2002), Thus, the Records Presented Here Repre- Sent Not Only A

March, 2006] NOTES 211

2002), thus, the records presented here repre- to Gujarat Ecological Education Research sent not only a significant eastern range exten- (GEER) Foundation, Gandhinagar. 7 pi. +18 sion for the species, but also new state and na- pp. tional records. RV thanks the Director, Gujarat Ecological Raju Vyas', Varad Giri^ and Aaron M. Bauer^ Education Research (GEER) Foundation, Gan- ' 505 Krishnadeep Tower, Mission Road, Fateh- dhinagar, India for logistic support and for per- gunj, Vadodara 390 002, Gujarat, India. mission for the study. Email: [email protected] 2 Bombay Natural History Society, Hombill LITERATURE CITED House, S. B. Singh Road, Mumbai 400 023, ANDERSON, S. C. 1999. The lizards of Iran. Soci- Maharashtra. India. ety for the Study of Amphibians and Reptiles, Email: [email protected] Oxford, Ohio, vii + 442 pp., 25 pis. CARRANZA, S. & E. N. ARNOLD. 2005. Systematics, ^ Department of Biology, Villanova University, biogeography, and evolution of Hemidactylus 800 Lancaster Avenue, Villanova, Pennsyl- geckos (Reptilia: Gekkonidae) elucidated us- vania 19085, U.S.A. ing mitochondrial DNA sequences. Molecular Email: [email protected] Phylogenetics and Evolution. 38(2):531-545. Received: 11 July 2005. CHAMPION, H. G. & S. K. SETH. 1968. A revised sur- Accepted: 31 July 2005. vey of the forest types of India. Manager of Publication, Government of India, New Del- hi. 7 map + 103 PI. + 404 pp. GAYEN, N. C. 1999. A synopsis of the reptiles of HamadryadYol. 30,1<¡os. 1 & 2, pp. 211 -213,2006. Gujarat, western India. Hamadryad 24:1-22. Copyright 2006 Centre for Herpetology, Madras Crocodile Bank Trust. MINTON, S. A. 1966. A contribution to the herpetology of West Pakistan. Bulletin of the Ameri- LEPIDODACmUS (SQUAMATA: GEKKONIDAE) IN can Museum of Natural History 134:27-184. ISLANDS ASIA: A L. AUREOUNEATUS FROM SULAWESI MURTHY, T. S. N. 1990. A field book of the lizards of India. Records of the Zoological Survey of Lepidodactylus geckos are modestly abundant India, Occasional Papers (115): 1-122. lizards in many Oceania herpetofaunas but be- SHARMA, R. C. 2000. Reptilia. In: State Fauna come increasingly uncommon to rare in the Series no. 8 Fauna of Gujarat (Part 1) Verte- faunas of the larger islands and island groups of brates, pp: 243-297. Q. H. Bagri (Ed). Zoo- the Pacific Rim, and even more uncommon in logical Survey of India, Calcutta. Islands Asia. This rarity has no current explana- . 2002. The fauna of India and adjacent tion, nor am I aware that anyone has attempted countries. Reptilia, Volume II (Sauria). Zoo- to explain it. Thus, a specimen of Lepidodac- logical Survey of India, Kolkata, India, xxv tylus from Sulawesi in the Naturalis collection + 430 pp. was unexpected. SMITH, M. A.1935. The fauna of British India, in- Presently, four species of Lepidodactylus are cluding Ceylon and Burma. Reptilia and Am- reported from Islands Asia. L. lugubris is the phibia. Vol. II. Sauria. Taylor and Francis, most widespread of these four and occurs in Cu London, xiii + 440 pp., 1 pi. Lao (Vietnam), Borneo, Sulawesi, Ambon, Hal- TIKADER, B. K. & R. C. SHARMA. 1992. Handbook. mahera. Témate, Komodo, and Lombok in this Indian lizards. Zoological Survey of India, area (Ineich, 1999). Because of its occurrence Calcutta. XV + 250 pp., 42 pis. also throughout the Pacific and coastally from VYAS, R. 2000. Comments on 'A synopsis of other Asian islands and mainland (de Rooij, the reptiles of Gujarat, India'. Hamadryad 1915), L. lugubris must be considered an exotic 25:203-207. species, probably beginning its human-mediated . 2005. Herpetofauna of the Jassore Wild- dispersal within the last five centuries through life Sanctuary. A technical report submitted European shipping. Its hybrid origin lies in east- 212 Hamadryad [Vol. 30, Nos. 1 & 2 em Micronesia and is relatively recent (Radtkey might have permitted natural dispersal; howev- et al., 1996). The other species are older island er, regular boat traffic occurs between these two residents, arriving and differentiating presum- coasts (C. A. Ross, pers. comm. Sept. 2005). As ably well before the arrival of humans in Asia. a regular resident of coconut palms and aerial These species have limited distributions: L. lom- ferns, individuals of Lepidodactylus aureolinea- bocensis, Lombok; L. intermedius, Komodo and tus might commonly be transported to Sulawesi. Rintja Islands.; L. ranauensis, Borneo (Sabah). Fortuitously, a single individual was captured Another species, L. listen, occurs to the west on and preserved by a Dutch biologist in the late Christmas Island in the Indian Ocean. 1930s. The preceding five taxa represent the three I wish to thank M. Hoogmoed and J. W. Am- phenetic groups (Brown and Parker, 1977) of tzen for assisting my study of the specimens in Lepidodactylus. L. lugubris is a Group III mem- the Netherlands' National Museum of Natural ber, L. intermedius and L. lombocensis Group II, History (Naturalis, RMNH). I also appreciate L. listen Group I, andi. ranauensis intermediate H. Ota's careful reading and comments for im- between Group I and II (Ota and Hikida, 1988; proving this manuscript; in review, A. M. Bauer, Bauer, 1994; Ota et al., 2000). Group III appears I. Ineich and R. M. Brown further clarified my to be most derived "clade" of Lepidodactylus, statements. My research into the biology and and with the exception of i. lugubris. Group III evolution of the Pacific and Australasian her- members occur on Pacific islands with the great- petofauna has been supported by the Smithso- est diversity in the Philippines. The Naturalis nian's Scholarly Studies Program, the Research specimen (RMNH 7341, Groot Sanghis, e/o [= Opportunity Fund of National Museum of Natu- vicinity of] Soemalata) from the north coast of ral History and the NMNH Department of Ver- Sulawesi was hidden under the lugubris epithet tebrate Zoology. but a close examination revealed its misidentifi- cation and dissection revealed that it was a ma- LITERATURE CITED ture male. My initial assumption was that this BAUER, A. M. 1994. Famiha Gekkonidae (Reptilia, specimen represented a new species, but further Sauria). Part I Australia and Oceania. Das examination revealed it to be a L. aureolineatus Tierreich 109. Berlin, Walter de Gruyter. xiii and a likely introduction from the Philippines. + 306 pp. Brown and Alcalá (1994:78-101) provide a BROWN, W. C. & A. C. ALCALÁ. 1977. Philippine thorough overview of the Philippine Lepidodac- lizards of the family Gekkonidae. Silliman tylus. They recognize two sections of Group III University, Dumaguete City, Philippines. 146 Lepidodactylus. Section A geckos have 4ToeL pp. (see Zug et al., 2003 for character abbreviations BROWN, W. C. & F. PARKER. 1977. Lizards of the and definitions) greater than 12, tail only mod- genus Lepidodactylus (Gekkonidae) from the erately depressed and no lateral skin flange, and Indo-Australian Archipelago and the islands combined femoral-precloacal pores greater than of the Pacific, with descriptions of new spe- 26. RMNH 7341 has 13 4ToeLm, moderately cies. Proceedings of the California Academy depressed tail without flange, and 29 FemPor + of Sciences, 4* ser. 41:253-265. PreclPor. Additionally, it has 115 Midb, which DE ROOlJ, N. 1915. The reptiles of the Indo-Aus- differentiates it from L. herrei, the other section tralian Archipelago. I. Lacertilia, Chelonia, K Lepidodactylus. In summary, RMNH 7341 is Emydosauria. E. J. Brill Ltd., Leiden, xiv + an adult male with large testes and epididymi- 384 pp. des, 35.7 mm SVL, 15.7 mm TrunkL, 42 mm INEICH, I. 1999. Spatio-temporal analysis of the TailL (regenerated tip), 9.4 mm HeadL, 6.5 mm unisexual-bisexual Lepidodactylus lugubris HeadW, 9 Suplab, 8 Inflab, 3 CircNa, 31 In- complex (Reptilia, Gekkonidae). In: Tropical torbS, 2 CloacS, 16 PreclP, and 8 4FingLm. island herpetofauna: origin, current diversity, The northern peninsula of Sulawesi is ca. 400 and conservation. pp:199-228. H. Ota (Ed). km from southern Mindanao and linked by an Elsevier Science B.V., Amsterdam. arc of islands (Kepulauan Sangihe). This arc OTA, H, R. I. S. DAREVSKY, I. INEICH & S. YAMASHIRO. March, 2006] NOTES 213

