Historia naturalis bulgarica, 23: 141-155, 2016

High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016)

Petar Beron

Abstract: After the publication of my monograph (Beron, 2008) appeared many new papers containing data on Arachnida and Myriapoda found in the Old World at or above 2200 m. In this supplement are added 12 sp. of Opilions, 37 sp. of , and 74 sp. of Diplopoda. Several changes occurred in the of some groups, following taxonomic revi- sions. Several genera of are no more in , but in , Fuhrmanodesmidae became synonyme of Trichopolydesmidae, etc. “Palpatores” are now distributed between the suborders and Dyspnoi. Several specialists, and especially Golovatch, published many new taxa of high mountain Diplopoda. Again Golovatch synonymized several genera of Diplopoda (Martensosoma, Nepalomorpha, Parorthomorpha, Armolites, Orophosoma), known from the high mountains, under Delartrum Attems. For many taxa has been “raised” the upper limit of distri- bution and they have been published from other mountains.

Key Words: Arachnida, Myriapoda, high altitude, Old World, supplementa, corrigenda

Underlined are taxa not included in the monograph of Beron (2008)

The mountain systems of the Old World and the Opilio lederi Roewer – up to 3000 m Isopoda, Arachnida and Myriapoda recorded therein O. parietinus (De Geer) – up to 3000 m Pyrenees and the mountains of Iberian (Azerbaijan) Peninsula O. redikorzevi Roewer – 2400 m Diplopoda Phalangium opilio L.– up to 3000 m New ref.: Gilgado et al. (2015) Ph. punctipes L. Koch – up to 3000 m Chordeumatida Araneae Fam. Opisthocheiridae New ref: Logunov & Demir (2006) Ceratosphys cryodeserti Gilgado, Mauriès et Fam. Thomisidae Enghoff – Sierra Nevada Xysticus bacurianensis Mcheidze – 3000 m Alps (Azerbaijan), 2900 m (Georgia) Arachnida Myriapoda Araneae Diplopoda New ref.: Tanasevitch (2011a, 2011b, 2013) New ref.: Golovatch, Evsyukov & Reip (2016) Fam. Fam. Polydesmidae nigripes (Simon) – 1900 – 2580 m Brachydesmus assimilis Lohmander – 2800 m Erigone remota L. Koch – 2650 m B. kalischewskyi Lignau – 2400 m E. tenuimana Simon – up to 2580 m Polydesmus lignaui Lohmander – 2800 m Caucasus Mountains of Turkey, Armenia, Lebanon, Iran, Arachnida Kopet Dagh and Arabian Peninsula Opiliones Arachnida New ref.: Snegovaya (1999, 2004, 2010, 2014) Araneae Fam. Phalangiidae New ref.: Logunov & Demir (2006), Seyyar, Odiellus bieniaszi Kulczynski – up to 3000 m Demir, Topçu & Taşdemir (2006) 142 Petar Beron

Fam. Gnaphosidae Tiso incisus Tanasevitch – 1300-2500 m Phaeocedus braccatus (L. Koch) – 2300 m (Karakorum) (Erciyes) Acari Fam. Thomisidae New ref.: Daniel (1973), Daniel, Stekolnikov, Xysticus bacurianensis Mcheidze – 2200 m Hakimitabar & Saboori (2010), Jing, Solhoy, X. cristatus (Clerck) – 2200 m Wang, Vollan & Xu (2005) Acari, known from the described area at or Mountains of Afghanistan, Pakistan, Karakorum, above 2200 m (suppl.): Tibet, Pamir, Kunlun and Tien Shan (from the border between Iran and Afghanistan to 120o E) Acariformes Prostigmata Arachnida Fam. Trombiculidae Opiliones Microtrombicula gratiosa Schluger et New or ommited ref.: Staręga (1986), Kudryashova – Hindu Kush, 2950 m Staręga & Snegovaya (2008a) M. similata Schluger et Amanguliev – Hindu Eupnoi Kush, 3080 m Fam. Phalangiidae M. vacillata Fernandes et Kulkarni – Hindu Homolophus gricenkoi Staręga et Snegovaya – Kush, 3080 m 2000 -3000 m (Tadjikistan) Neotrombicula lubrica Kudryashova – Hindu H. andreevae Staręga et Snegovaya – up to Kush, 2950-3990 m 4670 m (Pamir) Leptotrombidium silvaticum Hushcha et Pamiropilio naukat Staręga et Snegovaya – 3000- Schluger – Hindu Kush, 2782 m 3300 m (Kyrgyzstan, Alayskiy Hrebet) Montivagum dihumerale (Traub et Nadchatram) Dyspnoi – Hindu Kush, 3990 m Fam. Nemastomatidae Mediostoma ( ?) pamiricum Staręga, 1986- M. latum (Schluger et Kudryashova) – Hindu 3100 m (Pamir) Kush, 2950-3990 m Araneae M. raropinne (Schluger) – Hindu Kush, 3100 m New ref.: Kok, Lotz & Haddad (2004), Euschoengstia meshhedensis Kudryashova, Tanasevitch (2011b) Neronov et Farhang-Azad – Hindu Kush, 2950 – Fam. Linyphiidae 3990 m Acartauchenius himalayensis Tanasevitch – Brunehaldia lucida Schluger – Hindu Kush, 2500 m (Karakorum) 2950 m Agyneta nigripes (Simon) – 2635 m Shunsennia oudemansi (Schluger) – 2950 – (Karakorum) 4550 m Archaraeoncus prospiciens (Thorell) – 2700- Himalaya 3000 m (Karakorum) (East of Indus and South and West of Tzangpo, Caviphantes pseudosaxetorum Wunderlich – or Brahmaputra) 550-2500 m (Karakorum) New research: Ceratinella wideri (Thorell) – 2500-3100 m B. Petrov collected many different (Karakorum) during his several climbing expeditions in the Gongylidiellum confusum Thaler – 1600-2700 m Himalaya. (Karakorum) Notes on Isopoda, Arachnida and Myriapoda Maso sundevalli (Westring) – 2200 – 2300 m Arachnida (Karakorum) Scorpiones Pelecopsis indus Tanasevitch – 2300 m New ref.: Zambre, Sanap & Mirza (2014) (Karakorum) Fam. Euscorpiidae Piniphantes himalayensis (Tanasevitch) – Scorpiops spitiensis Zambre, Sanap et Mirza – 2800 m (Karakorum) Himalaya, 4200 m Porrhomma pygmaeum (Blackwall) – 2480 m Araneae (Karakorum) New ref.: Jastrzębski, 2010, 2011, Prinerigone vagans (Audouin) – 1700-2700 m Ovtchinnikov, Ahmad & Inayatullah (2008), (Karakorum) Tanasevitch (2011b), Wang & Zhu (2008) cotargus pilosus Simon – 1400-2450 m Some spiders in the Himalaya found at or above (Karakorum) 2200 m after 2008 are: High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 143

