Australasian Journal of Herpetology 32 a Revised

Home , Daru Island, Lepidodactylus, Luperosaurus

32 Australasian Journal of Herpetology Australasian Journal of Herpetology 38:32-64. Published 10 August 2018. ISSN 1836-5698 (Print) ISSN 1836-5779 (Online)

A revised taxonomy of the gecko genera Lepidodactylus Fitzinger, 1843, Luperosaurus Gray, 1845 and Pseudogekko Taylor, 1922 including the formal erection of new genera and subgenera to accommodate the most divergent taxa and description of 26 new species.

RAYMOND T. HOSER

488 Park Road, Park Orchards, Victoria, 3134, Australia. Phone: +61 3 9812 3322 Fax: 9812 3355 E-mail: snakeman (at) snakeman.com.au Received 29 March 2017, Accepted 15 June 2018, Published 10 August 2018.

ABSTRACT There have been numerous studies published on species in the genera Lepidodactylus Fitzinger, 1843, Luperosaurus Gray, 1845, and Pseudogekko Taylor, 1922 with a view to assigning correct genus-level placement. Some authors have split the genera into species groups, while others have reassigned species between genera. More recently in view of newly available molecular data, it has been proposed to merge one or more genera into an expanded genus Gekko Laurenti, 1768. Relying on ancient divergences between species groups ascertained via molecular studies and consistent morphological differences between them, this paper provides a new taxonomy for the relevant genera (excluding Gekko and Ptychozoon Kuhl and Van Hassett, 1822, which are dealt with in separate papers) to better reflect known phylogeny of the relevant species. The result is a division of Lepidodactylus including the groups 1-3 as defined by Brown and Parker (1977) are divided into 9 genera, eight formally named for the first time, as well as two newly named subgenera, Luperosaurus into four genera, two being named for the first time as well as a newly named subgenus and Pseudogekko retained as a single genus, but with a new subgenus formally named to accommodate the most divergent species. Two other subgenera within Lepidodactylus sensu lato are formally named for the first time. Twenty six obviously unnamed species are also formally described for the first time. Keywords: Reptile; Taxonomy; Gecko; Lizard; Asia; Oceana; South-east Asia; Nomenclature; Luperosaurus; Lepidodactylus; Gekko; Pseudogekko; Ptychozoon; new genus; Shireenhosergecko; Jackyhosergecko; Bobbottomcolotes; Martinekcolotes; Adelynhosergecko; Allengreercolotes; Borneocolotes; Rosssadliercolotes; Charlespiersoncolotes; Georgemarioliscolotes; new subgenus; Borealiscolotes; Solomoncolotes; Haroldcoggercolotes; Robwatsoncolotes; new species; shireenhoserae; robjealousi; dalegibbonsi; jarradbinghami; petewhybrowi; jackyhoserae; bobbottomi; potens; crusmaculosus; adelynhoserae; sloppi; huonensis; madangensis; judyfergusonae; haydnmcphiei; matteoae; brettbarnetti; stevebennetti; lucybennettae; lachlanmcpheei; allengreeri; pauldarwini; paulwoolfi; haroldcoggeri; daranini; jenandersonae.

INTRODUCTION There have been numerous studies published on species in the Relying on ancient divergences between species groups as genera Luperosaurus Gray, 1845, Lepidodactylus Fitzinger, 1843 determined by recent phylogenetic studies and consistent and Pseudogekko Taylor, 1922 over the past 200 years with a view morphological differences between the taxa, this paper provides a to assigning correct genus-level placement. new and improved taxonomy for the relevant genera (excluding Gekko and Ptychozoon Kuhl and Van Hassett, 1822, which are Some studies in more recent years have split the genera into dealt with in separate papers) to better reflect known phylogeny of morphologically and/or genetically divergent species groups (e.g. the relevant species. Brown and Parker 1977, Ineich 2008, Stubbs et al. 2017, Oliver et al. 2018 and many others), while other published papers have The result is a division of Lepidodactylus including the three reassigned species between genera. More recently in view of species groups as defined by Brown and Parker (1977) into 9 newly available molecular data, it has been proposed to merge one genera, eight formally named for the first time, as well as two newly or more genera into an expanded genus Gekko Laurenti, 1768. named subgenera, Luperosaurus into four genera, two being Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 33

named for the first time as well as a newly named subgenus and The divergence dates of each group alone as documented by Pseudogekko is retained as a single genus, but divided into two Heinicke et al. (2012), Pyron et al. (2013) and others is more than divergent subgenera, one being named for the first time. sufficient justification for the splitting of the genera as understood Each genus group assigned in this paper has a divergence from to date. The morphological divergences of each species group as their nearest relatives of in excess of 25 MYA based on the well documented in the cited literature further confirms the evidence provided in Fig. 2 of Heinicke et al. 2012 or Oliver et al. sensibility of these decisions. (2018), with the exception of one genus grouping (formerly The twenty six species newly named in this paper for the first time included in Lepidodactylus) that diverged from its nearest relatives are obviously different to any previously named forms and just over 20 million years ago, those species being included in synonyms I was able to identify. They do not reflect all that there is Luperosaurus (the type group) and so significantly different to be named in all the various genera. I am of the firm view that morphologically that a genus level division is more appropriate within most of the genera defined herein there remain further than the placement of the named species group within undescribed species. Luperosaurus. I also note that, notwithstanding the theft of relevant materials from The subgenus grouping within the genus Pseudogekko also has a this author in an illegal armed raid on 17 August 2011, which were divergence in excess of 25 MYA based on the same evidence, but not returned in breach of undertakings to the court (Court of Appeal due to the well-defined and ancient monophyly of this particular Victoria 2014 and VCAT 2015), I have made a decision to publish group of species, it is retained as a single genus. this paper. In the process of auditing the relevant species and genera, twenty This is in view of the conservation significance attached to the six obviously unnamed species have been identified and named in formal recognition of unnamed taxa at all levels and on the basis accordance with the rules set out in the International Code of that further delays may in fact put these presently unnamed or Zoological Nomenclature (Ride et al. 1999).. potentially improperly assigned taxa at greater risk of extinction. MATERIALS, METHODS AND RESULTS This comment is made noting the extensive increase in human These are inferred in both the abstract and introduction, but as a population in the relevant region and the general environmental matter of trite I spell them out in a little more explicit detail. destruction across the planet as documented by Hoser (1991), The available literature was examined relevant to the genera including low density areas without a large permanent human Lepidodactylus Fitzinger, 1843, Luperosaurus Gray, 1845 and population. Pseudogekko Taylor, 1922 as defined by most authors in the I also note the abysmal environmental record of various National, previous 200 years. Additional to this has been inspection of State and Local governments in the relevant region over the past specimens as required and possible in order to ascertain the 200 years as detailed by Hoser (1989, 1991, 1993 and 1996) in the classification of the genera. face of ongoing threats as diverse as introduced species, habitat Available information in the form of photos of specimens with good destruction and modification, introduced pathogens and other available locality data and other information was also utilized in this factors and combinations thereof. study. Published literature relevant to the taxonomy and nomenclature As mentioned already, two of the three genera were split into a adopted within this paper includes the following: total of thirteen genera, with ten being formally named according to Amarasinghe et al. (2009), Auffenberg (1980), Barnett and Emms the rules of the International Code of Zoological Nomenclature (1997), Bauer (1994), Bauer and Sadlier (1994, 2000), Bauer and (Ride et al. 1999) for the first time. In the case of the split Henle (1994), Bauer and Vindum (1990), Bauer et al. (1995, 2007), Lepidodactylus I have refrained from assigning subgenera, except Bavay (1869), Binaday et al. (2017), Bleeker (1859), Bobrov and when absolutely obviously necessary, even though a strong case Semenov (2008), Boettger (1897), Boissinot et al. (1997), can be made for doing so in some other groups. Boulenger (1883, 1884, 1885a, 1885b, 1886, 1887a, 1887b, 1889, The depth of divergence of the Philippine genus Pseudogekko 1897, 1920), Brongersma (1934, 1948), Brown (1964), Brown and Taylor, 1922 could have warranted a full genus-level split, because Alcala (1978), Brown and Parker (1977), Brown and Tanner the most divergent species had divergences of over 25 MYA from (1949), Brown et al. (1992), Brown and Diesmos (2000), Brown et the rest. They do however sit within a broadly monophyletic group. al. (2000, 2007, 2010, 2011, 2012, 2013), Buden (2007a, 2007b, For that reason I have moved two species in that genus to a new 2008, 2015a, 2015b), Buden and Taboroši (2016), Buden et al. subgenus. These are the species Lepidodactylus brevipes (2014), Cogger (2014), Cogger et al. (1983), Crombie and Menz Boettger, 1897, now known as Pseudogekko brevipes (Boettger, (2007), Crombie and Pregill (1999), Cuéllar and Kluge (1972), 1867) and the morphologically similar Pseudogekko atiorum Davis, Daan and Hillenius (1966), Darevsky (1964), Das (2004, 2005), Watters, Köhler, Whitsett, Huron, Brown, Diesmos and Siler, 2015. Das et al. (2008), Davis et al. (2015), de Rooij (1915), Dolino et al. In summary Lepidodactylus Fitzinger, 1843 has been divided as (2009), Duméril and Bibron (1836), Ferner et al. (2000), Fitzinger follows: (1843, 1861), Gardner (1985), Garman (1901, 1908), Gaulke The so-called group 1 of species as defined by Brown and Parker (2011, 2013), Gaulke and Attenbach (2006), Gaulke et al. (2003, (1977) has been split into four divergent genera, all formally named 2007), Gibbons and Brown (1988), Gibson-Hill (1947, 1950), Gill for the first time, with one of these groups split into two subgenera, (1993), Girard (1858), Goldberg (2017), Gray (1845), Grismer the new subgenus also being formally named. (2011a, 2011b), Grismer et al. (2002), Günther (1864), Han et al. The so-called group 2 of species as defined by Brown and Parker (2004), Heinicke et al. (2012), Henderson et al. (1996), Ineich (1977) has been split into four genera, all formally named for the (2008, 2011, 2009, 2015), Ineich and Ota (1993), Iskander and first time and including a new subgenus. Mumpuni (2002), Jean-Baptiste (2013), Kiehlmann (2014), Kluge The so-called group 3 of species as defined by Brown and Parker (1967, 1968), Koch (2011, 2012), Kopstein (1926), Lenort (2004), (1977) includes the type for the genus Lepidodactylus and the Loveridge (1948), Macleay (1877), Malkmus et al. (2002), Manthey synonym Amydosaurus Gray, 1845 (also for the species and Grossmann (1997), McCoy (2006, 2015), Meiri et al. (2017), Platydactylus lugubris Duméril and Bibron, 1836) and the so called Mertens (1922, 1929, 1930, 1967), Morrison (2003), Müller (1895), group 3 of species is now the sole content of the genus. Oliver and Hugall (2017), Oliver et al. (2018a, 2018b), Ota (1987, 1989), Ota and Crombie (1989), Ota and Hikida (1988), Ota et al. Amydosaurus cannot therefore be an available name for any other (1995, 1996, 1998, 2000), Pernetta and Black (1983), Peters species group. (1867, 1874), Peters and Doria (1878), Pianka and Vitt (2003), Lepidodactylus is also split into two subgenera, the new subgenus Pyron et al. (2013), Radtkey et al. (1995), Ride et al. (1999), Röll being named herein for the first time. (2002, 2006), Röll and von Düring (2008), Rösler (1995, 2000, The genus Luperosaurus Gray, 1845 has been divided four ways, 2017), Rösler et al. (2005, 2011, 2012), Russell (1979), Sadlier and with two genera formally named for the first time as well as a newly Bauer (1997), Sakai (2016), Sang et al. (2009), Sanguila et al. named subgenus. (2016), Savage (2002), Siler and Brown (2010), Siler et al. (2010, Pseudogekko Taylor, 1922 has been split into two subgenera. 2012, 2013, 2014, 2016, 2017), Slevin and Leviton (1956), Smith Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 34 Australasian Journal of Herpetology and Grant (1961), Somaweera and Somaweera (2009), Stejneger mandatory for all papers naming new taxa in terms of rules of the (1989, 1905, 1907), Stoliczka (1870), Stubbs et al. (2017), Supsup International Code of Zoological Nomenclature (future editions) et al. (2016), Taylor (1915, 1917, 1918, 1919, 1922a, 1922b, 1923, and this is one I support. As a matter of trite, I doubt anyone on 1944), Trautmann (1988), Turner and Green (1996), Volobouev et the planet would argue against the merits of peer review. However al. (1993), Wells and Wellington (1984, 1985), Werner (1900, this peer review must be done properly and not in name only, or by 1913), Wilson and Swan (2010), Yamashiro and Ota (1998, 2005), a means to ensure a corrupt cohort cannot usurp the role of the Yamashiro et al. (2000), Zug (1991, 2006), Zug and Kaiser (2014), ICZN in determining correct and legal nomenclature. Zug et al. (2003, 2011, 2012) and sources cited therein. GENUS LEPIDODACTYLUS FITZINGER, 1843 In terms of the nomenclature adopted within this paper, the Type species: Platydactylus lugubris Duméril and Bibron, 1843. following points should also be noted. Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, Spellings of names should not be altered in any way unless Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., absolutely mandatory according to the rules of the International Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., Code of Zoological Nomenclature (Ride et al. 1999). Gender or Adelynhosergecko gen. nov., Allengreercolotes gen. nov., alleged gender of names should not be altered. In the unlikely Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until event that a later author or so-called “first reviser” seeks to merge now treated as being of the single genus Lepidodactylus are named taxa, then the name to be used should be that first used in separated from all other geckos by the following suite of this paper, as dictated by page priority and order in the keywords of characters: Digits are moderately long, more or less dilated, free or the abstract. with a rudiment of web, inferiorly with a single or double series of Material may be repeated in sequential descriptions in order to transverse lamellae divided by a median groove and across the ensure that each complies wholly with the rules of the International entire pad, forming the scansor, with very short and free Code of Zoological Nomenclature (Ride et al. 1999). compressed distal non-retractile clawed joint raising from the Finally there are some other significant points worth mentioning extremity of the dilated and compressed digit; inner digit clawless. which will not be immediately obvious to readers of this paper or Body covered above with granular scales, inferiorly with juxtaposed the references cited herein. or subimbricate scales. Rostral and mental shields rounded; labials Without exception, all newly named species have been shown to are much larger than adjacent scales. Pupil vertical. Males with be separate species based on molecular data and depth of preanal pores. divergences. They also have morphological differences from one Lepidodactylus as defined herein are separated from the other another. genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Available information strongly indicates numerous more as yet Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., unnamed species. Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Invariably the named and unnamed species are separated by one Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by or other of lowland areas between hills and mountains or deep sea having a reduced number of scansors, with the terminal as well as trenches that remained submerged at times of glacial maxima. a few subterminal scansors divided and a significantly depressed The physical barriers themselves may not be the isolating body and flat and broad tail, versus a subcylindrical tail lacking mechanism in total, this also potentially including competing lateral flanges or spines in all the other genera. species, predators and other factors. This genus also consists of the entirety of the so-called group 3 of The molecular studies cited above do show clearly species and species as defined by Brown and Parker (1977). genus level differences between taxa. Geckos in the subgenus Borealiscolotes subgen. nov. type species However I must note that in the process of auditing these results, Lepidodactylus balioburius Ota and Crombie, 1989 being the only including inspection of relevant specimens subject of the other subgenus within Lepidodactylus are separated from the morphological and molecular studies I repeatedly found that nominate subgenus by the following suite of characters: less than authors had misidentified specimens at the species level and in 10 scansors confined to the distal half of broadly dilated and some cases even the genus level, and this being based on the strongly webbed digits; a strongly flattened tail with a broad flange genus-level taxonomy pre-dating this paper (i.e. the prevailing and of skin; the nostril is separated by a scale from the rostral (as used taxonomy and nomenclature at the relevant time). opposed to being in contact in specimens of the nominate As to why the misidentifications occurred, it is self evident that in subgenus). the molecular study papers at least, the publishing authors either Species of gecko in the genera Shireenhosergecko gen. nov., did not inspect the relevant specimens for which they put data in Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and their papers or alternatively, any inspections done were either Martinekcolotes gen. nov. all consist of geckos formerly placed in cursory, inadequate, or in the absence of knowing the diagnostic the so-called group 1 of species within Lepidodactylus as defined features of the alleged taxon. by Brown and Parker (1977). Collectively they are separated from These significant facts will explain why taxonomic statements and the other genera formerly included within Lepidodactylus (these conclusions in this paper will be at times at variance with the earlier being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko cited papers, which are still largely relied upon. gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. As this paper’s purpose it to name unnamed taxa before they and Rosssadliercolotes gen. nov.) by having numerous (genus) become extinct and not a critique of earlier papers by other authors Gekko-like undivided scansors on all digits and a subcylindrical tail. I will not dwell at length on this. Geckos in the genus Shireenhosergecko gen. nov. are separated However it is appropriate that I should also mention the serious from the other three genera Jackyhosergecko gen. nov., issue of editorial control, and associated alleged peer review, Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one including the serial offender, the online journal Zootaxa as dealt or other of the following five suites of characters: with in Hoser (2015a-f) and papers cited therein. 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, That journal (Zootaxa), described by many as PRINO (peer 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; reviewed in name only), by way of example relevant here, about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal published a paper Stubbs et al. (2017) complete with phylogram and femoral pores extending to the distal end of the thigh; bearing including erroneously identified species, rendering it largely a continuous series of 27-34 pores in males; length of hind limb useless and/or totally misleading to a casual reader who will not usually more than 70% of axilla-groin distance (S. mutahi), or: actually cross-check data with the relevant museum specimens. 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, Even a cursory review by a non-expert could have fixed the most 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at serious errors in that paper, but at even that level Zootaxa was an the base only between the third and fourth toes; 32-36 enlarged abject failure. preanal and femoral pores extending to the distal end of the thigh in a continuous series extending over the proximal 2/3 of the thigh; There have been calls from many quarters to make peer review

bearing 28-30 pores in males (S. oorti), or: medially. The subcylindrical tail lacks fringes or flanges. 3/ 37-43 mm snout-vent length in adults; digits long and slender; Geckos in the genus Adelynhosergecko gen. nov. are readily moderately dilated, limited interdigital webbing; 10-12 entire fourth separated from geckos in the genera Allengreercolotes gen. nov., toe scansors covering the distal 2/3 of the toe; 20-24 enlarged Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by scales in pore series confined to the preanal region only or only a having fewer than 25 enlarged preanal/femoral pores, snout-vent few on the base of the thigh, bearing about 19 preanal pores in length in adults of under 48 mm and of moderate to solid build (A. males (S. orientalis, S. petewhybrowi sp. nov.), or: novaeguineae, A. buleli, A. huonensis sp. nov., A. madangensis sp. 4/ 37-43 mm snout-vent length in adults; digits long and slender; nov., A. oligoporus, A. pulcher and A. sloppi sp. nov.). moderately dilated, extensive interdigital webbing; 10-12 entire Geckos in the genus Allengreercolotes gen. nov. are readily fourth toe scansors covering the distal 2/3 of the toe; 20-24 separated from geckos in the genera Adelynhosergecko gen. nov., enlarged scales in pore series confined to the preanal region only Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by or only a few on the base of the thigh, bearing about 19 preanal having 91-121 mid-body scale rows, 30 or more enlarged preanal / pores in males (S. shireenhoserae sp. nov., S. dalegibbonsi sp. femoral scales, chin secondary postmentals and less than 20% nov., S. jarradbinghami sp. nov., S. robjealousi sp. nov.), or: webbing on feet (A. allengreeri sp. nov., A. gardineri, A. guppyi, A. 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the intermedius, A. lombocensis, A. pauldarwini sp. nov., A. shebae, digits; more than 25 but fewer than 40 enlarged pre-anal-femoral A. tepukapili, A. vanuatuensis). scales in males, extending onto the distal portion of the thigh; Geckos in the genus Borneocolotes gen. nov. are readily enlarged preanal row not separated from scutes (S. browni). separated from geckos in the genera Adelynhosergecko gen. nov., Geckos in the genus Jackyhosergecko gen. nov. are separated Allengreercolotes gen. nov. and Rosssadliercolotes gen. nov. by from the other three genera, Shireenhosergecko gen. nov., having the following suite of characters: Slightly depressed, but not Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one as much as seen in Lepidodactylus (sensu stricto) as defined in or other of the following three suites of characters: this paper as found mainly in the Philippines; nine upper labials; 1/ Enlarged ventral scales limited to the preanal region and nine or ten lower labials; 100-110 mid body rows, long narrow digits proximal position of the thighs; fewer than 20 enlarged scales in with scansors underneath the distal four fifths; the terminal scansor pore series; males with 15 or fewer preanal pores; digits are is entire, one subterminal scansor is wholly entire, deeply notched broadly dilated, dark reddish-brown dorsal colouration with orange- or wholly divided; eight scansors beneath toe 4. Webbing absent red patches on the rear upper labials and neck (J. manni), or: between digits 1 and 2, and slight webbing between digits 3 and 4. 2/ Fewer than 15 scansors under digit 4 of the hind leg; more than Adult males have a continuous series of 36-38 pores in preanal 25 enlarged preanal/femoral scales, extending onto distal portions and femoral regions; femoral series extends almost to the distal of the thigh; 25 or more preanal/femoral pores in males; fewer than end of the thigh; male cloacal spur is single and large, versus 40 enlarged femoral scales; enlarged preanal scale row is medium in other species in the genera Adelynhosergecko gen. separated by a row of short scales from short rows of enlarged nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. preanal scutes webbing between digits 3 and 4 of hind leg less of the same size; tail is subcylindrical and without lateral than 1/5 of the digit length; eye diameter is usually 41% or less ornamentation; dorsal body colouration is grayish brown and than the head width (J. euaensis), or: ventrally grayish tan; there is a pair of dark spots on the dorsolateral part of the basal swollen area of the tail (B. 3/ More than 15 scansors under digit 4 of hind leg; more than 25 ranauensis). enlarged preanal/femoral scales, extending onto distal portions of the thigh; 25 or more preanal/femoral pores in males (J. Geckos in the genus Rosssadliercolotes gen. nov. are readily flaviocularis: Subgenus Solomoncolotes subgen. nov.). separated from geckos in the genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov. and Borneocolotes gen. nov. by having Geckos in the genus Bobbottomcolotes gen. nov. are separated one or other of the following two suites of characters: from the other three genera, Shireenhosergecko gen. nov., Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the 1/ Less than 35 enlarged scales in preanal/femoral region; sixteen following characters: More than 25 enlarged preanal/femoral or fewer scansors on toe IV; more than 25 enlarged scales in scales, extending onto distal portions of the thigh; 25 or more preanal/femoral region; less than 110 scales around midbody (R. preanal/femoral pores in males; fewer than 15 scansors under digit paurolepis), or: 4 of the hind leg: and one or other of: 2/ The unique combination of both divided terminal scansors on all 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ toes (including toe 4) and a nearly completely cylindrical tail without 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: fringes or evidence of dorsoventral compression (R. pantai). 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 Distribution: Most species of Lepidodactylus as defined herein the length of the digits; 40 or more enlarged preanal/femoral occur in the Philippines, but some occur north of the main scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. archipelago and also widely across the Pacific. magnus, B. bobbottomi sp. nov., B. potens sp. nov.). Content: Lepidodactylus lugubris (Duméril and Bibron, 1836) (type Geckos in the genus Martinekcolotes gen. nov. are separated from species); L. aureolineatus Taylor, 1915; L. balioburius Ota and the other three genera, Shireenhosergecko gen. nov., Crombie, 1989; L. christiani Taylor, 1917; L. herrei Taylor, 1923; L. Jackyhosergecko gen. nov. and Bobbottomcolotes gen. nov. by the moestus (Peters, 1867); L. planicaudus Stejneger, 1905; L. following characters: Enlarged ventral scales limited to the preanal woodfordi Boulenger, 1887; L. labialis (Peters, 1867); L. yami Ota, region and proximal position of the thighs; fewer than 20 enlarged 1987. scales in pore series; males with 15 or fewer preanal pores; digits SUBGENUS BOREALISCOLOTES SUBGEN. NOV. are broadly dilated, colour is pale grey-brown above, with a few Type species: Lepidodactylus balioburius Ota and Crombie, 1989. rather indistinct brown spots and scattered white dots and a brown Diagnosis: Geckos in the subgenus Borealiscolotes subgen. nov. streak running from the nostril to the eye; lower parts white with type species Lepidodactylus balioburius Ota and Crombie, 1989 small brown spots (M. listeri). being the only other subgenus within Lepidodactylus are separated Geckos in the genera Adelynhosergecko gen. nov., from the nominate subgenus by the following suite of characters: Allengreercolotes gen. nov., Borneocolotes gen. nov. and less than 10 scansors confined to the distal half of broadly dilated Rosssadliercolotes gen. nov. all consist of geckos formerly placed and strongly webbed digits; a strongly flattened tail with a broad in the so-called group 2 of species within Lepidodactylus as flange of skin; the nostril is separated by a scale from the rostral defined by Brown and Parker (1977). (as opposed to being in contact in specimens of the nominate They can be separated from all other species formerly placed in subgenus). Lepidodactylus sensu lato by the possession of well developed Geckos of the genera Lepidodactylus Fitzinger, 1843, scansors on almost all the undersurface of the digits and with one Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., or more, usually being a few, subterminal scansors divided Bobbottomcolotes gen. nov., Martinekcolotes gen. nov.,

