Saurolophini Saurolophus Anatotitan ? Tanius Edmontosaurus Phylogeny of Ornithodira Edmontosaurini Shantungosaurus Hadrosaurini Gryposaurus ? Thespesius ? Aralosaurus Microhadrosaurus ? Kritosaurus Hadrosaurinae Maiasaurini Maiasaura Hadrosaurus
Euhadrosauria Lambeosaurinae Pteropelyx Brachylophosaurus Parasaurolophus Telmatosaurus ? Gadolosaurus Arambourgiania Azhdarcho Gilmoreosaurus ? Arstanosaurus Diopocephalus Claosaurus ? Bactrosaurus Charonosaurus ? Bennettazhia Protohadros Unofficially named clades RANK Secernosaurus LEGEND ? Doratorhynchus Orthomerus Daughter clade Montanazhdarcho Mandschurosaurus Parasaurolophini ? Tsintaosaurus Above genus Quetzalcoatlus Hypsibema "Daughter clade" Azhdarchidae Zhejiangopterus Hadrosauridae ? Hironosaurus Tree root Daughter clade Lambeosaurini Nipponosaurus Genus and below ? Amurosaurus "Daughter clade" Tupuxuara Nanyangosaurus Jaxartosaurus Parent clade Daughter clade Ouranosaurus Lambeosaurus Daughter clade Azhdarchoidea Tapejara Hadrosauroidea Altirhinus ? Gnathosaurus Iguanodon Unnamed daughter clade Barsboldia Craspedodon Certain Daughter clade Dsungaripteridae Dsungaripterus Iguanodontoidea "Camptosaurus" sp. Corythosaurus Puntanipterus "Gnathosaurinae" Plataleorhynchus ? Lurdusaurus PLACEMENT WITHIN Less certain Daughter clade Pterodaustro Anhanguera Fukuisaurus Hypacrosaurus ? Noripterus Planicoxa Uncertain Cycnorhamphus "Phobetor" Ctenochasma Styracosterna ? Eolambia Diceratops PARENT CLADE Eosipterus Ctenochasmatidae Huanhepterus Camptosauridae Draconyx Very uncertain Kepodactylus Anhanguerinae Arthurdactylus Mesadactylus Pterodactylidae ? Haopterus "Coloborhynchus" clavirostris Bihariosaurus ? Normannognathus ? Cearadactylus Ankylopollexia Pararhabdodon Probactrosaurus Triceratops Herbstosaurus ? Germanodactylus Camptosaurus Dermodactylus Tropeognathus ? Anabisetia Wyomingopteryx Pterodactylus Criorhynchus Pterodactyloidea Nyctosaurus Anhangueridae Siroccopteryx Dryomorpha Dryosauridae Dryosaurus Torosaurus Istiodactylus Parapsicephalus ? Bogolubovia Gasparinisaura ? Kangnasaurus Pteranodon Angustinaripterus Ornithocheiroidea Pteranodontoidea Ornithostoma Euiguanodontia ? Loncosaurus Valdosaurus Arrhinoceratops ? "Mandschurosaurus" laosensis ? Muttaburrasaurus Rhamphorhynchidae Rhamphocephalus Ornithocheiridae Ornithocheirus Jinzhousaurus Anchiceratops ? Araripesaurus Lophorhothon Chasmosaurini Pentaceratops Brasileodactylus Iguanodontia Tenontosaurus ? Araripedactylus Ugrosaurus Chasmosaurus Campylognathoididae Eudimorphodon Rhamphorhynchus Santanadactylus Hypsilophodontia Hypsilophodon Ceratopsinae Polyonax Claorhynchus ? Ceratops Euornithopoda Parksosaurus Rhabdodon Agathaumas Centrosaurus Dysganus Dimorphodontidae Dimorphodon Campylognathoides Othnieliidae "Yandusaurus" multidens Drinker Ceratopsidae Centrosaurinae Avaceratops
