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080057-12.018.Pdf

080057-12.018.Pdf

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(8661)902/61:(z)z I ,7]l,,(']N N,r)hid Vol. 12, No. 2 (1998)

Taxonomy

Key to the speciesof subgents Centridium

I: Appendage(s)present on gynostcmium...... 2 1 Appendage(s)absent on 9ynostemium...... 6 2: Gynostemiumwith 2 appendages;corolla prcdominately orangc on adaxialsurface ...... S.diceratum 2 Gynostcmiumwith l appcndage;corolla white or pink on adaxialsurfacc...... 3 3: Gynostemiumbearing a recurvedhorn shaped appcndage on thebend...... 4 3 Gynostemiumappcndage not horn-shaped ...... 5 4: Throat appendages2; labellumclliptic, apex not emarginatc, irregularlyserrate .....S. calcaratum 4 Throatappendages 4; labellumlagcniforrn, apex emarginatc, not serrate...... S. weeliwolli 5: Gynostemiumappendlge reniform, recurved from thebend; nectaryspur prominent, cradled by the horizontalposterior sepal...... S.mimeticum 5 Gynostcmium appcndagcsquare, recurved from the bend, apex irregularly scrrate;ncctary spur absentor very poorly developed and hiddenbehind the alwaysvertical posterior sepal...... S. ecorne 6: Corolla orangc; posterior corolla lobes each deeply dividcd into 2 (soas to appearas 4 individuallobes)...... S.ceratophorum 6 Corolla white to pink; postcriorcorolla lobes undivided ...... 7 7: Plantsmostly 1.5-2.5crn high; nectaryspur abscnt...... S.perpusillum 7 Plantsmostly 4.5 25 cm high; ncctaryspur present ...... 8 8: Posteriorcorolla lobcs cuncale, the apexobtuse and unlobcd,with a distinctiveblunt lateraltooth at the basc...... S. edenlatum 8 Posterior corolla lobes either cuneatewith a tridentateapex or obovate with a crenateapex, lacking basal tooth...... 9 9: Plantsmostly 5 9 cm talli postcriorcorolla lobes cuneatc, apex tridentate; nectaryspur shortcr than the posteriorsepal ...... S. aceratum 9 Plantsmostly l0-25 cm tall; postedorcorolla lobes obovate, apex crenate; nectaryspur longcr than thc posteliorsepal ...... S.longicornu

Stylidium aceratum Lowrie & Kenneally,qr. /lor.

Stylidio calcqrqto R. Br. affinis sed cornu appendicise flexo gynostemii absenti.

?ypus. Great Northem Highway, north of Bullsbrook fpreciselocality withheld], Wcstern , 9 Novcmber 1991,A. Lowrie 496 (.holo: PERTH 04980336;lso: MEL).

A fibrous-rooted annual herb 5 9 cm high (including infloresccnce);stem translucent white, 2.54 mm long, 0.8 I mm diam.; basal rosetteof leavesflat, 5-12 mm diam. Leayes spathulate, 3 6 mm long, 0.7 1.5 mm widc nearapex, 0.3-0.6 mm wide at the base,flat in section,glabrous. Inflorescenceusually a simpledichasium but alsothc beginningsof a compounddichasium in older ,5-9cm long(including peduncle), glandular. Brqcts and. bracteole.r lanceolate, 1.5-2mmlons, '0noqnnolqt eleJras,{l oqs xede un1leqe1)eprs Jeqlrestueru8es eleuos rotrorls rltr^\ ctuprdsnoxodu tunlleqpl puu:(ulnrluelsou^3 eql lo pueq eql lu lueserdeScpuedd|) urn ruretsou^3 orlt Jo pu.q aqt tUo-UluesqE e8upuedde:(Fdas rouelsod ?ql ucqlre8uol) lrdes roualsod eqt uuqlieuoqs inds ,{rrlJou:(ruJo]rueJ sa8epucddeleorql Z) saqolllloror rouelsodeqt lo estq eql lt seSrpuaddetuorql lt;rruol Z Surlcrl ,{q (slseqlueredur ue,lrFare srelcp.ruqcSurlsurluoc csoq,lr) wnlDtDJlD) S uror:ls.lcl.lrp Lunu)JaJDS 'I I = UJO requnu eruosouroJqcl?puB seqol rlloJor ror:olsodctctuopr.rt Puu sccioll?llojor rorjotuB elelnctue8o,\sq serceds qlo1 wnnn)lDJ S sr wnJp.la)panlpllf,iS ot elrlulor lseJcauaqJ saqluu|

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A. Lowrie 1998

Figurc 1. .5/llidrr?rar.r'drrrr. A hrbit ol llolvcriDgplant; B - , eDlafgedsecli{tr lelt; C hypanthium and sepals; D laleralview ol-corullir,gynoslcrrinrn and hypanlhium;E corollar F throat appcndagcs; G - tabellurniH - laloralviclv of gyrostemium,anthcrs and slipilalcst;gma in lhe set non triggefedposition Scaie bar for all = | nnn. DrarvD lrc.|' A. It^rriz 4')6.