Adelynhosergecko gen. nov., Allengreercolotes gen. nov., moderately dilated, extensive interdigital webbing; 10-12 entire Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until fourth toe scansors covering the distal 2/3 of the toe; 20-24 now treated as being of the single genus Lepidodactylus are enlarged scales in pore series confined to the preanal region only separated from all other geckos by the following suite of or only a few on the base of the thigh, bearing about 19 preanal characters: Digits are moderately long, more or less dilated, free or pores in males (S. shireenhoserae sp. nov., S. jarradbinghami sp. with a rudiment of web, inferiorly with a single or double series of nov., S. dalegibbonsi sp. nov., S. robjealousi sp. nov.), or: transverse lamellae divided by a median groove and across the 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the entire pad, forming the scansor, with very short and free digits; more than 25 but fewer than 40 enlarged pre-anal-femoral compressed distal non-retractile clawed joint raising from the scales in males, extending onto the distal portion of the thigh; extremity of the dilated and compressed digit; inner digit clawless. enlarged preanal row not separated from scutes (S. browni). Body covered above with granular scales, inferiorly with juxtaposed Geckos of the genera Lepidodactylus Fitzinger, 1843, or subimbricate scales. Rostral and mental shields rounded; labials Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., are much larger than adjacent scales. Pupil vertical. Males with Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., preanal pores. Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Lepidodactylus as defined herein are separated from the other Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., now treated as being of the single genus Lepidodactylus are Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., separated from all other geckos by the following suite of Adelynhosergecko gen. nov., Allengreercolotes gen. nov., characters: Digits are moderately long, more or less dilated, free or Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by with a rudiment of web, inferiorly with a single or double series of having a reduced number of scansors, with the terminal as well as transverse lamellae divided by a median groove and across the a few subterminal scansors divided and a significantly depressed entire pad, forming the scansor, with very short and free body and flat and broad tail, versus a subcylindrical tail lacking compressed distal non-retractile clawed joint raising from the lateral flanges or spines in all the other genera. extremity of the dilated and compressed digit; inner digit clawless. This genus also consists of the entirety of the so-called group 3 of Body covered above with granular scales, inferiorly with juxtaposed species as defined by Brown and Parker (1977). or subimbricate scales. Rostral and mental shields rounded; labials Distribution: Northern Philippines and islands north of there. are much larger than adjacent scales. Pupil vertical. Males with Etymology: Borealiscolotes in Latin means northern gecko, which preanal pores. is in reflection of the relative distribution of these species relative to Lepidodactylus as defined herein are separated from the other others in the genus, or more widely in the clade of nine genera that genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., have until now been treated as within a greater Lepidodactylus. Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., Content: Lepidodactylus (Borealiscolotes) balioburius Ota and Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Crombie, 1989 (type species); Lepidodactylus (Borealiscolotes) Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by christiani Taylor, 1917; Lepidodactylus (Borealiscolotes) yami Ota, having a reduced number of scansors, with the terminal as well as 1987. a few subterminal scansors divided and a significantly depressed GENUS SHIREENHOSERGECKO GEN. NOV. body and flat and broad tail, versus a subcylindrical tail lacking lateral flanges or spines in all the other genera. Type species: Shireenhosergecko shireenhoserae sp. nov. (this paper). This genus also consists of the entirety of the so-called group 3 of species as defined by Brown and Parker (1977). Diagnosis: Species of gecko in the genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. Geckos in the subgenus Borealiscolotes subgen. nov. type species and Martinekcolotes gen. nov. all consist of geckos formerly placed Lepidodactylus balioburius Ota and Crombie, 1989 being the only in the so-called group 1 of species within Lepidodactylus as other subgenus within Lepidodactylus are separated from the defined by Brown and Parker (1977). Collectively they are nominate subgenus by the following suite of characters: less than separated from the other genera formerly included within 10 scansors confined to the distal half of broadly dilated and Lepidodactylus (these being genera Lepidodactylus Fitzinger, strongly webbed digits; a strongly flattened tail with a broad flange 1843, Adelynhosergecko gen. nov., Allengreercolotes gen. nov., of skin; the nostril is separated by a scale from the rostral (as Borneocolotes gen. nov. and Rosssadliercolotes gen. nov.) by opposed to being in contact in specimens of the nominate having numerous (genus) Gekko-like undivided scansors on all subgenus). digits and a subcylindrical tail. Distribution: New Guinea and nearby islands. Geckos in the genus Shireenhosergecko gen. nov. are separated Content: Shireenhosergecko shireenhoserae sp. nov. (type from the other three genera Jackyhosergecko gen. nov., species); S. browni (Pernetta and Black, 1983); S. dalegibbonsi sp. Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one nov.; S. jarradbinghami sp. nov.; S. mutahi (Brown and Parker, or other of the following five suites of characters: 1977); S. oortii (Kopstein, 1926); S. orientalis (Brown and Parker, 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, 1977); S. petewhybrowi sp. nov.; S. robjealousi sp. nov.. 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; GENUS JACKYHOSERGECKO GEN. NOV. about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal Type species: Lepidodactylus euaensis Gibbons and Brown, and femoral pores extending to the distal end of the thigh; bearing 1988. a continuous series of 27-34 pores in males; length of hind limb Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, usually more than 70% of axilla-groin distance (S. mutahi), or: Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at Adelynhosergecko gen. nov., Allengreercolotes gen. nov., the base only between the third and fourth toes; 32-36 enlarged Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until preanal and femoral pores extending to the distal end of the thigh now treated as being of the single genus Lepidodactylus are in a continuous series extending over the proximal 2/3 of the thigh; separated from all other geckos by the following suite of bearing 28-30 pores in males (S. oorti), or: characters: Digits are moderately long, more or less dilated, free or 3/ 37-43 mm snout-vent length in adults; digits long and slender; with a rudiment of web, inferiorly with a single or double series of moderately dilated, limited interdigital webbing; 10-12 entire fourth transverse lamellae divided by a median groove and across the toe scansors covering the distal 2/3 of the toe; 20-24 enlarged entire pad, forming the scansor, with very short and free scales in pore series confined to the preanal region only or only a compressed distal non-retractile clawed joint raising from the few on the base of the thigh, bearing about 19 preanal pores in extremity of the dilated and compressed digit; inner digit clawless. males (S. orientalis, S. petewhybrowi sp. nov.), or: Body covered above with granular scales, inferiorly with juxtaposed 4/ 37-43 mm snout-vent length in adults; digits long and slender; or subimbricate scales. Rostral and mental shields rounded; labials

are much larger than adjacent scales. Pupil vertical. Males with toe scansors covering the distal 2/3 of the toe; 20-24 enlarged preanal pores. scales in pore series confined to the preanal region only or only a Lepidodactylus as defined herein are separated from the other few on the base of the thigh, bearing about 19 preanal pores in genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., males (S. orientalis, S. petewhybrowi sp. nov.), or: Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., 4/ 37-43 mm snout-vent length in adults; digits long and slender; Adelynhosergecko gen. nov., Allengreercolotes gen. nov., moderately dilated, extensive interdigital webbing; 10-12 entire Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by fourth toe scansors covering the distal 2/3 of the toe; 20-24 having a reduced number of scansors, with the terminal as well as enlarged scales in pore series confined to the preanal region only a few subterminal scansors divided and a significantly depressed or only a few on the base of the thigh, bearing about 19 preanal body and flat and broad tail, versus a subcylindrical tail lacking pores in males (S. shireenhoserae sp. nov., S. dalegibbonsi sp. lateral flanges or spines in all the other genera. nov., S. jarradbinghami sp. nov., S. robjealousi sp. nov.), or: This genus also consists of the entirety of the so-called group 3 of 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the species as defined by Brown and Parker (1977). digits; more than 25 but fewer than 40 enlarged pre-anal-femoral Geckos in the subgenus Borealiscolotes subgen. nov. type species scales in males, extending onto the distal portion of the thigh; Lepidodactylus balioburius Ota and Crombie, 1989 being the only enlarged preanal row not separated from scutes (S. browni). other subgenus within Lepidodactylus are separated from the Geckos in the genus Bobbottomcolotes gen. nov. are separated nominate subgenus by the following suite of characters: less than from the other three genera, Shireenhosergecko gen. nov., 10 scansors confined to the distal half of broadly dilated and Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the strongly webbed digits; a strongly flattened tail with a broad flange following characters: More than 25 enlarged preanal/femoral of skin; the nostril is separated by a scale from the rostral (as scales, extending onto distal portions of the thigh; 25 or more opposed to being in contact in specimens of the nominate preanal/femoral pores in males; fewer than 15 scansors under digit subgenus). 4 of the hind leg: and one or other of: Species of gecko in the genera Shireenhosergecko gen. nov., 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: Martinekcolotes gen. nov. all consist of geckos formerly placed in 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 the so-called group 1 of species within Lepidodactylus as defined the length of the digits; 40 or more enlarged preanal/femoral by Brown and Parker (1977). Collectively they are separated from scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. the other genera formerly included within Lepidodactylus (these magnus, B. bobbottomi sp. nov., B. potens sp. nov.). being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko Geckos in the genus Martinekcolotes gen. nov. are separated from gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. the other three genera, Shireenhosergecko gen. nov., and Rosssadliercolotes gen. nov.) by having numerous (genus) Jackyhosergecko gen. nov. and Bobbottomcolotes gen. nov. by the Gekko-like undivided scansors on all digits and a subcylindrical tail. following characters: Enlarged ventral scales limited to the preanal Geckos in the genus Jackyhosergecko gen. nov. are separated region and proximal position of the thighs; fewer than 20 enlarged from the other three genera, Shireenhosergecko gen. nov., scales in pore series; males with 15 or fewer preanal pores; digits Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one are broadly dilated, colour is pale grey-brown above, with a few or other of the following three suites of characters: rather indistinct brown spots and scattered white dots and a brown 1/ Enlarged ventral scales limited to the preanal region and streak running from the nostril to the eye; lower parts white with proximal position of the thighs; fewer than 20 enlarged scales in small brown spots (M. listeri). pore series; males with 15 or fewer preanal pores; digits are Distribution: Jackyhosergecko gen. nov. species occur in Tonga broadly dilated, dark reddish-brown dorsal colouration with orange- (J. euaensis), Fiji Islands (J. manni and J. jackyhoserae sp. nov.) red patches on the rear upper labials and neck (J. manni), or: and Guadalcanal, Solomon Islands (J. flaviocularis). 2/ Fewer than 15 scansors under digit 4 of the hind leg; more than Etymology: Named in honour of my youngest daughter Jacky 25 enlarged preanal/femoral scales, extending onto distal portions Hoser of Park Orchards, Victoria, Australia in recognition of her of the thigh; 25 or more preanal/femoral pores in males; fewer than services to wildlife conservation over nearly 2 decades. For more 40 enlarged femoral scales; enlarged preanal scale row is detail see Hoser (2013) at page 5. separated by a row of short scales from short rows of enlarged Content: Jackyhosergecko euaensis (Gibbons and Brown, 1988) preanal scutes webbing between digits 3 and 4 of hind leg less (type species); J. jackyhoserae sp. nov.; J. manni (Schmidt, 1923); than 1/5 of the digit length; eye diameter is usually 41% or less J. flaviocularis (Brown, McCoy and Rodda, 1992). than the head width (J. euaensis), or: SUBGENUS SOLOMONCOLOTES SUBGEN. NOV. 3/ More than 15 scansors under digit 4 of hind leg; more than 25 Type species: Lepidodactylus flaviocularis Brown, McCoy and enlarged preanal/femoral scales, extending onto distal portions of Rodda, 1992. the thigh; 25 or more preanal/femoral pores in males (J. Diagnosis: Geckos in the subgenus Solomoncolotes subgen. nov. flaviocularis: Subgenus Solomoncolotes subgen. nov.). are separated from the nominate genus by the following suite of Geckos in the genus Shireenhosergecko gen. nov. are separated characters: More than 15 scansors under digit 4 of hind leg; more from the other three genera Jackyhosergecko gen. nov., than 25 enlarged preanal/femoral scales, extending onto distal Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one portions of the thigh; 25 or more preanal/femoral pores in males. or other of the following five suites of characters: Geckos of the genera Lepidodactylus Fitzinger, 1843, 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal Adelynhosergecko gen. nov., Allengreercolotes gen. nov., and femoral pores extending to the distal end of the thigh; bearing Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until a continuous series of 27-34 pores in males; length of hind limb now treated as being of the single genus Lepidodactylus are usually more than 70% of axilla-groin distance (S. mutahi), or: separated from all other geckos by the following suite of 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, characters: Digits are moderately long, more or less dilated, free or 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at with a rudiment of web, inferiorly with a single or double series of the base only between the third and fourth toes; 32-36 enlarged transverse lamellae divided by a median groove and across the preanal and femoral pores extending to the distal end of the thigh entire pad, forming the scansor, with very short and free in a continuous series extending over the proximal 2/3 of the thigh; compressed distal non-retractile clawed joint raising from the bearing 28-30 pores in males (S. oorti), or: extremity of the dilated and compressed digit; inner digit clawless. 3/ 37-43 mm snout-vent length in adults; digits long and slender; Body covered above with granular scales, inferiorly with juxtaposed moderately dilated, limited interdigital webbing; 10-12 entire fourth or subimbricate scales. Rostral and mental shields rounded; labials

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 38 Australasian Journal of Herpetology are much larger than adjacent scales. Pupil vertical. Males with moderately dilated, limited interdigital webbing; 10-12 entire fourth preanal pores. toe scansors covering the distal 2/3 of the toe; 20-24 enlarged Lepidodactylus as defined herein are separated from the other scales in pore series confined to the preanal region only or only a genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., few on the base of the thigh, bearing about 19 preanal pores in Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., males (S. orientalis, S. petewhybrowi sp. nov.), or: Adelynhosergecko gen. nov., Allengreercolotes gen. nov., 4/ 37-43 mm snout-vent length in adults; digits long and slender; Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by moderately dilated, extensive interdigital webbing; 10-12 entire having a reduced number of scansors, with the terminal as well as fourth toe scansors covering the distal 2/3 of the toe; 20-24 a few subterminal scansors divided and a significantly depressed enlarged scales in pore series confined to the preanal region only body and flat and broad tail, versus a subcylindrical tail lacking or only a few on the base of the thigh, bearing about 19 preanal lateral flanges or spines in all the other genera. pores in males (S. shireenhoserae sp. nov., S. dalegibbonsi sp. This genus also consists of the entirety of the so-called group 3 of nov., S. jarradbinghami sp. nov., S. robjealousi sp. nov.), or: species as defined by Brown and Parker (1977). 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the Geckos in the subgenus Borealiscolotes subgen. nov. type species digits; more than 25 but fewer than 40 enlarged pre-anal-femoral Lepidodactylus balioburius Ota and Crombie, 1989 being the only scales in males, extending onto the distal portion of the thigh; other subgenus within Lepidodactylus are separated from the enlarged preanal row not separated from scutes (S. browni). nominate subgenus by the following suite of characters: less than Geckos in the genus Bobbottomcolotes gen. nov. are separated 10 scansors confined to the distal half of broadly dilated and from the other three genera, Shireenhosergecko gen. nov., strongly webbed digits; a strongly flattened tail with a broad flange Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the of skin; the nostril is separated by a scale from the rostral (as following characters: More than 25 enlarged preanal/femoral opposed to being in contact in specimens of the nominate scales, extending onto distal portions of the thigh; 25 or more subgenus). preanal/femoral pores in males; fewer than 15 scansors under digit Species of gecko in the genera Shireenhosergecko gen. nov., 4 of the hind leg: and one or other of: Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ Martinekcolotes gen. nov. all consist of geckos formerly placed in 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: the so-called group 1 of species within Lepidodactylus as defined 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 by Brown and Parker (1977). Collectively they are separated from the length of the digits; 40 or more enlarged preanal/femoral the other genera formerly included within Lepidodactylus (these scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko magnus, B. bobbottomi sp. nov., B. potens sp. nov.). gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. Geckos in the genus Martinekcolotes gen. nov. are separated from and Rosssadliercolotes gen. nov.) by having numerous (genus) the other three genera, Shireenhosergecko gen. nov., Gekko-like undivided scansors on all digits and a subcylindrical tail. Jackyhosergecko gen. nov. and Bobbottomcolotes gen. nov. by the Geckos in the genus Jackyhosergecko gen. nov. are separated following characters: Enlarged ventral scales limited to the preanal from the other three genera, Shireenhosergecko gen. nov., region and proximal position of the thighs; fewer than 20 enlarged Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one scales in pore series; males with 15 or fewer preanal pores; digits or other of the following three suites of characters: are broadly dilated, colour is pale grey-brown above, with a few 1/ Enlarged ventral scales limited to the preanal region and rather indistinct brown spots and scattered white dots and a brown proximal position of the thighs; fewer than 20 enlarged scales in streak running from the nostril to the eye; lower parts white with pore series; males with 15 or fewer preanal pores; digits are small brown spots (M. listeri). broadly dilated, dark reddish-brown dorsal colouration with orange- Distribution: Known only from Guadalcanal, Solomon Islands (J. red patches on the rear upper labials and neck (J. manni), or: flaviocularis) from the type series. 2/ Fewer than 15 scansors under digit 4 of the hind leg; more than Etymology: The subgenus Solomoncolotes subgen. nov. is named 25 enlarged preanal/femoral scales, extending onto distal portions in reflection as to where it originates and the type of lizard. The of the thigh; 25 or more preanal/femoral pores in males; fewer than word Solomoncolotes in Latin literally means “Solomon Gecko”. 40 enlarged femoral scales; enlarged preanal scale row is GENUS BOBBOTTOMCOLOTES GEN. NOV. separated by a row of short scales from short rows of enlarged Type species: Bobbottomcolotes bobbottomi sp. nov. (this paper). preanal scutes webbing between digits 3 and 4 of hind leg less Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, than 1/5 of the digit length; eye diameter is usually 41% or less Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., than the head width (J. euaensis), or: Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., 3/ More than 15 scansors under digit 4 of hind leg; more than 25 Adelynhosergecko gen. nov., Allengreercolotes gen. nov., enlarged preanal/femoral scales, extending onto distal portions of Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until the thigh; 25 or more preanal/femoral pores in males (J. now treated as being of the single genus Lepidodactylus are flaviocularis: this being the entirety of the subgenus separated from all other geckos by the following suite of Solomoncolotes subgen. nov. as presently known). characters: Digits are moderately long, more or less dilated, free or Geckos in the genus Shireenhosergecko gen. nov. are separated with a rudiment of web, inferiorly with a single or double series of from the other three genera Jackyhosergecko gen. nov., transverse lamellae divided by a median groove and across the Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one entire pad, forming the scansor, with very short and free or other of the following five suites of characters: compressed distal non-retractile clawed joint raising from the 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, extremity of the dilated and compressed digit; inner digit clawless. 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; Body covered above with granular scales, inferiorly with juxtaposed about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal or subimbricate scales. Rostral and mental shields rounded; labials and femoral pores extending to the distal end of the thigh; bearing are much larger than adjacent scales. Pupil vertical. Males with a continuous series of 27-34 pores in males; length of hind limb preanal pores. usually more than 70% of axilla-groin distance (S. mutahi), or: Lepidodactylus as defined herein are separated from the other 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., the base only between the third and fourth toes; 32-36 enlarged Adelynhosergecko gen. nov., Allengreercolotes gen. nov., preanal and femoral pores extending to the distal end of the thigh Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by in a continuous series extending over the proximal 2/3 of the thigh; having a reduced number of scansors, with the terminal as well as bearing 28-30 pores in males (S. oorti), or: a few subterminal scansors divided and a significantly depressed 3/ 37-43 mm snout-vent length in adults; digits long and slender; body and flat and broad tail, versus a subcylindrical tail lacking

lateral flanges or spines in all the other genera. or other of the following three suites of characters: This genus also consists of the entirety of the so-called group 3 of 1/ Enlarged ventral scales limited to the preanal region and species as defined by Brown and Parker (1977). proximal position of the thighs; fewer than 20 enlarged scales in Geckos in the subgenus Borealiscolotes subgen. nov. type species pore series; males with 15 or fewer preanal pores; digits are Lepidodactylus balioburius Ota and Crombie, 1989 being the only broadly dilated, dark reddish-brown dorsal colouration with orange- other subgenus within Lepidodactylus are separated from the red patches on the rear upper labials and neck (J. manni), or: nominate subgenus by the following suite of characters: less than 2/ Fewer than 15 scansors under digit 4 of the hind leg; more than 10 scansors confined to the distal half of broadly dilated and 25 enlarged preanal/femoral scales, extending onto distal portions strongly webbed digits; a strongly flattened tail with a broad flange of the thigh; 25 or more preanal/femoral pores in males; fewer than of skin; the nostril is separated by a scale from the rostral (as 40 enlarged femoral scales; enlarged preanal scale row is opposed to being in contact in specimens of the nominate separated by a row of short scales from short rows of enlarged subgenus). preanal scutes webbing between digits 3 and 4 of hind leg less Species of gecko in the genera Shireenhosergecko gen. nov., than 1/5 of the digit length; eye diameter is usually 41% or less Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and than the head width (J. euaensis), or: Martinekcolotes gen. nov. all consist of geckos formerly placed in 3/ More than 15 scansors under digit 4 of hind leg; more than 25 the so-called group 1 of species within Lepidodactylus as defined enlarged preanal/femoral scales, extending onto distal portions of by Brown and Parker (1977). Collectively they are separated from the thigh; 25 or more preanal/femoral pores in males (J. the other genera formerly included within Lepidodactylus (these flaviocularis: Subgenus Solomoncolotes subgen. nov.). being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko Geckos in the genus Martinekcolotes gen. nov. are separated from gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. the other three genera, Shireenhosergecko gen. nov., and Rosssadliercolotes gen. nov.) by having numerous (genus) Jackyhosergecko gen. nov. and Bobbottomcolotes gen. nov. by the Gekko-like undivided scansors on all digits and a subcylindrical tail. following characters: Enlarged ventral scales limited to the preanal Geckos in the genus Bobbottomcolotes gen. nov. are separated region and proximal position of the thighs; fewer than 20 enlarged from the other three genera, Shireenhosergecko gen. nov., scales in pore series; males with 15 or fewer preanal pores; digits Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the are broadly dilated, colour is pale grey-brown above, with a few following characters: More than 25 enlarged preanal/femoral rather indistinct brown spots and scattered white dots and a brown scales, extending onto distal portions of the thigh; 25 or more streak running from the nostril to the eye; lower parts white with preanal/femoral pores in males; fewer than 15 scansors under digit small brown spots (M. listeri). 4 of the hind leg: and one or other of: Distribution: Known from central New Guinea, north and south of 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ there on the mainland and also some immediately adjacent islands 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: or landmasses. 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 Etymology: Named in honour of investigative journalist Robert the length of the digits; 40 or more enlarged preanal/femoral (Bob) Bottom OAM in recognition of his services to public welfare scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. in Australia. In the mid 1980’s he did a series of reports about magnus, B. bobbottomi sp. nov., B. potens sp. nov.). corruption involving fauna officials in New South Wales. In 1991 he Geckos in the genus Shireenhosergecko gen. nov. are separated reported on Police corruption in Victoria a full twelve months before from the other three genera Jackyhosergecko gen. nov., other “mainstream” newspaper journalists dared run with the story. Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one He authored numerous best-sellers about organized crime and or other of the following five suites of characters: corruption in Australian government including the following titles. 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, His ground-breaking books include: 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; 1/ Behind the Barrier. Gareth Powell Associates, Gladesville, about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal N.S.W.: published in 1969. and femoral pores extending to the distal end of the thigh; bearing 2/ The Godfather in Australia: Organised Crime’s Australian a continuous series of 27-34 pores in males; length of hind limb Connections. A. H. and A. W. Reed,Terrey Hills, N.S.W.: published usually more than 70% of axilla-groin distance (S. mutahi), or: in 1979. 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, 3/ Without Fear or Favour. Sun Books, South Melbourne: published 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at in 1984. the base only between the third and fourth toes; 32-36 enlarged 4/ Connections: Crime Rackets and Networks of Influence Down- preanal and femoral pores extending to the distal end of the thigh Under. Sun Books, South Melbourne: published in 1985. in a continuous series extending over the proximal 2/3 of the thigh; 5/ Connections II: Crime Rackets and Networks of Influence in bearing 28-30 pores in males (S. oorti), or: Australia. Sun Books, South Melbourne: published in 1987. 3/ 37-43 mm snout-vent length in adults; digits long and slender; 6/ Shadow of Shame: How the Mafia Got Away with the Murder of moderately dilated, limited interdigital webbing; 10-12 entire fourth Donald Mackay. Sun Books, South Melbourne: published in 1988. toe scansors covering the distal 2/3 of the toe; 20-24 enlarged 7/ Bugged! : Legal Police Telephone Taps Expose the Mr Bigs of scales in pore series confined to the preanal region only or only a Australia’s Drug Trade. Sun Books, South Melbourne: published in few on the base of the thigh, bearing about 19 preanal pores in 1989. males (S. orientalis, S. petewhybrowi sp. nov.), or: 8/ Fighting Organised Crime: Triumph and Betrayal in a Lifelong 4/ 37-43 mm snout-vent length in adults; digits long and slender; Campaign. BBP, Nelson Bay, N.S.W.: published in 2009. moderately dilated, extensive interdigital webbing; 10-12 entire fourth toe scansors covering the distal 2/3 of the toe; 20-24 The commercial success and public benefit caused by the enlarged scales in pore series confined to the preanal region only publication of Bottom’s earlier books inspired Sydney, Australia- or only a few on the base of the thigh, bearing about 19 preanal based publisher Charles Pierson to commission publication of the pores in males (S. shireenhoserae sp. nov., S. dalegibbonsi sp. book Smuggled: The Underground Trade in Australia’s Wildlife nov., S. jarradbinghami sp. nov., S. robjealousi sp. nov.), or: (Hoser, 1993), which in hindsight was probably the most significant contribution to wildlife conservation in Australia’s 200 history of 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the European settlement. digits; more than 25 but fewer than 40 enlarged pre-anal-femoral scales in males, extending onto the distal portion of the thigh; Content: Bobbottomcolotes bobbottomi sp. nov. (type species); B. enlarged preanal row not separated from scutes (S. browni). crusmaculosus sp. nov.; B. magnus (Brown and Parker, 1977); B. potens sp. nov.; B. pumilus (Boulenger, 1885). Geckos in the genus Jackyhosergecko gen. nov. are separated from the other three genera, Shireenhosergecko gen. nov., GENUS MARTINEKCOLOTES GEN. NOV. Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one Type species: Gecko listeri Boulenger, 1889.

Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, from the other three genera Jackyhosergecko gen. nov., Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., or other of the following five suites of characters: Adelynhosergecko gen. nov., Allengreercolotes gen. nov., 1/ 40-55 mm snout-vent length in adults; digits moderately dilated, Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until 10-11 entire fourth toe scansors covering the distal 2/3 of the toe; now treated as being of the single genus Lepidodactylus are about 1/4 webbed between toes 3 and 4; 28-36 enlarged preanal separated from all other geckos by the following suite of and femoral pores extending to the distal end of the thigh; bearing characters: Digits are moderately long, more or less dilated, free or a continuous series of 27-34 pores in males; length of hind limb with a rudiment of web, inferiorly with a single or double series of usually more than 70% of axilla-groin distance (S. mutahi), or: transverse lamellae divided by a median groove and across the 2/ 53-57 mm snout-vent length in adults; digits moderately dilated, entire pad, forming the scansor, with very short and free 12-14 entire fourth toe scansors; 9-10 first toe scansors; webbed at compressed distal non-retractile clawed joint raising from the the base only between the third and fourth toes; 32-36 enlarged extremity of the dilated and compressed digit; inner digit clawless. preanal and femoral pores extending to the distal end of the thigh Body covered above with granular scales, inferiorly with juxtaposed in a continuous series extending over the proximal 2/3 of the thigh; or subimbricate scales. Rostral and mental shields rounded; labials bearing 28-30 pores in males (S. oorti), or: are much larger than adjacent scales. Pupil vertical. Males with 3/ 37-43 mm snout-vent length in adults; digits long and slender; preanal pores. moderately dilated, limited interdigital webbing; 10-12 entire fourth Lepidodactylus as defined herein are separated from the other toe scansors covering the distal 2/3 of the toe; 20-24 enlarged genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., scales in pore series confined to the preanal region only or only a Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., few on the base of the thigh, bearing about 19 preanal pores in Adelynhosergecko gen. nov., Allengreercolotes gen. nov., males (S. orientalis, S. petewhybrowi sp. nov.), or: Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by 4/ 37-43 mm snout-vent length in adults; digits long and slender; having a reduced number of scansors, with the terminal as well as moderately dilated, extensive interdigital webbing; 10-12 entire a few subterminal scansors divided and a significantly depressed fourth toe scansors covering the distal 2/3 of the toe; 20-24 body and flat and broad tail, versus a subcylindrical tail lacking enlarged scales in pore series confined to the preanal region only lateral flanges or spines in all the other genera. or only a few on the base of the thigh, bearing about 19 preanal This genus also consists of the entirety of the so-called group 3 of pores in males (S. shireenhoserae sp. nov., S. jarradbinghami sp. species as defined by Brown and Parker (1977). nov., S. dalegibbonsi sp. nov., S. robjealousi sp. nov.), or: Geckos in the subgenus Borealiscolotes subgen. nov. type species 5/ Webbing between toes 3 and 4 is less than 1/4 the length of the Lepidodactylus balioburius Ota and Crombie, 1989 being the only digits; more than 25 but fewer than 40 enlarged pre-anal-femoral other subgenus within Lepidodactylus are separated from the scales in males, extending onto the distal portion of the thigh; nominate subgenus by the following suite of characters: less than enlarged preanal row not separated from scutes (S. browni). 10 scansors confined to the distal half of broadly dilated and Geckos in the genus Jackyhosergecko gen. nov. are separated strongly webbed digits; a strongly flattened tail with a broad flange from the other three genera, Shireenhosergecko gen. nov., of skin; the nostril is separated by a scale from the rostral (as Bobbottomcolotes gen. nov. and Martinekcolotes gen. nov. by one opposed to being in contact in specimens of the nominate or other of the following three suites of characters: subgenus). 1/ Enlarged ventral scales limited to the preanal region and Species of gecko in the genera Shireenhosergecko gen. nov., proximal position of the thighs; fewer than 20 enlarged scales in Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and pore series; males with 15 or fewer preanal pores; digits are Martinekcolotes gen. nov. all consist of geckos formerly placed in broadly dilated, dark reddish-brown dorsal colouration with orange- the so-called group 1 of species within Lepidodactylus as defined red patches on the rear upper labials and neck (J. manni), or: by Brown and Parker (1977). Collectively they are separated from 2/ Fewer than 15 scansors under digit 4 of the hind leg; more than the other genera formerly included within Lepidodactylus (these 25 enlarged preanal/femoral scales, extending onto distal portions being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko of the thigh; 25 or more preanal/femoral pores in males; fewer than gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. 40 enlarged femoral scales; enlarged preanal scale row is and Rosssadliercolotes gen. nov.) by having numerous (genus) separated by a row of short scales from short rows of enlarged Gekko-like undivided scansors on all digits and a subcylindrical tail. preanal scutes webbing between digits 3 and 4 of hind leg less Geckos in the genus Martinekcolotes gen. nov. are separated from than 1/5 of the digit length; eye diameter is usually 41% or less the other three genera, Shireenhosergecko gen. nov., than the head width (J. euaensis), or: Jackyhosergecko gen. nov. and Bobbottomcolotes gen. nov. by the 3/ More than 15 scansors under digit 4 of hind leg; more than 25 following characters: Enlarged ventral scales limited to the preanal enlarged preanal/femoral scales, extending onto distal portions of region and proximal position of the thighs; fewer than 20 enlarged the thigh; 25 or more preanal/femoral pores in males (J. scales in pore series; males with 15 or fewer preanal pores; digits flaviocularis: Subgenus Solomoncolotes subgen. nov.). are broadly dilated, colour is pale grey-brown above, with a few Distribution: Martinekcolotes listeri (Boulenger, 1889), the entirety rather indistinct brown spots and scattered white dots and a brown of the genus as known, is known only from Christmas Island in the streak running from the nostril to the eye; lower parts white with Indian Ocean, an island controlled by the imperialist colonial small brown spots (M. listeri being monotypic for the genus). Australian government. The species may now be extinct there due Geckos in the genus Bobbottomcolotes gen. nov. are separated to the recent introduction of feral pest species there combined with from the other three genera, Shireenhosergecko gen. nov., a general indifference to wildlife conservation by Australian Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the governments and bureaucracies. following characters: More than 25 enlarged preanal/femoral Etymology: Named in honour of Maryann Martinek of Bendigo, scales, extending onto distal portions of the thigh; 25 or more Victoria in recognition of her stellar contributions to wildlife preanal/femoral pores in males; fewer than 15 scansors under digit conservation in Victoria. This includes for her courageous role in 4 of the hind leg: and one or other of: terms of exposing the misconduct of the State government wildlife 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ department, calling itself DEWLP (at the time known as DSE) in 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: 2009 for their culpability in relation to causing the Black Saturday 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 Bushfire Holocaust (9 Feb), which besides killing 172 innocent the length of the digits; 40 or more enlarged preanal/femoral Victorians, destroyed countless other people’s lives and properties scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. and wiped out many Highlands Leadbeater’s Possums magnus, B. bobbottomi sp. nov., B. potens sp. nov.). (Gymnobelideus leadbeateri martinekae Hoser, 2018) by the Geckos in the genus Shireenhosergecko gen. nov. are separated removal of their habitat.

Martinek played a critical role in exposing the biggest ever “fake oligoporus, A. pulcher and A. sloppi sp. nov.). news” story created and executed by a criminally culpable State Geckos in the genus Allengreercolotes gen. nov. are readily Wildlife Department and their rorting staff, this being the “Sam the separated from geckos in the genera Adelynhosergecko gen. nov., Koala” scam, as detailed by Hoser (2010). See also Hoser (2018). Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by Content: Martinekcolotes listeri (Boulenger, 1889) (monotypic). having 91-121 mid-body scale rows, 30 or more enlarged preanal / GENUS ADELYNHOSERGECKO GEN. NOV. femoral scales, chin secondary postmentals and less than 20% Type species: Lepidodactylus novaeguineae Brown and Parker, webbing on feet (A. allengreeri sp. nov., A. gardineri, A. guppyi, A. 1977. intermedius, A. lombocensis, A. pauldarwini sp. nov., A. shebae, A. tepukapili, A. vanuatuensis). Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Geckos in the genus Borneocolotes gen. nov. are readily Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., separated from geckos in the genera Adelynhosergecko gen. nov., Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. by Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until having the following suite of characters: Slightly depressed, but not now treated as being of the single genus Lepidodactylus are as much as seen in Lepidodactylus (sensu stricto) as defined in separated from all other geckos by the following suite of this paper as found mainly in the Philippines; nine upper labials; characters: Digits are moderately long, more or less dilated, free or nine or ten lower labials. 100-110 mid body rows, long narrow digits with a rudiment of web, inferiorly with a single or double series of with scansors underneath the distal four fifths; the terminal scansor transverse lamellae divided by a median groove and across the is entire, one subterminal scansor is wholly entire, deeply notched entire pad, forming the scansor, with very short and free or wholly divided; eight scansors beneath toe 4. Webbing absent compressed distal non-retractile clawed joint raising from the between digits 1 and 2, and slight webbing between digits 3 and 4. extremity of the dilated and compressed digit; inner digit clawless. Adult males have a continuous series of 36-38 pores in preanal Body covered above with granular scales, inferiorly with juxtaposed and femoral regions; femoral series extends almost to the distal or subimbricate scales. Rostral and mental shields rounded; labials end of the thigh; male cloacal spur is single and large, versus are much larger than adjacent scales. Pupil vertical. Males with medium in other species in the genera Adelynhosergecko gen. preanal pores. nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. of the same size; tail is subcylindrical and without lateral Lepidodactylus as defined herein are separated from the other ornamentation; dorsal body colouration is grayish brown and genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., ventrally grayish tan; there is a pair of dark spots on the Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., dorsolateral part of the basal swollen area of the tail (B. Adelynhosergecko gen. nov., Allengreercolotes gen. nov., ranauensis). Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by having a reduced number of scansors, with the terminal as well as Geckos in the genus Rosssadliercolotes gen. nov. are readily a few subterminal scansors divided and a significantly depressed separated from geckos in the genera Adelynhosergecko gen. nov., body and flat and broad tail, versus a subcylindrical tail lacking Allengreercolotes gen. nov., and Borneocolotes gen. nov. by lateral flanges or spines in all the other genera. having one or other of the following two suites of characters: This genus also consists of the entirety of the so-called group 3 of 1/ Less than 35 enlarged scales in preanal/femoral region; Sixteen species as defined by Brown and Parker (1977). or fewer scansors on toe IV; More than 25 enlarged scales in preanal/femoral region; Less than 110 scales around midbody (R. Geckos in the subgenus Borealiscolotes subgen. nov. type species paurolepis), or: Lepidodactylus balioburius Ota and Crombie, 1989 being the only other subgenus within Lepidodactylus are separated from the 2/ The unique combination of both divided terminal scansors on all nominate subgenus by the following suite of characters: less than toes (including toe 4) and a nearly completely cylindrical tail without 10 scansors confined to the distal half of broadly dilated and fringes or evidence of dorsoventral compression (R. pantai). strongly webbed digits; a strongly flattened tail with a broad flange Distribution: New Guinea, Micronesia, Admiralty Islands and of skin; the nostril is separated by a scale from the rostral (as Espiritu Santo, an island in the Vanuatu Archipelago. opposed to being in contact in specimens of the nominate Etymology: Named in honour of my eldest daughter Adelyn Hoser subgenus). of Park Orchards, Victoria, Australia in recognition of her services Species of gecko in the genera Shireenhosergecko gen. nov., to wildlife conservation over nearly 2 decades. For more detail see Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and Hoser (2013) at pages 5 and 6. Martinekcolotes gen. nov. all consist of geckos formerly placed in Content: Adelynhosergecko novaeguineae (Brown and Parker, the so-called group 1 of species within Lepidodactylus as defined 1977) (type species); A. buleli (Ineich, 2008); A. huonensis sp. by Brown and Parker (1977). Collectively they are separated from nov.; A. madangensis sp. nov.; A. oligoporus (Buden, 2007); A. the other genera formerly included within Lepidodactylus (these pulcher (Boulenger, 1885): A. sloppi sp. nov.. being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko GENUS ALLENGREERCOLOTES GEN. NOV. gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. Type species: Lepidodactylus guppyi Boulenger, 1884. and Rosssadliercolotes gen. nov.) by having numerous (genus) Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, Gekko-like undivided scansors on all digits and a subcylindrical tail. Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Geckos in the genera Adelynhosergecko gen. nov., Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. and Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Rosssadliercolotes gen. nov. all consist of geckos formerly placed Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until in the so-called group 2 of species within Lepidodactylus as now treated as being of the single genus Lepidodactylus are defined by Brown and Parker (1977). separated from all other geckos by the following suite of They can be separated from all other species formerly placed in characters: Digits are moderately long, more or less dilated, free or Lepidodactylus sensu lato by the possession of well developed with a rudiment of web, inferiorly with a single or double series of scansors on almost all the undersurface of the digits and with one transverse lamellae divided by a median groove and across the or more, usually being a few, subterminal scansors divided entire pad, forming the scansor, with very short and free medially. The subcylindrical tail lacks fringes or flanges. compressed distal non-retractile clawed joint raising from the Geckos in the genus Adelynhosergecko gen. nov. are readily extremity of the dilated and compressed digit; inner digit clawless. separated from geckos in the genera Allengreercolotes gen. nov., Body covered above with granular scales, inferiorly with juxtaposed Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by or subimbricate scales. Rostral and mental shields rounded; labials having fewer than 25 enlarged preanal/femoral pores, snout-vent are much larger than adjacent scales. Pupil vertical. Males with length in adults of under 41 mm and of moderate build (A. buleli, A. preanal pores. huonensis sp. nov., A. madangensis sp. nov., A. novaeguineae, A. Lepidodactylus as defined herein are separated from the other

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 42 Australasian Journal of Herpetology genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., ventrally grayish tan; there is a pair of dark spots on the Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., dorsolateral part of the basal swollen area of the tail (B. Adelynhosergecko gen. nov., Allengreercolotes gen. nov., ranauensis). Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by Geckos in the genus Rosssadliercolotes gen. nov. are readily having a reduced number of scansors, with the terminal as well as separated from geckos in the genera Adelynhosergecko gen. nov., a few subterminal scansors divided and a significantly depressed Allengreercolotes gen. nov., and Borneocolotes gen. nov. by body and flat and broad tail, versus a subcylindrical tail lacking having one or other of the following two suites of characters: lateral flanges or spines in all the other genera. 1/ Less than 35 enlarged scales in preanal/femoral region; Sixteen This genus also consists of the entirety of the so-called group 3 of or fewer scansors on toe IV; More than 25 enlarged scales in species as defined by Brown and Parker (1977). preanal/femoral region; Less than 110 scales around midbody (R. Geckos in the subgenus Borealiscolotes subgen. nov. type species paurolepis), or: Lepidodactylus balioburius Ota and Crombie, 1989 being the only 2/ The unique combination of both divided terminal scansors on all other subgenus within Lepidodactylus are separated from the toes (including toe 4) and a nearly completely cylindrical tail without nominate subgenus by the following suite of characters: less than fringes or evidence of dorsoventral compression (R. pantai). 10 scansors confined to the distal half of broadly dilated and Distribution: Species within the genus Allengreercolotes gen. nov. strongly webbed digits; a strongly flattened tail with a broad flange occur in islands north and west of New Guinea including the of skin; the nostril is separated by a scale from the rostral (as Solomon Islands, Bismark Archipelago, Admiralty Islands, Vanuatu opposed to being in contact in specimens of the nominate Islands, Rotuma Island, Tuvalu and also the Lesser Sunda Islands. subgenus). Content: Allengreercolotes guppyi (Boulenger, 1884) (type Species of gecko in the genera Shireenhosergecko gen. nov., species); A. allengreeri sp. nov.; A. gardineri (Boulenger, 1897); A. Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and intermedius (Darevsky, 1964); A. lombocensis (Mertens, 1929); A. Martinekcolotes gen. nov. all consist of geckos formerly placed in pauldarwini sp. nov.; A. shebae (Brown and Tanner, 1949); A. the so-called group 1 of species within Lepidodactylus as defined tepukapili (Zug, Waitling, Alefaio, Alefaio and Ludescher, 2003); A. by Brown and Parker (1977). Collectively they are separated from vanuatuensis (Ota, Fisher, Ineich, Case, Radtkey and Zug, 1998). the other genera formerly included within Lepidodactylus (these Etymology: Named in honour of Dr. Allen E. Greer who for many being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko years was herpetology curator at the Australian Museum in gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. Sydney, New South Wales, Australia, for services to herpetology and Rosssadliercolotes gen. nov.) by having numerous (genus) well beyond the duties of his paid position. The name Gekko-like undivided scansors on all digits and a subcylindrical tail. Allengreercolotes means Allengreer Gecko in Latin. Geckos in the genera Adelynhosergecko gen. nov., GENUS BORNEOCOLOTES GEN. NOV. Allengreercolotes gen. nov., Borneocolotes gen. nov. and Type species: Lepidodactylus ranauensis Ota and Hikida, 1988. Rosssadliercolotes gen. nov. all consist of geckos formerly placed in the so-called group 2 of species within Lepidodactylus as Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, defined by Brown and Parker (1977). Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., They can be separated from all other species formerly placed in Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Lepidodactylus sensu lato by the possession of well developed Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until scansors on almost all the undersurface of the digits and with one now treated as being of the single genus Lepidodactylus are or more, usually being a few, subterminal scansors divided separated from all other geckos by the following suite of medially. The subcylindrical tail lacks fringes or flanges. characters: Digits are moderately long, more or less dilated, free or Geckos in the genus Allengreercolotes gen. nov. are readily with a rudiment of web, inferiorly with a single or double series of separated from geckos in the genera Adelynhosergecko gen. nov., transverse lamellae divided by a median groove and across the Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by entire pad, forming the scansor, with very short and free having 91-121 mid-body scale rows, 30 or more enlarged preanal / compressed distal non-retractile clawed joint raising from the femoral scales, chin secondary postmentals and less than 20% extremity of the dilated and compressed digit; inner digit clawless. webbing on feet (A. allengreeri sp. nov., A. gardineri, A. guppyi, A. Body covered above with granular scales, inferiorly with juxtaposed intermedius, A. lombocensis, A. pauldarwini sp. nov., A. shebae, A. or subimbricate scales. Rostral and mental shields rounded; labials tepukapili, A. vanuatuensis). are much larger than adjacent scales. Pupil vertical. Males with Geckos in the genus Adelynhosergecko gen. nov. are readily preanal pores. separated from geckos in the genera Allengreercolotes gen. nov., Lepidodactylus as defined herein are separated from the other Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., having fewer than 25 enlarged preanal/femoral pores, snout-vent Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., length in adults of under 41 mm and of moderate build (A. buleli, A. Adelynhosergecko gen. nov., Allengreercolotes gen. nov., huonensis sp. nov., A. madangensis sp. nov., A. novaeguineae, A. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by oligoporus, A. pulcher and A. sloppi sp. nov.). having a reduced number of scansors, with the terminal as well as Geckos in the genus Borneocolotes gen. nov. are readily a few subterminal scansors divided and a significantly depressed separated from geckos in the genera Adelynhosergecko gen. nov., body and flat and broad tail, versus a subcylindrical tail lacking Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. by lateral flanges or spines in all the other genera. having the following suite of characters: Slightly depressed, but not This genus also consists of the entirety of the so-called group 3 of as much as seen in Lepidodactylus (sensu stricto) as defined in species as defined by Brown and Parker (1977). this paper as found mainly in the Philippines; nine upper labials; Geckos in the subgenus Borealiscolotes subgen. nov. type species nine or ten lower labials. 100-110 mid body rows, long narrow digits Lepidodactylus balioburius Ota and Crombie, 1989 being the only with scansors underneath the distal four fifths; the terminal scansor other subgenus within Lepidodactylus are separated from the is entire, one subterminal scansor is wholly entire, deeply notched nominate subgenus by the following suite of characters: less than or wholly divided; eight scansors beneath toe 4. Webbing absent 10 scansors confined to the distal half of broadly dilated and between digits 1 and 2, and slight webbing between digits 3 and 4. strongly webbed digits; a strongly flattened tail with a broad flange Adult males have a continuous series of 36-38 pores in preanal of skin; the nostril is separated by a scale from the rostral (as and femoral regions; femoral series extends almost to the distal opposed to being in contact in specimens of the nominate end of the thigh; male cloacal spur is single and large, versus subgenus). medium in other species in the genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. Species of gecko in the genera Shireenhosergecko gen. nov., of the same size; tail is subcylindrical and without lateral Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and ornamentation; dorsal body colouration is grayish brown and Martinekcolotes gen. nov. all consist of geckos formerly placed in Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 43

