 2 database: Bridlington TA16, viii 1879 (WWF) (see Nat 1886:65). This species is an occasional 1 immigrant and I assume that such was the case 0 with this record. 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

griseus (Panzer, 1797) NO RECORDS

102 rufipes (De Geer, 1774) ruficornis (Fabricius, 1775) pubescens (Müller, O.F., 1776)

Harpalus rufipes (Degeer, 1774) 4 WATSONIAN 3 (35, 13, 73, 22, 5) A widespread species in YORKSHIRE Yorkshire with many records. Reported from 2 carrion traps, coastal dunes, woodland, gravel 1 pits, in pitfalls on riverside ings, at ex-colliery 0 sites, at light in houses, in damp herb-rich 9 pasture, in riverside shingle beds, on peat moors 8 and on arable margins. (iii-x, peaks vi and viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus HARPALUS Latreille, 1802 HAPLOHARPALUS Schauberger, 1926 affinis (Schrank, 1781) aeneus (Fabricius, 1775) non (De Geer, 1774)

Harpalus affinis (Schrank, 1781) 4 WATSONIAN (26, 11, 64, 19, 3) Widespread with many 3 YORKSHIRE records, a high proportion of these being recent. 2 Where habitat information is included in the 1 records, the species has been noted from dry, 0 open situations, including fields, grassland and 9 gardens. (i-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

103 anxius (Duftschmid, 1812)

Harpalus anxius (Duftschmid, 1812) 4 WATSONIAN 3 (1, 0, 0, 0, 0) We have a single record on our YORKSHIRE 2 database: Spurn Point TA3910, 20 ix 1986 (MLD) 1 (see Denton, 1995). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

attenuatus Stephens, 1828 consentaneus Dejean, 1829

Harpalus attenuatus Stephens, 1828 4 WATSONIAN 3 (1, 0, 0, 0, 0) A single record exists: Spurn YORKSHIRE (Kilnsea Warren) TA4114, 7 vi 1950 (SS), two 2 specimens (see Nat 1952:172). 1 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cupreus Dejean, 1829 NO RECORDS

dimidiatus (Rossi, 1790) NO RECORDS caspius sensu auctt. Brit. non (von Steven, 1806)

104 honestus (Duftschmid, 1812) NO RECORDS ignavus (Duftschmid, 1812) laevipes Zetterstedt, 1828 NO RECORDS seriepunctatus sensu Gyllenhal, 1827 non Sturm, 1818 quadripunctatus Dejean, 1829 montivagus Reitter, 1900 latus (Linnaeus, 1758) erythrocephalus (Fabricius, 1787) metallescens Rye, 1874

Harpalus latus (Linnaeus, 1758) 4 WATSONIAN 3 (9, 11, 7, 5, 2) Widespread but with few records. YORKSHIRE 2 Reported in Yorkshire from coastal dunes, dry grassland ,peat moors, upland heaths and moors, 1 and from lowland heaths. (iv-viii, peak v). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

neglectus Audinet-Serville, 1821

Harpalus neglectus Serville, 1821 4 WATSONIAN 3 (1, 0, 0, 0, 0) A species of sandy coastal dunes YORKSHIRE and cliffs (Luff, 1998 and Luff, 2007). We have 2 two records on the database: Saltaire SE1337, 1 1886 (JWC); Spurn TA3910, 15 v 1983 (MLD) (see 0 Denton, 1995 – “a single specimen located at the 9 Point”). Of these two records the former is 8 considered erroneous, and in respect of this there 7 is a comment in the (paper) Fordham records: 6 “This seems most improbable – JH Flint 8/2/63”. 5 The circumstances of this capture are not 4 recorded. 3 2 1 0

9

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105 pumilus Sturm, 1818 NO RECORDS vernalis sensu (Duftschmid, 1812) non (Panzer, 1796) picipennis sensu auctt. Brit. non Duftschmid, 1812 funestus Audinet-Serville, 1821

rubripes (Duftschmid, 1812) sobrinus Dejean, 1829

Harpalus rubripes (Duftschmid, 1812) 4 WATSONIAN 3 (3, 2, 13, 1, 0) This species is widespread but very YORKSHIRE 2 thinly recorded in Yorkshire, from habitats 1 including peat moors, in sedge litter, in a sand quarry, at post-idustrial sites and lowland sandy 0 heaths. (iv-ix, peak vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rufipalpis Sturm, 1818 rufitarsis (Duftschmid, 1812) non (Illiger, 1801) ignavus sensu auctt. Brit. non (Duftschmid, 1812)

Harpalus rufipalpis Sturm, 1818 4 WATSONIAN 3 (0, 0, 9, 1, 0) The accepted habitat for this species YORKSHIRE 2 is sandy soils, sand pits, heaths and dunes. We 1 have ten records. While most come from known sandy sites in the Hatfield Moor area and from 0 post-industrial sites in Sheffield, one or two are 9 suspect, e.g. Gateforth SE5628, 1945 (JRD), and a 8 recent record from the Magnesian limestone at 7 SE5007, this latter requiring confirmation. 6 (iv-viii, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

serripes (Quensel in Schönherr, 1806) NO RECORDS

106 servus (Duftschmid, 1812)

Harpalus servus (Duftschmid, 1812) 4 WATSONIAN (1, 0, 1, 0, 0) Notable B. There is only one record 3 YORKSHIRE on the YNU database: that from Spurn Chalk Bank 2 TA4112, 21 ix 1986 (MLD), under driftwood, 1 (Denton, 1995:15). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

smaragdinus (Duftschmid, 1812) SEE APPENDIX A discoideus Erichson, 1837

tardus (Panzer, 1796) rufimanus (Marsham, 1802) luteicornis sensu auctt. Brit. non (Duftschmid, 1812)

Harpalus tardus (Panzer, 1796) 4 WATSONIAN 3 (24, 1, 6, 3, 0) Infrequently recorded. Most of our YORKSHIRE 2 records are recent (post-1990), either from Spurn 1 or from the sandy/gravelly parts of the Hatfield Moors area. Also recorded from a sand quarry, 0 and a single records from carr land at Potteric 9 Carr, this latter requiring confirmation. 8 (iii-x, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

107 Subgenus CRYPTOPHONUS Brandmayr & Zetto Brandmayr, 1982 melancholicus Dejean, 1829 NO RECORDS tenebrosus Dejean, 1829 NO RECORDS ssp. centralis Schauberger, 1929

OPHONUS Dejean, 1821 HARPALUS sensu auctt. partim non Latreille, 1802

Subgenus OPHONUS Dejean, 1821 ardosiacus Lutshnik, 1922 obscurus sensu (Sturm, 1818) non (Fabricius, 1792) diffinis sensu Joy, 1932 non (Dejean, 1829) rotundicollis sensu auctt. Brit. non Kolenati, 1845

