Establishment of a Lectotype for the Species Plesiochelys Langii Rutimeyer
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A Phylogenomic Analysis of Turtles ⇑ Nicholas G
Molecular Phylogenetics and Evolution 83 (2015) 250–257 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A phylogenomic analysis of turtles ⇑ Nicholas G. Crawford a,b,1, James F. Parham c, ,1, Anna B. Sellas a, Brant C. Faircloth d, Travis C. Glenn e, Theodore J. Papenfuss f, James B. Henderson a, Madison H. Hansen a,g, W. Brian Simison a a Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA b Department of Genetics, University of Pennsylvania, Philadelphia, PA 19104, USA c John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA d Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA e Department of Environmental Health Science, University of Georgia, Athens, GA 30602, USA f Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA g Mathematical and Computational Biology Department, Harvey Mudd College, 301 Platt Boulevard, Claremont, CA 9171, USA article info abstract Article history: Molecular analyses of turtle relationships have overturned prevailing morphological hypotheses and Received 11 July 2014 prompted the development of a new taxonomy. Here we provide the first genome-scale analysis of turtle Revised 16 October 2014 phylogeny. We sequenced 2381 ultraconserved element (UCE) loci representing a total of 1,718,154 bp of Accepted 28 October 2014 aligned sequence. Our sampling includes 32 turtle taxa representing all 14 recognized turtle families and Available online 4 November 2014 an additional six outgroups. Maximum likelihood, Bayesian, and species tree methods produce a single resolved phylogeny. -
Universidad Nacional Del Comahue Centro Regional Universitario Bariloche
Universidad Nacional del Comahue Centro Regional Universitario Bariloche Título de la Tesis Microanatomía y osteohistología del caparazón de los Testudinata del Mesozoico y Cenozoico de Argentina: Aspectos sistemáticos y paleoecológicos implicados Trabajo de Tesis para optar al Título de Doctor en Biología Tesista: Lic. en Ciencias Biológicas Juan Marcos Jannello Director: Dr. Ignacio A. Cerda Co-director: Dr. Marcelo S. de la Fuente 2018 Tesis Doctoral UNCo J. Marcos Jannello 2018 Resumen Las inusuales estructuras óseas observadas entre los vertebrados, como el cuello largo de la jirafa o el cráneo en forma de T del tiburón martillo, han interesado a los científicos desde hace mucho tiempo. Uno de estos casos es el clado Testudinata el cual representa uno de los grupos más fascinantes y enigmáticos conocidos entre de los amniotas. Su inconfundible plan corporal, que ha persistido desde el Triásico tardío hasta la actualidad, se caracteriza por la presencia del caparazón, el cual encierra a las cinturas, tanto pectoral como pélvica, dentro de la caja torácica desarrollada. Esta estructura les ha permitido a las tortugas adaptarse con éxito a diversos ambientes (por ejemplo, terrestres, acuáticos continentales, marinos costeros e incluso marinos pelágicos). Su capacidad para habitar diferentes nichos ecológicos, su importante diversidad taxonómica y su plan corporal particular hacen de los Testudinata un modelo de estudio muy atrayente dentro de los vertebrados. Una disciplina que ha demostrado ser una herramienta muy importante para abordar varios temas relacionados al caparazón de las tortugas, es la paleohistología. Esta disciplina se ha involucrado en temas diversos tales como el origen del caparazón, el origen del desarrollo y mantenimiento de la ornamentación, la paleoecología y la sistemática. -
Comparative Bone Histology of the Turtle Shell (Carapace and Plastron)
Comparative bone histology of the turtle shell (carapace and plastron): implications for turtle systematics, functional morphology and turtle origins Dissertation zur Erlangung des Doktorgrades (Dr. rer. nat.) der Mathematisch-Naturwissenschaftlichen Fakultät der Rheinischen Friedrich-Wilhelms-Universität zu Bonn Vorgelegt von Dipl. Geol. Torsten Michael Scheyer aus Mannheim-Neckarau Bonn, 2007 Angefertigt mit Genehmigung der Mathematisch-Naturwissenschaftlichen Fakultät der Rheinischen Friedrich-Wilhelms-Universität Bonn 1 Referent: PD Dr. P. Martin Sander 2 Referent: Prof. Dr. Thomas Martin Tag der Promotion: 14. August 2007 Diese Dissertation ist 2007 auf dem Hochschulschriftenserver der ULB Bonn http://hss.ulb.uni-bonn.de/diss_online elektronisch publiziert. Rheinische Friedrich-Wilhelms-Universität Bonn, Januar 2007 Institut für Paläontologie Nussallee 8 53115 Bonn Dipl.