Transactions and Proceedings

of the Palaeontological Society of Japan

New Series No. 77

Palaeontological Society of Japan April lOth, 1970 Editor: Tokio SHIKAMA Associate Editor: Kiyotaka CHINZEI

Officers for 1969-1970

President: Fuyuji T AKAI Councillors (*Executives): Kiyoshi ASANO*, Tetsuro HANAI* (Secretary), Kotora HATAI, Itaru HAYAMI, Koichiro ICHIKAWA, Taro KANAYA, Kametoshi KAN­ MERA, Teiichi KOBAYASHI, Kenji KO NISHI, Tamio KOTAKA, Tatsuro MATSU­ MOTO*, Masao MrNATO, Hiroshi OZAKI* (Treasurer), Tokio SHIKAMA*, Fuyuji TAKA!* Executive Committee (Chairman: Fuyuji T AKAI) General Affairs: Tetsuro HANAI, Takashi HAMADA, Yasuhide IwASAKI Membership: Takashi HAMADA Finance: Hiroshi OzAKI, Saburo KANNO Planning: Hiroshi OZAKI, Hiroshi UJUE Publications Transactions: Tokio SHIKAMA, Kiyotaka CHINZEI Special Papers: Tatsuro MATSUMOTO, Tomowo OZAWA "Fossils": Kiyoshi ASANO, Yokichi TAKA YANAGI

Fossils on the cover is Globorotalia truncatulinoides (D '0RBIGNY, 1839). The photograph was taken on a scanning electron microscope, JEOL-JSM-2, x 100.

All communications relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN Geological Institute, Faculty of Science, University of Tokyo, Tokyo 113, Japan Sole agent: University of Tokyo Press Trans. Proc. Palaeont. Soc. Japan, N. S., No. 77, pp. 205-228, pl. 23, April 10, 1970

561. A MIOCENE MOLLUSCAN FAUNA IN THE PHILIPPINES

Y ASUHIDE IWASAKI

Department of Geology, University Museum, University of Tokyo

71 !I -:;l:::':,-~f.Jlb~O)I=j:l~Ji'i!t~1t.fi"Wf: 1966~fnllfi!i::.Jv'/ :,-lfb Tayabas !tk!KO)!l[\'.i(f\'4 1i!iiDij:J!l:~fT-:>f.:=*:ftlilt!$ • fRiii.IJPJrjiljj!!;f;J:, f*:ffv;J:.Li!. < t.;:v'i.l;ml: r.d~ili-t Mllifb;~ :Q c ,'(l:l,:b;h, :Q~1t.15r~*~· 2 -7FJftP;J:.U~Lt1f'011!l--:>f.:. ~/±II~Mr!lll Gumaca ~O)J.:!l!IL'$~Jf*lf0) m-e~"? t.::. ~ "Ctl\'l*~~;J:=t!(~ 20 w.. ~~ 16lmi.l;~t&)}1J~n. =1'117Jl~O) u W.H~v''"CT""­ "CP,!/;*rJ0)1t.Elll! • :ffl.itf.l!IC.fiij~~tl.'"C\t':Q. ~~J]':Jt.;:fjfic l....'"C, Vicarya callosa, Anadara multijormis, ]oannisiella cumingi, Paphia exarata t.;: c·~*J;., ..§.-:> '"( MARTIN, SMITH ~ICJ: -:>'"CfiUmi~tJ..t-: 71 !I -:;l:::':/,&~1 :.-' f::f.~7/JlliiiCJA< :$J':fni"":QI=j:l~Jj'Mi:r$ O)~J;R~vr.f*- :.J!~#u:ltfl1t.:fillf.lr.r~-t :Q c J!f4l( J....{~:Q. -/JO)ll[I:BJ:tk Pitogof;Ji[i:O) 2 ~iJ,; ~;tt.t-:: t~*~lHr. -::>v''"C f;J:, =1'3't::$J't.:MH-cv;J:~ :QiJ~ l!~fiiU;IJ-:Q 11[!±\;jj\@il'; §$_0)fillllf fi'ltliH~fjiJ[;""t:Q t. ci.I~L' ~ :Q. t.O)~!f;, :ffl.J:lli$.1Clli:v' Dosinia-Anacf.ara llf.~ci!I!Mi:~tt.t.: c,i(l:l,:btJ..:Q Batillaria llf.~O)jitij~*O)/~il-L.t.:JbO)cfi~~;h,:Q. ]!!lr.. t.O)l/if/Ji "Pitogo fauna" f;J: E*O)m.i~~~t.;: cvr.J;.;tl,.:Q Vicarya ~*tr~1t.15Uc i!lit@J'I':Jir. Jbj;,jlll)"C J: < ~w, 1.... '"Cv' :Q. Jitii~~l.mllfti'ltnX:O)J:il' ;.!U.!(-t :Q c, Jitii!lliO) f.it~O) itv' ~.&:~ J.... "C{!Mk 0) llf.~O)f./ltnX:lli!O))..;h,t< ~ l;l:, r, I iR~IC~It:Q~OCllbcf.l!-=f-7-llbJitii!l[\~O)~J: ~ t::k~v'i.l;, ~-:>t.::< fiiJ-O)it~!ikf_lll!Kvr.~-t:QcJ;."C;fJI'lv;J:t.:v'. 1/h"Cv':b~:Q "rViR~!Ilb~llf." c ll:ilif(L' ~ :Q, f;J: :Qi.l>ffi/J~:IltJ:ICff:fET :Q ~1t.15Uc 1.... '"C l!lit~:J.i!!fJI!O)fd/#.i.l,; J!it;tt.;: v'. & < c t l=j:l~Ji*ltr!lBO) 13*0) ~{t.1-illf0) itit~ttkJ:lll~~':Jn!t!J;Kv;J:, JRi¥i7 :/7!1!?.~~ Ht ~lCT:Q,Jl,~i.l;~;:, ?. *$&']5-"Cv;l:,~J'fv;l: KIMURA et al. (1968) IC~v'~. ~f,:~{t:Pllf c J...."CJ;.t.::~il"v;J:, ~~il13*0)-=ttJ..CJ:t!J.!(-t:Qcv'?.ll.t~-cJ&tlh'"Cv':Q. :!Ei ·~~ * ~

Introduction summarized with special emphasis on stratigraphy and geological structure In early 1966, a geological survey was (KIMURA, T., TOKUYAMA, A., GONZALES, made jointly by Japanese and Philippine B. A. & ANDAL, D. R., 1968). Marine geologists to contribute to the knowledge molluscan fossils were collected from the of geology and paleontology of Southeast lower Gumaca formation of the lower Asia. The surveyed area is the Tayabas Miocene near Pitogo and the Hondagua Isthmus district in the southern part of formation at southeast of Hondagua, and Luzon Island, the Philippines, where the were fowarded to the writer for paleont­ marine Neogene sediments are distri­ ological study. buted widely and cover the pre-Tertiary The fossil specimens are not well pre. rocks with unconformity. The ages of served and to some extent decalcified. these sediments ranges from the lower However, 36 species of gastropods and Miocene to Pleistocene. The result ob. bivalves are recognized; 25 of them are tained from their field survey has been identifiable with species already known Received June 4, 1969; read Jan. 25, 1969 but 11 are remained indeterminable. As at the Annual Meeting of the Society. already pointed out by DICKERSON, R. E. 205 206 Yasuhide IWASAKI graphy and faunal lists of the Philip­ pine Tertiary by SMITH, W. D. (1913, ..0 . 1924), DICKERSON (1921, 1922) and CORBY, G. W. et al. (1951) are referred fully for ecological evaluation as well as for taxo­ nomical comparison. The knowledge obtained from the study of the Japanese Miocene formations such as recognition of four assemblages in the Kadonosawa­ type fauna (CHINZEI, K. & IWASAKI, Y., 1967) is app1icable in all probability to assemblage analysis of the Pitogo fauna. The species association of the Pitogo fauna can be regarded as a mixture of two paleoecological elements, the Batil­ Fig. 1. Location of Pitogo and Tayabas laria and the Anadara-Dosinia assem­ Isthmus district in the Philippines. blages. The Ostrea assemblage may not be found near the locality of this fauna, (1921), all species are either identifiable but the Pitogo fauna is, in conclusion, to the living species of neighbourhood similar in ecological and taxonomical of the Philippines or common to the nature to the Japanese Kadonosawa-type fossil species of the Miocene of Java. fauna. In the early Miocene age, the Some of these species or their. allied same faunal province covered both the forms such as Vicarya callosa JENKINS, Japanese and Philippine Islands where Anadara multiformis (MARTIN) and ]oan­ the molluscs consist of the similar nisiella cumingi (HANLEY) are found also species association found in sediments in a certain Japanese Neogene fauna in the similar environment. The Pitogo which has tropical type species associ­ fauna is of shallow embayment facies, ation. Thus the Philippihe fauna, here which corresponds to the coal-measure called the Pitogo fauna provisionally, facies of CORBY et al. (1951), and makes provides rare opportunity for compara­ a contrast with the fauna of reef or tive study. of the Japanese lower .Mio­ calcareous facies. cene fauna with the Philippine fauna in Such species as Strombus tjilongan­ species to species basis. ensis MARTIN and Oliva cf. funebralis The discussions on paleoecology of LAMARCK are absent or rare in the the Pitogo fauna may be no more than Kadonosawa-type fauna. This differ­ mere general remarks because the re­ ence can be regarded as geographical cord on species association is available within the province and seems to re­ only at one locality, and the mode of present the difference between north­ occurrence, which is not autochthonous, ern margin and near center of a faunal does not permit any further inference distribution. Species association found on the association of species. Mono­ in the Miocene formation of Tanega­ graphs of the Tertiary molluscs of Java shima Island, southern Japan, which is by MARTIN, K. (1879), those of the Gaj the southernmost record of the Kado­ Miocene of Pakistan by .VREDENBURG, nosawa-type fauna, still shows a con­ E. (1928), and discussions on biostrati- siderable difference from that of the 561. Philippine Mioc ene Mollusca 207

T-'itogo fauna. In future, however, the t he Philippines making a correlation ·Occurrence of " Vicarya-bearing fauna" with the Tertiary of Java, and described wi ll connect the Kadonosawa-type and approxin; ately 60 species of gastropods the Pitogo faunas and wi ll make clear and bivalves chiefly from the Neogene ·t he · outline of geographical variations and Pleistocene formations! SMITH (1924) within one faunal province. and DICKERSO ' (1922) made further dis­ cussions oh :, biostratigraphy of the Philippines: · · ~hr e e molluscan fau nas of Fossil molluscs from the lower the· different ages i. e. fauna of the Gumaca· formation in the Miocene Vi go group or Batan formation, Tayabas Isthmus district of the Pliocene Malumbang formation and of the Pleistocene fo rmations were i) General remarks. recognized; and the · similarity ·of the It has been well known that t he molluscan faunas between the Philip­ Neogene molluscan fossi ls occur from pines and the Indonesia was also pointed various lo calities in the Philippines and out. DICKERSON (1924) presented a paper adjacent regions such as the Indonesia on the paleogeographical change of the and Burma. jENKI NS, H. M. (1863) re­ Philippine Islands during the Neogene. ported the Tertiary, probably Miocene, HAy AS AKA, I. (1943, 1944), in his reports fossil molluscs from java. The fauna on some Neogene molluscan fossi ls from -described in the report is composed of the Philippines, recorded the associate ~ 24 species contammg Conus, Oliva, occurrence of Anadara multifonnis and Murex, Vicarya and others. MARTI N Vicarya sp. in the Philippine Miocene. (1879) described fossil molluscs from Thus the Neogene molluscan fossils of many localities of java and discussed the Philippines and the Indonesia had the Tertiary stratigraphy. He proposed been described and their stratigraphical more than a hundred of "species", in­ value had also been discussed already ·cluding several "new species" of genera in the late 19th and early 20th centuries. ·Conu s, T ereb1'a , Strombus, Anadara, C01' ­ Recently CORB Y et al. (1951) publi shed .bula. Since then, voluminous monogra­ the reconnaissance report on geology of phs of molluscan fossils had been publi­ the Philippines. Although the main shed continuall y by MARTI N (1881- 1910). subject of their report is description Materials that he treated are chiefly of and correlation of stratigraphy of the the Neogene molluscs and other shell y Republic of Philippines, they sum­ fossils occurred from java and Sumatra marized briefly the standard sequence ·(1881- 1910) and the Philippines (1901), of molluscan fossi l occurrences in chro­ these include more than 1,000 species. nological order and made check lists ·Contemporaneously NOETLI NG, F. (1895, concerning localities and range of ap­ 1901) described the molluscan faunas pearance of each species. .and associated shell y remains of the Since Vicarya callos a had been des­ Miocene formations from Burma. His cribed orig inally by jENKI NS from java, report (1901) treated both systematic its occurrence has recorded from many descriptions and stratigraphical zon in gs. other lo calities in the Philippines and These were later revised by VREDE N­ japan as well as in the Indonesia, and BURG (1921). SMITH, W. D. (1913) sum­ its stratigraphical significance has also marized the Tertiary biostratigraphy of been discussed by several paleontolo- 208 Yasuhide IWASAKI

gists. Consequently, the species has Cultellus sp., Globularia ? sp., Apollon been regarded as an index fossil of the sp., Conus gene1'alis, Voluta? sp. and. lower Miocene. MARTIN's report on the Primovula rhodia are represented by stratigraphy of Java (1879), that of the only a single specimen. Philippines (1901) and SMITH's reports No corals, bryozoans, brachiopods,. of the Philippine Tertiary (1913, 1924) barnacles and other shelly groups ex­ discussed the stratig,r_aphical occurrence clusive of molluscs are included in the· of Vicarya callosa. Sign~ficance of oc­ Pitogo fauna. Smaller foraminifers. currence of V. callosa is also discussed cannot be found in the sediments con­ in the CORBY et a/'s report. According taining the Pitogo fauna. to MART IN, SMITH and CO RBY et a! . this Collection of the specimens was made species occurs from several Miocene at road side cliff along the road from localities throughout the Philippines, i. Pitogo to Gumaca, about 4 km north: e. Mindanao, Cebu, Batan of Albay and from Pitogo. The outcrop is irregular northern Luzon. alternation of sandstone and sandy mud­ The knowledge of the Pitogo fauna stone of the upper part of the lower may be fragmental and most of its Gumaca formation, which corresponds. species are identifiable · to those des­ to Loc. 11-3 of KIMURA et al. (1968).. cribed in the classical monographs. The other locality is the path side smalL However, apart from chronological dis­ outcrop of the Hondagua formation at cussion, the Pi togo fauna renders . an Loc. 18- 12, about 2.5 km southeast from interesting problem to the study of Hondagua, where Strombus cf. vittatus paleoecology and of the extent of a and one specimen of Clementia papyracea. faunal province. were collected (see * in the list). ii) List of fossils. Species identified include 20 spp. of Bi valvia bivalves and 16 spp. of gastropods as Anadara multiformis (MARTIN) listed in p. 208 and 209. Among bi­ j oannisiella cumingi (HANLEY) valves, venerids are the most abundant Vepricardiwn multispinosum in species number occupying 7 species. (SOWERBY ) Stromboids are abundant among gastro­ V asticardium sp. pods. In addition, the materials consist Circe intennedia REEVE of considerable amounts of shell frag­ Dosinia sp. ments hardly determining their taxo­ Paphia exarata (PHILIPPI) nomic position. Corbula sp. is the most Callista (Co stacallista) erycina (LINNE} abundant and exceeds 20 in individual Katelysia hiantina (LAMARCK) number. Species whose individual num­ Sunetta concinna DuNKER ber is more than 10 are Corbula sp., *Clementia papyracea (GRAY ) ]oannisiella cwningi, PajJhia exarata. .Mactra antiquata SPENGLER T ellinella vi7'gata, Gm'i ? sp., Vastica7'­ Lutraria arcuata R EEVE dium sp. have their individual number Azorinus scheepmalleri (DUNKER) a round 10. Species whose individual Cultellus sp. number exceed 5 are Anadm'a multifo7'­ Glauconome virens (LI NNE ) mis, Strombus tjilonganensis and Vicarya Macoma sp. ca{losa. Dosinia sp., Katelysia hiantina, Tellinella virgata (LI NNE ) Lutraria arcuata, Glauconome vi1'ens, Gari ? sp. 561. Philippine Miocene Mollusca 209