the so-called group 1 of species within Lepidodactylus as defined Diagnosis: Geckos of the genera Lepidodactylus Fitzinger, 1843, by Brown and Parker (1977). Collectively they are separated from Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., the other genera formerly included within Lepidodactylus (these Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko Adelynhosergecko gen. nov., Allengreercolotes gen. nov., gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. all until and Rosssadliercolotes gen. nov.) by having numerous (genus) now treated as being of the single genus Lepidodactylus are Gekko-like undivided scansors on all digits and a subcylindrical tail. separated from all other geckos by the following suite of Geckos in the genera Adelynhosergecko gen. nov., characters: Digits are moderately long, more or less dilated, free or Allengreercolotes gen. nov., Borneocolotes gen. nov. and with a rudiment of web, inferiorly with a single or double series of Rosssadliercolotes gen. nov. all consist of geckos formerly placed transverse lamellae divided by a median groove and across the in the so-called group 2 of species within Lepidodactylus as entire pad, forming the scansor, with very short and free defined by Brown and Parker (1977). compressed distal non-retractile clawed joint raising from the They can be separated from all other species formerly placed in extremity of the dilated and compressed digit; inner digit clawless. Lepidodactylus sensu lato by the possession of well developed Body covered above with granular scales, inferiorly with juxtaposed scansors on almost all the undersurface of the digits and with one or subimbricate scales. Rostral and mental shields rounded; labials or more, usually being a few, subterminal scansors divided are much larger than adjacent scales. Pupil vertical. Males with medially. The subcylindrical tail lacks fringes or flanges. preanal pores. Geckos in the genus Borneocolotes gen. nov. are readily Lepidodactylus as defined herein are separated from the other separated from geckos in the genera Adelynhosergecko gen. nov., genera Shireenhosergecko gen. nov., Jackyhosergecko gen. nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. by Bobbottomcolotes gen. nov., Martinekcolotes gen. nov., having the following suite of characters: Slightly depressed, but not Adelynhosergecko gen. nov., Allengreercolotes gen. nov., as much as seen in Lepidodactylus (sensu stricto) as defined in Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by this paper as found mainly in the Philippines; nine upper labials; having a reduced number of scansors, with the terminal as well as nine or ten lower labials. 100-110 mid body rows, long narrow digits a few subterminal scansors divided and a significantly depressed with scansors underneath the distal four fifths; the terminal scansor body and flat and broad tail, versus a subcylindrical tail lacking is entire, one subterminal scansor is wholly entire, deeply notched lateral flanges or spines in all the other genera. or wholly divided; eight scansors beneath toe 4. Webbing absent This genus also consists of the entirety of the so-called group 3 of between digits 1 and 2, and slight webbing between digits 3 and 4. species as defined by Brown and Parker (1977). Adult males have a continuous series of 36-38 pores in preanal Geckos in the subgenus Borealiscolotes subgen. nov. type species and femoral regions; femoral series extends almost to the distal Lepidodactylus balioburius Ota and Crombie, 1989 being the only end of the thigh; male cloacal spur is single and large, versus other subgenus within Lepidodactylus are separated from the medium in other species in the genera Adelynhosergecko gen. nominate subgenus by the following suite of characters: less than nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. 10 scansors confined to the distal half of broadly dilated and of the same size; tail is subcylindrical and without lateral strongly webbed digits; a strongly flattened tail with a broad flange ornamentation; dorsal body colouration is grayish brown and of skin; the nostril is separated by a scale from the rostral (as ventrally grayish tan; there is a pair of dark spots on the opposed to being in contact in specimens of the nominate dorsolateral part of the basal swollen area of the tail (B. subgenus). ranauensis). Species of gecko in the genera Shireenhosergecko gen. nov., Geckos in the genus Allengreercolotes gen. nov. are readily Jackyhosergecko gen. nov., Bobbottomcolotes gen. nov. and separated from geckos in the genera Adelynhosergecko gen. nov., Martinekcolotes gen. nov. all consist of geckos formerly placed in Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by the so-called group 1 of species within Lepidodactylus as defined having 91-121 mid-body scale rows, 30 or more enlarged preanal / by Brown and Parker (1977). Collectively they are separated from femoral scales, chin secondary postmentals and less than 20% the other genera formerly included within Lepidodactylus (these webbing on feet (A. allengreeri sp. nov., A. gardineri, A. guppyi, A. being genera Lepidodactylus Fitzinger, 1843, Adelynhosergecko intermedius, A. lombocensis, A. pauldarwini sp. nov., A. shebae, A. gen. nov., Allengreercolotes gen. nov., Borneocolotes gen. nov. tepukapili, A. vanuatuensis). and Rosssadliercolotes gen. nov.) by having numerous (genus) Geckos in the genus Adelynhosergecko gen. nov. are readily Gekko-like undivided scansors on all digits and a subcylindrical tail. separated from geckos in the genera Allengreercolotes gen. nov., Geckos in the genera Adelynhosergecko gen. nov., Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by Allengreercolotes gen. nov., Borneocolotes gen. nov. and having fewer than 25 enlarged preanal/femoral pores, snout-vent Rosssadliercolotes gen. nov. all consist of geckos formerly placed length in adults of under 41 mm and of moderate build (A. buleli, A. in the so-called group 2 of species within Lepidodactylus as huonensis sp. nov., A. madangensis sp. nov., A. novaeguineae, A. defined by Brown and Parker (1977). oligoporus, A. pulcher and A. sloppi sp. nov.). They can be separated from all other species formerly placed in Geckos in the genus Rosssadliercolotes gen. nov. are readily Lepidodactylus sensu lato by the possession of well developed separated from geckos in the genera Adelynhosergecko gen. nov., scansors on almost all the undersurface of the digits and with one Allengreercolotes gen. nov., and Borneocolotes gen. nov. by or more, usually being a few, subterminal scansors divided having one or other of the following two suites of characters: medially. The subcylindrical tail lacks fringes or flanges. 1/ Less than 35 enlarged scales in preanal/femoral region; Sixteen Geckos in the genus Rosssadliercolotes gen. nov. are readily or fewer scansors on toe IV; More than 25 enlarged scales in separated from geckos in the genera Adelynhosergecko gen. nov., preanal/femoral region; Less than 110 scales around midbody (R. Allengreercolotes gen. nov., and Borneocolotes gen. nov. by paurolepis), or: having one or other of the following two suites of characters: 2/ The unique combination of both divided terminal scansors on all 1/ Less than 35 enlarged scales in preanal/femoral region; Sixteen toes (including toe 4) and a nearly completely cylindrical tail without or fewer scansors on toe IV; More than 25 enlarged scales in fringes or evidence of dorsoventral compression (R. pantai). preanal/femoral region; Less than 110 scales around midbody (R. Distribution: Known only from the type locality of Ranau, Sabah, paurolepis), or: Malaysia, Borneo. 2/ The unique combination of both divided terminal scansors on all Content: Borneocolotes ranauensis (Ota and Hikida, 1988) (type toes (including toe 4) and a nearly completely cylindrical tail without species). fringes or evidence of dorsoventral compression (R. pantai). GENUS ROSSSADLIERCOLOTES GEN. NOV. Geckos in the genus Borneocolotes gen. nov. are readily Type species: Lepidodactylus paurolepis Ota, Fisher, Ineich and separated from geckos in the genera Adelynhosergecko gen. nov., Case, 1995. Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. by

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 44 Australasian Journal of Herpetology having the following suite of characters: Slightly depressed, but not between that seen in members of the genera as much as seen in Lepidodactylus (sensu stricto) as defined in Charlespiersoncolotes gen. nov.and Luperosaurus, most notably this paper as found mainly in the Philippines; nine upper labials; weakly spinose dorsal scales and it is placed in the genus nine or ten lower labials. 100-110 mid body rows, long narrow digits Luperosaurus, even though no other members of the genus with scansors underneath the distal four fifths; the terminal scansor Luperosaurus have spinose dorsal scales of any sort. It is likely it is entire, one subterminal scansor is wholly entire, deeply notched may need to be eventually assigned to a separate genus or or wholly divided; eight scansors beneath toe 4. Webbing absent subgenus. between digits 1 and 2, and slight webbing between digits 3 and 4. Georgemarioliscolotes gen. nov. is readily separated from Adult males have a continuous series of 36-38 pores in preanal Scelotretus Fitzinger, 1843 by having a small round to ovoid and femoral regions; femoral series extends almost to the distal auricular opening, versus a narrow elliptical or vertical slit-like end of the thigh; male cloacal spur is single and large, versus opening in Scelotretus Fitzinger, 1843. medium in other species in the genera Adelynhosergecko gen. Georgemarioliscolotes gen. nov. is further separated from nov., Allengreercolotes gen. nov., and Rosssadliercolotes gen. nov. Scelotretus Fitzinger, 1843 by having of the same size; tail is subcylindrical and without lateral 11-15 supralabials, versus 16 in Scelotretus Fitzinger, 1843 and ornamentation; dorsal body colouration is grayish brown and 10-14 infralabials, versus 15-18 in Scelotretus Fitzinger, 1843. ventrally grayish tan; there is a pair of dark spots on the Georgemarioliscolotes gen. nov. has round-hexagonal, flat convex dorsolateral part of the basal swollen area of the tail (B. dorsal body scales, versus hexagonal flat dorsal body scales in ranauensis). Scelotretus Fitzinger, 1843. Geckos in the genus Allengreercolotes gen. nov. are readily Georgemarioliscolotes gen. nov. has flat or convex dorsal body separated from geckos in the genera Adelynhosergecko gen. nov., tubercles, versus flat only in Scelotretus Fitzinger, 1843. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by having 91-121 mid-body scale rows, 30 or more enlarged preanal / Georgemarioliscolotes gen. nov. has 28-40 preanofemorals versus femoral scales, chin secondary postmentals and less than 20% 12 or less in Scelotretus Fitzinger, 1843, 11-13 scansors on toe 1, webbing on feet (A. allengreeri sp. nov., A. gardineri, A. guppyi, A. versus 10 in Scelotretus Fitzinger, 1843 and small anteriormost intermedius, A. lombocensis, A. pauldarwini sp. nov., A. shebae, A. chinshields, versus slightly enlarged in Scelotretus Fitzinger, 1843. tepukapili, A. vanuatuensis). Distribution: Philippines and adjacent islands and one species from Borneo. Geckos in the genus Adelynhosergecko gen. nov. are readily separated from geckos in the genera Allengreercolotes gen. nov., Content: Luperosaurus cumingii Gray, 1845 (type species); L. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by angliit Brown, Diesmos and Oliveros, 2011; L. corfieldi Gaulke, having fewer than 25 enlarged preanal/femoral pores, snout-vent Rösler and Brown, 2007; L. kubli Brown, Diesmos and Duya, 2007; length in adults of under 41 mm and of moderate build (A. buleli, A. L. macgregori Stejneger, 1907; L. palawanensis Brown and Alcala, huonensis sp. nov., A. madangensis sp. nov., A. novaeguineae, A. 1978; L. sorok Das, Lakim and Kandaung, 2008. oligoporus, A. pulcher and A. sloppi sp. nov.). CHARLESPIERSONCOLOTES GEN. NOV. Distribution: Known only from Palau and Maluku islands Type species: Luperosaurus joloensis Taylor, 1918. Content: Rosssadliercolotes paurolepis (Ota, Fisher, Ineich and Diagnosis: The genus Luperosaurus Gray, 1845 as recognized to Case, 1995) (type species); R. pantai (Stubbs, Karin, Arifin, date is a paraphyletic assemblage of morphologically similar Iskandar, Arida, Reilly, Bloch, Kusnadi and McGuire, 2017). species. Therefore it is split herein into four genera. GENUS LUPEROSAURUS GRAY, 1845 All four genera, formerly included in Luperosaurus, namely Type species: Luperosaurus cumingii Gray, 1845. Luperosaurus, Charlespiersoncolotes gen. nov. (described here), Scelotretus Fitzinger, 1843 and Georgemarioliscolotes gen. nov. Diagnosis: The genus Luperosaurus Gray, 1845 as recognized to are readily separated from all other geckos by the following suite of date is a paraphyletic assemblage of morphologically similar characters: Digits strongly dilated, half webbed, (excluding species. Therefore it is split herein into four genera. subgenus Haroldcoggercolotes subgen. nov. a subgenus of All four genera, formerly included in Luperosaurus, namely Scelotretus Fitzinger, 1843 which has only slight webbing between Luperosaurus, Charlespiersoncolotes gen. nov., Scelotretus the toes), with undivided, angularly curved lamellae below; all but Fitzinger, 1843 and Georgemarioliscolotes gen. nov. are readily thumb and inner toe with a very short, compressed, distal phalanx, separated from all other geckos by the following suite of with retractile claw ; legs bordered with cutaneous lobes; upper characters: Digits strongly dilated, half webbed, (excluding and lower surfaces covered with juxtaposed granular scales; pupil subgenus Haroldcoggercolotes subgen. nov. a subgenus of vertical; males with preanal pores. Scelotretus Fitzinger, 1843 which has only slight webbing between The genera Scelotretus Fitzinger, 1843 and Georgemarioliscolotes the toes), with undivided, angularly curved lamellae below; all but gen. nov. are readily separated from the other two genera thumb and inner toe with a very short, compressed, distal phalanx, (Luperosaurus and Charlespiersoncolotes gen. nov.) by the with retractile claw ; legs bordered with cutaneous lobes; upper presence of a distinctly elongate head, elongate versus robust and lower surfaces covered with juxtaposed granular scales; pupil body shape (in the other genera) and the presence of enlarged vertical; males with preanal pores. interstitial granules. The genera Scelotretus Fitzinger, 1843 and Georgemarioliscolotes The genera Luperosaurus and Charlespiersoncolotes gen. nov. are gen. nov. are readily separated from the other two genera separated from Scelotretus Fitzinger, 1843 and (Luperosaurus and Charlespiersoncolotes gen. nov.) by the Georgemarioliscolotes gen. nov. by the presence of beadlike, presence of a distinctly elongate head, elongate versus robust granular dorsals, a stout and robust, stout body and deeply body shape (in the other genera) and the presence of enlarged notched to divided penultimate subdigital scansors. interstitial granules. The genus Charlespiersoncolotes gen. nov. is most easily The genera Luperosaurus and Charlespiersoncolotes gen. nov. are separated from the genus Luperosaurus (as well as Scelotretus separated from Scelotretus Fitzinger, 1843 and Fitzinger, 1843 and Georgemarioliscolotes gen. nov.) by the Georgemarioliscolotes gen. nov. by the presence of beadlike, presence of strongly spinose dorsal tubercles. granular dorsals, a stout and robust, stout body and deeply notched to divided penultimate subdigital scansors. The species originally described as Luperosaurus palawanensis Brown and Alcala, 1978 has many characteristics intermediate The genus Charlespiersoncolotes gen. nov. is most easily between that seen in members of the genera separated from the genus Luperosaurus (as well as Scelotretus Charlespiersoncolotes gen. nov.and Luperosaurus, most notably Fitzinger, 1843 and Georgemarioliscolotes gen. nov.) by the weakly spinose dorsal scales and it is placed in the genus presence of strongly spinose dorsal tubercles. Luperosaurus, even though no other members of the genus The species originally described as Luperosaurus palawanensis Luperosaurus have spinose dorsal scales of any sort. It is likely it Brown and Alcala, 1978 has many characteristics intermediate may need to be assigned to a separate genus or subgenus. Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 45

Georgemarioliscolotes gen. nov. is readily separated from in Scelotretus Fitzinger, 1843 and 10-14 infralabials, versus 15-18 Scelotretus Fitzinger, 1843 by having a small round to ovoid in Scelotretus Fitzinger, 1843. auricular opening, versus a narrow elliptical or vertical slit-like Georgemarioliscolotes gen. nov. has round-hexagonal, flat convex opening in Scelotretus Fitzinger, 1843. dorsal body scales, versus hexagonal flat dorsal body scales in Georgemarioliscolotes gen. nov. is further separated from Scelotretus Fitzinger, 1843. Scelotretus Fitzinger, 1843 by having Georgemarioliscolotes gen. nov. has flat or convex dorsal body 11-15 supralabials, versus 16 in Scelotretus Fitzinger, 1843 and tubercles, versus flat only in Scelotretus Fitzinger, 1843. 10-14 infralabials, versus 15-18 in Scelotretus Fitzinger, 1843. Georgemarioliscolotes gen. nov. has 28-40 preanofemorals versus Georgemarioliscolotes gen. nov. has round-hexagonal, flat convex 12 or less in Scelotretus Fitzinger, 1843, 11-13 scansors on toe 1, dorsal body scales, versus hexagonal flat dorsal body scales in versus 10 in Scelotretus Fitzinger, 1843 and small anteriormost Scelotretus Fitzinger, 1843. chinshields, versus slightly enlarged in Scelotretus Fitzinger, 1843. Georgemarioliscolotes gen. nov. has flat or convex dorsal body The subgenus Scelotretus is further defined and separated from tubercles, versus flat only in Scelotretus Fitzinger, 1843. the subgenus Haroldcoggercolotes subgen. nov. and all other Georgemarioliscolotes gen. nov. has 28-40 preanofemorals versus geckos by the following suite of characters: Maximum SVL 140.0 12 or less in Scelotretus Fitzinger, 1843, 11-13 scansors on toe 1, mm; nares in contact with rostral; nasals 3-4; postmentals versus 10 in Scelotretus Fitzinger, 1843 and small anteriormost relatively small; dorsal tubercle rows 12-14; precloacal pores 14- chinshields, versus slightly enlarged in Scelotretus Fitzinger, 1843. 58; postcloacal tubercles 1-3; no webbing between fingers and Distribution: Philippines (Sulu Archipelago, Jolo Island, toes; fore and hind limbs with tubercles; lateral folds with tubercles; Mindanao) (C. joloensis) and Borneo (C. yasumai). subcaudals not enlarged; head unicolored, without pattern; Etymology: Named in honour of publisher Charles Pierson as nominate form with white, anteriorly bifurcated dorsal stripe (Rösler detailed in Hoser (2012) pages 67-68. et al. 2011). Content: Charlespiersoncolotes joloensis (Taylor, 1918) (type Distribution: Sulawesi and Palawan (subgenus species); C. yasumai (Ota, Sengoku and Hikida, 1996). Haroldcoggercolotes subgen. nov.), Indonesia, extending to the Solomon Islands and the Vanuatu Islands (subgenus Scelotretus GENUS SCELOTRETUS FITZINGER, 1843. Fitzinger, 1843). Type species: Gekko vittatus Houttuyn, 1782. Content: Scelotretus vittatus (Houttuyn, 1782) (type species): S. Diagnosis: The genus Luperosaurus Gray, 1845 as recognized to haroldcoggeri sp. nov.; S. daranini sp. nov.; S. gulat (Brown, date is a paraphyletic assemblage of morphologically similar Diesmos, Duya, Garcia and Rico, 2010); S. iskandari (Brown, species. Therefore it is split herein into four genera. Supriatna and Ota, 2000); S. jenandersonae sp. nov.; S. remotus All four genera, formerly included in Luperosaurus, namely (Rösler, Ineich, Wilms and Bo¨hme, 2012). Luperosaurus, Charlespiersoncolotes gen. nov., Scelotretus SUBGENUS HAROLDCOGGERCOLOTES SUBGEN. NOV. Fitzinger, 1843 (described here) and Georgemarioliscolotes gen. Type species: Luperosaurus iskandari Brown, Supriatna and Ota, nov. are readily separated from all other geckos by the following 2000. suite of characters: Digits strongly dilated, half webbed (excluding subgenus Haroldcoggercolotes subgen. nov. a subgenus of Diagnosis: The subgenus Haroldcoggercolotes subgen. nov. a Scelotretus Fitzinger, 1843 which has only slight webbing between subgenus of Scelotretus Fitzinger, 1843 is separated from the the toes or none), with undivided, angularly curved lamellae below; nominate subgenus Scelotretus by having only slight webbing all but thumb and inner toe with a very short, compressed, distal between the toes or none, versus half webbed in Scelotretus. phalanx, with retractile claw ; legs bordered with cutaneous lobes; The genus Luperosaurus Gray, 1845 as recognized to date is a upper and lower surfaces covered with juxtaposed granular scales; paraphyletic assemblage of morphologically similar species. pupil vertical; males with preanal pores. Therefore it is split herein into four genera. The genera Scelotretus Fitzinger, 1843 and Georgemarioliscolotes All four genera, formerly included in Luperosaurus, namely gen. nov. are readily separated from the other two genera Luperosaurus, Charlespiersoncolotes gen. nov., Scelotretus (Luperosaurus and Charlespiersoncolotes gen. nov.) by the Fitzinger, 1843 and Georgemarioliscolotes gen. nov. (described presence of a distinctly elongate head, elongate versus robust here) are readily separated from all other geckos by the following body shape (in the other genera) and the presence of enlarged suite of characters: Digits strongly dilated, half webbed, (excluding interstitial granules. subgenus Haroldcoggercolotes subgen. nov. a subgenus of The genera Luperosaurus and Charlespiersoncolotes gen. nov. are Scelotretus Fitzinger, 1843 which has only slight webbing between separated from Scelotretus Fitzinger, 1843 (described here) and the toes or none), with undivided, angularly curved lamellae below; Georgemarioliscolotes gen. nov. by the presence of beadlike, all but thumb and inner toe with a very short, compressed, distal granular dorsals, a stout and robust, stout body and deeply phalanx, with retractile claw ; legs bordered with cutaneous lobes; notched to divided penultimate subdigital scansors. upper and lower surfaces covered with juxtaposed granular scales; pupil vertical; males with preanal pores. The genus Charlespiersoncolotes gen. nov. is most easily separated from the genus Luperosaurus (as well as Scelotretus The genera Scelotretus Fitzinger, 1843 and Georgemarioliscolotes Fitzinger, 1843 and Georgemarioliscolotes gen. nov.) by the gen. nov. are readily separated from the other two genera presence of strongly spinose dorsal tubercles. (Luperosaurus and Charlespiersoncolotes gen. nov.) by the presence of a distinctly elongate head, elongate versus robust The species originally described as Luperosaurus palawanensis body shape (in the other genera) and the presence of enlarged Brown and Alcala, 1978 has many characteristics intermediate interstitial granules. between that seen in members of the genera Charlespiersoncolotes gen. nov.and Luperosaurus, most notably The genera Luperosaurus and Charlespiersoncolotes gen. nov. are weakly spinose dorsal scales and it is placed in the genus separated from Scelotretus Fitzinger, 1843 and Luperosaurus, even though no other members of the genus Georgemarioliscolotes gen. nov. by the presence of beadlike, Luperosaurus have spinose dorsal scales of any sort. It is likely it granular dorsals, a stout and robust, stout body and deeply may need to be eventually assigned to a separate genus or notched to divided penultimate subdigital scansors. subgenus. The genus Charlespiersoncolotes gen. nov. is most easily Georgemarioliscolotes gen. nov. is readily separated from separated from the genus Luperosaurus (as well as Scelotretus Scelotretus Fitzinger, 1843 by having a small round to ovoid Fitzinger, 1843 and Georgemarioliscolotes gen. nov.) by the auricular opening, versus a narrow elliptical or vertical slit-like presence of strongly spinose dorsal tubercles. opening in Scelotretus Fitzinger, 1843. The species originally described as Luperosaurus palawanensis Georgemarioliscolotes gen. nov. is further separated from Brown and Alcala, 1978 has many characteristics intermediate Scelotretus Fitzinger, 1843 by having 11-15 supralabials, versus 16 between that seen in members of the genera Charlespiersoncolotes gen. nov.and Luperosaurus, most notably Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 46 Australasian Journal of Herpetology weakly spinose dorsal scales and it is placed in the genus separated from the genus Luperosaurus (as well as Scelotretus Luperosaurus, even though no other members of the genus Fitzinger, 1843 and Georgemarioliscolotes gen. nov.) by the Luperosaurus have spinose dorsal scales of any sort. It is likely it presence of strongly spinose dorsal tubercles. may need to be eventually assigned to a separate genus or The species originally described as Luperosaurus palawanensis subgenus. Brown and Alcala, 1978 has many characteristics intermediate Georgemarioliscolotes gen. nov. is readily separated from between that seen in members of the genera Scelotretus Fitzinger, 1843 by having a small round to ovoid Charlespiersoncolotes gen. nov.and Luperosaurus, most notably auricular opening, versus a narrow elliptical or vertical slit-like weakly spinose dorsal scales and it is placed in the genus opening in Scelotretus Fitzinger, 1843. Luperosaurus, even though no other members of the genus Georgemarioliscolotes gen. nov. is further separated from Luperosaurus have spinose dorsal scales of any sort. It is likely it Scelotretus Fitzinger, 1843 by having 11-15 supralabials, versus 16 may need to be eventually assigned to a separate genus or in Scelotretus Fitzinger, 1843 and 10-14 infralabials, versus 15-18 subgenus. in Scelotretus Fitzinger, 1843. Georgemarioliscolotes gen. nov. is readily separated from Georgemarioliscolotes gen. nov. has round-hexagonal, flat convex Scelotretus Fitzinger, 1843 by having a small round to ovoid dorsal body scales, versus hexagonal flat dorsal body scales in auricular opening, versus a narrow elliptical or vertical slit-like Scelotretus Fitzinger, 1843. opening in Scelotretus Fitzinger, 1843. Georgemarioliscolotes gen. nov. has flat or convex dorsal body Georgemarioliscolotes gen. nov. is further separated from tubercles, versus flat only in Scelotretus Fitzinger, 1843. Scelotretus Fitzinger, 1843 by having 11-15 supralabials, versus 16 Georgemarioliscolotes gen. nov. has 28-40 preanofemorals versus in Scelotretus Fitzinger, 1843 and 10-14 infralabials, versus 15-18 12 or less in Scelotretus Fitzinger, 1843, 11-13 scansors on toe 1, in Scelotretus Fitzinger, 1843. versus 10 in Scelotretus Fitzinger, 1843 and small anteriormost Georgemarioliscolotes gen. nov. has round-hexagonal, flat convex chinshields, versus slightly enlarged in Scelotretus Fitzinger, 1843. dorsal body scales, versus hexagonal flat dorsal body scales in The subgenus Scelotretus is further defined and separated from Scelotretus Fitzinger, 1843. the subgenus Haroldcoggercolotes subgen. nov. and all other Georgemarioliscolotes gen. nov. has flat or convex dorsal body geckos by the following suite of characters: Maximum SVL 140.0 tubercles, versus flat only in Scelotretus Fitzinger, 1843. mm; nares in contact with rostral; nasals 3-4; postmentals Georgemarioliscolotes gen. nov. has 28-40 preanofemorals versus relatively small; dorsal tubercle rows 12-14; precloacal pores 14- 12 or less in Scelotretus Fitzinger, 1843, 11-13 scansors on toe 1, 58; postcloacal tubercles 1-3; no webbing between fingers and versus 10 in Scelotretus Fitzinger, 1843 and small anteriormost toes; fore and hind limbs with tubercles; lateral folds with tubercles; chinshields, versus slightly enlarged in Scelotretus Fitzinger, 1843. subcaudals not enlarged; head unicolored, without pattern; Distribution: Sumatra (G. brooksii) and West Malaysia (G. nominate form with white, anteriorly bifurcated dorsal stripe (Rösler browni). et al. 2011). Content: Georgemarioliscolotes brooksii (Boulenger, 1920) (type Distribution: Sulawesi and Palawan (subgenus species); G. browni (Russell, 1979). Haroldcoggercolotes subgen. nov.). GENUS PSEUDOGEKKO TAYLOR, 1922 Etymology: Named in honour of Dr. Harold G. Cogger, former Type species: Luperosaurus compresicorpus Taylor, 1915. deputy director and curator of herpetology at the Australian The genus Pseudogekko Taylor, 1922 is separated from all other Museum in Sydney, New South Wales, Australia in recognition of a geckos by the following suite of characters: Digits entirely dilated lifetime’s high achievement in herpetology. with infradigital lamellae extending to the base forming the scansor, Content: Scelotretus (Haroldcoggercolotes) gulat (Brown, those on tip divided; digits except inner with sessile retractile claw; Diesmos, Duya, Garcia and Rico, 2010); S. (Haroldcoggercolotes) a slight web between digits; no lateral skin fold; no enlarged chin iskandari (Brown, Supriatna and Ota, 2000); S. shields; pupil vertical. Body compressed, with a narrow abdominal (Haroldcoggercolotes) jenandersonae sp. nov.. region; 10-16 pre cloacal pores present in males in a single row. GEORGEMARIOLISCOLOTES GEN. NOV. Distribution: Philippines. Type species: Gekko brooksi Boulenger, 1920. Content: Pseudogekko compresicorpus (Taylor, 1915) (type Diagnosis: The genus Luperosaurus Gray, 1845 as recognized to species); P. atiorum Davis, Watters, Köhler, Whitsett, Huron, date is a paraphyletic assemblage of morphologically similar Brown, Diesmos and Siler, 2015; P. brevipes (Boettger, 1897); P. species. Therefore it is split herein into four genera. chavacano Siler, Welton, David, Watters, Davey, Diesmos, All four genera, formerly included in Luperosaurus, namely Diesmos and Brown, 2014; P. ditoy Siler, Welton, David, Watters, Luperosaurus, Charlespiersoncolotes gen. nov., Scelotretus Davey, Diesmos, Diesmos and Brown, 2014; P. isapa Siler, Davis, Fitzinger, 1843 and Georgemarioliscolotes gen. nov. (described Diesmos, Guinto, Whitsett and Brown, 2016; P. pungkaypinit Siler, here) are readily separated from all other geckos by the following Welton, David, Watters, Davey, Diesmos, Diesmos and Brown, suite of characters: Digits strongly dilated, half webbed, (excluding 2014; P. smaragdinus (Taylor, 1922); P. sumiklab Siler, Davis, subgenus Haroldcoggercolotes subgen. nov. a subgenus of Watters, Freitas, Griffith, Binday, Lobos, Amarga and Brown, 2017. Scelotretus Fitzinger, 1843 which has only slight webbing between SUBGENUS ROBWATSONCOLOTES SUBGEN. NOV. the toes), with undivided, angularly curved lamellae below; all but Type species: Lepidodactylus brevipes Boettger 1897, now known thumb and inner toe with a very short, compressed, distal phalanx, as Pseudogekko brevipes (Boettger, 1867). with retractile claw ; legs bordered with cutaneous lobes; upper Diagnosis: Robwatsoncolotes subgen. nov. can be separated and lower surfaces covered with juxtaposed granular scales; pupil from all other species in the genus Pseudogekko Taylor, 1915 (also vertical; males with preanal pores. being the nominate subgenus) by the following characters: Body The genera Scelotretus Fitzinger, 1843 and Georgemarioliscolotes size moderate with SVL 34.5-52 mm; total length 72-95.8 mm; gen. nov. are readily separated from the other two genera axilla-groin distance moderate being 17.8-29.8 mm; head length (Luperosaurus and Charlespiersoncolotes gen. nov.) by the moderate being 6.2-10.4 mm; midbody width 4.5-6.7 mm; head presence of a distinctly elongate head, elongate versus robust width 5.6-8.5 mm; snout length 3.8-6.0 mm; toe 4 scansors 15-20; body shape (in the other genera) and the presence of enlarged finger 3 scansors 12-16; paravertebrals 211-218; ventrals 96-123; interstitial granules. supralabials 13-16; infralabials 14-17; circumorbitals 33-46; The genera Luperosaurus and Charlespiersoncolotes gen. nov. are precloacal pores 12-16; femoral pores absent; conspicuous separated from Scelotretus Fitzinger, 1843 and dorsolateral spotting present; limb spotting absent; tail banding Georgemarioliscolotes gen. nov. by the presence of beadlike, absent; body striping absent; interorbital banding present. granular dorsals, a stout and robust, stout body and deeply Distribution: Central Phillippines. notched to divided penultimate subdigital scansors. Etymology: Named in honour of Brisbane wildlife controller and The genus Charlespiersoncolotes gen. nov. is most easily Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 47

