Supplementa and Corrigenda 2008-2016)
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Historia naturalis bulgarica, 23: 141-155, 2016 High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) Petar Beron Abstract: After the publication of my monograph (Beron, 2008) appeared many new papers containing data on Arachnida and Myriapoda found in the Old World at or above 2200 m. In this supplement are added 12 sp. of Opilions, 37 sp. of spiders, and 74 sp. of Diplopoda. Several changes occurred in the taxonomy of some groups, following taxonomic revi- sions. Several genera of Opiliones are no more in Phalangiidae, but in Sclerosomatidae, Fuhrmanodesmidae became synonyme of Trichopolydesmidae, etc. “Palpatores” are now distributed between the suborders Eupnoi and Dyspnoi. Several specialists, and especially Golovatch, published many new taxa of high mountain Diplopoda. Again Golovatch synonymized several genera of Diplopoda (Martensosoma, Nepalomorpha, Parorthomorpha, Armolites, Orophosoma), known from the high mountains, under Delartrum Attems. For many taxa has been “raised” the upper limit of distri- bution and they have been published from other mountains. Key Words: Arachnida, Myriapoda, high altitude, Old World, supplementa, corrigenda Underlined are taxa not included in the monograph of Beron (2008) The mountain systems of the Old World and the Opilio lederi Roewer – up to 3000 m Isopoda, Arachnida and Myriapoda recorded therein O. parietinus (De Geer) – up to 3000 m Pyrenees and the mountains of Iberian (Azerbaijan) Peninsula O. redikorzevi Roewer – 2400 m Diplopoda Phalangium opilio L.– up to 3000 m New ref.: Gilgado et al. (2015) Ph. punctipes L. Koch – up to 3000 m Chordeumatida Araneae Fam. Opisthocheiridae New ref: Logunov & Demir (2006) Ceratosphys cryodeserti Gilgado, Mauriès et Fam. Thomisidae Enghoff – Sierra Nevada Xysticus bacurianensis Mcheidze – 3000 m Alps (Azerbaijan), 2900 m (Georgia) Arachnida Myriapoda Araneae Diplopoda New ref.: Tanasevitch (2011a, 2011b, 2013) New ref.: Golovatch, Evsyukov & Reip (2016) Fam. Linyphiidae Fam. Polydesmidae Agyneta nigripes (Simon) – 1900 – 2580 m Brachydesmus assimilis Lohmander – 2800 m Erigone remota L. Koch – 2650 m B. kalischewskyi Lignau – 2400 m E. tenuimana Simon – up to 2580 m Polydesmus lignaui Lohmander – 2800 m Caucasus Mountains of Turkey, Armenia, Lebanon, Iran, Arachnida Kopet Dagh and Arabian Peninsula Opiliones Arachnida New ref.: Snegovaya (1999, 2004, 2010, 2014) Araneae Fam. Phalangiidae New ref.: Logunov & Demir (2006), Seyyar, Odiellus bieniaszi Kulczynski – up to 3000 m Demir, Topçu & Taşdemir (2006) 142 Petar Beron Fam. Gnaphosidae Tiso incisus Tanasevitch – 1300-2500 m Phaeocedus braccatus (L. Koch) – 2300 m (Karakorum) (Erciyes) Acari Fam. Thomisidae New ref.: Daniel (1973), Daniel, Stekolnikov, Xysticus bacurianensis Mcheidze – 2200 m Hakimitabar & Saboori (2010), Jing, Solhoy, X. cristatus (Clerck) – 2200 m Wang, Vollan & Xu (2005) Acari, known from the described area at or Mountains of Afghanistan, Pakistan, Karakorum, above 2200 m (suppl.): Tibet, Pamir, Kunlun and Tien Shan (from the border between Iran and Afghanistan to 120o E) Acariformes Prostigmata Arachnida