Contr. Tert. Quatern. Geol. 27(4) 117-123 1 pi. Leiden, December 1990

Lyria (Lyria) picturata (de Grateloup, 1834) from the

Miocene of Winterswijk-Miste, The Netherlands, with

notes on related taxa (, )

A.W. Janssen

NATIONAAL NATUURHISTORISCH MUSEUM

LEIDEN, THE NETHERLANDS

Janssen, A.W. Lyria (Lyria) picturata (de Grateloup, 1834) from the Miocene of Winterswijk-Miste, The Netherlands, with notes

on related taxa (Mollusca, Gastropoda). — Contr. Tert. Quatern. Geol., 27(4): 117-123, 1 pi. Leiden, December 1990.

A single specimen of the volutoid gastropod Lyria (Lyria) picturata (de Grateloup, 1834) is recorded from the Miocene (Hem-

moorian) of Winterswijk-Miste, The Netherlands. A revision of related European taxa leads to the conclusion that Voluta taurinia

1847 is of V. de Voluta oliva de 1847 be retained Michelotti, a junior subjective synonym picturata Grateloup. Grateloup, may as

of which taurinia Sacco, 1904 is a The distribution of in- a forma, Lyria var. magnoventrosa junior synonym. pattern Lyria picturata

cludes the North Sea Basin (forma oliva only), SW France, northern Italy, Turkey and the Paratethys.

words — Key Mollusca, Gastropoda, Volutoidea, Lyria, Miocene, .

A.W. Janssen, Nationaal Natuurhistorisch Museum (Palaeontology Department, Cainozoic Mollusca), P.O. Box 9517, 2300 RA

Leiden, The Netherlands

Contents 132). Mr van der Hoek assented to a study of this

and made specimen to having a plaster cast for the Introduction 117 p. RGM collections. 117 Systematic part p. The identificationof this specimen necessitated Lyria outside the North Sea Basin p. 119 with related forms from other Euro- a comparison Conclusions 120 p. which resulted in taxonomic pean localities, a References 120 p. revision.

Introduction

SYSTEMATIC PART Subsequent to the publication of my paper on the

Winterswijk-Miste molluscan fauna (Janssen, Phylum Mollusca several have been 1984) new species found, as Classis Gastropoda from time time made and to new exposures were Ordo large quantities of sediment processed. Since these SuperfamiliaV olutoidea new collections were not made by the Palaeon- Familia tology Department of the Nationaal Natuurhisto- Lyria Gray, 1847 s.str. risch Museum at Leiden (formerly 'Rijksmuseum

of the van Geologie en Mineralogie, RGM), most Type species: Lyria (Lyria) nucleus (Lamarck). housed in new taxa are private collections.

Several years ago, Mr B. van der Hoek (Twello, Lyria (Lyria) picturata (de Grateloup, 1834) The Netherlands) demonstrateda specimen of the PI. 1, Fig. 9 hitherto known from the North Sea genus Lyria,

Basin Miocene in one from Hem- only specimen Voluta — 8 1820 papillaris. Borson, p. 205, pi. 1, fig. (non

moor, F.R.G., published by Kautsky (1925, p. Gmelin). 118

Voluta taurinia (1825) Bonelli, Catalogus no. 2532 (un- 1982 Lyria taurinia (Michelotti, 1847, Bonelli m.s.). — Fer-

Mortara published). rero et al p. 173.

1827 Voluta de 16 1982 mitraeformis, Grateloup, p. (non Lyria taurinia (Michelotti, 1847, Bonelli m.s.) var.

Lamarck). perventrosa Sacco, 1890. — Ferrero Mortara et al. , p.

•1834 Voluta de 293. picturata [...] Nob., Grateloup, p. 173.

1837 Voluta Broc. var. — 11, 1982 magorum Pusch, p. 117, pi. Lyria taurinia (Michelotti, 1847, Bonelli m.s.) var.

fig. 2a-b (non Brocchi). magnoventrosa Sacco, 1890. — Ferrero Mortara et

1840 Voluta Grat. — de 39 Voluta 173. picturata. Grateloup, pi. ( al., p.

II), figs 5-11.