Fam. Amaurobiidae Myriapoda Himalmartensus ausobskyi Wang et Zhu – 1770- Chilopoda 3000 m Scolopendromorpha H. martensi Wang et Zhu – 1400-2900 m Fam. Scolopendridae H. nepalensis Wang et Zhu – 1000-2600 m Scolopendra canidens Newport – 2400 m Fam. Gnaphosidae Diplopoda Gnaphosa pakistanica Ovtchinnikov, Ahmad et New ref.: Golovatch (2014, 2015b, 2016)‚ Inayatullah – 4100 m Golovatch, Nzoko Fiemapong & VandenSpiegel Fam. Linyphiidae (2015)‚ Golovatch, VandenSpiegel & Semenyuk Anguliphantes nepalsioides Tanasevitch – 2500- (2016) 2600 m Polydesmida (restructuring) Gongylidiellum confusum Thaler – up to 3200 m Fam. Trichopolydesmidae (syn. Fuhrmanno- (Kashmir) desmidae) Indophantes tonglu Tanasevitch – 2500-2700 m Fam. Paradoxosomatidae Pelecopsis indus Tanasevitch – 2250 m Anoplocephalus martensi Golovatch – up to Tapinocyboides bengalensis Tanasevitch – 3100 m 3600 m () Tiso incisus Tanasevitch – 1900 – 2450 m Beronodesmoides lobatus Golovatch – 4600 m T.(?) indianus Tanasevitch – 2500 – 2600 m (Nepal) Walckenaeria martensi Wunderlich = W. nepa- B. montigena Golovatch 3550 m (Nepal) lensis Wund. (syn. by Tanasevitch, 2011b) B. typicus Golovatch – 3400 m (Nepal) Acari B. anteriporus Golovatch – 3350 m (Nepal) New and ommited ref.: Daniel (1973), Daniel B. martensi Golovatch – 2700 m (Nepal) & Stekolnikov (2009), Daniel, Stekolnikov, Beronodesmus latispinosus Golovatch – up to Hakimitabar & Saboori (2010), Ermilov & 3500 m (Nepal) Martens (2014), Fernandes & Kulkarni (2003) B. longispinosus Golovatch – up to 4270 m Some of the Acari, found in Himalaya at or (Nepal) above 2200 m after 2008, are: B. pallidus Golovatch – 4100 m (Nepal) Acariformes B. gorkhalis Golovatch – up to 3600 m (Nepal) Prostigmata B. latispinosus Golovatch – 3150 m (Nepal) Fam. Trombiculidae B. serratus Golovatch et al. – up to 3500 m Brunehaldia lucida Schluger – 2950 m (Nepal) Cheladonta ikaoensis Sasa et al. – 3450-4000 m B. sinuatospinus Golovatch – 2250 m (Nepal) Doloisia vlastae Daniel et Stekolnikov – 3450 m Delarthrum (= Armolites) chulingensis Euschoengastia meshhedensis Kudryashova et (Golovatch) – up to 3700 m al. – 2950-3990 m D. similis (Golovatch) – up to 2700 m Leptotrombidium angkamii Daniel et D. communicans (Golovatch) – up to 2650 m Stekolnikov – 3450-3900 m D. spiniger (Attems) – 2200 m () L. silvaticum Hushcha et Schluger – 2782 m Hirtodrepanum latigonopum Golovach – up to Microtrombicula gratiosa Schluger et 2700 m Kudryashova – 2950 m Kaschmiriosoma pleuroptera Attems – up to M. similata Schluger et Amanguliev – 3080 m 2800 m (Pakistan) M. vacillata Fernandes et Kulkarni – 3080 m Martensosoma silvestre Golovatch – up to 2600 m Montivagum dihumerale (Traub et Nadchatram) M. schawalleri Golovatch – up to 2150. m – 3990 m M. splendens Golovatch – up to 2150 m M. latum (Schluger et Kudryashova) – 2950-3990 m Delarthrum (= Nepalomorpha) hirsutum M. raropinne (Schluger) – up to 3100 m (Golovatch) – up to 4100 m Neotrombicula kounickyi Daniel et Stekolnikov D. kuznetzovi (Golovatch) – up to 3000 m – 3450-4000 m D. arunensis (Golovatch) – up to 2150 m N. lubrica Kudryashova – 1090-3990 m Delarthrum (= Orophosoma) simulans (Carl) – N. turkestanica Kudryashova – 3990 m up to 3700 m Shunsennia oudemansi (Schluger) – 1800-4100 m D. hingstoni (Carl) – up to 3400 m Trombiculindus mehtai Fernandes et Kulkarni D. fechteri (Golovatch) – up to 3150 m – 3450-4600 m Paranedyopus martensi Golovatch – up to 3600 m 144 Petar Beron

P. similis Golovatch – up to 3000 m Spirobolida P. cylindricus Carl – up to 2850 m Fam. Pachybolidae P. affinis Golovatch – up to 2700 m Trigoniulus brachysternus Attems – 1400 – 2400 P. schawalleri Golovatch – up to 2150 m m () Delarthrum (= Parorthomorpha) tuberculatum Siphonophorida (Golovatch) – up to 3300 m Fam. Siphonophoridae D. spectabilis (Golovatch) – up to 2650 m Teratognathus robusta Attems – 1000 – 2400 m D. tergalis Golovatch – up to 2650 m (Vietnam) D. nyakensis Golovatch – up to 2450 m Polydesmida D. philosophicum Golovatch – up to 2450 m Fam. Paradoxosomatidae Touranella himalayensis Golovatch – up to 2700 m Belousoviella kabaki Golovatch – 3360 m Substrongylosoma montigena Carl – up to (, Sichuan) 2300 m (India) Cleptomorpha sumatrana Golovatch – 1400 – 2600 m () Mountains of Southeast China (south of Gonobelus pentaspinus Golovatch – 2475 m Yang Tse), Thailand, Vietnam, , Philippine (China, Sichuan) Islands, and Hedinomorpha nigra Golovatch – 4000 m Opiliones (China, Sichuan) New ref.: Martens (2016) H. subnigra Golovatch – 3910 m (China, Yunnan) Fam. Nemastomatidae H. montana Golovatch – 3575 m (Yunnan) Sinostoma yunnanicum Martens – 3800 m H. proxima Golovatch – 3570 m (Yunnan) (China, Yunnan) H. yunnanensis Golovatch – 3480 m (Yunnan) Araneae H. reducta Golovatch – 2900 m (Sichuan) New ref.: Dankittipakul & Jocqué (2006), H. jeekeli Golovatch – 2600 m (China, Shaanxi) Zonstein & Marusik (2012) Inversispina erectispina Golovatch – 4036 m Fam. Zodariidae (China, Sichuan), 3130 m (Yunnan, idem) Cydrela pristina Dankittipakul et Jocqué – 1630 Kronopolites biagrilectus Hoffman – 3210 m – 2500 m (Thailand, Doi Inthanon) (China, Yunnan) Fam. Nemesiidae K. semirugosus Golovatch – 2955 m (China, Raveniola montana Zonstein et Marusik – up Sichuan) 4300 m (China, Yunnan) K. rugosus Golovatch – 2400 m (China, Yunnan) R. sangrila Zonstein et Marusik – up 4300 m Sellanucheza typica Golovatch – 3155 m (Sichuan) (China, Yunnan) Sigipinius grahami Hoffman – 4205 m (China, R. songi Zonstein et Marusik – up 4300 m Sichuan) (China, Yunnan) S. simplex Golovatch – 4195 m (China, Sichuan) R. yunnanensis Zonstein et Marusik – up 4300 S. complex Golovatch – 4120 m (China, Sichuan) m (China, Yunnan) S. campanuliformis Golovatch – 3910 m (China, Myriapoda Yunnan) Diplopoda S. dentiger Golovatch – 3570 m (China, Yunnan) New ref.: Golovatch (2015, 2016)‚ Korsós, S. kabaki Golovatch – 3550 m (China, Xinjiang Enghoff & Chang (2008) Uygur Prov.) Diplopoda found at or above 2200 m after 2008: Tectoporus beroni Golovatch – 1400-2600 m Glomerida (Sumatra) Fam. Glomeridae T. aberrans Golovatch – 1400-2600 m (Sumatra) Hyleoglomeris robusta Attems – 1800-2400 m T. beshkovi Golovatch – 1400-2600 m (Sumatra) (Vietnam) Tylopuso schawalleri Golovatch 2700 m (China, H. electa Silvestri – 2300 – 2400 m (Vietnam) Yunnan) Platydesmida Fam. Cryptodesmidae Fam. Platydesmidae Trichopeltis Pocock – up to 2400 m (T. kometis Platydesmus camptotrichus Attems – 700-2400 Attems, Vietnam) m (Vietnam) Fam. Polydesmidae P. variegatus Attems – 1500 – 2400 m Epanerchodus fuscus Golovatch – 3325 m (Vietnam) (China, Yunnan) High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 145