Corbula sp. submultiformis (V RE DEN BURG), A. ba la­ nensis (HAYAS AKA) or even to some other Gastropoda li ving species. A. multif onnis is named Cerithidea sp. on. a Neogene fossil occurred from java. Vicarya callos a JE NKI NS The species is characterized by g lo bose Strombus (Laevistrombus ?) and considerably variable shell form; tjilonganensis MARTIN often associated with shallow sea fauna S. cf. isabella LA JV! A l~ CK like the P i togo. A. my(j ensis was des­ -* 5. cf. vittalus LI NN E cribed from the ivlioc.ene of Burma. A. Natica cf. lineata LAMARCK submultifonnis was known from the Gaj Gl obularia ? sp. Miocene of Pakistan and the Miocene Primovula rhodia (A. ADAMS ) formation of Burma. The species is APollon sp. characterized by more or less longitudi­ Bursa· (Gyrineum) m argaritula nally elongated shell form. A. batan­ (DES HAY ES ) ensis was reported from the Miocene d:Vassarius crenulatus (LI NN E) formation of Bata n Island, Albay, the .Qliva cf.'fu,nebr alis LAMARCK Philippines. This species is associated Voluta (Volutoco7' ona) ? sp. with A. multifonnis, fl. submultifonnis 11 exillum s p. and A. bat anensis were morphologically ·Co nus (L eptoco nus) generalis LI NNE distinguished from A. multifonnis by .C (Cl eobula) minimus LI NN E the original authors. However, their shell forms resemble well with one an­ iii) Taxonomical remarks. other and the discrimination is often difficult owing to the overlapping nature in shell morphology, for example, shell A nadara multiformis (MARTI N) outline, number of radial ribs, orna­ Pl. 23, figs. 6a, 6b, text-fig. 2. mentation on shell surface etc. Further, their ecological characteristics including Seven isolated (six left and one right) faunal association are closely similar v alves are examined. Shell is small to with one another. In general, Anadara medium in size, thick and strongly species having such a globose shell inflated. It appears equilateral and is show considerable morphological vari­ inaequivalve judging from the asym­ ation probably due to the environmental metry of marginal crenulation on ventral modification. The Neogene shallow ma­ :side, Length is almost equal to height, rine associations with Vicarya species thus the shell form is globose in outline. frequently include such a g lobose form Beak is prominent. Hinge plate is well as fl. multiformis in · the Indonesia, .developed , wide, but not very long, Philippines and adjacent regions . ·shorter than shell length. Outer surface Three li ving species, A. ·rhombea (BORN), has 24 to 27 non-bifurcated radial ribs A . pilula (REEVE) and A. sabinae IREDAL E, on which granulation is distinct at least vvhich occur also from the Pliocene ·and in left valve. Pleistocene formation of the Philippines The individuals, which have morpholo­ and adjacent areas, are rhorphologicall y gical characters mentioned above, are similar to A . multifonnis. However, they referable either to Anadara multifonnis are diff'erent in such ecological chara­ .{MARTI N), A. myiiensis (NOETLI NG), A. cters as biogeographicaf distribution and 210 Yas~~hide I WASAKI

27. D (Left) ; 27.9, 28.4, 14.7, 26. E: (Left) ; 27.8, 28.3, 13.1, 24ca. F (Left) ; 22.1, 22.6, 11.5, 25. G (Ri g ht) ; 23.4, 22.1,. 11.6, 26 .

.foannisiella cumingi. (HANLEY )

Pl. 23, fig. 4.

More than 11 specimens are examined,. Fig. 2. Anadara multifonnis ( M ARTIN) most of them are inner mold. Ten speci­ after M r\RTI N, 1879 x 3;5. mens are bivalved. Shell is thin, stronly inflated, and nearly round in its outline .. faunal association. Habitat of Jl. multi­ Posterodorsal marg in is straig ht from jonnis appears to be similar to that of beak, and bents with obtuse angle at the li ving A. granosa (LIN NE). In con­ posterior end. Lunule is absent. Beak clusion, the specimens concerned are is prominent and antero-dorsal margin. assigned temporary to f l. multifonnis is concaved markedly in front of beak. which includes the similar fossil "spe­ Ventral margin is round. Shell surface cies " mentioned above, rather than to has no prominent sculpture except weak recent Jl. rhombeo, A. pilula or Jl. growth striations. Hinge plate is narrow, sabinae. Further examination, however, cardinal teeth is not stout. ··. may demonstrate the close relationship The specimens at hand are identi­ between Jl. pilula, A. rhombea and Jl. fiable easily with .foannisiella cumingi multifonnis. (HA NLE Y) or its allied species. f. cumingi A similar case can be seen in the rela­ is li ving in the southeast Asia and Ja­ tions among the following Japanese pan and is also found as fossils in the "species ": A. daitolzudoensis (MAKI­ Neogene and Quaternary sediments. The YAMA ), A. lwlwhataensis HATAI & Nrsr­ difference between the present speci­ YAMA, A. lwrosedaniensis HAT AI & mens and the fossil and recent f. cumingi NISI YAMA, A . ta/wyamai NODA and A. from japan can be found in the. follow­ yatsu.oensis NODA. Vicarya-Anadara fos­ ing characteristics: Large size in adult; sil association is also found in the strongly inflated shell; strong 'convexity Japanese lo wer Miocene. Here ·again of postero-dorsal margin. T he · form the Anadara species is called such va­ characterized by the above mentioned rious names as mentioned above. Shell features is differentiated occ asionally form of the Philippine Anadm-a multi­ from .f. cumingi and is assigned ,to·such fonnis is different slightly ffom the different species as .f. oblonga ·cH·ANLEY ) Japanese Anadara daitolwdoensis in shell or .f. alata (ADAMS & REEVE) ... or J. morphology, but the two species or spe­ semiasperoides NOMURA. However, ·these cies groups are considered to be quite morphological differences among ·· four similar in ecological .characters. species seem to be not essential. Unlike Measurements in mm.-A (-Left) ; 30.0 Anadam multifonnis, variations of shell (length), 29.8 (height), 14.4 (depth), 25ca form cover much · smaller range :> in (number of radial ribs). B (Left) ; 27.5, .f. cumingi, .f. oblonga, .f. alata or iri. ]. 27.0, 14.0; 27. C (Left)·; 26.8, 30.0, 15.3, semiasperoides. However, a considerabie 561. Philippine Miocene Mollusca 211

morphological variation is detectable in rubber cast. Shell is small in size, the present specimens. Actually it is, elongated oval in outline and inflated therefore, difficult to assign the present in comparison with other species of Pa­ specimens to one of these four " spe­ phia. Test is thin. Anterior margin is cies". ]. cumingi in the sense applied smoothly rounded and ventral margin is here includes four" species",]. cumingi, broadly arcuate. Shell surface is orna­ ]. oblonga, ]. alata and ]. semiasperoides. mented with fine and regularly spaced Morphological variation of fossil ]. concentric striations. Striations. are not cumingi from the Miocene Miyazaki undulated. They are, in general, indis­ group of the southern Kyushu was dis­ tinct in umbonal area. Lunule is. present cussed by SHUTO (1957b). It shows con­ but no escutcheon is observed. Pallial siderable variation in shell morphology. sinus is moderately deep. ]. cumingi from Pitogo have more con­ This species is smaller in size and vex shell form than that from Miya­ finer in surface sculpture than most liv­ zaki, but the range of variation is much ing spedes of Pap~ia from the north­ smaller. western Pacific except for P. exarata Recent form of ]. cumingi from the (PHILIPPI). However, several fossil spe~ Philippines is large and inflated in com­ cie~, such as P. protolirata (NOETLING), parison with the Japanese forms. There P. pseudoliratus (VREDENBURG), P. exilis is a tendency that the species becomes SHUTO, P. grata tsumaensis SHUTO ftnd consistently small and flat in shell form P. suzuensis MASUDA, have small shell northwardly throughout the Miocene in adult and fine concentric ribs without and the recent. weak undulations. SHUTO (1957a, c, 1961) Cytherea (Callista) everwijni described noted on P. exilis a wide range of varia­ by MARTIN (1883) from the Neogene of tion in size and in surface ornamenta­ Java is, regarded as the same species tion. Several subspecies was proposed with ]. cumingi. based on the differences of shell outline, Measurements in mm.-A. (Right & and of surface ornamentation. Size Left); 27.2 (length), 23.6 (height), 17.4 variation of the present specimens is (depth bivaled). B (R & L); 32.9, 29.0, small and falls in the domain of SHUTO's 21.5. C (R & L); 25.4, 21.6, 16.8. D (R P. exilis exilis, P. exilis takaokaensis and & L); 27.2, 22.5, 18.5ca. E (R & L); 28.9, P. grata tsuma.ensis. However, the sP,ell 25.3, 21.0. F (R & L); 26.0, 22.5, 17.7. G ornamentation of the pr~sent specimens (R & L); 22.9, 19.3, 14.9. H (R & L) ; is similar to that of P. exilis abbreviata 27.4, 23.6, 17.5. I (R & L); 26.9, 23.0, 19.1. and P. grata tsum0:ensis, but closest to J (R & L); 22.6, 21.1, 17.5. K (Right); that of P. exarata and P. lirata (PHILIPPI). 26.1, 24.5, ?. P. lir,ata is. a living species of the south­ east Asia. P. protolirata, P. pseudoli­ ratus, P. grata tsumaensis, P. exilis and Paphia exarata (PHILIPPI) P. suzuensis are the same at least in Pl. 23, figs. 14, 15. shell morphology with P. exarata · and P. lirata .. P. lirata was proposed origi­ Nine specimens are examined. Four nally for an .intermediate form between of them are bivalved. They are inner P. exarata and P. amabilis (PHILIPPI) in mold, and therefore surfac~ ornamenta­ shell size and in distinctness of concen­ tions and hinges are observed on silicon tric ornamentation.. Tbe present sped- 212 Yas~~hide IWASAKI

mens are identified tempOrary to P. guished from C. erycina solely by its exarata. P. suzuensis described by MA­ small shell size. Therefore, three small SUDA (1966) from the Miocene Higashi­ specimens could be identified to C. pha­ Innai formation is in all probability to sianella. However, there is no definite conspecific with the present species. reason to identify small specimens with The differences among ' the above men­ C. phasiim.ella and large one with C. tioned species are found only in their erycina. These four specimens are thus stratigraphical horizons and geographi- regarded as members of one species C. cal distributions.' · erycina. Cythe1'ea ventricola and C. Both Cytherea lilacina LAMARCK illus­ macra described by MARTIN (1880) from trated by MAkTIN (1883) from Sumatra the Miocene of java are In all proba­ and P. textrix· DESH AYES illustrated by bility con'specific with Callista erycina. DICKERSON (1922, pl. 7) of the Vigo fauna Venus sumatrCl1ta and V. astartaefonnis will be . conspecific vvith the . present described also by MARTIN (1883) from species. Sumatra are possibly the same with this Measurements in mm.- A (Right & species. Left) ; 42.6 (length), 26.5 (heig ht), 17.1 Measurements .in 111m .- A (Left) ; 41 {depth bi valved). B (R & L ) ; 32.1, 20.5, (leng th), 29 (height), 7.5ca (depth). B 12.9. C (R & L) ; 33.6, 20.0, 12.5 D (Left) ; (Left) ; 14.5, 10.3, ?. C (Rig ht) ; 12, 9.5, 32.0, .18.4, ?. E (Right) ; 33.8, 20.4, ?. F 2. D (Rig ht) ; 15ca, 10.5, 3.5. (R & L) ; 30.3, 19), 10.3. G (Right) ; 30.0, i8.5, ?. H (Right) ; 41.1, 24.2, ?. I (Left) ; 32.5, 20.5, ? . Sunetta co ncmna DUNKER Pl. 23, fi :s . 9.

Callista (Costacallista ) erycina (LI NNE ) One bivalved specimen and an inner Pl. 23 , fig s. 1, 8. mold of right valve are at hand. Shell is small sized, oval in shape and not Four specimens are . collected. All strongly inflated. Test is rather thin. except one specimen are very small in Beak is pointed at slightly anterior to size. Shell is oval shaped and is not the middle of dorsal margin. Shell thick. Anterior margin is regularly surface is smooth with no distinct scul­ rounded and posterior margin is sharply pture. There is no lunule but escutcheon rounded. Beak is pointed. Lunule is is prominent. Hinge plate is narrow, distinctive. Shell surface is ornamented cardinal teeth is distinct. Pallial sinus with roughly spaced concentric grooves. is shallow. Inner ventral margin cannot Hinge teeth are Callista-Pilar ty pe. be observed exactly. Stout marginal Ventral margin is not cren ulated. One crenulations seems to be absent. inner moid specimen is medium sized, The present species can be identified 41 mm in length. This is much larger w ith Sunetta concinna DUNKER in shell than the rest. outline especially by well developed Shell form and surface sculpture of escutcheon. S . so landerii (GRAY ) is dif­ the 'pr:esent species are very much like ferent from S. con: cinna in having stout those of Callista erycina (LI NNE). There marginal crenulations. S. menstrua/is is, however: . another ailied species C. (MENKE) has ~ound and flat shell outline, phasianella (DESHA YES ) which is distin- and, therefore, can be discriminated 561. Philippine Miocene Moll~~sca 21 3 from this species. S. ovalis MARTI N is middle part of dorsal margin. Shell is another close species, which is likely to well inflated, its anterior and posterior be conspecific with S. concinna. ends are arched narrowly. Surface is i\tfeasurements in mm.- A (Right & smooth near beak without any distinct Left) ; 25 .2 (l ength), 18.6 (height). B sculpture; but.is slightly undulated con­ (Right) ; 25.8, 19.8. centrically in ventral area. Hinge is, at least, mactrid ty pe, but its definite sculp­ ture is unknown. Cl ementia papyrqcea (GRAY ) The present species has a shell outline Pl. 23, fig . 5. close to J\!Ia ct-rct (Co elomactra) antiquata SF' ENGLER. It is vH'y difficult to discri­ Two Specimens of medium size are minate M. adanso ni PHILI PPI, M. cumingi examined. One occurred from Loc. 11- RE EVE and M. contrm'ia REEVE from the 3 iii association with most of other present species· especiall y in the case of species, and the other bivalved specimen fossils. Based on·sudace ornamentatron occurred from Loc. 17-12 in the Hon­ and shell si;ze, however, the present form dagua formation. The Hondagua speci­ may be rega1 i:l ed ·as young shell of M . men which consists of outer cast and antiqiwta. ·· ····.u: 1' • inner mold is preserved comparatively Mea sureme'nts in mm.- A (Left) ; 38 well. It is characterized by remarkable (length), 28 (height), 9 (depth). · B (Right) ; concentric undulation of shell surface. 30ca, 19ca, 8.5. C (Left) ; 33, 25.5, 9.5. This undulation is well developed in its adult stage. The former specimen from the ldwer Gumaca formation is Lutraria arcuata REEVE a left valve sli ghtly deformed. Undu­ Pl. 23, fig. 12. lation of shell surface is found in its younger stage. ·Outline is elongate-oval Only one specimen -is available for in shape and inflated well. Test is thin, study. Shell surface is slightly dissolved. Postero-dorsal margin is straight. BE!ak Laterally elongate outline is characteri­ is prominent. These features of shell stic. ·Shell is somewhat swollen and not form easily permit to identify the pre­ thick. Beak is situated at anterior one­ sent specimens to Clementia papyracea fourth of the length. Maximum shell (GR AY). height is measured at posterior one-thii-d JV!easw'ements in mm.- Hondagua spe­ of the length. Postero-dorsal margin is cimen; 41.0 (lehgth), 31.5 (height), 10.5 neariy straight or slightly concaved. (depth bi valved). Gumaca specimen; Shell surface is nearly smooth but · fine 40ca, 29.5, 7.0 (Left valve). concentric striation is observable. Inner surface and detail of hinge plate are not observed. Mactra antiquala SPENGL ER The present specimen is identifiable Pl. 23 , fig . 7. Lutra;'ia arcudla RE EV E by its shell outline. L. sieboldi REEVE and L. maxima Three specimens are examined. Shell jONA S differ sli ghtly in shell size and is sub-equilateral, medium size, thin and outline. Comparable species in shell compressed triangu1ar in lateral outline. outline· may als·o be find in genera Beak 'is nearly orthogyrous, pointed at Phaxus and Cultellus. 214 Yasuhide IWASAKI

Measurements in mm.- Right; 60 present species and Burmese T. foliacea (length), 28 (height), 6.5 (depth). of NOETLING's sense. Measurements in mm.-Right; 56 (length), 31.5 (height), 5ca (depth), illus­ Tellinella virgata (LINNE) trated specimen.

Pl. 23, fig. 16.