Ophonus ardosiacus Lutshnik, 1922 4 WATSONIAN 3 (5, 1, 0, 0, 0) Notable B. Rare, with six records YORKSHIRE 2 only; Marfleet TA12, 29 viii 1908 (TS), under 1 driftwood on the Humber shore; Spurn (Kilnsea Warren) TA4115, 19 viii 1949 (SS) and vi 1950 0 (SS); West Ayton SE9884, 1950 (AMR); Spurn 9 TA41, 21 vii 1963 (CJ). For the Kilnsea Warren 8 records see also Nat 1952:172. 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

azureus (Fabricius, 1775) NO RECORDS subquadratus sensu auctt. Brit. non (Dejean, 1829)

sabulicola (Panzer, 1796) NO RECORDS

stictus Stephens, 1828 NO RECORDS obscurus (Fabricius, 1792) non (Müller, O.F., 1776) monticola (Dejean, 1829)

Subgenus METOPHONUS Bedel, 1895 cordatus (Duftschmid, 1812) NO RECORDS

laticollis Mannerheim, 1825 NO RECORDS punctatulus (Duftschmid, 1812) non (Fabricius, 1792) nitidulus sensu Stephens, 1828 non (Schrank, 1781)

108 melletii (Heer, 1837/8) NO RECORDS melleti auctt. (misspelling) rectangulus Thomson, C.G., 1870 championi Sharp, 1912 rupicoloides Sharp, 1912 brevicollis sensu Jeannel, 1942 ? (Audinet-Serville, 1821)

parallelus (Dejean, 1829) NO RECORDS melleti sensu Jeannel, 1942 non (Heer, 1837/8) zigzag sensu auctt. Brit. non Costa, 1882 puncticeps Stephens, 1828 rectangulus sensu Sharp, 1912 non Thomson, C.G., 1870 angusticollis Müller, J., 1921

Ophonus puncticeps Stephens, 1828 4 WATSONIAN 3 (3, 0, 2, 0, 0) Rare in Yorkshire with only five YORKSHIRE records: Waterloo SE1817, 23 iv 1948 ((EWA); 2 Welwick TA3421, 15 vii 1948 (SS); Spurn TA4115, 1 vi 1950 (SS), in Marram on the Humber foreshore 0 near the Warren; Filey Brigg TA1381, 18 ix 1983 9 (JHF); Worsborough SE3403, 20 ix 1987 (JDC). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

puncticollis (Paykull, 1798)

Ophonus puncticollis (Paykull, 1798) 4 WATSONIAN 3 YORKSHIRE (2, 0, 2, 0, 0) RDB – Rare. A rare species in 2 Yorkshire with four records on the database, and 1 all are very old: Barnsley SE30, undated (EGB); 0 Hampole SE5010, 1908 (HHC); Filey TA08/TA18, 9 viii 1878 (WWF); Filey TA08/TA18, 31 v 1903 8 (MLT). The Filey records although locations are 7 imprecise appear in Luff (1998). The other two cannot be confirmed but remain in the records for 6 the time being. Hampole is on Magnesian 5 limestone. The Barnsley habitat remains unknown. 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

109 rufibarbis (Fabricius, 1792) brevicollis sensu auctt. partim non (Audinet-Serville, 1821) subpunctatus Stephens, 1828 seladon (Schauberger, 1926)

Ophonus rufibarbis (Fabricius, 1792) 4 WATSONIAN (28, 8, 29, 7, 4) Widely distributed and frequently 3 YORKSHIRE reported. From a wide range of habitats including 2 woodland margins, arable land, river banks, 1 gardens, roadside verges and coastal dunes. 0 (iv-x, peak v). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rupicola (Sturm, 1818) SEE APPENDIX A

schaubergerianus (Puel, 1937) rufibarbis sensu auctt. non (Fabricius, 1792) brevicollis sensu auctt. partim non (Audinet-Serville, 1821)

Ophonus schaubergerianus (Puel, 1937) 4 WATSONIAN 3 (4, 3, 10, 1, 1) Notable B. Some of the older YORKSHIRE records predating 1950 are questionable but not 2 invalidated on the database for the time being. 1 Most of our modern records are considered 0 reliable, bearing in mind that confusion is possible 9 with the common rufibarbis. Recorded from 8 grasslands, woodland, arable land and in coastal 7 dunes. (iv-x, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

subsinuatus Rey, 1886 NO RECORDS

110 ANISODACTYLUS Dejean, 1829 binotatus (Fabricius, 1787) spurcaticornis Dejean, 1829

Anisodactylus binotatus (Fabricius, 1787) 4 WATSONIAN 3 (2, 2, 49, 4, 0) Mainly recorded from a rather YORKSHIRE 2 restricted area of southern Yorkshire with a few 1 localities further afield, the species has been found to be widespread and very numerous in pitfall 0 traps on Thorne Moor during a survey in 2005, 9 indicating very large local populations. Noted 8 mostly from open, damp habitats, including peat 7 moors, wet heaths, gravel pits, damp woodland, 6 willow carr, and in wet areas on post-industrial 5 sites. (iii-ix, peak v). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nemorivagus (Duftschmid, 1812) NO RECORDS atricornis (Stephens, 1835)

poeciloides (Stephens, 1828) NO RECORDS pseudoaeneus sensu auctt. non Dejean, 1829

DIACHROMUS Erichson, 1837 germanus (Linnaeus, 1758) NO RECORDS

SCYBALICUS Schaum, 1862 oblongiusculus (Dejean, 1829) NO RECORDS

111 DICHEIROTRICHUS Jacquelin du Val, 1855 DICHIROTRICHUS auctt. (misspelling) gustavii Crotch, 1871 gustavi auctt. (misspelling) pubescens (Paykull, 1790) non (Müller, O.F., 1776)

Dicheirotrichus gustavii Crotch, 1871 4 WATSONIAN (12, 0, 1, 0, 0) All 13 records on the YNU 3 YORKSHIRE database are coastal or estuarine in the Humber 2 area, from saltmarsh or strand refuse. There are 1 no records predating 1946, but a spate of modern 0 records is the result of increased fieldwork in 9 those localities. (v-x, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

obsoletus (Dejean, 1829) SEE APPENDIX A

TRICHOCELLUS Ganglbauer, 1892 BRADYCELLUS sensu Fowler, 1886 partim non Erichson, 1837 cognatus (Gyllenhal, 1827)

Trichocellus cognatus (Gyllenhal, 1827) 4 WATSONIAN 3 (5, 18, 28, 17, 5) A widespread and fairly frequent YORKSHIRE species recorded from heaths and moors with 2 Calluna, usually but not always upland. Owing to 1 confusion with T. placidus a few records (of those 0 from the Vale of York and eastern VC63 ) may be 9 erroneous. (ii-xii, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