Fam. Trombiculidae Opiliones Microtrombicula gratiosa Schluger et New or ommited ref.: Staręga (1986), Kudryashova – Hindu Kush, 2950 m Staręga & Snegovaya (2008a) M. similata Schluger et Amanguliev – Hindu Eupnoi Kush, 3080 m Fam. Phalangiidae M. vacillata Fernandes et Kulkarni – Hindu Homolophus gricenkoi Staręga et Snegovaya – Kush, 3080 m 2000 -3000 m (Tadjikistan) Neotrombicula lubrica Kudryashova – Hindu H. andreevae Staręga et Snegovaya – up to Kush, 2950-3990 m 4670 m (Pamir) Leptotrombidium silvaticum Hushcha et Pamiropilio naukat Staręga et Snegovaya – 3000- Schluger – Hindu Kush, 2782 m 3300 m (Kyrgyzstan, Alayskiy Hrebet) Montivagum dihumerale (Traub et Nadchatram) Dyspnoi – Hindu Kush, 3990 m Fam. Nemastomatidae Mediostoma ( ?) pamiricum Staręga, 1986- M. latum (Schluger et Kudryashova) – Hindu 3100 m (Pamir) Kush, 2950-3990 m Araneae M. raropinne (Schluger) – Hindu Kush, 3100 m New ref.: Kok, Lotz & Haddad (2004), Euschoengstia meshhedensis Kudryashova, Tanasevitch (2011b) Neronov et Farhang-Azad – Hindu Kush, 2950 – Fam. Linyphiidae 3990 m Acartauchenius himalayensis Tanasevitch – Brunehaldia lucida Schluger – Hindu Kush, 2500 m (Karakorum) 2950 m Agyneta nigripes (Simon) – 2635 m Shunsennia oudemansi (Schluger) – 2950 – (Karakorum) 4550 m Archaraeoncus prospiciens (Thorell) – 2700- Himalaya 3000 m (Karakorum) (East of Indus and South and West of Tzangpo, Caviphantes pseudosaxetorum Wunderlich – or Brahmaputra) 550-2500 m (Karakorum) New research: Ceratinella wideri (Thorell) – 2500-3100 m B. Petrov collected many different animals (Karakorum) during his several climbing expeditions in the Gongylidiellum confusum Thaler – 1600-2700 m Himalaya. (Karakorum) Notes on Isopoda, Arachnida and Myriapoda Maso sundevalli (Westring) – 2200 – 2300 m Arachnida (Karakorum) Scorpiones Pelecopsis indus Tanasevitch – 2300 m New ref.: Zambre, Sanap & Mirza (2014) (Karakorum) Fam. Euscorpiidae Piniphantes himalayensis (Tanasevitch) – Scorpiops spitiensis Zambre, Sanap et Mirza – 2800 m (Karakorum) Himalaya, 4200 m Porrhomma pygmaeum (Blackwall) – 2480 m Araneae (Karakorum) New ref.: Jastrzębski, 2010, 2011, Prinerigone vagans (Audouin) – 1700-2700 m Ovtchinnikov, Ahmad & Inayatullah (2008), (Karakorum) Tanasevitch (2011b), Wang & Zhu (2008) cotargus pilosus Simon – 1400-2450 m Some spiders in the Himalaya found at or above (Karakorum) 2200 m after 2008 are: High Altitude Isopoda‚ Arachnida and Myriapoda in the Old World (supplementa and corrigenda 2008-2016) 143 Fam. Amaurobiidae Myriapoda Himalmartensus ausobskyi Wang et Zhu – 1770- Chilopoda 3000 m Scolopendromorpha H. martensi Wang et Zhu – 1400-2900 m Fam. Scolopendridae H. nepalensis Wang et Zhu – 1000-2600 m Scolopendra canidens Newport – 2400 m Fam. Gnaphosidae Diplopoda Gnaphosa pakistanica Ovtchinnikov, Ahmad et New ref.: Golovatch (2014, 2015b, 2016)‚ Inayatullah – 4100 m Golovatch, Nzoko Fiemapong & VandenSpiegel Fam. Linyphiidae (2015)‚ Golovatch, VandenSpiegel & Semenyuk Anguliphantes nepalsioides Tanasevitch – 2500- (2016) 2600 m Polydesmida (restructuring) Gongylidiellum confusum Thaler – up to 3200 m Fam. Trichopolydesmidae (syn. Fuhrmanno- (Kashmir) desmidae) Indophantes tonglu Tanasevitch – 2500-2700 m Fam. Paradoxosomatidae Pelecopsis indus Tanasevitch – 2250 m Anoplocephalus martensi Golovatch – up to Tapinocyboides bengalensis Tanasevitch – 3100 m 3600 m (Nepal) Tiso incisus Tanasevitch – 1900 – 2450 m Beronodesmoides lobatus Golovatch – 4600 m T.