Several specimens are examined. Most Vicarya callosa jENKINS of them are outer cast or inner mold Pl. 23, fig. 17. and are incomplete fragments. Shell is medium in size, elongated oval in shape. Five incomplete specimens are ex­ Test is not thick. Anterior end of shell amined. All specimens are broken in is rounded while posterior end is· curved their apical and basal ends. They are sharply. Beak is situated at slightly small in size. Shell form is elongated anterior to the middle. Posterior end conispiral with small apical angle, 27 to bends right-ward slightly.. ~hell surface 33•. Number of spires are estimated is ornamented with moder!J.tely spaced at nine to 13 in complete specimens. concentric rib which becomes sometimes Strong tubercles, eight to ten on one platy. A weak ridge runs from beak to whorl, arrange close to upper suture. posterior end_ Observable on inner They become weaker adaptically. Two surface are the cardinal teeth and a rows of spiral striae are present. Upper weak impression which corresponds to row corresponds with a row of tubercles, the ridge observed on outer surface. and runs connecting the tip of tubercles. Depth of pallial sinus cannot be confir­ The other row runs at middle of upper med. Cardinal teeth is small, weak ; two and lower sutures. A weak spiral ridge pieces are found in right valve. Both runs just below the lower striae. Weak anterior and posterior lateral teeth are and fine striations on shell surface are distinct in right valve. Ligamenta! plate prosoclinal in the upper half and opistho­ is short and narrow. clinal in the lower half of a whorl. The present species is, no doubt, a These specimens are identical with tellinid, and is most likely to be con­ Vicarya callosa JENKINS, a species which specific with Tellinella virgata (LINNE), is well known from the upper Paleogene although recent form has more round to lower Neogene formations of south­ shell outline, that is, short in length. east Asia and Japan. Difference of this Outline of cardinal teeth and surface species from V. callosa semperi described ornamentation coincide well with those from the Philippines by MARTIN (1901) of T. virgata. i\1/acoma sp. which occurs and SMITH (1913) in having strong tuber­ associated with the present species has cles and flattened whorl is hardly dis­ a shell form somewhat similar to T. cernible. In japan, several endemic virgata, but the differences can be found "species" and "subspecies" have been in shell outline and surface sculpture. reported by T AKEY AMA (1933), Y ABE & Tellina (Phylloda) foliacea REEVE report­ HA TAl (1938) and KAMADA (1960). Dis­ ed from the Miocene of Burma by crimination among these species has NOETLING (1901) is so similar in outline been made by the slightest difference to the present species that it is difficult in shape of cone or in surface orna­ to di~t.inguish the difference 'between the mentation.· Morphological differences 561. PhiliJJpine Miocene Mollusca 215 among the Japanese "species" seem to remarkable tubercles on shoulder of shell be not significant. It is said that at surface. Color pattern is not preserved. least two species, V . callosa japonica The present species is most likely to Y ABE & HAT AI and V. verneuili yo lw ­ be included in subgenus Laevistrombus yamai TAKEYAMA exist in the Japanese or Labiostrombus, because of its similar Miocene. The present species is closer shell morphology. Medium sized shell, in shell form and surface ornamentation row of tubercles on shoulder and simple to V . callo sa from the t ype loc ality in outer lip characterize this species, and Java, than that f rom Japan. are identical with those of Strombus V . callosa occurs commonly from the tjilonganensis MART IN, a fossil species lo wer Miocene coal-measure horizon of described from Java (T ext-fi g. 3). Other the Philippines, i. e., the Batan formation morpholog ical , ch.aracters appeared in of SMITH (1913) or the Vigo gro,up of the original <·description coincide fairly DICKERSO N (1921). Several localities are well with those: of the Philippine speci­ known as follows : The Cagayan and mens. There ' .. are no morpholog ically Ifgao districts of northern Luzon, the comparable recent species li ving in the Batangas district of central Luzon, southeast Asia. Mode of volutions ob- Bondoc Peninsula, Batan Island of Albay, eastern side of Cebu Island and the Zamboanga district of Mindanao Island. Measurements in mm.-A ; 57.5 (90) (shell height), 27.7 (36) (maximum di­ ameter), 2r (apical angle). B; 60.6 (80), 31.2 (34), 33 °. c ; 37.7 (58), 19.8 (24), 28 °.

D ; 45.5 (86), 24.6 (34), 28 o. ( ) : Estimated dimens.ions.

F ig . 3. Slrombus tjilonganensis M A ilTJ P: Strombus (Laevistrombus ?) aft~r M A ilTJ N , 1879 x 3;5. tjilonganensis MARTIN

P l. 23, figs. 2, 3; text-fig. 3

Five specimens are examined. Shell is strombiform with ver y thick test. Six spires are observable in two com­ paratively well preserved . specimens. Base of shell is slightly arcuated. Aper­ ture is compressed laterally. Outer lip is flared but not so. prominent as those of L ambis and E~tstrombus. .Test of outer li p is thickened extraordinally. Striations are not present on inner surface of the outer lip. Stromboidal notch exists near adapical end of the outer .lip.. Anterior siphonal canal bents Fig. 4. Axial section of Slron'!bu s obtusely. Each whor l has nine to 11 tjilonganensis. M Al~ TJ N . 216 Yasuhide IWASAKI served in axial section of this species that of S. isab ella especially of its fos­ resembles that seen in S. isabella sil fonn. The present specimens have LAMARCK (Text-fig. 4). close morphological resemblances to ·S. Measurements in mm.- A; 42ca (shell canarium LI NN E illustrated by DICKERSO N height), 35:0 (apertural height), 31.5 (1922, pl. 5), but the DICK ERS ON' S 5. (maximum. diameter). B; 39.0, 31.5, 26.5. canarium is more slender in having high C; 35ca, 27.5, 25.5. D; 37.0, 29.5, 25ca. spire. E; 35ca, 29.0, 22ca. 1\lfeasw·ements in mm.-A (6 spires) ; 34.0 (shell height), 23:5 (apertural height), Strombus d. isab ella LAM ARC.K 20ca (maximum diameter). B (5 spires) ; . ! . 33.0, 21.5, 18.0. Two · incomplete : specimens. . · a:re at hand: Test is thick, but,lthe · surface is . I''"·. ! , . . . . fl;tji'Sa (Gyrineum) margaritula dissolved away. Only ca:sts and molds are at our disposal. ShelP is ·small and ·, , (DES HAYES ) slender in its outline in cbn'lpari'son ·with P l. 23, fig. 15. that of the former specie's: Spire i's rather hi gh. Shoulder is round distinc­ Two specimens· are examined, but tively. Suture line is distinct and a one of them has some doubt in species narrow band runs along adapical margin identification. Shell is small in size, of whorl. Siphonal canal is not observ­ rather thin and fragile. General outline able. Outer lip is well developed and is is compressed fusiform. Whorls are lacking spines or tubercles. Adapical five in number and hig h. Shoulder is end of outer lip overlaps to adapical prominently arcuated. Surface is orna­ suture · of the spire. No striation or mented with many spiral ribs especially ornament is found inside of the lip. at lower half of whorl. There is a row Shell surfa:t e is smooth and no tubercles of tubercles on the periphery of body are found. whorl, eight tubercles in one whorls. Specimens at hand are characterized Another more or less indistinct row of by small size, and lack of spines and nodes which are smaller than tubercles tubercles on shell surface. Above men­ is found below periphery. Two varices tioned morphologicar characters are can be observed on the body whorl. similar to but not completely coincide They are wide and well developed. One with those of the re~e~1t specimens of is located' at opposite side of the other. S. isabella LAMARck in having rather Anterior canal bents slightly. Details slender form an.a ' small~r size. Fossil of aperture and posterior canal are not specimens des~ribe9 by MARtiN (1879) observable. as S. isabella · ~re v:ery 'close. to the pre­ · The preserit specimens are regarded sent form. M A RTIN '~ S. isabella var. as young individuals of Bursa , judging thersites and its allied form, S. varin­ from shell size and development of ginensis, have a 'certain r~semblance to varix. They. bear a certain resemblance the specimens at hand in having slender to Bu1'S a rana (LI NN E), B. margaritwla shell form and high spire. There re­ (DESH AY ES ) and "Ranella nobilis REEVE" mains some doubt in identification of of MARTI N (1889).. MARTI N (1899) des­ the present· form to S. isab ella, but cri·bed also B; margaritula as R. mar­ o verall sh ::! ll outline is comparable with garitula. Difference between R . mar- 561. Philippine Jl1iocene Moll~~sca 217 garitula and "R. nobilis " is represented : prominent apex, columella foldings and by number of spiral ribs, which is dif­ heavy test are still remained for ex­ ficult to count in the ill preserved spe· amination. The present specimen bears. cimens. Surface ornamentation of fossil similar shell outline to that found in and recent B. margaritula is, in spite 11 oluta scapha var. ponderosa described of their slender shell outline, similar to by MARTIN (1885) from the Neogene of that found on the present specimens. Java, but is too incomplete to be identi­ B. rana has prominent tubercles on its fied with this species. varix. Measw-ements in mm.-A; 31ca (shell Conu s (Cl eobula) minimus LINNE heig ht), 18.0 (maximum diameter). B; 22.5, 13.5. Pl. 23 , fig. 11.

Five specimens are at hand. Beside Oliva cf. funebralis L AMARCK the shell outline, section throug h axial plane is examined in one specimen (Text­ Pl. 23, fig. 10. fig. 5). Shell is rather small and cone­ Four ill preserved specimens are a t shaped not so t all. Test is thick. Side hand. They are olivid gastropods with is convex slightly. Spire is low and relatively high spiral slope and narrow spiral side is elevated slightly toward basal inductula and are sma ll in shell apex. Apex is projected abruptly and size. Four to five whorls are observed. pointed. Suture is smooth. Shoulder is Apex is projected. The color pattern rounded smoothly. Several faint spiral which is useful for identification of striations are found near base. A perture ali vid gastropods is not preserved. is narrow. In axial section, test appears The outline resembles that found in to be thick, and heavy in its outer part,. Oliva funebralis L AMARCK. High spire but columella is com paratively slender. and basal inductula suggest alliance of Secondary callus is well developed in the present specimens with 0. annulata parietal region, thus whorls of young GMELIN and 0. episcopalis LAMARCK. stage are completely filled with shell Absence of color pattern does not permit material. Protoconch is not observable. definite species identification. 0. June­ Color pattern on shell surface which is. bra/is illustrated by MARTIN (1895) from Java differs considerably from the pre­ sent specimens in shell outline. Mea surements in mm.-A; 23.0 (shell heig ht), 19.5 (apertural heig ht), 10.0 (maximum diameter). B; 20.5, 17.5, 10.5. C; 17.5, 15.5, 8.0. D ; ?, ?, ?.

Voluta ( Volutocorona) ? sp.

A fairly heavy coniformed shell is found. Preservation is incomplete and body whorl is broken. However, char­ Fig. 5. Axial section of Conus acteristic features of 11 oluta such as minims Li N NE . 218 Yastthide IWASAKI · a significant key ·for specific identifica" volcanic deposits,· that are widely · de­ tion of conid gastropods is not preserved veloped in the middle to southern region in fossil sp·ecimens at hand. of Luzon Island. Stratigraphy surround­ If one disregards i:he color pattern, ing fossil localities is as follows accord­ the present specimens are identifiable ing to KIMURA et al. (1968) : with Conus betulina LINNE or C. minimus Hondagua formation LINNE because they have similar shell f form especially in spiral slope and Miocene .Upper Gumaca formation round shoulder. Profile through axial l Lower Gumaca formation plane also suggests the same conclusion. Basement complex Only a reason why the present form is· identified with C. minimus is the differ­ The basement complex is composed ence in size. C. minimus is smaller in of older metamorphic rocks, and younger general than C. betulina. C. striatellus non-metamorphosed effusive rocks, sand­ JENKINS of MARTIN (1879) and C. glaucus stone and cherts, latter of which .are LINNE of MARTIN (1895), both from. java thought to be the Cretaceous in age. also have shell outline which is close to The lower Gumaca formation covers C. minimus. C. loroisii KIENER illustrated unconformably the basement and con­ by DICKERSON (1922, pl. 2) is closely sists of sandstone, mudstone, conglom­ similar to the present species. Further, erate, limestone and tuff. The lower the faunal association of C. loroisii listed part of the succession is siliceous and by DICKERSON is very close to the asso­ fine, but the upper part becomes coarser. ciation of the present species. Tuffaceous sediments and andesite lava Measurements in mm.-A; 35.5 (shell are found in the middle part. The height), 24.2 (maximum diameter). B; Eulepidina bearing reef limestone char­ 40.3, 28.5. c ; 32.5, 22.5. acterized by dominance of corals is a distinct horizon marker of the middle part. Fine sandstone which contains Geological outline molluscan fossils at the Locality 11-3 is upper in horizon than the Eulepidina The Tayabas Isthmus district, a nar­ limestone. The upper part is rich in row land extending from east to west coarse sediments and coal seams. This with width of less than 15 km, is located formation is widely distributed in the at ·the southern part of Luzon Island, Tayabas Isthmus district, and is char­ and connects the northern main part acterized by frequent lateral changes of of Luzon with the southern volcano-rich lithological facies and total thickness. region. Topographically the district is The Nephrolepidia limestone and cal­ a low land consisting of the widely dis­ careous sediments laying unconformably tributed Neogene and Quaternary sedi­ over the lower Gumaca formation are ments. Strikes of strata and trends of generally called the upper Gumaca for­ folding axes are .approximately parallel mation. The formation is distributed to the extension trend of the Isthmus. along the synclinal axis of the Isthmus, Stratigraphically~· .these Neogene ma­ ·and runs almost parallel to the middle rine and non-marine sediments overlies zone of the district. the pre-Tertiary metamorphic complex. The Hondagua formation is distributed There are no coverings of Quaternary in the eastern· area of the district. A 561. Philippine Miocene Mollusca 219 fault running in northwest to southeast neros conglomerate is composed of direction which is main trend of the coarse sand, conglomerate and large Philippine fault zone separates the area amount of clay and silt with sediments of the Hondagua formation from that of of volcanic origin. · Most part of the underlying formations. The lower part formation is considered to be non­ is characterized by conglomerate, sand­ marine origin and marine shell fossils stone and siltstone, while tuffaceous are scarcely found. Hondagua silt con­ beds and clean sands characterize the sists of bedded gray silt intercalating upper part. In the middle part, shell with massive sandstone. beds are intercalated. The second fossil The columnar sections presented by locality of this report, the Locality 17- KIMURA et al. and CORBY et al. do not 12 of KIMURA et al. (1968), is in the correspond exactly to each other because middle part. of the different interpretation on the The Neogene formations, especially geological structure of the district. KI­ the lower Gumaca formation, are gently MURA et al.'s lower Gumaca formation undulated into a syncline and an anti­ includes most of the non-calcareous cline. At the western area, the middle facies and Eulepidina limestone of CORBY part of the lower Gumaca formation, et al.'s Tayabas coalmeasure and Alone­ the Eulepidina limestone, is not undu­ ros conglomerate. KIMURA et al.'s upper lated, on the other hand, at the eastern Gumaca formation which is character­ side, even the overlying Hondagua for­ ized by the Nephrolepidina limestone mation is involved in gentle folding. seems to be equivalent to several lime­ In this report, the stratigraphical di­ stones of the Tayabas coalmeasure of visions and the formational names are CORBY et al. Upper silty part of the adopted from those of KIMURA et al. Hondagua formation of KIMURA et al. There is, however, at least another may correspond to the Hondagua silt. stratigraphical divisions proposed by According to CORBY et al., occurrence CORBY et al. (1951). CORBY et al.'s study of the larger foraminifers in the Philip­ on stratigraphy and on oil possibilities pines usually indicates W stage of the covers whole areas of the Republic of letter classification. Lepidocyclina in­ Philippines. The stratigraphical division eluding Nephrolepidina and Eulepidina, of the Tayabas Isthmus district as ap­ or Miogypsina characterizes the calcare­ peared in CORBY et al.'s report is to a ous member of the limestone-coalmeasure certain extent different from that of facies of the upper W stage. Vicarya KIMURA et al. callosa is said to occur from the coal­ measure facies, and is characteristic in f-~ calcareous member of the late Paleogene Miocene ~ to the early Neogene .in the Philippines l Tayabas coalmeasure as well as in the Indonesia. Thus the lower Gumaca formation which yields Basement complex the Pitogo fauna is likely to be older The Tayabas coalmeasure is composed Miocene in age, corresponding to upper chiefly of carbonaceous sand, silt, black W and/or lower X, and the age of the shales and limestone layers. Well-pre­ Hondagua formation is younger Miocene served molluscan shells found from this corresponding to the upper X and/or Y horizon are of marine origin. The Ala- of the Philippine's letter classification. _220 Yasuhide IWASAKI