-Geol. Torsten M. Scheyer Erklärung Hiermit erkläre ich an Eides statt, dass ich für meine Promotion keine anderen als die angegebenen Hilfsmittel benutzt habe, und dass die inhaltlich und wörtlich aus anderen Werken entnommenen Stellen und Zitate als solche gekennzeichnet sind. Torsten Scheyer Zusammenfassung—Die Knochenhistologie von Schildkrötenpanzern liefert wertvolle Ergebnisse zur Osteoderm- und Panzergenese, zur Rekonstruktion von fossilen Weichgeweben, zu phylogenetischen Hypothesen und zu funktionellen Aspekten des Schildkrötenpanzers, wobei Carapax und das Plastron generell ähnliche Ergebnisse zeigen. Neben intrinsischen, physiologischen Faktoren wird die -
THE GENERA of REPTILES. By
T he Genera of Re pt il e s. By BARON FRANCIS NOPCSA (Budapest). (Eingelangt am 11. Mai 1927.) Among all the Paleontologists living none has dealt with the recent and fossil reptiles with a wider grasp than Prof. L. D o l l o and at his anniversary it seems quite appropriate to review their whole array. The classification used in the following enumeration of all genera of reptiles is much the same as in my hook „Die Familien der Reptilien“; at this instance however an effort has been made to give a precise definition of every systematic unit. Alteration of the classification became necessary among the Dino- cephalians, the Nothosaurians, the Lacertilians (called here Sauroidea), the Coelurosauroidea and the Crocodilia. The new classification of the Sauroidea is intermediate between the classifications proposed by B o u l e n g e r and C a m p . Naturally as a rule only those fossil genera are referred to, that are more than simple catalogue numbers that facilitate the finding of the respective piece in a collection; indeterminable problematic genera of small interest have mostly been omitted. Extinct units and genera are marked in the lists with a cross ("j*). Difficulties of classification have been encountered in the Lacertilia and Ophidia, for in the new system the conventional families of recent Lizards and Serpents has been given subfamily rank and conse quently the different recent subfamilies had to be dropped. Instead of these subfamilies the minor units were separated by greek letters. — The tedious and complicated revision of the genera of recent Lizards and Snakes was done by Prof. -
The Girdles of the Oldest Fossil Turtle, Proterochersis Robusta, and the Age of the Turtle Crown Walter G Joyce1,2*, Rainer R Schoch3 and Tyler R Lyson4
Joyce et al. BMC Evolutionary Biology 2013, 13:266 http://www.biomedcentral.com/1471-2148/13/266 RESEARCH ARTICLE Open Access The girdles of the oldest fossil turtle, Proterochersis robusta, and the age of the turtle crown Walter G Joyce1,2*, Rainer R Schoch3 and Tyler R Lyson4 Abstract Background: Proterochersis robusta from the Late Triassic (Middle Norian) of Germany is the oldest known fossil turtle (i.e. amniote with a fully formed turtle shell), but little is known about its anatomy. A newly prepared, historic specimen provides novel insights into the morphology of the girdles and vertebral column of this taxon and the opportunity to reassess its phylogenetic position. Results: The anatomy of the pectoral girdle of P. robusta is similar to that of other primitive turtles, including the Late Triassic (Carnian) Proganochelys quenstedti, in having a vertically oriented scapula, a large coracoid foramen, a short acromion process, and bony ridges that connect the acromion process with the dorsal process, glenoid, and coracoid, and by being able to rotate along a vertical axis. The pelvic elements are expanded distally and suturally attached to the shell, but in contrast to modern pleurodiran turtles the pelvis is associated with the sacral ribs. Conclusions: The primary homology of the character “sutured pelvis” is unproblematic between P. robusta and extant pleurodires. However, integration of all new observations into the most complete phylogenetic analysis that support the pleurodiran nature of P. robusta reveals that this taxon is more parsimoniously placed along the phylogenetic stem of crown Testudines. All current phylogenetic hypotheses therefore support the basal placement of this taxon, imply that the sutured pelvis of this taxon developed independently from that of pleurodires, and conclude that the age of the turtle crown is Middle Jurassic. -