112 placidus (Gyllenhal, 1827)

Trichocellus placidus (Gyllenhal, 1827) 4 WATSONIAN 3 (32, 23, 109, 57, 3) Widely distributed with many YORKSHIRE 2 records, this is mainly a species of lowland fens, 1 marshes and damp woodland. Some records from 0 higher ground in the county may be referable to T. cognatus (owing to possible confusion with the 9 previous species. The far greater number of 8 placidus records over those of cognatus may 7 reflect higher collector effort in lowland localities. 6 (i-xii, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 20/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

BRADYCELLUS Erichson, 1837 caucasicus (Chaudoir, 1846) collaris (Paykull, 1798) non (Herbst, 1784)

Bradycellus caucasicus (Chaudoir, 1846) 4 WATSONIAN 3 (2, 6, 12, 9, 7) In Yorkshire this is an infrequent YORKSHIRE species of dry heaths with Calluna, and generally 2 from upland sites in the county. (ii-viii, peak viii). 1 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

csikii Laczó, 1912 NO RECORDS

113 harpalinus (Audinet-Serville, 1821)

Bradycellus harpalinus (Audinet-Serville, 1821) 4 WATSONIAN 3 (13, 24, 123, 36, 4) Many records with a wide YORKSHIRE 2 distribution in the county and from a variety of 1 habitats. These include open sites such as 0 reservoir and lake sides, gravel pits, peat moors, post-industrial sites, woodland margins and 9 lowland heaths. The species is also recorded from 8 light traps. (i-xii, peaks v and viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ruficollis (Stephens, 1828) similis (Dejean, 1829)

Bradycellus ruficollis (Stephens, 1828) 4 WATSONIAN (4, 31, 45, 14, 7) A species of Calluna heaths, 3 YORKSHIRE coastal dunes, peat moors and upland reservoir 2 margins, it is also recorded from light traps. Most 1 records are from higher ground with very few 0 records from VC61. VC65 is generally under- 9 recorded, so such may be the case for this species. 8 (ii-xi, peaks v and ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

114 sharpi Joy, 1912 distinctus sensu Fowler, 1886 non (Dejean, 1829)

Bradycellus sharpi Joy, 1912 4 WATSONIAN 3 (2, 5, 10, 5, 0) A rather infrequent species YORKSHIRE 2 recorded from a wide range of altitudes, but 1 none yet from VC65, and noted habitats include 0 gravel pits, woodland, lake margins, on clay cliffs on the east coast, and from upland heaths. 9 There is a very pronounced gap in the phenology 8 from vi-viii. (ii-x, peak iv). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

verbasci (Duftschmid, 1812)

Bradycellus verbasci (Duftschmid, 1812) 4 WATSONIAN (8, 3, 44, 10, 4) Frequent, from a wide variety of 3 YORKSHIRE habitats including coastal dunes, in light traps and 2 at lighted buildings, in gravel pits, on carr land, 1 cultivated field edges, peat moors, riverside flashes 0 and reed-beds and on post-industrial sites. (i-xii, 9 peak viii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

STENOLOPHUS Dejean, 1821 AGONODERUS Dejean, 1829 comma (Fabricius, 1775) NO RECORDS plagiatus Gorham, 1901

115 mixtus (Herbst, 1784) vespertinus (Panzer, 1796) ziegleri (Panzer, 1809)

Stenolophus mixtus (Herbst., 1784) 4 WATSONIAN 3 (21, 5, 88, 25, 1) Many records with a distribution YORKSHIRE 2 concentrated in the south and centre of the county, 1 this species is southern British insect and so is near the northern limit of its national distribution here in 0 Yorkshire. Recorded mainly from damp sites, e.g. 9 bogs, pond and lake margins, flooded gravel pits, 8 riverside flashes and ings, drain edges on peat 7 moors, in wet ground litter in hay meadows and on 6 carr land. (ii-x, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

skrimshiranus Stephens, 1828 NO RECORDS

teutonus (Schrank, 1781) NO RECORDS vaporariorum sensu (Fabricius, 1775) non (Linnaeus, 1758) anglicus Schiødte, 1861-63

116 ACUPALPUS Latreille, 1829 brunnipes (Sturm, 1825) SEE APPENDIX A brunneipes auctt. (misspelling)

dubius Schilsky, 1888 luridus sensu auctt. non Dejean, 1829 luteatus sensu Joy, 1932 non (Duftschmid, 1812)

Acupalpus dubius Schilsky, 1888 4 WATSONIAN 3 (14, 20, 154, 32, 0) A species with many records YORKSHIRE 2 but with a distribution confined to the south and 1 centre of our recording area. By far the most commonly encountered Acupalpus in damp 0 situations, recorded from ground litter in marshy 9 localities, also in gravel pits, damp woodland 8 margins, drain sides, riverside ings, reed-beds and 7 pond margins. Often found in large numbers in 6 trapping samples. (i-xi, peak v). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

elegans (Dejean, 1829) NO RECORDS

exiguus Dejean, 1829

Acupalpus exiguus Dejean, 1829 4 WATSONIAN 3 (1, 0, 3, 0, 0) Notable B. There are two confirmed YORKSHIRE 2 records on the YNU database: Sykehouse SE630162, 2 vi 2005 (RJM confirmed MLL), a 1 single specimen occurring by sweeping in marshy 0 grassland. The species turned up again at a site in 9 the Sykehouse area at SE644175, but this time in 8 some numbers. Males were obtained and 7 dissected to confirm the identification. See Freude, 6 Harde and Lohse (2004). Another two records are 5 held: Skipwith Common SE6537, vi 1923 (MLT), 4 and Thorne Moor SE71, iv 1914 (HHC). Both 3 would require voucher specimens for confirmation, 2 neither of which are forthcoming, and have not 1 been mapped here. The species was reported in pitfall traps on Thorne Moor in the early 1990s but 0 the material could not be traced or verified. A few 9 other specimens so named have been passed to 8 post-1990 records your recorder for identification in the past and pre-1990 records 7 Map produced by DMAP 14/4/2009 for Yorkshire Naturalists' Union 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 have all proved to be dark dubius.