In his descriptions of generalized during the Miocene Vigo stage. A stratigraphy of the Philippines, SMITH narrow land extends north to south (1924) classified the Miocene sedimentary covering an area to the east of the succession into the Batan formation,_the Tayabas Isthmus district of Luzon Island Vigo shale and Canguinsa sandstone in and the eastern part of Mindanao Island. ascending order. These formations are The other two islands were situated, one distributed widely in northern to. south­ at an area to the west of Luzon Island ern Luzon, Batan of Albay, Cebu and and the other at an area extending from Mindanao Islands. The Batan formation Marinduque to Panay Islands. Thus, is a coal-measure consisting of gray the Philippines were an archipelago in shale, sandstone and coal seams, and the Vigo stage. The Tayabas Isthmus, occupies generally the basal part of the especially neighbourhood of Pitogo, oc­ thick Neogene succession. This forma­ cupied a part of inland seas. tion is considered to be deposited in near shore to brackish environment, thus accompanied with Vicarya callosa, Notes on occurrence of the Vermetus javanus MARTI N, Corbula sp. Pitogo fauna and other molluscan fossils. Limestone facies of the formation yields Lepido­ The large outcrop of Loc. 11-3 consists cyclina sp. The overlying Vigo shale of steeply dipped alternation of sand­ is composed chiefly of thick shale which stone and siltstone which belongs to the also yields molluscan fossils such as upper part of the lower Gumaca forma­ Conus ornatissimus MARTI N and Ceri­ tion. Two layers yielding fossils are thium jenlzinsi MARTI N. found on the outcrop. The upper layer The lower Gumaca formation, at least is found in the sandstone and the layer molluscan bearing sandstone and silt­ itself consists also of weathered, brown­ stone alternation, seems to be equivalent ish, loose, clayey, fine sandstone of 10 em to the SMITH's Batan fo rmation and the thick. Aggregate occurrence of shell rest of the formation are probably to a fragments, broken gastropods and dis­ part of the Vigo shale. articulated valves, and the small size of KIMURA et al. discussed briefly the the shell characterize the upper layer. paleogeography or sedimentary environ­ Among Corbula sp., Gari ? sp., Ci1·ce ment of the Neogene formations based intennedia, Oliva cf. funeb1·alis, Apollon on lateral and vertical changes of litho­ sp., Vep1·icardium multispinosum, Bursa facies. Their conclusion is that the margaritula, Callista erycina, Conus ge­ sedimentary basin of the lower Gumaca neralis and others, which are found in formation had been reduced especiall y the upper layer, C01·bula sp. is the most in later stage into an embayment of the abundant. Non-fossiliferous massive southern area whereas the brackish to sandy mudstone of about 1m in thick­ non-marine environment dominates in ness estranges the two fossil layers. the northern area. The Honclagua for­ The lower layer consists of clark mation, at least its upper part, deposited brownish, hard, calcareous fine sand­ in shallow embayment near littoral zone. stone. Calcareous material has been In the study of the Philippine Tertiary, leached from molluscan shells. Shell DICKERSO N (1924) presented the paleo­ remains and their fragments are large geographical map of the Philippines in size compared with those of the 561. Philippine Miocene Mollusca 221 upper layer. Species contained are or if transported the distance short Vicarya callosa, Strombus tjilonganensis, from the habitat. Thus, the assemblage S. cf. isabella, Conus minimus, Anadara found in the lower layer is probably a multiformis, ]oannisiella cumingi, Paphia mixture of two different associations. exarata, Clementia papyracea, Lutraria V. callosa, A. multiformis and Corbula arcuata, Vasticardium sp., Tellinella vir­ sp. are transported to some distance gala and others. Fossils of the upper from their habitat and are deposited on and the lower layers are in general ill the muddy sand bottom of sublittoral preserved. In some specimens shell zone where the substratum has been materials have been leached away en­ settled by ]. cumingi, P. exarata and S. tirely, thus only their external casts or tjilonganensis. internal molds are available for study. Paleogeographically, the site of Loc. Shell carbonate, originally aragonite, 11-3 is estimated to be an interior of a has been altered into calcite. Thus shallow embayment, which opens south­ quite often, the original shell structure eastward, and is surrounded by a is not preserved. swampy area. Alternation of sandstone Except for a few examples, bivalved and mudstone with thin coal seams shell and gastropod with well preserved indicates oscillation between brackish aperture are found scarcely in the upper and marine environments. and the lower layers. In exceptional case, 10 of 14 individuals of ]oannisiella cumingi, and four out of 9 individuals Comparison between the Pitogo fauna of Paphia exarata are found in intact and the Kadonosawa-type fauna bivalve. No appreciable amount of worn out can be detectable either on their Stratigraphical significance of Vicarya­ shell surfaces or on external impressions. bearing molluscan fauna of the Philip­ Aperture or apex of most gastropods pines (" the Batan fauna " of SMITH are not preserved, except for Strombus (1913) and "the Vigo fauna" of DICK­ tjilonganensis, whose surface sculpture, ERSON (1921)) was discussed fully by apex or base are often preserved com­ MARTIN (1901), SMITH (1913) and DICK­ pletely. ERSON (1921). In general, the lower Species composition of both layers are part of the Philippine Miocene such as different slightly from each other. The the Batan formation is characterized by upper layer is characterized by clustered the Vicarya-bearing molluscan fauna and occurrence of Corbula sp., whereas the at the same time by the occurrence of lower layer is characterized by the coal seams just below or near to the Vicarya-Anadara association. Corbula molluscan fossil horizons. Distribution sp., Sunetta concinna, Callista erycina, of Vicarya-bearing fauna covers a wide Vasticardium sp. and Nassarius crenu­ area including the northern part of latus are common in both layers. Most Luzon Island; Batan Island of Albay; individuals of both the upper and the the Batangas district; Bondoc Peninsula; lower layers seem to be transported Cebu Island and the Zamboanga district from a distance. However, ]oannisiella of Mindanao Island. cumingi, Paphia exarata and probably Species associated with Vicarya callosa Strombus tjilonganensis of the lower in the northern Luzon listed by MARTIN layer are considered to be autochthonous (1901) are Terebra jenkinsi MARTIN, T. 222 Yast~hid e I WASAKI bandongensis MARTIN, FtLsus verbeehi the Japanese Miocene, and as far sout h MA RTIN, lV!urex g1'ooti JENK INS, Bw-sa as in Java. gyrina (LI NNE), Cardita decipiens MARTIN In the paleoecolog ical study of the and R ostellaria javana MARTI N. The Japanese Neogene molluscan fauna, other s pecies association g iven by CHINZEI & IWASAK I (1967) r ecognized DJCKERSON (1922) f rom Cebu Island is four different species assemblages in Cerithium jenhinsi MARTI N, C. herlzlotsi the lower Miocene Kadonosawa fauna, MARTIN, Voluta innexa REEVE, Natica i. e. the BatillaTia, Osh-ea, Dosinia­ s p., Conus sp. , Chione lacerata HANLEY Anadara and lllfacoma-Lucinoma assem­ and Pecten leopardus REEVE. T he former bl ages. T hey distribute zonally in assemblage includes T eTebra and CaTdita having a certain relation to depth and s pecies, whereas the latter has several the distr ibution of bottom sediments. cerithid and pectinid s pecies besides V. A molluscan fauna closely alli ed to that callosa. Although both assemblages are found in the Kadonosawa basin in spe­ somewhat different in species com­ cies composition is encountered also in position, overall characters seem to be several lower Miocene formations in similar to each other paleoecologicall y. Japan, i.e. the Kurosedani formation of T he Vicarya-Anadara association of the the Yatsuo g roup of Honshu, t he Sa kai Pitogo fauna does not s hare much formation of Kukinaga group of T ane­ similarity with these assemblages men­ gashima Island, and etc. The fauna of tioned above. Coexistence of VicaTya the Kurosedani formation is character­ sp., comparable to callosa, and AnadaTa ized by T elescopium telescopium LINNE, multifonnis is reported by HAY ASAKA Geloina luchuaca (PILSBRY), Vicw-ya cal­ (1943) from the northern part of Luzon. losa, Vicaryella ishiiana (YOKOYAMA ) T he distributions of 11icaTya-Anadara and "Anada1'a lwhehataensis HATA I & and Paphia-]oannisiella associations or NISIYAMA ". The fauna of the Sakai their ·ecolog icall y similar assemblages for mation is characterized by T . tele­ a re not restricted to the Philippines scopium, V. callosa, " A. lwlzehataensis ", b ut are traceable as far north as in Cyclina japonica KAMADA and ]oanni-

.Table 1. Ideali zed species association of near-shore 4 assemblages of the Japanese Kadonosawa-type fauna. (Modifie d after CHI NZEI & I wASAKI, 1967) Near shore Off -shore

T elescopium-G eloina Dosinia-Anadara assemblage Batillaria assemblage Ostrea assemblage assemblage

T elescopium Vicarya callosa Ostrea gravit:esta Anadam ninohensis telescopium Vicaryella ishiiana Clementia sp. Glycymeris cisshuensis Geloina luchu.ana Batillaria yamanarii Panopea japonica j oannisiella cumingi Nassarius sp. "Clinoca1·dium" Ringicula ninohensis shinjiense T apes siratoriensis .. "Anadara kalwhataensis " Dosinia nomurai Soletellina niinoensis Clementia papyracea Macoma sp. Polinices meisensis Conus tokunagai ,,· _ ( 561. Philippine Miocene Mollusca 223

.siella cumingi. The lo wer Miocene fauna others. KASENO's classification is rather which is characterized by those species rough and the A assemblage seems to is often called the Kaclonosa~a-type or be a mixture of the TelescojJium-Geloina, Yatsuo-Kaclonosawa fauna and has been Batillaria and Osl7-ea assemblages and used as a good indicator of the tropical the B assemblage includes the Dosinia­ shallow sea of the past. Anadar a and i\!Ja coma-Lucinoma assem­ Associate occurrence of Vicarya cal­ blages of (HINZE! & IWASAKI. .losa, Vicaryella ishiiana a nd " A. lw he­ The important species of the Kaclono­ hataensis " in the field indicate that sawa-type fauna are listed in the Table they all belong to the same, the Batil­ 1. Though some differences in species /.a ria assemblage of the Kaclonosawa-type composition appear between northern fauna. ]. cumingi is found commonly limit and the southernmost r ecords, with the species of the Dosinia-A nadara these can be regarded as slight geo­ :assemblage. T. telescopium and G. lu­ graphical variation influenced by eco­ dwana constitute yet another assem­ logical condition within the same bio­ blage which characterizes the inner-most geographical province. Combinations of :area of an embayment, perhaps the such particular species as " A. lw lzeha­ peculiar environment li ke mangrove taensis " and V. callosa of the Batillaria swamp. However, since mangrove assemblage, and Dosinia nomurai OTUKA swamp is not always found along the and " Clinocardium " shinjiense (YOKO­ tropical coast, the T elescojJ ium-Geloina YAMA ) of the Dosinia-Anadara assem­ .assemblage is naturally not a lways as­ blage are useful indices of the Kaclono­ sociate with the Kac!onosawa-type faunal sawa-ty pe fauna. :assemblages. The five, T elescopium­ The Amussiopecten-Ciementia-Paphia­ .Celoina, Batillaria, Ostrea, Dosinia-Ana­ ]oannisiella "assemblage" or the Amus­

Clementia papyracea constitute that of marcia nakamurai IKEBE, and sanguino­ the Dosinia-Anadara assemblage. Ab­ lariid species, Soletellina minoensis Yo­ sence of Ostrea sp. in the Pitogo fauna KOYAMA and Cult ell us izumoensis YOKO­ suggests the con(:lusion that the oyster YAMA of the Kadonosawa-type fauna bank or Ostrea assemblage usually found respectively. Thus, assemblage analysis in most examples of the Kadonosawa­ available for the Japanese Miocene fauna type fauna might not develop in this seems to be applicable also to the fauna particular area. of the Philippine Miocene. The marked combination of species, It is quite natural that the Pitogo V. callosa and "A. kakehataensis " of fauna has larger number of species the Kadonosawa-type fauna, is replaced comparable to those found in the Kado­ by that of V. callosa and A. multiformis nosawa-type fauna from Tanegashima in the Pitogo fauna. The combination than those from Okushiri Island. Such -of Dosinia nomurai and " Clinocardium" species as Ma.ctra antiquata, Strombus shinjiense of the Kadonosawa-type fauna tjilonganensis, S. cf. isabella, Oliva cf. is disappeared and is replaced by asso­ funebralis nnd Voluta ? sp. have not ciation of such species as Dosinia sp. been known from the Japanese area. or Paphia exarata and V asticardium sp. Since the recent representatives of these in the Pitogo fauna. However, Dosinia species or closely related species are sp. and Vasticardium sp. hold no longer inhabitants of tropical shallow sandy the place of the main constituents in substratum, their distributions are likely the Pitogo fauna. In the Miyazaki to be restricted to the south of the group, the Dosinia-Anadara assemblage Japanese Islands in the early Miocene of the Kadonosawa-type fauna may be age. In general, the Pitogo fauna can represented by the Clementia-Paphia­ be regarded as a southern extension, or joannisiella "assemblage". The conclu­ more exactly, a tropical representative, sion is .that the brakish sediment which of the Japanese Kadonosawa-type fauna. is characterized by the V. callosa-A. No molluscan fauna which includes multiformis combination of the Batillaria the Vicarya sp.-Anadara sp. combination assemblage becomes more muddy off­ has hitherto been reported from the shore, and it might be lived by the extensive area between the Philippines dwellers such as P. e:wrata and ]. cu­ and Kyushu. It can hardly be speculated mingi or in some places by Dosinia sp.. whether the fauna of this area is similar and V asticardium sp. to the Pitogo fauna in its species asso­ . Among 36 species of the Pitogo fauna, ciation or intermediate one between the V . . callosa, Clementia papyracea and ]. Pitogo and the Tanegashima faunas. cumingi are found also in the Japanese The lowe~ Miocene. sediments found in Kadonosawa-type fauna. A. multiformis Formosa and Iriomote Island of ,the and Conus mi.nimus have closely allied Yaeyama Islets consist exclusively of species in the . Kadonosawa,type fauna. thick coal bearing formations and the Such venerid species as Callista. ery.cina, shallow sea sediments including the Paphia exarata and If~telysia hiantina, lower Miocene molluscan fossils can and sanguinolariid sp~cies as Gari? sp., scarcely be expected to be found in this Cultellus sp. and A,zqrinus scheepmakeri area. are likely to,. fill .tp.e , places of venerid The Pitogo fauna is .also comparable specie~, .Dosinia . nomurai and Nippono- to the Miocene molluscan fauna from 226 Yas~~hide IWASA KI java, Indonesia, In the MART IN' s report Fauna of the upper . argillaceous part of (1879), molluscs with the V . callosa-A. the Gaj group is characterized by the multiformis combination were listed from occurrence of such species as Ostrea several localities. The lower Miocene multicast at a DESHA YES, Anadara sub­ bituminous sediments at one locality multifonnis (VREDENBURG), Omphalo­ near Liotjitjankang, western ] a va yield clathrum granosa (SOWERBY ), IJo sinia V . callosa, Strombus tjilonganensis, T ele­ pseudom-gu.s (D' ARCH IAC & HAIME), Pa­ scopium telescopium, C01'bula spp., Dosinia phia viTgata (SOWERBY), Vicarya verneuili spp., Paphia rimosa, Osh-ea spp. and A. (D'ARCHIAC) ana Turritella angulata multifonnis etc. The Liotjitjankang SOWERBY (BLA NF ORD, W. T., 1880; VRE­ fauna is very close to the Pitogo fauna DENBURG, 1928). Though the species. i·n its species composition and essentially are different to some ex tent from those the same in ecological characteristics. of Indonesia and the Philippines, ecolo­ Liotjitjankang is located. slightly to the gical characters of both faunas seem to south of equator, and thus the Pitogo resemble very closely to each other. fauna which is the southern extension Although the precise chronological cor­ of the Kadonosawa-type fauna extends relation of the faunas remains to be further south into the southern hemis­ studied, the Vicarya-bearing tropical phere. However, a certain kinds of fauna did exist almost contemporane­ species in the Liotjitjankang fauna are ously in the lower Miocene time along found rarely in the Kadonosawa-type the coasts of the western Pacific .and of including the Pitogo fauna. These are the · Indian Ocean. ~ rea of its distri­ Cypraea spp., Murex spp., Chama spp. bution is one of the largest among the and T ridacna spp. which are found Neogene molluscan faunas of the shal­ usually in coral reef facies. T hus, these low embayment. T1l.e Pi togo fauna · is species in the Liotjitjankang fauna are comprehensible as one of the southern likely to be derived from another Mio­ representatives of this widely distri­ cene shallow sea assemblage of parti­ buted fauna of the lndo:Pacific province. cular environment li ke coral reef facies. The assemblage attachin g to coral or Aclmowledgements.~ Fossil specimens bryozoan reef facies is not known in of the Pitogo fauna were supplied to­ the Kadonosawa-type fauna of the japa­ gether with the precise data on the nese area, although the fauna is li kely mode of occurrence and on the strati­ to have been survived in considerably. graphy of the Tayabas Isthmus district warm environment. The EulejJidina by Prof. Toshio KIMURA· of the Geologi­ limestone of the lower Gumaca forma­ cal Institute, University of Tokyo and tion, and certain elements of the Liot­ Dr. Akira TOK UY AMA of the Shizuoka jitjankang fauna suggest that .in the University. The study w.as. made un·der early Miocene, near shore reef facies the direction of Prof. Fuyuji T AKA ! of dominates in the area to the south of the Department of Geology, University the ] apanese area. Museum, University of Tokyo. Drs. The other molluscan fossils correlative Tetsuro HANA I and Kiyotaka CHI NZEI to the Pitogo and the Kadonosawa-type of the Ge9logical Institute of the same faunas are found from the Miocene Gaj University gave helpful advj ces espe­ g roup distributed . in the provinces of cially on paleoecology and · pal'eobio­ Sind, Pakistan and Cutch, western India. geography of the Miocene molluscs. The 561. Philippine Miocene Mollusca 227 writer wishes to express his sincere B. A. & A NDAL, D. R. (1968) : Geologic appreciations to these people. structures in the Tayabas Is thmus district, Philippines. Contributions to geology and palaeontology of southeast References cited Asia, 49. in Geology and Palaeontology BLA NFORD, W . T. (1880) : The geology of of Southeast Asia, KoBAY AS IH, T. & wes tern Sind. Mem. Ceo/. Surv. India, TORI YAM A, R. edit., No. 4, pp. 156- 178, Vol. 17, Art. 1 , pp. 1- 210, 1 map. Univ. Tokyo Press . CHI NZE I, K. & I wASAKI, Y . (1967): Paleo­ KOREED A, K. (1965) : Geological study of ecology of shallow sea molluscan faunae southern part of Tanegashima Island, in the Neogene deposits of northeast Kagoshima Prefecture. (MS in Japanese) Honshu, Japan. Trans. Proc. Palaeont. Cradulation thesis, Ceo/. I nst., Yokohama Soc . japan, N. S., No . 67, pp. 93- 113. Nat'/. Univ ., No. 44. CoRBY, G. W . & others (1951): Geology and MARTI N, K. (1879- 1880) : Die TerWirschi­ oi l possibilities of the Philippines. Repub. chten auf Java. 236 pp., 28 Taf., 1 Karte, Philip. , Dept. Agricul. Nat. Resources, Leiden. T echnic. Bull. No. 21, 363 pp., 57 pls. -- (1883) : Nachtrage zu den "Tertiar­ DI CKERSO N, R. E. (1921 ) : A fauna of the schichten auf Java". lter Nachtrag: Vigo group: Its bnring on the evolution Mollusken. Sammlung. Ceo/. Reichs-Mus. · of marine molluscan faunas. Philip. jour. Leiden , 1 te Ser., Beitriige Ceo/. Ost ­ Sci., Vol . 18, No.1, pp.1- 21, pl s. 1, 2. Asiens und Australiens, Ed. 1, pp. 194- -- (1.922) : Review of Philippine paleon­ 265, Taf. 9- 13. tology. Ibid., Vol. 20, No . 2, pp. 195- -- (1895- 1906) : Die Fossilien von Java. 229, pls. 1- 16. Sammlung. Ceo/. Reichs-Mus. Leiden, -- (1924) : Tertiary paleogeography of the Neue Folge, Ed. 1, Gastropoda, pp. 1- 332, Philippines. Ibid., Vol. 25, No. 1, pp. Tat. 1- 45. 11- 50, pls. 1-4. - -. (1801) : Concerning Tertiary Foss il s in H AY ASAKA, l. (1943) : Area multijonnis the Phili ppines. in Report on the geology M ARTI N from Luzpn. Trans. Nat. Hisl. of the Philippine Islands by BECKER, Soc. Taiwan, Vol . 33, No. 241, pp. 334- G.F., U. S . Ceo/. Surv., 21st Annual 340, pl. 11. Rept., pp. 487-625. -- (1944 ) : A new s pecies of Miocene Area -- (1909, 1910) : Die Fossilien von Java. from the Philippines. Ibid., Vol . .34, Sammlung. Ceo/. R eichs-Mus. L eiden, Nos. 246, 247, pp. 123- 126. Neue Folge, Ed. 1, 2te Abt., Ht. 1, 2, J ENKI NS , H. M. (1863): On some Tertiary pp. 334- 386, Taf. 46-54 . Mollusca from Mount Sela, in the Island M AS UD A, K. (196,6) : Molluscan fauna of the of Java. Quart. jou1·. Ceo/. Soc. London, Higashi-Innai formation of Noto penin­ Vol . 20, Pl. 1, pp. 45 - 73, pls. 6, 7. sula, Japan- Il; Remarks on molluscan KAM ADA, Y . (1960) : 9n the associated assemblage and description of species. occurrence of Vicarya and llicaryella in T rans. Proc. Palaeont. Soc. j apan, N. S ., the Japanese Tertiary, with the fi rst No. 64, pp. 517- 337, pls. 35, 36. description of a Paleogene species of NOETLI NG, F. (1895): On some marine fos­ Vicarya from Japan. Sci. Repts. , T ohoku sils from the Miocene of upper Burma. Univ., 2nd Ser. (Ceo/.), Spec. Vol. No. M em. Ceo/. Su1·v. I ndia, Vol. 27, Pt. 1, 4, pp. 281- 295, pls. 30, 31. pp. 1- 45+ 5, pl s. 1- 10. KASE NO , Y. (1964) : Biostratigraphic pro- - - (1901) : Fauna of the Miocene beds of blems of the Neogene strata in Hokuriku Burma. Palaeont. Indica, New Ser., Vol. region, central Japan. Fossils, No. 7, 1, Mem. Ceo/. Surv. h rdi a No. 3, 378 pp. pp. 27 - 35. 25 pis. KIMUR A, T ., T oi