A NEW LATE JURASSIC TURTLE from SPAIN: PHYLOGENETIC IMPLICATIONS, TAPHONOMY and PALAEOECOLOGY by BEN J
[Palaeontology, Vol. 54, Part 6, 2011, pp. 1393–1414] A NEW LATE JURASSIC TURTLE FROM SPAIN: PHYLOGENETIC IMPLICATIONS, TAPHONOMY AND PALAEOECOLOGY by BEN J. SLATER1, MATI´AS REOLID2, REMMERT SCHOUTEN3 and MICHAEL J. BENTON3 1School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, UK; e-mail: [email protected] 2Departmento de Geologı´a, Universidad de Jae´n, Campus Las Lagunillas sn, 23071 Jae´n, Spain; e-mail: [email protected] 3School of Earth Sciences, University of Bristol, Wills Memorial Building, Queen’s Road, Bristol BS8 1RJ, UK; e-mails: [email protected], [email protected] Typescript received 26 October 2010; accepted in revised form 27 May 2011 Abstract: The Jurassic was an important period in the evo- of the new taxon is hard to resolve, and it might be either a lution of Testudinata and encompasses the origin of many paracryptodire or a basal testudine, but it is distinct from clades, and this is especially true of Jurassic turtles from Wes- Plesiochelys. A complex taphonomic history is shown by a tern Europe. A new genus and species of Late Jurassic turtle, range of overlying grazing traces and bioerosion on the cara- Hispaniachelys prebetica gen. et sp. nov. from the upper Ox- pace. The carapace was subsequently overturned and buried fordian of the Prebetic (Southern Spain), is described on the ventrally up, terminating grazing activity, and was then bored basis of postcranial material. The specimen is the only known by sponges before final burial. Scanning electron microscopy tetrapod from the Mesozoic of the Prebetic and the oldest reveals phosphatic microspheroids associated with bacterial turtle from southern Europe. -
Nelby Wlned González Rodríguez. Matricula
UNIVERSIDAD AUTÓNOMA METROPOLITANA UNIDAD IZTAPALAPA Nombre: Nelby Wlned González Rodríguez. Matricula: 96338704 Teléfono: 54 41 76 02 Licenciatura: Biología División: C. B. S. Unidad: Iztapalapa Trimestre Lectivo: TITULO DEL PROYECTO: “PROYECTO DE ACTIVIDADES DE PRESTADORES DE SERVICIO SOCIAL DE LA DIVISIÓN DE C. B. S. PARA LA EVALUACIÓN Y PROTECCIÓN DE TORTUGAS MARINAS MEXICANAS”. Campamento Tortuguero Isla de Pájaros con registro en SEMARNAT e INE TITULO DEL TRABAJO: Estudio del centro de origen y distribución de las tortugas marinas que arriban a las costas de Guerrero, México. Asesor Interno: M. en C. Francisco Javier Olvera Ramírez. Departamento de Biología. Edificio AS. Cubículo 105. U A M – Iztapalapa, Av. San Rafael Atlixco 186 Col. Vicentina México DF. Tel. 58 04 46 00.ext.2794 Lugar de Realización: Campamento Tortuguero “Isla de Pájaros”. Playa Michigan, Domicilio conocido Municipio de Tecpan de Galeana, Guerrero, México. Clave de Registro: B.071.03 México D. F., a 23 de febrero de 2006 Dr. José Fco. Flores Pedroche Secretario Académico de la División de Ciencias Biológicas y de la Salud . Universidad Autónoma Metropolitana Iztapalapa PRESENTE: Por este conducto me permito manifestarle que el alumno Nelby Wlned González Rodríguez con matrícula 96338704 estudiante de la carrera de Biología, inicio el servicio social a partir del 10 de noviembre del 2003 y lo concluyó en septiembre del 2004. El informe de servicio final no fue terminado hasta ahora debido a causas personales extra escolares relacionadas con actividades laborales. Después de revisar el informe titulado "Estudio del centro de origen y distribución de las Tortugas Marinas que arriban a las costas de Guerrero, México". -
Biologie Historique Et Évolutionnisme M. Armand De Ricqlès
