POLYCHAETOUS ANNELIDS ERECTED BY TREADWELL, 1891 TO 1948, TOGETHER WITH A BRIEF CHRONOLOGY

OLGA HARTMAN

BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY VOLUME 109: ARTICLE 2 NEW YORK 1956 POLYCHAETOUS ANNELIDS ERECTED BY TREADWELL, 1891 TO 1948, TOGETHER WITH A BRIEF CHRONOLOGY POLYCHAETOUS ANNELIDS ERECTED BY TREADWELL, 1891 TO 1948, TOGETHER WITH A BRIEF CHRONOLOGY

OLGA HARTMAN

CONTRIBUTION NUMBER 162 OF THE ALLAN HANCOCK FOUNDATION, LOS ANGELES

BULLETIN OF THE AMERICAN MTYSETUM OF NATURAL HISTORY VOLUME 109 : ARTICLE 2 NEW YORK 1956 BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Volume 109, article 2, pages 239-310, plate 21 Issued April 23, 1956 Price: $1.00 a copy CONTENTS INTRODUCTION ...... 245 ALPHABETICAL LIST oF NAMES OF ANNELIDS ERECTED BY TREADWELL 247

ALPHABETICAL LIST OF VALID SPECIES, WITH SYNONYMS ...... 260 LIST OF SPECIES BY FAMILIES ...... 265

SYSTEMATIC DISCUSSION ...... 271 REFERENCES ...... 302

243

INTRODUCTION THE PURPOSE OF THE PRESENT REPORT is to field investigations at the Bermuda Biological assemble the contributions to annelidan litera- Station during 1916 and was stationed for a ture of the late Aaron Treadwell (1866-1947) time at the Dry Tortugas Laboratory in south- during the 58 years (1891-1948) of his publi- ern Florida. During the early part of 1920 cations on the polychaetous annelids. The type Treadwell was a member of the exploring party specimens on which these studies are based are of the Carnegie Institution of Washington to deposited largely in the American Museum of Samoa and Fiji islands, under the direction of Natural History in New York City and in the Alfred Goldsborough Mayor. United States National Museum in Washing- Treadwell held memberships in the Ameri- ton, D.C. Through the generosity of the ad- can Association for the Advancement of Sci- ministration of the American Museum of ence, of which he was a Fellow, the American Natural History it was possible to examine all Society of Zoologists, the American Society of the type collections that were deposited there, Naturalists, and the scholastic society, Phi Beta and those in the New York Zoological Society Kappa. which are now also in the American Museum. At least three students were inspired by Many of the types in the United States Na- Treadwell to publish on annelids: Ruth Hoag- tional Museum had been earlier examined by land, who published on the annelids of the me at Washington, as indicated in the biblio- Philippine Islands (1919) and Puerto Rico graphic citations. The lists below, therefore, (1920); Agnes Mullin, who described some include all the names erected by Treadwell and annelids (1923) of the Barbados-Antigua Ex- give such identities as are believed to be valid. pedition of the University of Iowa; and Jane Some indicated as not seen by me are here Winternitz who reported on some annelids given to complete the lists. (1936) from Florida. Aaron Louis Treadwell was born in Redding, Throughout his life Treadwell maintained a Connecticut, on December 23, 1866. He received youthful eagerness for the new or unknown. a baccalaureate degree in 1888 and master of He had the confidence of higher authority and science 'in 1890 from Wesleyan University in was entrusted with valuable collections from Middletown, Connecticut. From 1891 to 1900 many sources. Most of the types that are now he was Professor of Biology and Geology at preserved are in better than average condition. Miami University at Oxford, Ohio. In 1899 he The long list of titles, 71 in the list of was granted the degree of Doctor of Philoso- references, indicates a prolonged and genuine phy from the University of Chicago, where he interest in one of the more difficult groups of completed a valuable study on the cytogeny of marine invertebrates. Except for a rather rigid the direction of adherence to the classificaftons proposed by Podarke obscura Verrill under few Profs. C. 0. Whitman, C. M. Child, E. G. Chamberlin (1919), Treadwell consulted Conklin, and A. D. Mead. monographic works such as Fauvel's mono- From 1900 to 1938 Treadwell was Professor graph for France (1923 and 1927), McIntosh's of Biology and Zoology at Vassar College in monograph of the British annelids (1900 to 1924), or similar works. This resulted not Poughkeepsie, New York, and served as Chair- for man of its Zoology Department from 1914 to through disregard but from necessity, 1938. During most of these years, from 1909 Treadwell had only occasional access to large he library facilities, including those at Harvard to the time of his death on June 23, 1947, the of Con- also held honored positions as Curator of and Yale universities and Library Associate in the Ameri- gress. Vassar College maintained no large Annelids and Research there was an can Museum of Natural History. During sev- museum collections; thus only he was occasional chance to compare specimens with eral summers, between 1898 and 1907, To instructor and lecturer at the marine stations at those of other institutions. my knowledge, Treadwell kept no extensive catalogue or bibli- Woods Hole, Massachusetts, and Cold Spring from a Island, New York. He made ography, but was able to draw freely Harbor, Long 245 246 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 memory that had remarkable retentive powers. bellidae, nine; the Polyodontidae and Cirra- It was also my personal impression that the tulidae each, eight; and the other families have microscopic equipment and laboratory facilities fewer species. at his command were very meager for the ex- Among the genera newly elected only Sphae- anmination of the minute parts which must be ropomatus Treadwell, 1934, is valid, and distinguished for the characters of most of the Epitoka Treadwell, 1943, is questionable. Hali- marine annelids to be defined. planella, 1943, goes to Halyplanes Reibisch; The original list of 324 names (see below) is Monorchos, 1926, to Lygdamis Kinberg; Nu- here reduced to 266 valid species, of which 138 chubranchia, 1928, to Travisiopsis Levinsen; carry Treadwell's original name. They are dis- and Metamphinome, 1940, to Hipponoe tributed in 141 genera and 38 families. A sur- Audouin and Milne-Edwards. vey of the list shows that most species are in Names newly erected herein are Cirriformia the Polynoidae with 33 species. The Eunicidae hawaiensis for Audouinea branchiata Tread- have 30; the Nereidae, 24; the Lumbrineridae well, 1943, preoccupied; and Lumbrineris and Phyllodocidae each, 13; the Sabellidae and treadwelli for Lumbriconereis maculata Tread- Syllidae each, 12; the Terebellidae, 11; the well, 1901. They are discussed more fully Alciopidae and Serpulidae each, 10; the Ara- below. ALPHABETICAL LIST OF NAMES OF ANNELIDS ERECTED BY TREADWELL THE FOLLOWING LISTS alphabetically the names microcarunculata (Treadwell), 1901. See Hart- of annelids erected by Treadwell. Included are man, 1942c, p. 98. the original name, with its bibliographic cita- Anaitides minuta Treadwell, 1937b, p. 148, figs. tion, the family name, the locality, the 16-18. Phyllodocidae. Arena Bank, western Mex- type ico, in 35 fathoms. Holotype in A.M.N.H. Re- location of holotype or other authoritative ferred to Anaitides madeirensis Langerhans, specimen, revised name, if any, and references 1879. See Systematic Discussion. or comments (A.M.N.H. refers to the Ameri- Ancistrosyllis tentaculata Treadwell, 1941a, p. can Museum of Natural History; U.S.N.M., 1, figs. 1-3. Pilargiidae. Long Island, New York. to the United States National Museum). Holotype in A.M.N.H. See Hartman, 1947b, p. AcanthicolePis longicsrrata Treadwell, 1941d, 497. p. 26, figs. 14. Polynoidae. Planktonic off Ber- Anthostoma latacapitata Treadwell, 1901b, pp. muda in 600-800 fathoms. Holotype in A.M.N.H. 203-204, figs. 61-65. Orbiniidae. Puerto Rico. Referred to Eunoi purpurea Treadwell, 1936. See Holotype in U.S.N.M. Referred to Nainers Systematic Discussion. setosa (Verrill), 1900. See Hartman, 1951, pp. Acholoe orbiculata Treadwell, 1921a, pp. 1-3, 67-70, pl. 17, figs. 1-6. figs 1-8. Polynoidae. Belgian Congo. Holotype in Aphrodita magna Treadwell, 1925a, pp. 1-3, A.M.N.H. Referred to Acholoe astericola (delle figs. 1, 2. Aphroditidae. Cape Polonia, Uruguay. Chiaje), 1841. See Systematic Discussion. Holotype in U.S.N.M., not seen. Acoetes magnifica Treadwell, 1929c, pp. 1-4, Arabella attenuata Treadwell, 1906, p. 1172, fig. figs. 1-7. Polyodontidae. Montego Bay, Jamaica, 62. Arabellidae. Near Monterey Bay, California. British West Indies. Holotype in A.M.N.H. Re- Holotype in U.S.N.M. Referred to Notocirrus ferred to Panthalis pustulata Treadwell, 1924. See attenuatus (Treadwell), 1906. See Hartman, Hartman, 1939b, p. 83. 1944b, p. 174. Alciopa distorta Treadwell, 1943b, p. 35, figs. Arabella dubia Treadwell, 1922a, pp. 160-161, 16-18. Alciopidae. Planktonic east off Hawaii in fig. 52, pl. 7, figs. 11, 12, pl. 8, figs. 8, 9. Pago 50 meters. Holotype in U.S.N.M., not seen by me. Pago Harbor, Samoa. Holotype in A.M.N.H. Referred to Naiades cantrainii delle Chiaje, 1830. Referred to Arebella iricolor (Montagu), 1804. See Systematic Discussion. See Systematic Discussion. Alciopa mutilata Treadwell, 1934a, p. 8, pl. 2, Arabella iridescens Treadwell, 1906, p. 1171, figs. 20-21. Alciopidae. Off Puerto Rico in 100 figs. 59-61. Arabellidae. Off Hawaii in 127 fathoms. Holotype in U.S.N.M., not seen. From fathoms. Holotype in U.S.N.M. Redescribed in the very brief account, it seems to be Naiades Hartman, 1942a, pp. 124-125, fig. 13d. cantrainii delle Chiaje. Arabella pacifica Treadwell, 1941c, p. 23, figs. Ammotrypane bermudiensis Treadwell, 1936b, 18-21. Arabellidae. Sihuatanego, Mexico. Holo- pp. 60-61, figs. 24-26. Opheliidae. Nonsuch Island, type in A.M.N.H. Referred to Arabella semi- Bermuda, in 10-35 feet. Holotype in A.M.N.H. maculata (Moore), 1911. See Systematic Dis- Referred to Arnwndia maculata (Webster), 1884. cussion. See Systematic Discussion. Arabella setosa Treadwell, 1921d, pp. 113-114, Ampharete brevibranchiata Treadwell, 1926d, figs. 421-424, p1. 9, figs. 10, 11. Arabellidae. Buc- pp. 6-7, figs. 11-14. Ampharetidae. Bering Strait, coo Bay, Tobago, West Indies. Type in A.M.N.H. Alaska. Holotype in A.M.N.H. Referred to Am- Questionably Arabella iricolor (Montagu), 1804. pharete arctica Malmgren, 1866. See Systematic See Systematic Discussion. Discussion. Aracoda attenuata Treadwell, 1911, pp. 5-6, Ampharete seribranchiata Treadwell, 1926d, pp. figs. 8-11. Arabellidae. Fort Jefferson, Florida, 7-8, figs. 15-16. Ampharetidae. Bering Strait, shore. Holotype in A.M.N.H. Referred to Ara- Alaska. Holotype in A.M.N.H. Referred to Am- bella sp. See Systematic Discussion. pharete eupalea Chamberlin, 1920. See Berkeley Aracoda spatula Treadwell, 1911, pp. 6-7, figs. and Berkeley, 1942, p. 201. 12-14. Arabellidae. Fort Jefferson, Florida, shore. Amphinome microcarunculata Treadwell, 1901b, Holotype in A.M.N.H. Referred to Drilonereis pp. 194-195, fig. 32. Amphinomidae. Puerto Rico. spatula (Treadwell), 1911. See Systematic Dis- Holotype in U.S.N.M. Questionably Benthoscolex cussion. 247 24828BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109

Aricia cirrata Treadwell, 1901b, pp. 201-202, Hesionidae. Puerto Rico. Holotype in U.S.N.M. figs. 54-57. Orbiniidae. Puerto Rico. Holotype in Questionably Nereimyra mutilata (Treadwell), U.S.N.M. Referred to Scoloplos treadwelli Eisig, 1901. See Systematic Discussion. 1914, pp. 405-407, and discussed by Augener, Castalia oculata Treadwell, 1906, p. 1148, figs. 1933b, p. 318. 1-3. Hesionidae. Hawaii. Questionably Leoc-rates Dis- A4ricidea aGlaa Treadwell, 1901b, p. 202, figs. oculatus (Treadwell), 1906. See Systematic 58-60. Spionidae. Puerto Rico. Holotype in cussion. U.S.N.M. Referred to Laonice cirrata (Sars), Centrocorone spinifera Treadwell, 1939a, pp. 1851. See Hartman, 1941, p. 293. 1-3, figs. 1-9. Sabellariidae. Boca de Cangrejoo, to Audouinea branchiata Treadwell, 1943d, p. 1, Puerto Rico. Holotype in A.M.N.H. Referred figs. 1-3. Cirratulidae. Pearl Harbor, Hawaii. Phiragmatopoma lapidosa Kinberg, 1867. See Holotype in A.M.N.H. Referred to Cirriformia Hartman, 1944c, pp. 348-349, pls. 35, 36, 40. hawaiensis, new name. See Systematic Discus- Ceratonereis longicauda Treadwell, 1943a, pp. sion. 1-3, figs. 1-7. Nereidae. Gulf of Davao, Philip- Audouinea nmxculata Treadwell, 1929d, p. 10. pine Islands. Holotype in A.M.N.H. Referred to Cirratulidae. Berbera, British Somaliland. Holo- Ceratonereis anchylochaeta (Horst), 1924. See type in A.M.N.HL Referred to Cirriformia semi- Systematic Discussion. cincta (Ehlers), 1905. See Systematic Discussion. Ceratonereis singularis Treadwell, 1929d, pp. Audouinea oculata Treadwell, 1932b, pp. 17-18, 1-3, figs. 1-8. Nereidae. San Jose Island, Baja figs. 12-14. Cirratulidae. Sao Paulo, Brazil. Holo- California, Mexico. Holotype in A.M.N.H. Re- type in U.S.N.M. Referred to Cirriformia filigera ferred to Ceratonereis mirabilis Kinberg, 1866. (delle Chiaje), 1841. See Hartman, 1942a, p. 127- See Hartman, 1948b, p. 71. 128. Cirratulus branchiatus Treadwell, 1936a, pp. Audouinea pygidia Treadwell, 1936b, p. 64, figs. 276-277, fig. 20g. Cirratulidae. Near Amoy, China. 22, 23. Cirratulidae. Nonsuch Island, Bermuda, in Holotype in U.S.N.M. Referred to Cirriformia tidal pools. Paratype in A.M.N.H. Referred to chefooensis (Grube), 1877. See Systematic Dis- Cirriformia filigera (delle Chiaje), 1841. See cussion. Systematic Discussion. Cirratulus edongatus Treadwell, 1901b, p. 204, Autolytus bidens Treadwell, 1941d, pp. 28-29, fig. 67. Cirratulidae. Culebra, Panama. Holotype fig. 9. Syllidae. Nonsuch Island, Bermuda, plank- in U.S.N.M., not seen. Indeterminable as to tonic at surface. Holotype in A.M.N.H. Referred genus. to Odontosyllis enopla Verrili, 1900. See System- Cirratulus inhamatus Treadwell, 1937b, p. 153. atic Discussion. Cirratulidae. East of Cedros Island, western Autolytus pacificus Treadwell, 1943b, p. 40, figs. Mexico, in 38 fathoms. Referred to Tharyx mult- 28-30. Syllidae. Samoa. Holotype in U.S.N.M., filis Moore, 1909. See Systematic Discussion. not seen. Cirratulus multicirratus Treadwell, 1936b, pp. Autolytus variu-s Treadwell, 1914c, pp. 237-238, 63-64, figs. 34, 35. Cirratulidae. St. Davids Island, figs. 4-7. Syllidae. San Francisco Bay, California. Bermuda, on mud flats. Holotype in A.M.N.H. Types not known to exist. Redescribed by Berke- Referred to Cirriformia tentaculata (Montagu), ley and Berkeley, 1948, p. 70, fig. 102. 1808. See Systematic Discussion. Branchiomma disparoculatum Treadwell, 1914b, Cirratulus nigromaculata Treadwell, 1901b, p. pp. 223-224, pl. 12, figs. 44-46. Sabellidae. Off San 204, fig. 66. Cirratulidae. Culebra, Panama. Holo- Diego, California. Holotype in Los Angeles. Re- type in U.S.N.M. Referred to Cirriformia punc- ferred to Megatlonma splendida (Moore), 1905. tata (Grube), 1859. See Systematic Discussion. See Systematic Discussion. Cirratulus paUidus Treadwell, 1931f, pp. 1-2, Callizonella pigmenta Treadwell, 1943b, p. 38, figs. 1-3. Cirratulidae. Montego Bay, Jamaica, fig. 25. Alciopidae. Tropical mid-Atlantic Ocean. British West Indies. Holotype in A.M.N.H. Re- Holotype in U.S.N.M., not seen, appears to be ferred to Cirriformia tentaculata (Montagu), Krohnia sp. 1808. See Systematic Discussion. Capitellides teres Treadwell, 1939a, p. 6, figs. Corynocephalus magnachaetus Treadwell, 21-24. Capitellidae. Port Aransas, Texas. Holo- 1943b, p. 37, figs. 21, 22. Alciopidae. Equatorial type in A.M.N.H. See Hartman, 1951, p. 101. western Pacific Ocean. Holotype in U.S.N.M., Castalia longicirrata Treadwell, 1901b, p. 185, not seen. figs. 2, 3. Hesionidae. Puerto Rico. Holotype in Dasybranchus rectus Treadwell, 1901b, p. 207, U.S.N.M. Questionably Dalhousiella longicirrata fig. 72. Capitellidae. Culebra, Panama. Holotype (Treadwell), 1901. See Systematic Discussion. in U.S.N.M. Questionable, see Hartman, 1947a, Castalia mutilcata Treadwell, 1901b, p. 185. pp. 430-431. 1956196HARTMAN: ANNELIDS 249