? Nesodactylus Zephyrosauridae Orodromeus Centrosaurini Styracosaurus "Anoplosaurus" tanyspondylus Dorygnathus Peteinosaurus Notohypsilophodon Othnieliinae Othnielia Turanoceratops Pachyrhinosaurini Brachyceratops Atlascopcosaurus Gravitholus ? "Odontorhynchus" Fulgurotherium Zephyrosaurus Archaeoceratops Monoclonius Leaellynasaura ? Kulceratops Prenocephale Scaphognathus ? Sanpasaurus ? Chaoyangsaurus Ceratopsomorpha Zuniceratops Einiosaurus Qantassaurus Microceratops Tylocephale ? Stygimoloch Yandusaurus Graciliceratops ? Tichosteus Microcephale Sordes Nanosaurus Leptoceratops Ceratopsoidea Montanoceratops Achelousaurus Siluosaurus "Echizensaurus" Udanoceratops Protoceratops Pachycephalosaurinae Pachycephalosaurini Pachycephalosaurus Thescelosaurus Trachodon Magulodon Stegosaurus Phyllodon Asiaceratops Breviceratops Anurognathidae Anurognathus Ornithopoda Bugenasaura Neoceratopsia Coronosauria Bagaceratops Pachyrhinosaurus Comodactylus Marginocephalia Ceratopsia Psittacosauria Psittacosaurus Pachycephalosauridae Stegoceras Laopteryx ? Dendrorhynchoides Stenopelix Stegosaurini Wuerhosaurus Euoplocephalus Rhamphinion Notoceratops Lexovisaurus Pterosauria Preondactylus Batrachognathus Cerapoda Heterodontosauridae Lanasaurus Xuanhuasaurus ? Thecospondylus Geranosaurus Pachycephalosauria Wannanosaurus Homalocephaloidea Homalocephale ? Paranthodon Jeholosaurus Stegosaurinae Tuojiangosaurus Pinacosaurus ? Dianchungosaurus Yaverlandia Micropachycephalosaurus Amtosaurus Pterosauromorpha ? Sharovipteryx Echinodon Heterodontosaurinae Abrictosaurus Chialingosaurus Shanxia ? Avipes Taveirosaurus Goyocephalia Goyocephale Genasauria Thyreophora Scutellosaurus Huayangosauridae Huayangosaurus Chungkingosaurus Maleevus Scleromochlus Fabrosaurus Gigantspinosaurus Talarurus Ornithodira Dinosauromorpha Lagerpeton Lesothosaurus Bienosaurus Heterodontosaurini Heterodontosaurus "Katsuyamakensaurus" Syrmosaurini Saichania ? Gongbusaurus Regnosaurus Agilisaurus Thyreophoroidea Tatisaurus Kentrosaurus ? Nodocephalosaurus Pisanosaurus Emausaurus Changtusaurus Monkonosaurus Dinosauriformes Lagosuchus Pekinosaurus Lycorhinus Craterosaurus ? Ankylosaurus Tarchia Galtonia Lusitanosaurus ? Hesperosaurus Yingshanosaurus Marasuchus Lucianosaurus Camelotia Eurypoda Stegosauria Stegosauridae Dacentrurus Sauroplites Revueltosaurus Brachypodosaurus Pseudolagosuchus Technosaurus Ankylosaurinae ? Tsagantegia Gobisaurus Tecovasaurus Scelidosaurus Trimucrodon Azendohsaurus Melanorosauridae Melanorosaurus Ankylosauria Cryptosaurus Xiaosaurus Massospondylidae Massospondylus Minmi Dinosauria Ornithischia Alocodon ? Thecodontosaurus Lufengosaurus Heishansaurus Shamosaurinae ? Shamosaurus Luanpingosaurus Blikanasaurus Peishansaurus Gargoyleosaurus Mymoorapelta Sauropoda Cardiodon ? Coloradisaurus Priconodon Teyuwasu Saturnalia Lessemsaurus Ultrasaurus Yunnanosaurinae Yunnanosaurus Priodontognathus Saurischia Eshanosaurus Thotobolosaurus Asiatosaurus Tianzhenosaurus Cedarpelta "Gyposaurus" sinensis Euskelosaurus Gongxianosaurus ? Tyreophorus Polacanthinae Hylaeosaurus Sauropodomorpha ? Nyasasaurus Microdontosaurus Kunmingosaurus ? Stegosaurides Chinshakiangosaurus Tendaguriidae Tendaguria Jingshanosaurus ? Rhadinosaurus Tianchisaurus ? Acanthopholis Theropoda ? Arctosaurus Isanosaurus Ankylosauroidea Ankylosauridae Aletopelta ? Carnosaurus Yimenosaurus Polacanthini Gastonia ? Coelurosaurus Riojasaurus ? Sellosaurus ? Kotasaurus Stegopeltini Stegopelta ? Spinosuchus Mussaurus ? Velocipes Ruehleia