117 flavicollis (Sturm, 1825) NO RECORDS luridus Dejean, 1829

maculatus (Schaum, 1860) NO RECORDS

meridianus (Linnaeus, 1761)

Acupalpus meridianus (Linnaeus, 1761) 4 WATSONIAN 3 (4, 0, 12, 2, 0) Very infrequent and only recorded YORKSHIRE 2 from the southern half of the county. Generally noted in rather dry situations, e.g. a deer park, 1 drier areas on peat moors, ex-colliery sites, 0 woodland, and coastal dunes, but there are a few 9 records from carr land and reservoir margins. 8 (ii-x). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

parvulus (Sturm, 1825) dorsalis (Fabricius, 1787) non (Pontoppidan, 1763) derelictus (Dawson, 1854)

Acupalpus parvulus (Sturm, 1825) 4 WATSONIAN 3 (15, 3, 37, 8, 0) More frequently known to YORKSHIRE 2 coleopterists as the Acupalpus dorsalis of Fabricius, this species is fairly frequently recorded 1 from lowland sandy areas including disused and 0 flooded sand and gravel quarries, coastal dunes, 9 but also from pool edges on peat. There are 8 several recent records from light traps in suitable 7 locations. (ii-xi, peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

118 ANTHRACUS Motschulsky, 1850 ACUPALPUS sensu auctt. partim non Latreille, 1829 consputus (Duftschmid, 1812)

Anthracus consputus (Duftschmid, 1812) 4 WATSONIAN (7, 1, 3, 4, 0) Notable B. Scarce, with records 3 YORKSHIRE from a rather narrow central belt in Yorkshire, 2 with a few records from the east coast. Noted 1 from pitfalls in wet meadows, by reservoirs and in 0 marshy areas on ex-colliery sites, and from flood 9 refuse on riverside ings, by ponds in carr land and 8 at the edges of scrapes on bare peat. Most records 7 are post-1985 and so the species may be 6 increasing in range. (iv-x, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe CHLAENIINI Laporte, 1834

CHLAENIUS Bonelli, 1810 CHLAENIELLUS Reitter, 1908 nigricornis (Fabricius, 1787) melanocornis Dejean, 1826

Chlaenius nigricornis (F., 1787) 4 WATSONIAN 3 (13, 1, 1, 3, 1) Notable B. Very infrequent. Noted YORKSHIRE 2 from lake margins, bogs, riverside ings and flood 1 plains. Most records are from south central 0 Yorkshire with outliers from: Hornsea Mere TA1846, 1859 (WKB) in VC61; Semerwater 9 SD9186, 1905 (MLT) in VC65. (v-xii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nitidulus (Schrank, 1781) NO RECORDS schrankii (Duftschmid, 1812)

119 tristis (Schaller, 1783) SEE APPENDIX A holosericeus (Fabricius, 1787)

vestitus (Paykull, 1790)

Chlaenius vestitus (Paykull, 1790) 4 WATSONIAN (15, 1, 2, 0, 0) Recorded from very few sites. The 3 YORKSHIRE species is mostly coastal, especially at the base of 2 soft clay cliffs south of Bridlington around 1 Barmston, and was found repeatedly here during 0 YNU (and other) surveys from 2002 to 2006. In mud and clay cracks near water seems to be the 9 preferred habitat. Also recorded inland from 8 Barnby Dun SE6109, 1985 (PS), habitat not 7 recorded; Thixendale SE813019, 26 vi 2000 (MH 6 det MLD), habitat not recorded; Dunsville Sand 5 Quarry SE650077, 29 ix 2008 (RJM), on bare sand/gravel at edge of flooded gravel pit 4 surrounded by mounds of soil forming small clay 3 cliffs, an ideal, artificially created habitat. Luff 2 (1998) states that there is a recent record for 1 Wheldrake Ings SE74. In the Lower Derwent Ings Coleoptera survey (Denton and Marsh, 1995) the 0 species was never found. We are of the opinion 9 that the species referred to may apply to 8 nigricornis for which there are valid records (see post-1990 records 7 Map produced by DMAP 9/2/2009 for Yorkshire Naturalists' Union pre-1990 records above). The habitat requirements of the two 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 species are rather different. (v-ix).

CALLISTUS Bonelli, 1810 lunatus (Fabricius, 1775) NO RECORDS

120 Tribe OODINI LaFerté-Sénectère, 1851

OODES Bonelli, 1810 helopioides (Fabricius, 1792)

Oodes helopioides (Fabricius, 1792) 4 WATSONIAN (1, 1, 0, 1, 0) Notable B. There are three records 3 YORKSHIRE on the YNU database, and all of these very old: 2 Hornsea Mere TA1846, 1856 (WKB); Askham Bog 1 SE54 (indet location), c1870 (WCH); York SE65 0 (indet location), 1885 (JWE). The accepted habitat 9 preference is around edges of lakes, in fens and 8 by slow rivers in ground litter (Luff, 1998 and Luff, 7 2007). Nowadays only recorded from south of a 6 Wash/South Wales line. 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe LICININI Bonelli, 1810

LICINUS Latreille, 1802 depressus (Paykull, 1790)

Licinus depressus (Paykull, 1790) 4 WATSONIAN (9, 1, 1, 0, 0) Notable B. Rarely recorded in 3 YORKSHIRE Yorkshire with most records being from VC61. The 2 most recent record is: Cottam Well Dale SE9763, 1 14 ix 1983 (RGB) (see Nat 1988:69). (iv-ix). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

punctatulus (Fabricius, 1792) NO RECORDS punctulatus Kloet & Hincks, 1945 (error) silphoides sensu (Fabricius, 1792) non (Rossi, 1790)

121 BADISTER Clairville, 1806 Subgenus BADISTER Clairville, 1806 bullatus (Schrank, 1798) bipustulatus (Fabricius, 1792) non (Fabricius, 1775)

Badister bullatus (Schrank, 1798) 4 WATSONIAN (41, 9, 71, 21, 2) More familiarly known as 3 YORKSHIRE bipustulatus, this species is widely distributed with 2 many records, except in the far north. Recorded 1 from a wide variety of habitats including 0 deciduous woodland, lowland heaths, coastal 9 dunes, grassland, arable field edges, gravel pits, 8 urban gardens, pond margins and post-industrial 7 sites. (i-xii, peak vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

meridionalis Puel, 1925 NO RECORDS kineli Makolski, 1952

unipustulatus Bonelli, 1813

Badister unipustulatus Bonelli, 1813 4 WATSONIAN 3 (3, 0, 11, 0, 0) Notable B. Rather scarce with YORKSHIRE 2 records from only seven sites in the county. The species was recently located at Saltmarsh Delph 1 YWT reserve over several years, so it is apparently 0 well established there. There is a record for SE58 9 in (Luff, 1998), a record not on the YNU database. 8 (iv-vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