Dasychone loandensis Treadwell, 1943c, p. 4, 175, figs. 7-10. Phyllodocidae. Friday Harbor, figs. 21-25. Sabellidae. St. Paul de Loanda, Washington. Holotype in A.M.N.H. Referred to Africa. Holotype in A.M.N.H. Referred to Eteone flonga (Fabricius), 1780. See Berkeley Branchiomma nigrornaculata (Baird), 1865. See and Berkeley, 1945, p. 325. Systematic Discussion. Eulalia foliosa Treadwell, 1924c, p. 10, figs. Dasychone ponce Treadwell, 1901b, p. 209, figs. 16-18. Phyllodocidae. Barbados, West Indies. 76-78. Sabellidae. Puerto Rico. Holotype in Holotype in U.S.N.M., not seen. U.S.N.M. Referred to Branchiomma nigromacu- Eulalia magnapupula Treadwell, 1941c, pp. 21- lata (Baird), 1865. See Johansson, 1927, p. 162. 22, figs. 13-17. Alciopidae. Off Jicaron Island, Dasychonopsis arenosa Treadwell, 1924b, pp. Pacific Panama, in 500 fathoms. Holotype in 1-2, figs. 1-4. Sabellidae. San Juan, Puerto Rico. A.M.N.H. Referred to RhynchonereUa angelini Holotype in A.M.N.H. Referred to Branchiomma (Kinberg), 1866. See Systematic Discussion. nigromaculata (Baird), 1865. See Johansson, Eulalia quinquelineata Treadwell, 1901b, p. 1927, p. 162. 192, figs. 27-29. Phyllodocidae. Puerto Rico. Ho- Drieschia atlantica Treadwell, 1936b, pp. 52-53, lotype in U.S.N.M. Referred to Eulalia myriacy- figs. 7-9. Polynoidae. Nonsuch Island, Bermuda, clum (Schmarda), 1861. See Augener, 1925, pp. in 1000 fathoms. Holotype in A.M.N.H. Referred 24-25. to Drieschia pellucida Moore, 1903. See System- Eulepis fimbriata Treadwell, 1901b, pp. 190- atic Discussion. 191, figs. 23-24. Pareulepidae. Puerto Rico. Holo- Drilonereis brunnea Treadwell, 1921d, pp. 111- type in U.S.N.M. Referred to Pareulepis fimbri- 112. Arabellidae. Tobago, West Indies. Holotype ata (Treadwell), 1901. See Hartman, 1939b, pp. in A.M.N.H. Referred to Drilonereis spatula 79-80, pl. 23, figs. 280-288. Treadwell, 1921. See Systematic Discussion. Eulepis splendida Treadwell, 1901b, pp. 189- Drilonereis lumbricus Treadwell, 1922a, pp. 190, figs. 19-22. Pereulepidae. Puerto Rico. Holo- 161-162, figs. 53-55, pl. 7, figs. 13-15, pl. 8, fig. type in U.S.N.M. Referred to Pareulepis wyvillei 10. Arabellidae. Suva Harbor, Fiji. Holotype in (McIntosh), 1885. See Hartman, 1939b, p. 78. A.M.N.H. See Systematic Discussion. Eunereis africana Treadwell, 1943c, pp. 2-3, Drilonereis paucidentata Treadwell, 1922a, p. figs. 8-13. Nereidae. Lobito, Africa. Holotype in 162, fig. 56, pl. 7, figs. 16, 17, pl. 8, fig. 11. Ara- A.M.N.H. Referred to Platynereis dumerilii bellidae. Fiji. Holotype in A.M.N.H. Referred to (Audouin and Milne-Edwards), 1834. See Sys- Drilonereis planiceps (Grube), 1878. See System- tematic Discussion. atic Discussion. Eunice auriculata Treadwell, 1900; 1901, pp. Drilonereis pinnata Treadwell, 1921d, pp. 110- 196-197, figs. 33-36. Eunicidae. Puerto Rico. 111, figs. 412-413, pL. 8, fig. 11. Arabellidae. To- Holotype in U.S.N.M. See Hartman, 1944b, p. bago, West Indies. Holotype in U.S.N.M. Ques- 101. tionably Arabella iricolor (Montagu), 1804. See Eunice bilobata Treadwell, 1906, pp. 1168-1169, Systematic Discussion, as Arabella sp. figs. 47, 48. Eunicidae. Off Hawaii in 13.43 Drilonereis similis Treadwell, 1921d, p. 111, fathoms. Holotype in U.S.N.M., not seen. figs. 414-417. Arabellidae. Tobago, West Indies. Eunice culebra Treadwell, 1901b, p. 197, fig. 37. Holotype in A.M.N.H. Referred to Drilonereis Culebra, Panama. Holotype in U.S.N.M. Referred sp., juvenile stage. See Systematic Discussion. to Eunice (Nicidion) cariboea Grube, 1856. See Enipo cirrata Treadwell, 1925b, pp. 1-3, figs. Monro, 1933, p. 63. 1-4. Polynoidae. Kotzebue Sound, Alaska, in 12- Eunice hawaiensis Treadwell, 1906, pp. 1166- 17 fathoms. Holotype in U.S.N.M. Referred to 1167, figs. 42-44. Off Hawaii in 324 to 225 Polynoi gracilis Verrill, 1874. See Pettibone, fathoms. Holotype in U.S.N.M. See Hartman, 1954, p. 225, as Enipo. 1944b, p. 101. Epitoka pelagica Treadwell, 1943b, p. 40, fig. Eunice interrupta Treadwell, 1906, pp. 1167- 31. Questionably Pilargiidae or Syllidae. West of 1168, figs. 45-46. Off Hawaii in 43 to 70 fathoms. Peru, planktonic in 50 meters. Holotype in Holotype U.S.N.M., not seen. Probably Eunice U.S.N.M., not seen. Except for the reported ab- antennata Savigny, 1818 (Hartman, 1944b, pp. sence of a median antenna, this resembles species 115-117). of Ancistrosyllis McIntosh. Eunice nicidioformis Treadwell, 1906,- p. 1169, Eteone maculata Treadwell, 1922b, p. 174, figs. figs. 49-51. Off Hawaii in 95 to 152 fathoms. 3-6. Phyllodocidae. Friday Harbor, Washington. Holotype in U.S.N.M., not seen. Holotype in A.M.N.H. Referred to Eteone pa- Eunoe ezoculata Treadwell, 1923b, pp. 4-6, figs. cifica Hartman, 1936. See Hartman, 1936, p. 31. 1-4. Polynoidae. Off western Mexico in 475 Eteone tuberculata Treadwell, 1922b, pp. 174- fathoms. Holotype in U.S.N.M. Referred to Har- 250 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 mothoe teiebricosa Moore, 1910. See Hartman, U.S.N.M., not seen. 1938d, p. 118. Euratio punctata Treadwell, 1926a, p. 20. Lap- Eunoe purpurea Treadwell, 1936b, pp. 51-52, sus calami for Eurato punctata. figs. 1-6. Polynoidae. Nonsuch Island, Bermuda, Eurythoe oculata Treadwell, 1941c, p. 18, figs. in 600-1000 fathoms. Holotype in A.M.N.H. See 1-3. Amphinomidae. Balboa, Pacific Panama. Systematic Discussion. Holotype in A.M.N.H. Referred to Pseudeury- Eupanthalis evanida Treadwell, 1926b, pp. 186- thoe oculata (Treadwell), 1941. See Systematic 188, figs. 6-12. Polyodontidae. Philippine Islands. Discussion. Holotype in U.S.N.M. Referred to Panthalis Evarnella trimaculata Treadwell, 1924c, pp. 6-7, evanida (Treadwell), 1926. See Hartman, 1938d, figs. 1-4. Polynoidae. Barbados, West Indies. p. 127. Holotype in U.S.N.M. Referred to Harmothoi Eupanthalis oahuensis Treadwell, 1906, pp. trimaculata (Treadwell), 1924. See Hartman, 1155-1156, figs. 19-23. Polyodontidae. Off Hawaii 1938d, p. 118, figs. 38a, 39a, b. in 328 fathoms. Holotype in U.S.N.M. Referred Gattyana imbricata Treadwell, 1926d, pp. 2-4, to Eupanthalis mutilata (Treadwell), 1906. See figs. 1-5. Polynoidae. Bering Strait, Alaska. Hol- Hartman, 1938d, p. 123. otype in A.M.N.H. Referred to Gattyana cirrosa Eupholoe acuminata Treadwell, 1934a, pp. 3-4, (Pallas), 1766. See Pettibone, 1949, pp. 1, 4. pl. 1, figs. 7-8. Sigalionidae. Off Puerto Rico in Glycera abranchiata Treadwell, 1901b, pp. 200- 15 fathoms. Holotype in U.S.N.M. Referred to 201, fig. 49. Glyceridae. Puerto Rico. Holotype in Psammolyce flava Kinberg, 1855. See Hartman, U.S.N.M. Referred to Glycera tesselata Grube, 1942a, pp. 108-109, fig. 8. 1863. See Augener, 1922b, p. 205. Eupholoi cirrata Treadwell, 1934a, pp. 5-6, Glycera spadix Treadwell, 1943a, pp. 3-4, figs. figs. 9-16. Sigalionidae. Off Puerto Rico in 200- 811. Glyceridae, Gulf of Davao, Philippine 260 fathoms. Holotype in U.S.N.M. Referred to Islands. Holotype in A.M.N.H. Referred to Gly- Leanira hystricis Ehlers, 1874. See Hartman, cera tesselata Grube, 1863. See Hartman, 1950, 1942a, p. 105. 77. Eupholoi nuda p. Treadwell, 1936b, pp. 53-54, Goniada brunnea Treadwell, 1906, p. 1174, figs. figs. 10-14. Sigalionidae. Nonsuch Island, Ber- 67-70. Off Hawaii in 202-220 fathoms. Holotype muda, in 35 feet. Holotype in A.M.N.H. Referred in U.S.N.M. Redescribed in Hartman, to Psammolyce arenosa (delle Chiaje), 1841. See 1950, pp. Systematic Discussion. 17-19, pl. 1, figs. 1-6. Gonsada magna Treadwell, 1945, pp. 2-3, Euphrosyne branchiata Treadwell, 1939c, pp. 6-8. Goniadidae. figs. 170-172, figs. 46c-f. Georges Bank, Massachusetts, Euphrosinidae. Chesapeake in 30 meters. Holotype in A.M.N.H. Referred to Bay, Maryland, in 70 fathoms. Holotype i Goniada quinquelabiata Augener, 1906. See Hart- U.S.N.M., not seen. man, 1950, pp. 26-28. EuphrosYne longisetis Treadwell, 1939c, p. 172, Goniada oculata figs. 46g-i. Greenland. Holotype in U.S.N.M., not Treadwell, 1901b, p. 201, figs. seen. 50-53. Goniadidae. Puerto Rico. Holotype in U.S.N.M. Questionable, perhaps a species of Eupomatus decorus Treadwell, 1931c, pp. 4-5, See figs. 3a-d. Serpulidae. Grand Isle, Glycinde MuIller. Hartman, 1950, p. 14. Louisiana. Goniada teres Treadwell, 1931d, pp. 6-7, Holotype in U.S.N.M. Referred to Eupomatus 19-22. Goniadidae. figs. dianthus (Verrill), 1873, variety decorrus Tread- Montego Bay, Jamaica, West well, 1931. See Indies. Holotype in A.M.N.H. See Hartman, Hartman, 1951, p. 118. 1950, pp. 33-34. Eupomatus operculata Treadwell, 1929d, p. 12. Haliplanella Serpulidae. Berbera, British Somaliland. Holoo pacifica Treadwell, 1943b, p. 32, type in figs. 4-6. Phyllodocidae. Warm parts of Atlantic A.M.N.H. Referred to Eupomatus uncina- and Pacific oceans. tus Philippi, 1844. See Systematic Paratypes in A.M.N.H. Re- Discussion. ferred to Halyplanes gracilis Reibisch, 1893. See Eupomatus parvus Treadwell, 1901b, p. 210, Systematic figs. 79, 80. Serpulidae. Puerto Rico. Holotype in Discussion. U.S.N.M. Referred to Hydroides parvus (Tread- Halosydnta fusca-maculata Treadwell, 1924c, well), 1901. See Pixell, 1913, p. 73. pp. 5-6, figs. 5-9. Polynoidae. Barbados, West Indies. Holotype in U.S.N.M. Referred to EupomatUs similis Treadwell, 1929d, pp. 11-12, sydneUa Halo- fig. 31. Serpulidae. Baja California, Mexico. fuscamaculata (Treadwell), 1924. See Paratypes in A.M.N.H. Hartman, 1938d, pp. 111-112, figs. 36f, g. See Systematic Discus- Halosydna fuscomarginata Treadwell, sion. in 1924c, Eurato punctata Treadwell, 1926a, pp. 17-18, explanation of figures. Lapsus calams for figs. 28-30. Sabellidae. Samoa. Holotype in Halosydna fusca-maculata, quod vide. Halosydna grisea Treadwell, 1929a, pp. 1-3, 1956196HARTMAN: ANNELIDS 251 figs. 1-6. Polynoidae. Argentina. Holotype in (Augener), 1906. See Hartman, 1944c, pp. 333- U.S.N.M. Referred to Halosydnella grisea 334, pl. 31, figs. 30-33. (Treadwell), 1929. See Hartman, 1938d, p. 111; Hermodice pennata Treadwell, 1906, p. 1165, 1948b, p. 22. fig. 41. Amphinomidae. Off Hawaii in 24 fathoms. Halosydna obtusa-cirrata Treadwell, 1937b, pp. Holotype in U.S.N.M. Referred to Pherecardia 143-144, pl. 1, figs. 8-11. Polynoidae. Cedros Is- striata (Kinberg), 1857. See Augener, 1927a, land, western Mexico, in 45 fathoms. Holotype in p. 122. A.M.N.H. Referred to Halosydna latior Cham- .Hermodice pennata tutuilensis Treadwell, 1926a, berlin, 1919. See Hartman, 1938d, p. 110. p. 2. Amphinomidae. Pago Pago, Samoa. Holo- Halosydna oculata Treadwell, 1926a, pp. 8-10, type in U.S.N.M., not seen. Possibly Pherecardia figs. 9-13. Polynoidae. Pago Pago, Samoa. Holo- sp. type in U.S.N.M. Referred to Halosydnella ocu- Hipponoe elongata Treadwell, 1931d, pp. 3-4, lata (Treadwell), 1926. See Hartman, 1938d, figs. 10-12. Amphinomidae. Puerto Rico. Holotype pp. 112-113, figs. 36a-c. in A.M.N.H. Referred to .Pareurythoe elongata Haplobranchus atlanticus Treadwell, 1932a, pp. (Treadwell), 1931. See Systematic Discussion. 279-280, figs. 1-8. Sabellidae. St. Andrews, New Hyalinoecia branchiata Treadwell, 1934a, pp. Brunswick, Canada. Paratype in A.M.N.H. Re- 6-8, pl. 2, figs. 17-19. Onuphidae. Culebra Island, ferred to Fabricia sabeUa (Ehrenberg), 1836. Puerto Rico, in 300 fathoms. Holotype in See Rullier, 1954, p. 22. U.S.N.M., not seen. Haplosyllis gula Treadwell, 1924c, pp. 11-12, Hydroides californicus Treadwell, 1929d, p. 12, figs. 19-23. Syllidae. West Indies. Holotype in figs. 32, 33, Serpulidae. Western Mexico. Syn- U.S.N.M. Referred to Haplosylis spongicola types in A.M.N.H. Referred to Hydroides (Grube), 1855. See Monro, 1933, p. 34. crucigera Morch, 1863. See Rioja, 1944, p. 409. Harmothoe lanceocirrata Treadwell, 1928a, pp. Hypsicomus purpureus Treadwell, 1924c, pp. 454-455, figs. 5-9. Polynoidae. Holotype in 20-21, figs. 30-33. Sabellidae. Antigua, West A.M.N.H. Referred to Harmothoe crucis Indies. Holotype in U.S.N.M. Referred to Hypsi- (Grube), 1856. See Systematic Discussion. comus torquatus (Grube), 1878. See Hartman, Harmothoe lanceolata Treadwell, 1928a, p. 451. 1942a, p. 133, as Hypsicomus circumspiciens. Lapsus calami for Ha-rmothoe lanceocirrata, Iphionella elongata Treadwell, 1931a, pp. 315- quod vide. 317, figs. 2a-d. Polyodontidae. Marinduque Is- Harmothoe levis Treadwell, 1937a, pp. 2627, land, Philippine Islands, in 83-159 fathoms. figs. 1-5. Polynoidae. East Greenland. Holotype Holotype in U.S.N.M. Referred to Eupolyodontes in U.S.N.M. Referred to Harmothoe imbricata elongata (Treadwell), 1931. See Hartman, 1938d, (Linnaeus), 1767. See Hartman, 1938d, p. 129. pp. 125-126, figs. 41a-d. Harnothoe sylliformia Treadwell, 1928a, pp. Laetmonice nitida Treadwell, 1926b, pp. 183- 452-454, figs. 1-4. Polynoidae. Off Galapagos Is- 186, figs. 1-5. Aphroditidae. Near Jolo, Sulu lands in 300-700 fathoms. Holotype in A.M.N.H. Archipelago, in 20 fathoms. Holotype in U.S.N.M., See Systematic Discussion. not seen. Harmothoe tuberculata Treadwell, 1906, p. 1154. Lagisca crassa Treadwell, 1924a, pp. 1-3, figs. Polynoidae. Off Hawaiian Islands in 451 fathoms. 1-4. Polynoidae. Punta Arenas, Chile. Holotype Holotype in U.S.N.M. Referred to Arctonoe in U.S.N.M. Referred to Eunoe crassa (Tread- tuberculata (Treadwell), 1906. See Hartman, well), 1924. See Hartman, 1938d, p. 119, figs. 1938d, p. 117, figs. 37e, f. 38b-e. Harmothoe variegata Treadwell, 1917b, pp. Lanice expansa Treadwell, 1906, p. 1176. 260-261, pl. 1, figs. 17, 18, pl. 2, figs. 1-3. Poly- Terebellidae. Off Hawaii in 122-143 fathoms. noidae. Dry Tortugas, Florida. Holotype in Holotype in U.S.N.M., not seen. A.M.N.H. See Systematic Discussion. Laonome arenosa Treadwell, 1943d, p. 3, figs. Harmothoe' vilosa Treadwell, 1926a, pp. 10-12, 9-13. Terebellidae. Hawaii. Holotype in A.M.N.H. figs. 14-18. Polynoidae. Pago Pago, Samoa. Referred to Sabellastarte indica Savigny, 1820. Holotype in U.S.N.M. See Hartman, 1938d, p. 117. See Systematic Discussion. Hemipodia canadensis Treadwell, 1937c, pp. Laonome oculifera Treadwell, 1914b, pp. 222- 348-349, figs. 1-3. Glyceridae. Halifax Harbor, 223, pl. 12, figs. 39-43. Sabellidae. San Pedro, Nova Scotia, Canada. Referred to Glycera California. Holotype in Los Angeles. Referred ?capitata Oersted, 1843. See Hartman, 1950, p. 63. to Pseudopotamilla occelata Moore, 1905. See Hermella varians Treadwell, 1910b, p. 210, Hartman, 1938a, pp. 25-27, pl. 2, figs. 6, 7. fig. 81. Sabellariidae. Puerto Rico. Holotype in Laonome punctata Treadwell, 1906, p. 1179, U.S.N.M. Referred to Lygdamis asteriformis figs. 76, 77. Sabellidae. Oahu, Hawaii. Holotype 252 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 in U.S.N.M. Referred to Sabellastrate indica Leodice langi Treadwell, 1943c, p. 3, figs. 14-18. Savigny, 1820. See Systematic Discussion. Eunicidae. Capetown, Africa. Holotype in Laonome sanjuanensis Treadwell, 1941a, pp. 3- A.M.N.H. Referred to Eunice savigny- (Grube), 4, figs. 4-8. Sabellidae. East San Juan, Puerto 1878. See Systematic Discussion. Rico, in tidal pool. Holotype in A.M.N.H. Re- Leodice notata Treadwell, 1921d, pp. 52-54, ferred to Sabellastarte magnifica (Shaw), 1800. figs. 164-173, pl. 4, fig. 12. Eunicidae. West Indies. See Systematic Discussion. Referred to Eunice tafra Peters, 1854. See Hart- Leodice aciculata Treadwell, 1922a, pp. 143- man, 1944b, pp. 111-112, pl. 6, figs. 142-144; and 144, figs. 24, 25, pl. 3, figs. 7-13. Eunicidae. Samoa Systematic and Fiji Discussion. Holotype in A.M.N.H. Referred to Leodice rubrivittata Treadwell, 1921d, pp. 34- Eunice afra Peters, 1854. See Systematic Discus- 36, figs. 85-94, pl. 1, fig. 18. Eunicidae. Buccoo sion. Bay, Tobago, West Indies. Holotype in A.M.N.H. Leodice arcturi Treadwell, 1928a, pp. 475-477, Referred to Eunice vittata delle Chiaje, 1828. figs. 32-39. Eunicidae. Off New Jersey in 633 See Augener, 1927b, p. 40. fathoms. Holotype in A.M.N.H Referred to Leodice spongicola Treadwell, 1921d, pp. 25-27, Eunice norvegica (Linnaeus), 1767. See Systema- fig. 53. Eunicidae. tic Discussion. Montego Bay, Jamaica, West Indies. Holotype in A.M.N.H. Referred to Eunice Leodice argentinensis Treadwell, 1929a, pp. filamentosa Grube, 1856. See Systematic Discus- 3-5, figs. 7-12. Eunicidae. Mar del Plata, Ar- sion. gentina. Holotype in U.S.N.M. Referred to Leodice suviensi's Treadwell, 1922a, pp. 138-139, Eunice argentinensis (Treadwell), 1929. See figs. 12-16, pl. 2, figs. 8-13. Eunicidae. Suva Rioja, 1944, pp. 128-130, figs. 39-46. Harbor, Fiji. Referred to Eunice afra Peters, Leodice armillata Treadwell, 1922a, pp. 144- 1854. See Systematic Discussion. 146, figs. 26-29, p1. 3, figs. 14-19. Eunicidae. Pago Leodice Pago tenuis Treadwell, 1921d, pp. 51-52, Harbor, Samoa. Holotype in A.M.N.H. Re- figs. 154-163, pl. 4, 11. Eunicidae. Florida ferred to Eunice armillata (Treadwell), 1922. fig. See Systematic Keys. Holotype in A.M.N.H. Referred to Eunice Discussion. tenuis (Treadwell), 1921. See Systematic Discus- Leodice biformi-cirrata Treadwell, 1922a, pp. sion. 148-149, figs. 34, 35, pl. 4, figs. 6-11. Eunicidae. Leodice tubicola Treadwell, 1922a, Suva Harbor, Fiji. Holotype in A.M.N.H. Re- figs. 17-23, pl. 3, pp. 139-142, Eunice grubei Gravier, 1900. See Systematic figs. 1-6. Eunicidae. Pago Pago Discussion. Harbor, Samoa. Holotype in A.M.N.H. Referred to Eunice tubifex Crossland, 1904. See Leodice bucciensis Treadwell, 1921d, pp. 54-56, Discussion. Sytematic figs. 174-183. Eunicidae. Buccoo Bay, Tobago, Leodice viridis vernalis West Indies. Holotype in A.M.N.H. Referred to Treadwell, 1922a, pp. Eunice afra Peters, 1854. See Systematic 133-134, pl. 1, figs. 8-11. Suva Harbor, Fiji. Holo- Discus- type in A.M.N.H. Referred to Palolae siciliensis sion. (Grube), 1840. See Systematic Leodice crassi-tentacuata Treadwell, 1922a, Lepidasthenia Discussion. pp. 146-148, figs. 30-33, pl. 4, figs. 1-5. Eunicidae. elegans Treadwell, 1941c, pp. 19- Pago Pago Harbor, Samoa. 20, figs. 4-7. Polynoidae. Off Cedros Island, Holotype in Mexico, in 45 fathoms. Holotype in A.M.N.H. A.M.N.H. Referred to Eunice crassitentacutata Referred to (Treadwell), 1922. See Systematic Discussion. Arctonoe vittata (Grube), 1855. See Leodice Systematic Discussion. flava-punctata Treadwell, 1922a, pp. Lepidasthenia lactea Treadwell, 136-138, figs. 8-11, pl. 2, figs. 1-7. Eunicidae. figs. 1939a, pp. 3-5, Pago Pago Harbor, Samoa. Holotype in 13-15. Polynoidae. Galveston, Texas. Holo- A.M.N.H. Referred type in A.M.N.H. Referred to Lepidametria com- to Eunice afra Peters, 1854. mensalis Webster, 1879. See See Systematic Discussion. Lepidasthenia Hartman, 1951, p. 17. Leodice gracili-cirrata Treadwell, 1922a, pp. longicirrata Treadwell, 1928a, 149-150, figs. 36-38, pl. 5, figs. 1-8. Eunicidae. pp. 460-461, figs. 22-26. Polynoidae. West of Peru. Holotype in A.M.N.H. Referred to Suva Harbor, Fiji. Holotype in A.M.N.H. Re- thenia natans Lepidas- ferred to Eunice armillata (Treadwell), 1922. (Chamberlin), 1919. See Systematic See Systematic Discussion. Discussion. Leodice guanica Treadwell, 1921d, pp. 39-40, LePidasthenia occelata Treadwell, 1936a, pp. figs. 107-116, 264-265, figs. 18e-h. Polynoidae. Near pl. 2, figs. 9-12. Eunicidae. Sand China. in Amoy, Key, Florida. Holotype in A.M.N.H. Referred to Holotype U.S.N.M. Referred to Eunice afra Peters, 1854. See Systematic Discus- Lepidasthenia interrupta (Marenzeller), 1902. sion. See Hartman, 1938d, p. 114. Lepidasthenia ornata Treadwell, 1937b, pp. 145- 1956 HARTMAN: ANNELIDS 253 147, pl. 1, figs. 12-15. Polynoidae. Arena Bank, in A.M.N.H. Referred to Laeonereis culveri Mexico, in 35 fathoms. Holotype in A.M.N.H. (Webster), 1879. See Hartman, 1951, p. 44. Referred to Lepidasthenica virens (Blanchard), Loimia minuta Treadwell, 1929d, pp. 10-11, 1849. See Systematic Discussion. figs. 28-30. Terebellidae. Dry Tortugas, Florida. Lepidasthenia picta Treadwell, 1938a, pp. 456- Holotype in A.M.N.H. Referred to Loimia medusa 457, figs. 10-13. Polynoidae. Off Galapagos Is- (Savigny), 1820. See Systematic Discussion. lands in 15 feet. Holotype in A.M.N.H. Referred Lopadorhynchus varius Treadwell, 1943b, p. 32, to Lepidasthenia varius Treadwell, 1917. See figs. 7-10. Lopadorhynchidae. Between United Systematic Discussion. States and Hawaii, pelagic. Holotype in U.S.N.M., Lepidasthenia rufa Treadwell, 1928a, pp. 457- not seen. 458, figs. 14-17. Polynoidae. Puerto Rico. Holo- Lumbriconereis bilabiata Treadwell, 1901b, type in A.M.N.H. See Systematic Discussion. p. 199, figs. 45, 46. Lumbrineridae. Puerto Rico. Lepidasthenia variegata Treadwell, 1928a, pp. Holotype in U.S.N.M. Referred to Lumbrineris 458-459, figs. 18-21. Polynoidae. Off Puerto Rico bilabiata (Treadwell), 1901. See Hartman, 1942a, in 125 fathoms. Holotype in A.M.N.H. Referred p. 120. to Lepidasthenia rufa Treadwell, 1928. See Lumbriconereis grandis Treadwell, 1906, pp. Systematic Discussion. 1170-1171, figs. 52-56. Lumbrineridae. Off Hawaii Lepidasthenia varius Treadwell, 1917b, pp. in 257-319 fathoms. Holotype in US.N.M. Re- 259-260, pl. 1, figs. 11-16. Polynoidae. Dry Tor- ferred to Lumbrineris grandis (Treadwell), 1906. tugas, Florida. Holotype in A.M.N.H. See Sys- See Hartman, 1942a, pp. 114-116, figs. lOh, k, 1. tematic Discussion. Lumbriconereis maculata Treadwell, 1901b, pp. Lepidonotus atratus Treadwell, 1940, pp. 3-4, 198-199, figs. 42-44. Lumbrineridae. Puerto Rico. figs. 10-13. Polynoidae. Gulf of Devas, Philippine Holotype in U.S.N.M. Referred to Lumbrineris Islands. Holotype in A.M.N.H. Referred to treadwelli, new name. See Systematic Discussion. Thormora jukesii Baird, 1865. See Systematic Lumbriconereis minuta Treadwell, 1906, p. 1171, Discussion. figs. 57-58. Lumbrineridae. Off Hawaii in 238- Lepidonotus inquilinus Treadwell, 1917b, p. 258, 253 fathoms. Holotype in U.S.N.M. Referred to pl. 1, figs. 4-10. Polynoidae. Key West Harbor, Lumbrineris minuscula Moore, 1911. See Hart- Florida, shore. Holotype in A.M.N.H. Referred man, 1942a, p. 116, figs. 12e, f. to Lepidasthenia sp., juvenile stage. See Systema- Lumbriconereis parvapedata Treadwell, 1901b, tic Discussion. p. 198, figs. 38-40. Lumbrineridae. Culebra, Pana- Lepidonotus minutus Treadwell, 1936a, pp. 262- ma. Holotype in U.S.N.M. Referred to Lum- 264, figs. 18a-d. Polynoidae. Near Amoy, China. brineris parvapedata (Treadwell), 1901. See Holotype in U.S.N.M. Referred to Lepidonotus Hartman, 1942a, pp. 118-119, figs. lOi, j, m. caelorus Moore, 1903. See Hartman, 1938d, p. 108. Lumbrinereis bicirrata Treadwell, 1929b, pp. Lepidonotus pallidus Treadwell, 1939a, p. 3, 1-3, figs. 1-7. Lumbrineridae. Friday Harbor, figs. 10-12. Polynoidae. Freeport, Texas. Holo- Washington. Holotype in A.M.N.H. Referred to type in A.M.N.H. Referred to Lepidonotus sub- Lumbrineris bicirrata (Treadwell), 1909. See levis Verrill, 1873. See Hartman, 1951, p. 17. Hartman, 1944b, p. 156. Berkeley and Berkeley, Lepidonotus pilosus Treadwell, 1937b, pp. 141- 1948, p. 99, referred it to Lumbrineris bifurcata 143, pl. 1, figs. 1-7. Polynoidae. Arena Bank, (McIntosh), 1885. Mexico, in 45 fathoms. Holotype in A.M.N.H. Lumbrinereis branchiata Treadwell, 1912d, pp. Referred to Chaetacanthus magnificus (Grube), 94-95, figs. 333-343, pl. 8, figs. 5-6. Lumbrineridae. 1875. See Hartman, 1939b, pp. 28-29. Buccoo Bay, Tobago, West Indies. Holotype in Leptonereis egregicirrata Treadwell, 1924c, A.M.N.H. Referred to Lumbrineris branchiata pp. 13-14, fig. 24. Nereidae. Antigua, West Indies. (Treadwell), 1929. See Systematic Discussion. Holotype in U.S.N.M., not seen. Questionable, Lumbrinereis candida Treadwell, 1921d, pp. 96- description may be based on more than one 97, figs. 344-350, pl. 8, figs. 7-9. Lumbrineridae. species. Treadwell, 1939b, p. 233, referred it to British West Indies. Holotype in A.M.N.H. See Nereis. Systematic Discussion. Leptonereis mexicana Treadwell, 1942, pp. 1-4, Lumbbrinereis cervicalis Treadwell, 1922b, p. figs. 2-9. Nereidae. Off Baja California, Mexico. 176, figs. 14-21. Lumbrineridae. Friday Harbor, Holotype in A.M.N.H. Referred to Nicon mexi- Washington. Holotype in A.M.N.H. Referred to cana (Treadwell), 1942. See Systematic Discus- Lumbrineris inflata Moore, 1911. See Systematic sion. Discussion. Leptonereis nota Treadwell, 1941b, pp. 1-3, Lumbrinereis cingulata Treadwell, 1917b, p. figs. 7-10. Nereidae. Galveston, Texas. Holotype 263, pl. 2, figs. 7-12. Lumbrineridae. Bermuda. 254 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Holotype in A.M.N.H. Referred to Lumbrineris Maldanella coraUicola Treadwell, 1929d, p. 8, itnflata Moore, 1911. See Systematic Discussion. figs. 21-24. Maldanidae. Loggerhead Key, Florida. Lumbrinereis elongata Treadwell, 1931c, p. 3, Holotype in A.M.N.H. Referred to Euclymene figs. 2a-d. Lumbrineridae. Grand Isle, Louisiana. corallicola (Treadwell), 1929. See Systematic Holotype in U.S.N.M. Referred to Lumbrineris Discussion. parvapedata (Treadwell), 1901. See Hartman, Maldanella fimbriata Treadwell, 1934a, pp. 8-9, 1942a, pp. 118-119, fig. 10. pl. 2, figs. 22-24. Maldanidae. Off Puerto Rico Lumbrinereis nuchalis Treadwell, 1921d, pp. in 80-100 fathoms. Holotype in U.S.N.M. Re- 104-106, figs. 386-394, pl. 9, fig. 5. Lumbrineridae. ferred to Asychis fimbriata (Treadwell), 1934. Buccoo Bay, Tobago, West Indies. Holotype in See Systematic Discussion. U.S.N.M. See Hartman, 1944b, p. 140. Marphysa acicularum brevibranchiata Tread- Lumbrinereis paucidentata Treadwell, 1921d, well, 1921d, pp. 60-61, figs. 194-200, pl. 5, figs. pp. 99-100, figs. 357-364, pl. 9, figs. 1-4. Lum- 5-8. Eunicidae. Bermuda. Holotype in U.S.N.M. brineridae. Dry Tortugas, Florida. Holotype in Referred to Marphysa sanguinea (Montagu), A.M.N.H. Referred to Lumbrineris, near latreillii 1815. See Hartman, 1944b, p. 128. Audouin and Milne-Edwards, 1834. See Systema- Marphysa aransensis Treadwell, 1939a, p. 5, tic Discussion. figs. 16, 17. Eunicidae. Port Aransas, Texas. Lumbrinereis similabris Treadwell, 1926d, pp. 5- Holotype in A.M.N.H. See Systematic Discus- 6, figs. 6-10. Lumbrineridae. Bering Strait, sion. Alaska. Holotype in A.M.N.H. Referred to Lum- Marphysa bellii oculata Treadwell, 1921d, pp. brineris similabris (Treadwell), 1926. See 61-64, pl. 5, figs. 13, 14. Eunicidae. Key West Systematic Discussion. Harbor, Florida. Holotype in A.M.N.H. See Lumbrinereis singularisetis Treadwell, 1931d, Systematic Discussion. p. 1, figs. 1-3. Lumbrineridae. Monterey, Cali- Marphysa brevitentaculata Treadwell, 1921d, fornia. Holotype in A.M.N.H. Referred to Lum- pp. 69-70, figs. 235-243, pl. 6, figs. 13, 14. Euni- brineris zonata (Johnson), 1901. See Systematic cidae. West Indies. Holotype in A.M.N.H. See Discussion. Systematic Discussion. Lysidice fusca Treadwell, 1922a, pp. 154-155, Marphysa fragilis Treadwell, 1911, pp. 2-5, figs. 42-44, pl. 6, figs. 7-13. Eunicidae. Samoa figs. 1-7. Eunicidae. Loggerhead Key, Florida, and Fiji. Holotype in A.M.N.H. Referred to shore. Holotype in A.M.N.H. Referred to Mar- Lysidice ?collaris Grube, 1870. See Systematic physa regalis Verrill, 1900. See Systematic Dis- Discussion. cussion. Lysidice parva Treadwell, 1922a, pp. 155-156, Marphysa languida Treadwell, 1921d, pp. 73- figs. 45, 46, pl. 6, figs. 14-17. Eunicidae. Samoa. 75, figs. 257-268. Eunicidae. Guanica Harbor, Holotype in A.M.N.H. Referred to Lysidice Puerto Rico. Holotype in A.M.N.H. Referred ?coUaris Grube, 1870. See Systematic Discus- to Marphysa minima (Hansen), 1882. See Sys- sion. tematic Discussion. Lysidice sulcata Treadwell, 1901b, p. 200, figs. Marphysa nobilis Treadwell, 1917b, pp. 265-266, 47, 48. Eunicidae. Puerto Rico. Holotype in pl. 3, figs. 3-9. Eunicidae. Southern Florida. Holo- U.S.N.M. Referred to Lysidice collaris Grube, type in A.M.N.H. Referred to Marphysa san- 1870. See Fauvel, 1919, p. 477. guinea (Montagu), 1815. See Systematic Dis- Lysidice tortugae Treadwell, 1921d, pp. 85-86, cussion. figs. 298-304. Eunicidae. Florida. Holotype in Marphysa orientalis Treadwell, 1936a, pp. 266- A.M.N.H See Systematic Discussion. 268, figs. 18i-o. Eunicidae. Near Amoy, China. Macellicephala maculosa Treadwell, 1931a, pp. Holotype in U.S.N.M. See Systematic Discussion. 313-315, figs. la-g. Polyodontidae. Marinduque Marphysa simplex Treadwell, 1922a, pp. 151- Island, Philippine Islands, in 106 fathoms. Holo- 152, pL. 5, figs. 8-12. Eunicidae. Suva Harbor, type in U.S.N.M. Referred to Eupanthalis macu- Fiji. Holotype in A.M.N.H. Referred to Marphysa losa (Treadwell), 1931. See Hartman, 1938d, mossambica (Peters), 1854. See Systematic Dis- p. 125. cussion. Maldane cristata Treadwell, 1923b, pp. 9-10, Marphysa viridis Treadwell, 1917b, pp. 264-265, figs. 5-8. Maldanidae. Baja California, Mexico, pl. 2, figs. 13-18, pl. 3, figs. 1-2. Eunicidae. in 475 fathoms. Paratype in A.M.N.H. See Southern Florida, shore. Holotype in A.M.N.H. Systematic Discussion. Referred to Marphysa sanguinea (Montagu), Maldane philippinensis Treadwell, 1931a, pp. 1815. See Systematic Discussion. 320-321, figs. 4a-d. Maldanidae. Off Si Amil Is- Melaenis tropicus Treadwell, 1934a, pp. 1-2, land, Philippine Islands, in 890 fathoms. Holo- pl. 1, figs. 1-6. Aphroditidae. Off Virgin Islands type in U.S.N.M., not seen. in 300 fathoms. Holotype in U.S.N.M. Referred BULLETIN AMER. Mus. NAT. HIST. VOL. 109, PLATF. 21

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Aaron Louis Treadwell 1866-1947 From a photograph in the possession of Treadwell's daughter, Mrs. Beardsley Ruml 1956 HARTMAN: ANNELIDS 255 to Hermoine hystrix (Savigny), 1818. See Hart- in U.S.N.M. See Systematic Discussion. man, 1944a, p. 9. Nereis brevicirrata Treadwell, 1920b, pp. 467- Metalonome brunnea Treadwell, 1917b, p. 268, 468, figs. 1-4. Nereidae. Santos, Brazil. Holotype pl. 3, figs. 24-27. Sabellidae. Nassau Harbor, in U.S.N.M. Referred to Perinereis brevicirrata Bahamas. Holotype in A.M.N.H. Referred to (Treadwell), 1920. See Hartman, 1938b, p. 14. Sabellastarte magnifica (Shaw), 1800. See Sys- Nereis brevicirrata Treadwell, 1929d, pp. 3-5, tematic Discussion. figs. 9-14. Nereidae. Key Largo, Florida. Holo- Metamphinome multibranchiata Treadwell, type in A.M.N.H. Referred to Nereis pelagica 1940, pp. 1-2, figs. 1-3. Amphinomidae. Galveston, largoensis Treadwell, 1931. See Systematic Dis- Texas, on a floating log. Holotype in A.M.N.H. cussion. Referred to Hipponoe multibranchiata (Tread- Nereis (Ceratonereis) alaskensis Treadwell, well), 1940. See Hartman, 1951, p. 29. 1921c, pp. 1-3, figs. 1-4. Nereidae. Valdez Har- Monorchos philippinensis Treadwell, 1926b, bor, Alaska, in 200 fathoms. Holotype in pp. 190-193, figs. 18-20. Sabellariidae. Between U.S.N.M. Referred to Ceratonereis paucidentata Siquijor and Bohol Islands, Philippine Islands, (Moore), 1903. See Hartman, 1938b, p. 13. in 805 fathoms. Referred to Lygdamis philip- Nereis (Ceratonereis) bartletti Treadwell, pinensis (Treadwell), 1926. See Hartman, 1944c, 1937a, pp. 30-31, figs. 8-13. Nereidae. Off Green- pp. 334-335. land in 129 fathoms. Holotype in U.S.N.M. Re- Myriana cirrata Treadwell, 1931b, pp. 2-3, figs. ferred to Ceratonereis hircinicola (Eisig), 1870. 2a-c. Syllidae. Chesapeake Bay, Maryland, in 16 See Hartman, 1938b, p. 13. to 46 meters. Holotype in U.S.N.M., not seen. Nereis decora Treadwell, 1932b, pp. 15-17, Mysta maculata Treadwell, 1920a, pp. 593-594, figs. 6-11. Nereidae. S-ao Paulo, Brazil. Holotype figs. 1-4. Phyllodocidae. Sulade Island, Philippine in U.S.N.M. Referred to Nereis riisei Grube, Islands, in 24 fathoms. Holotype in U.S.N.M. 1856. See Hartman, 1938a, p. 7. Referred to Mysta ornata (Grube), 1878. See Nereis disparsetosa Treadwell, 1932b, pp. 13-15, Fauvel, 1933, p. 17. figs. 1-5. Nereidae. Sao Paulo, Brazil. Holotype Mystides gracilis Treadwell, 1941d, pp. 26-27, in U.S.N.M. Referred to Pseudonereis palpata figs. 5-7. Phyllodocidae. Off Bermuda, planktonic (Treadwell), 1923. See Hartman, 1938b, p. 14. at surface. Holotype in A.M.N.H. Referred to Nereis heterocirrata Treadwell, 1931c, pp. 1-2, Anaitides tmadeirensis (Langerhans), 1879. See figs. la-e. Nereidae. Near Chochi, Japan. Holo- Systematic Discussion. type in U.S.N.M., not seen. Nainereis mutilata Treadwell, 1931d, pp. 5-6, Nereis largoensis Treadwell, 1931d, p. 3. figs. 13-18. Orbiniidae. Montego Bay, Jamaica, Nereidae. Key Largo, Florida. Holotype in West Indies. Holotype in A.M.N.H. Referred to A.M.N.H. Referred to Nereis pelagica largoensis Naineris mutilata Treadwell, 1931. See Systematic Treadwell, 1931. See Systematic Discussion. Discussion. Nereis (Leptonereis) acuta Treadwell, 1923a, Namonereis kartaboensis Treadwell, 1926c, pp. pp. 3-5, figs. 1-7. Nereidae. Santos, Brazil. Holo- 101-103, fig. 33. Nereidae. Kartabo, British type in U.S.N.M. Referred to Laeonereis culveri Guiana. Holotype in A.M.N.H. Referred to (Webster), 1879. See Hartman, 1938b, p. 14. Lycastis kartaboensis (Treadwell), 1926. The type Nereis (Leptonereis) distorta Treadwell, 1936a, has been dried. pp. 273-275, figs. 20a-f. Nereidae. Near Amoy, Neanthes obscura Treadwell, 1928a, p. 472, China. Holotype in U.S.N.M., not seen. Based only figs. 50-53. Nereidae. Cocos Islands, west of Costa on a male in epitokous stage. Rica. Holotype in A.M.N.H. Referred to Per- Nereis (Neanthes) amoyensis Treadwell, 1936a, nereis helleri (Grube), 1878. See Systematic Dis- pp. 272-273, figs. 10j-m. Nereidae. Near Amoy, cussion. China. Holotype in U.S.N.M. Referred to Nereis Neosabellides alaskensis Treadwell, 1943e, pp. amoyensis (Treadwell), 1936. See Systematic 1-2, figs. 1-5. Ampharetidae. Nunivak Island, Discussion. Alaska. Holotype in A.M.N.H. Referred to Nereis (Neanthes) austratis Treadwell, 1923c, Asabellides sibirica (Wiren), 1883. See Systema- pp. 1-3, figs. 1-5. Nereidae. Off Punta del Este, tic Discussion. Maldonado, Uruguay. Holotype in U.S.N.M. Re- Nereis ambiguus Treadwell, 1937b, pp. 149-151, ferred to Neanthes succinea (Frey and Leuckart), pI. 2, figs. 19-24. Nereidae. Sulphur Bay, Clarion 1847. See Monro, 1938, p. 313. Island, Mexico, in 20 fathoms. Holotype in Nereis (Neanthes) linea Treadwell, 1936a, pp. A.M.N.H. Referred to Nereis riisei Grube, 1856. 268-270, figs. 19a-e. Nereidae. Near Amoy, China. See Systematic Discussion. Holotype in U.S.N.M. Referred to Peninereis Nereis arroyensis Treadwell, 1901b, pp. 193- aibuhitenssis Grube, 1878. See Hartman, 1938b, 194, figs. 30-31. Nereidae. Puerto Rico. Holotype p. 15. 256 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Nereis (Neanthes) orientalis Treadwell, 1936a, Nuchubranchia palmata Treadwell, 1928a, pp. pp. 270-272, figs. 19f-i. Nereidae. Near Amoy, 481-482, figs. 66-68. Typhloscolecidae. Off the China. Holotype in U.S.N.M. Referred to Peri- Galapagos Islands in 400-600 fathoms. Holotype nereis aibukitensis Grube, 1878. See Hartman, in A.M.N.H. Referred to Travisiopsis coniceps 1938b, p. 15. (Chamberlin), 1919. See Systematic Discussion. Nereis (Neanthes) palpata Treadwell, 1923a, Odonotosyllis octodentata Treadwell, 1917b, pp. 5-9, figs. 8-15. Nereidae. Sao Paulo, Brazil. p. 257, pl. 1, figs. 1-3. Syllidae. Dry Tortugas, Paratype in U.S.N.M. Referred to Pseudonereis Florida. Holotype in A.M.N.H. A valid species. palpata (Treadwell), 1923. See Hartman, 1938b, Oenone brevimaxillata Treadwell, 1931d, pp. p. 14. This may be identical with Pseudonereis 1-3, figs. 4-9. Lysaretidae, Mexico. Referred to atopodon Chamberlin, 1919 (Hartman, 1938b, Lysarete brasiliensis Kinberg, 1865. See Hartman, p. 15). 1942c, pp. 101-102. Nereis (Neanthes) paucidentata Treadwell, Onuphis branchiata Treadwell, 1931a, pp. 317- 1939a, p. 6, fig. 25. Nereidae. Charlestown, Massa- 320, figs. 3a-g. Onuphidae. Marinduque Island, chusetts. Holotype in A.M.N.H. Referred to Philippine Islands, in 106 fathoms. Holotype in Nereis riises Grube, 1856. See Systematic Discus- U.S.N.M., not seen. sion. Onuphis stigmatis Treadwell, 1922b, pp. 176- Nereis (Neanthes) varia Treadwell, 1941a, p. 178, figs. 22-34. Onuphidae. San Juan Island, 3. Nereidae. New name for Nereis (Neanthes) Washington, shore. Holotype in A.M.N.H. Re- paucidentata Treadwell, 1939. Referred to Nereis ferred to Nothria stigmatis (Treadwell), 1922. rissei Grube, 1856. See Systematic Discussion. See Hartman, 1944b, pp. 89-91, pl. 11, figs. 240- Nereis notomacula Treadwell, 1914b, pp. 191- 247. 192, pl. 11, figs. 8-12. Nereidae. Entrance to San Oophylax minuta Treadwell, 1937a, p. 29, figs. Francisco Bay, California. Syntype in A.M.N.H. 6, 7. Syllidae. Fox Channel, Greenland. Holotype Referred to Platynereis bicanaliculata (Baird), in not seen. Pettibone 1863. See U.S.N.M., (1954, p. 256) Systematic Discussion. referred it to Sphaerosyllis arenaceus Clapar6de, Nereis (Platynereis) integer Treadwell, 1920a, 1863. pp. 595-597, figs. 1-4. Nereidae. Philippine Islands. Ophelina magna Treadwell, 1914b, Holotype in A.M.N.H. Referred to Platynereis pp. 216-218, polyscalma Chamberlin, pl. 12, figs. 34-36. Opheliidae. off southern Cali- 1919. See Hartman, fornia in 3-40 fathoms. Paratype in A.M.N.H. 1938b, p. 15. Referred to Ophelia nagna (Treadwell), 1914. Nereis singularis Treadwell, 1943b, p. 41, figs. See 32. 33-37. Nereidae. East Hartman, 1936, p. of New England, plank- Ophelina mucronata Treadwell, 1914b, pp. 218- tonic at surface. Holotype in U.S.N.M. This ap- 219, pl. 12, figs. 37, 38. Opheliidae. La Jolla, pears to be an immature stage of Ceratonereis California, shore. Paratype in A.M.N.H. Re- mirabils Kinberg, 1866, known from the Gulf ferred to Euzonus Stream. Nereis singularis Wesenberg-Lund, 1949, (Thoracophelia) mucronata p. 278, (Treadwell), 1914. See Systematic Discussion. from Iran is another species and re- Panthalis oculea Treadwell, pp. sembles Neanthes oxypoda (Marenzeller), 1879. 1901b, 188-189, Nereis figs. 14-18. Polyodontidae. Puerto Rico. Holotype spisifera Treadwell, 1929d, pp. 5-6, figs. in U.S.N.M. Referred to Polyodontes oculea 15-20. Nereidae. Puget Sound, Washington. Holo- 1901. See type in A.M.N.H. Referred to Perinereis mon- (Treadwell), Hartman, 1939b, pp. 83- terea (Chamberlin), 1918. See 84, pl. 24, figs. 294-299. Systematic Discus- Panthalis pacifica Treadwell, 1914b, pp. 184- sion. 186, pI. 1-7. Polyodontidae. Nenine minuta Treadwell, 1939a, p. 5, figs. 18- 11, figs. Off San 20. Spionidae. Port Aransas, Diego, California, in 50 to 100 fathoms. Holotype Texas. Holotype in in Los Angeles. See Hartman, 1939b, pp. 87-88, A.M.N.H. Referred to Nerinides agilis (Verrill), pl. 26, figs. 309-312. 1873. See Systematic Discussion. Panthalis pustulata Nicidion fusca-fasciata Treadwell, 1922a, pp. Treadwell, 1924c, pp. 7-9, 156-157, figs. figs. 10-15. Polyodontidae. English Harbor, Anti- 47-50, pl. 7, fig. 5. Eunicidae. Pago qua, West Indies. Pago, Samoa. Holotype in A.M.N.H. Referred to Holotype in U.S.N.M. See Eunice (Nicidion) gracilis Crossland, 1903. See Hartman, 1939b, p. 87. Systematic Discussion. PParamarPhysa teres Treadwell, 1922a, pp. 153- Nicomache antiguensis 154, figs. 40, 41, pl. 6, figs. 2-6. Eunicidae. Pago Treadwell, 1924c, pp. Pago Harbor, Samoa. Holotype in 16-17, figs. 25-29. Maldanidae. Antilles, West See A.M.N.H. Indies. Holotype in U.S.N.M., not Systematic Discussion. seen. This may Parasabella minuta Treadwell, 1941a, p. 4, be referred to Nicomache antillensis Augener, 9-12. figs. 1922, from the same locality. Sabellidae. Sao Sebastio, Brazil. Holotype in A.M.N.H. Referred to Potamila minuta 1956 HARTMAN: ANNELIDS 257 (Treadwell), 1941. See Systematic Discussion. 69-71. Phyllodocidae. Galapagos Islands. Holo- Parasabella sulfurea Treadwell, 1917b, p. 267, type in A.M.N.H. Referred to .Anaitides pl. 3, figs. 16-23. Sabellidae. Dry Tortugas, madeirensis (Langerhans), 1879. See Systematic Florida. Holotype in A.M.N.H. Referred to Discussion. Hypsicomus torquatus (Grube), 1878. See Hart- Phyllodoce variegata Treadwell, 1926a, p. 12. man, 1951, p. 116. Lapsus calami for Phyllodoce violacea, quod vide. Perinereis diversidentata Treadwell, 1943c, Phyllodoce violacea Treadwell, 1926a, pp. 13- pp. 1-2, figs. 1-7. Nereidae. Cape Cross, Africa. 14, figs. 22-24. Phyllodocidae. Suva, Fiji. Holo- Holotype in A.M.N.H. Referred to Pseudonereis type in U.S.N.M. See Systematic Discussion. variegata (Grube), 1856. See Systematic Discus- Pionosyllis manca Treadwell, 1931b, pp. 1-2, sion. figs. la-e. Syllidae. Off Cape Henry, Virginia, in Phalacropisorus attenuatus Treadwell, 1943b, 20 fathoms. Holotype in U.S.N.M., not seen. This p. 34, fig. 14. Iospilidae. Marshall Islands, pelagic appears to be a representative of Exogoninae in 50 meters. Holotype in U.S.N.M. Resembles instead of Syllinae. Phalacrophorus uniformis Reibisch, 1895. See Pista groenlandica Treadwell, 1937a, pp. 33- Systematic Discussion. 35, figs. 14-16. Terebellidae. Cobourg Island, Phalacrophorus maculatus Treadwell, 1943b, p. Baffin Bay, Arctic Ocean. Holotype in U.S.N.M., 34, figs. 11-13. Iospilidae. Off Ecuador at surface. paratype in A.M.N.H. Referred to Pista maculata Holotype in U.S.N.M., paratype in A.M.N.H. (Dalyell), 1853. See Pettibone, 1954, p. 324 Referred to Phalacrophorus pictus Greeff, 1879. Placostegus calciferus Treadwell, 1929d, pp. See Systematic Discussion. 12-13, figs. 34-36. Serpulidae. Juba Cove, Guanica Phalacrophorus niger Treadwell, 1943b, pp. Harbor, Puerto Rico. Holotype in A.M.N.H. 33-34, fig. 15. Iospilidae. North central Atlantic Referred to Sclerostyla ctenactis M6rch, 1863. Ocean, pelagic in 50 meters. Holotype in U.S.N.M. See Systematic Discussion. Referred to Zospilus phalacroides Viguier, 1886. Plotobia paucichaeta Treadwell, 1943b, pp. 38- See Systematic Discussion. 39, fig. 26. Tropical southern Pacific Ocean, Phyllochaetopterus verrillii Treadwell, 1943d, planktonic at surface. Holotype in U.S.N.M. pp. 1-3, figs. 4-8. Chaetopteridae. Oahu, Hawaii. Referred to Travisiopsis benhami Monro, 1936. Holotype in A.M.N.H. A valid species. See Friedrich, 1950, p. 316. Phyllodoce fusca-cirrata Treadwell, 1926a, pp. Polydora californica Treadwell, 1914b, pp. 203- 14-15, figs. 25-27. Phyllodocidae. Pango Pango, 204, pl. 12, figs. 23-29. Spionidae. Locality un- Samoa. Holotype in U.S.N.M. Referred to known. Paratype in A.M.N.H. Referred to Anaitides tenuissima (Grube), 1878. See Syste- Boccardia sp. See Systematic Discussion. matic Discussion. Polynoe alba Treadwell, 1906, pp. 1149-1150, Phyllodoce magna-oculata Treadwell, 1901b, p. figs. 5-7. Polynoidae. Honolulu, Hawaii. Holotype 191, figs. 25, 26. Phyllodocidae. Puerto Rico. in U.S.N.M. Referred to Lepidasthenia alba Holotype in U.S.N.M. See Hartman, 1942a, pp. (Treadwell), 1906. See Hartman, 1942a, p. 103. 110-111, figs. 9c-d. Polynoe branchiata Treadwell, 1901b, p. 186, Phyllodoce nicoyensis Treadwell, 1928b, pp. figs. 5-7. Polynoidae. Puerto Rico. Holotype in 1-3, figs. 1-3. Phyllodocidae. Gulf of Nicoya, U.S.N.M. Referred to Chaetacanthus magnificus Costa Rica. Holotype in U.S.N.M., not seen. (Grube), 1875. See Seidler, 1924, pp. 97-98. PhyUodoce panamensis Treadwell, 1917a, pp. Polynoe lucida Treadwell, 1906, pp. 1150-1151, 428-430, figs. 1-2. Phyllodocidae. Chame Point, figs. 8-10. Polynoidae. Off Hawaii in 83-113 Panama. Holotype in U.S.N.M. Referred to fathoms. Holotype in U.S.N.M. Referred to Anaitides panamensis (Treadwell), 1917. See Lepidasthenia alba (Treadwell), 1906. See Hart- Monro, 1933, p. 24. man, 1942a, p. 103. Phyllodoce pulla Treadwell, 1926a, pp. 12-13, Polynoe mutilata Treadwell, 1906, pp. 1152- figs. 19-21. Phyllodocidae. Japan. Holotype in 1153, figs. 12-15. Polyodontidae. Off Hawaii in U.S.N.M., not seen. This may agree with Anaitides 328 and 319 fathoms. Holotype in U.S.N.M., madeirensis (Langerhans), 1879. paratype in A.M.N.H. Referred to Eupanthalis Phyllodoce stigmata Treadwell, 1925c, pp. 116- mutilata (Treadwell), 1906. See Hartman, 1938d, 118, fig. 11. Phyllodocidae. Off Hawaii, pelagic. p. 123. Holotype deposited in Hawaii, not seen. Polynoe nodosa Treadwell, 1901b, p. 187, figs. Phyllodoce tortugae Treadwell, 1917b, p. 262, 8, 9. Polynoidae. Puerto Rico. Holotype in pl. 2, figs. 4-6. Phyllodocidae. Loggerhead Key, U.S.N.M. Referred to Hermenia verruculosa Florida. Holotype in A.M.N.H. See Systematic Grube, 1856. See Treadwell, 1911, pp. 9-11, figs. Discussion. 23-26. Phyllodoce varia Treadwell, 1928a, p. 467, figs. Polynoe spicula Treadwell, 1906, pp. 1151-1152, 258 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 fig. 11. Polynoidae. Monterey Bay, California. Scoloplos rufa Treadwell, 1941b, p. 1, figs. Holotype in 1-6. U.S.N.M. Referred to Lepidonotus Orbiniidae. Galveston, Texas. Holotype in caelorus Moore, 1903. See Hartman, 1938d, p. A.M.N.H. Referred to Haploscoloplos robustu,s 108. (Verrill), 1873. See Systematic Discussion. Polynoella brunnea Treadwell, 1943b, pp. 31- Semiodera glabra Treadwell, 1928a, pp. 479- 32, figs. 1-3. Polynoidae. Off Ecuador, planktonic. 480, figs. 62-65. Flabelligeridae. Off Galapagos Holotype in U.S.N.M., not seen. Islands in 15 feet. Holotype in A.M.N.H. Re- Polyodontes californicus Treadwell, 1941c, pp. ferred to Semiodera cariboum (Grube), 1856. 20-21, figs. 9-12. Polyodontidae. Chamela Bay, See Systematic Discussion. Mexico, in 16 fathoms. Holotype in A.M.N.H. Sphaeropomatus miamiensis Treadwell, 1934b, Referred to Polyodontes panamensis (Chamber- pp. 338-344, 9 figs. Serpulidae. Miami River, lin), 1919. See Systematic Discussion. Florida, on carapace of shrimp. Holotype in Polyophthalmus incertus Treadwell, 1936b, pp. U.S.N.M. See Systematic Discussion. 61-62, figs. 27-29. Opheliidae. Nonsuch Island, Spio acuta Treadwell, 1914b, pp. 199-201, pl. Bermuda. Holotype in A.M.N.H. Referred to 11, figs. 14-20. Spionidae. San Diego, California. Polyophthalmus pictus (Du.jardin), 1839. See Sys- Holotype in Los Angeles. Referred to Nerinides tematic Discussion. acuta (Treadwell), 1914. See Hartman, 1941, Polyophthalmus papillatus Treadwell, 1943c, p. 3, figs. 19, 20. pp. 294-296, pl. 45, figs. 1-8. Opheliidae. Lobito, Africa. Holo- Spio hirsuta Treadwell, 1928a, pp. 478-479, type in A.M.N.H. Referred to Polyophthalmus figs. 54-57. Spionidae. East of Cocos Islands, pictus (Dujardin), 1839. See Systematic Discus- Pacific Panama. sion. Holotype in A.M.N.H. Referred to Nerine cirratulus hirsuta (Treadwell), 1928. Pomatoceros davaoensis Treadwell, 1942, pp. See Systematic Discussion. 4-5, figs. 12-14. Serpulidae. Gulf of Davao, Philip- Spirographis braziliensis Treadwell, 1932b, pp. pine Islands. Holotype in A.M.N.H. Referred to 18-19, figs. 15-19. Sabellidae. Pomatoceros caeruleus (Schmarda), 1861. See in Brazil. Holotype Systematic Discussion. U.S.N.M., not seen. Potamilla californica Sthenelais grubei Treadwell, 1901b, pp. 187-188, Treadwell, 1906, p. 1178. figs. 10-13. Sigalionidae. Puerto Rico. Sabellidae. Monterey Bay, California. Holotype Holotype in in U.S.N.M. Referred to Leanira grubei (Tread- U.S.N.M. Referred to Hypsicomus californicus well), 1901. See (Treadwell), 1906. See Hartman, 1942a, p. 133. Hartman, 1942a, pp. 106-107. Potamilla Streblosoma crassibranchia Treadwell, 1914b, elongata Treadwell, 1906, p. 1178, pp. 31. figs. 73-75. Sabellidae. Off Hawaii in 208-209, pl. 12, figs. 30, Terebellidae. San 256-277 Clemente Island, California. Holotype in Los An- fathoms. Holotype in U.S.N.M., paratype in geles. See A.M.N.H. Referred to Potamethus elongatus Systematic Discussion. (Treadwell), 1906. See Streblosoma crassibranchiata Treadwell, 1914b, Hartman, 1942a, pp. 134- p. 234. 135, figs. 15b-d; Berkeley and Berkeley, 1951, Lapsus calami for Streblosoma crassi- pp. 333-334. branchia, quod vide. Streblosoma magma 1937b, 155- Prionospio Plumosa Treadwell, 1931b, pp. 3-5, 156, Treadwell, pp. figs. 3a-d. Spionidae. Chesapeake Bay, Maryland. figs. 26-28. Terebellidae. Arena Bank, Holotype in U.S.N.M. Mexico, in 45 fathoms. Holotype in A.M.N.H. Referred to Prionospio Referred to treadwelli Hartman, 1951. See Hartman, 1951, Thelepus crispus Johnson, 1901. See P. 84. 1 Systematic Discussion. Sabella alba Treadwell, 1917b, pp. 266-267, pl. Streblosoma verrilli Treadwell, 1911, pp. 11- 3, figs. 10-15. Sabellidae. Dry Tortugas, Florida. 12, figs. 27-29. Terebellidae. Near Fort Jefferson, Holotype in A.M.N.H. Florida, on shore. Holotype in A.M.N.H. Re- Referred to Hypsicomus ferred to elegans (Webster), 1884. See Hartman, 1951, Thelepus setosus (Quatrefages), 1865. p. 115. See Systematic Discussions. Sabellaria setosa Treadwell, 1906, pp. Stylarioides dubius Treadwell, 1929a, pp. 1181, figs. 1180- figs. 25-27. 9-10, 78-81. Sabellariidae. Off Hawaii in ida. Flabelligeridae. Dry Tortugas, Flor- 253-382 fathoms. Holotype in U.S.N.M. Referred Paratypes in A.M.N.H. Referred to Pherusa to Phalacrostemma setosa (Treadwell), 1906. See dubia (Treadwell), 1929. See Systematic Dis- Hartmnan, 1944c, p. 330. cussion. Scolecolepis alaskensis Treadwell, 1914b, pp. Stylaroides glabra Treadwell, 1901b, p. 208, 201-203, p1. 12, figs. 21, 22. Spionidae. Shumagin figs. 73-75. Flabelligeridae. Puerto Rico. Holotype Islands, Alaska. Paratype in A.M.N.H. Referred in U.S.N.M. Referred to 7Semiodera glabra to Nerine alaskensis (Treadwell), 1914. See Hart- (Treadwell), 1901. See Systematic Discussion. man, 1948a, p. 8. Syllis complanata Treadwell, 1901b, p. 183, fig. 1. Syllidae. Puerto Rico. Holotype in 1956 HARTMAN: ANNELIDS 259