Fam. Xystodesmidae Vandenspiegel – 2600 m (Papua New Guinea) Riukiaria spatuliformes Golovatch – 2525 m Altai, the Mountains of Siberia, Mongolia and (China, Sichuan) the Far East Fam. Cryptodesmidae Araneae Trichopeltis kometis (Attems) – 1500 – 2400 m New ref.: Tanasevitch (2013) (Vietnam) Fam. Linyphiidae Polyzoniida Araeoncus crassiceps (Westring) – 1800-2800 m Fam. Siphonocryptidae (Altai) Siphonocryptus compactus Pocock – up to 2600 A. vorkutensis Tanasevitch – 500-2450 m m, Sumatra (Enghoff & Golovatch, 1995) (Altai) Hirudicryptus taiwanensis Korsós, Enghoff et Bathylinyphia major (Kul.) – 600-2500 m Chang – up to 2200 m, Taiwan, Korsós, Enghoff (Altai) & Chang, 2008) Bolyphantes distichus (Tanasevitch) – up to 2300 m (Altai) Mountains of New Guinea, Bismarck Archipelago Dicymbium facetum (L. Koch) – up to 3800 m and the Solomon Islands (Altai) Chilopoda Erigone arctica maritima Kul. – 2500-3000 m New ref.: Shileyko & Stoev (2016) E. arcticola Chamberlin et Ivie – 2400-3100 m Scolopendromorpha (Altai) Fam. Cryptopidae E. cristatopalpus Simon – up to 2400 m (Altai) Cryptops doriae Pocock – up to 3150 m (PNG) E. remota L. Koch – up to 3100 m (Altai) C. nepalensis Lewis, 1999-2700 m (PNG) Gonatium pacificum Eskov – 2400-3100 m C. spinipes Pocock – 2260-2600 m (PNG) (Altai) Cryptops sp. – 3150 m (PNG) Halorates altaicus Tanasevitch – 2000-2500 m Diplopoda (restructuring) (Altai) New ref.: Golovatch & Stoev (2010, 2011, Hilaila glacialis (Thorell) – 2400-3100 m 2014, 2014a), (Altai) Polydesmida H. meridionalis Tanasevitch – 200-3300 m Fam. Paradoxosomatidae (Altai) Caloma agametum Chamberlin – 2300 m (PNG) Improphantes improbulus (Simon) – 1400-2500 Eustrongylosoma curtipes Golovatch et Stoev – m (Altai) 2300 m (PNG) Incestophantes bonus Tanasevitch – 2100-2800 E. beroni Golovatch et Stoev – 2300 m (PNG) m (Altai) E. finimtel Golovatch et Stoev – [2300 m] (PNG) I. brevilamellus Tanasevitch – 2600-2800 m E. tumbuna Golovatch et Stoev – 2980 m (PNG) (Altai) E. masalai Golovatch et Stoev – [2300 m] (PNG) I. tuvensis Tanasevitch – 2600-3000 m (Altai) E. papua Golovatch et Stoev – [2300 m] (PNG) Islandiana falsifica (Keyserling) – 2400-3100 m E. tifalmin Golovatch et Stoev – [2300 m] (PNG) (Altai) Silvattia jeekeli Golovatch et Stoev – 2300 m Macrargus rufus (Wider) – 2000-2500 m (Altai) (PNG) Mughiphantes sobrioides Tanasevitch – 2600- Tectoporus fugilil Golovatch et Stoev – 2600 m 2800 m (Altai) (PNG) Oedothorax mongolensis (Heimer) – 2000- T. bispinosus Golovatch et Stoev – 2600 m (PNG) 800 m (Altai) T. moniliformis Golovatch et Stoev – 2600 m Oreoneta eskovi Saaristo et Marusik – 1700- (PNG) 2800 m (Altai) T. spiniger Golovatch et Stoev – 2300 m (PNG) Pelecopsis dorniana Heimer – 1600-2200 m T. wilhelmicus Golovatch et Stoev – 3610 m (Altai) (PNG) P. palmgreni Marusik et Esyunin – 1400-2200 Fam. Opisotretidae m (Altai) Opisotretus beroni Golovatch, Geoffroy, Stoev Porrhomma pallidum Jackson – 2100-2500 m et Vanden Spiegel – Papua New Guinea (Altai) Fam. Cryptodesmidae Scotinotylus antennatus (O. Pickard-Cambridge) Astrolabius hoffmani Golovatch, Stoev et – 2000-2300 m (Altai) 146 Petar Beron

S. protervus (L. Koch) – 1800-3000 m (Altai) Orobothriurus Maury, 1976 and made a list Semljicola angulatus (Holm) – 1650-2200 m of the Scorpion species recorded above 3000 m (Altai) in the Andes. This list contains 39 sp. of the gen- S. latus (Holm) – 1350-2200 m (Altai) era Orobothriurus, Bothriurus, Pachkutej, and Silometopus uralensis Tanasevitch – 1700 – Brachistosternus (Bothriuridae, up to 4910 m in 2200 m (Altai) Peru), Tityus (Buthidae, up to 4200 m in Bolivia), Stemonyphantes altaicus Tanasevitch – 1300- Hadruroides (Iuridae, up to 3379 m in Peru) and 2500 m (Altai) Teuthraustes (Chactidae, 3300 m in Ecuador). Tenuiphantes suborientalis Tanasevitch – 2200- According to these authors, the former world maxi- 2500 m (Tuva) mum altitude, recorded for “Orobothriurus” crassi- Thyreostenius parasiticus (Westring) – 500- manus – actually Pachakutej crassimanus (Maury, 2200 m (Altai) 1976) by Lourenço (1997) – 5560 m at Nevado Typhlochrestoides baikalensis Eskov – 1600- Huascaran (Peru) – is based on misidentification 2200 m (Altai) and errouneus locality data. These authors accept Walckenaeria capito (Westring) – 1700-2500 m as highest record of scorpions the newly described (Altai) Orobothriurus huascaran at 4910 m in Peru. In 1977 W. katanda Marusik, Hippa et Koponen – 1500- I have visited the area above Paron lake and climbed 2200 (Altai) on the slopes of Huandoy over 5000 m. The environ- W. esyunini Tanasevitch – 2600 m (Altai) ment was free of snow and ice, with forests of Polylepis and it is quite possible that scorpions will be found Mountains in West Equatorial Africa higher than 5000 m. In Asia scorpions have been Opiliones found (with certainty) up to 4600 m (Tibetiomachus New ref.: Staręga & Snegovaya (2008b) himalayensis Lourenço et Qi, 2006, Liochelidae), in Fam. Phalangiidae Africa – up to 3500 m (Uroplectes fischeri Tullgren, Camerobunus okucola Staręga et Snegovaya – 1910, Meru, Buthidae). 2200-2800 m (Cameroon) PSEUDOSCORPIONES found in the Old Diplopoda World at or above 2200 m and the highest found New ref.: Golovatch, Nzoko Fiemapong & Pseudoscorpions in the World (supplementa) VandenSpiegel (2015) No new information Fam. Pyrgodesmidae Monachodesmus armorum Golovatch et al. – OpilionES in the Old World known at or above 2320 m (Cameroon) 2200 m and the highest living Opiliones in the Udodesmus camerunensis Golovatch et al. – World (supplementa) 2320 m (Cameroon) New ref.: Das & Bastawade (2006), Martens Urodesmus cornutus Golovatch et al. – 2320 m (2016), Martens & Chemini (1988), Snegovaya (Cameroon (1999, 2004, 2010, 2014), Staręga (1986), Staręga ISOPODA in the Old World known at or above & Snegovaya (2008a, 2008b) 2200 m and the highest living Isopoda in the Eupnoi World (supplementa) Fam. Phalangiidae New ref.: no new information Camerobunus Staręga et Snegovaya – up to 2800 Arachnida m (C. okucola Staręga et Snegovaya, Cameroon) SCORPIONES in the Old World known at or Homolophus Banks (Euphalangium Roewer) – above 2200 m and the highest living Scorpiones in up to 4670 m (H. andreevae Staręga et Snegovaya, the World (supplementa) Pamir), 3000 m (H. gricenkoi Staręga et Snegovaya, New ref.: Ochoa et al. (2011), Zambre, Sanap Tadjikistan), 3000 m (H. [Euphalangium] martensi & Mirza (2014) (Das et Bastawade), Indian Himalaya) Order Scorpiones. Maximal altitude (correct- Odiellus Roewer – up to 3000 m (O. bieniaszi ed): 4910 m in Peru, 4600 m in the Himalaya. Kulczynski, Caucasus) Fam. Euscorpiidae, Scorpiopinae Opilio Herbst – up to 3000 m (O. lederi Roewer), Scorpiops Peters – up to 4200 m (S. spitiensis 2400 m (O. redikorzevi Roewer)(Caucasus) Zambre, Sanap et Mirza, Himalaya) Pamiropilio Staręga et Snegovaya – up to 3300 Ochoa, Ojanguren Affilastro, Mattoni m (P. naukat Staręga et Snegovaya, Kyrgyzstan, & Prendini (2011) described six new species of Alayskiy Hrebet) High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 147