1844 Voluta papillaris. Borson. — Deshayes & Milne Ed- Material studied — A single adult specimen with 431. wards, p. slightly corroded apical whorls in the collection of 1847 Voluta oliva. Grat., de Grateloup, pi. 46 (Supplement

Mr B. van der Hoek, Twello I), fig. 8. (The Netherlands),

1847 Voluta Taurinia. 319. M Bon., Michelotti, p. collected in 1983 (registration no. 02-339). A

1852 Voluta picturata, 1845. — Gratteloup (sic !), d'Or- plaster replica of this specimen is housed in the 9 bigny, (26e no. 142). p. etage RGM 799. collections, registration no. RGM 229

1852 Voluta Bonelli. — 53 Taurina, d'Orbigny, p. (26e Description — Shell solid, medium-sized, fusiform etage no. 885).

to 1 Voluta oliva. — mitraeform, about 2 4 times as as wide. [1852 d'Orbigny (26e etage no. 898) (men- high

tioned in the index on 188; the texts of nos The is only p. apex worn, somewhat more than eight post- 891 the to 899, obviously belonging to Mitra, The genus embryonal whorls are preserved. relatively are missing on page 53)]. whorls without high are slightly convex, any 1852 Voluta taurinia Bon. — Hoernes, p. 95, pi. 9, figs below the sutures. The whorl 14-16. thickening body

about three of the entire shell 1853 Voluta Bors. — Eichwald, 197. comprises quarters papillaris von p.

?1866 Voluta taurina Bon. var. — Below the the Speyer, p. 13, pi. 1, fig. height. periphery shell gradually 6a-d. tapers towards the relatively short and wide canal.

1880 Voluta taurinia Bon. — Hoernes 72. & Auinger, p. A There slightly thickened siphonal belt is present. 1890 Lyria taurinia (Bon.) var. A et B. — Bellardi & Sacco, is no 8, 5a-b. pseudumbilicus. p. pi. 1, fig. The is and widest in its 1890 taurinia C. — Bellardi Lyria (Bon.) var. & Sacco, p. 8, aperture long narrow,

6. third. There is distinct the pi. 1, fig. upper no boundary at

1896 Voluta taurinia Bon. — 58. Boettger, p. transition into the canal. The apertural margin is

1899 Voluta — 114. picturata Grat. Cossmann, p. slightly thickened externally. No internal thicken-

1899 Voluta taurinia Bon. — 114. Cossmann, p. columella is ing or denticles are present. The 1904 taurina Sacc. — Lyria (Bon.) var. subplanulata Sacco, p. with but 89. massive, a distinctly separated, un-

1904 taurina var. Sacc. — thickened callus. Three columellar folds Lyria (Bon.) perventrosa Sacco, p. are pres-

89. middle of which The ent, the one is the strongest.

1904 Lyria taurina (Bon.) var. magnoventrosa Sacc. — Sacco, lower one coincides with the left margin of the 89. p. canal. Two weak very columellar folds are vaguely 1906 Voluta taurinia Bon. — 5. Boettger, p. discernible above the and further 1912 taurinia Bon. — stronger ones, a Lyria Friedberg, p. 121, pi. 6, fig.

8a-b. the of the one near upper corner aperture.

1925 taurinia Bon. — 12. Lyria Kautsky, p. 132, pi. 9, fig. The collabral ornament of the shell is restricted

1928 Lyria picturata Grateloup. — Peyrot, 145, pi. 10, p. the ofthe to upper two preserved whorls, on which figs 38-44. some 10 to 12 ribs are rather indistinct 1928 present, Lyria picturata (Grateloup) mut. primitiva nov. mut., due to the the shell. This 9-10. poor condition of Peyrot, p. 147, pi. 11, figs apical

1928 oliva — the Lyria (Grateloup). Peyrot, p. 147, pi. 11, figs sculpture rapidly disappears on younger volu-

4-6. which tions, are entirely smooth. On the body

1938 taurinia Bon. — 135. Lyria Friedberg, p. near the whorl, especially canal, a very vague 1955 Lyria (Lyria) taurinia Bon. — Korobkov, pi. 77, fig. striation is visible in favourable la-b. spiral light.

and 1956 taurinia — narrow Lyria (Bonelli). Csepreghy-Meznerics, p. Orange-coloured, widely spaced spiral

10. lines 5 415, pi. 9, figs 9, are present on the two youngest whorls: on

1958 taurinia — Lyria (Bonelli). Eriinal-Erentoz, 86, pi. and 14 the p. the penultimate one on the body whorl, 15, figs 5-8. lower one being just above the siphonal belt.