U.S.N.M., not seen. This may be a species of fornia, in 10 fathoms. Holotype in Los Angeles. Typosyllis Langerhans, as it seems to have only Referred to Pherusa inflata (Treadwell), 1914. composite setae. See Hartman, 1952, p. 71. Terebella hiata Treadwell, 1931e, pp. 80-81, Trypanosyllis adamanteus Treadwell, 1914c, figs. 14c-f. Terebellidae. Haiti. Holotype in pp. 235-237, figs. 1-3. Syllidae. San Francisco, A.M.N.H. Referred to Polymniella au-rantiaca California, along shore. Paratype in A.M.N.H. Verrill, 1900. See Systematic Discussion. Redescribed by Rioja, 1941, pp. 697-698, pl. 2, Terebella parvabranchiata Treadwell, 1906, p. figs. 16-20. 1175, fig. 71. Terebellidae. Off Hawaii in 444 -478 Typosyllis aciculata Treadwell, 1945, pp. 1-2, fathoms. Holotype in U.S.N.M., not seen. figs. 1-5. Syllidae. Pacific Grove, California. Holo- Terebellides tentaculata Treadwell, 1906, p. type in A.M.N.H. Redescribed by Reish, 1950, 1177, fig. 72. Ampharetidae. Off Hawaii in 278- pp. 1-5, figs. 1-8. 473 fathoms. Holotype in U.S.N.M., paratype in Typosyllis crassicirrata Treadwell, 1925c, pp. A.M.N.H. Referred to Melinnexis tentaculata 113-115, fig. 10. Syllidae. Laysan Island, Hawaii. (Treadwell), 1906. See Systematic Discussion. Holotype in Hawaii, not seen. Thelepus branchiatus Treadwell, 1906, pp. 1176- Uncinereis lutea Treadwell, 1928a, pp. 469-471, 1177. Terebellidae. Off Hawaii in 122-143 figs. 40-49. Nereidae. Off the West Indies on fathoms. Holotype in U.S.N.M., not seen. sargassum. Paratypes in A.M.N.H. Referred to Thelepus crassibranchiatus Treadwell, 1901b, Platynereis dumerilii (Audouin and Milne-Ed- p. 206, figs. 69-71. Terebellidae. Puerto Rico. wards), 1834. See Systematic Discussion. Holotype in U.S.N.M. Referred to Thelepus Uncinereis trimaculosa Treadwell, 1940, p. 3, cincinnatus (Fabricius), 1780. See Augener, 1906, figs. 4-9. Nereidae. Galveston, Texas. Holotype in p. 182; Hessle, 1917, p. 213. A.M.N.H. Referred to Platynereis dumenrilii Thelepus haitiensis Treadwell, 1931e, pp. 79- (Audouin and Milne-Edwards), 1934. See Hart- 80, figs. 14a-b. Terebellidae. Haiti. Holotype in man, 1951, p. 47. A.M.N.H. See Systematic Discussion. Vanadis collata Treadwell, 1928a, pp. 462-463, Tomopteris longisetis Treadwell, 1936b, pp. figs. 27, 28. Alciopidae. Off Galapagos Islands in 58-59, figs. 18-21. Tomopteridae. Nonsuch Island, 400 fathoms. Holotype in A.M.N.H. Referred to Bermuda, pelagic in 500-1000 fathoms. Holotype Vanadis crystaUina Greeff, 1876. See Systematic in A.M.N.H. Referred to Tomopteris nisseni Discussion. Rosa, 1908. See Systematic Discussion. Vanadis fusca-punctata Treadwell, 1906, pp. Tomopteris opaca Treadwell, 1928a, pp. 463- 1159-1160, figs. 29-31. Alciopidae. Off Hawaii at 464, fig. 29. Tomopteridae. Off Galapagos and surface to 238 fathoms. Holotype in U.S.N.M. Cocos Islands in 550-700 fathoms. Holotype in Referred to Vanadis formosa Claparede, 1870. A.M.N.H. Referred to Tomopteris nisseni Rosa, See St0p-Bowitz, 1948, p. 25. 1908. See Systematic Discussion. Vanadis minuta Treadwell, 1906, pp. 1158- Tomopteris tentaculata Treadwell, 1928a, pp. 1159, figs. 25-28. Alciopidae. Off Hawaii, pelagic 464-465, figs. 30-31. Tomopteridae. off Galapagos in 4 fathoms. Holotype in U.S.N.M. Redescribed Islands, planktonic at surface. Holotype in by Stop-Bowitz, 1951, pp. 7-8. A.M.N.H. See Systematic Discussion. Vanadis uncinata Treadwell, 1943b, p. 36, figs. Torea fasciata Treadwell, 1943b, p. 35, figs. 23, 24. Alciopidae. Equatorial Pacific Ocean, 19, 20. Alciopidae. Between United States and planktonic in 50 meters. Holotype in U.S.N.M., Hawaii, planktonic in 50 meters. Holotype in not seen. U.S.N.M., not seen. 1936b, 65, 62- Vermilia glandulata Treadwell, p. Travisiopsis atlantica Treadwell, 1936b, pp. pp. 36-39. Serpulidae. Gurnets Rock, Bermuda, in 63, figs. 30-33. Typhloscolecidae. Nonsuch Island, 35 feet. Holotype in A.M.N.H. Referred to Bermuda, in 600-700 fathoms. Holotype in Vermiliopsis bermudiensis (Bush), 1904. See A.M.N.H. Referred to Travisiopsis lanceolata Hartman, 1942b, p. 90, figs. 154, 155. Southern, 1911. See Systematic Discussion. Vermiliopsis hawaiiensis Treadwell, 1943d, pp. Trophonia inflata Treadwell, 1914b, pp. 213- in A.M.N.H. Flabelligeridae. Southern Cali- 3-4, figs. 14, 15. Serpulidae. Holotype 214, pl. 12, fig. 33. Referred to Vermiliopsis multiannulata (Moore), fornia along shore to 28 fathoms. Paratype in Discussion. to Pherusa inflata (Tread- 1923. See Systematic A.M.N.H. Referred Vermiliopsis torquata Treadwell, 1943d, p. 4, well), 1914. See Hartman, 1952, pp. 71-74, figs. Hawaii. in 1, 2. figs. 16, 17. Serpulidae. Syntypes A.M.N.H. Referred to Vermiliopsis multianrnlata Trophonia minuta Treadwell, 1914b, p. 213, Discussion. pl. 12, fig. 32. Flabelligeridae. Off La Jolla, Cali- (Moore), 1923. See Systematic ALPHABETICAL LIST OF VALID SPECIES, WITH SYNONYMS Acholoe astericola (delle Chiaje), 1841, as Ceratonereis hircinicola (Eisig), 1870; as Nereis Acholoe orbiculata Treadwell, 1921 (Ceratonereis) bartletti Treadwell, 1937 Ampharete arctica Malmgren, 1866, as Ampharete Ceratonereis mirebilis Kinberg, 1866, as Cerato- brevibranchiata Treadwell, 1926 nereis singularis Treadwell, 1929, and perhaps Ampharete eupalea Chamberlin, 1920, as Nereis singulanris Treadwell, 1943 Ampharete seribranchiaea Treadwell, 1926 Ceratonereis paucidentata (Moore), 1903, as Anaitides madeirensis Langerhans, 1879, as Nereis (Ceratonereis) alaskensis Treadwell, Anaitides minuta Treadwell, 1937, Mystides 1921 gracil4s Treadwell, 1941, and PhyUodoce vaeia Chaetacanthus magnificus (Grube), 1875, as Treadwell, 1928 Lepidonotus pilosus Treadwell, 1937, and Anaitides panamensis (Treadwell), 1917, as Polynoe branchiata Treadwell, 1901 Phylodoce panamensis Treadwell, 1917 Cirratulus elongatus Treadwell, 1901, question- Anaitides pula (Treadwell), 1926, as Phyllodoce able pult Treadwell, 1926 Cirriformia ?chefooensis (Grube), 1877, as Anaitides tenuissima (Grube), 1878, as Phyllo- Cirratulus Treadwell, 1936 doce branchiatu-s fusce-cirrata Treadwell, 1926 Cirriformia filigere (delle Chiaje), 1841, as Ancistrosyllis tentaculata Treadwell, 1941 Audouinea oculaea Treadwell, 1932, and Aphrodita magna Treadwell, 1925 Audouinea 1936 Arabela pygidia Treadwell, 'iricolor (Montagu), 1804, as Arabella Cirriformia hawaiensis, new name for Audouinea dubie Treadwell, 1922, and questionably Ara- branchiata bela setosa Treadwell, 1943, preoccupied Treadwell, 1921, and Drilonereis Cirniformia punctata (Grube), 1859, as Cirratulus pinnata Treadwell, 1921 nigromaculata 1901 ArabeUa iridescens Treadwell, Treadwell, 1906 Cirriformia semicincta (Ehlers), 1905, as Audou- ArabeUa semimaculata (Moore), 1911, as Ara- inea maculata beUa pacifica Treadwell, Treadwell, 1929 1941 Cirriformia tentaculata (Montagu), 1808, as Cir- ArabeUa sp., as Aracoda attenuata Treadwell, ratulus pallidus Treadwell, 1931, and Cirratulus 1911 multicirratus Treadwell, 1936 Arctonoi tuberculkat (Treadwell), 1906, as Corynocephalus 1943 Harmothoe tuberculata Treadwell, 1906 magnachaeeus Treadwell, .Dalhousiella longicirrata (Treadwell), 1901, as Arctonoi vittata (Grube), 1855, as Lepidasthenia Castalia longicirrata Treadwell, 1901 elegans Treadwell, 1941 Dasybranchus rectus Treadwell, 1901, Armandia maculata (Webster), 1884, as Ammo- able question- trypane bermudiensis Treadwell, 1936 Drieschia pellucida Moore, 1903, as Dnieschia Asabellides sibirica (Wiren), 1883, as Neosabel- atlantica Treadwell, 1936 lides aleskensis Treadwell, 1943 Drilonereis lumbricus Asychis fimbriate (Treadwell), 1934, Treadwell, 1922 as Malda- Drilonereis planiceps as nella fimbriata Treadwell, 1934 nereis (Grube), 1878, Drilo- Autolytus pacificus Treadwell, 1943 paucidentata Treadwell, 1922 Autolytus varius Treadwell, 1914 Drilonereis spatula (Treadwell), 1911, as Aracoda spatula Treadwell, 1911, and Drilonereis brun- ?Benthoscoles microcarunculata (Treadwell), nea 1901, as Amphinome microcarunculata Treadwell, 1921 Tread- Drilonereis sp., juvenile, as Drilonereis similis well, 1,901 1921 Boccardie sp., as Polydora californica Treadwell, Treadwell, 1914 Epitoka pelagica Treadwell, 1943, questionable as to family status Branchiomma nigromaculata (Baird), 1865, as Eteone Dasychone ponce Treadwell, .longa (Fabricius), 1780, as Eteone 1901, Dasycho- tuberculata Treadwell, 1922 nopsis arenosa Treadwell, 1924, and Dasychone Eteone loandensis Treadwell, 1943 pacifica Hartman, 1936, as Eteone macu- Callizonelle pigmenta Treadwell, 1943, perhaps lata Treadwell, 1922 Krohnia sp. Euclymene corallicola (Treadwell), 1929, as Capitellides teres Treadwell, MaldaneUa corallicola Treadwell, 1929 1939 Eulalia foliosa 1924 Ceratonereis anchylochaeta (Horst), 1924, as Eulalia Treadwell, Ceratonereis longicauda Treadwell, 1943 myriacyclum (Schmarda), 1861, as Eulalia quinquelineata Treadwell, 1901 260 1956 HARTMAN: ANNELIDS 261 Eunice afra Peters, 1854, as Leodice aciculata Eurato punctata Treadwell, 1926, questionable Treadwell, 1922, Leodice flava-punctata Tread- Euzonus (Thoracophelia) mucronata (Tread- well, 1922, Leodice bucciensis Treadwell, 1921, well), 1914, as Ophelina mucronata Treadwell, Leodice guanica Treadwell, 1921, Leodice 1914 suviensis Treadwell, 1922, and questionably Fabricia sabella (Ehrenberg), 1836, as Haplo- Leodice notata Treadwell, 1921 branchus atlanticus Treadwell, 1932 Eunice antennata Savigny, 1818, as Eunice in- Gattyana cirrosa (Pallas), 1766, as Gattyana terrupta Treadwell, 1906 imbricata Treadwell, 1926 Eunice argentinensis (Treadwell), 1929, as Glycera ?capitata Oersted, 1843, as Hemipodia Leodice argentinensis Treadwell, 1929 canadensis Treadwell, 1937 Eunice armillata (Treadwell), 1922, as Leodice Glycera tesselata Grube, 1863, as Glycera abran- armnillata Treadwell, 1922, and Leodice gracili- chiata Treadwell, 1901, and Glycera spadix cirrata Treadwell, 1922 Treadwell, 1943 Eunice auriculata Treadwell, 1900 and 1901 Goniada brunnea Treadwell, 1906 Eunice bilobata Treadwell, 1906 Goniada oculata Treadwell, 1901, questionably Eunice crassi-tentaculata (Treadwell), 1922, as Glycinde sp. Leodice crassi-tentaculata Treadwell, 1922 Goniada quinquelabiata Augener, 1906, as Goni- Eunice filamentosa Grube, 1856, as Leodice ada magna Treadwell, 1945 spongicola Treadwell, 1921 Goniada teres Treadwell, 1931 Eunice grubei Gravier, 1900, as Leodice biformi- Halosydna latior Chamberlin, 1919, as Halosydna cirrata Treadwell, 1922 obtusa-cirrata Treadwell, 1937 Eunice hawajiensis Treadwell, 1906 Halosydnella fusca-maculata (Treadwell), 1924, Eunice nicidioformis Treadwell, 1906 as Halosydna fusca-maculata Treadwell, 1924, Eunice (Nicidion) cariboea Grube, 1856, as Eunice and Halosydna fuscomarginata Treadwell, culebra Treadwell, 1901 1924 Eunice (Nicidion) gracilis Crossland, 1903, as HalosydneUa grisea (Treadwell), 1929, as Nicidion fusca-fasciata Treadwell, 1922 Halosydna grisea Treadwell, 1929 Eunice norvegica (Linnaeus), 1767, as Leodice Halosydnella oculata (Treadwell), 1926, as arcturi Treadwell, 1928 Halosydna oculata Treadwell, 1926 Eunice savignyi (Grube), 1878, as Leodice langi Halyplanes gracilis Reibisch, 1893, as Haliplanela Treadwell, 1943 pacifica Treadwell, 1943 Eunice tenuis (Treadwell), 1921, as Leodice Haploscoloplos robustus (Verrill), 1873, as tenuis Treadwell, 1921 Scoloplos rufa Treadwell, 1941 Eunice tubifex Crossland, 1904, as Leodice tubi- Haplosyllis spongicola (Grube), 1855, as Hap- cola Treadwell, 1922 losyllis gula Treadwell, 1924 Eunice vittata (delle Chiaje), 1828, as Leodice Harmothoe crucis (Grube), 1856, as Harmothoe rubrivittata Treadwell, 1921 lanceocirrata Treadwell, 1928, and Harmothoe Eunoe crassa (Treadwell), 1924, as Lagisca lanceolata Treadwell, 1928 crassa Treadwell, 1924 Harmothoe imbricata (Linnaeus), 1767, as Eunoe purpurea Treadwell, 1936, as Acanthico- Harmothoe levis Treadwell, 1937 lepis longicirrata Treadwell, 1941 Harmothoe sylliformia Treadwell, 1928 Eupanthalis maculosa (Treadwell), 1931, as Harmothoe tenebricosa Moore, 1910, as Eunoe Macellicephala maculosa Treadwell, 1931 exoculata Treadwell, 1923 Eupanthalis mutilata (Treadwell), 1906, as Harmothoe trimaculata (Treadwell), 1924, as Polynoe mutilata Treadwell, 1906, and Eupan- Evarnella trimaculata Treadwell, 1924 thalis oahuensis Treadwell, 1906 Harmothoe variegata Treadwell, 1917 Euphrosine branchiata Treadwell, 1939, as Harmothoe villosa Treadwell, 1926 Euphrosyne branchiata Treadwell, 1939 Hermenia verruculosa Grube, 1856, as Polynoe Euphrosine longisetis Treadwell, 1939, as Euphro- nodosa Treadwell, 1901 syne longisetis Treadwell, 1939 Hermione hystrix (Savigny), 1818, as Melaenis Eupolyodontes elongata (Treadwell), 1931, as tropicus Treadwell, 1934 Iphionella elongata Treadwell, 1931 Hipponoe multibranchiata (Treadwell), 1940, as Eupomatus dianthus decorus Treadwell, 1931, as Metamphinom#e multibranchiata Treadwell, Eupomnatus decorus Treadwell, 1931 1940 Eupomatus similis Treadwell, 1929 Hyalinoecia branchiata Treadwell, 1934 Eupomatus uncinatus Philippi, 1844, as Eupoma- Hydroides crucigera Morch, 1863, as Hydroides tus operculata Treadwell, 1929 californicus Treadwell, 1929 262 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Hydroides parvus (Treadwell), 1901, as Eupom- Lumbrineris candida (Treadwell), 1921, as Lum- atus parvus Treadwell, 1901 brinereis candida Treadwell, 1921 Hypsicomus californicus (Treadwell), 1906, as Lumbrineris grandis (Treadwell), 1906, as Lum- Potamilla californica Treadwell, 1906 briconereis grandis Treadwell, 1906 Hypsicomus torquatus (Grube), 1878, as Sabella Lumbrineris inflata Moore, 1911, as Lumbrinereis alba Treadwell, 1917, Hypsicomus purpureus cervicalis Treadwell, 1922, and questionably Treadwell, 1924, and questionably Parasabella Lumbrinereis cingulata Treadwell, 1917 sulfurea Treadwell, 1917 Lumbrineeris, near latreilli Audouin and Milne- Iospilus phalacroides Viguier, 1886, as Phalacro- Edwards, 1834, as Lumbrinereis pauscidentata phorus niger Treadwell, 1943 Treadwell, 1921 Laeonerei-s culveri (Webster), 1879, as Nereis Lumbrineris minuscula Moore, 1911, as Lumbri- (Leptonereis) acuta Treadwell, 1923, and conereis minuta Treadwell, 1906 Leptonereis nota Treadwell, 1941 Lumbrineris nuchalis (Treadwell), 1921, as Laetmonice nitida Treadwell, 1926 Lumbrinereis nuchalis Treadwell, 1921 Lanice expansa Treadwell, 1906 Lumbrineris parvapedata (Treadwell), 1901, as Laonice cirrata (Sars), 1851, as Aricidea alata Lumbriconereis parvapedata Treadwell, 1901, Treadwell, 1901 and Lumbrinereis elongata Treadwell, 1931 Leanira grubei (Treadwell), 1901, as Sthenelais Lumbrineris similabris Treadwell, 1926, as grubei Treadwell, 1901 Lumbrinereis similabris Treadwell, 1926 Leanira hystricis Ehlers, 1874, as Eupholoe Lumbrineris treadwelli, new name for Lumbri- cirrata Treadwell, 1934 conereis maculata Treadwell, 1901, preoccupied fLeocrates oculatuss (Treadwell), 1906, as Castalia Lumbrineris zonata (Johnson), 1901, as Lumbri- oculata Treadwell, 1906 nereis singularisetis Treadwell, 1931 Lepidametria commensalis Webster, 1879, as Lycastis kartaboensis (Treadwell), 1926, as Lepidasthenia lactea Treadwell, 1939 Namonereis kartaboensis Treadwell, 1926 Lepidasthenia alba (Treadwell), 1906, as Polynoi Lygdamis asteriformis (Augener), 1906, as alba Treadwell, 1906, and Polynoe lucida Hermella varians Treadwell, 1901 Treadwell, 1906 Lygdamis philippinensis (Treadwell), 1926, as Lepidasthenia interrupta (Marenzeller), 1902, as Monorchos philippinensis Treadwell, 1926 Lepidasthenia occelata Treadwell, 1936 Lysarete brasiliensis Kinberg, 1856, as Oenone Lepidasthenia natans (Chamberlin), 1919, as brevimaxillata Treadwell, 1931 Lepidasthenia longicirrata Treadwell, 1928 Lysidice collaris Grube, 1870, as Lysidice sulcata Lepidasthenia rufa Treadwell, 1928, as Lepidas- Treadwell, 1901, Lysidice parva Treadwell, thenia variegata Treadwell, 1928 1922, and questionably Lysidice fusca Tread- Lepidasthenia varius Treadwell, 1917, as Lepidas- well, 1922 thenia picta Treadwell, 1928 Lysidice tortugae Treadwell, 1921 Lepidasthenia virens (Blanchard), 1849, as Maldane cristata Treadwell, 1923 Lepidasthenia ornata Treadwell, 1937 Maldane philippinensis Treadwell, 1931 Lepidasthenia sp., juvenile, as Lepidonotus in. Marphysa aransensis Treadwell, 1939 quilinus Treadwell, 1917 Marphysa bellii oculata Treadwell, 1921 Lepidonotus caelorus Moore, 1903, as Polynoe- Marphysa brevitentaculata Treadwell, 1921 spicula Treadwell, 1906, and Lepidonotss Marphysa minima (Hansen), 1882, as Marphysa minutus Treadwell, 1936 languida Treadwell, 1921 Lepidonotus sublevis Verrill, 1873, as Lepidonotus Marpliysa mossambica (Peters), 1854, as pallidus Treadwell, 1939 Marphysa simplex Treadwell, 1922 Leptonereis egregicirrata Treadwell, 1924, ques- Marphysa orientalis Treadwell, 1936 tionable Marphysa regalis Verrill, 1900, as Marphysa Loimia medusa fragilis Treadwell, 1911 (Savigny), 1820, as Losmia Marphysa sanguinea (Montagu), 1815, as minuta Treadwell, 1929 Marphysa acicularum brevibranchiata Tread- Lopadorhynchus varius Treadwell, 1943 well, 1921, Marphysa nobilis Treadwell, 1917, Lumbrineris bicirrata (Treadwell), 1929, as and Marphysa viridis Treadwell, 1917 Lumbrinereis bicirrata Treadwell, 1929 Megalomma splendida (Moore), 1905, as Branchi- Lumbrineris bilabiata (Treadwell), 1901, as ommna disparoculatum Treadwell, 1914 Lumbriconereis bilabiata Treadwell, 1901 Mfelinnexis tentaculata (Treadwell), 1906, as Lumbrineris branchiata (Treadwell), 1921, as Terebellides tentaculata Treadwell, 1906 Lumbrinereis branchiata Treadwell, 1921 Myriana cirrata Treadwell, 1931 1956 HARTMAN: ANNELIDS 263 Mysta ornata (Grube), 1878, as Mysta macltlata Pareulepis fimbriata (Treadwell), 1901, as Eulepis Treadwell, 1920 fimbriata Treadwell, 1901 Naiades cantraint&i delle Chiaje, 1828, as Alciopa Paresdepis wyvillei (McIntosh), 1885, as Eulepis distorta Treadwell, 1943, and perhaps Alciopa splendida Treadwell, 1901 mutilata Treadwell, 1934 ?Pareurythoe elongata (Treadwell), 1931, as Naineris mnutilata Treadwell, 1931, as Nainereis Hipponoe elongata Treadwell, 1931 mutilata Treadwell, 1931 Perinereis aibuhitensis Grube, 1878, as Nereis Naineris setosa (Verrill), 1900, as Anthostoma (Neanthes) linea Treadwell, 1936, and Nereis latacapitata Treadwell, 1901 (Neanthes) orientalis Treadwell, 1936 Areanthes succinea (Frey and Leuckart), 1847, Perinereis brevicirrata (Treadwell), 1920, as as Nereis (Neanthes) australis Treadwell, 1923 Nereis brevicirrata Treadwell, 1920 .Nereimyra mutilata (Treadwell), 1901, as Perinereis heUleri Grube, 1878, as Neanthes Castalia mutilata Treadwell, 1901 obscura Treadwell, 1928 Nereis amoyensis (Treadwell), 1936, as Nereis Perinereis monterea (Chamberlin), 1918, as (Neanthes) amoyensis Treadwell, 1936 Nereis spinifera Treadwell, 1929 Nereis arroyensis Treadwell, 1901 Phalacrophorus pictus Greeff, 1879, as Phala- Nereis heterocirrata Treadwell, 1931 crophorus maculatus Treadwell, 1943 Nereis (Leptonereis) distorta Treadwell, 1936, Phalacrophorus uniformis Reibisch, 1895, as questionable Phalacrophorus attenuatus Treadwell, 1943 Nereis petagica largoensis Treadwell, 1931, as Phalacrostemma setosa (Treadwell), 1906, as Nereis brevicirrata Treadwell, 1929, and Nereis Sabellaria setosa Treadwell, 1906 largoensis Treadwell, 1931 Pherecardia striata (Kinberg), 1857, as Hermo- Nereis -riisei Grube, 1856, as Nereis ambiguus dice pennata Treadwell, 1901 Treadwell, 1937, Nereis decora Treadwell, Pherecardia sp., as Hermodice pennata tutuilensis 1932, Nereis (Neanthes) paucidentata Tread- Treadwell, 1926 well, 1939, and Nereis (Neanthes) varia Tread- Pherusa dubia (Treadwell), 1929, as Stylarioides well, 1941 dubius Treadwell, 1929 Nerine alaskensis (Treadwell), 1914, as Scoleco- Pherusa infiata (Treadwell), 1914, as Trophonia lepis alaskensis Treadwell, 1914 inflata Treadwell, 1914, and Trophonia minuta Nerine cirratulus hirsuta (Treadwell), 1928, as Treadwell, 1914 Spio hirsuta Treadwell, 1928 Plhragmatopoma lapidosa Kinberg, 1867, as Nerinides acuta (Treadwell), 1914, as Spio acuta Centrocorone spinifera Treadwell, 1939 Treadwell, 1914 Phyllochaetopterus verrillii Treadwell, 1943 Nerinides agilis (Verrill), 1873, as Nerine Phyllodoce magna-oculata Treadwell, 1901 minuta Treadwell, 1939 Phyllodoce nicoyensis Treadwell, 1928 Nicomache antillensis Augener, 1922, as Nico- Phyllodoce stigmata Treadwell, 1925 mache antiguensis Treadwell, 1924 PhyUodoce tortugae Treadwell, 1917 Nicon mexicana (Treadwell), 1942, as Lepto- Phyllodoce violacea Treadwell, 1926, as PhyUo- nereis mexicana Treadwell, 1942 doce variegata Treadwell, 1926 Nothria stigmatis (Treadwell), 1922, as Onuphis Pionosyllis manca Treadwell, 1931, generic status stigmatis Treadwell, 1922 questionable Notocirrus attenuatus (Treadwell), 1906, as Pista maculata (Dalyell), 1853, as Pista groen- Arabella attentuata Treadwell, 1906 landica Treadwell, 1937 Odontosyllis enopla Verrill, 1900, as Autolytus Platynereis bicanaliculata (Baird), 1863, as bidens Treadwell, 1941 Nereis notomacula Treadwell, 1914 Odontosyllis octodentata Treadwell, 1917 Platynereis dumerilii (Audouin and Milne- Onuphis branchiata Treadwell, 1931 Edwards), 1834, as Eunereis africana Tread- Ophelia magna (Treadwell), 1914, as Ophelina well, 1943, Leptonereis maculata Treadwell, magna Treadwell, 1914 1928, Uncinereis lutea Treadwell, 1928, and Palola siciliensis (Grube), 1840, as Leodice Uncinereis trimaculosa Treadwell, 1940 viridis vernalis Treadwell, 1922 Platynereis polyscalma Chamberlin, 1919, as Panthalis evanida (Treadwell), 1926, as Eupan- Nereis (Platynereis) integer Treadwell, 1920 thalis evanida Treadwell, 1926 Polymniella autantiaca Verrill, 1900, as TerebeUa Panthalis pacifica Treadwell, 1914 hiata Treadwell, 1931 Panthalis pustulata Treadwell, 1924, as Acoetes Polynoe gracilis (Verrill), 1874, as Enipo cirrata magnifica Treadwell, 1929 Treadwell, 1925 Pa-rama.rphysa teres Treadwell, 1922 Polynoella brunnea Treadwell, 1943 264 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Polyodontes oculea (Treadwell), 1901, as Pantha- Sphaerosyllis arenaceus Claparede, 1863, as lis oculea Treadwell, 1901 Oophylas minuta Treadwell, 1937 Polyodontes panamensis (Chamberlin), 1919, as Spirographis braziliensis Treadwell, 1932 Polyodontes californicus Treadwell, 1941 Streblosoma crrassibranchia Treadwell, 1914, as Polyophthalmus pictus (Dujardin), 1839, as Streblosoma crassibranchiata Treadwell, 1914 Polyophthalmus incertus Treadwell, 1936, and Syllis complamata Treadwell, 1901 Polyophthalmus papilkatus Treadwell, 1943 Terebella parvabranchiata Treadwell, 1906 Pomatoceros caeruleus (Schmarda), 1861, as Tharyx multifilis Moore, 1909, as Cirratulus in- Pomatoceros davaoensis Treadwell, 1942 hamatus Treadwell, 1937 Potamethus elongatus (Treadwell), 1906, as Thelepus branchiatus Treadwell, 1906 Potamila elongata Treadwell, 1906 Thelepus cincinnatus (Fabricius), 1780, as The- Potamilla minuta (Treadwell), 1941, as Parasa- lepus crassibranchiatus Treadwell, 1901 bella minuta Treadwell, 1941 Thelepus crispus Johnson, 1901, as Streblosoma Prionospio treadwelli Hartman, 1951, as Priono- magna Treadwell, 1937 spio plumosa Treadwell, 1931 Thelepus haitiensis Treadwell, 1931 Psammolyce arenosa (delle Chiaje), 1841, as Thelepus setosus (Quatrefages), 1865, as Streb- Eupholoi nuda Treadwell, 1936 losoma verritli Treadwell, 1911 Psammolyce flava Kinberg, 1855, as Eupholoe Thornora jukesii Baird, 1865, as Lepidonotus acuminata Treadwell, 1934 atratus Treadwell, 1940 Pseudeurythoe oculata (Treadwell), 1941, as Tomopteris nisseni Rosa, 1908, as Tomopteris Eurythoe oculata Treadwell, 1941 longisetis Treadwell, 1936, and Tomopteris Pseudonereis palpata (Treadwell), 1923, as Ne- opaca Treadwell, 1928 reis (Neanthes) palpata Treadwell, 1923, and Tomopteris tentaculata Treadwell, 1928 Nereis disparsetosa Treadwell, 1932 Torea fasciata Treadwell, 1943 Pseudonereis variegata (Grube), 1856, as Peri- Travisiopsis benhami Monro, 1936, as Plotobia nereis diversidentata Treadwell, 1943 paucichaeta Treadwell, 1943 Pseudopotamilla occelata Moore, 1905, as Lao- Travisiopsis coniceps (Chamberlin), 1919, as nome oculifera Treadwell, 1914 Nuchubranchia palmata Treadwell, 1928 Rhynchonerella angelini (Kinberg), 1866, as Travisiopsis lanceolata Southern, 1911, as Travi- Eulalia nagnapupula Treadwell, 1941 siopsis atlantica Treadwell, 1936 Sabellastarte indica Savigny, 1820, as Laonome Trypanosyllis adamanteus Treadwell, 1914 punctata Treadwell, 1906, and Laonome arenosa Typosyllis aciculata Treadwell, 1945 Treadwell, 1943 Typosyllis crassicirrata Treadwell, 1925 Sabellastarte magnifica (Shaw), 1800, as Meta- Vanadis crystallina Greeff, 1876, as Vanadis col- laonome brunnea Treadwell, 1917, and Laonome lata Treadwell, 1928 sanjuanensis Treadwell, 1941 Vanadis formosa Claparede, 1870, as Vanadis Sclerostyla ctenactis M6rch, 1863, as Placostegus fusca-punctata Treadwell, 1906 calciferus Treadwell, 1929 Vanadis minuta Treadwell, 1906 Scoloplos treadwelli Eisig, 1914, as Aricia cirrata Vanadis uncinata Treadwell, 1943 Treadwell, 1901 Vermiliopsis bermudeissis (Bush), 1904, as Ver- Semiodera cariboum (Grube), 1856, as Semiodera milia glandulata Treadwell, 1936 glabra Treadwell, 1928 Vermiliopsis multiannulata (Moore), 1923, as .Semiodera glabra (Treadwell), 1901, as Styla- Vermiliopsis hawaiiensis Treadwell, 1943, and roides glabra Treadwell, 1901 Vermiliopsis torquata Treadwell, 1943 Sphaeropomatus miamiensis Treadwell, 1934 LIST OF SPECIES BY FAMILIES APHRODITIDAE Lepidasthenia natans (Chamberlin), 1919, as Aplirodita magna Treadwell, 1925 Lepidasthenia longicirrata Treadwell, 1928 Hermione hystrix (Savigny), 1818, as Melaenis Lepidasthenia rufa Treadwell, 1928, as Lepidas- tropicus Treadwell, 1934 thenia variegata Treadwell, 1928 Laetmonice nitida Treadwell, 1926 Lepidasthenia varius Treadwell, 1917, as Lepi- dasthenia picta Treadwell, 1928 POLYNOIDAE Lepidasthenia vsirens (Blanchard), 1849, as Lepi- Acholoe astericola (delle Chiaje), 1841, as Acho- dastlienia ornata Treadwell, 1937 loe orbiculata Treadwell, 1921 Lepidasthenia sp., juvenile, as Lepidonotus inqui- Arctonoe tuberculata (Treadwell), 1906, as Har- linus Treadwell, 1917 mothoe tuberculata Treadwell, 1906 Lepidonotus caelorus Moore, 1903, as Polynoe Arctonoe vittata (Grube), 1855, as Lepidasthe- spicula Treadwell, 1906, and Lepidonotus nsi- nia elegans Treadwell, 1941 nutus Treadwell, 1936 Chaetacanthus magnificus (Grube), 1875, as Lepidonotus ssublevis Verrill, 1873, as Lepidono- Polynoe branchiata Treadwell, 1901, and Lepi- tus pallidus Treadwell, 1939 donotus pilosus Treadwell, 1937 Polynoe gracilis (Verrill), 1874, as Enipo cirrata Drieschia pellucida Moore, 1903, as Drieschia Treadwell, 1925 atlantica Treadwell, 1936 Polynoella brunnea Treadwell, 1943 Eunoe crassa (Treadwell), 1924, as Lag'sca Thormora jukes'i Baird, 1865, as Lepidonotus crassa Treadwell, 1924 atratus Treadwell, 1940 Eunoe purpurea Treadwell, 1936, as Acanthico- lepis longicirrata Treadwell, 1941 POLYODONTIDAE Gattyana cirrosa (Pallas), 1776, as Gattyana Eupanthalis inaculosa (Treadwell), 1931, as imbricata Treadwell, 1926 Macellicephala maculosa Treadwell, 1931 Halosydna latior Chamberlin, 1919, as Halosydna Eupanthalis mutilata (Treadwell), 1906, as Poly- obtusa-cirrata Treadwell, 1937 noe mutilata Treadwell, 1906, and Eupanthalis HalosydneUa fusca-maculata (Treadwell), 1924, oahuensis Treadwell, 1906 as Halosydna fusca-maculata Treadwell, 1924, Eupolyodontes elongata (Treadwell), 1931, as and Halosydna fuscomarginata Treadwell, 1924 Iphionella elongata Treadwell, 1931 Halosydnella grisea (Treadwell), 1929, as Halo- Panthalis evanida (Treadwell), 1926, as Eupan- sydna grisea Treadwell, 1929 thalis evanida Treadwell, 1926 Halosydnella oculata (Treadwell), 1926, as Halo- Panthalis pacifica Treadwell, 1914 sydna oculata Treadwell, 1926 Panthalis pustulata Treadwell, 1924, as Acoetes Harmothoe crucis (Grube), 1856, as Harmothoe magnifica Treadwell, 1929 lanceocirrata Treadwell, 1928, and Harmo- Polyodontes oculea (Treadwell), 1901, as Pan- thoe lanceolata Treadwell, 1928 thalis oculea Treadwell, 1901 Harmothoe imbricata (Linnaeus), 1767, as Har- Polyodontes panamensis (Chamberlin), 1919, as motlhoe levis Treadwell, 1937 Polyodontes californicus Treadwell, 1941 Harmothoe sylliformia Treadwell, 1928 Harmothoe tenebricosa Moore, 1910, as Eunoe SIGALIONIDAE exoculata Treadwell, 1923 Leanira grubei (Treadwell), 1901, as Sthenelais Harmothoe trimaculata (Treadwell), 1924, as grubei Treadwell, 1901 EvarneUa trimaculata Treadwell, 1924 Leanira hysticis Ehlers, 1874, as Eupholoe cirrata Harmothoe variegata Treadwell, 1917 Treadwell, 1934 Harmothoe villosa Treadwell, 1926 Psammolyce arenosa (delle Chiaje), 1841, as Hermenia verruculosa Grube, 1856, as Polynoe Eupholoi nuda Treadwell, 1936 nodosa Treadwell, 1901 Psammolyce flava Kinberg, 1855, as Eupholog Lepidametria commensalis Webster, 1879, as acuminata Treadwell, 1934 Lepidasthenia lactea Treadwell, 1939 Lepidasthenia alba (Treadwell), 1906, as Poly- PAREULEPIDAE noe alba Treadwell, 1906, and Polynoe lucida Pareulepis fimbriata (Treadwell), 1901, as Treadwell, 1906 Eulepis fimbriata Treadwell, 1901 Lepidasthenia interrupta (Marenzeller), 1902, as Pareulepis wyviUei (McIntosh), 1885, as Eulepis Lepidasthenia occelata Treadwell, 1936 splendida Treadwell, 1901 265 266 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 AMPHINOMIDAE Vanadis crystallina Greeff, 1876, as Vanadis col- tBenthoscolex microcarunc4lata (Treadwell), lata Treadwell, 1928 1901, as Amphinome microcarunculata Tread- Vanadis formosa Claparede, 1870, as Vanadis well, 1901 fusca-punctata Treadwell, 1906 Hipponoe multibranchiata (Treadwell), 1940, as Vanadis minuta Treadwell, 1906 Metamphinome multibranchiata Treadwell, Vanadis uncinata Treadwell, 1943 1940 ?Pareurythog elongata (Treadwell), 1931, as LOPADORHYNCHIDAE Hipponoe elongata Treadwell, 1931 Halyplanes gracilis Reibisch, 1893, as Haliplanella Pseudeurythoe oculata (Treadwell), 1941, as pacifica Treadwell, 1943 Eurythoe oculata Treadwell, 1941 Lopadorhynchus varius Treadwell, 1943 Pherecardia striata (Kinberg), 1857, as Hermo- dice pennata Treadwell, 1901 IOSPILIDAE Pherecardia sp., as Hermodice pennata tutuilen- Iospilus phalacroides Viguier, 1886, as Phalacro- sis Treadwell, 1926 phorus niger Treadwell, 1943 Phalacrophorus pictus Greeff, 1879, as Phalacro- EUPHROSINIDAE phorus maculatus Treadwell, 1943 Euphrosine branchiata Treadwell, 1939, as Eu- Phalacrophorus uniformis Reibisch, 1895, as phrosyne branchiata Treadwell, 1939 Phalacrophorus attenuatus Treadwell, 1943 Euphrosine longisetis Treadwell, 1939, as Euphro- syne longisetis Treadwell, 1939 TYPHLOSCOLECIDAE Travisiopsis benhami Monro, 1936, as Plotobia PHYLLODOCIDAE paucichaeta Treadwell, 1943 Travisiopsis coniceps (Chamberlin), 1919, as Anaitides madeirensis Langerhans, 1879, as An- Nuchubranchia palmata Treadwell, 1928 aitides minuta Treadwell, 1937, Mystides graci- Travisiopsis lanceolata Southern, 1911, as Travisi- his Treadwell, 1941, and PhyUodoce varia opsis atlantica Treadwell, Treadwell, 1917 1936 Anaitides panamensis (Treadwell), 1917, as TOMOPTERIDAE Phyllodoce panamensis Treadwell, 1917 Tomopteris nisseni Rosa, 1908, as Tomopteris Anaitides tenuissitma (Grube), 1878, as Phyllo- opaca Treadwell, 1928, and Tomopteris tongi- doce fusca-cirrata Treadwell, 1926 setis Treadwell, 1936 Eteone nlonga (Fabricius), 1870, as Eteone Tomopteris tentaculata tuberculata Treadwell, 1922 Treadwell, 1928 Eteone pacifica Hartman, 1936, as Eteone macu- HESIONIDAE lata Treadwell, 1922 ?Dalhousiella longicirrata (Treadwell), 1901, as Eulalia foliosa Treadwell, 1924 Castalia longicirrata Treadwell, 1901 Eulalia myriacyclum (Schmarda), 1861, as Mysta ornata (Grube), 1878, as Mysta mxculata Eulalia quinquelineata Treadwell, 1901 Treadwell, 1920 PhyUodoce magna-oculata Treadwell, 1901 .Leocrates oculata (Treadwell), 1906, as Castalia Phyllodoce nicoyensis Treadwell, 1928 oculata Treadwell, 1906 Phyllodoce pulla Treadwell, 1926 ?Nereimyra mutilata (Treadwell), 1901, as Cas- Phyllodoce stigmata Treadwell, 1925 talia mutilata Treadwell, Phyllodoce tortugae Treadwell, 1917 1901 Phyllodoce violacea Treadwell, 1926, as PhyUo- PILARGIIDAE doce variegata Treadwell, 1926 AncistrosyUis tentaculata Treadwell, 1941 Epitoka pelagica Treadwell, 1943, questionably ALCIOPIDAE pilargiid or syllid Callizotnella pigmenta Treadwell, 1943, perhaps Krohnia sp. SYLLIDAE Corynocephalus magnachaetus Treadwell, 1943 Autolytus pacificus Treadwell, 1943 Naiades cantrainis delle Chiaje, 1828, as Alciopa Autolytus varius Treadwell, 1914 distorta Treadwell, 1943, and perhaps Alciopa Haplosyllis spongicola (Grube), 1855, as Haplo- mutilata Treadwell, 1934 syllis gula Treadwell, 1924 Rhynchonerella angelini (Kinberg), 1866, as Myriana cirrata Treadwell, 1931 Eulalia magnapupula Treadwell, 1941 Odontosyllis enopla Verrill, 1900, as Autolytus 2lorea fasciata Treadwell, 1943 bidens Treadwell, 1941 1956 HARTMAN: ANNELIDS 267 OdontosyUis octodentata Treadwell, 1917 Edwards), 1834, as Uncinereis lutea Treadwell, Pionosyllis manca Treadwell, 1931, questionable 1928, Leptonereis maculata Treadwell, 1928, Sphaerosyllis arenaceus Claparede, 1863, as Uncinereis trimnaculosa Treadwell, 1940, and Oophylax minuta Treadwell, 1937 Eunereis africana Treadwell, 1943 Syllis complanata Treadwell, 1901 Platynereis polyscalma Chamberlin, 1919, as Trypanosyllis adamanteus Treadwell, 1914 Nereis (Platynereis) integer Treadwell, 1920 Typosyllis aciculata Treadwell, 1945 Pseudonereis palpata (Treadwell), 1923, as Typosyllis crassicirrata Treadwell, 1925 Nereis (Neanthes) palpata Treadwell, 1923, and Nereis disparsetosa Treadwell, 1932 NEREIDAE Pseudonereis variegata (Grube), 1856, as Peri- Ceratonerets anchylochaeta (Horst), 1924, as nereis diversidentata Treadwell, 1943 Ceratonereis longicauda Treadwell, 1943 Ceratonereis hircinicola (Eisig), 1870, as Nereis GLYCERIDAE (Cerantonereis) bartletti Treadwell, 1937 Glycera ?capitata Oersted, 1843, as Hemipodia Ceratonereis mirabilis Kinberg, 1866, as Cerato- canadensis Treadwell, 1937 nereis singularis Treadwell, 1929, and perhaps Glycera tesselata Grube, 1863, as Glycera abran- Nereis singularis Treadwell, 1943 chiata Treadwell, 1901, and Glycera spadix Ceratonereis paucidentata (Moore), 1903, as Treadwell, 1943 Nereis (Ceratonereis) alaskensis Treadwell, 1921 GONIODIDAE Laeonereis culveri (Webster), 1879, as Nereis Goniada brunnee Treadwell, 1906 (Leptonereis) acuta Treadwell, 1923, and Lep- Goniada oculata Treadwell, 1901, questionably tonereis nota Treadwell, 1941 Glycinde sp. Leptonereis egregicirrata Treadwell, 1924, ques- Goniada quinquelabiata Augener, 1906, as Goni- tionable ada magna Treadwell, 1945 Lycastis kartaboensis (Treadwell), 1926, as Na- Goniada teres Treadwell, 1931 mxonereis kartaboensis Treadwell, 1926 Neanthes succinea (Frey and Leuckart), 1847, ONUPHIDAE as Nereis (Neanthes) australis Treadwell, 1923 Hyalinoecia branchiata Treadwell, 1934 Nereis amoyensis (Treadwell), 1936, as Nereis Nothnia stigmatis (Treadwell), 1922, as Onuphis (Neanthes) amoyensis Treadwell, 1936 stigmatis Treadwell, 1922 Nereis arroyensis Treadwell, 1901 Onuphsis branchiata Treadwell, 1931 Nereis heterocirrata Treadwell, 1931 Nereis (Leptonereis) distorta Treadwell, 1936, EUNICIDAE questionable Eunice afra Peters 1854, as Leodice aciculata Nereis pelagica largoensis Treadwell, 1931, as Treadwell, 1922, Leodice bucciensis Treadwell, Nereis brevicirrata Treadwell, 1929, and Nereis 1921, Leodice flava-punctata Treadwell, 1922, largoensis Treadwell, 1931 Leodice guanica Treadwell, 1921, Leodice suvi- Nereis riisei Grube, 1856, as Nereis ambiguus ensis Treadwell, 1922, and questionably Leodice Treadwell, 1937, Nereis decora Treadwell, 1932, notata Treadwell, 1921 Nereis (Neanthes) paucidentata Treadwell, Eunice antennata Savigny, 1818, as Eunice inter- 1939, and Nereis (Neanthes) varia Treadwell, rupta Treadwell, 1906 1941 Eunice argentinensis (Treadwell), 1929, as Leo- Nicon inexcicana (Treadwell), 1942, as Lepto- dice argentinensis Treadwell, 1929 nereis mexicana Treadwell, 1942 Eunice armillata (Treadwell), 1922, as Leodice Perinereis aibuhitensis Grube, 1878, as Nereis armillata Treadwell, 1922, and Leodice gracili- (Neanthes) linea Treadwell, 1936, and Nereis cirrata Treadwell, 1922 (Neanthes) orientalis Treadwell, 1936 Eunice auriculate Treadwell, 1900 and 1901 Perinereis brevicirrata (Treadwell), 1920, as Eunice bilobata Treadwell, 1906 Nereis brevicirrata Treadwell, 1920 Eunice crassi-tentaculata (Treadwell), 1922, as Perinereis helleri Grube, 1878, as Neanthes ob- Leodice crassi-tentaculata Treadwell, 1922 scura Treadwell, 1928 Eunice filamentosa Grube, 1856, as Leodice spons- Perinereis monterea (Chamberlin), 1918, as gicola Treadwell, 1921 Nereis spinifera Treadwell, 1929 Eunice grubei Gravier, 1900, as Leodice biformi- Platynereis bicanaliculata (Baird), 1863, as cirrata Treadwell, 1922 Nereis notomacula Treadwell, 1914 Eunice hawaiiensis Treadwell, 1906 Platynereis dumerilli (Audouin and Milne- Eunice nicidioformis Treadwell, 1906 268 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109