Phalangium L.– up to 3000 m (Ph. punctipes L. Odontobunus armatus (Sörensen)(Phalangiidae) Koch, Caucasus) – 4200 m (Kenya), 4000 m (Kilimanjaro) Dyspnoi Opilio almasyi Roewer (Phalangiidae) – 4200 m Fam. Nemastomatidae (Karakorum) Mitostoma Roewer – up to 3100 m (Mitostoma O. nigridorsum di Caporiacco (Phalangiidae) – ? – Pamir) 4200 m (Karakorum) Sinostoma Martens – up to 3800 m (S. Ereca undulata Sörensen (Assamiidae) – 4025 m yunnanicum Martens, China, Yunnan) (Ruwenzori) Fam. Trogulidae E. simulator Sörensen (Assamiidae) – 4000 m Anelasmocephalus Simon – up to 2450 m (A. (Kilimanjaro) osellai Martens et Chemini, Appenines), 2200 m (A. Metaereca abnormis Roewer (Assamiidae) – cambridgei Westwood, Alps) 4000 m (Ruwenzori) Hypoxestus patellaris Sörensen (Assamiidae) – Species of Opiliones, known from the Old World 4000 m (Ruwenzori) at and above 3500 m (restructuring of the list) Metabiantes punctatus Sörensen (Biantidae) – Homolophus Banks (syn. Euphalangium) 4000 m (Kilimanjaro) nordenskioeldi (L. Koch)(Phalangiidae) – 5600 m Guruia africana Karsch (Phalangiidae) – 4000 (Karakorum) m (Kilimanjaro) Himalphalangium palpale Roewer G. frigescens Loman (Phalangiidae) – 4000 m (Phalangiidae) – 5540 m (Nepal) (East Africa) Homolophus panpema Suzuki (Phalangiidae) – Rhampsinitus ? mesomelas Sörensen 5200 m (Nepal) (Phalangiidae) – 4000 m (Kilimanjaro) H. luteum Suzuki (Phalangiidae) – 5200 m Ereca maculata Roewer (Assamiidae) – 3975 m (Nepal) (Kilimanjaro) Octozaleptus harai Suzuki (Sclerosomatidae) – Dhaulagirius altitudinalis Martens 5200 m (Nepal) (Phalangodidae) – 3950 m (Nepal) Leiobunum mirum Roewer (Phalangiidae) – Cristina pachylomera Simon (Phalangiidae) – 5200 m (Nepal) 3870 m (Ruwenzori; 3658 m, Semien) Sabaconidae gen. sp. – > 5000 m (Nepal) Rhampsinitus discolor Karsch (Sclerosomatidae) ?Opilio sp. (Phalangiidae) – 4800 m – 3870 m (Ruwenzori) (Karakorum) Himaldroma altus Martens (Sclerosomatidae) – Homolophus andreevae Staręga et Snegovaya 3830 m (Nepal) (Phalangiidae) – 4670 m (Pamir) Rhampsinitus salti Roewer (Phalangiidae) – Hypoxestus accentuatus Sörensen (Assamiidae) 3800 m (Kilimanjaro) – 4600 m (Kilimanjaro) Gyoides maximus Martens (Sclerosomatidae) – Rhampsinitus bettoni Pocock (Sclerosomatidae) 3800 m (Nepal) – 4600 m (Kilimanjaro) Sinostoma yunnanicum Martens (Nemasto- ?Phalangium sp. (Phalangiidae) – 4500 m matidae) – 3800 m (Yunnan) (Karakorum) Odontobunus africanus Roewer (Phalangiidae) Egaenus (= Diabunus) laevipes (di Caporiacco) – 3770 m (Kivu) (Phalangiidae) – 4400 m (Karakorum) Gyoides gandaki Martens (Sclerosomatidae) – Sabacon dhaulagiri Martens (Sabaconidae) – 3760 m (Nepal) 4250 m (Nepal) Bunochelis spinifera Lucas (Phalangiidae) – Biantes pernepalicus Martens (Biantidae) – 4250 3711 m (Tenerife) m (Nepal) Eudasylobus infuscatus Lucas (Phalangiidae) – Himalphalangium dolpoense Martens 3650 m (Atlas) (Phalangiidae) – 4200 m (Nepal) Rilaena triangularis Herbst (Phalangiidae) – Micrassamula thak Martens (Assamiidae) – 3650 m (Atlas) 4200 m (Nepal) Biantes thakkhali Martens (Biantidae) – 3640 m Hypoxestus holmi Goodnight et Goodnight (Nepal) (Assamiidae) – 4200 m (East Africa) Randilea scabricula Roewer (Assamiidae) – Gyoides himaldispersus Martens 3630 m (Elgon) (Sclerosomatidae) – 4200 m (Nepal) Metabiantes trifasciatus Roewer (Biantidae) – 148 Petar Beron

3600 m (Meru) m (H. ausobskyi Wang et Zhu, 2900 m (H. martensi Mitopus glacialis Heer (Phalangiidae) – 3600 m Wang et Zhu), 2600 m (H. nepalensis Wang et Zhu), ( Alps) all from Nepal Dacnopilio scopulatus Lawrence (Phalangiidae) Fam. Linyphiidae – 3600 m (Meru) Acartauchenius Simon – up to 2500 m (A. hi- Egaenus tibetanus Roewer (Phalangiidae) – malayensis Tanasevitch, Karakorum) 3600 m (Karakorum) Agyneta Hull – up to 2635 m (A. nigripes Simon, Harmanda medioimmicans Martens (Scleroso- Karakorum) matidae) – 3600 m (Nepal) Anguliphantes Saaristo et Tanasevitch – up to Simienatus scotti Roewer (Assamiidae) – 3505 2600 m (A. nepalsioides Tanasevitch) m (Semien, Ethiopia) Araeoncus Simon – up to 2800 m (A. crassiceps Metabiantes convexus Roewer (Biantidae) – Westring, Altai), 2450 m (A. vorkutensis Tanasevitch, 3500 m (Ruwenzori) Altai) Ereca lata Sörensen (Assamiidae) – 3500 m Archaraeoncus Tanasevitch – up tp 3000 m (A. (Kilimanjaro) prospiciens Thorell, Karakorum) E. affinis Sörensen (Assamiidae) – 3500 m Bathylinyphia Eskov – up to 2500 m (B. major (Kilimanjaro) Kul., Altai) E. modesta Sörensen (Assamiidae) – 3500 m Bolyphantes C.L. Koch – up to 2300 m (B. dis- (Kilimanjaro) tichus Tanasevitch, Altai) Hypoxestus mesoleucus Sörensen (Assamiidae) Caviphantes Oi – up to 2500 m (C. pseudosaxe- – 3500 m (Kilimanjaro) torum Wunderlich, Karakorum) Harmanda latephippiata Martens (Scleroso- Ceratinella Emerton – up to 3100 m (C. wideri matidae) – 3500 m (Nepal) Thorell, Karakorum) H. nigrolineata Martens (Sclerosomatidae) – Dicymbium Menge – up to 3800 m (D. facetum 3500 m (Nepal) L. Koch, Altai) Pokhara yodai Suzuki (Sclerosomatidae) – 3500 Erigone Audoiin – up to 3100 m (E. arcticola m (Nepal) Chamberlin et Ivie, Altai; E. remota L. Koch, Altai; Rhampsinitus soerenseni Staręga (= Rh. pictus 2650 m, Alps), 3000 m (E. arctica maritima Kul., Sörensen)(Phalangiidae) – 3500 m (Meru) Altai), 2400 m (E. cristatopalpus Simon, Altai) Gonatium Menge – up to 3100 m (G. pacificum Maximal altitudes of Harvestmen (Opiliones), Eskov, Altai) living above 2200 m in the Old World Gongylidiellum Simon – up to 3200 m (G. confu- 1 – Triaenonychidae – up to 2500 m sum Thaler, Kashmir), 2700 m, Karakorum) 2 – Oncopodidae – up to 2600 m Halorates Hull – up to 2500 m (H. altaicus 3 – Podoctidae – up to 2410 m Tanasevitch, Altai) 4 – Biantidae – up to 4250 m Hilaila Simon – up to 3300 m (H. meridiona- 5 – Assamiidae – up to 4600 m lis Tanasevitch, Altai), 3100 m (H. glacialis (Thorell, 6 – Phalangiidae – up to 5600 m Altai) 7 – Sclerosomatidae – up to 5200 m Indophantes Tanasevitch et Saaristo – up to 8 – Ischyropsalididae – up to 2700 m 2700 m (I. tonglu Tanasevitch (Himalaya) 9 – Sabaconidae – more than 5000 m Improphantes Saaristo et Tanasevitch – up to 10 – Nemastomatidae – up to 3800 m 2500 m (I. improbulus Simon, Altai) ARANEAE found in the Old World at or above Incestophantes Tanasevitch – up to 3000 m 2200 m and the highest (I. tuvensis Tanasevitch, Altai), 2800 m (I. bonus found Araneae in the World (supplementa) Tanasevitch, I. brevilamellus Tanasevitch, Altai) New ref.: Dankittipakul & Jocqué (2006), Islandiana Braendegaard – up to 3100 m (I. fal- Logunov & Demir (2006), Jastrzębski, 2010a, sifica Keyserling, Altai) 2010b, 2011, Ovtchinnikov, Ahmad & Inayatullah Macrargus Dahl – up to 2500 m (M. rufus (2008), Seyyar, Demir, Topçu & Taşdemir (2006), Wider, Altai) Tanasevitch (2011b, 2013), Zonstein & Marusik Maso Simon – up to 2300 m (M. sundevalli (2012) Westring, Karakorum) Fam. Amaurobiidae Mughiphantes Saaristo et Tanasevitch – up to Himalmartensus Wang et Zhu – up to 3000 2800 m (M. sobrioides Tanasevitch, Altai) High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 149