122 Subgenus TRIMORPHUS Stephens, 1828 sodalis (Duftschmid, 1812) humeralis Bonelli, 1813

Badister sodalis (Duftschmid, 1812) 4 WATSONIAN 3 (1, 3, 15, 4, 0) An infrequent species, with nearly YORKSHIRE 2 all records from litter in damp woodland. Exceptions 1 to this trend are Thorne Moor records from 0 Phragmites litter and from drain sides and a Studley Park record “from moss by hillside stream”. (ii-xi, 9 peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus BAUDIA Ragusa, 1884 collaris Motschulsky, 1844 NO RECORDS anomalus (Perris, 1866) striatulus Hansen, 1944

dilatatus Chaudoir, 1837 NO RECORDS

peltatus (Panzer, 1797) NO RECORDS

123 Tribe PANAGAEINI Bonelli, 1810

PANAGAEUS Latreille, 1802 bipustulatus (Fabricius, 1775) quadripustulatus Sturm, 1815

Panagaeus bipustulatus (Fabricius, 1775) 4 WATSONIAN (13, 0, 0, 0, 0) Notable B. All our records are 3 YORKSHIRE from Spurn dating from 1930 (Denton, 1995:17) 2 to 2001 when the species was found frequently 1 during an arthropod survey (MLD). (v-viii). 0 9 8 7 6 5 4

3

2 1 0 9 8 post-1990 records 7 Map produced by DMAP 6/1/2009 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cruxmajor (Linnaeus, 1758)

Panagaeus cruxmajor (Linnaeus, 1758) 4 WATSONIAN 3 (1, 0, 2, 0, 0) RDB Endangered/Vulnerable. Very YORKSHIRE rare. We have three records, only one of these 2 being represented in (Luff, 1998). The records 1 are: Eastoft SE8016, 1910 (“Crawshaw”) 0 (Donisthorpe 1913:203); Thorne Moor SE71, 1946 9 (Anon); Wheldrake Ings, SE6944, 9-23 vi 1992 8 (DC teste MLD), in pitfall trap. The typical habitat 7 is well-vegetated fens and near water. The species 6 may be nocturnal, feeding on molluscs at the edge 5 of water bodies. 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

124 Tribe PERIGONINI Horn, 1881

PERIGONA Laporte, 1835 nigriceps (Dejean, 1831)

Perigona nigriceps (Dejean, 1831) 4 WATSONIAN (1, 0, 0, 2, 0) Very rarely recorded. There are 3 YORKSHIRE three records on the YNU database: Aberford 2 SE4337, 27 viii 1989 (RBA det PS); Bishop 1 Monkton Reserve SE345657, 5 ix 1998 (RJM), in 0 sedge litter at edge of lagoon; East Newton 9 TA2638, 1 ix 1999 (WRD), in heap of rotting wild 8 oat straw. There is a record on the NBN Gateway 7 for SE42 which requires investigation. This species 6 could easily be passed over as a small Trechus in the field; for an interesting note and description 5 see Allen (1950b). 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe MASOREINI Chaudoir, 1870

MASOREUS Dejean, 1821 wetterhallii (Gyllenhal, 1813) NO RECORDS wetterhalli auctt. (misspelling) laticollis (Sturm, 1825)

125 Tribe LEBIINI Bonelli, 1810

LEBIA Latreille, 1802

Subgenus LAMPRIAS Bonelli, 1810 chlorocephala (Hoffmann, J., 1803) chrysocephala (Motschulsky, 1864)

Lebia chlorocephala (Hoffmann, J., 1803) 4 WATSONIAN 3 (3, 6, 2, 0, 0) Notable B. Very scarce. The larvae YORKSHIRE 2 are ectoparasites of chrysomelid pupae (Luff, 1998:167). Recorded in Yorkshire from sand pits 1 on the coast, by grubbing among grasses on 0 sandy cliffs, and inland, with habitats including 9 under bark with remains of Chrysolina polita. Only 8 one recent record: Cayton Bay TA0685, 1990, RC. 7 (iii-vi). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cyanocephala (Linnaeus, 1758) SEE APPENDIX A

Subgenus LEBIA Latreille, 1802 cruxminor (Linnaeus, 1758) NO RECORDS

marginata (Geoffroy in Fourcroy, 1785) NO RECORDS haemorrhoidalis (Fabricius, 1787)

scapularis (Geoffroy in Fourcroy, 1785) NO RECORDS turcica (Fabricius, 1787)

126 DEMETRIAS Bonelli, 1810

Subgenus RISOPHILUS Leach, 1815 AETOPHORUS Schmidt-Göbel, 1846 imperialis (Germar, 1824) NO RECORDS

Subgenus DEMETRIAS Bonelli, 1810 atricapillus (Linnaeus, 1758)

Demetrias atricapillus (Linnaeus, 1758) 4 WATSONIAN 3 (76, 14, 119, 21, 3) Widespread and with many YORKSHIRE records from the central, southern and eastern 2 parts of the county. The species tends not to be 1 recorded in upland localities. In a very wide range 0 of habitats including woodland pitfalls, in flood 9 refuse, by beating various foliage, in gardens and 8 in arable land margins, in Glyceria beds, compost 7 heaps, under stones, and by ponds and rivers. 6 (i-xii, peak vi). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

monostigma Samouelle, 1819 unipunctatus (Germar, 1824)

Demetrias monostigma Samouelle, 1819 4 WATSONIAN 3 (7, 1, 0, 0, 0) Notable B. Very scarce and mainly YORKSHIRE 2 (but not exclusively) coastal in Yorkshire. The species has occurred in several localities on the 1 Spurn peninsula from 1946 to 2006, also “Goole” 0 (indet location), 1902 (HHC); Bubwith Bridge 9 SE707364, 16 iv 1990 (MLD). (iv-viii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

127 CYMINDIS Latreille, 1806

Subgenus CYMINDIS Latreille, 1806 axillaris (Fabricius, 1794) NO RECORDS macularis Mannerheim in Fischer von Waldheim, 1824 NO RECORDS

Subgenus TARULUS Bedel, 1906 vaporariorum (Linnaeus, 1758) basalis Gyllenhal, 1810

Cymindis vaporariorum (Linnaeus, 1758) 4 WATSONIAN 3 (0, 9, 2, 0, 0) Notable B. Scarce, with only 11 YORKSHIRE 2 records on the database. A northern and boreo- 1 montane species, it has been recorded in 0 Yorkshire from bogs and wet moorland on high ground. There are no recent records. (vi-ix). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PARADROMIUS Fowler, 1887

DROMIUS sensu auctt. partim non Bonelli, 1810 linearis (Olivier, 1795)

Paradromius linearis (Olivier, 1795) 4 WATSONIAN 3 (100, 29, 159, 44, 6) Widespread with many YORKSHIRE 2 records, this is one of our most abundant and 1 frequently recorded carabids. The wide habitat range includes peat moors, lowland heaths, bogs, 0 canal and river sides, in gravel pits, all kinds of 9 ground litter, in dry grassland by sweeping, in 8 gardens, reed-beds and on arable margins. 7 Records indicate a reluctance to venture onto the 6 higher ground. (i-xii, peaks v and viii). 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