Eunice (Nicidion) cariboea Grube, 1856, as brinereis nuchalis Treadwell, 1921 Eunice culebra Treadwell, 1901 Lumbrineris parvapedata (Treadwell), 1901, as E-unice (Nicidion) gracilis Crossland, 1903, as Lumbriconereis parvapedata Treadwell, 1901, Nicidion fusca-fasciata Treadwell, 1922 and Lumbrinereis elongata Treadwell, 1931 E-unice norvegica (Linnaeus), 1767, as Leodice Lumbrineris similabris (Treadwell), 1926, as arcturi Treadwell, 1928 Lumbrinereis similabris Treadwell, 1926 Eunice savignyi (Grube), 1878, as Leodice langi Lumbrineris treadwelli, new name, for Lumbri- Treadwell, 1943 conereis maculata Treadwell, 1901 Eunice tenuis (Treadwell), 1921, as Leodice Lumbrineris zonata (Johnson), 1901, as Lumbri- tenuis Treadwell, 1921 nereis singularisetis Treadwell, 1931 Eunice tubifex Crossland, 1904, as Leodice tubi- cola Treadwell, 1922 ARABELLIDAE Eunice vittata (delle Chiaje), 1828, as Leodice Arabella iricolor (Montagu), 1804, as ArabeUa rubrivittata Treadwell, 1921 dubia Treadwell, 1922, and questionably Dri- Lysidice collaris Grube, 1870, as Lysidice sulcata lonereis pinnata Treadwell, 1921 and Arabella Treadwell, 1901, Lysidice parva Treadwell, setosa Treadwell 1922, and questionably Lysidice fusca Tread- Arabella iridescens Treadwell, 1906 well, 1922 Arabella semimaculata (Moore), 1911, as Ara- Lysidice tortugae Treadwell, 1921 bella pacifica Treadwell, 1941 Marphysa aransenis Treadwell, 1939 Arabella sp., as Aracoda attenuata Treadwell, Marphysa bellii oculata Treadwell, 1921 1911 Marphysa brevitentaculata Treadwell, 1921 Drilonereis lumbricus Treadwell, 1922 Marphysa minima (Hansen), 1882, as Marphysa Drilonereis planiceps (Grube), 1878, as Drilo- languida Treadwell, 1921 nereis paucidentata Treadwell, 1922 Marphysa mossambica (Peters), 1854, as Mar- Drilonereis spatula (Treadwell), 1911, as Ara- physa simplex Treadwell, 1922 coda spatula Treadwell, 1911, and Drilonereis Marphysa orientalis Treadwell, 1936 brunnea Treadwell, 1921 Marphysa regalis Verrill, 1900, as Marphysa fra- Drilonereis sp., juvenile, as Drilonereis simi;is gilis Treadwell, 1911 Treadwell, 1921 Marphysa sanguinea (Montagu), 1815, as Mar- Notocirrus attenuatus (Treadwell), 1906, as Ara- physa acicularum brevibranchiata Treadwell, beUa attenuata Treadwell, 1906 1921, Marphysa nobilis Treadwell, 1917, and Marphysa viridis Treadwell, 1917 LYSARETIDAE Palola siciliensis (Grube), 1840, as Leodice yin- Lysarete brasiliensis Kinberg, 1865, as Oenone dis vernalis Treadwell, 1922 brevimaxillata Treadwell, 1931 Paramarphysa teres Treadwell, 1922 ORBINIIDAE LUMBRINERIDAE Haploscoloplos robustus (Verrill), 1873, as Lumbrineris bicirrata (Treadwell), 1929, as Scoloplos rufa Treadwell, 1941 Lumbrinereis bicirrata Treadwell, 1929 Naineris mutilata Treadwell, 1931, as Nainereis Lumbrineris bilabiata (Treadwell), 1901, as mutilata Treadwell, 1931 Lumbriconereis bilabiata Treadwell, 1901 Naineris setosa (Verrill), 1900, as Anthostoma Lumbrineris branchiata (Treadwell), 1921, as latacapitata Treadwell, 1901 Lumbrinereis branchiata Treadwell, 1921 Scoloplos treadwelli Eisig, 1914, as Aricia cirrata Lumbrineris candida (Treadwell), 1921, as Treadwell, 1901 Lumbrinereis candida Treadwell, 1921 Lumbrineris grandis (Treadwell), 1906, as SPIONIDAE Lumbriconereis grandis Treadwell, 1906 Boccardia sp., as Polydora californica Treadwell, Lumbrineris inflata Moore, 1911, as Lumbrinereis 1914 cervicalis Treadwell, 1922, and questionably Laonice cirrata (Sars), 1851, as Aricidea alata Lumbrinereis cingulata Treadwell, 1917 Treadwell, 1901 Lumbrineris, near latreilli Audouin and Milne- Nerine alaskensis (Treadwell), 1914, as Scole- Edwards, 1834, as Lumbrinereis paucidentata colepis alaskensis Treadwell, 1914 Treadwell, 1921 Nerine cirratulus hirsuta (Treadwell), 1928, as Lumbrineris mtniuscul Moore, 1911, as Lumbri- Spio hirsuta Treadwell, 1928 conereis minuta Treadwell, 1906 Nerinides acuta (Treadwell), 1914, as Spio acuta Lumbrineris nuchalis (Treadwell), 1921, as Lum- Treadwell, 1914 1956 HARTMAN: ANNELIDS 269 Nerinides agilis (Verrill), 1873, as Nerine minuta Euclymene corallicola (Treadwell), 1929, as Treadwell, 1939 Maldanella coralticola Treadwell, 1929 Prionospio treadwelli Hartman, 1951, as Priono- Maldane cristata Treadwell, 1923 spio plumosa Treadwell, 1931 Maldane philippinensis Treadwell, 1931 Nicomache antillensis Augener, 1922, as Nico- CHAETOPTERIDAE mache antiguensis Treadwell, 1924 Phyllochaetopterus verriii Treadwell, 1943 CIRRATULIDAE SABELLARIIDAE Cirratulus elongatus Treadwell, 1901, questionable Lygdamis asteriformsis (Augener), 1905, as Cirriformia tchefooensis (Grube), 1877, as Cir- HermeUa varians Treadwell, 1901 ratulus branchiatus Treadwell, 1936 Lygdamis philippinensis (Treadwell), 1926, as Cirriformia filigera (delle Chiaje), 1841, as Monorchos philippinensis Treadwell, 1926 Audouinea oculata Treadwell, 1932, and Audo- Phalacrostemma setosa (Treadwell), 1906, as uinea pygidia Treadwell, 1936 Sabellaria setosa Treadwell, 1906 Cirriformia hawaiensis, new name for Audouinea Phragmatopoma lapidosa Kinberg, 1867, as branchiata Treadwell, 1943 Centrocorone spinifera Treadwell, 1939 Cirriformia punctata (Grube), 1859, as Cirratu- lus nigromaculata Treadwell, 1901 AMPHARETIDAE Cirriformia semicincta (Ehlers), 1905, as Audo- Ampharete arctica Malmgren, 1866, as Ampharete uinea maculata Treadwell, 1929 brevibranchiata Treadwell, 1926 Cirriformia tentaculata (Montagu), 1808, as Ampharete eupalea Chamberlin, 1920, as Am- Cirratulus multicirratus Treadwell, 1936, and pharete seribranchiata Treadwell, 1926 Cirratutus pallidus Treadwell, 1931 Asabellides sibirica (Wiren), 1883, as Neosabel- Tharyx multifilis Moore, 1909, as Cirratulus in- lides alaskensis Treadwell, 1943 hamatus Treadwell, 1937 Melinnexis tentaculata (Treadwell), 1906, as Terebellides tentaculeta Treadwell, 1906 OPHELIIDAE Armandia maculata (Webster), 1884, as Ammo- TEREBELLIDAE trypane bermudiensis Treadwell, 1936 Euzonus (Thoracophelia) mnucronata (Tread- Lanice expansa Treadwell, 1906 well), 1914, as Ophelia mucronata Treadwell, Loimia medusa (Savigny), 1820, as Loimia 1914 minuta Treadwell, 1929 Ophelia magna (Treadwell), 1914, as Ophelina Pista maculata (Dalyell), 1853, as Pista groen- magna Treadwell, 1914 landica Treadwell, 1937 Polyophthalmus pictus (Dujardin), 1839, as Polymniella aurantiaca Verrill, 1900, as TerebeUa Polyophthalmus incertus Treadwell, 1936, and hiata Treadwell, 1931 Polyophthalmus papillatus Treadwell, 1943 Streblosoma crassibranchia Treadwell, 1914, as Streblosoma crassibranchiata Treadwell, 1914 FLABELLIGERIDAE Terebella parvabranchiata Treadwell, 1906 Pherusa dubia (Treadwell), 1929, as Stylarioides Tlielepus branchiatus Treadwell, 1906 dubius Treadwell, 1929 Thelepus cincinnatus (Fabricius), 1780, as Thele- Pherusa inflata (Treadwell), 1914, as Trophonia pus crassibranchiatus Treadwell, 1901 inflata Treadwell, 1914, and Trophonia minuta Thelepus crispus Johnson, 1901, as Streblosoma Treadwell, 1914 magna Treadwell, 1937 Semiodera cariboum (Grube), 1856, as Semiodera Thelepus haitiensis Treadwell, 1931 glabra Treadwell, 1928 Thelepus setosus (Quatrefages), 1865, as Stre- PSemitodera glabra (Treadwell), 1901, as Sty- blosoma verrilli Treadwell, 1911 laroides glabra Treadwell, 1901 SABELLIDAE CAPITELLIDAE Brenchiomma nigromaculata (Baird), 1865, as Capitellides teres Treadwell, 1939 Dasychone loandensis Treadwell, 1943, Dasy- Dasybranchus rectus Treadwell, 1901, questiona- chone ponce Treadwell, 1901, and Dasychonop- ble sis arenosa Treadwell, 1924 MALDANIDAE Eurato punctata Treadwell, 1926, questionable Asychis fimbriata (Treadwell), 1934, as Malda- Fabricia sabeUa (Ehrenberg), 1836, as Haplo- nella fimbriata Treadwell, 1934 branchus atlanticus Treadwell, 1932 270 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109