herpetologist Robert Watson for services to wildlife conservation sp. nov. are charaterised by the presence of spots or peppering, and education in the south-east Queensland region. versus banding in S. orientalis. Content: Pseudogekko (Robwatsoncolotes) brevipes (Boettger, S. robjealousi sp. nov. is separated from S. dalegibbonsi sp. nov. 1867) (type species); P. (Robwatsoncolotes) atiorum Davis, by a darkish and well-marked dorsum, versus relatively indistinct Watters, Köhler, Whitsett, Huron, Brown, Diesmos and Siler, 2015. markings in S. dalegibbonsi sp. nov.. SHIREENHOSERGECKO SHIREENHOSERAE SP. NOV. Distribution: Known only from the type locality, but presumably Holotype: A preserved female specimen at the American Museum from other suitable rocky habitats in the general region, noting that of Natural History, New York City, USA, specimen number: AMNH the holotype was collected on a cliff face. 76766 collected from Mount Riu on Sudest Island, Louisiadae Etymology: Named in honour of Rob Jealous of Strathdale, Archipelago, Papua New Guinea. This facility allows access to its Bendigo, Victoria, Australia, in recognition of many decades of holdings. services to herpetology in Australia, in particular with regards to his Diagnosis: Shireenhosergecko shireenhoserae sp. nov. is support of scientists and keepers working with dangerously morphologically similar in most respects to the species S. orientalis venomous elapid snakes as well as his services to the Brown and Parker, 1977 from Papua New Guinea (as defined by conservation of these often killed and mistreated reptiles. those authors), but is readily separated from that species by SHIREENHOSERGECKO DALEGIBBONSI SP. NOV. extensive webbing on the feet. Holotype: A preserved specimen at the Bernice P. Bishop S. shireenhoserae sp. nov. is readily separated from all other Museum at Honolulu, Hawaii, USA specimen number: BPBM Herp- species in the genus Shireenhosergecko gen. nov. as defined BPBM 42860, collected from Umwate, Papua New Guinea, within this paper by the following suite of characters: 37-43 mm Latitude -9.26 S., Longitude 148.28 E. at an elevation of about 943 snout-vent length in adults; digits long and slender; moderately metres. This facility allows access to its holdings. dilated, extensive interdigital webbing; 10-12 entire fourth toe Diagnosis: Both Shireenhosergecko robjealousi sp. nov. and S. scansors covering the distal 2/3 of the toe; 19-24 enlarged scales dalegibbonsi sp. nov. are morphologically similar to S. orientalis in pore series confined to the preanal region only or only a few on Brown and Parker (1977) and would be defined as that using the the base of the thigh, bearing about 19 preanal pores in males. criteria of Brown and Parker (1977), but are readily separated from S. shireenhoserae sp. nov. is separated from the morphologically that taxon by their larger size (40-48 mm snout-vent in adults, similar S. jarradbinghami sp. nov. by having a grayish-brown as versus 37-43 mm) and of more stocky build (stocky vs moderate). opposed to dark brownish dorsal colouration. S. shireenhoserae The forelimbs of both S. robjealousi sp. nov. and S. dalegibbonsi sp. nov. is also slightly more robust in build than S. jarradbinghami sp. nov. are charaterised by the presence of spots or peppering, sp. nov.. versus obvious banding in S. orientalis. Distribution: Restricted to Sudest Island, Louisiadae Archipelago, S. robjealousi sp. nov. is separated from S. dalegibbonsi sp. nov. Papua New Guinea. by a darkish and well-marked dorsum, versus relatively indistinct Etymology: Named in honour of my wife, Shireen Hoser of markings in S. dalegibbonsi sp. nov.. Melbourne, Victoria, Australia for decades of service to wildlife Shireenhosergecko robjealousi sp. nov. and S. dalegibbonsi sp. conservation in Australia. nov. are clearly more like one another than they are to any other SHIREENHOSERGECKO PETEWHYBROWI SP. NOV. species in the genus. Holotype: A preserved specimen at Louisiana State University Distribution: Known only from in and around the type locality, but Museum of Natural Science. LSUMZ Herps Collection, specimen presumably from other suitable habitats in the general region. number: LSUMZ Herps 95824, collected at “Amau River Camp #1, Etymology: Named in honour of Dale Gibbons of Maiden Gully, about a 4 hour walk form Amau Village, good uncut primary Bendigo, Victoria, Australia, in recognition of many decades of forest.”, Latitude -9.98 S., Longitude 148.46 E. This facility allows services to herpetology in Australia, in particular with regards to his access to its holdings. support of scientists and keepers working with dangerously Diagnosis: Shireenhosergecko petewhybrowi sp. nov. is similar in venomous elapid snakes as well as his services to the most respects to S. orientalis Brown and Parker (1977), as defined conservation of these often killed and mistreated reptiles, as well by those authors, from which it is most readily differentiated by its as often unacknowledged services on countless wildlife field brownish ground colour, versus grayish or grayish tan in S. surveys in various parts of Australia, for which he never sought or orientalis. The forelimbs of S. petewhybrowi sp. nov. are got any payment. He has also made significant financial characterized by dark blotches versus banding in S. orientalis. contributions to poor and impoverished people in Thailand at great Distribution: Known only the vicinity of the type locality, but financial cost and hardship to himself. presumably in similar appropriate habitat further afield. This and SHIREENHOSERGECKO JARRADBINGHAMI SP. NOV. most other species referred to in the paper are presumably under Holotype: A preserved specimen at the Bernice P. Bishop threat from deforestation and generalized human created habitat Museum at Honolulu, Hawaii, USA specimen number: BPBM Herp- destruction, as well as human movement of so-called “weedy BPBM 17229, collected from Misima Island, Papua New Guinea, species” that may outcompete for resources. Latitude -10.66 S., Longitude 152.69 E. This facility allows access Etymology: Named in honour of Peter (Pete) Whybrow of to its holdings. Taggerty, north-east Victoria, Australia for services to herpetology Diagnosis: S. shireenhoserae sp. nov. is separated from the spanning some decades. morphologically similar S. jarradbinghami sp. nov. by having a SHIREENHOSERGECKO ROBJEALOUSI SP. NOV. grayish-brown as opposed to dark brownish dorsal colouration. It is Holotype: A preserved specimen at the Bernice P. Bishop the only species likely to be confused with S. jarradbinghami sp. Museum at Honolulu, Hawaii, USA specimen number: BPBM Herp- nov.. S. shireenhoserae sp. nov. is also slightly more robust in build BPBM 25943, collected from the Kamiali Wildlife Management than S. jarradbinghami sp. nov. and the two can be separated on Area, 1.3 km N and 6.2 km W of Cape Dinga, Morobe Province, that basis also. Papua New Guinea, Latitude -7.30 S., Longitude 147.09 E. This Shireenhosergecko shireenhoserae sp. nov. is otherwise facility allows access to its holdings. morphologically similar in most respects to the species S. orientalis Diagnosis: Both Shireenhosergecko robjealousi sp. nov. and S. Brown and Parker, 1977 from Papua New Guinea (as defined by dalegibbonsi sp. nov. are morphologically similar to S. orientalis those authors), but is readily separated from that species by Brown and Parker (1977) and would be defined as that using the extensive webbing on the feet (and same applies for S. criteria of Brown and Parker (1977), but are readily separated from jarradbinghami sp. nov.). that taxon by their larger size (40-48 mm snout-vent in adults, S. shireenhoserae sp. nov. is readily separated from all other versus 37-43 mm) and of more stocky build (stocky vs moderate). species in the genus Shireenhosergecko gen. nov. as defined The forelimbs of both S. robjealousi sp. nov. and S. dalegibbonsi within this paper by the following suite of characters: 37-43 mm snout-vent length in adults; digits long and slender; moderately

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 48 Australasian Journal of Herpetology dilated, extensive interdigital webbing; 10-12 entire fourth toe Bobbottomcolotes gen. nov.. scansors covering the distal 2/3 of the toe; 20-24 enlarged scales Geckos in the genus Bobbottomcolotes gen. nov. are separated in pore series confined to the preanal region only or only a few on from the other three genera, Shireenhosergecko gen. nov., the base of the thigh, bearing about 19 preanal pores in males. Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the Distribution: Known only from the type locality, Misima Island and following characters: More than 25 enlarged preanal/femoral presumably restricted to Misima Island, but this is by no means scales, extending onto distal portions of the thigh; 25 or more certain. preanal/femoral pores in males; fewer than 15 scansors under digit Etymology: Named in honour of Jarrad Bingham of Bacchus 4 of the hind leg: and one or other of: Marsh, Victoria, Australia, a councilor at the Moorabool Shire 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ Council also in Victoria, for services to local government and 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: wildlife conservation, in particular that of venomous snakes. 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 JACKYHOSERGECKO JACKYHOSERAE SP. NOV. the length of the digits; 40 or more enlarged preanal/femoral Holotype: A preserved specimen at the Australian Museum in scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. Sydney, New South Wales, Australia, specimen number: R.30511, magnus, B. bobbottomi sp. nov., B. potens sp. nov., B. pumilus). collected from Namuana, Kadavu Island, Fiji, Latitude -19.00 S., Distribution: Believed to be confined to the Adelbert Mountains in Longitude 178.25 E. The Australian Museum in Sydney, New South the Madang District of Papua New Guinea. Wales, Australia allows access to its holdings. Etymology: Named in honour of Robert (Bob) Bottom OAM, one Paratypes: 1/ A preserved specimen at the Australian Museum in of the foremost investigative journalists in Australian history. He Sydney, New South Wales, Australia, specimen number: R.30512, authored numerous best-selling books about government collected from Namuana, Kadavu Island, Fiji, Latitude -19.00 S., corruption and organized crime in Australia in the 1970’s and Longitude 178.25 E. 1980’s and years since then. For more detail, see also the 2/ A preserved specimen at the California Academy of Sciences. etymology for the genus Bobbottomcolotes gen. nov.. USA, CAS Herpetology collection, specimen number: 190626, BOBBOTTOMCOLOTES POTENS SP. NOV. collected from about 3 km. south of Richmond High School at Holotype: A preserved specimen at the Museum of Comparative Kadavu Island, Fiji, Latitude -19.09 S., Longitude 178.08 E. Zoology, Harvard University, USA, specimen number: MCZ Herp Diagnosis: Jackyhosergecko jackyhoserae sp. nov. is similar in R-54246, collected from Ebabaang, Huon Peninsula, Papua New most respects to J. manni (Schmidt, 1923), a species most Guinea, Latitude -6.49 S., Longitude 147.46 S. This facility allows commonly known to date as Lepidodactylus manni. access to its holdings. While colouration of both species is variable, J. jackyhoserae sp. Paratype: A preserved specimen at the Royal Belgian Institute of nov. is readily separated from J. manni by a general lack of well- Natural Sciences, Brussels, Belgium, specimen number: RBINS defined white pepeering in the region of the snout and upper labials Reptilia 423150, collected at Papua New Guinea, Latitude -5.85 S., in particular. J. jackyhoserae sp. nov. is generally a grayish brown, Longitude 145.92 E. versus a dark or reddish-dark brown in J. manni, irrespective of Diagnosis: Until now the species Bobbottomcolotes potens sp. dorsal markings or patterning on the specimens. nov. has been treated as an outlier population of B. magnus Brown Distribution: J. jackyhoserae sp. nov. is currently only known from and Parker, 1977 as defined by them in that paper. the Island of Kadavu, Fiji. J. manni is currently only known from While morphologically similar in most respects to B. magnus which and therefore restricted to Viti Levu (the main Fiji island) and it has been treated as until now, it is most readily separated from nearby Ovalau. that taxon by colouration being grayish tan in dorsal colouration as Etymology: Named in honour of my youngest daughter Jacky opposed to dark brown, as well as a sightly more slender build. Hoser of Park Orchards, Victoria, Australia in recognition of her Dorsal markings in B. potens sp. nov. are relatively indistinct, services to wildlife conservation over nearly 2 decades. See also versus clearly visible, albeit variable in B. magnus. Hoser (2013) at page 5 for more detail.. The species Bobbottomcolotes bobbottomi sp. nov. would until now BOBBOTTOMCOLOTES BOBBOTTOMI SP. NOV. have been defined as B. magnus Brown and Parker, 1977 as Holotype: A preserved male specimen in the American Museum of defined by them in that paper, and is separated from that taxon and Natural History, New York City, USA, specimen number: AMNH B. potens sp. nov. by its smaller adult size 38-44 mm snout vent, 185087 collected from the Adelbert Mountains in the Madang versus 50-70 mm snout vent in B. magnus and B. potens sp. nov., District of Papua New Guinea. This facility allows access to its as well 25-28 femoral / preanal pores in males versus 40-50 holdings. noticeably enlarged ones in B. magnus and B. potens sp. nov.. Paratype: A preserved female specimen in the American Museum B. bobbottomi sp. nov. is separated from the also morphologically of Natural History, New York City, USA, specimen number: AMNH similar B. pumilus (Boulenger, 1885) and B. crusmaculosus sp. 185088 collected from the Adelbert Mountains in the Madang nov. by having digits of moderate length versus relatively short and District of Papua New Guinea. dilated digits in B. pumilus and B. crusmaculosus sp. nov.. Diagnosis: Bobbottomcolotes bobbottomi sp. nov. would until now It is further separated from that species by having about 20% have been defined as B. magnus Brown and Parker, 1977 as webbing between toes 3 and 4, versus almost 50% webbing defined by them in that paper, but is separated from that taxon by between the toes in B. pumilus and B. crusmaculosus sp. nov.. its smaller adult size 38-44 mm snout vent, versus 50-70 mm There are no other described species in the genus snout vent in B. magnus, as well 25-28 femoral / preanal pores in Bobbottomcolotes gen. nov.. males versus 40-50 noticeably enlarged ones in B. magnus. Geckos in the genus Bobbottomcolotes gen. nov. are separated The taxon described in this paper Bobbottomcolotes potens sp. from the other three genera, Shireenhosergecko gen. nov., nov. is similar in most respects to B. magnus which it has been Jackyhosergecko gen. nov. and Martinekcolotes gen. nov. by the treated as until now, but is most readily separated from that taxon following characters: More than 25 enlarged preanal/femoral by colouration being grayish tan in dorsal colouration as opposed scales, extending onto distal portions of the thigh; 25 or more to dark brown. preanal/femoral pores in males; fewer than 15 scansors under digit B. bobbottomi sp. nov. is separated from the also morphologically 4 of the hind leg: and one or other of: similar B. pumilus (Boulenger, 1885) and B. crusmaculosus sp. 1/ Webbing between digits 3 and 4 of the hind legs is more than 1/ nov. by having digits of moderate length versus relatively short and 3 the length of the digit (B. pumilus, B. crusmaculosus sp. nov.), or: dilated digits in B. pumilus. 2/ Webbing between digits 3 and 4 of the hind legs is less than 1/4 It is further separated from that species by having about 20% the length of the digits; 40 or more enlarged preanal/femoral webbing between toes 3 and 4, versus almost 50% webbing scales; 10 or more dubdigital scansors on digit 1 of hind foot (B. between the toes in B. pumilus and B. crusmaculosus sp. nov.. magnus, B. bobbottomi sp. nov., B. potens sp. nov.). There are no other described species in the genus Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 49

Distribution: B. potens sp. nov. is apparently restricted to the Longitude 161 E. Mountains of the Huon Peninsula only. Diagnosis: Allengreercolotes pauldarwini sp. nov. is similar in Etymology: Potens means strong in Latin and reflects the general most respects to A. guppyi Boulenger, 1884, but is most easily size and build of this taxon relative to other similar related species. separated from that taxon by colouration. Both species have about BOBBOTTOMCOLOTES CRUSMACULOSUS SP. NOV. six smallish blackish spots along each flank. In A. guppyi there is Holotype: A preserved specimen at the Louisiana State University peppering in a line between the spots and/or the borders of each Museum of Natural Science. LSUMZ Herps Collection, (USA), spot is irregular and is not well defined. In A. pauldarwini sp. nov. specimen number: LSUMZ Herps 97472, collected from Daru the spots are well-defined and there is no black pigment joining Island at the mangroves at the South-east end of the airstrip, these spots in the intervening spaces. Western Province, Papua New Guinea. In A. pauldarwini sp. nov. the dorsal surface is characterized by a The Louisiana State University Museum of Natural Science. well-defined pattern of light yellowish-brownish-grey triangles along LSUMZ Herps Collection allows access to its holdings. the mid-dorsal line, surrounded by relatively unmarked darker Paratype: A preserved specimen at the Museum of Comparative grayish brown. By contrast in A. guppyi the dorsal markimgs are ill Zoology, Harvard University, USA, specimen number: MCZ Herp defined and invariably peppered with black speckling. R-137586, collected at Daru Island, Western Province, Papua New The iris of A. pauldarwini sp. nov. is reddish, versus brownish to Guinea, Latitude -9.08 S., Longitude 143.21 E. bluish-grey in A. guppyi. Diagnosis: Bobbottomcolotes crusmaculosus sp. nov. has until A. pauldarwini sp. nov. is depicted in life on plate 2, image F of now been treated as a population of B. pumilus (Boulenger, 1885), McCoy 1989. A. guppyi in life is depicted in plates 19 and 20 of which it is clearly most closely related to. It is readily separated McCoy (2006). The species Allengreercolotes allengreeri sp. nov. from B. pumilus by the presence of patches of dark brown is similar in most respects to A. guppyi Boulenger, 1884, (and A. speckling on the fore and hind limbs as well as patches of dark pauldarwini sp. nov.) which it would otherwise key as, using the key brown pigment on the anterior upper labials. The dorsal dark and in Brown and Parker (1977). light markings posterior to the pelvis are distinct and clear in B. A. allengreeri sp. nov. is however separated from these species by crusmaculosus sp. nov., versus ill defined in B. pumilus. its adult snout-vent length of 31-38 mm, versus 45-50 mm in A. Distribution: Mainland and immediately adjacent islands on the guppyi and A. allengreeri sp. nov. and by having 34-46 femoral / south-west coast of Papua New Guinea in the general vicinity of preanal pores in males versus 39-52 in A. guppyi and A. the type locality. allengreeri sp. nov.. Etymology: Crusmaculosus literally means spotted or speckled The femoral / preanal pores in A. allengreeri sp. nov. are moderate, legs in Latin, which differentiates this taxon from its most closely versus prominent in A. guppyi and A. pauldarwini sp. nov.. related congeneric species. It is worth noting that within the known populations of A. guppyi ALLENGREERCOLOTES ALLENGREERI SP. NOV. colouration also varies significantly between the islands and the retention of the northwestern populations in the Solomon Islands Holotype: A preserved mature male specimen at the Museum of and Bougainville as a single species A. guppyi is based on a lack Comparative Zoology (MCZ) at Harvard University, Cambridge, of molecular evidence separating them and the fact that ice-age Massachusetts, USA, specimen number: MCZ 135433 collected sea levels meant that most relevant islands were a single from Ndrova Island in the Admiralty Islands. connected landmass, theoretically allowing free movement Paratypes: Two preserved mature male specimens at the between them. Museum of Comparative Zoology (MCZ) at Harvard University, Distribution: Allengreercolotes pauldarwini sp. nov. occurs in the Cambridge, Massachusetts, USA, specimen numbers: MCZ main eastern islands and nearby smaller islands of the Solomon 135434 and MCZ 139418 collected from Ndrova Island in the Islands, including the type locality Malaupaina, Malaita and San Admiralty Islands. Cristobel. Diagnosis: Allengreercolotes allengreeri sp. nov. is similar in most Etymology: Named in honour of Paul Darwin, manager of the respects to A. guppyi Boulenger, 1884 which it would otherwise Brush Ski Lodge, Mount Hotham, Victoria, for some years key as, using the key in Brown and Parker (1977). preceding 2018, in recognition for his services to the downhill snow It is however it is separated from that species by its adult snout- skiing industry, including actively diverting young Australians and vent length of 31-38 mm, versus 45-50 mm in A. guppyi, and by foreigners away from their mobile phones and computers and having 34-46 femoral / preanal pores in males versus 39-52 in A. actually engaging in outdoor sport, exercise and appreciation of guppyi. their natural environment. The femoral / preanal pores in A. allengreeri sp. nov. are moderate, ALLENGREERCOLOTES PAULWOOLFI SP. NOV. versus prominent in A. guppyi. Holotype: A preserved specimen at the United States National A. guppyi Boulenger, 1884 and A. pauldarwini sp. nov. (this paper) Museum, better known as the Smithsonian, at Washington, DC, are most readily separated from one another by colour as United States of America, specimen number: USNM 533293, explained in the description following this one, but both otherwise collected from Temotu Village at Mahele Point, Taumako Island, visibly share the same features as for A. guppyi as explained Duff Islands, Solomon Islands, Latitude -9.88 S., Longitude above. 167.175 S. This facility allows access to its holdings. Distribution: A. allengreeri sp. nov. is known only from the type Diagnosis: A. paulwoolfi sp. nov. is similar in most respects to A. locality Ndrova Island in the Admiralty Islands, but presumably vanuatuensis (Ota, Fisher, Ineich, Case, Radtkey and Zug, 1998), occurs on adjacent islands. as defined by Ota, Fisher, Ineich, Case, Radtkey and Zug (1998) Etymology: Named in honour of Dr. Allen E. Greer, curator of from which it is separated by having a moderately well-defined row herpetology for many years at the Australian Museum in Sydney, of 4-6 well-spaced dark grey to black spots on the lower flanks. New South Wales, Australia, (now retired) in recognition of his A. paulwoolfi sp. nov. is further separated from A. vanuatuensis by services to herpetology globally. the presence of one or more well defined scattered dark blotches ALLENGREERCOLOTES PAULDARWINI SP. NOV. on both fore and hind limbs. Holotype: A preserved specimen at the Australian Museum in Distribution: Known only from the type locality. Sydney, New South Wales, Australia, specimen number R.93499 Etymology: Named in honour of Paul Woolf of Walloon, collected at Malaupaina, Solomon Islands, Latitude -10.15 S., Queensland, Australia, in recognition of many years services to Longitude 161.58 E. The Australian Museum is a government- herpetology including as foundation president of the Herpetological owned facility that allows access to its holdings. Society of Queensland Incorporated. Paratype: A preserved specimen at the Museum of Comparative ADELYNHOSERGECKO ADELYNHOSERAE SP. NOV. Zoology, Harvard University, USA, specimen number: MCZ Herp Holotype: A preserved specimen at the South Australian Museum R-115559 collected from Malaita, Solomon Islands, Latitude -9 S, in Adelaide, South Australia, Australia, specimen number: SAMA