Vulcanodon Nodosauridae ? Struthiosaurus Sinosaurus Anchisaurus ? Anoplosaurus Hoplitosaurus Protoavis Plateosauridae Plateosaurus ? Dracopelta Glyptodontopelta Eoraptor ? Nodosaurus Herrerasauria Herrerasauridae Chindesaurus Eusauropoda ? Oshanosaurus ? Palaeoscincus Herrerasaurinae Herrerasaurus Shunosaurus "Pleurocoelus" altus ? Texasetes Polacanthus Barapasaurus Euhelopus ? Sarcolestes Staurikosauridae Staurikosaurus Dachongosaurus Pawpawsaurus ? Guaibasauridae Guaibasaurus Caseosaurus ? Ohmdenosaurus Protognathosaurus ? Liaoningosaurus ? Zizhongosaurus "Morosaurus" agilis Edmontoniinae ? Niobrarasaurus Animantarx Datousaurus Tienshanosaurus Panoplosaurinae Panoplosaurus Lancanjiangosaurus Neotheropoda Magnosaurus ? Aliwalia Lapparentosaurus Edmontonia ? Alwalkeria Tehuelchesaurus Mamenchisaurus Sauropelta Limaysaurus ? Liassaurus Chuanjiesaurus ? Hudiesaurus ? Merosaurus Klamelisaurus ? Sinocoelurus Rhoetosaurus ? Nurosaurus ? Teinurosaurus Mamenchisauridae Omeisaurus Silvisaurus Rebbachisaurinae Rebbachisaurus ? Calamospondylus Amygdalodon ? Histriasaurus Altispinax ? Volkheimeria ? Cetiosaurus Capitalsaurus Rayososaurus Amargasaurus Chuangdongocoelurus Rebbachisauridae Nigersaurus Coeluroides ? Megacervixosaurus "Elaphrosaurus" gautieri ? "Rebbachisaurus" tasmenensis ? Mongolosaurus Dystylosaurus "Elaphrosaurus" iguidensis Diplodocoidea Dicraeosauridae Dicraeosaurus Embasaurus Jobaria Amphicoelias Supersaurus Inosaurus Losillasaurus Kagasaurus ? Patagosaurus Neosauropoda Diplodocimorpha ? Hisanohamasaurus Seismosaurus Katsuyamasaurus Cetiosauriscus Diplodocidae Dinheirosaurus Kelmayisaurus Syntarsus Qinlingosaurus Dystrophaeus Diplodocus Podokesaurus Agustiniidae Agustinia Dyslocosaurus "Megalosaurus" andrewsi ? Pterospondylus Eucoelophysis "Titanosaurus" rahioliensis Diplodocinae Barosaurus "Megalosaurus" chubutensis ? Shuvosaurus Coelophysis Haplocanthosaurus Apatosaurinae Eobrontosaurus "Megalosaurus" insignis Halticosaurus Coelophysidae Camposaurus Macronaria Sugiyamasaurus ? Aragosaurus "Megalosaurus" pannoniensis Gojirasaurus Chondrosteosaurus "Megalosaurus" pombali Dolichosuchus Bashunosaurus Camarasaurus "Megalosaurus" terquemi Dilophosaurus ? Abrosaurus Apatosaurus Ceratosauria Coelophysoidea Coelophysidae Procompsognathinae Procompsognathus Camarasauromorpha Camarasauridae ? Lourinhasaurus Brachiosaurus Mifunesaurus ? Ligabueino Cedarosaurus Newtonsaurus Elaphrosaurus Noasaurus Ngexisaurus Giraffatitan "Poekilopleuron" schmidti Neoceratosauria Sarcosaurus Liliensternus Ilokelesia Segisaurus Velocisaurus Titanosauriformes Brachiosauridae Sauroposeidon Prodeinodon Genusaurus Compsosuchus Noasauridae Masiakasaurus ? Austrosaurus Quilmesaurus Dryptosauroides ? Bellusaurus "Sidormimus" ? Dandakosaurus Genyodectes ? Damalasaurus Sigilmassasaurus Ceratosaurus Bothriospondylus "Tsuchikurasaurus" Abelisauria Abelisauridae Abelisaurinae Abelisaurus "Ischyrosaurus" Wakinosaurus Ornithomimoides Carnotaurinae Majungasaurus Ornithopsis Walgettosuchus Jubbulpuria Betasuchus Aucasaurus Pelorosaurus Tetanurae "Allosaurus" sp. Abelisauroidea Laevisuchus Spinosaurus Carnotaurus Venenosaurus Becklespinax Xenotarsosaurus