Hypsicomus californicus (Treadwell), 1906, as SERPULIDAE Potamilla californica Treadwell, 1906 Eupomatus dianthus (Verrill), 1873, decorus Hypsicomus torquatus (Grube), 1878, as Sabella Treadwell, 1931, as Eupomatus decorus Tread- alba Treadwell, 1917, HyPsicomus purpureus well, 1931 Treadwell, 1924, and questionably Parasabella Eupomatus similis Treadwell, 1929 sulfurea Treadwell, 1917 Eupomatus uncinatus Philippi, 1844, as Eupoma- Megalomma splendida (Moore), 1905, as tus operculata Treadwell, 1929 Branchiomma disparoculatum Treadwell, 1914 Hydroides crucigera Morch, 1863, as Hydroides Potamethus elontgatus (Treadwell ), 1906, as californicus Treadwell, 1929 Potamilla elongata Treadwell, 1906 Hydroides parvus (Treadwell), 1901, as Eupo- Potamilla minuta (Treadwell), 1941, as Parasa- matus parvus Treadwell, 1901 bella minuta Treadwell, 1941 Pomatoceros caeruleus (Schmarda), 1861, as Pseudopotamilla occelata Moore, 1905, as Pomatoceros davaoensis Treadwell, 1942 Laomome oculifera Treadwell, 1914 Sclerostyla ctenactis M6rch, 1863, as Placostegus Sabellastarte indica Savigny, 1820, as Laonome calciferus Treadwell, 1929 punctata Treadwell, 1906, and Laonome arenosa Sphaeropomatus miamiensis Treadwell, 1934 Treadwell, 1943 Vermiliopsis bermudensis (Bush), 1904, as Ver- Sabellastarte magnifica (Shaw), 1800, as Meta- milia glandulata Treadwell, 1936 lonome brunnea Treadwell, 1917, and Laonome Vermiliopsis multiannulata (Moore), 1923, as sanjuanensis Treadwell, 1941 Vermiliopsis hazwaijensis Treadwell, 1943, and Spirographis braziliensis Treadwell, 1932 Vermiliopsis torquata Treadwell, 1943 SYSTEMATIC DISCUSSION POLYNOIDAE cida occurs in deep waters of the Gulf Stream. Acholoe astericola (delle Chiaje), 1841 Fauvel (1939, p. 30) regards it as the same as D. Polynoe astericola DELLE CHIAJE, 1841, p. 106. pelagica Michaelsen (1892) from Ceylon. Acholoe orbicuZata TREADz.L, 1921a, pp. 1-3, figs. 1-8. Lepidasthenia virens (Blanchard), 1849 Polynoi virens BLANC3ARD, 1849, p. 16, pl. 1, The holotype of Acholoe orbiculata comes fig. 2. from the Belgian Congo. It is a slender speci- Lepidasthenia ornata TREADWELL, 1937b, pp. men about 50 mm. long. Many pairs of elytra 145-147, figs. 12-15. cover the dorsal side of the body. Cirriferous parapodia have unique nail-headed processes The holotype of Lepidasthenia ornata comes shown by Treadwell (fig. 5). The specimen from Arena Bank, western Mexico, in 35 fath- agrees fully with Acholoe astericola (delle oms; it has entangled terebellid tentacles, indi- Chiale) which is fully diagnosed by Fauvel cating that it was associated with a host species. (1923a, p. 94, fig. 36). Augener (1918, p. 152) It is a typical representative of Lepidasthenia has recorded the species from western Africa. virens (Blanchard) known to be a commensal of Thelepus crispus Johnson. It is noteworthy Arctonoe vittata (Grube), 1855 that both the polynoid and its host species (as Polynoe vittata GRUBE, 1855, pp. 82-83. Streblosoma magna) are recorded in the same Lepidasthenia elegans TREADwEI, 1941c, pp. report as coming from the same station. Lepi- 19-20, figs. 4-7. dasthenia virens is more fully described (Hart- The holotype of Lepidasthenia elegans comes man., 1939b, pp. 46-47, pl. 8 as Lepidametria). from east of Cedros Island, western Mexico, in Lepidasthenia varius Treadwell, 1917 45 fathoms. It agrees fully with Arctonoe Lepidasthenia varius TREADwEI., 1917b, pp. vittata (Grube). Anteriormost notosetae are 259-260, figs. 11-16. delicately bidentate at the distal end. The speci- Lepidasthenia picta TEADwEix, 1928a, pp. 456- men has been preserved in alcohol; conse- 457, figs. 10-13; 1941C, pp. 18-19. quently the typical broad, dark, transverse bar The type of Lepidasthenia varius comes from on the dorsum of the anterior end is faded. Dry Tortugas, Florida, and that of Lepidas- Variations in color pattern are discussed else. thenia where (Hartman, 1948a, p. 11). The species is picta from the Galapagos Islands in 15 common throughout the northeast Pacific and feet. Parapodia are distally rounded and com- is a frequent commensal with mollusks, echino- pressed. Neurosetae are deeply incised at the tip derms, and other invrertebrates (Hartman, in all segments; the superiormost one or two 1939b, pp. in a fascicle are somewhat slenderer than sub- 29-30, pl. 3, figs. 33-37). acicular ones but otherwise similar. Setal fas- Drieschia pellucida Moore, 1903 cicles are fuller in anterior segments, with 15 to 20 setae in a Drieschia pellucida Moo=, 1903, pp. 794-798, bundle, and more spinous along pl. 55, figs. 1-2. their free length than those in badk where they Drieschia attantica TREADWEL, 1936b, pp. number about seven im a bundle. Treadwell 52-53, figs. 7-9. (1941c, p. 19) gave corrections and additions to the original account and recorded the species The holotype of Drieschia atlantica was taken more extensively from Port Parker, Costa Rica, off Bermuda in 1000 fathoms. It agrees fully in 132 to 4 fathoms. with D. pellucida Moore (1903), originally de- scribed from 70 miles off Massachusetts in the Lepidasthenia rufa Treadwell, 1928 Gulf Stream. In the parapodia the two or three Lepidasthenia rufa TREADw mL, 1928a, pp. 457- larger inferiormost setae resemble those of 458, figs. 14-17. species of Scalisetosus McIntosh, but the basal- Lepidasthenia variegata TREADwEIL, 1928a, pp. most scale is less pronounced. Drieschia pellu- 458-560, figs. 18-21. 271 272 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Types of both Lepidasthenia rufa and L. long, slender lobes; they have embedded acic- variegata originate from Puerto Rico; they are ula and one or two slender notosetae. Neuro- identical. The notopodium is a long, digitate, podia are much larger and have a prolonged and very slender lobe arising from the dorsal superior lobe. The supra-acicular fascicle has a midlength of the neuropodium; it is supported few slenderer setae with a long spinous region. by a single aciculum. The neuropodium has Subacicular setae are more numerous, coarser, setae of two kinds; supra-acicular ones are and gradually shorter at the lowermost end of longer and slenderer than subacicular ones and the fascicle. The presence of a few notosetae is have an entire tip. Subacicular setae are typical of Lepidametria commensalis Webster, abruptly thicker and have a distal end that is in- which is known to be present in southern Flor- cised, with the accessory tooth short and thick ida. Lepidametria Webster is hardly distin- at the base and shorter than the main tooth or guishable from Lepidasthenia Malmgren, except fang. The species is known only from Puerto for the presence of notosetae in the first, and Rico. their absence in the second, genus, a character that varies with the of the individuals. Lepidasthenia natans (Chamberlin), 1919 age Harmopsides natans CHAMBERLIN, 1919, pp. 48- Thormora jukesii Baird, 1865 51, p1. 6, figs. 1-5. Tliormora jukesii BAIRD, 1865, pp. 199-200. Lepidasthenia longicirrata TREADWELL, 1928a, Lepidonotus atratus TiEADwELL, 1940, pp. 3-4, pp. 460-461, figs. 22-26. figs. 10-13. The holotype of Lepidasthenia longicirrata The type of Lepidonotus atratus comes from comes from west of Peru and may thus be pe- Digos, Gulf of Devas, Philippine Islands. It lagic. It agrees in all details with Lepidasthenia retains a few of the kinds of natans two notosetae (Chamberlin) described from west of characteristic of Thormora Baird; some are Peru, southwest coast of Central America, and coarser and betwveen spinous along their free length; Peru and Easter Island. The body is others are slenderer, smooth along their free unusually short for species of this genus, but and hastate in the length, shape. Elytra lack margi- absence of notopodial fascicles and the lep- nal Thormora is a common In- idonotoid prostomium are fringe. jjukesii characters of Lepi- do-Pacific species (Pruvot, 1930, pp. 9-11, dasthenia (Hartman, 1942a, pp. 102-103). The pl. 1, so-called notopodial setae described for L. longi- figs. 11-15). cirrata are embedded acicula. Lepidasthenia na- Harmothoe crucis (Grube), 1856 tans has unique neurosetae; in the superior part Polynoe crucis GRUBE, 1856, p. 47. of the fascicle they are very long, distally at- Harmothoi lanceocirrata TREADWELL, 1928a, tenuated, and delicately spinous; farther down pp. 454-455, figs. 5-9. they gradually shorten, become more distinctly The holotype of Harmothoe lanceocirrata spinous, and have a bifid tip. The species is not comes from known outside the type localities. the Leeward Islands east of Puerto Rico. It agrees in all details with the West In- Lepidasthenia sp., juvenile stage dian Harmothoi crucis (Grube). Diagnostic features Lepidonotus inquilinus TnADwmu, 1917, p. 258, include the elytra that are conspicu- figs. 4-10. ously fringed along their outer lateral margins and have macrotubercles on The holotype and cotype collections contain their dorsal sur- face. The tubercIes are spines, some with en- individuals small to about 15 mm. in length, tire others with taken July 6, 1915, on a Marphysa tip, bifid or trifid tip. More species at numerous ones are much finer spines. Neuro- Mangrove Key in Key West Harbor, Florida. setae are about as They are immature specimens of a coarse as notosetae; all are in species of full, dense fascicles and project laterally. Infe- Lepidasthenia. The number of elytral scars riormost setae terminate varies, but none has more than 15 pairs; in a bifid tip in which pos- the accessory tooth is at a noticeable distance terior segments are immature, prostomial an- from tennae are the main fang. Uppermost neurosetae lepidonotoid. Parapodia are long, are and an laterally projecting, with notopodia reduced to longest have entire tip. Tips of setae in the middle of the fascicle intergrade from 19561956HARTMAN: ANNELIDS 273 an entire to a bifid one. Acicular lobes of both cles, all of one kind and closely spaced; they notopodia and neuropodia are prolonged later- are limited to this area. Other parts of the ely- ally as slender processes, largely concealed by trum are smooth except for a few widely the full setal fascicles. The prostomial eyes are strewn, digitate tubercles that are delicately at- small, or one or two of the four may be ab- tached and perhaps easily lost. sent. Harmothoe crucis is a common intertidal Harmothoe sylliformia is known only from polynoid in West Indian seas. Ehlers (1887, pp. deep waters off the Galapagos Islands. 49-50, p1. 10, figs. 9-10, pl. 11, fig. 1, as Poly- noe polytricha) has given a comprehensive ac- Harmothoe variegata Treadwell, 1917 count, and Augener (1925, pp. 22-23) a partial Harmothoe variegata TREADwELL, 1917b, pp. synonymy. 260-261, p1. 1, figs. 17-18, p1. 2, figs. 1-3. The holotype comes from Dry Tortugas, Harmothoe sylliformia Treadwell, 1928 Florida, taken June to July, 1909. There are 15 Harmothoe sylliformia T=ADwzrL, 1928a, pp. pairs of elytra, and the last seven segments are 452-453, figs. 1-4. exposed. Elytra have a pigment pattern as The holotype comes from Tagus Cove, Al- shown by Treadwell (fig. 17). The prosto- bemarle Island, Galapagos Islands, in 300 to mium has four small, widely spaced eyes. Pro- 700 fathoms. It is accompanied with a calanoid stomial peaks are little developed. Notosetae are copepod and may thus be a pelagic species. The very coarse, distally slightly bifid and weakly 20 or more specimens in the collection are in pectinated along their free length. Neurosetae various stages of development; one of them are slenderer and of two kinds; most are shows traces of the fifteenth elytrophoral scar deeply bidentate at the tip and pectinated be- and the others are less fully developed. yond their thickest region; a single lowermost The prostomium is broadly bilobed; its fron- one is distally entire as shown by Augener tal peaks are very slender, papillar, and con- (1927b, p. 46, figs. A-D, as Paralepidonotus tinuous with the broad ocular lobes. The four boholensis curacaoensis Horst). This author eyes are moderately large; the anterior pair is used the confused generic name Malmgrenia located at the sides and just in front of the McIntosh, and noted that Harmothoe variegata widest part of the prostomium; the posterior Treadwell was perhaps the same as Horst's eyes are near the posterior margin of the pro- species from Curarao. He further referred stomium and also lateral. The median prostomial Harmothoe roberti Augener (1922c, pp. 40-41) furrow is little marked. The base of the median from Tortugas, Florida, to the same species antenna is large and broad; it fills the space and reviewed the confusion of generic names. between the slender prostomial peaks. Palpi and Treadwell's specific name is the oldest. cirri are smooth. Notosetae are more than twice Harmothoe variegata Treadwell is known as thick as neurosetae but much shorter; they from littoral areas of the West Indies. are transversely spinous and terminate in an entire, blunt tip. They range from nearly Eunoe purpurea Treadwell, 1936 straight to arcuate ones. Neurosetae are much Eunoe purpurea TREADWELL, 1936b, pp. 51-52, slenderer and longer; they range from longer figs. 1-6. spinous ones in the superiormost part of the Acanthicolepis longicirrrata TREADWELL, 1941d, fascicle, to short, thick spinous ones below. p. 26, figs. 1-4. All neurosetae are delicately bifid at the tip, The holotype of Eunoe purpurea comes from with the accessory tooth short and widely diver- off Bermuda in 500 to 900 fathoms; that of gent from the main fang. Typical neuropodia Acanthicolepis longicirrata is from off Ber- have about 25 or more setae in a bundle; others muda in 500 to 800 fathoms. The two are iden- have fewer. Elytra are largely lacking from the tical. The prostomium is harmothoid; the body collection. A detached one in the bottom of a consists of fewer than 31 segments; elytro- paratype vial is broadly oval and weakly exca- phores number 15 pairs. Notosetae greatly ex- vate at its anterior margin; its scar of attach- ceed in thickness the neurosetae. On the pro- ment is eccentric. The excavate region in front stomium the anterior eyes are at the sides; all of the scar has a lunar patch of microtuber- are very large with the two of a side nearly 274 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 coalescent Eunoe purpurea is known only from Eurythoe oculata TREADwELL, 1941c, p. 18, figs. great depths off Bermuda; it may be a bathy- 1-3. pelagic species. The type specimen is from Balboa, Panama, at the dock. The body is vermiform and meas- SIGALIONIDAE ures about 180 mm. long; it was sexually ma- Psammolyce arenosa (delle Chiaje), 1941 ture in July. The prostomial caruncle is a very Sigalion arenosum DELLE CHIAJE, 1841, pp. 58, small papillar lobe at the postmedian margin 107, p1. 98, figs. 4-5. of the prostomium as typical of Pseudeurythoe Psammnolyce rigida TREADWELL, 1901, p. 188; Fauvel. Branchiae are present from the third not Grube, 1868. segment to near the posterior end of the body; Eupkoloe nuda TREADWELL, 1936b, pp. 53-54, they are dendritically branched and located at figs. 10-14. the posterior face of notopodia, behind the in- The holotype of Eupholoi nuda comes from sertion of the dorsal cirrus. Notopodia have Gurnets Rock, Bermuda, in 35 feet from coral. full, spreading fascicles of long, slender, nearly Although all elytra are missing, the scars are colorless setae and an accompanying single present on segments 2, 4, 5, 7 and all odd- transverse series of several yellow acicula that numbered segments to 29, and on every segment terminate distally in an elongated knob. Neuro- thereafter to the posterior end. The four pro- podia have similar full spreading fascicles; stomial eyes are as first described. The second they are accompanied with a transverse series segment has the unique composite setae that of five or six thicker, darker spines, each with characterize Psamnmolyce arenosa (delle a short lateral boss some distance from the tip Chiaje); these setae have a long spinous shaft (about as shown by Treadwell, fig. 3, except and a short falcate appendage. The third seg- that in most setae the lateral process is smaller ment has a very long slender dorsal cirrus. In than shown). typical neuropodia the most numerous composite Pseudeurythoe oculata approaches P. ambig- setae are thick falcigers with short appendages; uus (Monro) recorded from the Pacific side the lowermost are much slenderer and have of Panama and North Carolina (Hartman, longer appendages. Psammolyce arenosa has 1945, p. 12); the latter lacks the conspicuous been widely reported from tropical parts of the prostomial eyes. West Indian and Atlantic regions. Augener (1933a, p. 193) has established some of the ?Pareurythoe elongata (Treadwell), 1931 synonymy. Hipponoe elongata TtDAIwELL, 1931d, pp. 3-4, figs. 10-12. POLYOLONTIDAE Pareurythoei elongata HARTMAN, 1951, p. 28. Polyodontes panamensis (Camberlin), 1919 The holotype comes from Puerto Rico. It is Panthalis panamensis CHAMBERLIfl, 1919, pp. large, measures about 185 mm. long, and 86-89, pl. 11, figs. 4-8, pl. 12, figs. 1-6. grossly resembles a larger individual of Eury- Polyodontes californicus TREBDWEL, 1941c, thoe pp. 20-21, figs. 9-12. complanata (Pallas) in size and propor- tions. It is at once distinguished from the The holotype of Polyodontes californicus latter for its prostomial parts. The prostomiurn comes from Chamela Bay, Mexico, in 16 fath- is subquadrate; its posterior and lateral mar- oms. It is posteriorly incomplete. Elytra are gins are largely concealed by a caruncle that colored brown along their inner halves and fade has the form of a broadly divergent V with to lighter brown at the sides; they are sur- spreading branches and its base slightly pro- rounded with a paler border. The individual longed so as to lie at the middorsum of the agrees with P. panamensis (Chamberlin) first first setigerous segment. The distal ends of described from the Pacific side of Panama and the caruncle extend forward and nearly conceal recorded also from the Galapagos Islands, from the two pairs of large dark eyes. The caruncle shore to 89 fathoms (Hartman, 1939b, p. 84). is flat depressed and thus easily overlooked, but its outlines are visible with subdued illumi- AMPHINOMIDAE nation or when the prostomium is turned to Pseudeurythoe oculata (Treadwell), 1941 the side. The anterior pair of eyes is located at 1956 HARTMAN: ANNELIDS 275 the sides of the prostomium immediately at the Islands, probably at the surface; it is a small im- sides of the posterior paired antennae. The mature stage of Anaitides and its proboscidial posterior eyes can be seen from the side or characters are those of A. madeirensis (Lang- when the overlying caruncle is pushed aside. erhans). In the original illustration (Treadwell, The base of the median antenna is visible fig. 69) the nuchal papilla is not shown; it slightly behind the line that would connect the should be indicated at t-he postmedian incision posterior paired eyes. of the prostomium. Anaitides madeirensis has The first segment is setigerous and medially a circum-mundane distribution in warm littoral divided by the posterior base of the caruncle. seas (Fauvel, 1923a, pp. 150-151, fig. 53, as The second segment extends across the body Phyllodoce). and is broader. Branchiae are first present from the third segment; they are large, dendritically ?Anaitides pulla (Treadwell), 1926 branched, and continued on segments to near PhyUodoce pulla TREADWELL, 1926a, pp. 12-13, the posterior end of the body. Notopodia have figs. 19-21. thick fascicles of yellow, very slender, distally This comes from Japan; its type is deposited pointed setae. Neuropodia have similar setae, in the United States National Museum. Its accompanied with fascicles of somewhat thicker affinities are with species of Anaitides Czerni- spines that are distally blunt and have a short awsky, and it may be close to, if not the samne lateral spur, as first described. as, A. madeirensis (Langerhans) (see above). The generic name, Pareurythoe Gustafson, is questioned because of the unique caruncle Anaitides tenuissima (Grube), 1878 which is unlike that of any other amphinomid. PP. 95-96. of Pareurythoe are dis- PhyUodoce tenuuissima GRUBE, 1878, The various species Phyllodoce fusca-cirrata TiEADWELL, 1926a, cussed elsewhere (Hartman, 1951, p. 28). PPar- eurythoe oculata differs from others in having pp. 14-15, figs. 25-27. an overlapping caruncle. The type of Phyllodoce fusca-cirrata comes from Samoa and is deposited in the United PHYLLODOCIDAE States National Museum. From its description Anaitides madeirensis (Langerhans), 1879 it can be referred to Anaitides tenuissima (Grube) first described from Bohol, Philippine PhyUodoce (Anaitis) madeireisss LANGERHANS, Islands, and well illustrated as Phyllodoce ma- 1879, pp. 307-308, p1. 17, fig. 44. by Willey (1905, p. 265, pL. 3). Phyllodoce varia TREApwELL, 1928a, p. 467, crolepidota figs. 69, 70. Fauvel (1932a, p. 70, as Phyllodoce) recorded Anaitides minuta TREADWELL, 1937b, p. 148, if from Indo-Pacific seas and gave synonymies. pl. 2, figs. 16-18. Mystides gracilis TREADWELL, 1941d, pp. 27-28, Phyllodoce violacea Treadwell, 1926 figs. 5-7. Phyllodoce violacea TREADWELL, 1926a, pp. 13- The type of Anaitides minuta is from Arena 14, figs. 22-24. Bank, western Mexico, in 35 fathoms; that of Phyllodoce variegata TREADWELL, 1926a, p. 12. Mystides gracilis is from off Bermuda at the Phyllodoce pruvoti FAUvEL, 1930a, pp. 512-514, surface. Both are representatives of Anaitides fig. 1. HARTMAN, 1954c, pp. 631-633, fig. 175D, Czerniawsky. The proboscis has, on either side G-I. near the base, about six longitudinal rows of The type of Phyllodoce violacea originates papillae with six or more in a row. The pro- from Suva, Fiji, and is deposited in the United stomium has a nuchal papilla at its postmedian States National Museum. Striking features in- incision but no median antenna. The first two clude its great length (attaining about 1000 tentacular segments are fused, thus not as in mm.) and the brilliant coloring of iridescent to Mystides Theel in which they are distinct. In dark brown. Dorsal cirri are dark brown and posterior segments the dorsal cirri are sub- have very pale borders. The prostomium has tetragonal as typical for Anaitides madeirensis two very small eye spots. Composite setae have (Langerhans). a short, slightly curved, smooth appendage Phyllodoce varia comes from the Galapagos (Hartman, 1954c, fig. 175). These characters 276 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 agree with those of Phyllodoce pruvoti Fauvel; Halyplanes gracilis is known from northern, the two are considered identical. The species southern equatorial, and Guinea currents (Rei- has a distribution that includes Fiji, the Loyalty bisch) and in tropical equatorial regions of the Islands (Fauvel, 1930), and the northern Mar- Pacific Ocean (Treadwell, 1943, map 1). shall Islands (Hartman, 1954). IOSPILIDAE Phyllodoce tortugae Treadwell, 1917 Iospilus phalacroides Viguier, 1886 PAyllodoce tortugae TREADwELL, 1917b, p. 262, figs. 4-6. lospilus phalacroides VIGUIER, 1886, p. 392, pl. 23, figs. 1-6. FAUVEL, 1923a, p. 194, fig. 72e. This originates from Loggerhead Key, Dry Phalacrophorus niger TREADWELL, 1943b, pp. Tortugas, Florida. On the proboscis the papil- 33-34, fig. 15. lae are dispersed, but they leave bare a broad The type of Phalacrophorus niger originates middorsum and midventrum as first shown. from the north central Atlantic Ocean, pelagic The body is greatly prolonged; dorsal cirri are in 50 meters. It is considered identical with foliaceous, long, and triangular and they have lospilus phalacroides Viguier, especially be- pinnately arranged markings. The species is cause segments 3 and 4 are without dorsal and known only through its original discovery. ventral cirri, the pigmentation pattern is identi- LOPADORHYNCHIDAE cal, and the pharynx is presumably unarmed. These characters are those of Iospilus Viguier, Halyplanes gracilis Reibisch, 1893 not Phalacrophorus Greeff. lospilus phalacro- Halyplanes gracilis REIBISCH, 1893, p. 252. ides occurs in warmer parts of the Atlantic Haliplanes gracilis REIBISCH, 1895, pp. 24-26, Ocean and off southern pl. 2, figs. 10-13. BERGSTR6M, 1914, P. 90. Europe (Fauvel, 1923a, Haliplanella pacifica TREADWELL 1943b, p. 32, p. 194). p1. 1, figs. 4-6. Phalacrophorus pictus Greeff, 1879 The holotype of Haliplanella pacifica is in the Phalacrophorus pictus GREEFF, 1879, p. 249, United States National Museum. Its specific p1. 14, figs. 25-30. REIBISCH, 1895, P. 10, p1. 1, name suggests that it came from the Pacific figs. 4-7. FAUVEL, 1923a, p. 196, fig. 72f. Ocean, but it originates from the middle At- Phalacrophorus maculatus TREADwELL, 1943b, lantic Ocean in 100 meters. A paratype col- p. 34, figs. 11-13. lected from an adjacent station in about the A paratype of maculatus in same depth is in the Phalacrophorus American Museum of Nat- the American Museum of Natural History com- ural History. The latter is a small individual pares fully with P. pictus Greeff (Reibisch, that measures less than 1.5 mm. long and is 1895, p. 10). very darkly stained. The segments with tentac- Phalacrophorus pictus is recorded ular cirri are completely free from equa- from one another. torial parts of the Pacific and Atlantic oceans, The two pairs of prostomial antennae are very between latitude 400 N. and latitude 400 S. slender and short. The first segment has two pairs of slender cirri and a setigerous fascicle Phalacrophorus uniformis Reibisch, 1895 with simple spine-like setae. The second seg- Phalacrophorus uniformis REIBISCH, 1895, pp. ment has a pair of very large tentacular cirri 15-17, p1. 1, figs. 10-16. and fascicles of composite setae. Segments ? Phalacrophortis attenuatus TREADWELL, 1943b, farther back have dorsal cirri that are short p. 34, fig. 14. and compressed cylindrical. These features are This species is distinguished from P. pictus identical with those of Halyplanes gracilis Rei- Greeff bisch for having an attenuate, instead of short, (1893) known from the same general The anterior or localities. The original generic name, Haly- body. eight more, instead of only planes Reibisch (1893, p. 252), three, parapodia are rudimentary. was later Phalacrophorus attenuatus agrees with P. uni- spelled Haliplanes (Reibisch, 1895, p. 24; Berg- formiis in these respects. stro6m, 1914, p. 90). The first comes from Haliplanella Treadwell the Marshall Islands; the second, from the Indo- (1943, p. 32) was first used by Treadwell al- Pacific South erroneously and Atlantic areas (Reibisch, thouzgh attributed to Reibisch. 1895, p. 17). Their identity is suggested. 1956 HARTMAN: ANNELIDS 277 ALCIOPIDAE 600 fathoms. Prostomial, parapodial, and anal Rhynchonerella angelini (Kinberg), 1866 structures are considerably damaged but are about as first shown (Treadwell, figs. 66-68). Kronia angelini KINBERG, 1866, p. 242. This is identical Travisiopsis Eulalia magnapupula TREADWELL, 1941c, pp. 21- with coniceps 22, figs. 13-17. (Chamberlin) described from about the same locality, off Peru and the Galapagos Islands Two individuals are labeled Eulalia magna- from surface to 300 fathoms. they originate from Stn6p-Bowitz Pupula; the Pacific side of (1948, p. 61) has designated the synonymy Panama in 50 fathoms in plankton. They are and representatives, not of Eulalia Savigny, but of geographic distribution. an alciopid, Rhynchonerella angelini (Kin- Travisiopsis lanceolata Southern, 1911 berg), that is common in very the warmer Travisiopsis lanceolata SOUTHERN, 1911, pp. 30- parts of the eastern Pacific Ocean. Stf6p-Bo- 32, pl. 1, figs. 3, 5,6. witz (1948, pp. 34-36) has given a diagnostic Travisiopsis atlantica Ti ADwFLL, 1936, pp. 62- account and synonymy. 63, figs. 30-33; 1941c, p. 22. The of Naiades cantrainii delle Chiaje, 1830, holotype Travisiopsis atlantica comes St0p-Bowitz, 1948 from Nonsuch Island, Bermuda, pelagic in 600 fathoms; later it is recorded from the Pacific Naicades cantrainitS DELLE CHIAJE, 1830, se&swu side of Panama St0p-Bowitz, 1948, p. 24. in 500 fathoms (Treadwell, Alciopa distorta TiJEADwE.L, 1943b, p. 35, figs. 1941c). The type agrees fully with recent ac- 16-18. counts of T. lanceolata (St0p-Bowitz, 1948, p. 58; Friedrich, 1950, p. 317). Monro (1939, The holotype of Alciopa distorta is at the pp. 347-348) has recorded the species off United States National Museum and originates southern New England in 0 to 750 meters and from east of the Hawaiian Islands in 50 meters. off Brazil in about the same The description, and geographic range agree depth. fully with published accounts of Naiades can- trainii delle Chiaje (see St,p-Bowitz, 1948, TOMOPTERIDAE pp. 24-25, fig. 15). Tomopteris nisseni Rosa, 1908 Tomopteris nisseni RosA, 1908, p. 1. Vanadis crystallina Greeff, 1876 Tomopteris opaca TnRADwEu., 1928a, pp. 463- Vanadis crystallina GREEFF, 1876, pp. 68-69, pl. 464, fig. 29. 3, figs. 35-39. Tomopteris longisetis TREADwELL, 1936b, pp. Vanadis collata TREADWELL, 1928a, pp. 462-463, 58-59, figs. 18-21. figs. 27, 28. Tomopteris opaca comes from the Galapagos The holotype of Vanadis collata was collected and Cocos Islands. Its most conspicuous fea- off the Galapagos. Islands in 400 fathoms. The hure is the frilled condition;of the parapodial first six or more parapodial pairs are very small, flanges. The posterior end is without a tail but as characteristic, of V. crystallina Greeff. The is frayed and thick, and appears as if a tail had two agree also in prostomial and parapodial been present and was broken off. In other re- parts. Treadwell (1928a, fig. 27) shows the spects it agrees fully with.Tomopteris nisseni proboscis partly everted. Rosa as described by Stop-Bowitz (1948, pp. .44-46, fig. 29). Tomopteris longisetis comes TYPRLOSCOLECIDAE from Nonsuch Island, Bermuda, in 500 to Travisiopsis coniceps (Chamberlin), 1000 fathoms and is at once recognizable as a 1919 large individual of T. nisseni Rosa. In this re- Plotobia coniceps CHAMBERLIN, 1919, pp. 156- spect, however, it is exceeded by one measuring 158, pl. 66, figs. 2-4. 105 mm. long, 5 mm. and with 39 Nuchubranchia palmata TREADWELL, 1928a, wide, para- pp. 481-482, figs. 66-68. podial segments described by St0p-Bowitz (1948, p. 45). This species is very common in The holotype of Nuchubranchia palmata northern Atlantic waters in depths from sur- comes from the Galapagos Islands in 400 to face to 3239 meters (StSp-Bowitz, 1948). 278 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Tomopteris tentaculata Treadwell, 1928 ?Nereimyra mutilata (Treadwell), 1901 Tomopteris tentaculata TRi-iEAW., 1928a, pp. Castalia mutilata TREADwULL, 1901b, p. 185, 464-465, figs. 30-31. fig. 4. This comes from the Galapagos Islands in Irma mutilata TREADWELL, 1939b, pp. 213-214, 1100 fathoms. It is only about 6 mm. long and fig. 41. may be immature. Conspicuous features include Two very imperfect specimens come from the paired epaulettes that form a raised ridge San Antonio Bridge, San Juan, Puerto Rico. on either side of the prostomium. The first They have lost dorsal and tentacular cirri and pair of cirri are largely concealed beneath the setal appendages. The species cannot belong to larger second pair. The chromophile glands Irma Grube, which is a synonym of Podarke are located at the inferior part of the neuro- Ehlers (Horst, 1921, p. 78), because a median podial flange. Small gonadial elements are vis- prostomial antenna is absent. Castalia Savigny ible in the upper part of the neuropodial lobe. is preoccupied and is replaced by Nereimyra There is no tail. This may represent a juvenile Blainville stage of a (St6p-Bowitz, 1948, p. 61). better-known species. The body is about 17 mm. long and consists HESIONIDAE of 53 segments. The prostomium is quadrate, wider than and has a ?Dalhousiella longicirrata (Treadwell), 1901 long, pair of large, bi- articulated and a of small fron- Castatia palpi pair very longicirrata TREADWEL, 1901b, p. 185, tal antennae. There are four eyes in quadrate figs. 2,3. Tentacular cirri number six Hesione longicirrata TRDwi&LL, 1939b, pp. arrangement. pairs. 216-217, fig. 44. Parapodia are uniramous. The generic name is questioned because the character of setal ap- This comes from Fish Hawk Station 6079, is not known. off St. Thomas, Virgin Islands, Sail Rock, in 20 pendages to 23 fathoms in coral, and is deposited in the ?Leocrates oculata United States National Museum. It cannot be- (Treadwell), 1906 long to Hesione Savigny which is characterized Castalia oculata TREADwrLr, 1906, p. 1148, by having no palpi. It is not Castalia Savigny (a figs. 1-3. synonym of Nereimyra Blainville) which has The holotype specimen is in the United only six pairs of tentacular cirnr. The body is States National Museum; it comes from the short and consists of about 19 segments. Para- vicinity of Hawaii in 49 to 176 fathoms on a podia are uniramous. The prostomium has a pair rocky bottom. The body is short, fusiform, and of articulated palpi, a pair of slender frontal consists of 16 parapodial segments. There are antennae, and a large facial tubercle. There are eight pairs of tentacular cirri. The frontal mar- four eyes in trapezoidal arrangement. Tentacu- gin of the prostomium has a thick, short, me- lar cirri number eight pairs. Setae are composite dian antenna. The facial tubercle is a minute falcigers in which the appendage is long, knife- papilla. A median antenna is lacking but was like, and distally bifid. These characters are those perhaps lost from the specimen. Parapodia are of Daihoiella McIntosh (see Fauvel, 1923a, p. biramnous; notopodia have slender setae, and 234). neuropodia have composite falcigers with a bi- Three other species of Dalhousiella have been fid tip. These characters agree with those of described from West Indian localities. These Leocrates chinensis Kinberg (Hartman, 1948b, include D. longisetis (Grube, 1856, p. 15) from p. 47) except that the anterior eyes are greatly St. Croix which is incompletely known; D. enlarged and a median antenna is absent. The carpenteri McIntosh, recorded from the Carib- identity of the two cannot therefore be deter- bean Sea in 720 to 800 meters (Monro, 1939, mined. p. 348); and D. hesionides (Augener, 1906, p. 157, as Castalia) off the West Indies in SYLLIDAE 164 and 399 fathoms. Dalhousiella carpenteri, Odontosyllis 1900 including probably D. hesionidens, is character- enopla Verrill, ized for having composite falcigers which are Odontosyllis enopla VmuLL, 1900, pp. 627-628. Autolytus bidens TREIADW., 1941d, pp. 28-29, deeply bifid at the tip. fig. 9. 1956 HARTMAN: ANNELIDS 279 The type of Autolytus bidens comes from Nereis decora TREADWELL, 1932b, pp. 15-17, Bermuda off shore at the surface. It is a typical figs. 6-11. representative of Odontosyllis enopla Verrill, Nereis ambiguus TiEADwELL, 1937b, pp. 149- first described from the same place (Verrill, 151, pl. 2, figs. 19-24. 1900, 627). Ventral cirri are com- Nereis (Neanthes) paucidentata TREADWEL, p. triangular, 1939a, p. 6, fig. 25. pressed lobes. The prostomium is overlain at Nereis (Neanthes) varia TPEADWEmL, 1941a, its posterior margin by a nuchal hood; com- p.3. posite setae are identical. Galloway and Welch (1911, pp. 13-39, pls. 1-5) have described the The type of Nereis decora comes from Villa luminescence of this species, and Huntsman Bella, Ilha de Sao Sebastiao, Brazil; that of N. (1948, pp. 363-369) has described the condi- ambiguus originates from Sulphur Bay, Clarion tions for spawning at Bermuda. Island, in 20 fathoms. Nereis (Neanthes) pau- cidentata is from Charlestown, Massachusetts. NEREIDAE The name Nereis (Neanthes) varia was pro- Nicon mexicana (Treadwell), 1942 posed to replace the last-named, presumed to be Leptonereis mexicana TREADWELL, 1942, pp. 1-4, preoccupied. figs. 2-9. The proboscis of the specimen of Nereds The holotype comes from outside Topolo- ambiguus which I examined had not been ob- bampo Bay, Baja Califomia, Mexico; others served, as it was withdrawn and not dissected. are from Grande Point, San Jose Bay, Baja Area I has three small teeth in tandem; II has California. All are in the epitokous reproduc- a patch of larger, dark amber paragnaths; III tive stage and include both male and female is similar to II, IV resembles VI in having individuals. Parapodia have no foliaceous dor- about 10 to 12 paragnaths in two or three ir- sal lobe which characterizes Leptonereis Kin- regular rows; V has none; VI has about 12 berg (Hartman, 1948b, p. 56, pl. 8). In the cones in a transverse patch of three irregular male epitoke there is a long pre-epitokal re- rows; VII and VIII (continuous) have a gion of at least 55 segments and a long post- single row of 10 larger and smaller cones. epitokal or third region with normal parapodia. Areas II and III are the most conspicuous. Prostomial eyes are large; each is round and The original description of the proboscis stated has a reddish brown lens occupying most of the that paragnaths are absent from areas I and II, upper disk. Notopodia have no homogomph fal- III has a tuft of about eight on a side, IV has cigers. Acicula are black and occur singly in three in a transverse row, V has none, VI has parapodial rami. The prostomium is subrec- three or four, VII has seven or eight very long tangular in its preocular part instead of tri- and blunt-pointed cones on either side, having angular as originally shown. The paired pro- irregular ones in front of them, and VIII a stomial antennae are large and digitate; they circular patch of about 20. I am unable to fit extend distally nearly to the end of the palpo- this formula, especially as it concerns areas VII style. In the male epitoke the posterior para- and VIII, to the specimen examined by me. podia have a long, slender, dorsal cirrus extend- The jaws are large and pale except for the den- ing distally beyond the notopodial lobe; the tate edge, which has about nine light brown cirrus is inserted at the upper base of the lobe. teeth along the cutting edge. Homogomph fal- The latter is long and slender; it resembles the cigers are first present from about segment 26, middle lobe but is slenderer. The everted pro- or the middle of the body. Acicula are black boscis is smooth; it is without pharyngeal proc- and occur singly in a ramus (Hartman, 1940a, esses. The distal paired jaws are dark horny pp. 221-222). brown and have crenulated teeth at the cutting The holotype of Nereis (Neanthes) pauci- edge. dentata measures about 40 mm. long and 5 mm. Nicon mexicana is known only from the wide; peristomial tentacles are moderately western side of Mexico. thick and short. In median parapodia the distal lobes are dark; the dorsal lobe is very large Nereis riisei Grube, 1856 and has its dorsal cirrus inserted at the upper Nereis ri'isei GRuB3E, 1856, pp. 162-163. base. On the pharynx area I has a single cone, 280 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109