1960 1833. — 60. Lyria picturata Grateloup, sp. Glibert, p. The distinct growth lines are but not accen- 1960 taurinia Bonelli, 1825. — 60. Lyria sp. Glibert, p. tuated. almost 1966 taurinia — run or are Lyria Bonelli, 1826. Strausz, p. 374, pi. 41, They perpendicular only

figs 11-14. slightly curved backwards between the sutures. 119

NORTH SEA BASIN On the body whorl they are flexuous, curving LYRIA OUTSIDE THE

backwards on the canal, with a distinct separate Forms related to the shells from Miste and Hem- the belt. curve on siphonal been mentioned moor have frequently from — here is the Remarks The shell described only one

1 localities outside the North Sea Basin from the Miste A further (Aquitaine known locality . speci- Basin, northern Italy, Turkey, and various men from the Miocene of the North Sea Basin was

localities: from the famous Paratethys Romania, Hungary, mentioned, as Lyria taurinia, Austria, Poland). Hemmoor locality by Kautsky (1925, pi. 9, fig. From Saubrigues (Aquitaine Basin), Lyria oliva 12). This specimen (judging from the illustration)

was described based with the Miste shell in size and (de Grateloup) (PL 1, Fig. 7), agrees perfectly This in on a single shell. specimen (see photograph overall form, but apparently the colour pattern is Peyrot, 1928, pi. 11, figs 4-6) well with the not preserved. It is undoubtedly conspecific with agrees North Sea Basin in well in the Miste shell. shells, sculpture as as

overall form. The French specimen might be Additional records of Miocene Lyria from the but is has North slightly broader, a similarly elongate Sea Basin were given by Gripp (1915, p. this L. oliva is taurinia several apical part. According to Peyrot 26, as Voluta Bon.) from Early

related to another from the Miocene localities. These records could not be closely Lyria Aquitaine viz. L. of the material Basin, picturata (de Grateloup) (PI. 1, Figs checked, as the whereabouts are which in several also known from of these localities 1, 2a-c), is, varieties, unknown. Specimens one from Saubrigues and a number of other localities. (Segeberg) studied by Kautsky (1925, p. 132), The differences between L. oliva and L. another of picturata however, proved to belong to group be found in the shell's are to mainly proportions, volutoid species, related to V. suturalis Nyst. shell L. picturata having a relatively shorter apical The Late Oligocene Lyria (Lyria) tumida (von in thick-set The part, resulting a more shell form. Koenen, 1909) (PI. 1, Fig. 12) is known from shell wall in L. picturata is also stated to be slightly several localities (Freden, Volpriehausen and below the thus the This well il- produced sutures, making spira Ahnetal, all F.R.G.). species was somewhat scalariform. Judging from Peyrot's il- lustrated by R. Janssen (1979, pi. 16, fig. 33). It lustrations, however, this latter feature is hardly differs from the Miocene form by its slender shell, recognisable. its somewhat shorter apical shell part and, From Italy a related Lyria species has been re- especially, by the development of the collabral

corded under the name of L. taurinia from several sculpture, which frequently persists until the body Miocene localities in the Torino hills. Bonelli whorl. Also, the columellar folds seem to be some- 1826 is cited (1825, or even 1827) usually as being what better developed in L. tumida.

the author of this taxon. Bonelli's paper, however, A further Late Oligocene Lyria record concerns of the Torino described being a catalogue museum, was a juvenile specimen (PI. 1, Fig. lla-c) The first valid introductionof the Voluta never published. by Speyer (1866, p. 13, pi. 1, fig. 6a-d) as ‘taurinia’ Voluta is Michelotti who name (as taurinia) by taurina Bon. var. R. Janssen (1979), com- (1847), who recorded the species from 'La colline pared this shell with Miocene specimens of Lyria de Turin, Dax et la Pologne'. The first illustration taurinia, found several differences and regarded of L. taurinia specimens from the type area is given Speyer's specimen to be not conspecific with that in Bellardi Sacco This referred it aff. & (1890, pi. 1, fig. 5a-b). species, and to as Lyria (Lyria) sp. ex form (compare PI. 1, Fig. 3a-b) corresponds close- taurinia (Bonelli 1825). ly with L. picturata. The relationship was recognis-

ed also other by authors, e.g. Peyrot (1928, p.

147). The differences between the two taxa, as

that incor- given by author, are partly based on an

rect translation of Bellardi & Sacco's Latin

diagnosis:

1 before this I that Just submitting paper experienced a second Lyria specimen was found at the same locality. It is in '[...] Superficies [...], in anfractubus mediis et ultimo the private collection ofMr W. Peletier, at Winterswijk, The levis, in vel Netherlands (Dr P.A.M. Gaemers, personal communi- primis quatuor quinque longitudinaliter cation). costata\ [...]'. 120

conclusion in that of variation. The same was an- Strausz (1966, p. 375) was correct stating range

that there four five radial Strausz in this does not mean are or ticipated by (1966, p. 374) and, fact,

first four & Milne Edwards ribs on the first whorls, but that the or already by Deshayes (1844, p. five whorls have radial sculpture. This could be 431), who stated sub Voluta papillaris: 'Fossile de

several L. from de la Turin'. verified on specimens of taurinia Dax, et Superga, près the in the RGM collections. In The laws in this case that the cor- type area present priority enjoin these collabral of 10-13 ribs rect of this is L. shells a sculpture some specific name complex picturata (de

the volutions. This means that A was by is present on upper Grateloup, 1834). lectotype designated

should be Type of L. taurinia (Michelotti, 1847) synonym- Peyrot (1928, pi. 11, figs 38-40). locality ised with L. picturata (de Grateloup, 1834). this taxon is Saubrigues in SW France (dep.