Indosaurus ? Astrodon Eustreptospondylus Tarascosaurus Majungatholus Giganotosaurus ? Atlasaurus Kaijiangosaurus Indosuchus ? Eucamerotus Metriacanthosaurus Spinosaurinae Irritatorini Angaturama Carcharodontosaurus ? Pleurocoelus Piatnitzkysaurus Sonorasaurus Streptospondylus ? Bahariasaurus Titanosauria ? Pleurocoelus sp. Unquillosaurus Aegyptosaurus Walkersaurus Spinosauridae Baryonychinae Cristatusaurus Irritator Algoasaurus Valdoraptor Campylodoniscus ? Saltriosaurus Fukuiraptor Suchomimus ? Carcharodontosaurinae ? Neovenator ? Bruhathkayosaurus ? "Zupaysaurus" Cryolophosaurus Spinosauria ? Siamosaurus Baryonyx Allosaurinae Allosaurus ? Iuticosaurus Torvosauridae Brontoraptor Monolophosaurus ? Clasmodosaurus Erectopus Poekilopleuron Allosauroidea Allosauridae Acrocanthosaurus Wyomingraptor ? Aepisaurus Megalosaurus Chubutisaurus Chilantaisaurus Torvosaurus Sinraptoridae Szechuanosaurus ? Antrodemus ? Macrurosaurus ? Afrovenator Edmarka Yangchuanosaurus ? Huabeisauridae Huabeisaurus Neotetanurae Carnosauria ? Xuanhanosaurus Lourinhanosaurus ? "Titanosaurus" falloti Tugulusaurus ? Gasosaurus Sinraptor ? Phuwiangosaurus Marshosaurus ? Tangvayosaurus Coelurosauria "Aniksosaurus" Andesaurus Beelemodon ? Araucanoraptor "Chilantaisaurus" maourtensis Struthiomimus Paralititan "Ornithomimus" affinis Sinornithoides Ozraptor Ornithomimus Pellegrinisauridae Pellegrinisaurus Piveteausaurus Tochisaurus Janenschia Bagaraatan Byronosaurus Ornithomimini Dromiceiomimus Epachthosaurus Deltadromeus Magyarosaurus Maniraptoriformes ? "Aristosuchus" sp. Borogovia Anserimimus Troodontidae ? Koparion Saurornithoides Jainosaurus ? Diplotomodon ? Paronychodon Ornithomiminae Gallimimus Hypselosaurus ? Labocania Archaeornithomimus Ampelosaurus Archaeornithoides ? Deinocheirus Lirainosaurus Nemegtosaurus Bradycneme Troodon ? Euronychodon ? Sanchusaurus Argentinosaurus Quaesitosaurus Avimimidae Avimimus Orcomimus Albertosaurus Daspletosaurus Calamosaurus Ornithomimidae Garudimimus Tarbosaurus ? Nemegtosauridae Rapetosaurus Elopteryx Gorgosaurus Tyrannosaurus Laplatasaurus Harpymimus ? Shanshanosaurus Elmisaurus ? Malawisaurus Heptasteornis ? Kakuru Tyrannosaurini ? Aublysodon ? Antarctosaurus ? Gondwanatitan Arctometatarsalia Ornithomimosauria Pelecanimimus Conchoraptor Argyrosaurus ? Coeluridae Coelurus Alioramus Oviraptor Alamosaurus ? Compsognathidae ? Aristosuchus Citipati Caenagnathinae Chirostenotes Titanosaurus "Ornithomimus" sp. Tyrannosaurinae Deinodon Ingenia Aeolosaurus Ricardoestesia Compsognathus Oviraptoridae Khaan Saltasauridae ? Opisthocoelicaudinae Opisthocoelicaudia Phaedrolosaurus Chingkankousaurus Ginareemimus Sinosauropteryx Caenagnathidae Caenagnathasia Enigmosaurus Tyrannosauroidea Tyrannosauridae Alectrosaurus Erlikosaurus ? Arkansaurus ? Futabasaurus Oviraptorosauria ? Nomingia Nanshiungosaurus Saltasaurinae Neuquensaurus ? Dryptosaurus ? Tonouchisaurus Segnosaurus ? Nqwebasaurus ? Siamotyrannus Alxasauridae Alxasaurus Therizinosaurinae Therizinosaurus ? Nedcolbertia ? Ornitholestes ? Stokesosaurus Microvenator ? "Nanshiungosaurus" bohlini Saltasaurini Rocasaurus ? Proceratosaurus ? Iliosuchus ? Santanaraptor Eotyrannus ? Caudipteryx Therizinosauroidea Therizinosauridae Nothronychus ? Scipionyx ? Itemirus "Chilantaisaurus" zheziangensis Maniraptora ? Palaeopteryx Thecocoelurus Alvarezsaurus Patagonykus Mononykus Saltasaurus ? Timimus "Neimenggusaurus" "Ornithomimus" minutus Therizinosauria ? Beipiaosaurus Protarchaeopteryx Parvicursor ? Shanyangosaurus Alvarezsauridae Alvarezsaurinae Shuvuuia