II has three to six in a cluster, III has an ir- Nereis arroyensis Treadwell, 1901 regular double row, IV has a diagonal double Nereis arroyensis TREADWELL, 1901b, pp. 193- row, V and VI have one each, and VII and 194, figs. 30-31. HOAGLAND, 1919, p. 574. TnAD- VIII are continuous and have a double alter- WELL, 1939b, p. 223. nating row. These several individuals are believed iden- The holotype is in the United States Na- tical and are referred to the common West In- tional Museum and comes from Arroyo, Puerto dian. Nereis riisei Grube. It has been further Rico. The pharyngeal armature has not previ- reported from Haiti, Florida, and French Gui- ously been described. The paragnaths of the ana (Fauvel, 1932b, pp. 28-30, fig. 4), also maxillary ring are larger than those on the oral the Gulf of California to Ecuador (Hartnan, one. Area I has three pointed cones in tan- 1940a, pp. 221-222, pl. 33, fig. 37). Its geo- dem; II is a crescent-shaped patch with about graphic range extends along both sides of 12 larger cones; III has a broad patch of 25 middle America and the coastal fringes of the to 30 cones; IV has about 20 pointed cones. On -Gulf Stream. the oral ring V is bare; VI has a cluster of six pointed cones; VII and VIII are continuous Nereis pelagica largoensis Treadwell, 1931 and have many smaller and a few larger cones Nereis brevicirrata T1.ADWELL, 1929d, pp. 3-5, in three to five irregular rows. The jaws are figs. 9-14. dark brown; each has five coarse, blunt teeth. Nereis largoensis TREADWELL, 1931, p. 3. HART- The parapodial lobes resemble those of Nereis MAN, 1951, pp. 45-46. irrorata (Malmgren) (Fauvel, 1923a, p. 342). The holotype comes from Key Largo, Flor- Nereis arroyensis is not known ouside ida. It has much resemblance to Nereis pelagica Puerto Rico. Linnaeus, especially in its prostomial and para- podial parts. On the pharynx area I has two Nereis amoyensis (Treadwell), 1936 small cones in tandem (originally given as Nereis (Neanthes) amoyensis TREADWEL, one) ; area V has a single small cone (originally 1936b, pp. 272-273, figs. 19j-m. described as absent); VI has four large para- The holotype is in the United States gnaths in diamond arrangement; areas VII National Museum and comes from and VIII (continuous) have three or four rows Amoy, China. It is to in a transverse the row on referred Nereis, sensu stricto, instead of band, with outer -the Neanthes maxillary side having the largest ones. Kinberg because the notopodia have In median and posterior noto- fglcigerous setae. The appendage resembles that parapodia the of podial lobe increases in length going back to a Nereis pelagica Linnaeus, with which this rectangular shape; the dorsal cirrus is inserted species may be allied. at the upper base of this lobe. In posterior seg- ments notopodia have homogomph falcigers in Ceratonereis anchylochaeta (Horst), 1924 which the appendage is about six times as long Nereis (Lycoris) anchylochaeta HORsT, 1924, as wide, and slightly curved in its distal half pp. 155-157, pl. 30, figs. 8, 9. when seen in profile; the cutting edge is smooth, Ceratonereis longicauda TREADWELL, 1943a, and the free part is about as long as the em- pp. 1-3, figs. 1-7. bedded part. The holotype of Ceratonereis longicauda Nereis pelagica largoensis Treadwell differs -comes from Padado, Gulf of Davao, Philippine from the stem species chiefly for having the Islands. Notopodia have only spinigerous setae. posterior notopodial lobes longer, and area V On the pharynx the oral ring has paragnaths has a paragnath instead of lacking it. It differs only on areas VII and VIII; there are three from N. pelagica occidentalis Hartman (1945, cones in a transverse row and each one is on a p. 20) from North Carolina in that the noto- *brown, weakly chitinized patch. Neuropodia :podial falciger has an appendage of another have composite falcigers in anterior and median kind. -Nereis pelagica largoensis is recorded segments. Posterior neuropodia have a charac- from areas in the Gulf of Mexico (Hartman, teristic simple falciger (Horst, 1924, figs. 8-9). 1951,p.46). Ceratonereis anchylochaeta is widely re- 1956 HARTMAN: ANNELIDS 281 corded from Indo-Pacfic regions (Fauvel, east Pacific. It has been more widely recorded 1932, pp. 88-89). from Hawaii and Australia (Hartman, 1954b, pp. 36-37, figs. 38, 39). Platynereis dumerilii (Audouin and Milne-Edwards), 1834 Perinereis helleri Grube, 1878 Nereis dumerilii AUDOUIN AND MLNE-ED- Nereis (Perinereis) helleri GRuBE, 1878, pp. 81- wARDs, 1834, p. 296. 82. Uncinereis lutea TREADWELL, 1928a, pp. 469- Neanthes obscura TiEADwELL, 1928a, p. 472, 471, figs. 40-49. figs. 50-53. Leptonereis maculata TREADWELL, 1928a, p. 469. Uncinereis trimaculosa T1xADwEzL, 1940, p. 3, The holotype of Neanthes obscura comes figs. 4-9. from west of Costa Rica. In the original ac- Eunereis africana TREADWELL, 1943c, pp. 2-3, count of the pharyngeal armature, the formula figs. 8-13. should read area VI with a single transverse The holotype of Uncinereis lutea comes from piece on either side, areas VII and VIII con- the Sargasso Sea on sargassum. Leptonereis tinuous with a double irregular row of para- maculata (not a holotype, as it was presumed gnaths. This is a typical representative of Peri- to be the same as Nicon maculata Kinberg) is nereis and agrees with P. helleri Grube. The from the vicinity of the Galapagos Islands. distal ends of the peristomial cirri are pseudo- Uncinereis trimaculosa comes from a floating annulated, as is the case for many nereids log at Galveston, Texas, and Eunereis africana from southern Pacific regions. Fauvel (1932, p. was taken at Lobito, Africa. All are typical rep- 104) has given a diagnostic key of the group resentatives of Platynereis dumerilii (Audouin of species to which P. helleri Grube belongs. and Milne-Edwards) and come from localities wvithin the known geographic range. Specimens Perinereis monterea (Chamberlin), 1918 labeled Leptonereis maculata are in the epito- Nereis (Neanthes) monterea CHAMBERLIN, kous stage; the pharyngeal pectinae are present 1918, pp. 174-175. but faded. Some have paler, others darker, pos- Nereis spinifera TREADWELL, 1929d, pp. 5-6, terior parapodial lobes. Unique features of all figs. 15-20. are the reduced pharyngeal pectinae, the com- The holotype individual of Nereis spinifera posite notopodial falciger of .median and pos- comes from Puget Sound, Washington; it terior parapodia, and the long tentacular cirri agrees fully with Perinereis monterea (Cham- (Hartnian, 1945, p. 22). berlin) which is known from other parts of Platynereis dumerilii is cosmopolitan in warm the northeastern Pacific Ocean (Hartnan, seas and associated with algae. In epitokous 1940a, p. 229). stages .the pharyngeal parts tend to change in the direction of loss or darker pigmentation of Pseudonereis variegata (Grube), 1856 the basal areas. Nereitepas variegata GRUBE, 1856, p. 165. Perinereis diversidentata TREADWEL, 1943c, Platynereis bicanaliculata (Baird), 1863 pp. 1-2, figs. 1-7. Nereis bicanaliculata BAIPD, 1863, P. 109. Pseudonereis variegata DAY, 1953, p. 425. Nereis notomacula TREADWELL, 1914b, pp. 191- The holotype indivridual of Perinereis diver- 192, p1. 11, figs. 8-12. sidentata comes from Cape Cross, Africa. The A syntype of Nereis notomacula comes from everted pharynx has pectinae on the maxillary San Francisco Bay, Califomia. It is charac- ring and chitinized cones on the oral ring; this terized especially for the presence of simple, is therefore referred to Pseudonereis Kinberg. dark brown falcigers in median and posterior Posterior dorsal lobes are elongate to rectilinear notopodia, occuring usually singly in a para- and have the dorsal cirrus attached so as to podium. In other respects also this compares leave a free margin at its inferior side which with Platynereis bicanaliculata (Baird), earlier is about as broad as the base of the dorsal cir- known as P. agassizi (Ehlers) and as such re- rus. Pseudonereis variegata (Grube) has been ported from many littoral regions of the north- recorded from South Africa (Day, 1951, p. 32), -2-82 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 and the synonymy has been established (Day, about segment 11 or not before 20 and have 1953, p. 425). filaments that number from one to five at maxi- EUNICIDAE mum development (rarely they may number eight or nine filaments). Acicula are dark Eunice armillata (Treadwell), 1922 brown to black, subacicular hooks are distally Leodice armillata TRADwEI, 1922a, pp. 144- hooded and have a bifid tip, with the lateral 147, figs. 26-29, pl. 3, figs. 14-19. tooth about at right angles to the shaft. Pro- Leodice gracili-cirrata TREADWELL, 1922a, pp. stomial 149-150, figs. 36-38, 5, figs. antennae are short and smooth or ir- p1. 1-8. regularly articulate. Hartman (1944b, pp. Leodice armillata comes from Aua and Utile 110-111) gives a partial synonymy. Eunice reefs in Pago Pago Harbor, Samoa; Leodice afra is widely distributed in Indo-Pacific and gracili-cirrata is from Suva Harbor, Fiji. In West Indian seas, in littoral zones. both specimens the prostomial antennae are neatly articulated; dorsal cirri are prolonged. Eunice crassi-tentaculata (Treadwell), 1922 Branchiae are first present from about the Leodice crassi-tentaculata Treadwell, 1922a, third or fourth segment and have up to seven pp. 146-148, figs. 30-33, pl. 4, figs. 1-5. filaments; branchiae are absent from posterior The comes from segments. Acicula are holotype Pago Pago Harbor, yellow; subacicular hooks Samoa. It is near E. afra Peters (see above) occur singly in parapodia; they terminate dis- in the tally in a bifid tip having first branchiae present far back, in which the accessory tooth in case not before is at right angles to the fang. The two this segment 30. Branchial are be- filaments are limited to two at maximum lieved to agree in all specific details. The species devel- is cdose to, if not identical Eunice opment. Acicula and subacicular hooks are with, min- dark to the latter are danevensis McIntosh (1885) from the Philip- black; distally bifid. The pine Islands and specimen differs from typical E. afra in having other parts of the Southern antennae Hemisphere (see Ehlers, 1908, pp. 90-92, p1. prostomial that are proportionately 12). very large, long, and conspicuous. Eunice afra Peters, 1854 Eunice grubei Gravier, 1900 Eunice afra PrEns, 1855, p. 39. Eunice grubei GRAVIER, 1900, pp. 258-261, figs. tLeodice notata TinADWI-., 1921a, pp. 52-54, 125-129, pl. 14, figs. 87, 88. figs. 164-173, p1. 4, fig. 12. Leodice biformi-cirrata TPADwRLL, 1922a, pp. Leodice bucciensis TuADWEZ, 1921d, pp. 54- 148-149, figs. 34, 25, p1. 4, figs. 6-11. 56, figs. 174-183. The holotype of Leodice biformi-cirrata came Leodice guanica TwADwx, 1921d, PP. 3940, 1920. figs. 107-116, 2, figS. from Suva, Fiji, April, Branchiae are p1. 9-12. present from the third setigerous segment and Leodice aciculata TR14DWEII, 1922a, pp. 143- have to 144, figs. 24, 25, p1. 3, figs. 7-13. up seven filaments in pectinate arrange- Leodice flava-punctata T-AiiW , 1922a, P. ment. Prostomial antennae are moniliform. Aci- 136-138, figs. 8-11, p1. 2, figs. 1-7. cula are dark to black. Subacicular hooks are Leodice suviensi TISADWEIL., 1922a, pp. 138- distally bifid, with the subdistal one oblique to 139, figs. 12-16, p1. 2, figs. 8-13. the shaft. Composite setae are bidentate and distally The type specimen of Leodice aciculata and falcigerous. This agrees fully with Eunice that of L. grubei Gravier, first described from the Red suzriensis originate in Suva Harbor, Sea and Fiji; that of L. flava punctata comes from well known from Indo-Pacific areas Pago Pago Harbor, Samoa; L. guanica comes (Fauvel, 1932, p. 136). from Sand Key, Florida; L. bucciensis comes Eunice tubifex 1904 from Buccoo Bay, Tobago, West Indies, and Crossland, L. notata is also from the West Indies. All are EUnice tubifex CROSSLAND, 1904, pp. 303-310, considered figs. 52-55, p1. 21, figs. 14. identical with the circum-mundane, Leodice tubicola TREADWELL, tropical Eunice afra Peters. Characteristic fea- 142, 1922a, pp. 139- tures include the figs. 17-23, pl. 3, figs. 1-6. distribution and kind of The branchial branching; they are first present from holotype of Leodice tubicola comes from Pago Pago Harbor, Samoa. It is accompanied 19561HARTMAN: ANNELIDS 2832 by fragments of the tube. Characteristic fea- Eunice savignyi was first described from the tures include two kinds of composite setae (fal- Philippine Islands and has been recognized as cigerous and spinigerous), the sharply recurved a representative of the fauna of the Agulhas yellow subacicular hooks, and the unique tube. Current (Fauvel, 1932, p. 136). Treadwell recognized the similarities of the two but considered tubicola different from Eunice norvegica (Linnaeus), 1767 tubifex, especially because of differences in Nereis norvegica LINNA&Eus, 1767, p. 1086. size. Eunice tubifex is recorded from Indo- Leodice arcturi TREADWELL, 1928a, pp. 475-477, Pacific seas (Fauvel, 1930b, p. 26), the China figs. 32-39. Sea (Monro, 1924), and southern Africa (Day, The holotype of Leodice arcturi was taken 1934, p. 50). off New Jersey in 633 fathoms. It is a slender individual, measuring about 80 mm. long and Eunice savignyi Grube, 1878 2 mm. wide. The prostomial antennae are Eunice savignyi GRUBE, 1878, pp. 150-151. smooth and cirriform. Branchiae are present Leodice langi TREADWELL, 1943c, p. 3, figs. from the sixth or seventh setiger and have up 14-18. to 14 pectinately arranged filaments; they end The holotype of Leodice langi comes from abruptly after about segment 42. Subacicular Cape Town, South Africa. The mandibles were hooks are yellow and distally bifid, hooded. lost in dissection but are illustrated (Treadwell, Composite hooks are bifid falcigers. These fig. 18). The five prostomial antennae are torn characters agree with those of Eunice norvegica off at the base, so that it is not known whether (Linnaeus) (as E. pennata in Fauvel, 1923a, they were smooth or articled. Prostomial eyes pp. 400401, figs. 56h-o). The species is known are large, dark, and visible between the bases to be a deep-water commensal of polyps in of the two outer pairs of antennae. One of the north Atlantic regions. peristomial cirri remains; it extends back to the first setigerous segment and has four arti- Eunice filamentosa Grube, 1856 cles, a longest basal and two shorter more dis- Eunice filamentosa GRuBEr, 1856, p. 56. tal; the distal-most one is again longer. Bran- Leodice sspongicola TREwWELL, 1921d, pp. 25- chiae are first present from the third setigerous 27, fig. 53. segment as a minute filament located near the The holotype of Leodice spongiola comes upper base of the long dorsal crrus. At the from Montego Bay, Jamaica, taken in July, sixth or seventh segment each branchia is bifid, 1931. Branchiae are first present from the with the two branches about equally long but twenty-fourth setiger. Acicula are yellow; sub- greatly surpassed in length by the larger dorsal acicular hooks are yellow and have the unique cirrus. The eighth segment has a trifid branchia. structure known for Eunice filamentosa (Hart- The number of filaments increases to about man, 1944b, p. 107, pl. 6, figs. 123-126). The eight at parapodium 19 and decreases again species is known mainly from West Indian lo- thereafter. Branchiae are absent from a long calities in littoral zones. posterior end. Ventral cirri are dtick, cirriform through ?Eunice tenuis (Treadwell), 1921 the sixth setiger; thereafter a broad, glandular pad covers most of the ventral cirrus to near Leodice tenuis TREADwELL, 1921d, pp. 51-52, the slenderer tip. Acicula and subacicular hooks figs. 154-163, pl. 4, fig. 11. are pale yellow. The first are slightly curved at The holotype comes from Dry Tortugas, the tip and taper to a slender end. The latter Florida. The prostomium and jaws are lacking. occur singly or by two's in a parapodium; they Branchiae are first present between segments 80 terminate in a bifid tip in which the main fang to 90 and have single long filaments on many is nearly at right angles to the shaft and the segments, or successive segments have none. secondary tooth is mudh smaller. Composite Maximum number of filaments is three. Acic- falcigers are bifid, with the main tooth only ula are yellow; subacicular hooks are pale, dis- slightly curved to the appendage and the sec- tally bifid, and hooded. Composite setae are cnidary tooth much smaller than the main one. distally bifid falcigers and hooded. This indi- 284 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 vidual resembles species of Eunice (Nicidion) Lysidice collaris Grube, 1870 mainly for the great reduction of its branchiae. Lysidice collaris GRUBE, 1870, pp. 495-496. It may approach E. (Nicidion) cariboea Grube, Lysidice sulcata TREADWELL, 1901, p. 200, figs. also known from West Indian seas. 47, 48. .Lysidice fusca TREADWELL, 1922a, pp. 154-155, Eunice (Nicidion) gracilis Crossland, 1904 figs. 42-44, P1. 6, figs. 7-13. Nicidion gracilis CROSSLAND, 1904, pp. 327-329, fLysidice parva TREADWELL, 1922, pp. 155-156, figs. 65, 66, p1. 22, figs. 10, 11. figs. 45, 46, p1. 6, figs. 14-17. Nicidion fusca-fasciata TREADWELL, 1922a, pp. The type of Lysidice sulcata is from Puerto 156-157, figs. 47-50, p1. 7, fig. 5. Rico; that of L. fusca is from Samoa; and that The holotype of Nicidion fusca-fasciata of L. parva, from Samoa. The last two differ (given as Nicidion flava-fasciata in the records from each other chiefly in that the former is of the American Museum of Natural History) larger than the latter. All are referable to comes from Pago Pago Harbor, Samoa, 1928- Lysidice collaris Grube, known from cosmo- 1929. It agrees fully with Eunice (Nicidion) politan tropical littoral seas. Okuda (1937, pp. gracilis Crossland, first described from Zanzi- 288-289, figs. 34a-e) has illustrated the diag- bar and widely recorded from Indo-Pacific seas nostic parts. (Fauvel, 1932, pp. 140-141, figs. 20a-f). KEY TO SPEcIES OF Marphysa DESCRIBED -Y Palola siciliensis 1840 TREADWELL (Grube), 1. Composite setae absent. .Marphysa mossambica Eunice siciliensis GRuBE, 1840, p. 83. Composite setae present...... 2 Leodice viridis vernalis TREADWELL, 1922a, pp. 2. Composite setae spinigerous...... 3 133-134, pl. 1, figs. 8-1. Composite setae falcigerous...... 5 The holotype of Leodice viridis vernalis Composite setae both falcigerous and spini- comes from Suva Harbor, Fiji, April, 1920. gerous; branchiae abrupt and limited to about It is a small individual and measures about segments 10 to 22; eyes large.. 2.5 mm. across. There are no ...... Marphysa bellii oculata subacicular hooks. 3. Composite spinigers few and limited to anterior Branchiae are first present from about seg- segments; subacicular setae absent ...... ment 91 and occur irregularly on following ...... Marphysa brevitentaculata segments; they are simple filaments or some- Composite spinigers more numerous; subacicu- times divided. This agrees with Palola sicilien- lar setae present...... 4 sis (Grube) kmown from circum-mundane 4. Subacicular setae distally bifid; acicula dark to tropical seas, especially in coral reefs. Fauvel black; furcate setae conspicuous and of two (1932, p. 139) has given a diagnosis and kinds, coarse and fine-toothed in posterior synonymy. segments . Marphysa sanguinea Subacicular setae distally acicular; acicula yel- Lysidice tortugae Treadwell, 1921 low; furcate setae absent...... Marphysa aransensis Lysidice tortugae TREADwELL, 1921d, pp. 85-86, 5. Subacicular setae distally bifid; acicula pale figs. 298-304. brown or dark yellow; branchiae with few The holotype of Lysidice tortugae comes (2 or 3) filaments...... Macrphysa minima from Florida, and was associated with sponge. Subacicular setae distally entire; acicula dark It differs from other species of the genus nota- or black; branchiae with filaments more nu- bly for its very long, slender body, resembling merous and in pectinate arrangement...... a drilonereid in that respect. The prostomium ...... Marp hysa regalis is deeply bilobed and overlain by three slender, Marphysa orientalis Treadwell (1936) from filiform antennae. The two eyes are dark and China (not seen by me) resembles Marphysa crescentic in shape. The first two segments sanguinea, but the presence of subacicular have no parapodia, and the first segment is setae was not noted. only about a third as long as the second. The Marphysa sanguinea (Montagu), species is 1815 known only through its original dis- Nereis sanguinea MONTAGU, 1815, pp. 20-21, covery. pl. 2, pl. 3, fig. 1. 1956 HARTMAN: ANNELIDS 285 Marphysa viridis TREADXVELL, 1917b, pp. 264- On the dissected pharynx (which had not 265, p1. 2, figs. 13-18, p1. 3, figs. 1-2. been previously examined), the maxillary parts Marphysa nobilis TREADWELL, 1917b, pp. 265- are pale or yellow; I (forceps) are very long 266, pl. 3, figs. 3-9. and slender; II has five teeth on each side; The holotype of Marphysa nobilis is from III has seven on one, none on the other, side; Mangrove Key, Florida, June, 1915, and that IV has four and seven teeth, and V is a plate. of Marphysa viridis is from Boca Grande Key, The mandibular plates are now decalcified. Florida, July, 1915. Both are easily referable Marphysa aransensis differs from M. to the older, widely distributed Marphysa sanguinea by having subacicular setae that are sanguinea (Montagu). Branchiae are first pres- distally acicular, not bifid; furcate setae are ent from segments 24 and 22; the mailm absent instead of present; acicula are yellow, number of filaments is five in palmate-pectinate not dark. The maxillary formulas differ in arrangement; they are continued back to near that maxillae II had five teeth on a side instead the posterior end of the body. Dorsal cirri of of three or four. Earlier (Hartman, 1944b, branchial segments are proportionately very p. 126) I regarded the two as identical, without small. Acicula are dark to black and fade dis- having seen the type specimen from the Gulf tally. Parapodia have composite spinigers. Sub- of Mexico. acicular setae are distally bifid. Marphysa Marphysa aransensis is known only from the sanguinea is known from the east and west type locality at Aransas Pass, Texas. coasts of America, in warm littoral seas, and from cosmopolitan areas. Marphysa brevitentaculata Treadwell, 1921 Marphysa brevitentaculata TREiADwL, 1921d, Marphysa aransensis Treadwell, 1939 pp. 69-70, figs. 235-243, pl. 6, figs. 13, 14. Marphysa aransensis TRE LLw, 1939a, p. 5, The holotype specimen comes from Tobago, figs. 16, 17. British West Indies, taken in April, 1928. As The holotype comes from Aransas Pass, originally stated, it is very long, measuring Texas. The proportions of the single individual about 600 mm. long and 2 to 4 mm. wide, and agree with the description. I find the first consists of more than 800 segments. The pros- branchiae present on setigerous segment 24 as tomium is almost semicircular and has only a a single filament and so continued to 29 where very slight emargination at its frontal margin. double filaments occur. There are three fila- Prostomial antennae are very short. Anterior ments on segment 38 and the number increases parapodia are very small and inconspicuous. to four or five filaments in postmedian seg- The jaws of the holotype have been removed ments. The filaments are in unipinnate arrange- and are not in the vial. Another individual was ment, with all pinnae about equally long and dissected and found to have the following each about twice as long as the much thicker, formula: I (forceps) are long and falcate; II triangular dorsal cirrus. Branchiae are present has six teeth on the left and five on the right through a long region. Composite setae are side, with the largest tooth distalmost and de- spinigerous, with the appendage long and creasing in size basally; III has eight equally acutely pointed at the tip. Acicula are dark small crenulated teeth on the left, and none on yellow and occur singly or doubly in a para- the right side; IV has four small teeth like podium; they do not project from the fleshy those on III, on the left, and nine similar teeth lobe or they may be slightly emerging. Sub- on the right side; V is a small chitinized plate acicular setae are present in branchial seg- on each side. This differs from the original ments; they are yellow and occur singly in a account in that II has five teeth on a side, III ramus; they terminate distally in a blunt spine has six, IV has three and nine. and are weakly curved along their length. Parapodia have broadly rounded foliaceous Median and posterior segments have no post-setal lobes, especially in prebranchial and furcate setae. The position which would be anterior branchial segments. Anterior segments taken by such setae has a supra-acicular fascicle have composite spinigers with a short ap- of long, slender, slightly limbate setae that pendage; they are located in small fascicles in number 10 or more in a bundle. a subacicular position. Farther back they are 286 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 replaced by simple setae. Furcate setae are Marphysa languida TREADWELL, 1921d, pp. 73- present, though few in number, in branchial 75, figs. 257-258. segments. Subacicular setae are absent; their The holotype and only specimen of Marphysa position is taken over by simple, narrowly languida comes from Guanica Harbor, Puerto limbate, distally pointed setae. Acicula are dark Rico, taken in May, 1915. As originally stated, to black and occur singly in median and pos- the pharyngeal parts are missing and were not terior segments. described. The prostomium is deeply bilobed at Marphysa brevitentaculata resembles M. its frontal margin. Branchiae are first present mossambica (Peters) which also inhabits deep at about segment 60 and come to have at most burrows, has very short parapodial lobes, and two or three filaments; they are absent from becomes very long (to 600 mm.) (Crossland, about the last 50 segments. Acicula are dark 1903, p. 139). The latter, however, has a deeply yellow or pale brown at the base and number incised prostomium, and composite setae are about three in a parapodium in anterior seg- altogether absent. Other species with an un- ments, and one or two in median and posterior divided prostomium are M. mortenseni Monro segments. Subacicular setae are yellow, distally (1928, p. 88), M. strangulum (Grube, 1878), bifid, and occur singly in a ramus. Furcate setae M. bellii (Audouin and Milne-Edwards, 1834), are present, located in the upper end of the M. adenesis (Gravier, 1900), and M. macintoshi setigerous fascicle, in median and posterior seg- (Crossland, 1903). Marphysa brevitentaculata ments. Composite setae are falcigerous. These and M. macintoshi agree in having composite characters agree with those of M. minima spinigers and branchiae present on many seg- (Hansen), first described from Brazil. Another ments. The others named have composite falc- nearly related species is M. atlantica (Kinberg) gers and branchiae limited to a few anterior from the La Plata region in 30 to 40 fathoms. segments. Marphysa brevitentaculata differs It differs in that branchiae are first present from M. macintoshi in that the latter has sub- from about segment 20. As M. languida, this acicular setae, the former has none. has been further described and recorded from Marphysa brevitentaculata is known only western Mexico (Rioja, 1941, pp. 712-715, from the West Indies in littoral sands. pl. 5, figs. 6-15). Its geographic range thus in- Marphysa mossambica (Peters), 1854 cludes both coasts of tropical America. Eunice mossambica PETERS, 1854, p. 612. Marpliysa simplex TREAwERL, 1922a, pp. 151- Marphysa regalis Verrill, 1900 152, fig. 39, p1. 5, figs. 8-12. Marphysa regalis VERRILL, 1900, p. 636. The holotype specimen of Marphysa simplex Marphysa fragilis TREADWELL, 1911, pp. 2-5, comes from Fiji. It is characterized by having figs. 1-7. no composite setae; branchiae are pectinately The holotype of Marphysa fragilis comes divided; the prostomium is incised at its frontal from Loggerhead Key, Dry Tortugas, Florida, margin. Branchiae are first present from about taken in July, 1910. It is characterized for hav- segment 29 to segment 32, rapidly come to ing composite falcigers; subacicular setae are have four or five filaments each, and are con- acicular and dark. It agrees fully with M. regalis tinued on segments to the posterior end of the Verrill (Hartman, 1944b, p. 126). The species body. In anterior segments the acicula number is known from Bermuda and the West Indies. four or five in a ramus and are black; in pos- terior segments they diminish in number to Marphysa bellii oculata Treadwell, 1921 only one. These characters agree with those Marphysa bellii oculata TREADwEzI, 1921d, of Marphysa mossambica (Peters) as described pp. 61-64, figs. 201-211, pl. 5, figs. 13-14. by Crossland (1903, p. 139). Fauvel (1932, p. 142) has indicated its geographic distribu- This comes from Mangrove Key in Key tion in Indo-Pacific seas. West Harbor, Florida. It is unique in having both falcigerous and spinigerous composite Marphysa minima (Hansen), 1882 setae; subacicular spines are distally bifid. Nausicaa mzinima HANSEN, 1882, p. 8, pl. 2, Branchiae are very large and abrupt in their figs. 22-25. occurrence; they are present only on segments 1956 HARTMAN: ANNELIDS 287 10 to 22 where they have 10 or 11 pinnae each. Lumbrineris zonata (Johnson), 1901 The variety differs from the stem species for Lumbriconereis zontta JOHNSON, 1901, pp. 408- having, instead of lacking, prostomial eyes, al- 409, pl. 9, figs. 93-100. though Fauvel (1923a, p. 410) described them Lumbrinereis singularisetis TIEADWELL, 1931d, as present. It is possible therefore that there p. 1, figs. 1-3. is no specific distinction separating the variety The holotype of Lumbrinereis singularisetis from the stem form. Marphysa bellii (Audouin comes from Monterey, California. On the dis- and Milne-Edwards) is known from southern sected pharynx maxilla III has two teeth on and southwestern Europe (Fauvel, 1923a, p. each side, IV has a single tooth on each side. 410). Parapodial lobes of posterior segments are Paramarphysa teres Treadwell, 1922 short. This agrees fully with Lumbrineris zonata (Hartman', 1944b, p. 144) extensively Paramarphysa teres TREADWELL, 1922a, pp. 153- known from littoral zones of the northeastern 154, figs. 40-41, pl. 6, figs. 2-6. Pacific Ocean. This species comes from Samoa. Its resem- blance to P. orientalis Willey (1905, p. 283) Lumbrineris branchiata (Treadwell), 1921 Manaar is noteworthy. Okuda from the Gulf of Lumbrinereis branchiata TREADWELL, 1921d, (1937, pp. 287-288, figs. 32-33) concluded that 8, figs. 5, 6. the two differ in prostomial parts. In other re- pp. 94-95, figs. 333-343, p1. spects they seem to be similar. The holotype of Lumbrinereis branchiata comes from Buccoo Bay, Tobago, West Indies, LUMBRINERIDAE taken in April, 1918. The body is very long, 1926 slender, and resembles that of a drilonereid. A Lumbrineris similabris Treadwell, long posterior end is presumed missing from it. Lumbrinereis similacbris TREADWELL, 1926d, p. 5, The prostomium is conical, a little longer than figs. 6-10. wide, and has no eyes. Anterior parapodia are The single specimen of Lumbrinereis simi- very small and inconspicuous. Simple hooded labris comes from Bering Strait between King hooks are present from t-he first one and accom- Island and the two Diomedes. It is a large, panied with about an equal nunber of pointed fragmented specimen, resembling Lumbrineris limbate setae. Acicula are yellow and embedded. fragilis (Muller). Most setae and hooks are In median or perhaps later segments the pointed broken off close to the body, so that their pres- setae are completely replaced by simple hooded ence cannot be determined in anterior segments. hooks. Posterior parapodia have characteristic According to Berkeley and Berkeley (1942, fleshy, alate processes located behind and below p. 195) simple hooks are present from the the parapodial base; they agree with those seventh setiger. Acicula are dark brown to shown for Lumbrineris alata Hartmnan (1951, black and number to five in a ramus; setae are pl. 15, fig. 1) for specimens described from the dark amber to translucent brown. The post-setal Gulf of Mexico. In other respects, also, there lobe is broadly rounded, but in far posterior is agreement with L. branchiata, so that the two segments it elongates somewhat but continues must undoubtedly be regarded as the same. Far as a broadly rounded lobe. posterior segments are present in collections as On the dissected pharynx maxillae III and L. alata; they show that the post-setal lobe IV have each a single broad, short tooth; II elongates considerably to form a digitate lobe has four teeth on the right and four on the directed laterally. left side, with the teeth increasing in size Lumbrineris branchiata is known from the distally and the third and fourth teeth widely West Indies and the Gulf of Mexico. Lumbri- separated. Maxillary carriers are longer than conereis branchiata Fauvel (1943, pp. 22-24, broad. fig. 2) from the Gulf of California is a dif- Lumbrineris similabris is abundant off Alaska ferent species; it is unique in havring palmately in 12 to 15 fathoms and Vancouver Island, tufted branchiae near the base of parapodia Canada, in 23 fathoms (Berkeley and Berkeley, and the prostomium broadly depressed. The 1942, p. 195). specific name is a homonym. 288 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Lumbrineris candida (Treadwell), 1921 The holotype comes from Dry Tortugas, Lumbrinereis candida TXEADwEL, 1921d, pp. Florida, taken in July, 1914. The pharyngeal 96-97, figs. 344-350, pI. 8, figs. 7-9. parts have been dissected out and are missing. The holotype comes from Buccoo Bay, Anterior parapodia have composite hooded Tobago, West Indies. The body is very long hooks in which the articulation is complete. and slender, as first described. The prostomium Acicula are yellow. In posterior parapodia the is conical, slightly depressed, and about as long post-setal lobe does not elongate; these segments as broad. Anterior segments have simple hooded have simple hooded hooks without pointed hooks in addition to pointed setae. In posterior setae. The individual resembles Lumbrineris segments the post-setal lobe is digitate, with the latreilli Audouin and Milne-Edwards (Hart- lobe about as long as the prolonged parapodial man, 1944b, pp. 158-159, pl. 9, figs. 213-216) base; it extends distally beyond the tips of the except that the mandibles were described as hooded hooks. Embedded acicula are yellow. very broad and short, and maxillae as nearly The maxillae are dentate, thus not as originally edentate. described; II has about four teeth on each side, III has two short teeth on each side, and the Lumbrineris treadwelli, new name last is a broad thin plate. Lumbriconereis maculata TREADWELL, 1901b, Lumbrineris candida belongs to the L. pp. 198-199, figs. 4244. tetraura group (Hartman, 1944b, pp. 147- Lumbrineris maculata TREADWELL, 1942a, pp. 150), from which it may be distinguished by 119-120, figs. llh, 14d, e. its greatly attenuated form. It is known only This species comes from Puerto Rico and from the type locality. Florida (Treadwell, 1921d, p. 103). It is re- described in Hartman Lumbrineris inflata (Moore), 1911 (1942a, pp. 119-120). The preoccupied specific name is replaced by Lumbrinereis inflata MooRn, 1911, pp. 289-291, L. pl. 19, figs. 128-132, pl. 20, figs. 133, 134. treadwelli, new name. Lumbrinereis cingulata TREADWELL, 1917b, p. 263, pl. 2, figs. 7-12. ARABELLIDAE Lumbrinereis ceruicalis TREADWELL, 1922b, p. Arabella iricolor (Montague), 1804 176, figs. 14-21. Nereis iricolor MONTAGU, 1804, p. 82. The holotype of Lumbrinereis cingulata Arabella dubia TREADWELL, 1922a, pp. 160-161, comes from Bermuda in crevices of coral rock fig. 52, p1. 7, figs. 11, 12, p1. 8, figs. 8, 9. Not Han- near low-water mark; that of Lumbrinereis sen, 1882. cervicalis is from Friday Harbor, Washing- .Arabella setosa TREADWELL, 1921d, pp. 113- ton, taken in February, 1920. The first is a 114, figs. 421-424, p1. 9, figs 10-11. slender, nearly cylindrical individual measuring The holotype of Arabella dubia comes from about 37 mm. long and 1 mm. wide. Acicula Pago Pago, Samoa; that of Arabella setosa is and setae are yellow or nearly colorless. An- from Buccoo Bay, Tobago, West Indies. The terior parapodia have pointed limbate setae and pharyngeal armature has been removed from one or two hooded hooks that are incompletely the former and is missing. Most setae are or imperfectly composite; posterior hooded broken off. The prostomium has four eyes hlooks are entire. Posterior parapodial lobes are arranged in a transverse row. Parapodial lobes short and not noticeably different from those of and other parts that have been described agree median parapodia. It is believed identical with with those of Arabella iricolor (Montagu). Lumbrinteris inflata (Moore). The specimen The specimen labeled Arabella setosa has from Friday Harbor is much larger but agrees post-setal parapodial lobes that are in all longer than details with Lumbrineris inflata (see Hart- is typical, but in other respects it agrees with man, 1944b, p. 161, for synonymy and distribu- A. iricolor, which is known from cosmopolitan tion). areas in littoral seas. Lumbrineris paucidentata (Treadwell), 1921 Arabella semimaculata Luimbrinereis Paucidentata TREADWELL, 1921a, (Moore), 1911 pp. 99-100, figs. 357-364, pl. 9, figs. 1-4. Aracoda semimaculata MOORE, 1911, pp. 295- 297, pl. 20, figs. 143-149. 