(?) indianus Tanasevitch – 2500 – 2600 m (Nepal) Walckenaeria martensi Wunderlich = W. nepa- B. montigena Golovatch 3550 m (Nepal) lensis Wund. (syn. by Tanasevitch, 2011b) B. typicus Golovatch – 3400 m (Nepal) Acari B. anteriporus Golovatch – 3350 m (Nepal) New and ommited ref.: Daniel (1973), Daniel B. martensi Golovatch – 2700 m (Nepal) & Stekolnikov (2009), Daniel, Stekolnikov, Beronodesmus latispinosus Golovatch – up to Hakimitabar & Saboori (2010), Ermilov & 3500 m (Nepal) Martens (2014), Fernandes & Kulkarni (2003) B. longispinosus Golovatch – up to 4270 m Some of the Acari, found in Himalaya at or (Nepal) above 2200 m after 2008, are: B. pallidus Golovatch – 4100 m (Nepal) Acariformes B. gorkhalis Golovatch – up to 3600 m (Nepal) Prostigmata B. latispinosus Golovatch – 3150 m (Nepal) Fam. Trombiculidae B. serratus Golovatch et al. – up to 3500 m Brunehaldia lucida Schluger – 2950 m (Nepal) Cheladonta ikaoensis Sasa et al. – 3450-4000 m B. sinuatospinus Golovatch – 2250 m (Nepal) Doloisia vlastae Daniel et Stekolnikov – 3450 m Delarthrum (= Armolites) chulingensis Euschoengastia meshhedensis Kudryashova et (Golovatch) – up to 3700 m al. – 2950-3990 m D. similis (Golovatch) – up to 2700 m Leptotrombidium angkamii Daniel et D. communicans (Golovatch) – up to 2650 m Stekolnikov – 3450-3900 m D. spiniger (Attems) – 2200 m (India) L. silvaticum Hushcha et Schluger – 2782 m Hirtodrepanum latigonopum Golovach – up to Microtrombicula gratiosa Schluger et 2700 m Kudryashova – 2950 m Kaschmiriosoma pleuroptera Attems – up to M. similata Schluger et Amanguliev – 3080 m 2800 m (Pakistan) M. vacillata Fernandes et Kulkarni – 3080 m Martensosoma silvestre Golovatch – up to 2600 m Montivagum dihumerale (Traub et Nadchatram) M. schawalleri Golovatch – up to 2150. m – 3990 m M. splendens Golovatch – up to 2150 m M. latum (Schluger et Kudryashova) – 2950-3990 m Delarthrum (= Nepalomorpha) hirsutum M. raropinne (Schluger) – up to 3100 m (Golovatch) – up to 4100 m Neotrombicula kounickyi Daniel et Stekolnikov D. kuznetzovi (Golovatch) – up to 3000 m – 3450-4000 m D. arunensis (Golovatch) – up to 2150 m N. lubrica Kudryashova – 1090-3990 m Delarthrum (= Orophosoma) simulans (Carl) – N. turkestanica Kudryashova – 3990 m up to 3700 m Shunsennia oudemansi (Schluger) – 1800-4100 m D. hingstoni (Carl) – up to 3400 m Trombiculindus mehtai Fernandes et Kulkarni D. fechteri (Golovatch) – up to 3150 m – 3450-4600 m Paranedyopus martensi Golovatch – up to 3600 m 144 Petar Beron P. similis Golovatch – up to 3000 m Spirobolida P. cylindricus Carl – up to 2850 m Fam. Pachybolidae P. affinis Golovatch – up to 2700 m Trigoniulus brachysternus Attems – 1400 – 2400 P. schawalleri Golovatch – up to 2150 m m (Vietnam) Delarthrum (= Parorthomorpha) tuberculatum Siphonophorida (Golovatch) – up to 3300 m Fam. Siphonophoridae D. spectabilis (Golovatch) – up to 2650 m Teratognathus robusta Attems – 1000 – 2400 m D. tergalis Golovatch – up to 2650