ACCEPTED MANUSCRIPT MANUSCRIPT ACCEPTED ACCEPTED MANUSCRIPT 1 Taxonomic revision of the genus Ichthyosarcolites Demarest, 1812, and description of a 2 new canaliculate rudist from the Cenomanian of Slovenia: Oryxia sulcata gen. et sp. 3 nov. (Bivalvia, Hippuritida). 4 Valentin Rineau and Loïc Villier 5 UMR 7207 CR2P, Sorbonne Université/CNRS/MNHN/UPMC, 43 rue Buffon, F-75231 6 Paris cedex 05, France 7 Corresponding author: Valentin Rineau. E-mail:
[email protected] MANUSCRIPT ACCEPTED 1 ACCEPTED MANUSCRIPT 8 Taxonomic revision of the genus Ichthyosarcolites 9 Desmarest, 1812, and description of Oryxia sulcata gen. 10 et sp. nov. (Bivalvia, Hippuritida), a new canaliculate 11 rudist from the Cenomanian of Slovenia: 12 Abstract 13 Ichthyosarcolites was amongst the first rudist genera to be described. Nineteen species 14 have been assigned to this genus since its introduction in 1812, all from shallow-marine 15 tropical carbonates of Albian and Cenomanian age (mid-Cretaceous). Almost all 16 nominal species suffer from vague original descriptions, and some forms were 17 inaccurately assigned to the genus. Several species were defined on the basis of the 18 number of flanges along the shell, such as Ichthyosarcolites rotundus , I. monocarinatus , 19 I. triangularis , I. bicarinatus , I. tricarinatus , I.MANUSCRIPT alatus and I. polycarinatus . An analysis 20 of the relative position of the flanges on the shell by hierarchical clustering helps with 21 taxon definition. Two species with a single flange are here recognised, one with a dorsal 22 flange ( I. monocarinatus ), the other with a ventral one (I. triangularis ). There is no 23 consistency in flange distribution on the shells whatever their number is, i.e., two or 24 more, and homology of individual flanges cannot be demonstrated.