128 longiceps (Dejean, 1826)

Paradromius longiceps (Dejean, 1826) 4 WATSONIAN 3 (7, 0, 5, 0, 0) Notable A. Very scarce, and YORKSHIRE 2 recorded from saltmarshes and Phragmites beds. Our records are mainly estuarine (Humber) but 1 not exclusively. There are also records for Thorne 0 Moor SE71, 1950 (Anon); Inkle Moor SE7116, viii 9 1990 (RJM), in pitfalls and water traps in reed 8 beds; Shirebrook SK4284, 1982 (ST det AL). 7 (ii-ix). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

DROMIUS Bonelli, 1810 agilis (Fabricius, 1787) bimaculatus Dejean, 1825

Dromius agilis (F., 1787) 4 WATSONIAN 3 (2, 3, 10, 12, 0) Widely distributed in central and YORKSHIRE 2 southern Yorkshire but very infrequently recorded. 1 Noted from woodland situations, both coniferous and deciduous. (i-xii, peak vii). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

129 angustus Brullé, 1834

Dromius angustus Brulle, 1834 4 WATSONIAN (1, 6, 2, 2, 0) Another very infrequent Dromius, 3 YORKSHIRE this species is recorded from conifers, and in the 2 case of the Yorkshire records, Scots Pine is often 1 mentioned. The majority of our records are from 0 the winter months. (x-i). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

meridionalis Dejean, 1825 discus Puel, 1919

Dromius meridionalis Dejean, 1825 4 WATSONIAN (10, 4, 5, 8, 0) Widespread but with few records, 3 YORKSHIRE this species is arboreal, with Yorkshire records 2 from oak, ash, elm, alder and elder. Also reported 1 being beaten from pines: Skipwith Common SE63, 0 1937 (WJF) (see Nat 1938:97). One recent record 9 gives “shaken from gorse at roadside”. 8 (i-x, peak vi). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

130 quadrimaculatus (Linnaeus, 1758)

Dromius quadrimaculatus (Linnaeus, 1758) 4 WATSONIAN 3 (9, 19, 40, 23, 3) A very widespread and fairly YORKSHIRE 2 frequent arboreal species. Athough we have a few records from light traps, a malaise trap, by 1 general sweeping and in flood refuse, the majority 0 of the records indicate under bark of oak, hazel, 9 lime, beech, Scots Pine, alder, birch and apple. 8 (i-xii, peak ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

CALODROMIUS Reitter, 1905 DROMIUS sensu auctt. partim non Bonelli, 1810 spilotus (Illiger, 1798) quadrinotatus (Zenker in Panzer, 1800) non (Fabricius, 1798)

Dromius spilotus (Illiger, 1798) 4 WATSONIAN 3 (14, 12, 29, 8, 2) Very widespread and fairly YORKSHIRE 2 frequent, although less so than quadrimaculatus. 1 In pitfalls in woodland, in flood refuse, in piles of old straw at woodland edges and by old 0 hedgerows, on bramble, and subcortical on oak, 9 ash, ?elm, crab apple, beech and birch. Only one 8 record mentions conifers specifically: Eggborough 7 SE5724, 19 v 1993 (RJM); strange in the light of 6 the usually accepted range of host tree species. 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

131 PHILORHIZUS Hope, 1838 DROMIUS sensu auctt. partim non Bonelli, 1810 DROMIOLUS Reitter, 1905 melanocephalus (Dejean, 1825)

Philorhizus melanocephalus (Dejean, 1825) 4 WATSONIAN (59, 10, 101, 26, 0) Widespread with many 3 YORKSHIRE records in the southern half of the county, and 2 from a wide range of habitats that are not very 1 wet: gravel pits, woodland, grasslands, roadside 0 verges, post-industrial sites, coastal dunes, peat 9 moors, arable margins, straw heaps, decaying 8 vegetable refuse and mowings. 7 (i-xii, peak v). 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

notatus (Stephens, 1827) nigriventris (Thomson, C.G., 1857)

Philorhizus notatus (Stephens, 1827) 4 WATSONIAN 3 (12, 4, 0, 1, 0) Very infrequent and recorded YORKSHIRE 2 mainly from coastal sites, among dunes and dry 1 grassland. The few inland sites include Pannal 0 Ash SE3052, 1939 (RRUK); and Allerthorpe Common SE74, 30 iv 1927 (WJF). (iv-x). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

132 quadrisignatus (Dejean, 1825)

Philorhizus quadrisignatus (Dejean, 1825) 4 WATSONIAN (0, 0, 1, 0, 0) RDB Endangered. There is a single 3 YORKSHIRE record on the YNU database: Denaby Ings, 2 SE5000, iii 1966 (PS). The accepted habitat is in 1 ground litter associated with deciduous trees and 0 shrubs (Luff, 1998). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sigma (Rossi, 1790)

Philorhiz us sigma (Rossi, 1790) 4 WATSONIAN 3 (0, 1, 6, 4, 0) Notable A. Rare in Yorkshire with YORKSHIRE 11 records only. Recorded from bogs and around 2 flooded gravel pits. The four known localities in 1 the county include Askham Bog SE5748; Elland 0 Gravel Pits SE1121; Thorne Moor SE71; Throxenby 9 Mere TA0089 undated therefore not mapped. 8 (iii-ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

vectensis (Rye, 1873) NO RECORDS insignis sensu auctt. non (Lucas, 1846)

133 MICROLESTES Schmidt-Göbel, 1846 BLECHRUS Motschulsky, 1847 maurus (Sturm, 1827) glabratus sensu auctt. non (Duftschmid, 1812)

Microlestes maurus (Sturm, 1827) 4 WATSONIAN 3 (17, 0, 1, 0, 0) Predominantly estuarine and YORKSHIRE 2 coastal in our region with only two inland records 1 (in flood refuse and in a sand quarry). Generally in 0 flood and strand line refuse, reed-beds and dunes. (ii-ix). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 25/8/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

minutulus (Goeze, 1777) NO RECORDS glabratus (Duftschmid, 1812)

LIONYCHUS Wissmann, 1846 quadrillum (Duftschmid, 1812) SEE APPENDIX A bipunctatus (Heer, 1838) unicolor Schilsky, 1888

134 SYNTOMUS Hope, 1838 METABLETUS Schmidt-Göbel, 1846 foveatus (Geoffroy in Fourcroy, 1785) foveola (Gyllenhal, 1810)

Syntomus foveatus (Geoffroy in Fourcroy, 1785) 4 WATSONIAN (20, 6, 14, 1, 0) Our records indicate sandy 3 YORKSHIRE heaths, coastal dunes and a sand quarry in the 2 range of habitats. The species seems to favour 1 inland and lowland localities in Yorkshire. The 0 species can occur in large numbers when located. 9 (iii-ix, peak v). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

obscuroguttatus (Duftschmid, 1812) SEE APPENDIX A atratus (Dejean, 1825)

truncatellus (Linnaeus, 1761)