Oedothorax Bertkau – up to 2800 m (Oe. mon- Phaeocedus Simon – up to 2300 m (Ph. bracca- golensis Heimer, Altai) tus L. Koch, Erciyes, Turkey) Oreoneta Kulczynski – up to 2800 m (O. eskovi Fam. Thomisidae Saaristo et Marusik, Altai) Xysticus Walckenaer – up to 3000 m (X. bacu- Pelecopsis Simon – up to 2300 m (P. indus rianensis Mcheidze, Azerbaijan), 2900 m (Georgia), Tanasevitch, Karakorum), 2200 m (P. dorniana 2200 m (X. cristatus Clerck, Turkey) Heimer, P. palmgreni Marusik et Esyunin, Altai) Species of Araneae, known from the Old World at Piniphantes Saaristo et Tanasevitch – up to 2800 and above 3500 m (supplement to the list) m (P. himalayensis Tanasevitch, Karakorum) Raveniola montana Zonstein et Marusik Porrhomma Simon – up to 2500 m (P. pallidum (Nemesiidae) – 4300 m (China, Yunnan) Jackson, Altai), 2480 m (P. pygmaeum Blackwall, R. shangrila Zonstein et Marusik (Nemesiidae) Karakorum) – 4300 m (China, Yunnan) Prinerigone Wunderlich – up to 2700 m (P. va- R. songi Zonstein et Marusik (Nemesiidae) – gans Audouin, Karakorum) 4300 m (China, Yunnan) Scotargus Simon – up to 2450 m (S. pilosus R. yunnanensis Zonstein et Marusik Simon, Karakorum) (Nemesiidae) – 4300 m (China, Yunnan) Scotinotylus Simon – 3000 m (S. protervus Gnaphosa pakistanica Ovtchinnikov, Ahmad et L. Koch, Altai), 2300 m (S. antennatus O. Pickard- Inayatullah (Gnaphosidae) – 4100 m (Pakistan) Cambridge, Altai) Semljicola Strand – 2200 m (S. angulatus Holm, ACARI found in the Old World at or above 2200 S. latus Holm Altai) m and the highest found Acari in the World Silometopus Simon – up to 2200 m (S. uralensis (supplementa) Tanasevitch, Altai) New or ommited ref.: Daniel (1973, Stemonyphantes Menge – 2500 m (S. altaicus 2015), Daniel & Stekolnikov (2009), Daniel, Tanasevitch, Altai) Stekolnikov, Hakimitabar & Saboori (2010), Tapinocyboides Wiehle – up to 3100 m (T. ben- Ermilov & Martens (2014), Fernandes & galensis Tanasevitch) Kulkarni (2003) Tenuiphantes Tanasevitch – up to 2500 m (T. Acariformes suborientalis Tanasevitch, Tuva) Sarcoptiformes Thyreostenius Simon – up to 2200 m (Th. para- Acaridida – correction: the highest Prostigmata siticus Westring, Altai) found is on 3500 m (Himalaya) Tiso Simon – up to 2500 m (T. incisus Oribatida – correction: the highest Oribatida is Tanasevitch, Karakorum) found on 5800 m (Himalaya) T.(?) indianus Tanasevitch – 2500 – 2600 m Fam. Scheloribatidae Typhlochrestoides Eskov – up to 2200 m (T. Perscheloribates Hammer – up to 2800 m (P. baikalensis Eskov, Altai) nepalensis Ermilov et Martens, Nepal) Walckenaeria Blackwall – up to 2600 m (W. Trombidiformes esyunini Tanasevitch, Altai), 2500 m (W. capito Prostigmata – correction: the highest Westring, Altai), 2200 m (W. katanda Marusik, Prostigmata is found on 5100 m (Himalaya) Hippa et Koponen, Altai) Fam. Trombiculidae Walckenaeria martensi Wunderlich = W. nepa- Aboriginesia Kudryashova becomes synonyme lensis Wund. (syn. by Tanasevitch, 2011b) of Xinjiangsha Wen et Shao Fam. Zodariidae Brunehaldia Vercammen-Grandjean et Cydrela Thorell – up to 2500 m (C. pristina Kolebinova – up to 2950 m (B. lucida Schluger, Dankittipakul et Jocqué, Thailand, Doi Inthanon) Hindu Kush) Fam. Nemesiidae Cheladonta Lipovsky, Crossley et Loomis – up Raveniola Zonstein – up to 4300 m (R. mon- to 4000 m (Ch. ikaoensis Sasa et al., Nepal) tana Zonstein et Marusik, R. sangrila Zonstein et Doloisia Oudemans – up to 3450 m (D. vlastae Marusik, R. songi Zonstein et Marusik, R. yunnanen- Daniel et Stekolnikov, Nepal) sis Zonstein et Marusik, China, Yunnan) Euschoengastia Ewing – up to 3990 m (E. mes- Fam. Gnaphosidae hhedensis Kudryashova et al., Hindu Kush) Gnaphosa Latreille – up to 4100 m (G. pakistani- Leptotrombidium Nagayo et al. – up to 3900 m ca Ovtchinnikov, Ahmad et Inayatullah, Pakistan), (L. angkamii Daniel et Stekolnikov, Nepal), 2782 m 150 Petar Beron

(L. silvaticum Hushcha et Schluger, Hindu Kush) Chordeumatida Microtrombicula Ewing – up to 3080 m (M. sim- Fam. Opisthocheiridae ilata Schluger et Amanguliev, M. vacillata Fernandes Ceratosphys cryodeserti Gilgado, Mauriès et et Kulkarni), 2950 m (M. gratiosa Schluger et Enghoff – (Sierra Nevada) Kudryashova, all from Hindu Kush) Platydesmida Montivagum Kudryashova – up to 3990 m (M. Fam. Platydesmidae dihumerale Traub et Nadchatram, M. latum (Schluger Platydesmus Cook – up to 2400 m (P. et Kudryashova)(Hindu Kush), 3100 m (M. raropin- camptotrichus Attems, P. variegatus Attems, ne (Schluger, Hindu Kush) Vietnam) Neotrombicula Hirst – up to 4000 m (N. kou- Polydesmida nickyi Daniel et Stekolnikov, Nepal), 3990 m (N. Fam. Fuhrmannodesmidae becomes synonyme lubrica Kudryashova, Hindu Kush, N. turkestanica of Trichopolydesmidae (after Golovatch, 2013) Kudryashova) Fam. Paradoxosomatidae – up to 4600 m Shunsennia Jameson et Toshioka – up to 4550 (Nepal) m (Shunsennia oudemansi Schluger, Hindu Kush; Anoplocephalus Schaeffer – up to 3600 m (A. 4100 m, ) martensi Golovatch, Nepal) Trombiculindus Radford – up to 4600 m (T. me- Belousoviella Golovatch – up to 3360 m (B. htai Fernandes et Kulkarni, Nepal) kabaki Golovatch, Sichuan) Species of Prostigmata (Trombidiformes), Beronodesmoides Golovatch – up to 4600 m (B. known from the Old World at and above 3500 m lobatus Golovatch, Nepal), 3550 m (B. montigena (supplement to the list) Golovatch, Nepal), 3400 m (B. typicus Golovatch, Trombiculindus mehtai Fernandes et Kulkarni Nepal), 3350 m (B. anteriporus Golovatch, Nepal), (Trombiculidae) – 4600 m (Nepal) 2700 m (B. martensi Golovatch, Nepal) Shunsenia oudemansi Schluger (Trombiculidae) Beronodesmus Golovatch – up to 4270 m (B. – 4100 m (Hindu Kush) longispinosus Golovatch, Nepal), 4100 m (B. pallidus Neotrombicula kounickyi Daniel et Stekolnikov Golovatch, Nepal), 3600 m (B. gorkhalis Golovatch, (Trombiculidae) – 4000 m (Himalaya) Nepal), 3500 m (B. serratus Golovatch et al., B. lat- Ch. ikaoensis Sasa et al. (Trombiculidae) – 4000 ispinosus Golovatch et al., Nepal), 2650 m (B. mar- m (Nepal) tensi Golovatch et al., Nepal), 2250 m (B. sinuat- Euschoengastia meshhedensis Kudryashova et ospinus Golovatch, Nepal) al. (Trombiculidae) – 3990 m (Hindu Kush) Caloma Chamberlin – up to 2300 m (C. agame- Montivagum dihumerale Traub et Nadchatram tum Chamberlin, Papua New Guinea) (Trombiculidae) – 3990 m (Hindu Kush) Cleptomorpha Golovatch – up to 2600 m (C. su- M. latum (Schluger et Kudryashova) matrana Golovatch, Sumatra) (Trombiculidae) – 3990 m (Hindu Kush) Delarthrum Attems (syn. Nepalomorpha Leptotrombidium angkamii Daniel et Golovatch, Parorthomorpha Golovatch, Martensosoma Stekolnikov(Trombiculidae) – 3900 m (Nepal) Golovatch, Armolites Golovatch, Orophosoma Jeekel) – up to 3250 m (D. tenuitergale Golovatch, Nepal), DIPLOPODA found in the Old World at or above 3100 m (D. typicum Golovatch, Nepal); according to 2200 m and the highest found Diplopoda in the Golovatch (2014), in the list in Beron (2008) should World (supplement) be changed the following species: New Ref.: Enghoff & Golovatch (1995), Delartrum [Martensosoma] silvestre (Golovatch), Golovatch (2013, 2014, 2015a, 2015b, 2016), Delartrum [Martensosoma] schawalleri (Golovatch), Golovatch, Evsyukov & Reip (2016), Golovatch Delartrum [Martensosoma] splendens (Golovatch), & Stoev (2009, 2010, 2011, 2014a, 2014b), Delartrum [Nepalomorpha] hirsutum (Golovatch), Golovatch, VandenSpiegel & Semenyuk (2016)‚ Delartrum [Nepalomorpha] kuznetzovi (Golovatch), Golovatch, Nzoko Fiemapong & VandenSpiegel Delartrum [Nepalomorpha] arunense (Golovatch), (2010, 2015)‚ Korsós, Enghoff & Chang (2008), Delartrum [ Parorthromorpha] tuberculatum (Golovatch), Nefediev, Nefedieva & Jankowski (2015) Delartrum [Parorthromorpha] spectabile (Golovatch, Glomerida 2650 m, Nepal, Golovatch, 2016), Delartrum Fam. Glomeridae [Parorthromorpha] tergale (Golovatch), Delartrum Hyleoglomeris Verhoeff – up to 2400 m (H. [Parorthromorpha] nyakense (Golovatch), Delartrum robusta Attems, H. electa Silvestri, Vietnam) [Parorthromorpha] philosophica (Golovatch), Delartrum High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 151