1956 HARTMAN: ANNELIDS 289 Arabella pacifica TREADWELL, 1941c, p. 23, figs. setae project directly from the body wall. Para- 18-21. podia are first distinct from about segment 30. The holotype of Arabella pacifica comes Acicula are very large, thick, and project from from Sihuantanejo, Mexico, in coral. It is a parapodia for a considerable distance. On the moderately large individual of Arabella semi- maxillary pieces, the forceps have teeth at the maculata (Moore). Posterior segments have base; II has four teeth on a side. Mandibles prolonged post-setal lobes. Prostomial eyes are are present and black. Drilonereis logani Cross- hardly visible and seem to be absent. The species land (1924, pp. 64-70, figs. 80-88) from east- is known from southern California to western ern Africa shows remarkable resemblance and Mexico in littoral zones. may be the same. Arabella sp. Drilonereis planiceps (Grube), 1878 Aracoda attenuata TREADWELL, 1911, pp. 5-6, Arabella planiceps GRuBE, 1878, pp. 174-175, figs. 8-11. pl. 8, figs. 4-4b. Drilonereis attenuata TREADWELL, 1921d, pp. Drilonereis paucidentata TREADWELL, 1922a, 107-108, figs. 395-399, p1. 9, figs. 6-9. p. 162, fig. 56, p1. 7, figs. 16, 17, p1. 8, fig. 11. The holotype comes from Dry Tortugas, The holotype of Drilonereis paucidentata F?lorida. Parapodia have no projecting acicula (labeled D. pacifica in the records of the Mu- as is typical for species of Drilonereis. On the seum) comes from Suva, Fiji, taken in April, maxillae the forceps are dentate only at the 1920. Another specimen, which was used for base and not along the entire cutting edge as the description, is from Pago Pago Harbor, in Notocirrus. It is, therefore, not the same as Samoa. On the holotype the pharyngeal anna- Arabella attenuata Treadwell (1906) which is ture has been dissected out and is missing. referred to Notocirrus (Hartman, 1944b, p. The prostomium is greatly depressed, broadly 176). The prostomium is oval and has a deep rounded in front, and has no eyes. The first middorsal longitudinal groove and eye spots two segments have no parapodia. Farther back at its posterior margin. Anterior parapodia are they are very small and in posterior segments very small. In far posterior segments the the post-setal lobe elongates about as originally post-setal lobe does not elongate. The specimen shown. Acicula are yellow; they extend from may be an immature stage of another species the parapodium at their distal ends and are of Arabella. The specific name is preoccupied. accompanied by two or three broadly limbate, geniculate, distally pointed setae without dentic- Arabella sp. ulations at the cutting edge. In addition, there Drilonereis pinnata TREADWELL, 1921d, pp. 110- is a thick fascicle of many slenderer setae 111, figs. 412-413, p1. 8, fig. 11. which terminate in a long, very slender tip This species comes from Buccoo Bay, To- (shown by Crossland, 1924, p. 61, for Drilo- bago, West Indies, taken in April, 1918. Para- nereis major, and by Grube, 1878, pl. 8, for podia have no projecting acicula, and on the Arabella planiceps). Mandibles are absent. maxillae the forceps are falcate. It is there- Crossland (1924, p. 63) suggested the pos- fore a species of Arabella. Anterior parapodia sible identity of his Drilonereis major with are not reduced. In posterior segments the D. planiceps (Grube). However, thfe similarities post-setal lobe is not noticeably prolonged. On of D. planiceps from the Philippine Islands, the jaw pieces, II has five teeth on a side, III D. paucidentata from Fiji and Samoa, and has four on a side, and IV has two on each D. major from Suez and Zanzibar are so great side. This may be a representative of Arabella that there can be little doubt of their identity. iricolor (Montagti) (see above). All are very long, slender forms and have a depressed, semicircular prostomium without Drilonereis lumbricus Treadwell, 1922 eyes. Anterior parapodia are reduced; far Drilonereis lumbricus TREADWELL, 1922a, pp. posterior parapodia have post-setal lobes only 161-162, figs. 53-55, pl. 7, figs. 13-15, pl. 8, fig. 10. slightly elongated. Mandibles are usually ab- The holotype is from Fiji. Anterior para- sent. Maxillary plates are about the same. podia are reduced so as to appear absent; the Parapodial setae are of two kinds, one or two 290 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 in a parapodium are shorter, geniculate, distally are continued on all successive segments to pointed, and have a smooth cutting edge; many the end of the body. In this respect the species others terminate in a very long, slender tip. is unique. Thoracic neurosetae are largely long An individual reported from Bikini Atoll, pointed setae in full fascicles; they are ac- northern Marshall Islands (Hartman, 1954c, companied with a few acicular spines in an p. 634), is believed conspecific. The geographic inferior position; when perfect they have a range thus includes a large area in the Indo- distal hood. Pacific regions. Furcate setae are present in abdominal noto- the two tines are Drilonereis spatula podia; unequally long. Ab- (Treadwell), 1911 dominal neuropodia have four or five yellow Aracoda spatula TREADWELL, 1911, pp. 6-7, figs. acicula that emerge from the neuropodial lobe 12-14. for a considerable in a Drilonereis spatula TREADWELL, 1921d, pp. 108- distance, spreading 109, figs. 400-406. fascicle. They are accompanied by many distally Drilonereis brunnea TREADwELL, 1921d, pp. pointed, longer setae. 111-112, figs. 418-420. Naineris mutilata differs from N. laevigata in As Aracoda spatula (Grube) having branchiae present from the this was collected from instead of a a reef near Fort Jefferson, Florida, in second, later, segment. Thoracic June, neuropodia have a broadly foliaceous post-setal 1909. The body is slender, about 170 mm. long lobe and and 2.5 mm. mostly pointed setae instead of a wide at most. The prostomium is smaller lobe and setae of another kind. broadly rounded, longer than wide, has a mid- Naineris mutilata is from the dorsal longitudinal groove, and has no eyes. known only Maxillae I (forceps) are falcate; II has seven type locality. teeth on a side. Anterior parapodia are very Haploscoloplos robustus (Verrill), 1873 small. Acicula are yellow, heavy, and project Anthostoma robustum VERiLL, 1873, pp. 597- for a considerable distance from the parapodial 598, pl. 1, fig. 76. lobe. In posterior segments the post-setal lobe Scoloplos rufa TREADWELL, 1941b, p. 1, figs. 1-6. is elongate and erect. Drilonereis spatula is be- The holotype of Scoloplos rufa comes from lieved to include D. brunnea which comes from Offats Buccoo Bay, Bayou, Galveston, Texas. It is a slender, Tobago, West Indies. It is not immature Branchiae are first known outside West Indian localities. specimen. present though small on setigerous segment 23 and Drilonereis sp., juvenile gradually increase in size from about segment 26, or in abdominal segments where they are Drilonereis similis TREADWELL, 1921d, p. 111, and erect. An figs. 414-417, p1. 8, fig. 12. large interramal cirrus is pres- ent, the first from posterior thoracic segments, This comes from Buccoo Bay, Tobago, West and it is continued for about 12 anterior ab- Indies. It is perhaps an immature stage of dominal segments. It was originally described Drilonereis spatula (see above). as a "much smaller lobe . . . on the dorsal neuropodial surface," but not illustrated. ORBINIIDAE Branchiae are simple throughout, not double Naineris mutilata Treadwell, 1931 as originally shown (Treadwell, fig. 5). The Nainereis mutilata TREADWELL, 1931d, pp. 5-6, illustration shows both branchiae of a segment, figs. 13-18. and the midpoint of the dorsum is shown be- tween the bases of the paired branchiae. Ab- This comes from Montego Bay, Jamaica, dominal West Indies. It is broken in several pieces neuropodia have long, transverse but ridges, not divided lobes; they are like those is believed to represent parts of a single indi- for vidual. The prostomium is broadly rounded at Haploscoloplos robustus (Hartman, 1951, its pl. 21, figs. 4-6), and not as for H. anterior margin; there are no visible eyes. with fragilis It is followed by a single smooth ring or peris- (Verrill), which I had earlier thought tomium. Branchiae are present from the sec- Scoloplos rufa identical (Hartman, 1951, p. ond setigerous segment and already large; 76). they The geographic distribution of Haploscolop- 1956 HARTMAN: ANNELIDS 291 los robustus is about the same as that of H. The holotype comes from east of Cocos fragilis (Verrill); both occur along the east- Island, Pacific Panama. Length of 80 segments ern shores of the United States and in the is 18 mm. Branchiae are first present from the Gulf of Mexico. second setiger, and hooded hooks are present in both notopodia and neuropodia. This is SPIONIDAE therefore referred to Nerine Johnston. The Nerinides agilis (Verrill), 1873 prostomium is rounded in front and has four Nerine agilis VERRILL, 1873, p. 600. eyes in a transverse row; they are located on Nerine minuta TREADWELL, 1939a, p. 5, figs. the prostomial ridge, not on the peristomium 18-20. as originally shown (Treadwell, fig. 54). The Nerine agilis HARTMAN, 1951, P. 81. lateral margins of the peristomium are elevated This species is here referred to Nerinides as a pair of folds. Mesnil instead of Nerine Johnston because it Branchiae are fused to the notopodial lobe lacks hooded hooks in the notopodia, whereas for the entire length of the lobe. Their general Nerine has such hooks in both notopodia and outlines are about as first shown (Treadwell, neuropodia of posterior segments. In Nerinides fig. 55). Hooded hooks are first present at agilis the neuropodial hooks are distally biden- about segment 28 in both notopodia and neuro- tate, with the main fang at an obtuse angle to podia and are present only in median segments; the shaft. Branchiae are first present from the they are distally bidentate. Tufts of white second setigerous segment. On the prostomium setae first appear in neuropodium 37, and in the four eyes are in a straight transverse row. notopodium 45; they are conspicuous where The species is known from New England south present and distinguish this subspecies from into the Gulf of Mexico, in littoral sands. the stem, Nerine cirratulus (delle Chiaje). The prostomial eyes have the arrangement shown Boccardia sp. for Nerine cirratulus saipanensis Hartman Polydora californica TEADW.ELL, 1914b, pp. (1954a, p. 230, fig. 2a) but the latter has an 203-204, p1. 12, figs. 23-29. Not Fewkes, 1889. acutely pointed prostomium. A paratype specimen labeled as coming from Nerine cirratulus hirsuta is known only from California has branchiae on setigerous seg- Cocos Island. ments 2 to 4 and from 7, continued back on many segments; this specimen is therefore a CIRRATULIDAE representative of Boccardia and agrees with B. proboscidea Hartman (1940b, pp. 382-387). Tharyx multifilis Moore, 1909 The original account was based perhaps on Tharyx multifilis Mool., 1909, pp. 267-268, another specimen which is not known to exist; pl. 9, fig. 43. it was said to have come from an unknown Cirratulus i-nhamatus TRiADwaLL, 1937b, p. 153, locality, taken from a tangled mass of tubes pl.2, fig. 25. apparently constructed by the Polydora. Such The type of Cirratulus inhamatus comes tubes are not constructed by Boccardia probo- from east of Cedros Island, Baja California, scidea Hartman, which inhabits vertical bur- Mexico, in 38 to 40 fathoms. It now lacks all rows in shaley rocks. tentacular structures, including the larger The specific name of Polydora californica paired palpi which characterize Tharyx Treadwell is a homonym of Spio californica Webster, but their bases are visible. The pros- Fewkes, 1889, which has been referred to tomium has no eyes. Notopodia and neuro- Polydora, sensu latior (Hartman, 1940b, pp. podia have only slender, capillary setae of one 386-387). It seems best, therefore, to drop kind in all segments; in postmedian segments Polydora californica Treadwell from the list they number 20 to 30 in a fascicle and are of valid species. arranged in a double transverse series. Setae are directed laterally in long, silky, flowing Nerine cirratulus hirsuta (Treadwell), 1928 tufts. Notopodia and neuropodia of a side are Spio hirsuta TREADWELL, 1928a, pp. 478-479, separated from each other by a small interval. figs. 54-57. The bases of the lateral branchiae, to be seen 292 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 on a few segments, are immediately above the setae to the end of the body. Notopodial spines notopodial ridge. are first present a little farther back and in- in a The first, or palpal-bearing, segment shows crease in number to two or three fascicle; setae. the bases of a pair of slender tentacular fila- they are also accompanied with capillary ments. This segment is followed by the first Lateral branchiae, in middle and posterior at a setigerous segment, but the two are not clearly segments, are inserted dorsolaterally, point separated from each other except by lateral above the notopodium for a distance exceeding first setiger has biramous para- that between the branches of a parapodium. grooves. The recorded from podia and the base of a slender filament im- Cirriformia punctata was first more ex- the notopodial elevation. These West Indies and is known from mediately above the America. characters agree with those of Tharyx multifilis tensive tropical regions of eastern California and Moore, known from southern 1877 western Mexico. Cirriformia chefooensis (Grube), Cirratulus chefooensis GRTBE, 1877, pp. 50-51. Cirriformia semicincta (Ehlers), 1905 ?Cirratulus branchiatus TREADWELL, 1936a, pp. Cirratulus semicinctus EHLERS, 1905, pp. 292- 276-277, fig. 20g. 293, p1. 9, figs. 11-14. The single type specimen of Cirratulus Audouinea maculata TREAnwEL, 1929d, p. 10. branchiatus, deposited in the United States Na- The holotype of Audouinea maculata comes tional Museum, comes from the vicinity of from Berbera, British Somaliland, tropical east- Amoy, China. I have not seen it, but am ques- ern Africa. It agrees fully with Cirriformia tionably referring it to the older C. chefooensis semicincta (Ehlers), first described from (Grube) because of the agreement in the Hawaii and more widely recorded from cir- descriptions of both. Tentacular structures cumtropical regions. Notable features include arise from the dorsum of the fifth or sixth the presence of dorsal tentacles on the fourth setigerous segment. Posterior parapodia have setigerous segments; notopodia and neuropodia transverse rows of curved acicular spines that have acicular spines; the pigment pattern of alternate with slender pointed setae. The geo- dispersed dark spots over the surface persists graphic distributions agree. through preservation. Cirriformia filigera (delle Chiaje), 1841 Cirriformia punctata (Grube), 1859 Cirrathulus filigerus DELLE CHIAJE, 1841, p. 85. 64, Cirratulus punctatus GRUBE, 1859, p. 107. Audouinea pygidia TREADWELL, 1936b, p. Cirratulus nigromaculata TREADwEL, 1901b, figs. 22, 23. p. 204, fig. 66; 1939b, p. 273. The type of Audouinea pygidia comes from Cirratulus niger HARTMAN, 1939a, pp. 17-18, Nonsuch Island, Bermuda, in tidal pools. Dorsal fig. 31. tentacles are inserted on the sixth setigerous The body is grayish yellow flecked with segments. Acicular spines are first present in black; the dorsal tentacles are transversely the neuropodium of segment 18; they number barred with yellow and black, and the lateral two or three in a row. Farther back they are branchiae are lightly barred or plain. Dorsal thicker, distally more curved, and occur singly tentacles are attached on about the third in a ramnus. Notopodial spines are first present setigerous segment, where they form a crowded from about the anterior third of the body, paired bundle, each just above the insertion where there are two to four in a series. These of the lateral branchia. This species is thus characters are those of Cirriformia filigera referred to Cirriformia Hartman instead of (delle Chiaje) earlier recorded from southeast- Cirratulus Savigny, in which dorsal tentacles ern United States (Hartman, 1945, p. 35). Neuropodial are on the first setigerous segment. 1808 spines are first present in about the sixth Cirriformia tentaculata (Montagu), neuropodium; the first are single in a ramus, Terebella tentacuiata MONTAGU, 1808, pp. 110- slender pointed setae; lll, pl. 6, fig. 2. accompanied with 1931f, pp. 1-2, farther back they increase to three in a Cirratulus pallidus TREADWELL, fascicle and are accompanied with a few pointedI figs. 1-3. 1956 HARTMAN: ANNELIDS 293 Cirratulus multicirratus Ta:EADwELL, 1936b, pp. is from Nonsuch Island, Bermuda, in 10 to 63-64, figs. 34, 35. 35. feet in sand. It is now dark so that the ar- The holotype of Cirratulus pallidus comes rangement of lateral eye spots is not easily from Montego Bay, Jamaica, West Indies; seen. In the original description they are noted that of Cirratulus multicirratus is from St. as 21 pairs, first present from between seg- Davids Island, Bermuda, on mud flats. Both ments 5 and 6, and continued back to between are species of Cirriformia, because dorsal segments 25 and 26. This is thus referred to tentacles are present on a segment behind the Armandia Filippi. The prostomium terminates first setigerous one. Posterior parapodia have in a slender palpode. Large nuchal organs are yellow, distally curved acicular spines in trans- in front of the mouth which is in line with the verse series. Lateral branchiae are inserted a first pair of setigerous bundles. Branchiae are short distance above the notopodial base. The present from the second segment and con- distribution for the Atlantic shores of North tinued to the end of the body; they number 30 America is given in Hartman (1945, p. 35). pairs. The pygidial funnel is broken and its long median filament lost; there is a semicircu- Cirriformia hawaiensis, new name lar midventral flange at the base to which a Audouinia branchiata TREADWELL, 1943d, p. 1, long filament might have been attached. The figs. 1-3. dorsal distal margin of the funnel has a long, The holotype of Audouinia branchiata is longitudinal fissure with crenulate margin and from Pearl Harbor, Hawaii. It is unique in perhaps was the base of terminal papillae. One having a prosomium that is dusky at its frontal can distinguish about 11 pairs of crenulations. margin; this region is continued laterally to These features agree with those of Armandia the paired ocular patches, immediately above maculata (Webster) first described from Ber- the oral groove. Anterior parapodia have only muda (see Hartman, 1942a, pp. 129-130, fig. pointed setae. Distally curved acicular spines 14a). are first present in neuropodia from segment 23. Notopodial spines first appear much farther Euzonus (Thoracophelia) mucronata back and are never so thick or so curved as (Treadwell), 1914 neuropodial ones. In posterior median seg- Ophelina mucronata TREADWELL, 1941b, pp. ments the neuropodial spines number five or 218-219, pl. 12, figs. 37, 38. six in a transverse row; they alternate with This species has been earlier referred to slender pointed setae and are dark amber in Thoracophelia Ehlers (Hartman, 1938c, p. color. In posterior segments their number 107). Thoracophelia is more conveniently re- diminishes to two or three in a series. Lateral garded as a subgenus of Euzonus Grube, 1866. branchiae are inserted above the notopodial A generic diagnosis follows. The body con- ridge; in posterior segments their origin is sists of three regions: a cephalic, a thoracic, progressively higher so as to be dorsolateral. and an abdominal. The cephalic region is set Cirriformia hawaiensis differs from other off from the thorax by a muscular constriction species of the genus for its unique prostomial behind the second setigerous segment. The parts. The specific name is changed because thorax is separable from the abdomenl by a branchiatus Treadwell, 1943, is preoccupied by pair of prominent glandular ridges which are Cirratulus branchiatus Treadwell, 1936, which part of the thorax; it consists of eight setiger- is also a species of Cirriformia (see above). ous segments. The abdomen is slenderer, con- sists of 28 or 29 segments, and tapers to a OPHELIIDAE pygidial area. Total number of segments is Armandia maculata (Webster), 1884 about 38 or 39. The prostomium is acutely Ophelina maculata WEBSTER, 1884, p. 322, pl. pointed and merges in an inflated part com- 11, figs. 54-55. prising the first two segments. The first or Ammotrypane bermudiensis TREADWELL, 1936b, peristomial segment has the oral aperture on p. 60. its ventral side, a pair of nuchal organs at the The holotype of Ammotrypane bermudiensis sides and the first setal tufts. 294 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 The thorax includes eight biramous seg- Polyophthalmus pictus (Dujardin), 1839 ments; paired sensory organs are located be- Nais picta DUJARIIN, 1839, p. 293, pl. 7, figs. tween the rami. The abdomen has a deep mid- 9-12. ventral longitudinal groove along most of its Polyophthalmus incertus TREADWELL, 1936b, length. Branchiae are present on a limited pp. 61-62, figs. 27-29. number of segments. The posterior end termi- Polyophthalmus papillatus TREADWELL, 1943c, nates in a pygidium with a large, triangular p. 3, figs. 19, 20. midventral lobe and many pairs of lateral fila- The of ments. holotype Polyophthalmus incertus The genus is conveniently recognized comes from Nonsuch Island, Bermuda, taken for two subgenera: Euzonus, sensu stricto, at the surface at of and Thoracophelia Ehlers. In the first night; that Polyophthalmus the papillatus comes from Lobito, Africa. Both are branchiae are pectinately divided; in the sec- referable to the ond they are easily widely known P. pictus bifurcated. (Dujardin). In P. incertus the setae of the last few segments are considerably prolonged. KEY TO SPEcIEs oF Euzonus GRUBE, EMENDED In P. papillatus I find 28 setigerous segments; 1. Branchiae deeply bifurcated...... the anal papillae have fallen off but their bases ...... Subgenus Thoracophelia, 2 can be distinguished. I would say that the Branchiae pectinately divided or secondarily everted proboscis is labiate, with a deep mid- dichotomously branched...... ventral groove, not globular as originally ...... Subgenus Euzonus, 4 stated. 2. Each branch of the bifurcation without lateral Polyophthalmus pictus has a known geo- processes ...... 33 graphic range that includes Bermuda and Each branch of the bifurcation with lateral western processes ...... Euzonus Africa. (Thoracophelia) williamsi (Hartman) 3. With 20 pairs of branchiae; anus with five pairs FLABELLIGERIDAE of dorsal papillae. Eusonus (Thoracophelia) furcifera Ehlers Semiodera cariboum (Grube), 1856 With 18 pairs of branchiae; anus with seven Siphonostomum cariboum GRUBBE, 1856, pp. 108- pairs of dorsal papillae...... Euzonus 109. (ThoracoPhelia) mucronata (Treadwell) 4. Semiodera glabra TREADWELL, 1928a, pp. 479- Branchiae unipectinately branched ..5...... 5 Branchiae pectinate at the base and dichoto- 480, figs. 62-65. mous more distally...... d 6 The type of Semiodera glabra comes from 5. Branchiae number 15 pairs...... southeast of Islands in 15 feet. The Euzonus (Euzonus) dillonensis (Hartman) Galapagos Branchiae number 19 pairs...... large tongue-like process, characteristic of .. .Euzonus (Euzonus) ezoensis (Okuda) Semiodera Chamberlin, is fully everted. The 6. Branchiae number 17 pairs...... epithelial covering of the body is largely ...... Euzonus (Euzonus) arcticus Grube sloughed off, so that the body appears nearly Branchiae number 15 pairs...... smooth. Small bits of the papillated cover are .....Euzonus (Euzonus) yasudai (Okuda) present about the parapodial bases and near the end. The everted oral Euzonus Grube has been emended by posterior organ shows both the bases of tentacular filaments, strewn over Augener (1912, pp. 176-178) and Annenkova most of the lower (1935, pp. 235-236). The or ventral side of the organ, latter regarded except for a median bare ridge or stripe. Its Thoracophelia Ehlers as identical with Euzonus lateral Grube. The group is at margins are crenulated as shown by present limited largely Ehlers (1887, pl. 42). This agrees fully with to the North Pacific Ocean, except for E. Semiodera cariboum (Grube). (E.) arcticus Grube, which is Arctic. The The anatomy and biology of type locality, originally given as Gala- Euzonus pagos Islands, was changed to Haiti by Tread- (Thoracophelia) mucronata (Treadwell) have well Semiodera been described by McConnaughey and Fox, (1929d, p. 9). cariboum was 1949. first described from St. Croix, West Indies, in 6 fathoms. Ehlers (1887) reported it from 1956 HARTMAN: ANNELIDS 295 Key West, Florida, in 1 to 2 fathoms. MALDANIDAE Stylarioides capensis americana Monro (1928, Asychis fimbriata (Treadwell), 1934 pp. 96-97) from Taboga, Panama, shore under MaldaneUa fimbriata TREADwwLL, 1934a, pp. -9, stones, may refer to the same species. pl. 2, figs. 22-24. Stylarioides glabra Treadwell (1901b, p. 208) from Puerto Rico, is also perhaps refer- The holotype is at the United States National able to Semiodera Chamberlin, as it is said to Museum and comes from off southern Puerto have the unique oral process. The type speci- Rico in 80 to 100 fathoms. I regard it as a men is in the United States National Museum. species of Asychis Kinberg rather than Malda- Its distinction from Semiodera cariboum nella McIntosh; the former is characterized (Grube) is not possible from the meager de- by having an anal plaque, the latter by having scription. none. MaldanelUa fimbriata is said to have 20 setigerous segments with a collar at the frontal Pherusa dubia (Treadwell), 1929 margin of the first segment; the anal plaque is bilobed at the dorsal margin and has a few Stylarioides dubius TEADwwLL, 1929d, pp. 9-10, lateral fringes. The first four segments are figs. 25-27. short, the next five longer, and the last are A paratype collection comes from Dry again shorter. As the cephalic plaque is shown Tortugas, Florida. There are three small in- without conspicuous nuchal ridges, the species dividuals with proportions as first described. seems to belong to Asychis Kinberg and not In general appearance they resemble Pherusa Maldane Grube. It shows resemblance to inflata (Hartian, 1952, p. 72) but differ in Asychis collariceps (Augener, 1906, pp. 163- the character of setae and oral tentacles. The 167, pl. 6, figs. 114-120), also from West cephalic cage consists of a few very long, Indian localities, in 169 to 395 fathoms, but slender, distally pointed setae which are weakly the anal plaques are not the same, according transversely barred; most appear smooth and to the illustrations. Asychis fimbriata (Tread- translucent. The bars are widely spaced, so well) has not been recorded outside its type that each bar is about twice as long as wide, locality. or longer. The anterior end of the body is obliquely truncate, with the ventral or oral end Maldane cristata Treadwell, 1923 the more distally prolonged. surrounding the prostomium Maldate cristata TREADWEL, 1923b, pp. 9-10, The appendages figs. 5-8. include four pairs of long, slender tentacles Maldane carinata MooRm, 1923, pp. 233-235. inserted on a semicircular ridge over the pros- tomium, with the lateralmost about in line As M. cristata this comes from off Cape San with the bases of the paired palpi. A fifth pair Lucas, Baja California, Mexico, in 630 fathoms of tentacular bases is visible between the outer (Albatross Station D 5698). As M. carinata row and the outer base of the palpi. The bases Moore it was taken off San Clemente Island, of the uppermost or medial pair are largest, in 654 to 704 fathoms (Albatross Station 4405), and there is gradual decrease in size lat- off Santa Rosa Island, in 31 to 45 fathoms erally. (Albatross Station 4431), and off Point Pinos Parapodia are biramous and have notosetae Lighthouse in 1073 fathoms (Albatross Station and neuropodial hooks. The first few segments 4547), California. have slender, long, distally pointed setae, which The margins of cephalic and anal plaques were originally overlooked, and a fascicle of are smooth. The body attains a length of about pointed neurosetae. In typical parapodia the 130 mm. and a width of 2.5 mm. On the cephalic ventral hooks are coarse, yellow, and number plaque the crest is narrow, elevated, promi- three or four in a transverse row; they are nent, and extends lengthwise along most of the accompanied with slender, hair-like setae. Each length. The first two setigerous segments have hook is simple, distally curved, and tapers to thick body walls; farther back they are thin- a blunt tip. Pherusa dubia is known only from ner. The first setiger has only slender setae; Florida. uncini are present from the second one. Un- 296 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 cinial rows are short at first and much longer to 743 fathoms. Grossly examined, these in- farther back. dividuals resemble Melinna Malmgren, but In addition to the localities named above, they have no heavy nuchal spines, and a very this form has been described, as Maldane large median oral tentacle is conspicuously ex- cristata (Monro, 1937, pp. 306-307, fig. 23), tended forward in front of the prostomium. from the Gulf of Oman, Arabian Sea, in 195 Branchiae number four pairs; all are similar, meters. tapering cylindrical. Thoracic and abdominal Euclymene corallicola uncini are short, avicular, with three or four (Treadwell), 1929 teeth in a single row, the largest tooth at the MaldanelUa coraUicola TiEADIwrL, 1929d, p. 8, base. figs. 21-24. The similarity of these features to those of The single specimen comes from Loggerhead Sosanopsis armipotens Moore (1923, pp. 215- Key, Dry Tortugas, Florida, in coral rock. It 216, pI. 18, figs. 26-29) from off Santa Cata- measures about 100 mm. long; greatest width lina Island in 2228 fathoms, red mud, is at is about 4 mm. at the fifth segment. The body once obvious and suggests their identity. is abruptly narrower at the ninth segment and Furthermore, Melinna monocera Augener remains so to the posterior end. There are 19 (1906, pp. 177-180, pL. 6, figs. 121-125) from setigerous segments and two ante-anal ones. off the West Indies in 116 and 170 fathoms The anal funnel has 29 marginal lobes which is nearly allied. are about equally small but some are irregu- Annenkova (1931, pp. 269-270, fig. 1) has larly bifid and others broken off. The cephalic characterized the genus Melinnexis and re- plaque is distinct and has a raised median viewed some of the literature. ridge which extends forward to form the prostomial lobe. Nuchal ridges, not previously Asabellides sibirica (Wir6n), 1883 described, are visible as a pair of slightly raised Sabellides sibirica WIR*fN, 1883, pp. 418-419. ridges parallel to the median one; they extend Neosabellides alaskensis TR:J3ADw, 1943e, pp. about halfway through the cephalic plaque. No 1-2, figs. 1-5. prostomial ocelli are visible. The dorsal margin The holotype of Neosabellides alaskensis of the cephalic plaque is entire, not crenulate comes from Nash Harbor, Nunivak Island, as in Euclymene coronata Verrill; there are Alaska, taken in July, 1937. Over-all length is two deeply emarginate notches at the sides. about 18 mm., width 1 mm. The anterior The first three setigerous segments are simi- margin of the prostomium is prolonged be- lar to one another in that each neuropodium has cause of the everted condition of the oral a single large, yellow, distally curved spine tentacles. The bases of the four pairs of projecting from the neuropodium. More pos- branchiae are like those of Asabellides sibirica terior neuropodia have rows of rostrate yellow (Wiren) (see Annenkova, 1929, pp. 494-495, hooks. These characters are those of Euclymene pl. 38, figs. 50-51, pl. 39, figs. 60-65, for Verrill. Euclymene corallicola (Treadwell) ap- illustrated diagnosis), with three pairs form- proaches E. grossa newporti Berkeley and ing a transverse series and a fourth pair in- Berkeley (1941) described from southern Cali- serted behind the others at the middle of each fornia. series. The paired nephridial papillae are visible at the middorsum, behind the innermost branch- AMPHARETIDAAE ial bases. The thorax consists of 14 setigerous Melinneis tentaculata (Treadwell), 1906 segments; uncini are present from the third Terebellides tentaculata T1READWELL, 1906, p. segment. The abdomen consists of about 22 1177, fig. 72. segments. In abdominal parapodia, the dorsal cirri are small and inconspicuous. Paratypes of Terebellides tentaculata in the The synonymy is believed to include, in American Museum of Natural History origi- addition to Neosabellides alaskensis: Asabel- nate from Hawaii. They agree fully with the lides orientalis Annenkova, 1929, from the brief original account and may have come from northwestern Pacific Ocean (fide Annenkova, the original collection, taken off Hawaii in 278 1938, p. 201), and Pseudosabellides littoralis 1956 HARTMAN: ANNELIDS '297 Berkeley and Berkeley, 1943, from Ungava, Loimia minuta TREADWELT., 1929d, pp. 10-11, Hudson Bay, and western Canada. figs. 28-30. Asabellides sibirica may be considered a com- The holotype of Loimia minuta comes from mon Arctic intertidal species. It constructs a Dry Tortugas, Florida. It agrees fully with L. fragile, loose-fitting tube, two or three times medusa (Savigny) as revised by Fauvel (1914, as long as the occupant, coated with sand and pp. 145-146, pl. 7, figs. 6-9) and reported from mud particles (Berkeley and Berkeley, 1943). other West Indian localities by Monro (1928, pp. 99-100; 1939, pp. 347-348). The species Ampharete arctica Malmgren, 1866 has a cosmopolitan distribution in warm seas. Ampharete arctica MALMGREN, 1866, pp. 364- 365, pl. 26, fig. 77. Thelepus crispus Johnson, 1901 Ampharete brevibranchiata TREADWELL, 1926d, Thelepus crispus JOHNSON, 1901, p. 428, pl. 17, pp. 6-7, figs. 11-14. figs. 175-178. The holotype of Ampharete brevibranchiata Streblosoma magna TREADwELL, 1937b, pp. 155- comes from Bering Strait, betwveen King 156, figs. 26,28. Island and the two-Diomedes. It is a moderately The type of Streblosoma magna comes from large, typical indivridual of Ampharete arctica Arena Bank, Baja California, Mexico, in 35 to Malmgren. Berkeley and Berkeley (1952, pp. 45 fathoms. It has setae from the second branch- 65-66, figs. 133-35) have given a diagnostic ial segment and uncinigerous tori from the account for individuals from the northern Pa- third, or first postbranchial, segment. Branchiae cific. Paleae are conspicuous, number about 30 number three pairs and form filamentous tufts. pairs; each is slender and distally pointed. Notopodial fascicles of pointed setae are present Branchiae number four pairs; they are cirri- on many segments, to near the posterior end of form and arranged in anterior and posterior the body. These characters are those of Thele- rows. The body consists of 14 thoracic setiger- pus Leuckart. Thoracic ventral scutes are fur- ous, and 13 abdominal, segments. Uncini are rowed and folded, not well defined as in Thele- present from the third setiger; they have about pus setosus (Quatrefages). Uncinigerous pin- 10 teeth in a single row at the cutting edge. nules are wart-like, not prominent. These fea- Ampharete arctica is known from circum- tures distinguish the species Thelepus crispus polar areas in moderate depths (Berkeley and Johnson, as diagnosed by Berkeley -and Berk- Berkeley, 1952, p. 66). eley (1942, p. 204). Thelepus crispus is a common littoral tere- TEREBELLIDAE bellid in the northeastern Pacific. It frequently Polymniella aurantiaca Verrill, 1900 harbors a polynoid commensal, Lepidasthenia virens (Blanchard), in its southern ranges and Eupolymnia (Polymniella) aurantiaca VERRiLL, Halosydna brevisetosa Kinberg farther to the 1900, pp. 660-661. Terebella hiata TREADWELL, 1931e, pp. 80-81, north. fig. 14. Thelepus setosus (Quatrefages), 1865 The holotype of Terebella hiata comes from Phenacia setosa QuATREFAGES, 1865, pp. 376- Haiti. It agrees fully with Polymniella auranti- 377. aca Verrill, first described from Bermuda, in Streblosoma verrilli TPEADwELL, 1911, pp. 11- having the peculiar distribution of branchiae 12, figs. 27-29. (see Hartman, 1942b, p. 74). Uncinial ridges Streblosoma verrilli was collected from a are present from the second setiger, and thor- reef near Fort Jefferson, Florida. Setae are acic notosetae are continued posteriorly through first present from the second branchial seg- a long region. Uncini are in double rows. Noto- ment and uncini from the third setigerous one. setae are distally denticulate. The species is Branchiae are in tufts on three successive seg- known only from Bermuda and Haiti. ments. Notosetae are present on about 40 seg- ments and absent from somewhat more than Loimia medusa (Savigny), 1820 half of the body length. The peristomial ridge Terebella medusa SAVIGNY, 1820, p. 85. has many eye spots in dispersed arrangement. 