Miyazaki group. japan. jour. Geol. Vicarya, with description of two Japanese Geogr., Vol. 28, Nos. 1-3, pp. 138-160, forms. japan. jour. Geol. Geogr., Vol. 10, pl. 12. Nos. 3, 4, pp. 129-144, pl. 13. -- (1957b) : Polymorphism in mollusks and UozuMr, S. & FuJIE, T. (1966) : Neogene facies differentiation. (Palaeontological molluscan fauna in Hokkaido, Part II. .basis. for the consideration of speciation­ Description of the Okushiri fauna asso­ !). jour. Geol. Soc. japan, Vol. 63, No. ciated with Vicarya, from Okushiri Is­ 745, pp. 565-585. land, southwest. Hokkaido. jour. Fac. -- (1957c) : Palaeontological aspects of Sci., Hokkaido Univ., Ser. 4, Geol. speciation. (Ditto-Il). Ibid., Vol. 63, Mineral., Vol. 13, No. 2, pp. 139-163, No. 746, pp. 636-647. pls. 11-13. -- (1961) : Palaeontological study of the VREDENBURG, E. (1921) : . Results of a revi­ Miyazaki group.-A general account of sion of some portions of Dr. NoETLING's the faunas. Mem. Fac. Sci., Kyushu second monograph on the Tertiary fauna Univ., Ser. D. Geol., Vol. 10, No.2, pp. of Burma. Rec. Geol. Surv. India, Vol. 73-206, pis. 11-13. 51, Pt. 3, pp. 224-302. SMITH, W. D. (1913) : Contributionii to the -- (1928) : Descriptions of Mollusca from stratigraphy and foiisil invertebrate the post-Eocene Tertiary formation of fauna of the Philippine Islands. Philip. north-western Inida. M em. Geol. Surv. jour. Sci., Vol. 8, No. 4, Sec. A, pp. India, Vol. 50, Pts. I, 2, 506+21 pp., 235-300, pls. 1-20. 33 pl3. -- (1924) : Geology and mineral resources YABE, H. & HAT AI, K. M. (1938): On the of the Philippine Islands. Bureau Sci. Japanese species of Vicarya. Sci. Repts., Manila, Publ. No. 19, 539 pp., 39 pb. Tohoku Imp. Univ., 2nd Ser. (Geol.), T AKEY AMA, T. (1933) : Notes on the genus Vol. 19, No. 2, pp. 149-172, pl. 21.

(TJ Higashi-Innai F-~ P'3 Kurosedani ffi'o itn t1- Tanegashima fiJFf'r~ Iriomoto *® ~ Miyazaki '8 ~ilf Tsurikake ~<] 1~ Kadonosawa f'l / iR Okushiri ~ fK Yaeyama i\ :ill: UJ Kukinaga 3if 7k Sakai ~ :j:j: Yatsuo ;\.. ~

Explanation of Plate 23. (All figures are natural size unless otherwise stated) Fig. 1. Callista (Costacallista) erycina (LINNE) ; specimen B. Figs. 2, 3. Strombus (Laevistrombus?) tjilonganensis MARTIN; fig. 2, specimen A; fig. 3, specimen B. Fig. 4. ]oannisiella cumingi (HANLEY) ; specimen A. Fig. 5. Clementia papyracea (GRAY); Hondagua specimen. Fig. 6a, b. Anadara multiformis (MARTIN); a; shell surface; b, inner surface; specimen A. Fig. 7. Mactra antiquata SPENGLEER; specimen A. Fig. 8. Callista (Costacallista) erycina (LINNE); Silicon,rubber cast of specimen, not measured, x2. Fig. 9. Sunetta concinna DuNKER; specimen A. Fig: 10. Oliva cf. funebralis LAMARCK; specimen A.· Fig. 11. Conus (Cleobula) minimus LINNE; specimen A. Fig. 12. Lutraria arcuata REEVE. Fig. 13, Bursa (Gyrineum) margaritula (DESHAYEs); specimen A. Figs. 14,, 15. Paphia exarat.a (PHILIPPI); fig. 14, .specimen A; fig. 15, specimen D. Fig. 15. Tellinella virgata (LINNE); inner mold. Fig. 17. Vicarya callosa JENKINS; specimen A. IWASAKI: Philippine Miocene Mollusca Plate 23

4

1

9

13 Trans. Proc. Palaeont. Soc. Japan, N.S., No. 77, pp. 229-234, pl. 24, April 10, 1970

562. A NEW SPECIES OF SAGENOPTERIS FROM NARIWA, SOUTHWEST HONSHU, JAPAN*

· KAZUO HUZIOKA

Institute of Mining Geology, Mining College, Akita University

Jtz~;J,.[ Sagenopteris (ry-tJTH1: Sagenopteris nariwaensis c ,;,,cg; l t::. rV<:~~~(ry tTrHll fi!JX:~~~hc'f:f)):(ry :*~~~~ !tJ £\: G ~;: ~ -=> -c JR J:Z(rytfrilHU fJ l*!$ ~ ;h,t::.

In 1905, YOKOYAMA first described OISHI (1930, 1931, 1932, 1938, 1940) and some fossil plants from the Nariwa OISHI and HUZIOKA (1935, 1936, 1938), district in Okayama Prefecture and who described about one hundred considered this plant-bearing bed to be species of plants. These plants are Upper Triassic in age. The fossil stated to be closely related to the plants, called by the name of the Na­ Rhaetic floras of Europe and Greenland. riwa flora, was studied in detail by The Nariwa Formation which bears the

~ Locality of Sa~nopteris nariwaensis

~ Suzuriishi Group

•

Fig. 1. Localities of Fossil Plants in the Eda area.

* Received June 7, 1969; read June 15, 1969. 229 230 Kaz~w HUZIOKA

Nariwa flora overlies the ]ito Formation Filicales incertae sedi s which yielded Entomonotis ochotica of 19. Sphenopteris gracilis Or s i-II the Norian stage with conformity, and 20. S. sp. both of the formations are regarded by 21. Cladophlebidium ? olzayamaensis Orsi-Ir· Japanese geologists as representing the et H UZ I OK A 22. CladojJhl ebis bitchuensis Or SI·II Norian stage of the Upper Triassic. 23. C. denticulata (BR ON GNIA RT) KOBATAKE (1954) and KON'NO (1962) 24. C. gigantea Or s HI added two new species of Equisetales 25. C. haiburnensis ( LI NDLEY et H UTTON). to the Nar iwa flora. The Nariwa flora BRONGN IAR T as now known consists of 104 valid 26. C. nariwaensis Or sHI et H uZ IOK A plants which are distributed in the 27. C. nebbensis (BRON GNI A RT) Pteridophyta and the Gymnospermae, 28. C. pseudodelicatula OISH I including the present nevv species of 29. C. racibors/lii ZEILLER Sagenopteris. The following is a com­ 30. C. raciborskii forma integra Or s HI el plete list of the fossil plants hitherto T A K AHAS I known from 96 localities of the Nariwa 31. C. (OsmundojJsis ?) subplectrop!wra district : Or s i-I r et H uz iOKA 32. C. tenue OI SHI et H u zroKA Table 1. List of the fossil plants 33. C. sp. from Nariwa. 34. Spiropteris sp. PTERIDOPHYTA Equisetales Marattiaceae 35. A nnulariopsis inopinata ZEILLER ?- 1. Marattiopsis muensleri (GOE PP ERT) 36. Pseudolobatannularia densiflora Sc HIMPER K OBAT AKE Osmundaceae 37. Equisetites multidentatus OISHI 2. Todites goeppertianus (MONS T ER) 38. E. nariwaensis KON'NO K RASS ER 39. E. sp. 3. T. prince ps (PRESL) GoTH AN 40. Neocalamites carrerei (ZEILLER) <1. T. williamsoni ( BRONGN IART) H A LLE SEWARD 41. N. hoerensis (Sc HI MPER) H A LLE Dipteridaceae 42. Phyllotheca sp. 5. Clathropteris elegans OISHI GYMNOSPERMAE 6. C. meniscoides BRONGNIA RT Cycadophyta 7. C. obo vata OI SHI Cycad ales 8. Dictyophylium muensteri (GOEPPERT) 43. Ctenis japonica OISHI NATHORST 44. C. talzamiana OISHI et H u zroK A 9. D. nilssoni (BRONGNI ART) GoEPPERT 45. C. yabei 0ISI·I I 10. D. spectabile NATHORST 46. Nilssonia acuminata ( P R ESL) 11. Goepperlella varida OI SHI et H uz i OKA GOEPPERT 12. H ausmannia (Protorhipis) crenata 47. N. brevis BRONGN I ART ( N ATHORST) MoELLER 48. N. muensteri (PRESL) SCI·II MPER 13. H . (P.) nariwaensis 0ISI·II 49. N. orienta/is H EER 14. H . (P.) dentata OI SHI 50. N. simplex OISHI 15. Thaumatopteris elongata 0ISI·II 51. Cfr. N. tenuicaulis ( PHI LLIPS) 16. T. kochibei (YoKOYAMA) O ISHI et Fo_x -STR AN GW A YS Y A M AS ITA Bennettitales 17. T. nipponica 0ISI·II 52. Otozamites huzisawae OI SHI et 18. T. pusilla (NATHORST) O rs r-rr et H UZ I OK A Y AM AS ITA 53. 0. landfolius OI SHI 562. N ew Sagenopteris 231

54. 0. molinianu. s ZI GNO 95. P . schenki H EER 55 . P terophyllu.m aequ.ale (BI~O NGN I A RT) 96. Cfr. Storgaardia spectabilis H AR R I S NAT HORST 97. S tenorachis bi tch!te1\Sis OI SHI 56. P. angustu.m (BRAUN) G oT H AN 98. S . elegans OI SHI 57. P . ctenoides OI S HI 99. S . (Ixostrobus ?) kon1:an.us O ISHI et 58. P. cfr . distans M oR RI S H UZ I OKA 59. P. jaegeri B RONGN I A RT 100. Swedenb01·gia cryptomerio1:des 60. P . schenki Z E I LLER N A T H ORST • 61. P. sen·at u. m OI S HI et H uz i OKA 101. 5. major H ARR I S 62. P. sp. a Gymnos permae? incertae sedis 63. P. sp. b 102. Campyrophyllum hoermanni G OTH AN ? 64. P. sp. c 103. CzekanowsiJurassic. Compared with Productive organs likely in connection those two species, the present one has with this leaf have 'never been found. narrower lanceolate leaflets. The leaf figured in Pl. 24, fig. 1 ·is a Five valid and two undeterminable well preserved specimen though its species of Sagenopteris have been known petiole is missing. The median leaflets from the Japanese Mesozoic formations, are 6 em in. length and 1 em in breadth. as shown in Table 2. Pl. 24, figs. 2-6, show petiolated leaves Occurrence: Loc. 96, Higashi-Eda, with a stout petiole, which is 1 mm Nariwa, Nariwa Town, Kawakami-gun, broad and more than 2 em long, with Okayama Prefecture; Nariwa Formation six leaflets at its top, somewhat thick­ (Norian, Triassic). ened. The median leaflet in Fig. 2 is Plants associated with S. nariwaensis more than. 6 em , long . and 1.6 em wide. at Loc. 96: Clathropteris meniscoides The hteral veins are generally qbscure BRONGNIART, Cladophlebis denticulata 562. New Sagenopteris 233

Table 2. Species of Sagenopteris in Japan.

Species Occurrences

S. nariwaensis HuziOKA Nariwa Formation, Okayama Pref.; Norian, Upper Triassic.

S. ni/ssoniana (BRONGNIART) WARD Yamanoi Formation of the Mine Group, Yama­ (syn. S. rhoifolia PRESL, S. sp. guchi Pref. (OISHI and TAKAHASI, 1936, OISHI, YoKOYAI\·IA, 1905; S. sp. OISHI and 1940) ; Carnian, Upper Triassic. HUZIOKA, 1938) Nariwa Formation, Okayama Pref.; Norian, Upper Triassic. Kuruma Formation of Toyama and Nagano Prefs. (OISHI, 1940) ; Lias, Lower Jurassic.