[Armolites] chulingensis (Golovatch), Delartrum Trichopeltis Pocock – up to 2400 m (T. kometis [Armolites] simile (Golovatch), Delartrum [Armolites] Attems, Vietnam) communicans (Golovatch), Delartrum [Armolites] spini- Fam. Pyrgodesmidae ger (Attems, India), Delartrum [Orophosoma] simu- Monachodesmus Silvestri – up to 2320 m (M. lans (Carl), Delartrum [Orophosoma] hingstoni (Carl), armorum Golovatch et al., Cameroon) Delartrum [Orophosoma] fechteri (Golovatch) (all oth- Udodesmus Cook – up to 2320 m (U. camerun- ers from Nepal) ensis Golovatch et al., Cameroon) Eustrongylosoma Silvestri – up to 2980 m (E. Urodesmus Porat – up to 2320 m (U. cornutus tumbuna Golovatch et Stoev), 2300 m (P. beroni Golovatch et al., Cameroon) Golovatch et Stoev, E. curtipes Golovatch et Stoev, Fam. Polydesmidae Golovatch et Stoev, E. masalai Golovatch et Stoev, E. Brachydesmus Heller – up to 2800 m (B. assi- finimtel Golovatch et Stoev, E. maculatum Golovatch milis Lohmander, 1936, Caucasus), 2400 m (B. ka- et Stoev, E. papua Golovatch et Stoev, E. tifalmin lischewskyi Lignau, Caucasus) Golovatch et Stoev), all from Papua New Guinea Epanerchodus Attems – up to 3325 m (E. fuscus Gonobelus Attems – up to 2475 m (G. pen- Golovatch, China, Yunnan) taspinus Golovatch, China, Sichuan) Polydesmus Latreille – up to 2800 m (P. lignaui Hedinomorpha Verhoeff – up to 4000 m (H. ni- Lohmander, Caucasus) gra Golovatch, China, Sichuan), 3910 m (H. subnigra Fam. Xystodesmidae Golovatch, China, Yunnan), 3575 m (H. montana Riukiaria Attems – up to 2525 m (R. spatuli- Golovatch, Yunnan), 3570 m (H. proxima Golovatch, formes Golovatch, China, Sichuan) Yunnan), 3480 m (H. yunnanensis Golovatch, Fam. Opisotretidae Yunnan), 2900 m (H. reducta Golovatch, Sichuan), Opisotretus beroni Golovatch, Geoffroy, Stoev 2600 m (H. jeekeli Golovatch, China, Shaanxi) et Vanden Spiegel (Papua New Guinea) Hirtoderpanum Golovatch – up to 2600 m (H. Order Polyzoniida latigonopum Golovatch, Nepal) Fam. Siphonocryptidae Inversispina Zhang – up to 4036 m (I. erectispina Siphonocryptus Pocock – up to 2600 m (S. com- Golovatch, China, Sichuan), 3130 m (Yunnan, idem) pactus Pocock (Sumatra, Enghoff & Golovatch, Kronopolites Attems – up to 3210 m (K. biagri- 1995) lectus Hoffman, China, Yunnan), 2955 m (K. semiru- Hirudicryptus Enghoff et Golovatch – up to gosus Golovatch, China, Sichuan), 2400 m (K. rugo- 2200 m (H. taiwanensis Korsós, Enghoff et Chang sus Golovatch, China, Yunnan) (Taiwan, Korsós, Enghoff & Chang, 2008) Sellanucheza Enghoff, Golovatch et Nguyen Species of DIPLOPODA, known from the Old Duc – up to 3155 m (S. typica Golovatch, Sichuan) World at and above 3500 m (restructuring of the Sigipinius Hoffman – up to 4205 m (S. graha- list in Beron, 2008) mi Hoffman, China, Sichuan), 4195 m (S. simplex Golovatch, China, Sichuan), 4120 m (S. complex Nepalmatoiulus ivanloebli Enghoff (Julida, Golovatch, China, Sichuan), 3910 m (S. campanuli- Julidae) – 4800 m (Nepal) formis Golovatch, China, Yunnan), 3570 m (S. den- Beronodesmoides lobatus Golovatch tiger Golovatch, China, Yunnan), 3550 m (S. kabaki (Polydesmida, Paradoxosomatidae) – 4600 m Golovatch, China, Xinjiang Uygur Prov.) (Nepal) Tectoporus Carl – up to 3610 m (T. wilhelmicus Uixenus sp. (Polyxenida, Polyxenidae) – 4550 Golovatch et Stoev, Papua New Guinea), 2980 m (T. m (Nepal) fugilil Golovatch et Stoev, Papua New Guinea), 2600 Hingstonia variata Golovatch (Polydesmida, m (T. aberrans Golovatch, T. beroni Golovatch, T. Trichopolydesmidae = Fuhrmannodesmidae) – 4500 beshkovi, Sumatra; T. fugilil Golovatch et Stoev, T. m (Nepal) bispinosus Golovatch et Stoev, T. moniliformis Papua Anaulaciulus niger Korsós (Julida, Julidae) – New Guinea), 2300 m (T. spiniger Golovatch et Stoev, 4500 m (Nepal) Papua New Guinea) A. bilineatus Korsos (Julida, Julidae) – 4300 m Tylopus Jeekel – up to 2700 m (T. schawalleri (Nepal) Golovatch, China, Yunnan) Beronodesmus longispinosus Golovatch Fam. Cryptodesmidae (Polydesmida, Paradoxosomatidae) – 4270 m (Nepal) Astrolabius Verhoeff – up to 2600 m (A. hoffmani Usbekodesmus sp. (Polydesmida, Polydesmidae) Golovatch, Stoev et Vandenspiegel, Papua New Guinea) – 4250 m (Nepal) 152 Petar Beron