Herpetology R64666, collected from New Britain, Papua New shaped dorsal crossbands of dark and light grey, the dark bands Guinea, Latitude -5.61 S., Longitude 151.41 E. The South becoming darker on the lower flanks. Australian Museum in Adelaide, South Australia, Australia allows Adelynhosergecko huonensis sp. nov. is similar in most respects to access to its holdings. A. novaeguineae Brown and Parker (1977), but is separated from Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that species by having a grey dorsal colouration versus reddish- Britain, is similar in most respects to the species A. novaeguineae brown in both A. novaeguineae and A. madangensis sp. nov. as (Brown and Parker, 1977) and would be identified as this taxon described within this paper. A. huonensis sp. nov. has vague as using the criteria given in that paper. A. adelynhoserae sp. nov. is opposed to moderately distinct dorsal markings as seen in the however readily separated from A. novaeguineae by the general other two species. absence of reddish-brown or red flecks and markings on the dorsal A. madangensis sp. nov. is separated from A. novaeguineae and surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 A. huonensis sp. nov. and others in the genus by the presence of scattered yellow dots on each of the lower flanks above where the an indistinct dorsal pattern with distinctly jagged edges on the grayish upper body turns to the lighter undersurface. The well- upper surface, these sometimes consisting of bands or markings defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. indicative of this. are whiteish and grayish in colour versus tan and reddish-brown in Geckos in the genus Adelynhosergecko gen. nov. are readily A. novaeguineae. separated from geckos in the genera Allengreercolotes gen. nov., The species A. judyfergusonae sp. nov. from Unea Island, is Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by separated from A. adelynhoserae sp. nov. by the relative lack of having fewer than 25 enlarged preanal/femoral pores, snout-vent black peppering in the lighter bands as compared to that seen in A. length in adults of under 41 mm and of moderate build. adelynhoserae sp. nov., which is readily noticeable. The limited Exceptional to this in terms of size, Adelynhosergecko sloppi sp. number of blackish spots on the dorsum and flanks of A. nov. does exceed 41 mm in snout-vent length. adelynhoserae sp. nov. are indistinct, versus prominent in A. Distribution: Known only from New Britain in the Bismarck judyfergusonae sp. nov.. Archipelago of Papua New Guinea. Otherwise both species are similar. Etymology: Named in honour of my eldest daughter Adelyn Hoser The species A. haydnmcphiei sp. nov. from Ambitle Island, is of Park Orchards, Victoria, Australia in recognition of her services readily separated from both A. adelynhoserae sp. nov.and A. to wildlife conservation over nearly 2 decades. judyfergusonae sp. nov.by an absence of yellow mottling on the ADELYNHOSERGECKO SLOPPI SP. NOV. back of the jaw and sides of neck. Striping behind the eye is also Holotype: A preserved specimen at the Museum of Natural noticeable in A. haydnmcphiei sp. nov.. History, London, UK, specimen number: BMNH 1974.3027 The species A. matteoae sp. nov. from the Kei Islands, is similar in collected from Waigue Island (=Waigeo), north-west New Guinea. most respects to A. novaeguineae (Brown and Parker, 1977), but This is a facility that allows access to its holdings. is separated from it by a diffuse dull beige indististinctly marked Diagnosis: The species Adelynhosergecko sloppi sp. nov. from colouration with scattered yellowish flecks on the dorsum. Waigeo Island, Irian Jaya, Indonesia, is separated from all other The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, geckos in the genera Adelynhosergecko gen. nov., Irian Jaya, Indonesia, is separated from all other geckos in the Allengreercolotes gen. nov., Borneocolotes gen. nov. and genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Rosssadliercolotes gen. nov. by having fewer than 25 enlarged Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by preanal/femoral pores and an adult snout-vent length in excess of having fewer than 25 enlarged preanal/femoral pores and an adult 42 mm and robust build. It is morphologically similar in most snout-vent length in excess of 42 mm and robust build. It is respects to species within the genus Adelynhosergecko gen. nov. morphologically similar in most respects to species within the in particular Adelynhosergecko novaeguineae (Brown and Parker, genus Adelynhosergecko gen. nov. in particular Adelynhosergecko 1977), but differs from this taxon by its larger adult size (in excess novaeguineae (Brown and Parker, 1977), but differs from this taxon of 42 mm snout-vent, versus under 41 mm) and more robust build. by its larger adult size (in excess of 42 mm snout-vent, versus Otherwise it woukd key out as that species according to that in under 41 mm) and more robust build. Brown and Parker (1977). All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. Distribution: Known only from the type locality, but potentially haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily more widespread in North-west New Guinea. separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis Etymology: Named in honour of the family Great Dane, named sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. Slopp, in recognition of his services in protecting the scientific stevebennetti sp. nov, A. lucybennettae sp. nov. and A. research facilities of this author for the nearly six years preceding lachlanmcpheei sp. nov. by having no webbing or less than 1/5 the publication of this paper. webbing between digits 3 and 4 of the hind leg, versus from 1/5 to 1/4 in the other species. ADELYNHOSERGECKO HUONENSIS SP. NOV. The species A.brettbarnetti sp. nov. from Woodlark Island and A. Holotype: A preserved specimen at the National Museum of stevebennetti sp. nov. from Boiaboiawaga Island, both in south- Natural History, Smithsonian Institution, Washington, DC, USA, east Papua New Guinea have until now been identified as A. specimen number: USNM Amphibians and Reptiles 119248, novaeguineae (Brown and Parker, 1977) and are similar to it. The collected at Gusika, Morobe Province, Papua New Guinea, can both be separated from that taxon by marbling on the nape Latitude -6.42 S., Longitude 147.83 E. The National Museum of and anterior dorsum as well as tail bands that have the darker Natural History, Smithsonian Institution allows access to its segments darker posteriorly and fading anteriorly. A. stevebennetti holdings. sp. nov. is further separated from the other species (A.brettbarnetti Paratypes: Two preserved specimens at the American Museum of sp. nov.) by scattered dark flecks on the whitish surface of the Natural History, New York, USA, specimen numbers: AMNH 66665 lower flanks. and 66667, collected at Gusika, Morobe Province, Papua New The species A. lucybennettae sp. nov. from the vicinity of Utai Guinea, Latitude -6.42 S., Longitude 147.83 E. Village, Sandaun Province, in northern Papua New Guinea and A. Diagnosis: Adelynhosergecko huonensis sp. nov. is similar in lachlanmcpheei sp. nov. from the vicinity of Weewak in East Sepik most respects to A. novaeguineae Brown and Parker (1977), but is Province, Papua New Guinea are similar in most respects to A. separated from that species by having a grey dorsal colouration novaeguineae (Brown and Parker, 1977) and until now would have versus reddish-brown in both A. novaeguineae and A. been identified as that taxon, but can be separated from it by madangensis sp. nov. as described within this paper. A. huonensis longer and narrower digits (long and narrow versus moderate). A. sp. nov. has vague as opposed to moderately distinct dorsal lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. markings as seen in the other two species. nov. by the presence of about four well defined and irregular A. madangensis sp. nov. is further separated from A. novaeguineae and A. huonensis sp. nov. by the presence of an Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved Australasian Journal of Herpetology 51

indistinct dorsal pattern with distinctly jagged edges on the upper ADELYNHOSERGECKO JUDYFERGUSONAE SP. NOV. surface sometimes consisting of bands or markings indicative of Holotype: A preserved specimen at the Royal Belgian Institute of this. Natural Sciences, Brussells, Belgium, specimen number: RBINS The eight species A. novaeguineae, A. sloppi sp. nov., A. Reptilia 380100 collected at Unea Island (near New Britain) in huonensis sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. Papua New Guinea. This facility allows access to its holdings. nov., A. stevebennetti sp. nov, A. lucybennettae sp. nov., and A. Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New lachlanmcpheei sp. nov. are separated from the other species in Britain, is similar in most respects to the species A. novaeguineae the genus by having webbing between digits 3 and 4 of the hind leg (Brown and Parker, 1977) and would be identified as this taxon being less than a quarter of the length of the digits and more than using the criteria given in that paper. A. adelynhoserae sp. nov. is four fifths as well as having 16 or less subdigital scansors on digit however readily separated from A. novaeguineae by the general four of the hind leg. absence of reddish-brown or red flecks and markings on the dorsal Geckos in the genus Adelynhosergecko gen. nov. are readily surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 separated from geckos in the genera Allengreercolotes gen. nov., scattered yellow dots on each of the lower flanks above where the Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by grayish upper body turns to the lighter undersurface. The well- having fewer than 25 enlarged preanal/femoral pores, snout-vent defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. length in adults of under 41 mm and of moderate build. are whiteish and grayish in colour versus tan and reddish-brown in Exceptional to this in terms of size, Adelynhosergecko sloppi sp. A. novaeguineae. nov. does exceed 41 mm in snout-vent length. The species A. judyfergusonae sp. nov. from Unea Island, is Distribution: Apparently restricted to the Huon Peninsula, New separated from A. adelynhoserae sp. nov. by the relative lack of Guinea. black peppering in the lighter bands as compared to that seen in A. Etymology: Named in reflection fo where the species occurs, adelynhoserae sp. nov., which is readily noticeable. The limited being the Huon Peninsula. number of blackish spots on the dorsum and flanks of A. ADELYNHOSERGECKO MADANGENSIS SP. NOV. adelynhoserae sp. nov. are indistinct, versus prominent in A. judyfergusonae sp. nov.. Holotype: A preserved specimen at the American Museum of Natural History, New York, USA, specimen number: AMNH Otherwise both species are similar. 100209, collected from Mount Nibo, Sepik District, Papua New The species A. haydnmcphiei sp. nov. from Ambitle Island, is Guinea. The American Museum of Natural History allows access to readily separated from both A. adelynhoserae sp. nov.and A. its holdings. judyfergusonae sp. nov.by an absence of yellow mottling on the Paratypes: 1/ A preserved specimen at the American Museum of back of the jaw and sides of neck. Striping behind the eye is also Natural History, New York, USA, specimen number: AMNH noticeable in A. haydnmcphiei sp. nov.. 100210, collected from Mount Nibo, Sepik District, Papua New The species A. matteoae sp. nov. from the Kei Islands, is similar in Guinea. most respects to A. novaeguineae (Brown and Parker, 1977), but 2/ A preserved specimen at the Museum of Vertebrate Zoology, is separated from it by a diffuse dull beige indististinctly marked University of California, Berkeley, California, USA, specimen colouration with scattered yellowish flecks on the dorsum. number: MVZ Amphibian and reptile specimens 89674, collected at The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, Wanuma, Adelbert Mountains, Madang Province, Papua New Irian Jaya, Indonesia, is separated from all other geckos in the Guinea. genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Diagnosis: The newly described species Adelynhosergecko Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by huonensis sp. nov. is similar in most respects to A. novaeguineae having fewer than 25 enlarged preanal/femoral pores and an adult Brown and Parker (1977), but is separated from that species by snout-vent length in excess of 42 mm and robust build. It is having a grey dorsal colouration versus reddish-brown in both A. morphologically similar in most respects to species within the novaeguineae and A. madangensis sp. nov. as described herein. genus Adelynhosergecko gen. nov. in particular Adelynhosergecko A. huonensis sp. nov. has vague as opposed to moderately distinct novaeguineae (Brown and Parker, 1977), but differs from this taxon dorsal markings as seen in the other two species. by its larger adult size (in excess of 42 mm snout-vent, versus under 41 mm) and more robust build. A. madangensis sp. nov. is further separated from A. novaeguineae and A. huonensis sp. nov. by the presence of an All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. indistinct dorsal pattern with distinctly jagged edges on the upper haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily surface sometimes consisting of bands or markings indicative of separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis this. sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. stevebennetti sp. nov, A. lucybennettae sp. nov. and A. The eight species A. novaeguineae, A. sloppi sp. nov., A. lachlanmcpheei sp. nov. by having no webbing or less than 1/5 huonensis sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. webbing between digits 3 and 4 of the hind leg, versus from 1/5 to nov., A. stevebennetti sp. nov, A. lucybennettae sp. nov., and A. 1/4 in the other species. lachlanmcpheei sp. nov. are separated from the other species in the genus by having webbing between digits 3 and 4 of the hind leg The species A.brettbarnetti sp. nov. from Woodlark Island and A. being less than a quarter of the length of the digits and more than stevebennetti sp. nov. from Boiaboiawaga Island, both in south- four fifths as well as having 16 or less subdigital scansors on digit east Papua New Guinea have until now been identified as A. four of the hind leg. novaeguineae (Brown and Parker, 1977) and are similar to it. The can both be separated from that taxon by marbling on the nape Geckos in the genus Adelynhosergecko gen. nov. are readily and anterior dorsum as well as tail bands that have the darker separated from geckos in the genera Allengreercolotes gen. nov., segments darker posteriorly and fading anteriorly. A. stevebennetti Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by sp. nov. is further separated from the other species (A.brettbarnetti having fewer than 25 enlarged preanal/femoral pores, snout-vent sp. nov.) by scattered dark flecks on the whitish surface of the length in adults of under 41 mm and of moderate build. lower flanks. Exceptional to this in terms of size, Adelynhosergecko sloppi sp. The species A. lucybennettae sp. nov. from the vicinity of Utai nov. does exceed 41 mm in snout-vent length. Village, Sandaun Province, in northern Papua New Guinea and Distribution: Believed to be limited to the ranges in the north of and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East Madang and immediately west of the Sepik River mouth, wholly Sepik Province, Papua New Guinea are similar in most respects to within Papua New Guinea. A. novaeguineae (Brown and Parker, 1977) and until now would Etymology: Named in reflection of where the species occurs, have been identified as that taxon, but can be separated from it by being Madang, Papua New Guinea. longer and narrower digits (long and narrow versus moderate). A. lachlanmcpheei sp. nov. is separated from A. lucybennettae sp.

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 52 Australasian Journal of Herpetology nov. by the presence of about four well defined and irregular most respects to A. novaeguineae (Brown and Parker, 1977), but shaped dorsal crossbands of dark and light grey, the dark bands is separated from it by a diffuse dull beige indististinctly marked becoming darker on the lower flanks. colouration with scattered yellowish flecks on the dorsum. Adelynhosergecko huonensis sp. nov. is similar in most respects to The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, A. novaeguineae Brown and Parker (1977), but is separated from Irian Jaya, Indonesia, is separated from all other geckos in the that species by having a grey dorsal colouration versus reddish- genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., brown in both A. novaeguineae and A. madangensis sp. nov. as Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by described within this paper. A. huonensis sp. nov. has vague as having fewer than 25 enlarged preanal/femoral pores and an adult opposed to moderately distinct dorsal markings as seen in the snout-vent length in excess of 42 mm and robust build. It is other two species. morphologically similar in most respects to species within the A. madangensis sp. nov. is further separated from A. genus Adelynhosergecko gen. nov. in particular Adelynhosergecko novaeguineae and A. huonensis sp. nov. by the presence of an novaeguineae (Brown and Parker, 1977), but differs from this taxon indistinct dorsal pattern with distinctly jagged edges on the upper by its larger adult size (in excess of 42 mm snout-vent, versus surface sometimes consisting of bands or markings indicative of under 41 mm) and more robust build. this. All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. Adelynhosergecko huonensis sp. nov. is similar in most respects to haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily A. novaeguineae Brown and Parker (1977), but is separated from separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis that species by having a grey dorsal colouration versus reddish- sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. brown in both A. novaeguineae and A. madangensis sp. nov. as stevebennetti sp. nov, A. lucybennettae sp. nov. and A. described within this paper. A. huonensis sp. nov. has vague as lachlanmcpheei sp. nov. by having no webbing or less than 1/5 opposed to moderately distinct dorsal markings as seen in the webbing between digits 3 and 4 of the hind leg, versus from 1/5 to other two species. 1/4 in the other species. A. madangensis sp. nov. is further separated from A. The species A.brettbarnetti sp. nov. from Woodlark Island and A. novaeguineae and A. huonensis sp. nov. by the presence of an stevebennetti sp. nov. from Boiaboiawaga Island, both in south- indistinct dorsal pattern with distinctly jagged edges on the upper east Papua New Guinea have until now been identified as A. surface sometimes consisting of bands or markings indicative of novaeguineae (Brown and Parker, 1977) and are similar to it. The this. can both be separated from that taxon by marbling on the nape Geckos in the genus Adelynhosergecko gen. nov. are readily and anterior dorsum as well as tail bands that have the darker separated from geckos in the genera Allengreercolotes gen. nov., segments darker posteriorly and fading anteriorly. A. stevebennetti Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by sp. nov. is further separated from the other species (A.brettbarnetti having fewer than 25 enlarged preanal/femoral pores, snout-vent sp. nov.) by scattered dark flecks on the whitish surface of the length in adults of under 41 mm and of moderate build. lower flanks. Exceptional to this in terms of size, Adelynhosergecko sloppi sp. The species A. lucybennettae sp. nov. from the vicinity of Utai nov. does exceed 41 mm in snout-vent length. Village, Sandaun Province, in northern Papua New Guinea and Distribution: Known only from the type locality Unea Island (near and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East New Britain) in Papua New Guinea. Etymology: Named in honour Sepik Province, Papua New Guinea are similar in most respects to of Judy Feguson of Taggerty, north-eastern Victoria, Australia in A. novaeguineae (Brown and Parker, 1977) and until now would recognition of many decades services to herpetology in Australia. have been identified as that taxon, but can be separated from it by longer and narrower digits (long and narrow versus moderate). A. ADELYNHOSERGECKO HAYDNMCPHIEI SP. NOV. lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. Holotype: A preserved specimen at the Museum of Comparative nov. by the presence of about four well defined and irregular Zoology, Harvard University, Cambridge, Massachusetts, USA, shaped dorsal crossbands of dark and light grey, the dark bands specimen number: MCZ Herp R-153102 collected from Waramung becoming darker on the lower flanks. Plantation, Ambitle Island, New Ireland Province, Papua New Adelynhosergecko huonensis sp. nov. is similar in most respects to Guinea. This facility allows access to its holdings. A. novaeguineae Brown and Parker (1977), but is separated from Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that species by having a grey dorsal colouration versus reddish- Britain, is similar in most respects to the species A. novaeguineae brown in both A. novaeguineae and A. madangensis sp. nov. as (Brown and Parker, 1977) and would be identified as this taxon described within this paper. A. huonensis sp. nov. has vague as using the criteria given in that paper. A. adelynhoserae sp. nov. is opposed to moderately distinct dorsal markings as seen in the however readily separated from A. novaeguineae by the general other two species. absence of reddish-brown or red flecks and markings on the dorsal A. madangensis sp. nov. is further separated from A. surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 novaeguineae and A. huonensis sp. nov. by the presence of an scattered yellow dots on each of the lower flanks above where the indistinct dorsal pattern with distinctly jagged edges on the upper grayish upper body turns to the lighter undersurface. The well- surface sometimes consisting of bands or markings indicative of defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. this. are whiteish and grayish in colour versus tan and reddish-brown in A. novaeguineae. Adelynhosergecko huonensis sp. nov. is similar in most respects to A. novaeguineae Brown and Parker (1977), but is separated from The species A. judyfergusonae sp. nov. from Unea Island, is that species by having a grey dorsal colouration versus reddish- separated from A. adelynhoserae sp. nov. by the relative lack of brown in both A. novaeguineae and A. madangensis sp. nov. as black peppering in the lighter bands as compared to that seen in A. described within this paper. A. huonensis sp. nov. has vague as adelynhoserae sp. nov., which is readily noticeable. The limited opposed to moderately distinct dorsal markings as seen in the number of blackish spots on the dorsum and flanks of A. other two species. adelynhoserae sp. nov. are indistinct, versus prominent in A. judyfergusonae sp. nov.. A. madangensis sp. nov. is further separated from A. novaeguineae and A. huonensis sp. nov. by the presence of an Otherwise both species are similar. indistinct dorsal pattern with distinctly jagged edges on the upper The species A. haydnmcphiei sp. nov. from Ambitle Island, is surface sometimes consisting of bands or markings indicative of readily separated from both A. adelynhoserae sp. nov.and A. this. judyfergusonae sp. nov. (the two species it would otherwise be Geckos in the genus Adelynhosergecko gen. nov. are readily confused with) by an absence of yellow mottling on the back of the separated from geckos in the genera Allengreercolotes gen. nov., jaw and sides of neck. Striping behind the eye is also noticeable in Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by A. haydnmcphiei sp. nov.. having fewer than 25 enlarged preanal/femoral pores, snout-vent The species A. matteoae sp. nov. from the Kei Islands, is similar in