S. petiolata OISHI (1940) Kiyosue Formation of the Toyonishi Group, Yama­ guchi Pref.; Upper Jurassic. Utano Formation, Yamaguchi Pref. (TAKAHASI, 1957) ; Middle Jurassic.

S. phillipsi (BRONGNIART) PRESL Kuzuryu subgroup of the Tetori Group, Fukui (syn. S. paucifo/ia PRESL (OISHI, Pref. (OISHI, 1940, KIMURA, 1958); Upper 1940) Jurassic.

S. ? inequilateralis OISHI (1940) Yuasa Formation of the Ryoseki Series, Wakayama Pref. ; Lower Cretaceous.

S. sp. YoKOYAMA (1889) Tetori Group, Ishikawa Pref.; Upper Jurassic.

S. sp. KIMURA (1958-a) Kizaki Plant Bed, Nagano Pref.; Upper Jurassic.

(BRONGNIART), C. nebbensis (BRONG­ of Caytonia. Ann. Bot., N.S. Vol. V, NIART), Taeniopteris minensis OISHI, No. 17, pp. 47-58. Podozamites lanceolatus (LINDLEY et KIMURA, T. (1958-a): Mesozoic Plants from HUTTON), and etc. the Kizaki District, Nagano Prefecture, Japan. Jubilee Pub/. Commem. Prof. H. Collectors: Messrs. Hisao OTSUKA, FUJIMOTO, pp. 135-138. Takeo 0GA WA and Hideshi HIRAMA TSU. - (1958-b): On the Tetori Flora (Part Depository: The Nariwa Museum, 3). Mesozoic Plants from the Kuzuryu Nariwa Town, Okayama Pref. Sub-Group, Tetori Group, Japan. Bull. Sen. High School attached to Tokyo Univ. References Educ., No. Il-2, pp. 1.-47. KoBATAKE, N. (1954): On the Mesozoic Fo:-ITAINE, W.M. (1889): The Potomac or Verticulate Leaves from Japan and Korea younger Mesozoic Flora. U.S. G.S. Mon. with some Views of Lobatannularia and 15, pp. 1-377. its Affinities. Sci. Rep. No. 3, South HARRIS, T.M. (1932) : The Fossil Flora of Col/. and North Coli., Osaka Univ., pp. Scoresby Sound East Greenland. Part 3. 71-80. Caytoniales and Bennettitales. Med. om KoN'NO, E. (1962) : Some Species of Neo­ Greenland, Ed. 85, No. 5, pp. 1-133. calamites and Equisetites in Japan and -- (1940-a): Caytonia. Ann. Bot., N.S. Korea. Sci. Rept. Tohoku Univ., 2nd Vol. IV, No. 16, pp. 713-734. ser. (Geology), Special Volume, No. 5 -- (1940-b) : On some Jurassic Specimens (KON'NO Memorial Volume), pp. 1-47. of Sagenopteris. Ann. Mag. Nat. Hist., OISHI, S. (1930) : Notes on some Fossil Ser. II, Vol. VI, pp. 249-265. Plants· from the Upper Triassic Beds of -- (1941) : Caytonanthus, the Microphyll Nariwa. Jap. Jour. Geol. Geogr., Vol. 234 Kazuo HUZIOKA

VII, No. 2, pp. 49-58. daceae. jour. Fac. Sci., Hokkaido Imp. -- (1931): Yabeiella sp. from the Japanese Univ., Ser. IV, Vol. III, No. 2, pp. 135- Triassic. Ibid., Vol. VII I, No. 4, pp. 184. 357-359. -- and -- (1938) : Fossil Plants from -- (1932) : The Rhaetic Plants from the Nariwa. A Supplement. Ibid., Vol. IV, Nariwa District, Okayama Prefecture, Nos. 1-2, pp. 69-101. Japan. jour. Fac. Sci., Hokkaido Imp. OISHI, S. and E. T AKAHASI (1936) : The Univ., Ser. IV, Vol. II, Nos. 3-4, pp. Rhaetic Plants from Province Nagato. 257-379. A Supplement. Ibid., Vol. III, No. 2, -- (1938) : The Japanese Equivalents of SEwARD, A. C. (1910) : Fossil Plants. Vol. the Lepidopteris and Thaumatopteris Zones II. of East Greenland. Proc. Imp. Acad. T AKAHASI, E. (1957) : Fossil Flora of the Tokyo, Vol. 14, pp. 77-80. Toyora and the Toyonishi Groups, Yama­ -- (1940): The Mesozoic Floras of Japan. guchi Prefecture (in Japanese). Yama­ jour. Fac. Sci., Hokkaido Imp. Univ., Ser. guchi jour. Sci., Vol. 8, pp. 79-82. IV, Vol. V, Nos. 2-4, pp. 123-480. YoKOYAMA, M. (1889): Jurassic Plants from OISHI, S. and K. HuziOKA (formerly YAMA­ Kaga, Hida and Echizen. jour. Coil. Sci., SITA) (1935) : On the Genus Sweden­ Imp. Univ. Tokyo, Vol. IV, Art. 1, pp. borgia NATHORST and its Occurrence in 1-89. the Nariwa Bed,. Okayama Pref., Japan. -- (1905): Mesozoic Plant,s from Nagato Proc. Imp. Acad. Tokyo, Vol. 11, pp. and Bitchu. Ibid., Vol. XX, Art. 5, pp. 438-440. 1-13. -- and -- (1936) : On the Fossil Dipteri-

Higashi-Eda !ft: f}{ Nariwa Town JjJ(:~jji!IJ Eda t:t. Kawakami-gun ) I UJtl; Yamamoto UJ :;js:

Explanation of Plate 24

Figs. 1, 2, 3, 4, 5, q, 6a, 7, 7a. Sagenopteris nariwaensis, sp. nov. Loc. 96, Higashi-Eda, Na­ riwa, Nari.wa-town, Kawakami-gun, Okayama Prefecture. HUZIOKA : New Sagenopteris Plate 24

6

x3

SHIBATA photo. Trans . Proc. Palaeont. Soc. Japan, N .S., No. 77, pp. 235 - 2<12, pl. 25, April 10, 1970

563. FRESHWATER MOLLUSCS FROM THE COAL-BEARING OWADA FORMATION, SOUTHEAST RUMOI, HOKKAIDO, JAPAN*

HIROSHI NOD A

Institute of Geology and Paleontology, Faculty of Science, Tohoku University, Sendai, Japan

:ll:.ifiJiD: W/M rtl· ri'HRi~ll :7c ;f 11 E13~&.1~iilli i*7J<: :f~~ ft.Ci fc-::>v,-c : ::ll:.i4ij ill f.!l pJinn Wi:r!Hm 1c 5J1fi -9- ~ :k;fQEl3 ~@U~ J:: fJ 1£ Lt.: ii.bJcl1ii f\1 U:J :a: t§!n·JL.t.: *~i 51~ 1.7< 0) -Wfilill :a: 15- tr 1i.fift u n i o uryuensis Suzu KI, L anceolaria piscifonnis (YoKOYAMA), M argaritifera perdahurica Y oKOYAM A, Margaritijera owadaensis NoDA, n. sp., Viviparus cf. uryuensis Y or

Introduction Acknowledgements

During the stratigraphic work in the The writer here wishes to express his Owada coal-field (SUGAI, 1968), the writer hearty thanks to Professor Kotora HAT AI collected some fresh-water molluscan of the Institute of Geology and Paleont­ fossils from the coal bearing Owada ology, Faculty of Science, Tohoku U ni­ Formation exposed around the Owada versity for his kind supervision and Colliery, southeast of Rumoi City, Hok­ encouragement during the present study. kaido (Fig. 1). Acknowledgements are also due to As­ The stratigraphy, geology and pale­ sociate Professor Tamio KoT AKA of the ontology of the Rumoi and Uryu coal­ Tohoku University for his kind sugges­ fields situated near the present area tions and discussions on the biostrati­ have been studied by many workers, graphy, Mr. Kimiji KUMAGAI for his yet there still remain some problems photographic work and to the Ministry ·Concerning the stratigraphy, geological of Education of the Japanese Govern­ age and correlation based upon the · ment for financial support. lithological, structural and paleontologi­ ·cal evidences of the Owada Formation, which is covered by the fossiliferous Historical Review Miocene Yudoro and . Togeshita forma­ tions with unconformity. YAMANE (1912a, b), liZUKA and UEMURA (1920a, b) reported on the geology around the Owada coal-field (SUGAI, 1968). END O * Received Jul y 26, 1969; read June H , 1969 (1931) in his study of the plants from at Yakohama. the Japanese Paleogene proposed the

235 236 Hiroshi NODA name of the Owada coal-bearing Forma­ age suggested by T ANAl (1956). tion but without definition; the strati­ graphic name was adopted by NAGAO (1933). Remarks on the Fresh-water Molluscs HASHIMOTO (1950) studied the strati­ from the Coal-bearing graphy around the Owada coal-field and Owada Formation defined the Owada coal-bearing Forma­ tion. He (HASHIMOTO, 1950) recognized The coal-bearing Owada Formation a stratigraphic break between the Owada covers with unconformity the unknown Formation and the Yudoro Formation Paleozoic Kumaneshiri Formation which and this was accepted by TsuSHIMA is mainly composed of schalstein and and YAMAGUCHI (1952), who considered slate and is covered by the Yudoro and the formation to be Miocene in age Togeshita formations with unconformity. based upon the geology and good quality The formation distributed around the of the coal. HASHIMOTO (1950) collected Owada Colliery is composed of pebble a freshwater molluscan fossil, Corbicula conglomerate and medium grained sand­ atrata tokudai (YOKOYAMA), at Baba­ stone which yielded the fresh-water Tanzan-no-sawa where the Owada For­ molluscs described in the present article. mation is distributed. Fresh-water molluscan fossils collected In 1956, T ANAl stated that the Owada and discriminated from a pale brownish Formation is characterized by Paleogene gray tuffaceous medium grained sand­ plants as Onoclea, Osmunda, Equisetum, stone exposed along the national road Ulmus, Planera, Mallotus, Pueraria, Pla­ side near Owada and on the opposite tanus, J\!Jaelea, etc., and he correlated the side of the Rumoi River (see locality formation with the upper part of the map) are Unio uryuensis SUZUKI, Lanceo­ Ishikari Group. From the heavy mineral laria pisciformis (YOKOYAMA), Margariti­ association, T ANAl (1956), IrJIMA and fera perdahurica YOKOYAMA, Margariti­ T ANAl (1955) and li]IMA (1957) recognized fera owadaensis NODA, n. sp., and Vivi­ similarity between the Owada Formation pants cf. uryuensis SUZUKI. These fossils and the Ishikari Group. Recently, HITO­ are restricted in geological and geo­ SUGI and SASAKI (1959) supported their graphical distributions. For example, opinion. Unio uryuensis SuzuKI had been record­ The age of the fresh-water mollusc ed from only the Upper Tachibetsu Corbicula atrata tokudai (YOKOYAMA) Formation* (YOKOYAMA, 1932; SUZUKI, collected by HASHIMOTO (1950) from a 1941a, 1942), Margaritijera pisciformis rolled block in the area of distribution (YOKOYAMA) is known from the Lower of the formation was questioned at that and Upper Tachibetsu Formation (YOKO­ time as to whether it indicated the YAMA, 1932), the coal-bearing Yubari Paleogene or the Neogene. Subsequent­ Formation (SuzuKI, 1941a), Lower Corbi­ ly, TSUSHIMA and YAMAGUCHI (1953) cula bearing Formation (SUZUKI, 1942, reported Viviparus sp., Lanceolaria sp. 1944), Middle Uryu Group (SUZUKI, 1942), and Margaritijera sp. from the forma­ Margaritijera perdahurica YoKOYAMA tion and based upon them they consid­ occurs from the coal-bearing Yubari ered the formation to be Miocene in age and this was later accepted by * =Stratigraphic names quoted; strati­ TAKEDA (1954), contrary to the Paleogene graphic nomenclature not undertaken. 563. F1·eshwater Molluscs j?·om Rt~mo i 237

Fig. 1. Map s howing the positions of the fossil lo calities.

Formation (SUZUKI, 1942, 1944), Yubetsu semblage cited above is known only Formation (SUZUKI, 1942), Lower Corbi­ from the central part of western Hok­ cula bearing Formation (SUZUKI, 1942, kaido in Japan. This fauna comprises 1944), Tachibetsu Formation (YOKOYAMA, warm water dwellers different from the 1932; SUZUKI, 194lb), Wakkanappe For­ freshwater molluscan assemblage from mation (SUZUKI, 1942) and Viviparus the Sasebo Group in North Kyushu uryuensis SuzuKI is known from the (UEJI, 1934), the coal-bearing Jinbu For­ Upper Tachibetsu and Lower Tachibetsu mation in Mie Prefecture (SUZUKI, and Formation (YOKOYAMA, 1932; SU ZUK I, OYAMA, 1946), the Kaura Formation in 194lb), and Middle Uryu Group (SUZUKI, Shimane Prefecture (SUZUKI, 1949) and 194lb) all in Hokkaido. from other sporadic occurrences in Na­ From the above cited data, the coal­ gano Prefecture (OMORI and lBARAGI, bearing Owada Formation is considered 1966 ; SUZUKI, 1949). to correspond with the Upper Shiroki The T appu and Shimokine formations to Tachibetsu Formation of the Uryu are not developed in the Owada area Group. Because the lower and upper where the Owada Formation is offiapped parts of the Owada Formation are by the Miocene Yudoro and Togeshita missing in the present area, exact cor­ formations. This distribution of the relation with other areas is difficult. different strata seems to indicate that However, in broad sense, the formation the Owada area may have been a mar­ may be said to be a correlative of the ginal area favourable to the dwelling of Uryu or Ishikari Group. The geological freshwater molluscan assemblages and age of the Uryu Group was considered adjacent to the Kabato Massif which is to be Eocene by T ANAl (1950), ASANO mainly composed of Paleozoic rocks. (1962), OH ARA (1966), OHARA and KA N'NO It is thought that the Miocene Yudorv (1969) and SUGAI (1968). and Togeshita formations were deposit­ The fossil fresh-water molluscan as- ed after a long period of eros.iori · sub- 238 Hi1·oshi NODA sequent to the deposition of the coal­ teeth along the posterior dorsal side. bearing Owada Formation. This view The posterior ridge is rather blunt. is also upheld by the unconformity be­ The shell is sculptured with blunt ele­ tween the formations lying unconform­ vated concentric growth lines. ably on the Unknown Paleozoic Kumane­ The species is distinguishable from shiri Formation. It is considered that the Recent species Unio biwae KOBELT, those elevated land areas were distri­ 1879 or Uuio douglasiae GRIFFITH. and buted in the Owada area during the PIDGEON, 1834 (fide KUROD A, 193i) in Miocene Yudoro stage. · From such as­ having rather shouldered dorsal margin sumptions, the present writer is inclined and more inflated shell. Among the to the view that the coal-bearing Owada fossil species, M argaritifera perdahurica Formation was deposited during the YoKOYA MA (1932) resembles the present Paleogene, and the age may be Eocene species in shell form but the latter dif­ based upon the freshwater molluscan fers from the former in having a pos­ fossils mentioned above contrary to the terior eleva ted ridge. Miocene age stated by TSUSHIMA and The shell collected from the coal­ YAMAGUCHI (1953) and T AKED A (1954). bearing Owada Formation is rather of medium size and measures about 5 em in length and 3 em in height. Description of Species The specimens of intact and isolated valves were collected from a tuffaceous, Family U nionidae FLEMI 1G, 1828 medium grained sandstone (Loc. no. 1) where they are rather common. Subfamily Unioninae FLEMING, 1828 Locality and formation : Locality nos. Genus Unio PHILIPSON, 1788 1, 2, Owada Formation. Depository: IGPS* coil. cat. no. 86890. Subgenus Unio s. s.