Sigipinius grahami Hoffman (Polydesmida, H. serrata Golovatch (Polydesmida, Paradoxosomatidae) – up to 4205 m (China, Trichopolydesmidae = Fuhrmannodesmidae) – 3600 Sichuan) m (Nepal) Sigipinius simplex Golovatch (Polydesmida, Tectoporus wilhelmicus Golovatch et Stoev Paradoxosomatidae) – 4195 m (China, Sichuan) (Polydesmida, Paradoxosomatidae) – 3610 m (Papua Sphaeroparia petarberoni Mauriès et New Guinea) Heymer (Polydesmida, Trichopolydesmidae = Beronodesmus gorkhalis Golovatch (Polydesmida, Fuhrmannodesmidae) – 4200 m (Ruwenzori) Paradoxosomatidae) – 3600 m (Nepal) Sigipinius complex Golovatch (Polydesmida, Anoplocephalus martensi Golovatch (Polydesmida, Paradoxosomatidae) – 4120 m (China, Sichuan) Paradoxosomatidae) – 3600 m (Nepal) Beronodesmus pallidus Golovatch (Polydesmida, Dolichoiulus canariensis Brölemann (Julida, Paradoxosomatidae) – 4100 m (Nepal) Julidae) – 3600 m (Nepal) Delarthrum (= Nepalomorpha) hirsutum Nepalmatoiulus mauriesi Enghoff (Julida, (Golovatch) (Polydesmida, Paradoxosomatidae) – Julidae) – 3600 m (Nepal) 4100 m (Nepal) Polydesmus (Polydesmida, Polydesmidae) – Kashmireuma nepalensis Mauriès (Chordeumatida, 3600 m (Tenerife) Kashmireumatidae) – 4100 m (Nepal) Hedinomorpha montana Golovatch (Polydesmida, Nepalella sp. (Chordeumatida, Megalotylidae) Paradoxosomatidae) – 3575 m (China, Yunnan) – 4100 m (Nepal) Hedinomorpha proxima Golovatch (Polydesmida, Inversispina erectispina Golovatch (Polydesmida, Paradoxosomatidae) – 3570 m (China, Yunnan) Paradoxosomatidae) – 4036 m (China, Sichuan) Sigipinius kabaki Golovatch (Polydesmida, Hedinomorpha nigra Golovatch (Polydesmida, Paradoxosomatidae) – 3550 m (China, Xinjiang Paradoxosomatidae) – 4000 m (China, Sechuan) Uygur Prov.) Usbekodesmus sacer Golovatch (Polydesmida, Beronodesmoides montigena Golovatch Polydesmidae) – 4000 m (Nepal) (Polydesmida, Paradoxosomatidae) – 3550 m (Nepal) Hingstonia gogonema Golovatch (Polydesmida, Beronodesmus serratus Golovatch, Trichopolydesmidae = Fuhrmannodesmidae) – 4000 VandenSpiegel et Semenyuk (Polydesmida, m (Buthan) Paradoxosomatidae) – 3500 m (Nepal) Beronodesmus latispinosus Golovatch Anaulaciulus acaudatus Korsós (Julida, Julidae) (Polydesmida, Paradoxosomatidae) – 3500 m (Nepal) – 3990 m (Sikkim) Nepalella deharvengi Mauriès (Chordeumatida, Sigipinius campanuliformis Golovatch Megalotylidae) – 3500 m (Nepal) (Polydesmida, Paradoxosomatidae) – 3910 m (China, Stemmiulus sjostedti Attems (Stemmiulida, Yunnan) Stemmiulidae) – 3500 m (Meru) Hedinomorpha subnigra Golovatch (Polydesmida, Hingstonia beatae Golovatch (Polydesmida, Paradoxosomatidae) – 3910 m (China, Yunnan) Trichopolydesmidae = Fuhrmannodesmidae) – 3500 Tianella daamsae Shear (Chordeumatida, m (Nepal) Cleidogonidae) – 3900 m (Nepal) Sphaeroparia minuta Attems (Polydesmida, Nepalella gunsa Shear (Chordeumatida, Trichopolydesmidae = Fuhrmannodesmidae) – 3500 Megalotylidae) – 3800 m (Nepal) m (Meru) Nepalmatoiulus hyalilobus Enghoff (Julida, Hedinomorpha yunnanensis Golovatch (Polydesmida, Julidae) – 3800 m (Nepal) Paradoxosomatidae) – 3480 m (China, Yunnan) Delarthrum (= Armolites) chulingensis (Golovatch) Ceratosphys simoni Ribaut (Craspedosomatida, (Polydesmida, Paradoxosomatidae) – 3700 m (Nepal) Opisthocheiridae) – 3460 m (Sierra Nevada) Sigipinius dentiger Golovatch (Polydesmida, Janetschekella valesiaca (Faës) (=nivalis Paradoxosomatidae) – 3570 m (China, Yunnan) Schubart) (Craspedosomatida, Craspedosomatidae) Anaulaciulus tibetanus Korsós (Julida, Julidae) – 3450 m (Alps) – 3700 m (Tibet) Hingstonia fittkaui Golovatch (Polydesmida, Re-calculated figures for the number of some Trichopolydesmidae = Fuhrmannodesmidae) – 3650 Isopoda, Arachnida and Myriapoda recorded in m (Nepal) the Old World at or higher than 2200 m H. sympatrica Golovatch (Polydesmida, Scorpiones – from 28 to 29 sp. Trichopolydesmidae = Fuhrmannodesmidae) – 3650 Opiliones – from 294 to 305 sp. m (Nepal) Araneae – from 1340 to 1381 sp. High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 153

Acari Trombidiformes – added 12 sp. of and 24 genera. Many corrections are made also in the Trombiculidae lists of high altitude taxa in different areas. Chilopoda – from 159 to 160 sp. Total inhabitants of altitudes over 2200 m in the Diplopoda – from 294 to 365 sp.; added are also Old World: 3927 (formerly 3789) sp. the order Polyzoniida, the families Platydesmidae, The tendency, called by us “taxogradient”, re- Xystodesmidae, Opisotrettidae and Siphonocryptidae mains valid.

References

Arachnida lipede from Taiwan (Diplopoda, Siphonocryptida). – Acta Daniel M. 1973. Chiggers in the high altitude region of the Zoologica Academiae Scientiarum Hungaricae, 54 (2): east Hindu-Kush. – Proceedings of the 3rd International 151-157. Congress of Acarology, Prague, 1971. Academia, Prague, Logunov D.V., Demir H. 2006. Further faunistic notes on Co- 393-396. zyptila and Xysticus from Turkey (Araneae, Thomisidae). Daniel M. 2015. Expansion of small terrestrial mammals and their – Arachnologische Mitteilungen, 31: 38-43. parasites into the Barun Valley (Makalu Mt. Region, Nepal Martens J. 2006. Weberknechte aus den Kaukasus (Arachnida, Himalaya) linked with changes in glaciation and human Opiliones, Nemastomatidae). – Senckenbergiana biologica, activities. – Journal of Mountain Science, 12 (1): 14-29. 86 (2): 145-210. Daniel M., Stekolnikov A.A. 2009. Chigger mites (Acari, Martens J. 2016. Sinostoma yunnanicum, the first nemastomatine Trombiculidae) from Makalu Region in Nepal Himalaya, harvestman in China (Arachnida: Opiliones: Nemastoma- with a description of three new species. – Journal of Medical tidae). – Zootaxa, 4126 (3): 444-450. Entomology, 46: 753-765. Martens J., Chemini C. 1988. Die Gattung Anelasmocephalus Daniel M., Stekolnikov A.A., Hakimitabar M., Saboori A. Simon 1879 – Biogeographie, Artengrenzen und Biospecies- 2010. Chigger mites (Acari, Trombiculidae) parasitizing Konzept (Opiliones: Trogulidae). – Zoologischer Jahrbuch small mammals in the Eastern Hindu Kush and some other für Systematik, 115: 1-48. Afghan areas. – Parasitology Research, 107: 1221-1233. Ochoa J.A., Ojanguren Affilastro A.A., Mattoni C.I., Prendini Dankittipakul P., Jocqué R. 2006. Two new species of Cydrela L. 2011. Systematic revision of the Andean scorpion genus Thorell (Araneae: Zodariidae) from Thailand. – The Raffles Orobothriurus Maury, 1976 (Bothriuridae), with discus- Bulletin of Zoology, 54(1): 93-101. sion of the altitude record for scorpions. – Bulletin of the Das N.P.I., Bastawade D.B. 2006. A new species of genus Eupha- American Museum of Natural History, 359: 90 pp. langium Roewer (Opiliones: Phalangiidae) from Western Ovtchinnikov S.V., Ahmad B., Inayatullah M. 2008. De- Himalayan region of India. – Zoo’s Print Journal, 21 (4): scription of a new species of the genus Gnaphosa 2207-2209. (Araneae, Gnaphosidae) from Pakistan. – Vestnik zoologii, Enghoff H., Golovatch S.I. 1995. A revision of the Sipho- 42 (1): 81-83. nocryptidae (Diplopoda, Polyzoniida). – Zoologica Scripta, Seyyar O., Demir H., Topçu A., Taşdemir A. 2006. Phaeocedus 24 (1): 29-41. is a new genus of ground spider (Araneae, Gnaphosidae) in Ermilov S.G., Martens J. 2014. A new species of Perschelorib- Turkey. – Scientific Research and Essay, 1 (1): 26-27. ates (Acari, Oribatida, Scheloribatidae) from Nepal with Shileyko A.A., Stoev P.E. 1016. Scolopendromorpha of New a key to all species of the genus from the Oriental Region. Guinea and adjacent islands (Myriapoda, Chilopoda). – – Acarina, 22 (1): 14-19. Zootaxa, 4147 (3): 247-280. Fernandes S., Kulkarni S.M. 2003. Studies on the trombiculid Snegovaya N.Yu. 1999. Contribution to the Harvest Spider mite fauna of India. – Records of the Zoological Survey of (Arachnida, Opiliones) Fauna of the Caucasus. – Turkish India, Occasional Paper 212. Journal of Zoology, 23: 453-459. Jastrzębski P. 2010. Salticidae from the Himalayas. The genus Snegovaya N.Yu. 2004. Preliminary notes on the harvestman Hasarius Simon, 1871 (Araneae: Salticidae). – Genus, 21: fauna of Azerbaijan. – Arthropoda Selecta, Special Issue, 319-323. 1: 307-318. Jastrzębski P. 2011. Salticidae from the Himalayas. The genus Snegovaya N.Yu. 2010. Further studies on harvestmen of the Pancorius Simon, 1892 (Arachnida: Araneae). – Genus, genus [Opilio] Herbst, 1798 (Arachnida: Opiliones: Pha- 22: 181-190. langiidae) from the Caucasus. – Caucasian Entomological Jing Shen, Solhoy T., Wang Huifu, Vollan Th.I., Xu Rumei. Bulletin, 6 (1): 3-18. 2005. Differences in Soil Communities along a Snegovaya N.Yu. 2014. Survey of the genus Phalangium Linnaeus, High Altitude Gradient at Shergyla Mountain, Tibet, China. 1758 (Phalangiidae: Opiliones) from the Caucasus with descrip- – Arctic, Antarctic, and Alpine Research, 37 (2): 261-266. tion of two new species. – Fragmanta Faunistica, 57 (1): 1-19. Kok O.B., Lotz L.N., Haddad C.R. 2004. Diversity and Ecology Staręga W. 1986. Eine neue Art der Nemastomatidae (Opiliones) of Spiders (Arachnida: Araneae) of the Deosai Plateau, aus dem Pamir nebst nomenklatorisch-taxonomischen An- Northern Pakistan. – Pakistan Journal of Biological Sci- merkungen. – Bulletin of the Polish Academy of Sciences, ences, 7 (10): 1689-1694. Biological Sciences, 34 (10-12): 301-05. Korsós Z., Enghoff H., Chang H. W. 2008. A most unusual Staręga W., Snegovaya N.Y. 2008a. New species of Opilioninae distribution pattern: a new Siphonocryptid mil- (Opiliones: Phalangiidae) from the mountains of Kyrgyz- 154 Petar Beron