Syntomus truncatellus (Linnaeus, 1761) 4 WATSONIAN (7, 1, 5, 0, 0) With very few records, this species 3 YORKSHIRE has been noted from drain sides in farmland, on 2 coastal dunes, on dry heaths with Calluna, drier 1 parts of peat moors and in sand/gravel workings. 0 (iii-ix). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records 7 Map produced by DMAP 9/11/2008 for Yorkshire Naturali sts' Union pre-1990 records 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

135 Tribe ODACANTHINI Laporte, 1834

ODACANTHA Paykull, 1798 COLLIURIS sensu auctt. non De Geer, 1774 melanura (Linnaeus, 1767) NO RECORDS

Tribe DRYPTINI Schaum, 1857

DRYPTA Latreille, 1796 dentata (Rossi, 1790) NO RECORDS emarginata (Olivier, 1790)

Tribe ZUPHIINI Bedel, 1895

POLISTICHUS Bonelli, 1810 POLYSTICHUS auctt. (misspelling) connexus (Geoffroy in Fourcroy, 1785) NO RECORDS fasciolatus sensu auctt. non (Rossi, 1790) vittatus Brullé, 1834

APPENDIX A - Species considered inappropriate for inclusion in the Yorkshire list or for deletion from the Yorkshire list.

The following species appear on the Recorder 6 database and were transcribed from the paper records or have been received since that project was completed. These should be: a) considered for deletion from our county list on the basis that the records are anomalous with the known national or regional distribution and/or habitat requirements, and that voucher specimens which would be needed for confirmation are not forthcoming, or b) are considered by me to be errors in identification or interpretation of nomenclature (flagged as invalid in Recorder and hence not mapped), or c) have been received from bona fide sources but owing either to their rarity or the anomalous habitat associations, vouchers have not been seen by me for inclusion in this atlas (flagged as requiring confirmation in Recorder and therefore not mapped in this edition of the atlas).

These records remain on the database but flagged as invalid or requiring confirmation. Known introductions remain on the database. Comments on this policy in general, or on the species in particular, will be welcomed.

Tachys scutellaris - (0, 1, 0, 0, 0) There is only one record on the YNU database, Raincliffe Woods SE9988, before 1950 (GBW). The typical habitat is on mud in saltmarshes around the English coast from Devon to Norfolk (Luff, 1998) (Luff, 2007). In the absence of a voucher specimen this record must be considered an error, possibly for bistriatus, or a manuscript error where another species was intended, and should be deleted from the Yorkshire list. The distribution map has been retained in this report.

Bembidion schuppelii - (0, 0, 1, 1, 0) Notable A. There are two records on the YNU database: Barnsley SE30, date unspecified (EGB) and therefore not mapped; Lindley Wood Reservoir SE2249, 29 vii 1947 (Anon) {but see Nat 1948:??]

[CHECK]. Both of these records are rather unlikely judging from the accepted national distribution (Luff, 1998). The habitat is vegetated sand/gravel by upland streams and rivers.

Harpalus smaragdinus - (0, 1, 0, 0, 0) Notable B. The single record on the YNU database from Hutton-le-Hole SE7089, 1972 (DD) is considered invalid unless a voucher specimen is forthcoming. Two regionally local records are indicated in Luff (1998), one for SK69 which would require investigation to confirm whether this is a Yorkshire record, and for SE91 which lies in North Linclolnshire.

Ophonus rupicola – (1, 0, 0, 0, 0) Notable B. The single record on the YNU database, Brough SE9326, c.1880 (EBW) is considered unreliable and invalid, unless a voucher specimen is forthcoming. For this reason no map has been generated. [Luff, 1998:140 states that Moore (1957) includes Yorkshire and Lancashire so Moore may have seen the Wrigglesworth record]

136 Dicheirotrichus obsoletus - (1, 0, 0, 0, 0) Notable B. We have a single record for “Spurn” with no other details, 1901 (HEJ). To this record Fordham adds the comment “mud on Humber foreshore – a vague and possibly erroneous record”. The habitat is similar to gustavi but the species is much more restricted to south and south-east England. Old records exist for north Lincolnshire opposite Spurn, so the species could well occur on the northern bank of the Humber. A voucher for the above record would be very welcome: the record remains on the database for the time being.

Bradycellus distinctus - (0, 2, 0, 0, 0) Notable A. Rare in Yorkshire with only two records. Sandsend NZ8613, date unknown (HB); Upgang NZ8811, 1934 (HB). In the light of the now accepted distribution of validated records (Luff, 1998) both these instances are considered unlikely and may be referable to sharpi in the absence of vouchers.

Acupalpus brunnipes - (0, 0, 1, 0, 0) Notable A. The single record is from subfossil peat deposits: Thorne Moor SE7117, 1993 (NW), in a 1993 peat sampling survey. Nowadays this is an insect from the far south of Britain.

Chlaenius tristis - (1, 0, 0, 0, 0) RDB – Endangered. There is a single record on the YNU database: Hornsea TA24, 1826 (“Grimston”) “specimens being taken in 1826”. [The site was probably around Hornsea Mere.] Now the only reliable locality for this species is the Llyn peninsula, North Wales (Luff, 1998 and Luff, 2007). However it was known from the East Anglian fens before 1900, so Hornsea was perhaps a possibility (but where are the specimens?).

Lebia cyanocephala - (0, 0, 0, 1, 0) RDB Endangered. There is but a single record on the YNU database: Malham Cove SD8964, 1 ix 1883 (Anon), with the comment “Error – WJF”. In the absence of a supporting voucher specimen or other evidence the record must be considered invalid, but is nonetheless mapped here.

Lionychus quadrillum - (0, 1, 0, 0, 0) RDB Rare. We have only a single record on the YNU dadabase: Raincliffe Woods SE98, 1870 (RLa). Nationally the species is one of sand and shingle near running water (Luff, 1998).

Syntomus obscuroguttatus - (0, 0, 1, 0, 0,) A single record of this species exists in the YNU database: Thorne Moor SE71, 1978 (Anon). I do not know where this record came from. It does not figure in (Luff, 1998) or in (Skidmore, 2006), and lies well north of the northern limit of its accepted range in Britain. Do any fellow coleopterists know why this record should be maintained in our list?