298 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Uncini have a large fang that is surmounted by from St. Paul de Loanda, Africa. The color two larger teeth side by side, or there may be has faded, but the body retains the dispersed another smaller one above these. Notosetae are dark flecks characteristic of Branchiomma ni- distally somewhat expanded and have smooth gromaculata. The thoracic collar is entire; it has cutting edges. There are 13 ventral scutes in a pair of midventral lobes projecting forward the thoracic region. These characters are those to cover the base of the crown. The collar is of Thelepus setosus (Quatrefages), previously continued laterally as a continuous band to the recorded from southern Florida and in the Gulf dorsal side of the body, within the notopodial of Mexico (Hartman, 1951, p. 113). bases. There are no distinct dorsal lappets, but there are thickened paired areas where they Thelepus haitiensis Treadwell, 1931 would be. This differs from the original figure Thelepus haitiensis TREADwELL, 1931e, pp. 79- in that the 80, (Treadwell, fig. 21) ventral lobes figs. a-b. are rounded at the end, not pointed; the dorsal The holotype comes from Haiti. Except for lobes are not sharply set off. the absence of eye spots on the peristomium, The tentacular crown has paired filaments on this species seems indistinguishable from The- the outer sides of the radioles, characteristic of lePus setosus (see above). It is known only Branchiomma K5lliker, 1858 (not Claparede, from the original account. 1869). The filaments at the base of the radioles are smaller and Streblosoma crassibranchia have brown tips; those near Treadwell, 1914 the middle are broader and longer. Paired eye Streblosoma crassibranchia TR ADwEi.L, 1914b, spots occur between successive pairs of proc- pp. 208-209, pl. 12, figs. 30,31. esses, nearer the outer Streblosoma crassibranchiata TRADwiELL, proximal ones, not mid- 1914b, p. 234. way between them. Dasychone loandensis is regarded as a possi- The locality and date, originally questioned, ble variety of B. nigromaculata (Day, 1955, are given in the vial, labeled as type, as San 445). p. Clemente Island, California, June, 1896. Branchiomma nigromaculata is widely dis- Branchiae are present on three segments; the on both number persed shores of the middle Atlantic of filaments in each is five, four, and Ocean (Johansson, 1927, p. 164).. three on one side and five, four, and seven on the other one. Notosetae are present from the Megalomma splendida (Moore), 1905 second branchial segment, and uncini are first Pseudopotamilla splendida MooRE., 1905, pp. present on the fourth setigerous one. The peri- 564-566, p1. 37, figs. 23-27. stomium has many small eye spots. The thorax Branchiomtm disparoculatum TREADWELL, includes at least 18 segments provided with 1914b, Pp. 223-224, figs. 44-46. notopodia. Thoracic uncini are subrectangular, The with a type of Branchiomma disparoculatum major fang surmounted by subequal was off San teeti arranged in a crescent. dredged Diego, California. It is referred to Megalomma Johansson, because the Streblosoma crassibranchia has been found radioles have more abundantly in terminal compound eyes. In this the vicinity of the Channel case are limited to Islands of southern California, in moderate they the dorsalmost pair of depths. radioles, and the eyes are large and spiralled about the subdistal part of the stem. The tho- SABELLIDAE racic collar is conspicuous and has large, dorsal Branchiomma lappets. It agrees fully with Megalomma nigromaculata (Baird), 1865 dida splen- Sabella nigro-maculata BAThD, 1865, p. 159, pl. 5, (Moore), recorded from dredged depths figs. 5, 6. in the northeastern Pacific Ocean (Moore, Dasychone loandensis TRi4waEri, 1943c, p. 4, 1923, p. 242) and as Branchiomma burrardum figs. 21-25. Berkeley and Berkeley (1941, p. 55). Dasychone nigromaculata DAY, 1955, p. 445, fig. 7. Potamilla minuta (Treadwell), 1941 The type Parasabella minuta TREADwELL, 1941a, p. 4, of Dasychone loandensis comes figs. 9-12. 1956 HARTMAN: ANNELIDS 299 The holotype comes from Sao Sabastiao The thoracic collar has no dorsal lappets. It is Island, Brazil. It measures about 10 mm. long, referred to S. magnifica (Shaw), widely re- and the branchiae are 3 mm. long. The tentacular ported from West Indian localities. crown consists of eight (not seven) pairs of The holotype of Metalonome brunnea comes radioles. I am unable to find eye spots, but there from Nassau Harbor, Bahamas, and a lot of are pigmented bars at wide intervals along the paratypes comes from the same place. Portions radiolar length. The thoracic collar conceals the of the tube are white, somewhat chitinized, and peristomium except at the dorsal side; it con- externally covered with fine white silt. The sists of a pair of lobes with midventral incision thoracic collar has a pair of prolonged tri- and is widely separated dorsally; the ventral angular lobes on its ventral ends; it extends lappets are overlapping. laterally as a shorter membrane to the dorso- The thorax has only four (not five) setiger- lateral side and ends above the first notosetal ous segments; the first segment is uniramous fascicles. Thoracic notosetae are of a single and has only pointed setae. Thoracic notosetae kind and include longer and shorter limbate are of two kinds; the uppermost are longer kinds. Thoracic neurosetae are entirely avicular. and limbate; the lower are abruptly shorter This is believed to be identical with Sabellas- and mucronate. Neurosetae include avicular tarte magnifica (Shaw). and pennoned kinds. In the abdomen the notopodial uncini are in Sabellastarte indica Savigny, 1820 single rows of seven or more, and the neuro- Sabellastarte indica SAvGNwY, 1820, p. 77. setae are in conspicuous fascicles of 24 or more Laonome punctata TREADWELL, 1906, p. 1179. long, limbate, distally pointed setae. The fecal Laonome arenosa TimADwEu, 1943d, p. 3, figs. groove runs forward midventrally along the 9-13. abdomen, turns right between the fifth and The holotype of Laonome arenosa comes sixth ventral shields, and is continued forward from Pearl Harbor, Hawaii. The thoracic col- middorsally from the fourth segment. The tube lar consists of a pair of semicircular dorsal is thin walled, externally covered with a layer lobes; it has a pair of dorsolateral clefts, lesser of fine sand grains. ventrolateral clefts, and a deep midventral fis- This individual may represent an immature sure that divides the two longest paired ventral stage, as its author suggested, but it cannot be lappets. Thoracic setae are like those of Sabel- referred to a known species chiefly because of lastarte Savigny. its low thoracic count of only four setigers and Nine paratype indiv'iduals of Laonome punc- because of the presence of a peculiar foliaceous, tata are from Waialua Reef, Oahu, Hawaii, frilled membrane extending along the inner taken in 1902. They retain much of the dark base of the tentacular crown. pigment on the tentacular crown and thoracic Potamilla minuta is known only through its collar; the pale body is dorsally speckled with original find. small dark spots, and a larger spot is present in the space between notopodia and neuropodia. Sabellastarte magnifica (Shaw), 1800 These characters agree with those of Sabella- Tubularia magnifica SHAw, 1800, pp. 228-229, starte indica Savigny, which is often regarded pl. 9, figs. 1-6. as the same as S. magnifica (Shaw) (Johans- Metalonome brunnea TREADWELL, 1917b, p. 268, son, 1926, pp. 15-16, figs. 1-5). Sabellastarte pl. 3, figs. 24-27. indica is at most a Pacific form of the West Laonome sanjuanensis TREADwEIL, 1941a, pp. Indian S. magnifica (Shaw). 3-4, figs. .4-8 The holotype of Laonome sanjuanensis SERPULIDAE comes from east San Juan, Puerto Rico, in a Sclerostyla ctenactis M8rch, 1863 tidal pool. It is a pale, small individual, per- Sclerosstyla ctenactis M6RCH, 1863, p. 386. haps immature, representative of Sabellastarte WRIGLEY, 1951, p. 184, fig. 38. Savigny. Thoracic notosetae are of one kind, Placostegus calciferus TREADWELL, 1929d, pp. tapering limbate, longer above and shorter be- 12-13, figs. 34-36. low. Thoracic neurosetae are entirely avicular. Spirodiscus calciferus TREADWELL, 1939b, p. 301. 300 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 The holotype of Placostegus calciferus comes Eupomatus uncinatus Philippi, 1844 from Julia Cove, Guanica Harbor, Puerto Rico, Eupomatus uncinatus PHILIPPI, 1844, P. 195, from a calcareous tube grown firmly on a shell. pl.6, fig. Q. Unique features include the completely calcified Eupomatus operculata T1EADnwLL, 1929d, p. 12. opercular stalk and disk and the tranversely The holotype of Eupomatus operculata comes barred radioles of the tentacular crown. This from Berbera, British Somaliland. It is unique form agrees with Sclerostyla ctexactis M6rch, in two first described from St. Thomas in the having nearly equally large opercula, as Antilles, first described. Each operculum has a long cy- and later named Sclerostyla differens Augener lindical basal (1922, p. 50) from the Barbados and stalk; the chalice has about 38 Col6n, denticulations of equal size in a circular ar- Panama. Another individual in the collections The distal consists of of the Allan Hancock Foundation was rangement. cup six (on collected one operculum) and seven (on the other one) by the "Velero III" and comes from Octavia terminal Bay, Colombia; it is illustrated curved fangs which are nearly equally by Wrigley large and similarly developed. tach fang has (1951, fig. 38). a small pointed spur visible within the cup, to Sphaeropomatus mianiensis Treadwell, 1934 be seen when the spines are pushed apart. The Sphaeropomatus miamiensis TREADWELL, 1934b, presence of two opercula, instead of a single pp. 339-341, figs. 1-5. operculum, is abnormal and has no specific sig- In other The* tubes are white, nificance. respects, this form agrees massed, erect, and re- with Eupomatus uncinatus'Philippi, well known semble those of the Filograninae. The thorax from has seven tropical parts of the Atlantic Ocean and setigerous segments; the first or colb its seas. lar segment is uniramous. Its notosetae are of adjacent two kinds: some* are transversely spinous, Eupomatus similis Treadwell, 1929 others are slender capillary setae. The next six Eupomatus similis TEA^DwErL., 1929d, pp. 11-12, thioracic segments have smooth notosetae and fig. 31. neurosetae which are uncinate with many teeth This along the cutting edge. Tentacular radioles species originates from Baja Califor- number eight on the left, and nine on the right, nia, Mexico. It measures only about 12 mm. side; they terminate in blunt, abrupt tips. long and is free from its tube. The operculum Each has a radiole has eight or nine transverse paired red basal chalice longest on its ventral side spots, possibly eyes resembling those of Sabella and bordered with about 40 crenulations. The crassicornis Sars, but the spots tend to merge distal cup is also asYmmetrical, with the long- crosswise. est spine on the dorsalmost end and the shortest The collar and thoracic membranes are dis- one on the opposite side; the spines number tinct. The operculum is on the left side; it has about 20 and are curved inward at their distal a straight stalk surmounted by a soft, vesicular end. At the inner base of each spine is a re- cap, largest on its ventral side; its distal end curved hook directed inward and down so that is flat or only slightly convex; its basal part is it points towards the center. Some of the lateral cup-like or slightly grooved, possibly through spines of the distal cup have a projecting boss shrinking. The operculum has no chitinized or at their outer distal side, but there are no lateral horny parts. teeth. The accessory operculum is represented Specimens in the American Museum of Nat- by a small, fleshy, club-shaped knob on the left ural History come from Indian River, Florida, side. and were collected by Mr. J. C. Armstrong. This individual agrees with the description Those of the type collection in the United of Hydroides recurvispina Rioia, 1941, from States National Museum come from the Miami Acapulco, western Mexico, except that there is River, Florida, from the carapace of a crab. greater asymmetry in the operculum of Eupo- The discrepancies therefore in the original matus similis than in H. recurvispina. account and the diagnosis herein given may be multiannulata the result of actual differences in local popula- Vermiliopsis (Moore), 1923 tions. Metavermilia multiannulata MOORE, 1923, pp. 251-253, pl. 18, fig. 48. 1956 5HARTMAN: ANNELIDS 301 iermiliopsis hawaiiens's T1mADwELL, 1943d, believed to be the same as Vermiliopsis multi- pp. 3-4, figs. 14, 15. annulata (Moore), first described from Vermiliopsis torquata T1EADWELL, 1943d, p. 4, dredged areas of California, and later from figs. 16, 17. Acapulco, Mexico (Rioja, 1941, p. 734, pl. 9, The holotypes of Vermiliopsis hawaiensis figs. 27-31). and V. torquata come from Hawaii. In the first one the operculum is conspicuous for its large Pomatocerus caeruleus (Schmarda), 1861 size and very dark color. The body is only Placostegus caeruleus SCHMARDA, 1861, pp. 29- about 4.5 mm. long and the operculum with 30, pl. 21, fig. 178. stalk another 3 mm. The thoracic collar is four- Pomatoceros davaoensis TREADW, 1942, pp. lobed, with the dorsolateral lobes larger than 4-5, figs. 12-14. the others and widely separated on the dorsal The holotype of Pomatoceros davaones's margin; the ventrolateral lobes are in contact Treadwell comes from Padada, Gulf of Davao, midventrally. The tentacular crown has four Philippine Islands. It is a typical individual of pairs of radioles in which the distal ends are P. caeruleus as described by Ehlers (1905, pp. free. The thorax has six uncinigerous segments. 67-70, pI. 9, figs. 11-19, as Pomatoceros strigi- The tube is thick and longitudinally ridged. ceps) for individuals from New Zealand. Ab- Vermiliopsis torquata measures about 9 mm. dominal uncini have 10 or 11 teeth in a single long and 0.5 mm. wide. The operculum has a row. The opercular stalk is very thick and has heavy stalk and terminates in an operculum conspicuous lateral wings; its distal end is pro- resembling that of V. hawaiensis. Radioles vided with a tri-pronged process far larger than number about 12 on a side and also have a long, originally shown (Treadwell, fig. 12). Mesnil free tip. The thoracic collar has a pair of prom- and Fauvel (1939, p. 35) have indicated a inent dorsal lobes. These two individuals are partial synonymy and geographic range. REFERENCES ANNENKOVA, N. P. 1927a. Polychaeten von Neu-Pommern. Sitzber. 1929. Beitrage zur Kenntnis der Polychaeten- Gesell. Naturf. Fr. Berlin, for 1926, Fauna der USSR. Fam. Pectinariidae pp. 119-152, 1 p1. Quatrefages (Amphictenidae Malm- 1927b. Bijdragen tot de Kennis der Fauna van gren) und Ampharetidae Malmgren. Cura9ao. Resultaten eener Reis van Dr. Ann. Mus. Zool. Acad. Sci. USSR, pp. C. J. van der Horst in 1920. Polychaeten 477-502, pls. 37-39. von Curagao. Zool. Genoots. Nat. Artis 1931. Zur Polychaetenfauna von Franz- Magistra, Amsterdam, vol. 15, pp. 39-82, Joseph-Land (Melinnexis gen. nov. arc- 9 figs. tica sp. nov.). Zool. 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Board Canada, Toronto, pp. 1-100, 160 Part 2. Errant species from Cape shores figs. and estuaries. Ibid., vol. 12, pt 3, pp. 1951. A second record of the polychaetous an- 397-441, 6 figs. nelid Potamethus elongatus (Tread- 1955. The Polychaeta of South Africa. Part 3. well). Jour. Washington Acad. Sci., vol. Sedentary species from Cape shores and 41, no. 10, pp. 333-334, figs. 1-4. estuaries. Jour. Linnean Soc. London, 1952. Annelida, Polychaeta sedentaria. Ca- vol. 42, pp. 407-452, 8 figs. nadian Pacific Fauna, no. 9b (2), Fish. DuJAwIN, FELIX Res. Board Canada, Toronto, pp. 1-139, 1839. Observations sur quelques ann6lides 292 figs. marins. Ann. Sci. Nat. Paris, ser. 2, BLANCHARD, EML.c vol. 11, pp. 287-194, pl. 1, figs. 1-12. 1849. Anelides du Chile. In Gay, Claude, His- EHLES, ERNST toria fisica y politica de Chile. Paris, 1887. Report on the annelids of the dredging Zoologia, vol. 3, pp. 9-52, pls. 1, 2. expedition of the U. S. coast survey CHAMBERLIN, R. V. steamer Blake. Mem. 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Annelida polychaeta of the Indian Mu- 304 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 seum, Calcutta. Mem. Indian Mus., vol. bekannter von Hrn. Ehrenberg gesam- 12, no. 1, pp. 1-262, 40 figs., 9 pls. melter Anneliden des rothen Meeres. 1933. Ann6Iides polychetes du Golfe du Pei Monatsber. Akad. Wiss. Berlin, pp. Tchen Ly de la collection de Musee 484-521. Hoang ho Pai ho. Publ. Mus. Hoangho 1877. Ueber eine Sanumlung von wirbellosen Pai ho Tien-Tsein, vol. 15, pp. 1-67, Seethieren, welche Herr Dr. Eugen 6 figs. Reimann dem hiesigen zoologischen Mu- 1939. Annelida polychaeta. Resultats scien- seum zum Geschenk gemacht. Jahrb. tifiques des croisieres du Navire-4cole Ges. Vaterl. Kultur, Breslau, vol. 54, Belge "Mercator." Mem. Mus. Hist. Nat. pp. 48-51. Belgique, ser. 2, fasc. 15, pp. 1-37. 1878. Annulata Semperiana. Mem. Acad. Sci. 1943. Annelides polych6tes de Californie re- St. Petersburg, vol. 25, ix+300 pp., 15 cueillies par L. Diguet. Medm. Mus. Hist. PIS. Nat. 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Actinien, Echinodermen und Wiirmen Museum and the description of a new des Adriatischen und Mittelmeers. K6- genus. Proc. U. S. Natl. Mus., vol. 86, nigsberg, J. H. Bon, pp. 61488, 1 pi. pp. 107-134, 7 figs. 1855. Beschreibung neuer oder wenig bekann- 1939a. The polychaetous annelids collected by ter Anneliden. Arch. Naturgesch., Ber- the Presidential Cruise of 1938. Smith- lin, vol. 21, no. 1, pp. 81-128, pls. 3-4. sonian Misc. Coll., vol. 98, pp. 1-22, 1856-1859. Annulata Oerstediana. Enumeratio 3 figs. Annulatorum, quae in itinere per In- 1939b. Polychaetous annelids. Aphroditidae to diam occidentalem et Americam cen- Pisionidae. Allan Hancock Pacific Ex- tralem annis 1845-1848 ... in itinere ad peditions. Los Angeles, vol. 7, pp. 1-156, Americam Meridionalem collectis. Vi- pls. 1-28. densk. Medd. Naturhist. For., Copen- 1940a. Polychaetous annelids. Chrysopetalidae hagen, pt. 1, pp. 44-62; pt. 2, pp. 158- to Goniadidae. Allan Hancock Pacific 186; pt. 3, pp. 105-120. Expeditions. Los Angeles, vol. 7, pp. 1870. Beschreibungen neuer oder weniger 173-287, pls. 31-44. 1956 HARTMAN: ANNELIDS 305 1940b. Boccardia proboscidea, a new species of no. 1, pp. 1-181, 3 figs., 19 pls. spionid worm from California. Jour. 1951. The littoral marine annelids of the Gulf Washington Acad. Sci., vol. 30, pp. 382- of Mexico. Publ. Inst. Marine Sci., vol. 387, 1 fig. 2, no. 1, pp. 7-124, pls. 1-27. 1941. Some contributions to the biology and 1952. The identity of Stylarioides inflata life history of Spionidae from Cali- (Treadwell). Pacific Sci., Univ. Hawaii, fornia. Allan Hancock Pacific Expedi- vol. 6, no. 1, pp. 71-74, 2 figs. tions. Los Angeles, vol. 7, pp. 289-324, 1954a. New species of polychaetous annelids pls. 45-48. from the Marshall and Gilbert Islands. 1942a. The identity of some marine annelid Jour. Washington Acad. Sci., vol. 44, worms in the United States National pp. 228-232, 2 pls. Museum. Proc. U. S. Natl. Mus., vol. 92, 1954b. Australian Nereidae. Trans. Roy. 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Zeitschr. pedition 1933-34, Scientific reports. Wiss. Zool. Leipzig, vol. 33, pp. 267-316, London, British Museum (Natural His- pls. 14-18. tory), vol. 4, Zoology, pt. 8, pp. 243-321, LINNAEUS, CAROLUS 28 figs. 1767. Systema naturae. Editio duodecima, re- 1938. On a small collection of Polychaeta formata. Stockholm, p. 1086. from Uruguay. Ann. Mag. Nat. Hist., MCCONNAUGHEY, BAYARD H., AND DENNIS L. ser. 11, vol. 2, pp. 311-314, 3 figs. Fox 1939. Polychaeta of the Rosaura Expedition. 1949. The anatomy and biology of the marine Novitates Zool., vol. 41, pp. 345-354, polychaete Thoracophelia mucronata figs. 280-283. (Treadwell), Opheliidae. Univ. Cali- MONTAGU, GEORGE fornia Publ. Zool., vol. 47, pp. 319-340, 1804. Descriptions of several marine animals pls. 26-30. found on the south coast of Devon- MCINTOSH, WILLIAM CA1MECHA shire. Trans. Linnean Soc. London, vol. 1885. Report on the Annelida Polychaeta col- 7, pp. 80-84, pl. 7. lected by H.M.S. Challenger during the 1808. New and rare animals found on the years 1873-76. In Report on the scien- south coast of Devonshire. Ibid., vol. 9, tific results of the voyage of H.M.S. pp. 108-111, pls. 6-8. Challenger. London, Zoology, vol. 12, 1815. Descriptions of several new or rare pp. 1-554, pls. 1-55, la-39a. animals principally marine found on 1901. Notes from the Gatty Marine Labora- the south coast of Devonshire. Ibid., tory, St. Andrews. Ann. Mag. Nat. Hist., vol. 11, pp. 18-21, pls. 3-5. ser. 7, vol. 8, pp. 216-232, 1 pl. MOORX, JOHN PERCY MALMGREN, ANDERS JOHAN 1903. Some pelagic Polychaeta new to the 1866. Nordiska Hafs-Annulater. Ofvers. K. Woods Hole fauna. Proc. Acad. Nat. Vetensk.-Akad. Forhandl., Stockholm, Sci. Philadelphia, vol. 55, pp. 793-801, vol. 22, pp. 355-410, pls. 18-29. pl. 55. MESNIL, FELIX, AND PIRE FAUVEL 1905. Five new species of Pseudopotamilla 1939. Polychetes sedentaires de l'expedition from the Pacific coast of. North America. du "Siboga." Maldanidae, Cirratulidae, Ibid., vol. 57, pp. 555-569. Capitellidae, Sabellidae et Serpulidae. 1909. Polychaetous annelids from Monterey Monographie 242 de Uitkomsten op Bay and San Diego, California. Ibid., zoologisch, botanisch, oceanographisch- vol. 61, pp. 235-295, pls. 7-9. en en geologisch gebied verzameld in 1911. The polychaetous annelids dredged by Nederlandsch Oost-Indie 1899-1900 an the U.S.S. Albatross off the coast of boord H.M. Siboga onder commando southern California in 1904. Euphro- van . .. Tydeman. Leiden, livr. 133, pp. synidae to Goniadidae. Ibid., vol. 63, 1-42, 12 figs. pp. 234-318, pls. 15-21. MoRcH, OTTo A. L. 1923. The polychaetous annelids dredged by 1863. Revisio critica Serpulidarum. Naturhist. the U.S.S. Albatross off the coast of Tidsskr., Copenhagen, ser. 3, vol. 1, southern California in 1904. Spionidae pp. 347-470, 1 pI. to Sabellariidae. Ibid., vol. 75, pp. 179- MONRO, C. C. A. 259, pls. 17, 18. 1924. On the Polychaeta collected by H.M.S. OKUDA, SHIRO Alert, 1881-1882. Jour. Linnean Soc. 1937. Polychaetous annelids from the Palau London, vol. 36, pp. 37-64, 24 figs. Islands and adjacent waters, the South 1928. On the Polychaeta collected by Dr. Th. Sea Islands. Bull. Biogeogr. Soc. Japan, Mortensen off the coast of Panama. vol. 7 (12), pp. 257-316, 59 figs. 1956 HARTMAN: ANNELIDS 307 PETERS, WILHELM C. H. RIOJA, ENRIQUE 1854. Naturwissenschaftliche Reise nach Mos- 1941. Datos para el conocimiento de la fauna sambique in 1842 bis 1848 ausgefiihrt. de poliquetos de las costas del Pacifico Monatsber. Akad. Wiss. Berlin, pp. 610- de Mexico. An. Inst. Biol. Mexico, vol. 614. 12, pp. 669-746, 11 pls. 1855. Ueber die Gattung Bdella (Sav.) und 1944a. Estudio de algunos poliquetos del Museo die in Mosambique beobachteten Anne- Argentino de ciencias naturales. Ibid., liden. Arch. Naturgesch., vol. 21, no. 1, vol. 15, pp. 115-138, 62 figs. pp. 35-42. 1944b. Observaciones acerca del operculo de PETTIBONE, MARIAN FT. Hydroides crucigera M6rch y descrip- 1949. Polychaetous annelids of the Poly- cion de un caso de duplicidad de este noidae from the northeastern Pacific, organo. Ibid., vol. 15, pp. 409-414, figs. with a description of a new species. 1-5. Amer. Mus. Novitates, no. 1414, pp. 1-6, RoSA, DANIELI figs. 1-9. 1908. Nouve specie di Tomopteridi. Diagnosi 1954. Marine polychaete worms from Point preliminari. Boll. Mus. Zool. Anat. Barrow, Alaska, with additional records Comp. Torino, vol. 23, no. 588, p. 1. from the North Atlantic and North RULLIER, FRANgOIS Pacific. Proc. U. S. Natl. Mus., vol. 103, 1954. Essai de revision du genre Fabricia pp. 203-356, figs. 26-39. (annelides polychetes). Bull. Soc. Zool. PHILIPPI, A. France, vol. 79, no. 1, pp. 14-29, figs. 1844. Einige Bemerkungen uiber die Gattung 1, 2. Serpula, nebst Aufza.hlung der von mir SAVIGNY, JULES CUSAR im Mittelmeer mit dem Thiere beobach- 1820. Systeme des annelides. In Description teten Arten. Arch. Naturgesch., vol. 10, de l':gypte. Paris, Panckouche, Histoire no. 1, pp. 186-198, pl. 6. naturelle, vol. 1, pt. 3, pp. 1-128. PIXELL, HELEr L. M. SCHMARDA, LUDWIG K. 1913. Polychaeta of the Indian Ocean, to- 1861. Neue wirbellose Thiere beobachtet und gether with some species from the Cape gesammelt auf einer Reise um die Erde Verde Islands. The Serpulidae, with a 1853 bis 1857. Leipzig, vol. 1, pt. 2, classification of the genera Hydroides pp. 1-164, 22 pls. and Eupomatus. Trans. Linnean Soc. SEIDLER, HANS J. London, vol. 16, pp. 69-92, pls. 8, 9. 1924. Beitrage zur Kenntnis der Polynoiden. PRUVOT, G. 89A, no. 11, 1930. Annelides polych&tes de Nouvelle- Arch. Naturgesch., vol. M. pp. 1-217, 22 figs. Caledonie recueillies par Frangois. SHAW, G. Arch. Zool. Exp. Gen. Paris, vol. 70, 1800. Descriptions of Mus bursarius and pp. 1-94, 8 figs., 3 pls. Tubularia magnifica, from drawings QUATREFAGES, ARMAND DE communicated by Major-GeneralThomas 1865. Histoire naturelle des anniles marina Trans. Linnean et d'eau douce. Annelides et gephyriens. Davies, F.R.S. and L.S. Paris, Roret, vol. 2, pt. 2, pp. 337-794. Soc. London, vol. 5, pp. 227-229, 2 pls. REIBIscH, JOHANNES SOUTHERN, ROWLAND 1893. Die Phyllodociden der Plankton-Expedi- 1911. Polychaeta of the coasts of Ireland. tion. Zool. Anz., vol. 16, pp. 248-255, The Alciopinae, Tomopteridae and 2 figs. Typhloscolecidae. Fish. Ireland Sci. 1895. Die pelagischen Phyllodociden und Invest., Dublin, no. 3, pp. 1-37, 3 pls. Typhloscoleciden der Plankton-Expedi- ST0P-BOWITZ, C. tion. In Hensen, Victor (ed.), Ergeb- 1948. Polychaeta from the "Michael Sars" nisse der . .. Plankton-Expedition der North Atlantic deep-sea expedition 1910. Humboldt-Stiftung. Kiel and Leipzig, In Report on the scientific results of vol. 2H, pt. c, pp. 1-63, 5 pls., 3 charts. the "Michael Sars" North Atlantic deep- REISH, DONALD JAmES sea expedition 1910. Bergen, vol. 5, 1950. Redescription of the polychaetous anne- no. 8, pp. 1-91, 51 figs., 5 tables. lid Typosyllis aciculata Treadwell. Amer. 1951. Polychetes pelagiques de l'Expedition Mus. Novitates, no. 1466, pp. 1-5j 8 Suedoise Antarctique 1901-1903. In figs. Further zoological results of the Swed- 308 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 ish Antarctic Expedition 1901-1903. U. S. Natl. Mus., vol. 100, pp. 589-602, Stockholm, vol. 4, no. 7, pp. 1-14. 8 figs. TREADWELL, AARON Louis 1920b. A new polychaetous annelid of the genus 1891. A preliminary note on the anatomy and Nereis from Brazil. Proc. U. S. Natl. histology of Serpula dianthus (Verrill). Mus., vol. 58, pp. 467-468, 4 figs. Zool. Anz., vol. 14, pp. 276-280, 2 pls. 1921a. Polychaetous annelids collected at St. .1900. Lateral line organs in Eunice auriculata Paul de Loanda by the American Mu- n. sp. Science, vol. 12, pp. 342-343, 2 seum Belgian Congo Expedition. Amer. figs. Mus. Novitates, no. 8, pp. 1-3. 1901a. The cytogeny of Podarke obscura 1921b. Report on the annelids of Puget Sound, (Verrill). A dissertation submitted to Fiji and Samoa. Yearbook Carnegie the faculties of the Graduate Schools Inst. Washington, no. 19, pp. 199-200. of Arts, Literature, and Science, in 1921c. Nereis (Ceratonereis) alaskensis, a new candidacy for the degree of Doctor of polychaetous annelid from Alaska. Proc. Philosophy, the University of Chicago. U. S. Natl. Mus., vol. 60, art. 2, pp. 1-3, Jour. Morph., vol. 17, pp. 399-486, 8 4 figs. figs., 5 pls. 1921d. Leodicidae of the West Indian region. 1901b. The polychaetous annelids of Porto Rico. Publ. Carnegie Inst. Washington, no. Bull. U. S. Fish Comm., vol. 20, pp. 181- 293, pp. 1-131, 467 figs., 9 pls. 210, 81 figs. 1922a. Leodicidae from Fiji and Samoa. Ibid., 1905. On the heteronereis stage of Nereis no. 312, pp. 127-170, 168 figs., 8 pls. kobiensis McIntosh. Biol. Bull., Woods 1922b. Polychaetous annelids collected at Fri- Hole, vol. 9, pp. 226-231, 5 figs. day Harbor, State of Washington, in 1906. Polychaetous annelids of the Hawaiian February and March, 1920. Ibid., no. Islands, collected by the Steamer Al- 312, pp. 171-181, 37 figs. batross in 1902. Bull. U. S. Fish Comm., 1923a. Two new species of polychaetous anne- vol. 23, pp. 1145-1181, 81 figs. lids of the genus Nereis from Brazil. 1909. Haplosyllis cephalata as an ectoparasite. Rev. Mus. Paulista, vol. 13, pp.- 1-10, Bull. Amer. Mus. Nat. Hist., vol. 26, 15 figs. pp. 359-360, 2 figs. 1923b. Polychaetous annelids from Lower Cali- 1911. Polychaetous annelids from the Dry fornia with description of new species. Tortugas, Florida. Ibid., vol. 30, pp. 1- Amer. Mus. Novitates, no. 74, pp. 1-11, 12, 29 figs. 8 figs. 1914a. Researches upon annelids at Tortugas. 1923c. The heteronereis phase of a new species Yearbook Carnegie Inst. Washington, of polychaetous annelid from Uruguay. vol. 13, pp. 220-222. Proc. U. S. -Natl. Mus., vol. 64, art. 9, 1914b. Polychaetous annelids of the Pacific pp. 1-3, 1 pl. coast in the collection of the zoological 1924a. A new marine annelid from Chile. Ibid., museum of the University of California. vol. 65, art. 23, pp. 1-3, 4 figs. Univ. California Publ. Zool., vol. 13, 1924b. Dasychonopsis arenosa, a new species pp. 175-234, 2 pls. of polychaetous annelid from Porto 1914c. New Syllidae from San Francisco Bay, Rico. Amer. Mus. Novitates, no. 107, collected by the U.S.S. Albatross. Ibid., pp. 1-2, 4 figs. vol. 13, pp. 235-238, figs. 1-7. 1924c. Polychaetous annelids, collected by. the 1917a. A new species of polychaetous annelid Barbados-Antigua expedition from the from Panama, with notes on an Ha- University of Iowa in 1918. Univ. Iowa waiian form. Proc. U. S. Natl. Mus., vol. Studies, vol. 10, no. 4, pp. 1-23, 2 pls. 1925a. A new species of polychaetous annelid 52, pp. 427-430, 5 figs. from 1917b. Polychaetous annelids from Florida, Uruguay, Aphrodita magna. Proc. U. S. Natl. Mus., vol. 67, art. 12, pp. Porto Rico, Bermuda and the Bahamas. 1-3, 3 figs. Publ. Carnegie Inst. Washington, no. 1925b. list 251, pp. 255-272, 3 pls. A of the annelids collected by Capt. 1920a. Polychaetous R. A. Bartlett in Alaska, in 1924, with annelids collected by the description of a new vol. United States Fisheries steamer Alba- species. Ibid., tross in 67, art. 29, pp. 1-3, 4 figs. the waters adjacent to the 1925c. Polychaetous annelids of tropical central Philippine Islands in 1907-1910. Bull. Pacific. Bull. Bernice P. Bishop Mus., 1956 HARTMAN: ANNELIDS 309 HoInolulu, vol. 27, pp. 113-119. 1931d. New species of polychaetous annelids 1926a. Polychaetous annelids from Fiji, Samoa, from California, Mexico, Porto Rico, China and Japan. Proc. U. S. NatL Mus., and Jamaica. Amer. Mus. Novitates, vol. 69, art. 15, pp. 1-26, pls. 1-2. no. 482, pp. 1-7,21 figs. 1926b. Contributions to the zoology of the 1931e. New trebellid annelids, Thelepus haiti- Philippine Archipelago and adjacent re- ensis and Terebella hiata. Zoologica, gions. Additions to the polychaetous vol. 12, pp. 79-81, 1 fig. annelids collected by the U.S. fishery 1931f. A new West Indian polychaetous anne- steamer Albatross. Bull. U. S. Natl. Mus., lid. Amer. Mus. Novitates, no. 461, 2 pp., vol. 100, art. 6, pp. 183-193, 20 figs. 3 figs. 1926c. A new polychaetous annelid from Kar- 1932a. Haplobranchus atlanticus, a new species tabo, British Guiana, genus Namoneresis. of polychaetous annelid from St. An- Zoologica, vol. 7, pp. 101-104, 1 fig. drews, N. B. Contrib. Canadian Biol., 1926d. Polychaetous annelids collected by Cap- Toronto, new ser., vol. 7, pp. 277-281, tain R. A. Bartlett in Alaska in 1924, 8 figs. with descriptions of new species. Amer. 1932b. New species of polychaetous annelids Mus. Novitates, no. 223, pp. 1-8, 17 figs. from Ilha de Sao Sebastiao, together 1928a. Polychaetous annelids from the Arcturs with a list of other annelids determined oceanographic expedition. Zoologica, from Brasil. Rev. Mus. Paulista, vol. vol.8, pp. 449-489, 3 figs. 17, art. 2, pp. 1-20, 1 pl. 1928b. A new polychaetous annelid of the 1934a. New polychaetous annelids. Smithsonian genus Phyllodoce from the west coast Misc. Coll., vol. 91, no. 8, pp. 1-9, 2 pls. of Costa Rica. Proc. U. S. Natl. Mus., 1934b. Sphaeropomatus miamiensis, a new vol. 74, pp. 1-3, 5 figs. genus and species of serpulid polychaete. 1929a. Two new species of polychaetous anne- Jour. Washington Acad. Sci., vol. 24, lids from the Argentina coast. Ibid., pp. 338-341, 9 figs. vol. 75, art. 26, pp. 1-5, 6 figs. 1936a. Polychaetous annelids from Amoy, 1929b. Lumbrinereis bicirrata, a new poly- China. Proc. U. S. Natl. Mus., vol. 83, pp. chaetous annelid from Puget Sound. 261-279, figs. 18-20. Amer. Mus. Novitates, no. 338, pp. 1-3, 1936b. Polychaetous annelids from the vicin- 7 figs. ity of Nonsuch Island, Bermuda. Zoo- 1929c. Acoetes magnifica, a new species of logica, vol. 21, pp. 49-68, 3 pls. polychaetous annelid from Montego Bay, 1937a. Polychaetous annelids collected by Cap- Jamaica, British West Indies. Ibid., no. tain Robert A. Bartlett in Greenland, 355, pp. 14, 7 figs. Fox Basin and Labrador. Jour. Wash- 1929d. New species of polychaetous annelids ington Acad. Sci., vol. 27, pp. 23-26, in the collections of the American Mu- 16 figs. seum of Natural History, from Porto 1937b. The Templeton Crocker Expedition. Rico, Florida, Lower California, and Polychaetous annelids from the west British Somaliland. Ibid., no. 392, pp. coast of Lower California, the Gulf of 1-13, 36 figs. California and Clarion Island. Zoologica, 1931a. Contributions to the biology of the vol. 22, pp. 139-160, 2 pls. Philippine Archipelago and adjacent re- 1937c. Hemipodia canadensis, a new polychae- gions. Four new species of polychaetous tous annelid from Nova Scotia. Jour. annelids collected by the United States Biol. Board Canada, Toronto, vol. 3, fisheries steamer Albatross during the pt. 4, pp. 348-349, 3 figs. Philippine Expedition of 1907-1910. 1939a. New polychaetous annelids from New Bull. U. S. Natl. Mus., vol. 100, pp. 313- England, Texas and Puerto Rico. Amer. 321, 4 figs. Mus. Novitates, no. 1023, pp. 1-7, 25 1931b. Three new species of polychaetous figs. annelids from Chesapeake Bay. Proc. 1939b. Polychaetous annelids of Porto Rico U. S. Natl. Mus., vol. 79, art. 1, pp. 1-5, and vicinity. Scientific survey of Porto 3 figs. Rico and the Virgin Islands. New York, 1931c. Three new species of polychaetous New York Academy of Sciences, vol. annelids in the collections of the United 16, pt. 2, pp. 151-318, 118 figs. States National Museum. Ibid., vol. 80, 1939c. New species of polychaete worms of art. 2, pp. 1-5, 3 figs. the genus Euphrosyne, with notes on 310 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 109 Euphrosyne borealis Oersted. Proc. U. S. from Massachusetts and California. Natl. Mus., vol. 86, pp. 169-173, 1 fig. Ibid., no. 1280, pp. 1-3, 8 figs. 1940. A new genus and two new species of 1948. Polychaeta. Canadian Atlantic Fauna, polychaetous annelids from Texas and Toronto, no. 9b, pp. 1-69, 49 figs. one new species from the Philippine VERRLL, ADDISON EMERY Islands. Amer. Mus. Novitates, no. 1089, 1873. Report upon the invertebrate animals of pp. 1-4, 13 figs. Vineyard Sound and the adjacent 1941a. Polychaetous annelids from the New waters, with an account of the physical England region, Porto Rico and Brazil. characters of the region. Rept. U. S. Ibid., no. 1138, pp. 1-4, 12 figs. Fish Comm., pp. 295-778. 1941b. New species of polychaetous annelids 1900. Additions to the Turbellaria, Nemertina from the vicinity of Galveston, Texas. and Annelida of the Bermudas, with re- Ibid., no. 1139, pp. 1-3, 10 figs. visions of some New England genera 1941c. Eastern Pacific expeditions of the New and species. Trans. Connecticut Acad. York Zoological Society. Polychaetous Sci., Vol. 10, pp. 595-671, 1 pl. annelids from the west coast of Mexico VIGUIER, CAMILLE and Central America. Zoologica, vol. 26, 1886. Atudes sur les animaux inferieurs de la pp. 17-24, 21 figs. Baie d'Alger. Recherches sur les anne- 1941d. Plankton of the Bermuda oceanographic lides pelagiques. Arch. Zool. Exp. Gin. expeditions. Polychaetous annelids from Paris, ser. 2, vol. 4, pp. 347-442, pls. Bermuda plankton, with eight shore 21-27. species and four from Haiti. Ibid., vol. WEBSTER, HARRISON EDWIN 26, pp. 25-30, 9 figs. 1884. Annelida from Bermuda. Bull. U. S. 1942. Polychaetous annelids from Lower Cali- Natl. Mus., no. 25, pp. 305-327, pls. 7-12. fornia and the Philippine Islands in the WESENBEEG-LUND, ELISE collections of the American Museum of 1949. Polychaetes of the Iranian Gulf. In Natural History. Amer. Mus. Novitates, jessen, Knud, and Ragnar Spirck (eds.), no. 1172, pp. 1-5, 15 figs. Danish scientific investigations in Iran. 1943a. Polychaetous annelids from the Philip- Copenhagen, pt. 4, pp. 247-400, 47 figs., pine Islands in the collections of the 3 maps, 3 tables. American Museum of Natural History. WILLEY, ARTHUR Ibid., no. 1220, pp. 1-4, 7 figs. 1905. Report on the Polychaeta collected by 1943b. Biological results of the last cruise of Professor Herdman, at Ceylon, in 1902. the Carnegie. Polychaetous annelids. In In Herdman, W. A., Report to the gov- Scientific results of Cruise VII of the ernment of Ceylon on the pearl oyster Carnegie during 1928-1929 under com- fisheries of the Gulf of Manaar. London, mand of Captain J. P. Ault. Washing- Royal Society, pt. 4, pp. 243-324, pls. ton, pp. 30-59, 1 pl., 4 maps, table. 1-8. 1943c. Polychaetous annelids from Africa in WIRfN, AxEL the collections of the American Museum 1883. Chaetopoder fran Sibiriska Ishafvet och of Natural History. Amer. Mus. Novi- Berings haf, insamlade under Vega- tates, no. 1221, pp. 1-6, 25 figs. expeditionen 1878-1879.In Nordenskiold, 1943d. New species of polychaetous annelids A. E., Vega-expeditionens vetenskapliga from Hawaii. Ibid., no. 1233, pp. 1-4, 17 iakttagelser. Stockholm, pp. 381-428, figs. pls. 27-32. 1943e. Neosabellides alaskensis, a new species WRIGLEY, ARTHUR of polychaetous annelid from Alaska. 1951. Some Eocene serpulids. Proc. Geol. Ibid., no. 1235, pp. 1-2, 5 figs. Assoc. London, vol. 62, pt. 3, pp. 177- 1945. New species of polychaetous annelids 202, 66 figs.