Biologie historique et évolutionnisme M. Armand de Ricqlès, professeur Enseignement Cours : L’évolution secondaire des tétrapodes vers les milieux aquatiques. 4. Les diapsides (suite et fin) ; les anapsides Mercredi 6, 13, 20 et 27 janvier, 3, 10, 17 et 24 février 2010 (16 heures) Le cours a débuté par un résumé rapide des problèmes généraux de phylogénie des Squamates (lézards [Sauria], amphisbènes, et serpents [Ophidia]), déjà abordés l’année dernière. Compte tenu de la diversité anatomique et de la répartition stratigraphique des divers squamates fossiles du Jurassique et du Crétacé, les recherches ont abouti à des phylogénies contradictoires, en particulier en ce qui concerne l’origine et l’apparentement des serpents (voir Annuaire 2008-2009, pp. 433-436). On en a d’abord terminé, cette année, avec la présentation historique et descriptive des groupes pertinents, avant de revenir aux problèmes généraux de phylogénie, prenant cette fois en compte les données moléculaires. 1. Les Mosasaures Ils constituent un grand groupe de Squamates ayant vécu dans les mers du Crétacé supérieur, après la disparition des ichthyosaures. Il s’agit d’un groupe naturel (clade) très efficient dans son adaptation à des niches écologiques variées dans les mers épicontinentales et peut-être les océans. Son évolution est bien documentée, en particulier dans les phosphates sud-tethysiens (Maroc) et dans la « continental seaway » nord-américaine. La répartition du groupe a été mondiale, mais il demeure rare en Asie. La découverte des Mosasaures constitue historiquement un des épisodes fondateurs de la paléontologie des vertébrés. On a relaté en détail les circonstances pittoresques de ces découvertes, de 1766 à 1780, dans les carrières de Maestricht et les études ultérieures de Cuvier, aboutissant à la notion d’« espèces perdues » du 424 ARMAND DE RICQLÈS fait des « Révolutions du globe » (1808), bien qu’il ait noté les fortes affinités anatomiques du fossile avec les lézards actuels. -
Qt147611bv.Pdf
UC Berkeley PaleoBios Title Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia Permalink https://escholarship.org/uc/item/147611bv Journal PaleoBios, 32(1) ISSN 0031-0298 Authors Cadena, Edwin A Parham, James F Publication Date 2015-09-07 DOI 10.5070/P9321028615 Supplemental Material https://escholarship.org/uc/item/147611bv#supplemental Peer reviewed eScholarship.org Powered by the California Digital Library University of California PaleoBios 32:1–42, September 7, 2015 PaleoBios OFFICIAL PUBLICATION OF THE UNIVERSITY OF CALIFORNIA MUSEUM OF PALEONTOLOGY Edwin A. Cadena and James F. Parham (2015). Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia. Cover illustration: Desmatochelys padillai on an early Cretaceous beach. Reconstruction by artist Jorge Blanco, Argentina. Citation: Cadena, E.A. and J.F. Parham. 2015. Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia. PaleoBios 32. ucmp_paleobios_28615. PaleoBios 32:1–42, September 7, 2015 Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia EDWIN A. CADENA1, 2*AND JAMES F. PARHAM3 1 Centro de Investigaciones Paleontológicas, Villa de Leyva, Colombia; [email protected]. 2 Department of Paleoherpetology, Senckenberg Naturmuseum, 60325 Frankfurt am Main, Germany. 3John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA; [email protected]. Recent studies suggested that many fossil marine turtles might not be closely related to extant marine turtles (Che- lonioidea). The uncertainty surrounding the origin and phylogenetic position of fossil marine turtles impacts our understanding of turtle evolution and complicates our attempts to develop and justify fossil calibrations for molecular divergence dating. -
The Oldest Pre-Podocnemidid Turtle (Chelonii, Pleurodira), from the Early Cretaceous, Ceará State, Brasil, and Its Environment
Treb. Mus. Geol. Barcelona, 9: 43-95 (2000) The oldest pre-Podocnemidid turtle (Chelonii, Pleurodira), from the early Cretaceous, Ceará state, Brasil, and its environment France de LAPPARENT de BROIN* RESUMEN LAPPARENT de BROIN F. de, La tortuga pre-Podocnemidid más antigua (Chelonii, Pleurodira), Cretácico inferior, estado de Ceará, Brasil, y su entomb. Se describe una nueva forma de tortuga pleurodira, Brasilemys josai, n.g., n. sp. Su posición filética se examina en relación con las filogenias establecidas anterior- mente. Se sitda también Brasilemys n.g. en relación con algunas formas reciente- mente descritas y adn no bien posicionadas en Ia filogenia de los Pleurodira, como son los Dortokidae, Hamadachelys y otra forma de la Chapada do Araripe sin nombre. En los Podocnemidoidea, Brasilemys n.g. aparece como el primer repre- sentante de los Podocnemidoidae, inmediatamente después de la derivación de los Bothremydidae. La aparición de este nuevo genero forma parte de la extraordinaria radiación que ocurrió en los Pelomedusoides al inicio del Cretácico, cuando se sepa- raron America del Sur y Africa. La fauna de tortugas de la Chapada do Araripe incluye cuatro taxones de Pelomedusoides de agua dulce, representantes de la fauna de la parte norte del antiguo continente Gondwana, además de una tortuga crypto- dira, posiblemente litoral y perteneciente a un grupo más cosmopolita. De acuerdo con los datos de la estratigrafIa y con los del estudio de los cocodrilos fósiles, la edad de la fauna de tortugas se situarIa entre el Aptiense superior de Gadoufaoua (Niger) y el Albiense superior - base del Cenomaniense del Sahara. Palabras dave: Quelonios, Nuevo taxón, Rejaciones filéticas, Cretácico, Brash, Consideraciones paleoecológicas y estratigráficas. -