Amara anthobia - (0, 0, 2, 0, 0) There are only two records for this species on the YNU database: Beighton Ponds SK4483, 1988 (AL); Lindholme SE6805, 22 v 2007 (AL). The normally accepted habitat is open sandy sites and dunes. Vouchers require examination before a map is produced. ly HELOPHORIDAE Leach, 1815 57

Acknowledgements

The recording system could not function effectively without the input from Yorkshire coleopterists (and other workers further afield) past and present. I am very grateful to a „hard core‟ of present members of the YNU Entomological Section for passing their records to me for inclusion in the database, and for keeping the database up to date. In this respect I thank Bill Dolling, Frank Kenington, Peter Kendall, Mike Denton, and Roy Crossley. I am also grateful to Colin Johnson and the late Dr. Peter Skidmore for giving patiently of their time and expertise in identifying/checking many specimens (some in very battered condition) presented to them over many years. Their help and encouragement in my study of coleopterology has been outstanding. I also thank Derek Whitely and Eric Smith of the Sorby Natural History Society, Sheffield, for passing on records to me regularly. To Dmitri Logunov and Philip Rispin of Manchester University Museum for use of facilities, access to Coloeoptera collections and searching for specimens and literature references for me. Martin Limbert for continued access to Coleoptera collection at Doncaster Museum. To Mike Denton for carrying out a proof-reading exercise for me and pointing out numerous anomalies and inconsistencies in the text, and providing advice and sharing of his expertise over many years. Any omissions or mistakes in the text or in the mapping are my responsibility. Any errors which come to the notice of readers will, it is hoped, be fed back to the author.

References

Allen, A. A., 1950a. A second English capture of Trechus subnotatus Dej. (Col., Carabidae). Entomologist‟s Mon. Mag. 86 (1950):38.

Allen, A.A. 1950b. Two species of Carabidae (Col.) new to Britain. Entomologist‟s Mon. Mag. (1950) 86:89-90.

Anderson, R. & Luff, M.L. 1994. Calathus cinctus Motschulsky, a species of the Calathus melanocephalus/mollis complex (Col., Carabidae) in the British Isles. Entomologist‟s Mon. Mag. 130:131-135.

Cooter, J. 1999.. On „English‟ names for British Coleoptera. Coleopterist 8 (1): 43-45.

137

Crowson, R.A. 1942. Leistus rufomarginatus Duft. (Col., Carabidae) new to Britain. Entomologist‟s Mon. Mag. 78:281-282.

Coulson, J. & Butterfield, J.E.L. 1979. Geographical characterisation of moorland using invertebrates. Dept of Zoology, University of Durham.

Dandy J.E. 1969. Watsonian Vice-counties of Great Britain. The Ray Society, London

Denton, M.L. 1990. Entomogical Reports for 1988-1989. Coleoptera: Staphylinidae (Aleocharinae). Naturalist 115 (1990): 97-102.

Denton, M.L. 1995. The Beetles of Spurn Peninsula. Yorkshire Museum.

Denton, M.L. 2003. The Aleocharinae (Coleoptera: Staphylinidae) of Yorkshire. Naturalist 128 (2003): 103-125.

Denton, M.L. 2008. Entomological records for 2003-2007. (Coleoptera:Staphylinidae (Aleocharinae). Naturalist 133 (2008): 96-100 .

Denton, M.L. & Marsh, R.J. 1995. Lower Derwent Coleoptera survey – unpublished.

Denton, M.L. & Marsh, R.J. 2006. The Coleoptera (Beetles) of the Lower Derwent Valley. In Land Use, Ecology and Conservation in the Lower Derwent Valley. Ed. Tim Milsom. PLACE, York.

Duff, A.G. 2006. Coleoptera Checklist of the British Isles. Coleopterist website www.coleopterist.org.uk

Edelsten, H.M. 1941. Brief note on Carabus auratus. Entomologist‟s Mon. Mag. 77 (1941): 107.

Flint, J.H. 1988. Entomological reports for 1984-1986: Coleoptera. Naturalist 113 (1988): 69-72.

Fowler W.W. 1882. (Entomologist 15: 149-152) Natural Localities of British Coleoptera: No. V – the sea-coast.

Freude, Harde and Lohse (2004). Die Kafer Mitteleuropas 2 (2004):403. Elsevier, Munchen.

Hodge, P. J. & Jones, R.A. 1995. New British Beetles – Species not in Joy‟s practical handbook. BENHS, Reading.

JNCC, 1992. Hyman, P.S., and Parsons, M.S. A Review of the scarce and threatened Coleoptera of Great Britain. Part 1. Peterborough.

JNCC, 1994. Hyman, P.S., and Parsons, M.S. A Review of the scarce and threatened Coleoptera of Great Britain. Part 2. Peterborough.

Key, R.S. 1999. More on „noddy names‟ for British beetles. Coleopterist 8 (2): 94.

Lindroth, C.H. 1974. Handbks. Ident. Brit. Insects 4 (2). Coleoptera – Carabidae. Royal Entomological Society, London.

Luff, M.L. 1998. Provisional Atlas of the ground beetles (Coleoptera, Carabidae) of Britain. Huntingdon: Biological Records Centre.

Luff, M.L. 2007 The Carabidae (ground beetles) of Britain and Ireland (2nd edition). Handbks. Ident. Brit Insects vol 4 pt. 2.

Marsh, R.J. 1991. Coleoptera records for Yorkshire: A review and update. Naturalist 116 (1991): 101-106.

Marsh, R.J. 2002. Coleoptera report for 1995-2001. Naturalist 127 (2002): 69-80.

Moore B.P. 1957. The British Carabidae (Coleoptera), part II: the county distribution of the species. Entomologist‟s Gazette 8:171-180.

Parry, J.A. 1975. Dyschirius angustatus Ahrens (Col., Carabidae) in East Sussex. Entomologist‟s Mon. Mag. 111:160.

Price, A. 1968. Brief note on Carabus auratus. Entomologist‟s Mon. Mag. 104 (1968): 229.

Pope, R.D. 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbks. Ident. Brit. Insects 11(3), pp. xiv+105. Royal Entomological Society, London.

Shirt, D.B., ed. 1987. British Red Data Books: 2. Insects. Nature Conservancy Council, Peterborough.

Skidmore, P. 1971. The Insect fauna of a bog oak found near Askern. Naturalist 171:111-112.

138

Skidmore, P. et al 1985. The Insects of Thorne Moor. Sorby Record, No 23.

Skidmore, P. 2006. An inventory of the Invertebrates of Thorne and Hatfield Moors. Thorne and Hatfield Conservation Forum, Doncaster.Proc. Trans. BENHS. 19:17-21.

Speight, M.C.D., Martinez, M. & Luff, M.L. 1986. The Asaphidion (Col.:Carabidae) species occurring in Great Britain and Northern Ireland.

Watson, H.C. 1883. 2nd. ed. Topographical Botany. Quaritch, London.

R J Marsh 18 October 2009.

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