stan, Tadjikistan and Uzbekistan. – Acta Arachnologica, Paradoxosomatidae (Diplopoda Polydesmida), XVII. – Ar- 57 (2): 75-85. thropoda Selecta, 24 (2): 127-168. Staręga W., Snegovaya N.Y. 2008b. A new harvestman from Golovatch S.I. 2016. On several new or poorly-known Oriental Cameroon (Arachnida: Opiliones: Phalangiidae). – Polish Paradoxosomatidae (Diplopoda Polydesmida), XX. – Ar- Journal of Entomology, 77: 321-327. thropoda Selecta, 25 (3): 219-240. Tanasevitch A.V. 2011a. On linyphiid spiders (Araneae) from Golovatch S.I., Akkari N. 2016. Identity of the Millipede, the Eastern and Central Mediterranean kept at the Muséum Pseudoniponiella kometis (Attems, 1938)(Diplopoda: d’histoire naturelle, Geneva. – Revue suisse de Zoologie, Polydesmida: Cryptodesmidae). – Tropical Natural His- 118 (1): 49-91. tory, 16 (1): 1-6. Tanasevitch A.V. 2011b. Linyphiid spiders (Araneae, Linyphii- Golovatch S.I., Evsyukov A., Reip H. 2016. The millipede fami- dae) from Pakistan and India. – Revue suisse de Zoologie, ly Polydesmidae in the Caucasus (Diplopoda: Polydesmida). 118 (3): 561-598. – Zootaxa, 4085 (1): 1-51. Tanasevitch A.V. 2013. The Linyphiid spiders of the Altais, Golovatch S.I., Nzoko Fiemapong A.R., VandenSpiegel D. southern Siberia (Araneae, Linyphiidae). – Arthropoda 2015. Notes on Afrotropical Pyrgodesmidae, 2 (Diplopoda: Selecta, 22 (3): 267-306. Polydesmida). – Arthropoda Selecta, 24 (4): 387-400. Wang Xin-Ping, Ming-Sheng Zhu. 2008. Himalmartensus, a Golovatch S.I., Stoev P. 2009. New or poorly-known millipedes new genus of the spider family Amaurobiidae from Nepal (Diplopoda) from Papua New Guinea, 1. – Arthropoda (Araneae). – The Journal of Arachnology, 36: 241-250. Selecta, 18 (3-4): 125-130. Zambre A., Sanap R.V., Mirza Z.A. 2014. A new high-elevation Golovatch S.I., Stoev P. 2010. New or poorly-known millipedes scorpion species of the genus Scorpiops Peters, 1861 (Scor- (Diplopoda) from Papua New Guinea, 2. – Arthropoda piones: Euscorpiidae: Scorpioninae) from the Himalayas, Selecta, 19 (3): 125-143. India. – Comptes Rendus Biologies, 337: 399-404. Golovatch S.I., Stoev P. 2014a. On five new species of the Zonstein S., Marusik Yu.M. 2012. A review of the genus Raveniola millipede family Paradoxosomatidae from Papua New (Araneae, Nemesiidae) in China, with notes on allied genera Guinea (Diplopoda: Polydesmida). – Arthropoda Selecta, and description of four new species from Yunnan. – ZooK- 23 (1): 21-32. eys, 211: 71-99. Golovatch S.I., Stoev P. 2014b. Review of the Papuan millipede genus Silvattia Jeekel, 2009, with descriptions of three new Myriapoda species (Diplopoda: Polydesmida: Paradoxosomatidae: Eustrongylosomatini). – In: Telnov D. (ed.): Biodiversity Gilgado J.D., Enghoff H., Tinaut A., Mauries J.-P., Ortuño , Biogeography and Nature Conservation in Wallacea and V.M. 201 . Sierra Nevada (Granada, Spain): a high-altitude New Guinea, vol. II: 149-156, 4 Plates. biogeographical crossroads for millipedes (Diplopoda), Golovatch S.I., Stoev P., VanDenSpiegel D. 2010. New or with first data on its MSS fauna and description of a new poorly-known millipedes (Diplopoda) from Papua New species of the genus Ceratosphys Ribaut, 1920 (Chordeuma- Guinea, 3. – Arthropoda Selecta, 19 (3): 145-152. tida: Opisthocheiridae). – Zootaxa, 4044 (3): 391-410. Golovatch S.I., VanDenSpiegel D., Semenyuk I.I. 2016. On Golovatch S.I. 2013a. On several new or poorly-known Oriental several new or poorly-known Oriental Paradoxosomatidae Paradoxosomatidae (Diplopoda Polydesmida), XII. – Ar- (Diplopoda Polydesmida), XXI. – Arthropoda Selecta, 25 thropoda Selecta, 21 (1): 1-12. (4): 335-354. Golovatch S.I. 2013b. On several new or poorly-known Oriental Nefediev P.S., Nefedieva J.S., Jankowski K. 2015. The first Paradoxosomatidae (Diplopoda Polydesmida), XIII. – Ar- record of a julid millipede (Diplopoda: Julida: Julidae) from thropoda Selecta, 22 (1): 1-31. the Altai Mountains of Mongolia. – Invertebrate Zoology, Golovatch S.I. 2013c. On several new or poorly-known Oriental 12 (2): 213-214. Paradoxosomatidae (Diplopoda Polydesmida), XIV. – Ar- thropoda Selecta, 22 (4): 307-332. General papers Golovatch S.I. 2014. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda Polydesmida), XVI. – Ar- Beron P. 2008. High-mountain Isopoda Oniscidea, Arachnida thropoda Selecta, 23 (3): 227-251. and Myriapoda in the Old World. – Bureschiana, Sofia, 1: Golovatch S.I. 2015a. Two new species of the millipede order Pensoft & Nat. Mus. Natur. Hist. Sofia, 556 pp. Polydesmida from Southern China (Diplopoda). – Zoolog- Franz H. 1975. Die Bodenfauna der Erde in biozönotischer icheskii zhurnal, 94 (9): 1023-1028. Betrachtung. Teil I. – Erdwissenschaftliche Forschung, X: Golovatch S.I. 2015b. On several new or poorly-known Oriental 549-554.

Author’s Address:

Petar Beron, National Museum of Natural History, 1Tsar Osvoboditel, 1000 Sofia, Bulgaria. E-mail: [email protected] High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 155 Високопланински Isopoda‚ Arachnida и Myriapoda в Стария свят (допълнения и поправки 2008-2016)

Петър Берон

(резюме)

След излизането на моята монография (Beron, 2008) бяха публикувани много нови данни и из- вършени някои корекции на по-старата информация върху високопланинските Isopoda‚ Arachnida и Myriapoda в Стария свят. Това наложи настоящото първо допълнение. С така ревизираните данни броят на известните видове от някои групи във високите планини се изменя както следва: Scorpiones – от 28 на 29 в. Opiliones – от 293 на 306 в. Araneae – от 1340 на 1381 в. Acari Trombidiformes – добавят се 12 в. oт сем. Trombiculidae Diplopoda – от 294 на 365 в. Общ брой на изоподите, паякоподобните и многоножките в стария свят, познати над 2200 m: 3927 вида (според Beron, 2008-3789)