Chelonological Studies of L.I. Khosatzky with His Annotated Bibliography on Turtles
Proceedings of the Zoological Institute RAS Vol. 317, No. 4, 2013, рр. 382–425 УДК 568.13 + 598.13 + 57(09) CHELONOLOGICAL STUDIES OF L.I. KHOSATZKY WITH HIS ANNOTATED BIBLIOGRAPHY ON TURTLES I.G. Danilov1*, G.O. Cherepanov2 and N.S. Vitek3 1Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia; e-mail: [email protected] 2Department of Vertebrate Zoology, Faculty of Biology and Soil Sciences, Saint Petersburg State University, Universitetskaya Emb. 7/9, 199034 Saint Petersburg, Russia; e-mail: [email protected] 3Department of Biology, University of Florida, P.O. Box 118525, Gainesville, FL 32611, USA; e-mail: [email protected] ABSTRACT This paper presents a review of the chelonological research of L.I. Khosatzky (1913–1992), a well-known Soviet zoologist-herpetologist and paleontologist from Leningrad (now Saint Petersburg). It provides an annotated bibliography of his 115 publications on turtles. Khosatzky’s turtle research was devoted to the origin of turtles, systematics, phylogeny and biogeography of fossil and recent turtles, biomechanics, morphology and evolution of the turtle shell, and the ecological physiology of turtles. Khosatzky supervised numerous student works (course projects, graduate and candidate theses) on different aspects of turtle biology, the results of which were only partially published. Some of Khosatzky’s students became well-known specialists on turtles. Key words: biogeography, biomechanics, ecological physiology, evolution, history of science, morphology, origin, phylogeny, shell, systematics, turtles ХЕЛОНОЛОГИЧЕСКИЕ ИССЛЕДОВАНИЯ Л.И. ХОЗАЦКОГО С ЕГО АННОТИРОВАННОЙ БИБЛИОГРАФИЕЙ ПО ЧЕРЕПАХАМ И.Г. Данилов1*, Г.О. Черепанов2 и Н.С. Витек3 1Зоологический институт Российской академии наук, Университетская наб. -
In Search of the Tree of Life for Turtles
IVERSON ET AL. – Supertrees 85 Defining Turtle Diversity: Proceedings of a Workshop on Genetics, Ethics, and Taxonomy of Freshwater Turtles and Tortoises H. Bradley Shaffer, Nancy N. FitzSimmons, Arthur Georges, and Anders G.J. Rhodin, Eds. Chelonian Research Monographs 4:85–106 • © 2007 by Chelonian Research Foundation In Search of the Tree of Life for Turtles JOHN B. IVERSON1, RAFE M. BROWN2, THOMAS S. AKRE3, THOMAS J. NEAR4, MINH LE5, ROBERT C. THOMSON6, AND DAVID E. STARKEY7 1Department of Biology, Earlham College, Richmond, Indiana 47374 USA [[email protected]]; 2Natural History Museum and Biodiversity Research Center, Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence 66045 USA [[email protected]]; 3Department of Biological and Environmental Sciences, Longwood University, Farmville, Virginia 23909 USA [[email protected]]; 4Department of Ecology and Evolutionary Biology and Peabody Museum of Natural History, Yale University, New Haven, Connecticut 06520 USA [[email protected]]; 5Department of Herpetology and Center for Biodiversity and Conservation, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024 USA, and Department of Ecology, Evolution, and Environmental Biology, Columbia University, 2960 Broadway, New York, New York 10027 USA [[email protected]]; 6Section of Evolution and Ecology, University of California, Davis, California 95616 USA [[email protected]]; 7Department of Biology, University of Central Arkansas, Conway, Arkansas 72035 USA [[email protected]] ABSTRACT. – Based on a thorough review of the literature, we provide a bibliography of papers featuring phylogenetic hypotheses for living turtles, a composite tree of all turtle species based on those hypotheses, a compilation of the most rigorously derived trees from those papers (i.e., using contemporary methods with bootstrapping), and supertrees for selected families of turtles using input trees from those most rigorous trees.