Unio (Unio) uryuensis SUZUKI, 1941 Genus L anceolaria CONRAD, 1853

Pl. 25, figs. 1, 2, 4, 5, 7, 13 Lanceolaria piscifo nnis (YOKOYAMA, 1932)

194lb. Unio uryuensis SuzuKI, jour. Fac . Pl. 25, fig . 10 Sci., Imp. Univ. , Tokyo, S ec . 2, vol. 6, pt. 2, p. 25 - 26, pl. 1, figs. lla-c. 1932. Nodularia piscif ormis YOKOYAMA , jour. 1942. Un io uryuensis SuzuKI, Su zuKI, japa­ Fa c. Sci. Imp. Univ ., Tokyo, S ec. 2, vol. n ese j our. Ceo!. Ceogr., vol. 18, no. 4, 3, pt. 6, p. 243, pl. 3, fig s. 1- 2. p. 151- 152, pl. 17, fig s. 3, 6. 194la. Lanceolaria piscifonnis (YOKOY Alv!A) , SuzuKI , Ibid., vol. 6, pt. 1, p. 8- 9, pl. The present species was originally 1, figs. lOa- c. described based upon the specimen 194lb. Lanceolaria pisciformis (YoKO YAMA), from the Upper Tachibetsu Formation, SuzuKI, Ibid., vol. 6, pt. 2, p. 26- 28, Uryu coal field by SuzuKI in 194lb. pl. 1, fig s. 12-13. The species is characterized by the 1942. Lanceolaria pisc if onnis (YOKOYAMA), SuzuKI, japanese jour. Ceo !. Ceogr., transversely elliptical form, with both anterior and posterior dorsal parts * =Abbreviation for Institute of Geology rather shouldered. The beak is situated and Paleontology, Faculty of Science, Tohoku anteriorly and there are long lateral University, Sendai. 5 63. F1·eshwate1· Moll~~scs fron~ Rumoi 239

vol. 18, no. 4, p. 152- 153, pl. 18, figs . 6. (fide S UZUKI, 1941c). 1- 5, pl. 19, figs. 1- 2. 1932. Nodularia biwae Y oKOYAMA (not Ko­ 1944. Lanceolaria pis6formis (YoKOYAMA) , BEL T, 1879), jour. Fac. S ci., I mp. Univ ., SuZUK I, Trans. P roc. Paiaeonl. So c. Tolzy o, Sec. 2 , vol. 3, pt. 6, p. 243- 244, j apan, no. 181, p. 7. pJ. 4, fi g. L!_ 1932. M argarilifera perdahurica Y orAnimal Pl. 25, fig. 6 Kingdom, vol. 12, p. 1-601, pls. 1-15 .. (fide KuRODA, 1931). Compared with: HASHIMOTO, W., 1950, Geology of the en-· 1932. Viviparus uryuensis YoKOYAMA, jour. virons of the Kabato Massif, Part 1, On Fac. Sci., Imp. Univ., Tokyo, Sec. 2, the base of the Upper Mizuho To in the 563. Freshwater Molluscs from Rumoi 241

Rumoi oil and coal fields (in japanese). Gesel., vol. 11, p. 1-171, .pis. 1-23. (fide Bull. Geol. Commit. Hokkaido, no. 13, p. KuRODA, 1931). 1-8, 1 map, 1 table. KuRODA, T., 1931, An illustrated catalogue HITOSUGI, T. and SASAKI, M., 1959, Geolog­ of the Japanese shells (in japanese). ical map of the coal-fields of Japan III. Venus, voi. 2, no. 6, append. p. 27-76, Explanatory text of the Owada district, figs. 43-93. Rumoi coal field (in japanese with English NAGAO, T., 1933, Paleogene (in japanese), abstract). Geol. Surv. japan, 28 pp. Iwanami Koza, Iwanami Shoten, 120 pp. liJIMA, A., 1957, Preliminary note on the OHARA, S., 1966, Stratigraphy and geological heavy mineral association of the Ishikari structures of the Tertiary deposits in the Series (Eocene-Oligocene) in the Ishikari, Uryu coal-field, Hokkaido, Japan (in japa­ Kabato and Rumoi coal fields in Hokkai­ nese with English abstract). jour. Colt. do, Japan. jour. Geol. Soc. japan, voi. Arts. Sci., Chiba Univ., vol. 4, p. 617- 63, no. 737, p. 67-81, pl. 3, figs. 1-4, tabs. 630, figs. 1-6, tabs. 1-2. 1-3. -- and KAN'NO, S., 1969, Mid-Tertiary liJIMA, A. and T ANAl, T., 1955, On the molluscan fauna of the Uryu coal field heavy mineral association of the Ishikari (in japanese). Fossils, no. 17, p. 41-49, Group, Part 1. Especially on the geo­ figs. 1-2. logic age of the Kabato and Owada coal­ OMORI, M. and IBARAGI, N., 1966, On the bearing Formation (japanese abstract fresh-water molluscan fauna collected only). jour. Geol. Soc. japan, voi. 61, from the Komoro Group, Nagano Prefec­ no. 718, p. 332. ture (in japanese). jour. Soc. Earthsci. IrzuKA, Y. and UEMURA, M., 1919, Report Amat. japan, no. 17, p. 41-43, pis. 1-2, of the geology of Rumoi and Tomamae figs. 1-4. oil and coal fields (in japanese). Min. SASA, Y ., 1938, On the stratigraphic posi­ Surv. Rep., Geol. Surv. japan, no. 76, p. tion of the coal deposits in the Uryu 74-90, pl. 1. coal fields in Hokkaido (in japanese). -- and -- 1920, Report of the geology Trans. Hokkaido Coal-mining Soc., no. of the Rumoi coal and oil fields, Teshio 291, p. 1-16, pl. 1. Province (in japanese). Ibid., no. 30, p. SUGAI, K., 1968, Geological maps of the coal 1-68, pis. 1-2, figs. 1-2, tabs. 1-10. fields of Japan VIII. Study on the strati­ JIMBO, K, 1890, Hokkaido Chishitsu Rya­ graphy and geological structure of Uryu­ kuron (in japanese). Hokkaido Prefec­ Rumoi coal fields, Hokkaido, Japan (in ture, p. 1-94, pls. 1-7, figs. 1-24, tabs. japanese with English abstract). Geol. 1-22 (fide SUZUKI, 1941c). Surv. japan, 61 pp. KrRA, T. 1959. C:>loured illustrations of the SuzuKI, K., 1941a, Some non-marine shells shells of Japan (in japanese). Hoikusha, from the Oligocene Ishikari Series in the 23 pp., 71 pis. Ishikari coal-field, Hokkaido. jour. Fac. KoBELT, W., 1879, Fauna molluscorum ex­ Sci., Imp. Univ., Tokyo, Sec. 2, vol. 6, tramarinorum Japoniae, nach den von pt. 1, p. 1-11, pis. 1:-2, figs. 1-2. Prof. Rein gemachten Sammlungen bear­ -- 1941b, Notes on the Tertiary non-ma­ beitet von. Abhand. Senckenberg. Naturf. rine Mollusca from the coal-field of Uryu,

Baba-Tanzan-no-sa wa .~t~ mw 0) iR Shiroki s Ishikari ::£i ~;t Tappu iJ! *:fli Jimbu r'l' lft Tachibetsu ~ 7J j)lj Kumaneshiri I!Jl! j;J~ i

Hokkaido. Ibid., vol. 6, pt. 2, p. 13-37, TAKEDA, H., 1954, Geological interpretation pis. 1-2, figs. la-b. of the Rumoi oil field (in Japanese with -- 194lc, On the three Tertiary non-ma­ English abstract). jour. japan. Assoc. rine shells illustrated inK. JIMBO's "Hok­ Petrol. Tech., vol. 19, no. 5, p. 155-163,. kaido Tisitu Ryakuron (Geological sketch ftgs. 1-9. of Hokkaido), 1890 ". Trans. Pro c. ?alae- TANAI, T., 1956, On the geologic age of the ant. Soc. japan, art. 138, p. 520-525, figs. Owada and Kabato coal-bearing forma­ 1-6. tions in Hokkaido, Japan (in japanese -- 1942, The Tertiary naids of Hokkaido with English abstract). Bull. Geol. Surv. in the collection of the Department of japan, vo!. 7, no. l, p. 11-20, figs. 1-3,. Geology and Mineralogy, Hokkaido Im­ table l. perial University. japanese jour. Geol. TsusHIJ\IA, K. and YAMAGUCI.II, s:, 1954, Ex­ Geogr., val. 18, no. 4, p. 141-156, pis. planatory text of the geological map of 18-19. Japan, Scale 1: 50,000, Rumoi (in japa­ -- 1944, Notes on some Tertiary non-ma­ nese with English abstract). Ceo!. Surv. rine Mollusca from North Nippon (in japan, 16 pp. japanese with English abstract). Trans. UEJI, T., 1934, Fresh-water fos::;il molluscs Proc. Palaeont. Soc. japan, art. 181, p. from Kitamatsuura coal-field, Northern 100-109, pis. 50-51. Kyushu, Japan (in japanese with English -- 1949, Development of the fossil non­ abstract). Venus, vol. 4, no.· 6, p. 341- marine molluscan faunas in Eastern Asia. 350, pls. 5-7. japanese jour. Geol. Geogr., vol. 21, nos. YAMAC>~E, S., 1912a, Report of the geology 1-'1, p. 91-133, tabs. 1-23. of Rumoi and Tomamae areas, Teshio· -- and OYAMA, K., 1948, Fresh-water shells Province (in japanese). Min. Surv. from the coal bearing Tertiary formations Rep., Geol. Surv. japan, no. 10, p. 1-84, in Mie-ken, Japan (in japanese with Eng­ pis. 1-2, figs. 1-2, tabs. 1-5. lish abstract). Venus, val. 15, nos. 1-4, -- 1912b, Report on the geology of Rumoi p. 36-44, figs. 1-5. and Tomamae areas, Teshio Province (in T AKAI, F., 1950, A1nynodon watanabei from Japanese). Ibid., no. 10, p. 85-122, pls. the Latest Eocene of Japan with a brief 1-3, figs. 1-2. summary of the Latest Eocene Mam­ YoKOYAMA, M., 1932, Tertiary Mollusca from malian faunule in Eastern Asia. Rep. the coal-field of Uryu, Ishikari. jour. G2o!. Surv. japan, no. 131, p. 1-14, pl. Fac. Sci., Imp. Univ., Tokyo, Sec. 2, vol. 1, tabs. 1-2. 3, pt. 6, p. 221-247, pls. 1-4.

Explanation of Plate 25

(All figures in natural size)

Figs. 1, 2, 4, 5, 7, 13. Unio uryuensis SuzuKI, p. 238, Loc. no. 1, Owada Formation, TGPS coil. cat. no. 86890. Fig;;. 3a-c. Margaritijera owadaensis NooA, n. sp., p. 240, Loc. no. l, Holotype, 3a: right valve, 3b: left valve, 3c: umbilical view, Owada Formation, IGPS col!. cat. no. 86893. Fig. 6. Viviparus cf. uryuensis YoKOYAMA, p. 240, Loc. no. 1, Owada Formation, IGPS coil. cat. no. 86889. Figs. 8a-b, 9, 11, 12. Margaritijera perdahurica YoKOYAMA, p. 239-240, Loc. no. 1, Owada Formation, IGPS coil. cat. no. 8(;)892. Fig. 10. Lanceolaria piscijormis (YoKOYAMA), p. 238-239, Loc. no. 1, Owada Formation, IGPS coil. cat. no. 86891. (All specimens are preserved in the Institute of Geology and Paleontology, Faculty of Science, Tohoku Univenity, Sendai, Japan) NODA : FTeshwate1· Molluscs /Tom Rumoi Plate 25

3a

1

3b

Sa 6

9

10

KUMAGAI photo. Trans. Proc. Palaeont. Soc. Japan, N.S., No . 77, pp. 243- 248, pl. 26, April 10, 1. 970

564. HALOBIA STYRIACA, UPPER TRIASSIC PELECYPOD, DISCOVERED IN OKIN A W A-JIMA, THE RYUKYU ISLANDS*

TEIICHI KOBAYASHI

Univers ity of Tokyo

and

T AKASHI ISHIBASHI

Department of Geology, Kyusbu Univers ity

ff,iJ;J (i'i'K:lllil<::lj~I:J-~""Th:r:c r.):(.)] Halobia styriaca O)i)EJil_ : (_jji(i;n:l 'i' K:1ffiUh7l\:l11\ ~ l = ,l i\l;-c'·Th .l'i!. L t::1 t T:i 0) ? 1? 1:: H alobia styriaca ;9: ;Jo -:d:. o Styriaca group 0)" P 1::' 7 v'i tJ - .::. " {I F>l!rt;\'i!['"(· iJb IJ , H alobia styriacav'i l"lf.'tliT l'.·l\1:: t~H'I!d i '·JfJ:{'l)-c· ;Jo ~ 0)'"(' , ff,if.j( O) styriaca /[;i\~

ffi '! -c· 111; -? --c v, t:. ~r nq'IJ -? t.::. 0 +~*t1- · T:i1;ri; ~

It was some years ago that new fossil Upper Triassic fossil beds in Okinawa localities were discovered in the Nakijin is indeed an important addition to the Formation by the junior author (1969) at stratigraphic column of the Ryukyu is­ Nakijin and Kamimotobu villages, Mo­ lands. tobu peninsula, Okinawa-jima and a small Setting aside obscure occurrences in lot of fossils was submitted the senior a few places of the Pacific province, the author through Dr. KONISHI for identifi­ distribution of the styriaca group has so cation. As the preliminary observation far been restricted to the region from has already been reported in the 94th the Alps to Indonesia. Therefore the meeting of the Palaeontological Society Okinawa styriaca indicates the eastern of japan at Akita, 1966, it included Ha­ limit of the distribution of this Tethys lobia styriaca and a few other pelecypods. species. The styriaca horizon is definitely Car­ Since that time the junior author nic and most probably lower Carnic in found additional new localities and great­ age, because the known occurrences of ly amplified the collection of pelecypods, the Halo bia styriaca group are exclusive­ ammonoids and other fossils by repeated ly in the Carnic stage and Ha lobia sty­ explorations so much that it requires riaca as a species is typical of the lower more time to be worked out. Therefore Carnic fauna. The discovery of such an only Halobia styriaca is described here as an advanced report. *Received Aug. 29, 1969; read Nov. 29, Here the authors record their best 1969 at Kagosbima. thanks to Professor Kenji KoNISHI of

243 244 Teiichi KOBAYASHI and Takashi ISHIBASHI the Kanazawa University for his cour­ is written here with reference to not tesy and assistance which they received only KrTTL's (1912), but also later publi­ in connection with this study. cations. (See DIENER, 1923, KUTASSY, 1930, KOBAYASHI and MASA T ANI, 1968). All of them were described from the Family Halobiidae KITTL Carnic stage of the Alps and the Medi­ terranean region or Southeast Europe Genus Halobia BRONN except for the isolated occurrences of The Halobia styriaca Group H. styriaca in Indonesia (VOLZ, 1899, KRUMBECK, 1921, 24, WANNER, 1931). KITTL (1912) classified Halobia species Halobia cf. styriaca and H. aff. styriaca into 11 groups. The styriaca group is are known to occur respectively at Basi­ one of them having the round or oval liaca, Italy and the Himalaya (DIENER, outline of the shell, scarcely prosogyral 1908). Of H. cf. styriaca from the Ma­ umbo, simple fiat anterior ear, indistinct layan frontier of Thailand (KOBAYASHI or flattened posterior triangular field and and TOKUYAMA, 1959), CHEN (1964) iden­ radial ribs broad, simple generally and tified it with Halobia substyriaca nov. sometimes once bifurcated, but twice bi­ from the Carnic of Western Szechuan. furcation is rare. He referred the fol­ Halobia styriaca has once been report­ lowing 10 species to the styriaca group : ed by PrROUTET (1908) from the isle of Sonde, New Caledonia in association Halobia areata KITTL: North Alps, Car­ with Halobia kwaluana in the basal part nic. of the Upper Triassic formation, but no Halobia arthaberi KrTTL: North Alps, palaeontological work has since been upper (?) Carnic. published of the species. ARTHABER Halobia beyrichi (Mo;s.): North Alps, Carnic. suggested the possible occurrence of the same species in Mexico, but this sug­ Halobia bosniaca KrTTL: Dinarid, Car­ nic. gestion has presumably been derived from the confusion of the species with Halobia landlensis KrTTL: North Alps, Carnic. H. austriaca (FRECH, 1907, KITTL, 1912). Halobia (?) lenticularis (GEMM.) : Italy KITTL (1912, p. 94) quoted that "Die (Basilicata and Sicily), North Alps (?), Art (H. styriaca) ist geradezu ein Leit­ lower Carnic. fossil fUr die unterkarnischen Schich­ ten." In North America Halobia artha­ Halobia (?) lepsiusi (GEMM.): North Alps, Carnic. beri and H. aff. lepsiusi were reported Halobia marmorea KrTTL: North Alps, to occur in western Canada in the low­ lower Carnic. er Noric stage (TOZER, 1961, McLEARN, Halobia (?) .richthofeni (Mo;s.): South 1960), but a further information is need­ Alps, lower Carnic 1St. Cassian). ed to confirm their specific identification. Halobia styriaca (MOJS.): North Alps, Hungary, Roumania (Dobrudscha), Di­ Halobia styriaca (Mo;srsovrcs) narid (Bosnia, Dalmatia), Greece, Sici­ ly, Indonesia (Timor, North Sumatra), Pl. 26, figs. 1-10, text-figs. a, b. lower Carnic. 1874. Daonella styriaca MoJS. Abhandl. k. k. The distribution of the above species geol. R.-Anst., Bd. 7, p. 10, pl. 1, figs. 564. Halobia from Okinawa-Jima 245