Sacco Stratum the 'Marnes de When describing Lyria taurinia, Bellardi & Fandes). typicum are bleus de Mar- distinguished three main varieties, A, B and C. Of Saubrigues' ('Faluns de Saint-Jean

C of Miocene these, only var. was illustrated (pi. 1, fig. 6). sac') (Burdigalian/Fanghian) age,

form PI. named which constitute of fill. This (compare 1, Fig. 8) was var. part a palaeocanyon

It resembles The forms of L. should magnoventrosa by Sacco (1904). closely more elongate picturata

The forma oliva L. oliva from Saubrigues, SW France. be referred to as (de Grateloup, 1847),

features of which L. 1904 is distinguishing as given by Peyrot (1928, magnoventrosa Sacco, a junior

in do the mainte- The fact that the forma oliva p. 148), my opinion, not justify synonym. apparently

co-occurred with the form nance of two separate taxa. typical indicates that

various known these when the Among the Lyria forms from are not subspecies, even only

the European Oligocene and Early Miocene deposits elongate form has so far become known from

described from the North Sea Basin. especially the one North Sea

Basin, L. (L.) tumida (von Koenen, 1909) (PI. 1,

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4761-736, pis 46-52 (1856). cepted 12 November 1990 122

PLATE 1

Outlines (colour patterns omitted !) of Lyria species from various European Late Oligocene and Miocene localities.

Lyria (Lyr6ia) picturata (de Grateloup, 1834)

Fig. 1. Saubrigues (France, dep. Landes); Miocene (Burdigalian/Langhian). After de Grateloup (1834, pl. 39 = Voluta II,

fig. 5; as Voluta picturata. Grat.).

Fig. 2. Lectotype. Saubrigues (France, dép. Landes); Miocene (Burdigalian/Langhian). After Peyrot [1928, pl. 11, figs

38-40; as Lyria picturata (Grat.)].

Fig. 3. Colli Torinesi (Italy); Miocene (‘Helvetiano’). After Bellardi & Sacco [1890, pl. 1, fig. 5a-b; as Lyria taurinia

(Bon.)].

After Strausz Bonelli, Fig. 4. Szob (Hungary), Miocene (‘Tortonian’). (1966, pl. 41, figs 11, 12; as Lyria taurinia 1826)

(the same specimen was also illustrated in Csepreghy-Meznerics, 1956, pl. 9, figs 9, 10).

Fig. 5. Lapugy (Rumania), Miocene (‘Badenian’). After Strausz (1966, pl. 41, figs 11, 12; as Lyria taurinia Bonelli, 1826); specimen showing internal columellar folds.

Steinabrunn Miocene After Hoernes 14a; Voluta taurinia Fig. 6. (Austria); (‘Badenian’). (1852, pl. 9, fig. as Bon.).

Lyria (Lyria) picturata (de Grateloup, 1834) forma oliva (de Grateloup, 1847)

Fig. 7. Holotype. Saubrigues (France, dep. Landes); Miocene (Burdigalian/Langhian). After de Grateloup (1847, pl. 46 =

Voluta oliva. suppl. I, fig. 8; as Grat.).

Fig. 8. Rio della Batteria (Italy); Miocene (‘Helvetiano’). After Bellardi & Sacco [1890, pl. 1, fig. 6; as Lyria taurinia (Bon.)

var. C].

Fig. 9. Winterswijk-Miste (The Netherlands). Miocene (Hemmoorian). After plaster replica in the Nationaal

Natuurhistorisch Museum, dept. Cainozoic Mollusca no. RGM 229 799 (original in private collection of B. van der

Hoek).

10. Hemmoor Miocene After taurinia Fig. (F.R.G.), (Hemmoorian). Kautsky (1925, pl. 9, fig. 12; as Lyria Bon.).

Lyria (Lyria) sp.

Fig. 11. Göttentrup (F.R.G.), Oligocene (Chattian). After Speyer (1866, pl. 1, fig. 6a-c; as Voluta taurina Bon.).

Lyria (Lyria) tumida (von Koenen, 1909)

12. Freden Alfeld/Leine After Fig. near (F.R.G.), Oligocene (Chattian). R. Janssen (1979, pl. 16, fig. 33).

Fig. 11c x 4, all other figures natural size. 123\1

PLATE 1