length in adults of under 41 mm and of moderate build. 1/4 in the other species. Exceptional to this in terms of size, Adelynhosergecko sloppi sp. The species A.brettbarnetti sp. nov. from Woodlark Island and A. nov. does exceed 41 mm in snout-vent length. stevebennetti sp. nov. from Boiaboiawaga Island, both in south- Distribution: Only known from the type locality of Ambitle Island, east Papua New Guinea have until now been identified as A. New Ireland Province, Papua New Guinea. novaeguineae (Brown and Parker, 1977) and are similar to it. The Etymology: Named in honour of Haydn McPhie of Mirboo North, can both be separated from that taxon by marbling on the nape West Gippsland, Victoria, Australia, in recognition of many and anterior dorsum as well as tail bands that have the darker decades of important contributions to Australian herpetology. segments darker posteriorly and fading anteriorly. A. stevebennetti Included in this has been in apprehending thieves seeking to steal sp. nov. is further separated from the other species (A.brettbarnetti reptiles from hands on wildlife displays by Snakebusters: sp. nov.) by scattered dark flecks on the whitish surface of the Australia’s best reptiles. Dressed in plain clothes, he was able to lower flanks. observe and stop thieves after allowing them to leave the display The species A. lucybennettae sp. nov. from the vicinity of Utai area with reptiles they had stolen. Village, Sandaun Province, in northern Papua New Guinea and Significantly, on one occasion he caught out a female thief and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East unsuccessfully attempting to steal a Woma Python (Aspidites Sepik Province, Papua New Guinea are similar in most respects to ramsayi (Macleay, 1882)) from a display at the Traralgon A. novaeguineae (Brown and Parker, 1977) and until now would Agricultural Show. The woman happened to be a female employee have been identified as that taxon, but can be separated from it by of the State wildlife department! longer and narrower digits (long and narrow versus moderate). A. ADELYNHOSERGECKO MATTEOAE SP. NOV. lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. nov. by the presence of about four well defined and irregular Holotype: A preserved specimen at the Museum Zoologicum shaped dorsal crossbands of dark and light grey, the dark bands Bogoriense, Bogor, Indonesia, specimen number: MZB 14063 becoming darker on the lower flanks. collected from the Kei Islands, Indonesia. This facility allows access to its holdings. Adelynhosergecko huonensis sp. nov. is similar in most respects to A. novaeguineae Brown and Parker (1977), but is separated from Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that species by having a grey dorsal colouration versus reddish- Britain, is similar in most respects to the species A. novaeguineae brown in both A. novaeguineae and A. madangensis sp. nov. as (Brown and Parker, 1977) and would be identified as this taxon described within this paper. A. huonensis sp. nov. has vague as using the criteria given in that paper. A. adelynhoserae sp. nov. is opposed to moderately distinct dorsal markings as seen in the however readily separated from A. novaeguineae by the general other two species. absence of reddish-brown or red flecks and markings on the dorsal surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 A. madangensis sp. nov. is further separated from A. scattered yellow dots on each of the lower flanks above where the novaeguineae and A. huonensis sp. nov. by the presence of an grayish upper body turns to the lighter undersurface. The well- indistinct dorsal pattern with distinctly jagged edges on the upper defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. surface sometimes consisting of bands or markings indicative of are whiteish and grayish in colour versus tan and reddish-brown in this. A. novaeguineae. Adelynhosergecko huonensis sp. nov. is similar in most respects to The species A. judyfergusonae sp. nov. from Unea Island, is A. novaeguineae Brown and Parker (1977), but is separated from separated from A. adelynhoserae sp. nov. by the relative lack of that species by having a grey dorsal colouration versus reddish- black peppering in the lighter bands as compared to that seen in A. brown in both A. novaeguineae and A. madangensis sp. nov. as adelynhoserae sp. nov., which is readily noticeable. The limited described within this paper. A. huonensis sp. nov. has vague as number of blackish spots on the dorsum and flanks of A. opposed to moderately distinct dorsal markings as seen in the adelynhoserae sp. nov. are indistinct, versus prominent in A. other two species. judyfergusonae sp. nov.. A. madangensis sp. nov. is further separated from A. Otherwise both species are similar. novaeguineae and A. huonensis sp. nov. by the presence of an indistinct dorsal pattern with distinctly jagged edges on the upper The species A. haydnmcphiei sp. nov. from Ambitle Island, is surface sometimes consisting of bands or markings indicative of readily separated from both A. adelynhoserae sp. nov.and A. this. judyfergusonae sp. nov.by an absence of yellow mottling on the back of the jaw and sides of neck. Striping behind the eye is also Geckos in the genus Adelynhosergecko gen. nov. are readily noticeable in A. haydnmcphiei sp. nov.. separated from geckos in the genera Allengreercolotes gen. nov., Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by The species A. matteoae sp. nov. from the Kei Islands, is similar in having fewer than 25 enlarged preanal/femoral pores, snout-vent most respects to A. novaeguineae (Brown and Parker, 1977), but length in adults of under 41 mm and of moderate build. is separated from it and others in the genus by a diffuse dull beige indististinctly marked colouration with scattered yellowish flecks on Exceptional to this in terms of size, Adelynhosergecko sloppi sp. the dorsum. nov. does exceed 41 mm in snout-vent length. The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, Distribution: Only known from the type locality of Kei Islands, Irian Jaya, Indonesia, is separated from all other geckos in the located south-west of Irian Jaya, Indonesia. genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Etymology: Named in honour of Cathryn Matteo of Hawthorn, Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by Victoria, Australia in recognition to her significant logistical having fewer than 25 enlarged preanal/femoral pores and an adult contributions to herpetological research and conservation in snout-vent length in excess of 42 mm and robust build. It is Australia over some decades, in particular with regard to morphologically similar in most respects to species within the information technology (IT), sorting data and similar matters. genus Adelynhosergecko gen. nov. in particular Adelynhosergecko ADELYNHOSERGECKO BRETTBARNETTI SP. NOV. novaeguineae (Brown and Parker, 1977), but differs from this taxon Holotype: A preserved specimen at the Bernice P. Bishop by its larger adult size (in excess of 42 mm snout-vent, versus Museum, Honolulu, Hawaii, USA, specimen number: BPBM Herp- under 41 mm) and more robust build. BPBM 39879 collected at Gisabwai, Woodlark Island, Papua New All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. Guinea, Latitude -9.08 S., Longitude 152.77 E. This facility allows haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily access to its holdings. separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis Paratypes: Two preserved specimens at the Bernice P. Bishop sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. Museum, Honolulu, Hawaii, USA, specimen numbers: BPBM Herp- stevebennetti sp. nov, A. lucybennettae sp. nov. and A. BPBM 39880 and BPBM Herp-BPBM 39881 collected at Gisabwai, lachlanmcpheei sp. nov. by having no webbing or less than 1/5 Woodlark Island, Papua New Guinea, Latitude -9.08 S., Longitude webbing between digits 3 and 4 of the hind leg, versus from 1/5 to 152.77 E.

Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that species by having a grey dorsal colouration versus reddish- Britain, is similar in most respects to the species A. novaeguineae brown in both A. novaeguineae and A. madangensis sp. nov. as (Brown and Parker, 1977) and would be identified as this taxon described within this paper. A. huonensis sp. nov. has vague as using the criteria given in that paper. A. adelynhoserae sp. nov. is opposed to moderately distinct dorsal markings as seen in the however readily separated from A. novaeguineae by the general other two species. absence of reddish-brown or red flecks and markings on the dorsal A. madangensis sp. nov. is further separated from A. surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 novaeguineae and A. huonensis sp. nov. by the presence of an scattered yellow dots on each of the lower flanks above where the indistinct dorsal pattern with distinctly jagged edges on the upper grayish upper body turns to the lighter undersurface. The well- surface sometimes consisting of bands or markings indicative of defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. this. are whiteish and grayish in colour versus tan and reddish-brown in Adelynhosergecko huonensis sp. nov. is similar in most respects to A. novaeguineae. A. novaeguineae Brown and Parker (1977), but is separated from The species A. judyfergusonae sp. nov. from Unea Island, is that species by having a grey dorsal colouration versus reddish- separated from A. adelynhoserae sp. nov. by the relative lack of brown in both A. novaeguineae and A. madangensis sp. nov. as black peppering in the lighter bands as compared to that seen in A. described within this paper. A. huonensis sp. nov. has vague as adelynhoserae sp. nov., which is readily noticeable. The limited opposed to moderately distinct dorsal markings as seen in the number of blackish spots on the dorsum and flanks of A. other two species. adelynhoserae sp. nov. are indistinct, versus prominent in A. A. madangensis sp. nov. is further separated from A. judyfergusonae sp. nov.. novaeguineae and A. huonensis sp. nov. by the presence of an Otherwise both species are similar. indistinct dorsal pattern with distinctly jagged edges on the upper The species A. haydnmcphiei sp. nov. from Ambitle Island, is surface sometimes consisting of bands or markings indicative of readily separated from both A. adelynhoserae sp. nov.and A. this. judyfergusonae sp. nov.by an absence of yellow mottling on the Geckos in the genus Adelynhosergecko gen. nov. are readily back of the jaw and sides of neck. Striping behind the eye is also separated from geckos in the genera Allengreercolotes gen. nov., noticeable in A. haydnmcphiei sp. nov.. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by The species A. matteoae sp. nov. from the Kei Islands, is similar in having fewer than 25 enlarged preanal/femoral pores, snout-vent most respects to A. novaeguineae (Brown and Parker, 1977), but length in adults of under 41 mm and of moderate build. is separated from it by a diffuse dull beige indististinctly marked Exceptional to this in terms of size, Adelynhosergecko sloppi sp. colouration with scattered yellowish flecks on the dorsum. nov. does exceed 41 mm in snout-vent length. The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, Distribution: Only known from the type locality of Woodlark Island, Irian Jaya, Indonesia, is separated from all other geckos in the Papua New Guinea. genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., Etymology: Named in honour of Brett Barnett of Ardeer, Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by Melbourne, Victoria, Australia in recognition of his immense having fewer than 25 enlarged preanal/femoral pores and an adult contributions to herpetology over some decades, including snout-vent length in excess of 42 mm and robust build. It is contributions to the sustainability of the Victorian Herpetological morphologically similar in most respects to species within the Society, at times hijacked by dishonest people trying to misuse genus Adelynhosergecko gen. nov. in particular Adelynhosergecko society funds for their own financial self-gratification. novaeguineae (Brown and Parker, 1977), but differs from this taxon He has also provided critically important support for the care and by its larger adult size (in excess of 42 mm snout-vent, versus maintenance of captive reptiles at facilities across Australia, under 41 mm) and more robust build. including scientific research institutions, private and public zoos All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. and private hobbyists. haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily ADELYNHOSERGECKO STEVEBENNETTI SP. NOV. separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis Holotype: A preserved specimen at the Bernice P. Bishop sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. Museum, Honolulu, Hawaii, USA, specimen number: BPBM Herp- stevebennetti sp. nov, A. lucybennettae sp. nov. and A. BPBM 15843 collected at Boiaboiawaga Island, Milne Bay lachlanmcpheei sp. nov. by having no webbing or less than 1/5 Province, Papua New Guinea, Latitude -10.21 S., Longitude webbing between digits 3 and 4 of the hind leg, versus from 1/5 to 150.90 E. This facility allows access to its holdings. 1/4 in the other species. Paratypes: Four preserved specimens at the Bernice P. Bishop The species A.brettbarnetti sp. nov. from Woodlark Island and A. Museum, Honolulu, Hawaii, USA, specimen number: BPBM Herp- stevebennetti sp. nov. from Boiaboiawaga Island, both in south- BPBM 15844-15847 also collected at Boiaboiawaga Island, Milne east Papua New Guinea have until now been identified as A. Bay Province, Papua New Guinea, Latitude -10.21 S., Longitude novaeguineae (Brown and Parker, 1977) and are similar to it. The 150.90 E. can both be separated from that taxon and others in the genus by marbling on the nape and anterior dorsum as well as tail bands Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that have the darker segments darker posteriorly and fading Britain, is similar in most respects to the species A. novaeguineae anteriorly. A. stevebennetti sp. nov. is further separated from the (Brown and Parker, 1977) and would be identified as this taxon other species (A.brettbarnetti sp. nov.) by scattered dark flecks on using the criteria given in that paper. A. adelynhoserae sp. nov. is the whitish surface of the lower flanks. however readily separated from A. novaeguineae by the general absence of reddish-brown or red flecks and markings on the dorsal The species A. lucybennettae sp. nov. from the vicinity of Utai surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 Village, Sandaun Province, in northern Papua New Guinea and scattered yellow dots on each of the lower flanks above where the and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East grayish upper body turns to the lighter undersurface. The well- Sepik Province, Papua New Guinea are similar in most respects to defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. A. novaeguineae (Brown and Parker, 1977) and until now would are whiteish and grayish in colour versus tan and reddish-brown in have been identified as that taxon, but can be separated from it by A. novaeguineae. longer and narrower digits (long and narrow versus moderate). A. lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. The species A. judyfergusonae sp. nov. from Unea Island, is nov. by the presence of about four well defined and irregular separated from A. adelynhoserae sp. nov. by the relative lack of shaped dorsal crossbands of dark and light grey, the dark bands black peppering in the lighter bands as compared to that seen in A. becoming darker on the lower flanks. adelynhoserae sp. nov., which is readily noticeable. The limited number of blackish spots on the dorsum and flanks of A. Adelynhosergecko huonensis sp. nov. is similar in most respects to adelynhoserae sp. nov. are indistinct, versus prominent in A. A. novaeguineae Brown and Parker (1977), but is separated from judyfergusonae sp. nov..

Otherwise both species are similar. indistinct dorsal pattern with distinctly jagged edges on the upper The species A. haydnmcphiei sp. nov. from Ambitle Island, is surface sometimes consisting of bands or markings indicative of readily separated from both A. adelynhoserae sp. nov.and A. this. judyfergusonae sp. nov.by an absence of yellow mottling on the Geckos in the genus Adelynhosergecko gen. nov. are readily back of the jaw and sides of neck. Striping behind the eye is also separated from geckos in the genera Allengreercolotes gen. nov., noticeable in A. haydnmcphiei sp. nov.. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by The species A. matteoae sp. nov. from the Kei Islands, is similar in having fewer than 25 enlarged preanal/femoral pores, snout-vent most respects to A. novaeguineae (Brown and Parker, 1977), but length in adults of under 41 mm and of moderate build. is separated from it by a diffuse dull beige indististinctly marked Exceptional to this in terms of size, Adelynhosergecko sloppi sp. colouration with scattered yellowish flecks on the dorsum. nov. does exceed 41 mm in snout-vent length. The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, Distribution: Only known from the type locality of Boiaboiawaga Irian Jaya, Indonesia, is separated from all other geckos in the Island, Milne Bay Province, Papua New Guinea, but probably also genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., occurs on the immediately adjacent larger islands and/or nearby Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by New Guinea mainland. having fewer than 25 enlarged preanal/femoral pores and an adult Etymology: Named in honour of Steve Bennett of Narre Warren, snout-vent length in excess of 42 mm and robust build. It is Melbourne, Victoria, Australia in recognition of his immense morphologically similar in most respects to species within the contributions to herpetology and aviculture over some decades, genus Adelynhosergecko gen. nov. in particular Adelynhosergecko including logistical support for the wildlife education business novaeguineae (Brown and Parker, 1977), but differs from this taxon Snakebusters: Australia’s best reptiles. by its larger adult size (in excess of 42 mm snout-vent, versus ADELYNHOSERGECKO LUCYBENNETTAE SP. NOV. under 41 mm) and more robust build. Holotype: A preserved specimen at the Louisiana State University All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. Museum of Natural Science, Baton Rouge, USA, LSUMZ Herps haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily Collection, specimen number: LSUMZ 95849, collected from Utai separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis Village, 2 km south of the airfield, Sandaun Province, Papua New sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. Guinea, Latitude -3.40 S., Longitude 141.58 E. This facility allows stevebennetti sp. nov, A. lucybennettae sp. nov. and A. access to its holdings. lachlanmcpheei sp. nov. by having no webbing or less than 1/5 Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New webbing between digits 3 and 4 of the hind leg, versus from 1/5 to Britain, is similar in most respects to the species A. novaeguineae 1/4 in the other species. (Brown and Parker, 1977) and would be identified as this taxon The species A.brettbarnetti sp. nov. from Woodlark Island and A. using the criteria given in that paper. A. adelynhoserae sp. nov. is stevebennetti sp. nov. from Boiaboiawaga Island, both in south- however readily separated from A. novaeguineae by the general east Papua New Guinea have until now been identified as A. absence of reddish-brown or red flecks and markings on the dorsal novaeguineae (Brown and Parker, 1977) and are similar to it. They surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 can both be separated from that taxon and others in the genus by scattered yellow dots on each of the lower flanks above where the marbling on the nape and anterior dorsum as well as tail bands grayish upper body turns to the lighter undersurface. The well- that have the darker segments darker posteriorly and fading defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. anteriorly. A. stevebennetti sp. nov. is further separated from the are whiteish and grayish in colour versus tan and reddish-brown in other species (A.brettbarnetti sp. nov.) by scattered dark flecks on A. novaeguineae. the whitish surface of the lower flanks. The species A. judyfergusonae sp. nov. from Unea Island, is The species A. lucybennettae sp. nov. from the vicinity of Utai separated from A. adelynhoserae sp. nov. by the relative lack of Village, Sandaun Province, in northern Papua New Guinea and black peppering in the lighter bands as compared to that seen in A. and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East adelynhoserae sp. nov., which is readily noticeable. The limited Sepik Province, Papua New Guinea are similar in most respects to number of blackish spots on the dorsum and flanks of A. A. novaeguineae (Brown and Parker, 1977) and until now would adelynhoserae sp. nov. are indistinct, versus prominent in A. have been identified as that taxon, but can be separated from it by judyfergusonae sp. nov.. longer and narrower digits (long and narrow versus moderate). A. Otherwise both species are similar. lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. The species A. haydnmcphiei sp. nov. from Ambitle Island, is nov. by the presence of about four well defined and irregular readily separated from both A. adelynhoserae sp. nov.and A. shaped dorsal crossbands of dark and light grey, the dark bands judyfergusonae sp. nov.by an absence of yellow mottling on the becoming darker on the lower flanks. back of the jaw and sides of neck. Striping behind the eye is also Adelynhosergecko huonensis sp. nov. is similar in most respects to noticeable in A. haydnmcphiei sp. nov.. A. novaeguineae Brown and Parker (1977), but is separated from The species A. matteoae sp. nov. from the Kei Islands, is similar in that species by having a grey dorsal colouration versus reddish- most respects to A. novaeguineae (Brown and Parker, 1977), but brown in both A. novaeguineae and A. madangensis sp. nov. as is separated from it by a diffuse dull beige indististinctly marked described within this paper. A. huonensis sp. nov. has vague as colouration with scattered yellowish flecks on the dorsum. opposed to moderately distinct dorsal markings as seen in the other two species. The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, Irian Jaya, Indonesia, is separated from all other geckos in the A. madangensis sp. nov. is further separated from A. genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., novaeguineae and A. huonensis sp. nov. by the presence of an Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by indistinct dorsal pattern with distinctly jagged edges on the upper having fewer than 25 enlarged preanal/femoral pores and an adult surface sometimes consisting of bands or markings indicative of snout-vent length in excess of 42 mm and robust build. It is this. morphologically similar in most respects to species within the Adelynhosergecko huonensis sp. nov. is similar in most respects to genus Adelynhosergecko gen. nov. in particular Adelynhosergecko A. novaeguineae Brown and Parker (1977), but is separated from novaeguineae (Brown and Parker, 1977), but differs from this taxon that species by having a grey dorsal colouration versus reddish- by its larger adult size (in excess of 42 mm snout-vent, versus brown in both A. novaeguineae and A. madangensis sp. nov. as under 41 mm) and more robust build. described within this paper. A. huonensis sp. nov. has vague as All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. opposed to moderately distinct dorsal markings as seen in the haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily other two species. separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis A. madangensis sp. nov. is further separated from A. sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. novaeguineae and A. huonensis sp. nov. by the presence of an stevebennetti sp. nov, A. lucybennettae sp. nov. and A.

Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 56 Australasian Journal of Herpetology lachlanmcpheei sp. nov. by having no webbing or less than 1/5 Britain, is similar in most respects to the species A. novaeguineae webbing between digits 3 and 4 of the hind leg, versus from 1/5 to (Brown and Parker, 1977) and would be identified as this taxon 1/4 in the other species. using the criteria given in that paper. A. adelynhoserae sp. nov. is The species A.brettbarnetti sp. nov. from Woodlark Island and A. however readily separated from A. novaeguineae by the general stevebennetti sp. nov. from Boiaboiawaga Island, both in south- absence of reddish-brown or red flecks and markings on the dorsal east Papua New Guinea have until now been identified as A. surface and sides. A. adelynhoserae sp. nov. instead has 4 to 6 novaeguineae (Brown and Parker, 1977) and are similar to it. They scattered yellow dots on each of the lower flanks above where the can both be separated from that taxon by marbling on the nape grayish upper body turns to the lighter undersurface. The well- and anterior dorsum as well as tail bands that have the darker defined dorsal cross-bands on the tail of A. adelynhoserae sp. nov. segments darker posteriorly and fading anteriorly. A. stevebennetti are whiteish and grayish in colour versus tan and reddish-brown in sp. nov. is further separated from the other species (A.brettbarnetti A. novaeguineae. sp. nov.) by scattered dark flecks on the whitish surface of the The species A. judyfergusonae sp. nov. from Unea Island, is lower flanks. separated from A. adelynhoserae sp. nov. by the relative lack of The species A. lucybennettae sp. nov. from the vicinity of Utai black peppering in the lighter bands as compared to that seen in A. Village, Sandaun Province, in northern Papua New Guinea and adelynhoserae sp. nov., which is readily noticeable. The limited and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East number of blackish spots on the dorsum and flanks of A. Sepik Province, Papua New Guinea are similar in most respects to adelynhoserae sp. nov. are indistinct, versus prominent in A. A. novaeguineae (Brown and Parker, 1977) and until now would judyfergusonae sp. nov.. have been identified as that taxon, but can be separated from it by Otherwise both species are similar. longer and narrower digits (long and narrow versus moderate). A. The species A. haydnmcphiei sp. nov. from Ambitle Island, is lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. readily separated from both A. adelynhoserae sp. nov.and A. nov. by the presence of about four well defined and irregular judyfergusonae sp. nov.by an absence of yellow mottling on the shaped dorsal crossbands of dark and light grey, the dark bands back of the jaw and sides of neck. Striping behind the eye is also becoming darker on the lower flanks. noticeable in A. haydnmcphiei sp. nov.. Adelynhosergecko huonensis sp. nov. is similar in most respects to The species A. matteoae sp. nov. from the Kei Islands, is similar in A. novaeguineae Brown and Parker (1977), but is separated from most respects to A. novaeguineae (Brown and Parker, 1977), but that species by having a grey dorsal colouration versus reddish- is separated from it by a diffuse dull beige indististinctly marked brown in both A. novaeguineae and A. madangensis sp. nov. as colouration with scattered yellowish flecks on the dorsum. described within this paper. A. huonensis sp. nov. has vague as The species Adelynhosergecko sloppi sp. nov. from Waigeo Island, opposed to moderately distinct dorsal markings as seen in the Irian Jaya, Indonesia, is separated from all other geckos in the other two species. genera Adelynhosergecko gen. nov., Allengreercolotes gen. nov., A. madangensis sp. nov. is further separated from A. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by novaeguineae and A. huonensis sp. nov. by the presence of an having fewer than 25 enlarged preanal/femoral pores and an adult indistinct dorsal pattern with distinctly jagged edges on the upper snout-vent length in excess of 42 mm and robust build. It is surface sometimes consisting of bands or markings indicative of morphologically similar in most respects to species within the this. genus Adelynhosergecko gen. nov. in particular Adelynhosergecko Adelynhosergecko huonensis sp. nov. is similar in most respects to novaeguineae (Brown and Parker, 1977), but differs from this taxon A. novaeguineae Brown and Parker (1977), but is separated from by its larger adult size (in excess of 42 mm snout-vent, versus that species by having a grey dorsal colouration versus reddish- under 41 mm) and more robust build. brown in both A. novaeguineae and A. madangensis sp. nov. as All of adelynhoserae sp. nov., A. judyfergusonae sp. nov., A. described within this paper. A. huonensis sp. nov. has vague as haydnmcphiei sp. nov. and A. matteoae sp. nov.are also readily opposed to moderately distinct dorsal markings as seen in the separated from A. novaeguineae, A. sloppi sp. nov., A. huonensis other two species. sp. nov., A. madangensis sp. nov., A. brettbarnetti sp. nov., A. A. madangensis sp. nov. is further separated from A. stevebennetti sp. nov, A. lucybennettae sp. nov. and A. novaeguineae and A. huonensis sp. nov. by the presence of an lachlanmcpheei sp. nov. by having no webbing or less than 1/5 indistinct dorsal pattern with distinctly jagged edges on the upper webbing between digits 3 and 4 of the hind leg, versus from 1/5 to surface sometimes consisting of bands or markings indicative of 1/4 in the other species. this. The species A.brettbarnetti sp. nov. from Woodlark Island and A. Geckos in the genus Adelynhosergecko gen. nov. are readily stevebennetti sp. nov. from Boiaboiawaga Island, both in south- separated from geckos in the genera Allengreercolotes gen. nov., east Papua New Guinea have until now been identified as A. Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by novaeguineae (Brown and Parker, 1977) and are similar to it. They having fewer than 25 enlarged preanal/femoral pores, snout-vent can both be separated from that taxon by marbling on the nape length in adults of under 41 mm and of moderate build. and anterior dorsum as well as tail bands that have the darker Exceptional to this in terms of size, Adelynhosergecko sloppi sp. segments darker posteriorly and fading anteriorly. A. stevebennetti nov. does exceed 41 mm in snout-vent length. sp. nov. is further separated from the other species (A.brettbarnetti Distribution: Only known from the type locality and immediate sp. nov.) by scattered dark flecks on the whitish surface of the surrounds of the vicinity of Utai Village, 2 km south of the airfield, in lower flanks. Sandaun Province, Papua New Guinea The species A. lucybennettae sp. nov. from the vicinity of Utai Etymology: Named in honour of Lucy Bennett of Narre Warren, Village, Sandaun Province, in northern Papua New Guinea and Melbourne, Victoria, Australia in recognition of her immense and A. lachlanmcpheei sp. nov. from the vicinity of Weewak in East contributions to herpetology over some decades, including Sepik Province, Papua New Guinea are similar in most respects to logistical support for the wildlife education business Snakebusters: A. novaeguineae (Brown and Parker, 1977) and until now would Australia’s best reptiles. have been identified as that taxon, but can be separated from it by ADELYNHOSERGECKO LACHLANMCPHEEI SP. NOV. longer and narrower digits (long and narrow versus moderate). A. Holotype: A preserved specimen at the Louisiana State University lachlanmcpheei sp. nov. is separated from A. lucybennettae sp. Museum of Natural Science, Baton Rouge, USA, LSUMZ Herps nov. by the presence of about four well defined and irregular Collection, specimen number: LSUMZ 97474, collected from SIL shaped dorsal crossbands of dark and light grey, the dark bands guest house grounds and vicinity, Kreer Heights, Wewak, Papua becoming darker on the lower flanks. New Guinea, Latitude -3.59 S., Longitude 143.64 E. This facility Adelynhosergecko huonensis sp. nov. is similar in most respects to allows access to its holdings. A. novaeguineae Brown and Parker (1977), but is separated from Diagnosis: Adelynhosergecko adelynhoserae sp. nov. from New that species by having a grey dorsal colouration versus reddish-