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Egure 2. S/l/i./i! di.erutun A. habit of nolvering ; B leal enlarged seclion left; C hypanlhium and sepalsi D cololla; E labcllur]r:F fiont view ofgynostemium, antheN and stipitate stigma in the tdggercd position; G latefal view of gynostemium, antheN and stipitate stigma in the tdggered position. Scale bar for all = 1 nnn. Drawn lioln M.D. Bltrret 235 & A. Lo)ttie 1526. (t arn?rg) aso8nr'rutrrp Z 0 91 q'8uol utu g 9 97 9'prosd11e+'u^\orq {rvp 'esoqol8 '8uol 'sJoqtuu rpaas uerp ulru t-Z aTzsdn3 aIl-qsn:q xodl urru E l r aq] uoo,{\teqaletrdrts €ru8lls:,tolle,{IJ€p sreqlueiunrualsou{3 aqt;o puaqeqt uo 3uo1urur 7 ii r oicpucddee41-u:oq e qlIA\'8uolulul / t , unlnquunt o^rlrsucsaql'8uol urutg l J uurnJoi{esrq c^rtisu?s uou tcolc.ql '3uo1urut7 '?laulSre 'cpr,r '3uo1urui -r 'u:oyucSe1 g c wntwalsou'lg snorqulS utc xcdp uur 8 0 J 1 1 'eAEJuoJ'soqol alloJocJouelup eqtJo snurs eql ,roleqpeuoltrsod wn11aqo7 3uo1 urur 1 1 r'osEq ol 'q13ue1 eargsa8upuedde :olrelsod :3uo1uru g 1 r :reql 3uo1u pesnl seiupueddu,rouetuu l seqolelloJoJ 'so8upucddc roualsod 1oeseq eqt lt Z puc saqolBlloioo louatuu Jo cs€qcql tE Z peduqr--po.ry Suuuaq 'SuoIwuJ 'epr,t toorllJ Vdes Jouelsod,{q pelperc g Z Z tnds ttotcatrl etruerJr.rt+ xedu ruur g g ! '8uol 'epr^\ '8uol urul 9 gS'elEeunJ seqol rorrelsod:elEurSffura xed]l urul Z 8l Lrlrug8-g'/ 'ruroJrJnJr 'et€lnf,ruei 'snorqutB'.srq fllcuqsrps^€,^dle seqolJor.r€tua:parrlrd,{llDcrlra^ seqol cq} tr 'lptuozuorl s{.Iuurqsrppor rllr,ll ec€JJnslElxepu uo1urdyep o11oLo3Suolrttw 9 Z-i4 I lDdesroualsod 'scqol :8uol u[u g Z-Z lrora rrDdelppru :8uol rutx 9 Z Z EIloJo3rouotuE orlt tepun sprs,\\lnope,{Elds puElEluozuoqrredroueluE:lElnpuElS'otBloocuul'cs[qcqlo]o?rJIIr?'9.r/rdasrulnputlS'srseqluu 'esoqo18 '8uol le ulurp rurtr l l 7 1 unryrynd(p JelnpuulS urtrtOZ OI slntpad :r:1npuu13{1csrvd,'^ 'elnaE '3uo1 xed"'epr,rruwZZ VI ururg tr-7 7'cudrIla-ctr^ocloseloa\rD.tq ptrc sTrn:g re1npuel8 '(elJunpedSurpnlcur) Suol rxc gZ-01 lunrs€qcrppunodruoc pcqeuc:q qcnur uodouE uaJsa.loLf:ul 'etncE 'snoJqelS'uoDces ro csntqo xadp ur t€U'esuq aqt tu epr,\\ruur (80-L'0,{ltsou) I-€'0 'xedE '8uol 'etu J€ouepr,r ruru (g g t ,{nsou) g t-t ruu (t I - 11 ,{psour)/Z-E 9 toe:)uDI ro o1u1nq1ed:^ 'Suol ,ra,1ra? ulurpulru (97-t1 fpsour) gg 0l l€U so^rolJocllcsoJ Ixseq : ururprutrl 1 6 g 6 ruurg 1 'ellq,\\ ruets :(eJuecse.rolJurSurpnlcur) q?rq ulc AZ-01 q./a{ lrnuuu petoor snorqrt V

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y1,1 nror3 arrr21&5 .r.u ..IIt rltr.uri) C) ?9 oll^\o'I V Nnlr.rinVol. 12, No. 2 (1998)

Gordon Uline for providing the interpretationofthe place nameWeeli Wolli; JohnThompspn for his photographsand data on S. weeliwolli at Mt Augustus; Gordon Graham for his companionshipon expeditionsto the Kimberley; Matthew and RussellBanett for their collection and field obseryations of S. dicerdtum; the leadersand membersof the 1996lnndscope Expedition to the Kimberley;Dr Sid Jamesfor the chromosomecounts; Paul Wilson forhis assistancewiththe Latin diagnoses;Dr Barbara Rve for her comments.and the staff of the Westem AustralianHerbarium.

References

Farell, P.G. & James,S.H. (1979). Srylidiun ecoaie (F. Muell. ex Ericksonand willis) comb.et stat. not. (Stylidi^ceae). A strolian Journal o/ Bottnt 27:3945. Mildbraed,J. (1908). .1r.' Engler,A. (ed.) "DasPflanzenreich." Vol. IV, 278 (35). pp. l-98 (Engelmann: L,eipzig.\