2000. Réévaluation of the taxonomic status of uamata: genus Lepidodactylus) from Tuvalu, two Lepidodactylus species (Squamata: Gek- south-central Pacific. Proceedings of the Bio- konidae) from the Lesser Sunda Archipelago, logical Society of Washington 116:38-46. Indonesia. Copeia 2002(4): 1109-1113. & T. HIKIDA. 1988. A new species oí Lepido- George R. Zug dactylus (Sauria: Gekkonidae) from Sabah. Department of Vertebrate Zoology, National Copeia 1988:616-621. Museum of Natural History, Smithsonian RADTKEY, R. R., B. BECKER, R. D. MILLER, R. RIBLET, Institution, Washington, D.C. 20013-7012, & T. J. CASE. 1996. Variation and evolution of U.S.A. class I Mhc in sexual and parthenogenetic Email: [email protected] geckos. Proceedings of the Royal Society of Londons 263:1023-1032. Received: 7 October 2005. ZUG, G. R., D. WATLING, T. ALEFAIO, S. ALEFAIO & C. Accepted: 27 October 2005. LUDESCHER. 2003. A new gecko (Reptilia: Sq-

REVIEWERS FOR HAMADRYAD VOL. 30, NOS. 1 & 2.

R. Bain, A. M. Bauer, W. Branch, R. M. Brown, A. S. Captain, R David, S. K. Dutta, V. Giri, I. Ineich, R. F. Inger, T.-M. Leong, C. J. McCarthy, H. Ota, S. Platt, B. Stuart, J. Sukumaran, R R van Diik, J. H. van Wyk, V. Wallach, G. Watkins-Colwell, R. Whitakerand E.-M. Zhao.