4-5. obscure in the umbonal part where the 1899. Daonella styriaca VoLz, Zeitschr. deut­ concentric sculptures are strong. Some sch. geol. Gesell., Bd. 51, p. 27, pl. 1, 15 to 20 ribs are countable in the medi­ fig. 1. an part of the shell in the middle stage 1906. Daonella styriaca RE;\;Z, Neues ]ahrb. of growth. They are very broad, fiat, f. Min. etc., 1906, I, p. 30, pl. 3, figs. 1-2, non 3. straight and separated from one another 1907. Daonella styriaca WANNER, N eues by narrow grooves. Some posterior ribs ]ahrb. f. Min. etc., Beil. Bd. 24, p. 194, are, however, slightly arcuate with back­ pl. 9, fig. 3. ward convexity. These ribs are partly 1912. Halobia styriaca KITTL, Result, wiss. bifurcated and very rarely even twice Erforsch. Balaton-Sees, 1 Bd., 1 Th. bifurcated by insertion of similar nar­ Pal. Bd. 2, p. 9, pl. 6, figs. 3-7. row grooves. 1924. Halobia styriaca KRUMBECK, Palaeont. Figure 2 and 1 are respectively an von Timor, Lief. 13, p. 132, pl. 9, fig. external mould of a left valve and its 8, pl. 10, figs. 1-6. rubber cast respectively. It measures 1925. Halobia styriaca DIENER, Leitfossilien, 19 mm in height and its outline is 4 Lief. p. 27, text-fig. 5. thought less deformed than other spe­ 1930. Halobia styriaca KuTASSY, Foeldtani Koezleony, Bd. 60, p. 205, pl. 3, fig. 2. cimens. The shell is nearly as long as high; umbo almost median and only a None of the specimens before hand little projected above the hinge margin :shows a complete outline of the shell, which is straight and horizontal in front but evidently it is fairly tall and nearly of the umbo, but the margin is gradu­ -equilateral. The umbo is located sub­ ally descending behind the umbo and medially, but a little anteriorly, moder­ more distinctly slant than in other spe­ ately convex, slightly projected above cimens. The other margins are well the hinge margin and apparently short­ rounded. The shell is gently inflated -er than the shell-length; The propor­ and the convexity is strengthened in tional height to the length is not exact­ the umbonal region where some 10 ly determinable, because all specimens concentric folds are regularly disposed. are deformed. The anterior ear is simple, depressed The anterior ear is very fiat and and fiat, but somewhat thickened at the ·smooth, but it is not clearly separated hinge margin. The preumbonal angle from the inflated main part of the shell. of the ear is about 20 degrees. Therefore it appears very indistinct, The Posidonia or Bositra stage trans­ when the shell is flattened. The pos­ mits into the Halobia stage at the height terior triangular area is narrower than of about 7 mm. Radial ribs are broad, the ear and ill-defined. There radial fiattopped and separated from one an­ sculptures are obscure or absent. other by narrow grooves, some of which Some concentric grooves or folds are are bifurcated by insertion of a groove. well marked in the umbonal one-third Anterior ribs are straight, while pos­ -or one-fourth of the shell which is more terior ones are gently arcuate with inflated than the remaining part. They backward convexity. About 20 ribs are .are generally stronger on the posterior countable near the ventral margin, but lateral side. Additional concentric folds on the lateral sides of the shell the tropoden der HMO : Motobu high school, Motobu­ Trias von Timor, 2. Palaeo1i to i." von T i­ cho, Okinawa. mor, Lief. 13. HNa-P : 200m west of the ruin of the K uTASSY, A. (1928): Die Ausbildung der 248 Teiichi KOBAYASHI and Takashi ISHIBASHI

Trias im Morna Gebirge (Ungaren-Sieben­ REED, F.R.C. (1927): Palaeozoic and Meso­ biirgen). Centralbl. f. Min. etc. ]ahrg. zoic Fossils from Yunnan. Pal. Indica, 1928, Abt. B, No. 5. N.S., Vol. 10, No. 1. -- (1930) : Triadische Fossilien von Portu­ RENZ, C. (1906) : Ueber Halobien und Dao­ _giesischen Timor. Foezldtani Koezloeny, nellen aus Griechenland nebst asiatischen Bd. 60. Vergleichstucken. Neues ]ahrb. f. Min. -- (1931) : Lamellibranchiata triadica, 2. etc. jahrg., 1906, Bd. 1. Fossilium Catalogus, 1, Animalia, Pars TozER, E.T. (1961): The Sequence of Ma­ 51. rine Triassic Fauna in Western Canada. McLEARN, F.H. (1960) : Ammonoid Faunas Geol. Surv. Canada, Paper 61-6. of the Upper Triassic Pardonat Forma­ VoLz, W. (1899): Beitrage zur geologischen tion, Peace River Foothills, British Co­ Kenntniss von Nord-Sumatra. Zeitschr. lumbia. Geol. Surv. Canada, Mem. 31. deutsch. geol. Gesell., Bd. 51. MoJsrsovrcs, E.V. (1874): Ueber die Pele­ WANNER, ]. (1907) : Triaspetrefakten der cypodengattungen Daonella und Halobia. Mollukken und des Timorarchipels. Neues Abhandl. geol. Reichesanst. 7 (2). ]ahrb. f. Min. Bd. 24. PrROUTET, M. (1908) : Note sommaire sur -- (1931) : Mesozoikum. Leidsch geologi le Trias de Ia Nouvelle Caledonie. Bull. sche Medd., Dee! 5. Soc. geol. France, 4 Ser., t. 8.

Explanation of Plate 26

Halobia styriaca (MoJsrsovrcs) from Okinawa, Ryukyu Islands Loc. HNa-P: 200m west of ruin of Nakijin castle, Nakijin-son, Okinawa. Loc. HMO: Motobu high school, Motobu-cho, Okinawa. Fig. 1. . Rubber cast of a left valve from HNa-P. x 2 Fig. 2. External mould of the same valve as the preceding. x 3 Fig. 3. A left valve cut by a vein; HNa-P. x 1! Fig. 4. Two deformed valves disposed almost rectangularly; HNa-P. x 1! Fig. 5. Internal mould of a left valve laterally compressed: HNa-P. x 1! Fig. 6. Rubber cast of a left valve diagnonally compressed; HNa-P. x 1! Fig. 7. Rubber cast of a left (?) valve laterally compressed; HNa-P. x 1! Fig. 8. Rubber cast of a right valve laterally compressed; HMO. x 2 Fig. 9. Rubber cast of a right valve; HMO. x 1! Fig. 10. Rubber cast of a left (?) valve; HMO. x 2. KoBAYASHI and ISHIBASHI: Halobia styriaca Plate 26 249

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY OF JAPAN

B:;$:[l)~1tJ'¥:S<1lf~ 103 @f9US "L 11ilJ1'.~:*~7: !l!l~tJIJ:fu~f:~~ ':!:: Sjs:i!Ji!i> JltiJll.,~jm 3'z:ftt Y !l-1::: ~!IS•i ~ :hJ::.;j,JII~iSE\: On the distribution of Dictyozamites and qJl~0)7 Y-'E-7-1 r ...... tL;:$:i~Ii!rs • :;~c~r&z its palaeobotanical significance ...... Permian brachiopods from Khao Phrik, ...... KIMURA, T. Thailand (ftjj/C) ...... YANAGIDA, J. lli~1'ffL!hllf~O):fffiMfl rJ;~tf-OJ~Hifll~- ~<'n: -9 :lj. n1 ® ~ t.tmitl i!IA, wrr ut 't'f !lt 1Ji ...... ;<£ 7M.n -r bft · C. tf:lj.JJ!.~OJfiiHJTIC."?v'"L .. ;Nf\tf:£ Addition to the Permian Bryozoa from Ahermatypic corals from the Pleistocene Komuk, Peninsular Thailand ...... Ninomiya Formation in Kanagawa Pre- ...... SAKAGAMI, s. fecture, central Japan ...... HAMADA, T. iti1i.liil i1Jt,~*£j J~M\Hi)](£0) ::1 ; r· Y r O):i:i!l'f!t Discovery of Calceola from the Fukuji Jl~.fttO)~fii·. · · · · · · · · · · · -~..l:.~Jt:t:: · i¥JJII~:iJI1:: Series, Gifu Prefecture, Japan ...... Elaphurus shikamai OTSUKA O)!*.=OJt)'!(:;$:1::: ...... HAMADA, T. "?v'--c ...... ·*~:faz 4f'Jv ;f-. r; ~Ji OJ 1i!z: lH lliJ:$:~ ...... :5 Mi 31Hli Fossil Mollusca from Teshima, Shodo-gun, :/Y;f-~rj.b. ljL)I-!-[O)!*fi9:iF-J fii3:;$:J:t!l'f!t'¥:Si- Molluscan fauna of the Kukinaga Group in lf~~~e~~d.mJm~~~~::J70)~•M~ Tane-ga-shima, South Kyushu, Japan .. ~:IYlliJfJr:-c. ' 0 7-.IC."?v'"L ...... ·IIlftlEZ: ...... :;lc;;t~r,?; . .'f'-~1'¥':::::. A review of some Cretaceous corbiculids YML'l+JO) :/7 ;;<. IC. "?v'L · .'fifi-1-EE]EIJ)J · '§'J:i!!:t\~ in North America ...... OTA, Y. .7L"l·I·JO)JlP.;fi9*c'tffff1lih¥!J~f ...... :;~c~r&z The hinge structure of Trigonoides, with i¥fl!§f.'fi,l6j~II9:1F-O)Ji!z:M~tt~ ...... :*ttaAMi; • ,J,®WJ\.= description of Trigonoides mifunensis sp. nov., from upper Cretaceous Mifune H.~lml~O)/li'f!t C.1'f1L!TI ...... £\:~ 7'.i Group, Kumamoto Prefecture, Japan .. .7L'l+I®:1J~l:9.X.OJ1wnX:Jl'f.l'l1i!JJf10)~1i!JI.H~ll1~ ...... TAMURA, M. · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · ·lUi:EE3fi'r,?; ~ J':!i Jf11:.t\:O) "i!fi1fft'l" .=tx:fUC."?v'"L .... )Ifll]( t% 7P->' r 0-0)j!J:t!liPi?:lj.f::. 2 ~~~'f!tO)~~ ...... ~rll~nA0 · Jili7Jc 1% 12 R 1 13 (R) IC. fi, :!:i!!'f!tilli~ IJJ!!JC,l6ji~itft!l.ml Halobia styriaca (MOJSISOVICs), upper Tri­ illOJ~II9:1F-J ;0;-Wfr~~t::., c~i*lif e~;;tfiJ~ • assic pelecypods, discovered in Okinawa- .'f'-~1'¥:::::. • *~r&z · **~0) 250

a*~~~~~ ~ro ~~~ · A~~~~ . ~ro On the Spongoplegma antarcticum HAEC KEL. il-'- 1 FJ 19 E1 (FJ) · 20 o C*) O) jiijjEJ, WJd~J!X: "' ...... N AKASE KO, K. t:::. fili i:~ rn 1'!· * w0) JR;It:.* "J!: JlH"Ji:ifll Jilin ~f: · 1'1 ~:. i ''J $Jiif.\ .'S~ ;!I::. fi!IO):.i:Ji"1Ji·\ :=: if,Ji.l£1 'iX1lH:. Ei l:. -::>v'-c · · · · ~~~ -C 1'Tbht..: (~1Jn:tf 55 1:",) o ...... ~~ ~*~~ · * W V- Radiolarian fossils from the Oidawara for- mation in Gifu Prefecture, Japan ...... l EDA , K. North American Paleontological Convention, An interesting s pecies of Ca tania from the Rat Buri Limestone, central Thailand .. 1969 ...... - ...... JJ!i llnifij•;J;i: . t!li i :J:t. !fJ!£ :ff~ ...... OzA- w A, T ., K ANME RA, K. & T oR IYA MA, R. Recent and fossil Rapana from the Pacific coast of North America ...... K ANNO, S. Note on the molluscan assemblage in Usuki Bay, Kyushu ...... SH UTO, T. Lower Triassic ammonoids from the Kita- kami Massif ...... BAN DO, Y · Evolution and classification of Sphenophyi­ Uncommon keeled ammonites from the up­ lales in upper Paleozoic Cathaysia Flora per Cretaceous of Hokkaido -and Sagha- ...... ASAMA, K. lien ...... M ATSUMOTO, T. Some palynomorphs from the upper Creta- Early Devonian brachiopods from the Lesser ceous sediments of Hokkaido ...... Khingan di strict of Northeast China ...... T AKAHAS HI, K ...... H AMA DA, T . Evolutionary trend of Acer in East Asia Occurrence of marine reptile bones from and North America ...... T ANA I, T. the upper Cretaceous Futaba Group . ... Miocene coccolithophorids from Noto, Japan ...... 0 BATA, I. & HASEGAWA, Y ...... N ISHID A, s. A new elasmosaur from the upper Cretace- Nannofossils from the Nohori and Ananai ous Futaba Group ...... formations, Shikoku, Japan .. N iSH IDA, S...... H ASE GAWA , Y. & OBATA, I. *•~~m~~0)~~~70)&a~~~~~• 7 1 "/ :/ a :/ • r 7 "/ ~ li~ 1:. J:: 6 JJ!.~ :/ •\' ~ J~t 3<~1Wi f} ~ 0) 1'1J1iH::. -::> v' -c ( -=J-~fl) · · .. {til- *~II v'"( .. · · · · · · · · · · · ·. · · Miocene Foraminifera from the Sandakan ...... -...... -... * t"tAJl t(jf~ · 'L ···· · ··· · ··· blages in the Southeast Indian Ocean ...... l:jl ill {!§= • "'i1§ fill= . ~~i ;):j: :It;] ...... N AGASE , K. & UJIIE, H. On some species of Thecosphaera from the liMiAIHl? r 1t T:i ''I=•0) ITrl fli {:;j;: J li 1 Fl 20 1=1LH!i: 2 ·Neogene formations, Japan ...... A

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NEWS

O :;js;: -'9- 0){>H ff< c L -c , Index of Transactions of the Palaeontological Society of Japan, Article No. 1 to No. 515 ~~ff L , ~~ IC ili ~:ffiT ~ .::. c ct.~: -:> t.::o 0 t;),JJ!I -'9- No. 14, The Litho- and Bio-facies of Carbonate Sedimentary Rocks, A Symposium iJ> 1969 {)~ 11 F1 25 13 1 CThff~tct.:: o 5EW!i 2000 p:j, ~~~'J: 1 fj:.O)I 'M I 1700 p:j o 0 11:11 18 1o} iJ> , 1969 "'~'- 10 Ji 1c Thff~ tct.::o Ji ~0 0 ~ 24 @1 7J !;illt l!l~ $:~~1H'J: , 1972 11~ 8 f'l 21 ~30 13 Canada, Montreal -c·F,rJ(f!i ~ tc ~ o iili*~:?ti'J: , Secretary-General, 24th International Geological Congress, 601 Booth Street, Ottawa 4, Ontario, Canada. 0 :=.~ ntB'I J: ~ UB;fQ 45 i.f.!J£ 13 YM.ii ~~!Ufj't;ll)JJlX.~)JIHc-? '-' ' -c !H!~iJ ; t.:: o 1 1'1= 3 'f7Jp:j.CJrkJ, *':J 15 1'1' ~f*ffi.o HJJ fi~ I'J:!JU IJc L-c 1 fJ'.o J:t:.: ~:j;tt';~tfi'J: , *trf1'lii'TttEEIIR;I:L/ i*J 2-6-2o :;1.1/ i*Jl\.mi~~C:!ld09 -'%~ . ME'fl f.t; A :=.~ M BJ&.> -c, f.G:;tn~r.i'i.toJ:tf.fflK;R~~Jlf;J<~*of.:: v ' o Jt~fl\rlitrJI'J: , f!B ;fll45 :if. 5 Ji

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1970 {j:'-4FJ 5 13 s J6 fi ~ B '!5 ~ ¥.J "f- ~ 1 9 7 0 :if. 4 Fl 10 13 ***$:fJII#~mtl!l'Iil#~~* 1*1 ~jjj ~ :?3 Jll Fs~ ~ X. 13 :;js;: r!J1:. ~J.J ¥:~¥11 ~ · *c ~J r ;<,g WJrATI'l1f,7 7 % (th'€ 1-l ~ }!~ * 8 4 7 8 0 i!f) k",li ;It 2 ; 13 700 p:j r:!J .lllO :?3 * trr t~ .~ 1R :llli: :r.: #vi~~~~Pll\IJ¥~5t~t± ~ EEl 7G Transactions and Proceedings of the Palaeontological Society of japan

New Series No. 77 April 10, 1970

CONTENTS

TRANSACTIONS 561. IwASAKI, Yasuhide: A Miocene molluscan fauna in the Philippines ...... 205

562. HuziOKA, Kazuo: A new species of Sagenopte1'is from Nariwa, Southwest Honshu, Japan ...... 229

563. NonA, Hiroshi: Freshwater molluscs from the coal-bearing Owada forma- tion, Southeast Rumoi, Hokkaido, Japan ...... 235

564. KOBAYASHI, Teiichi and ISHIBASHI, Takashi: Halobia styriaca, Upper Tri- assic pelecypod, discovered in Okinawa-Jima, the Ryukyu Islands ...... 243

PROCEEDINGS ...... 249