Dromaeosauridae Dromaeosaurinae Dromaeosaurus Paraves Alvarezsauria ? Rapator ? Koreanosaurus Avisaurus ? Megaraptor ? Ornithodesmus Adasaurus Velociraptorinae Deinonychus Eumaniraptora Rahonavis Achillobator Soroavisaurus Sinornithosaurus Nuthetes Saurornitholestes Microraptor Bambiraptor Pyroraptor Velociraptor Unenlagia Utahraptor Avisauridae Neuquenornis Deinonychosauria Variraptor Avialae ? Wyleyia ? Concornis Aves Archaeopterygiformes Archaeopteryx
Yungavolucris
? Hulsanpes Changchengornis ? Largirostrisornis ? Yandangornis Confuciusornis Lectavis ? Iberomesornis Pygostylia Confuciusornithidae Proornis Cuspirostrisornithidae Cuspirostrisornis Kizylkumavis Catenoleimus Lenesornis Gobipteryx Sazavis Zhyraornithidae Zhyraornis Longchengornis Explorornis Alexornis Abavornis Gobipterygidae ? Nanantius Holboltia Boluochia Sinornis Incolornis Eoalulavis Horezmavis ? Jibeinia Gurilynia Hesperornis ? Longipteryx Sinornithiformes Cathayornis ? Noguerornis Eoenantiornithiformes Eoenantiornis ? Protopteryx Enantiornithiformes Enantiornis Ornithothoraces Enantiornithes ? Liaoxiornis Chaoyangiidae Chaoyangia Hesperornithinae Parahesperornis Platanavis Parascaniornis
? Vorona Asiahesperornis Euornithes Chaoyangiiformes ? Songlingornithidae Songlingornis Hesperornithidae Coniornis Patagopterygiformes Patagopteryx Gansuiformes Gansus Judinornis Ambiortiformes Ambiortus ? Kuszholia Baptornis ? Liaoningornis ? Gargantuavis ? Yanornis Struthioniformes ? Yixianornis ? Otogornis Ornithurae Hesperornithiformes Pasquiaornis
Rheiformes
Apsaravis Enaliaornis ? Apatornis Casuariiformes Psittaciformes
Carinatae Ichthyornithiformes Ichthyornis
Struthiones Aepyornithiformes Columbiformes
Limenavis
Ratitae Apteryges Apterygiformes ? Pteroclidiformes Tinamiformes ? Lithornithiformes ? Eleutheronithidae ? Dinornithiformes Eremopezus ? Dromornithiformes Turnicidae Limenavis ? Sternidae Neornithes Palaeognathae Stromeria ? Anseriformes ? Phalaropodidae "Charadriida" "Scolopacida" Laridae REFERENCES Neognathae Galloanserae ? Galliformes Charadriiformes Stercorariidae Sphenisciformes • Keesey, TM. "The Dinosauricon", http://www.dinosauricon.com (DEFUNCT). Retrieved 2005-05-16. Mesitornithidae Burnithidae Dromadidae Ciconiimorphae Phoenicopteriformes Gaviiformes
Charadriomorphae Graculavidae ? Ardeidae ? Podicepiformes ? Rhynchoceti Ciconiiformes Procelariiformes Eurypygoidea Otides Gruimorphae Gruiformes Cariamae ? Falconiformes Pelecaniformes
? Opisthocomiformes
? Cuculiformes ? Musophagiformes Caprimulgiformes
? Accipitriformes Apodiformes ? Pandionidae ? Sagittariidae Anomalogonatae Strigiformes Coliiformes Zygodactylus Sandcoleiformes Coraciiformes Renato Filipe Vidal Santos Trogoniformes Bucerotimorphae Bucerotiformes
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