brown in both A. novaeguineae and A. madangensis sp. nov. as and Böhme, 2012, also placed in the subgenus of Scelotretus described within this paper. A. huonensis sp. nov. has vague as Fitzinger, 1843, namely Scelotretus is separated from other opposed to moderately distinct dorsal markings as seen in the species complex members (the subgenus) by a pattern consisting other two species. of an irregularly flecked pattern on the head, body and limbs. A. madangensis sp. nov. is further separated from A. Sometimes a light vertebral stripe densely interspersed with darker novaeguineae and A. huonensis sp. nov. by the presence of an flecks is discernible. Two dark parallel paravertebral sacral stripes indistinct dorsal pattern with distinctly jagged edges on the upper framing a lighter sacral are characteristic. Tail above has short surface sometimes consisting of bands or markings indicative of irregular dark stripes and flecks, below monochromatic, whitish, this. medially with narrow, dark streaks. Subadults with all characters of Adelynhosergecko huonensis sp. nov. is similar in most respects to the adults. All nuchal tubercles round to oval, conical, pointed and A. novaeguineae Brown and Parker (1977), but is separated from irregularly arranged. that species by having a grey dorsal colouration versus reddish- The species S. haroldcoggeri sp. nov. with a centre of distribution brown in both A. novaeguineae and A. madangensis sp. nov. as being the eastern Solomon Islands, is morphologically similar in described within this paper. A. huonensis sp. nov. has vague as most respects to S. remotus, including in colouration and size, opposed to moderately distinct dorsal markings as seen in the shape and configuration of nuchal tubercles. It is separated from other two species. that species however by the presence of large irregular light A. madangensis sp. nov. is further separated from A. greenish blotches on the dorsal surface of the body, a head that is novaeguineae and A. huonensis sp. nov. by the presence of an darker than the body (versus lighter in S. remotus) and scattered indistinct dorsal pattern with distinctly jagged edges on the upper large light green spots on the snout, versus none in S. remotus. surface sometimes consisting of bands or markings indicative of The species S. daranini sp. nov. known only from the Kei Islands in this. the Moluccas, Indonesia conforms with the so called S. vittatus Geckos in the genus Adelynhosergecko gen. nov. are readily phenotype 3 as broadly defined by Rösler et al. (2012). separated from geckos in the genera Allengreercolotes gen. nov., This species is similar in most respects to nominate S. vittatus, but Borneocolotes gen. nov. and Rosssadliercolotes gen. nov. by is readily separated from it by having large nuchal tubercles having fewer than 25 enlarged preanal/femoral pores, snout-vent (versus medium) and in S. daranini sp. nov. they are distinctly length in adults of under 41 mm and of moderate build. conical shaped and pointed. Exceptional to this in terms of size, Adelynhosergecko sloppi sp. The species S. jenandersonae sp. nov. known only from Nissan nov. does exceed 41 mm in snout-vent length. Atoll, Green Islands, Papua New Guinea, more-or-less conforms Distribution: Only known from the type locality of the vicinity of with the so called S. vittatus phenotype 4 as broadly defined by Wewak, Papua New Guinea. Etymology: Named in honour of Rösler et al. (2012). This species is superficially similar to S. Lachlan McPhee of Frankston, Victoria, Australia in recognition of haroldcoggeri sp. nov. and was mistakenly identified as this taxon his significant contributions to the welfare of underprivileged and by Rösler et al. (2012), who included both as one phenotype in disabled people in Melbourne, Victoria, Australia over many years. their description of their phenotype 4. SCELOTRETUS (HAROLDCOGGERCOLOTES) However S. jenandersonae sp. nov., including as depicted by HAROLDCOGGERI SP. NOV. Rösler et al. (2012) differs from S. haroldcoggeri sp. nov. (as Holotype: A preserved specimen in the Australian Museum in depicted by McCoy (2006) at plate 15) by having a brownish Sydney, New South Wales, Australia, specimen number: R.42095, dorsum with numerous grayish black flecks, versus grayish-brown collected at Santa Ana Island, Solomon Islands, Latitude -10.83 S., with irregular green spots and not the peppered grayish black Longitude 162.46 E. This government-owned facility allows access flecks. to its holdings. At the rear of the body and the base of the tail in S. jenandersonae Paratype: A preserved specimen in the Australian Museum in sp. nov. is an obvious dark blackish mid-vertebral stripe formed by Sydney, New South Wales, Australia, specimen number: a concentration of dark flecks. This is not seen in S. haroldcoggeri R.127285, collected at Makira Island, San Cristobal Island, sp. nov.. Solomon Islands, Latitude -10.83 S., Longitude 162.46 E. The limbs and tail of S. jenandersonae sp. nov. are clearly Diagnosis: The species Gekko vittatus Houttuyn, 1782, herein peppered with black flecks. This is not the case in S. placed in the genus Scelotretus Fitzinger, 1843 subgenus haroldcoggeri sp. nov. or for that matter any other species in the Scelotretus conforms broadly with the so called G. vittatus subgenus. The closest similarity to the flecks on the limbs and tail phenotype 1 as broadly defined by Rösler et al. (2012) and as is dark brown marbling on the legs of S. daranini sp. nov.. described in Boulenger (1885) at pages 185-186. Distribution: S. haroldcoggeri sp. nov. is restricted to the Eastern The subspecies described as “Var. bivittatus” on pages 186-187 of Solomon Islands and the Vanuatu Islands. Boulenger (1885) describes a composite of species and the name Etymology: Named in honour of Dr. Harold G. Cogger, former bivittatus as originally applied by Duméril and Bibron in 1836 deputy director and curator of herpetology at the Australian applies either to typical S. vittatus or to a north-west New Guinea Museum in Sydney, New South Wales, Australia in recognition of a specimen of a different species-level taxon (Rösler et al. 2012) and lifetime’s high achievement in herpetology. is therefore not applicable or available for the three newly named SCELOTRETUS (HAROLDCOGGERCOLOTES) DARANINI SP. species in this subgenus as formally described in this paper. NOV. A modified diagnosis of the species S. vittatus is as follows: Holotype: A preserved male specimen in the Zoologisches S. vittatus is separated from the others in the species complex Museum, Berlin, now Museum für (subgenus) by a colouration including a distinct light vertebral Naturkunde, Berlin, Germany, specimen number: ZMB 48737 stripe, several millimetres wide and bordered with dark brown, collected from the Kei islands, Moluccas, Indonesia. This facility bifurcating in the neck region and usually reaching the posterior allows access to its holdings. margin of the eyes. The stripe may be interrupted or shortened, but Paratype: A preserved male specimen in the Zoologisches even when faded is discernable. The vertebral stripe slightly Museum, Berlin, now Museum für widens more or less saddle-like in the anterior caudal region. The Naturkunde, Berlin, Germany, specimen number: ZMB 48738 tail is annulated with sharply defined light and dark rings, the dark collected from the Kei islands, Moluccas, Indonesia. rings are usually twice as broad as the light ones. Juveniles are Diagnosis: The species Gekko vittatus Houttuyn, 1782, herein similar to adults, but with more intensive caudal colouration. placed in the genus Scelotretus Fitzinger, 1843 subgenus Roundish to oval pointed tubercles present only within the white Scelotretus conforms broadly with the so called G. vittatus rami of the vertebral band, all remaining nuchal tubercles are blunt phenotype 1 as broadly defined by Rösler et al. (2012) and as and slightly convex. described in Boulenger (1885) at pages 185-186. The species described as Gekko remotus Rösler, Ineich, Wilms The subspecies described as “Var. bivittatus” on pages 186-187 of Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.net Copyright- Kotabi Publishing - All rights reserved 58 Australasian Journal of Herpetology

Boulenger (1885) describes a composite of species and the name Etymology: Named in honour of Dara Nin, of Ringwood, bivittatus as originally applied by Duméril and Bibron in 1836 Melbourne, Victoria, Australia in recognition of his contributions to applies either to typical S. vittatus or to a north-west New Guinea herpetology in Australia, in particular through his work with specimen of a different species-level taxon (Rösler et al. 2012) and Snakebusters, Australia’s best reptiles shows, educational displays is therefore not applicable or available for the three newly named and reptile parties. See Hoser (2014) for details. species in this subgenus as formally described in this paper. SCELOTRETUS (HAROLDCOGGERCOLOTES) A modified diagnosis of the species S. vittatus is as follows: JENANDERSONAE SP. NOV. S. vittatus is separated from the others in the species complex Holotype: A preserved male specimen at the (subgenus) by a colouration including a distinct light vertebral Senckenbergmuseum, Frankfurt am Main, now Forschungsinstitut stripe, several millimetres wide and bordered with dark brown, Senckenberg, Frankfurt, Germany, specimen number: SMF 9157, bifurcating in the neck region and usually reaching the posterior collected from Nissan atoll, Green Islands, Papua New Guinea. margin of the eyes. The stripe may be interrupted or shortened, but This facility allows access to its holdings. even when faded is discernable. The vertebral stripe slightly Paratypes: Two preserved male specimens at the widens more or less saddle-like in the anterior caudal region. The Senckenbergmuseum, Frankfurt am Main, now Forschungsinstitut tail is annulated with sharply defined light and dark rings, the dark Senckenberg, Frankfurt, Germany, specimen numbers: SMF 9158 rings are usually twice as broad as the light ones. Juveniles are and SMF 9159 collected from Nissan atoll, Green Islands, Papua similar to adults, but with more intensive caudal colouration. New Guinea. Roundish to oval pointed tubercles present only within the white Diagnosis: The species Gekko vittatus Houttuyn, 1782, herein rami of the vertebral band, all remaining nuchal tubercles are blunt placed in the genus Scelotretus Fitzinger, 1843 subgenus and slightly convex. Scelotretus conforms broadly with the so called G. vittatus The species described as Gekko remotus Rösler, Ineich, Wilms phenotype 1 as broadly defined by Rösler et al. (2012) and as and Böhme, 2012, also placed in the subgenus of Scelotretus described in Boulenger (1885) at pages 185-186. Fitzinger, 1843, namely Scelotretus is separated from other The subspecies described as “Var. bivittatus” on pages 186-187 of species complex members (the subgenus) by a pattern consisting Boulenger (1885) describes a composite of species and the name of an irregularly flecked pattern on the head, body and limbs. bivittatus as originally applied by Duméril and Bibron in 1836 Sometimes a light vertebral stripe densely interspersed with darker applies either to typical S. vittatus or to a north-west New Guinea flecks is discernible. Two dark parallel paravertebral sacral stripes specimen of a different species-level taxon (Rösler et al. 2012) and framing a lighter sacral are characteristic. Tail above has short is therefore not applicable or available for the three newly named irregular dark stripes and flecks, below monochromatic, whitish, species in this subgenus as formally described in this paper. medially with narrow, dark streaks. Subadults with all characters of A modified diagnosis of the species S. vittatus is as follows: the adults. All nuchal tubercles round to oval, conical, pointed and S. vittatus is separated from the others in the species complex irregularly arranged. (subgenus) by a colouration including a distinct light vertebral The species S. haroldcoggeri sp. nov. with a centre of distribution stripe, several millimetres wide and bordered with dark brown, being the eastern Solomon Islands, is morphologically similar in bifurcating in the neck region and usually reaching the posterior most respects to S. remotus, including in colouration and size, margin of the eyes. The stripe may be interrupted or shortened, but shape and configuration of nuchal tubercles. It is separated from even when faded is discernable. The vertebral stripe slightly that species however by the presence of large irregular light widens more or less saddle-like in the anterior caudal region. The greenish blotches on the dorsal surface of the body, a head that is tail is annulated with sharply defined light and dark rings, the dark darker than the body (versus lighter in S. remotus) and scattered rings are usually twice as broad as the light ones. Juveniles are large light green spots on the snout, versus none in S. remotus. similar to adults, but with more intensive caudal colouration. The species S. daranini sp. nov. known only from the Kei Islands in Roundish to oval pointed tubercles present only within the white the Moluccas, Indonesia conforms with the so called S. vittatus rami of the vertebral band, all remaining nuchal tubercles are blunt phenotype 3 as broadly defined by Rösler et al. (2012). and slightly convex. This species is similar in most respects to nominate S. vittatus, but The species described as Gekko remotus Rösler, Ineich, Wilms is readily separated from it by having large nuchal tubercles and Böhme, 2012, also placed in the subgenus of Scelotretus (versus medium) and in S. daranini sp. nov. they are distinctly Fitzinger, 1843, namely Scelotretus is separated from other conical shaped and pointed. species complex members (the subgenus) by a pattern consisting The species S. jenandersonae sp. nov. known only from Nissan of an irregularly flecked pattern on the head, body and limbs. Atoll, Green Islands, Papua New Guinea, more-or-less conforms Sometimes a light vertebral stripe densely interspersed with darker with the so called S. vittatus phenotype 4 as broadly defined by flecks is discernible. Two dark parallel paravertebral sacral stripes Rösler et al. (2012). This species is superficially similar to S. framing a lighter sacral are characteristic. Tail above has short haroldcoggeri sp. nov. and was mistakenly identified as this taxon irregular dark stripes and flecks, below monochromatic, whitish, by Rösler et al. (2012), who included both as one phenotype in medially with narrow, dark streaks. Subadults with all characters of their description of their phenotype 4. the adults. All nuchal tubercles round to oval, conical, pointed and However S. jenandersonae sp. nov., including as depicted by irregularly arranged. Rösler et al. (2012) differs from S. haroldcoggeri sp. nov. (as The species S. haroldcoggeri sp. nov. with a centre of distribution depicted by McCoy (2006) at plate 15) by having a brownish being the eastern Solomon Islands, is morphologically similar in dorsum with numerous grayish black flecks, versus grayish-brown most respects to S. remotus, including in colouration and size, with irregular green spots and not the peppered grayish black shape and configuration of nuchal tubercles. It is separated from flecks. that species however by the presence of large irregular light At the rear of the body and the base of the tail in S. jenandersonae greenish blotches on the dorsal surface of the body, a head that is sp. nov. is an obvious dark blackish mid-vertebral stripe formed by darker than the body (versus lighter in S. remotus) and scattered a concentration of dark flecks. This is not seen in S. haroldcoggeri large light green spots on the snout, versus none in S. remotus. sp. nov.. The species S. daranini sp. nov. known only from the Kei Islands in The limbs and tail of S. jenandersonae sp. nov. are clearly the Moluccas, Indonesia conforms with the so called S. vittatus peppered with black flecks. This is not the case in S. phenotype 3 as broadly defined by Rösler et al. (2012). haroldcoggeri sp. nov. or for that matter any other species in the This species is similar in most respects to nominate S. vittatus, but subgenus. The closest similarity to the flecks on the limbs and tail is readily separated from it by having large nuchal tubercles is dark brown marbling on the legs of S. daranini sp. nov.. (versus medium) and in S. daranini sp. nov. they are distinctly Distribution: S. daranini sp. nov. is restricted to the the Kei conical shaped and pointed. islands, Moluccas, Indonesia. The species S. jenandersonae sp. nov. known only from Nissan

Atoll, Green Islands, Papua New Guinea, more-or-less conforms Boettger, O. 1897. Neue Reptilien und Batrachier von den with the so called S. vittatus phenotype 4 as broadly defined by Philippinen. Zool. Anz. 20:161-166. Rösler et al. (2012). This species is superficially similar to S. Boissinot, S., Ineich, I., Thaler, L. and Guillaume, C. 1997. Hybrid haroldcoggeri sp. nov. and was mistakenly identified as this taxon origin and clonal diversity in the parthenogenetic gecko, by Rösler et al. (2012), who included both as one phenotype in Lepidodactylus lugubris in French Polynesia. Journal of their description of their phenotype 4. Herpetology 31(2): 295-298. However S. jenandersonae sp. nov., including as depicted by Bottom, R. 1969. Behind the Barrier. Gareth Powell Associates, Rösler et al. (2012) differs from S. haroldcoggeri sp. nov. (as Gladesville, N.S.W., Australia. depicted by McCoy (2006) at plate 15) by having a brownish Bottom, R. 1979. The Godfather in Australia: Organised Crime’s dorsum with numerous grayish black flecks, versus grayish-brown Australian Connections. A. H. and A. W. Reed, Terrey Hills, with irregular green spots and not the peppered grayish black N.S.W., Australia. flecks. Bottom, R. 1984. Without Fear or Favour. Sun Books, South At the rear of the body and the base of the tail in S. jenandersonae Melbourne, Australia. sp. nov. is an obvious dark blackish mid-vertebral stripe formed by Bottom, R. 1985. Connections: Crime Rackets and Networks of a concentration of dark flecks. This is not seen in S. haroldcoggeri Influence Down-Under. Sun Books, South Melbourne, Australia. sp. nov.. Bottom, R. 1987. Connections II: Crime Rackets and Networks of The limbs and tail of S. jenandersonae sp. nov. are clearly Influence in Australia. Sun Books, South Melbourne, Australia. peppered with black flecks. This is not the case in S. Bottom, R. 1988. Shadow of Shame: How the Mafia Got Away with haroldcoggeri sp. nov. or for that matter any other species in the the Murder of Donald Mackay. Sun Books, South Melbourne, subgenus. The closest similarity to the flecks on the limbs and tail Australia. is dark brown marbling on the legs of S. daranini sp. nov.. Bottom, R. 1989. 7/ Bugged! : Legal Police Telephone Taps Distribution: S. jenandersonae sp. nov. is only known from the Expose the Mr Bigs of Australia’s Drug Trade. Sun Books, South Nissan atoll, Green Islands, Papua New Guinea. Melbourne, Australia. Etymology: Named in honour of Jen Anderson, of Ringwood, Bottom, R. 2009. Fighting Organised Crime: Triumph and Betrayal Melbourne, Victoria, Australia in recognition of her contributions to in a Lifelong Campaign. BBP, Nelson Bay, N.S.W. Australia. herpetology in Australia, in particular through her work with Snakebusters, Australia’s best reptiles shows, educational displays Boulenger, G. A. 1883. On the geckos of New Calendonia. Proc. and reptile parties. Also see Hoser (2014) for relevant details. Zool. Soc. London 1883:116-130. REFERENCES CITED Boulenger, G. A. 1884. Diagnoses of new reptiles and batrachians from the Solomon Islands, collected and presented to the British Amarasinghe, A. A. T., Bauer, A. M., Ineich, I., Rudge, J., Bahir, M. Museum by H. B. Guppy, Esq., M. B., H. M. S. ‘Lark.’. Proc. Zool. M. and Gabadage, D. E. 2009. The original descriptions and Soc. London 1884:210-213. figures of Sri Lankan gekkonid lizards (Squamata: Gekkonidae) of the 18th, 19th and 20th centuries. Taprobanica 1(2):83-106. Boulenger, G. A. 1885a. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Auffenberg, W. 1980. The herpetofauna of Komodo, with notes on Pygopodidae, Agamidae. London: 450 pp. adjacent areas. Bulletin of the Florida State Museum 25(2):39-156. Boulenger, G. A. 1885b. Descriptions of three new species of Barnett, L. K. and Emms, C. 1997. Herpetological observations in geckoes. Ann. Mag. nat. Hist. (5)16:473-475. the Chagos Archipelago, British Indian Ocean Territory. British Herpetological Society Bulletin (59):6-12. Boulenger, G. A. 1886. On the reptiles and batrachians of the Solomon Islands. Trans. Zool. Soc. London 12:35-62. Bauer, A. M. 1994. Liste der rezenten Amphibien und Reptilien: Gekkonidae I (Australia). Das Tierreich 108, W. de Gruyter und Co. Boulenger, G. A. 1887a. Catalogue of the Lizards in the British (Berlin). Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London:575pp. Bauer, A. M. and Henle, K. 1994. Familia Gekkonidae (Reptilia, Sauria). Part I. Australia and Oceania. Das Tierreich, 109. Walter Boulenger, G. A. 1887b. Second contribution to the herpetology of de Gruyter, Berlin. the Solomon islands. Proc. Zool. Soc. London 1887:333-340. Bauer, A. M. and Sadlier, R. A. 1994. The terrestrial herpetofauna Boulenger, G. A. 1889. On the reptiles of Christmas Island. Proc. of the Ile des Pins, New Caledonia. Pacific Science 48(4):353-366. Zool. Soc. London 1888:534-536. Bauer, A. M. and Sadlier, R. A. (eds.) 2000. The herpetofauna of Boulenger, G. A. 1897. On the reptiles of Rotuma Island, New Caledonia. Contributions to Herpetology, 17; Society for Study Polynesia. Ann. Mag. Nat. Hist. (6)20:306-307. Amphibians and Reptiles, Ithaca, New York. Boulenger, G. A. 1920. Description of a new gecko and a new Bauer, A. M. and Vindum, J. V. 1990. A checklist and key to the snake from Sumatra. Ann. Mag. nat. Hist. (9) 5:281-283. herpetofauna of New Caledonia, with remarks on biogeography. Brongersma, L. D. 1934. Contributions to Indo-Australian Proc. Cal. Acad. Sci. 47(2):17-45. herpetology. Zoologische Mededelingen 17: 161-251. Bauer, A. M., Günther, R. and Klipfel, M. 1995. The herpetological Brongersma, L. D. 1948. Lizards from the island of Morotai contributions of Wilhelm C.H. Peters (1815-1883). 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Two new species of True Cobra in the genus Pseudogekko Taylor. The Philippine Journal of Science 95:331- Boulengeria Dollo, 1886 from West Africa and South Africa 352. (Serpentes: Elapidae). Australasian Journal of Herpetology 20:3-7. Kopstein, P. F. 1926. Reptilien von den Molukken und den Hoser, R. T. 2014. A logical new taxonomy for the Asian subfamily benachbarten Inseln. Zoologische Mededelingen 1:71-112. Draconinae based on obvious phylogenetic relationships and morphology of species (Squamata: Sauria: Agamidae: Koch, A. 2011. The Amphibians and Reptiles of Sulawesi: Draconinae). Australasian Journal of Herpetology 22:9-59. Underestimated Diversity in a Dynamic Environment. In: Zachos, F.E. and Habel, J.C. (eds.), Biodiversity Hotspots. Springer, Berlin, Hoser, R. T. 2015a. Dealing with the “truth haters” ... a summary! p. 383-404. Introduction to Issues 25 and 26 of Australasian Journal of Herpetology. Including “A timeline of relevant key publishing and Koch, A. 2012. Discovery, Diversity, and Distribution of the other events relevant to Wolfgang Wüster and his gang of thieves.” Amphibians and Reptiles of Sulawesi and its offshore islands. and a “Synonyms list”. Australasian Journal of Herpetology 25:3- Edition Chimaira:374 pp. 13. Lenort, J. 2004. Haltung und Nachzucht des philippinischen Hoser, R. T. 2015b. The Wüster gang and their proposed “Taxon Smaragdgeckos Pseudogekko smaragdinus (Taylor, 1922). Sauria, Filter”: How they are knowingly publishing false information, Berlin 26(3):7-12. recklessly engaging in taxonomic vandalism and directly attacking Loveridge, A. 1948. New Guinean reptiles and amphibians in the the rules and stability of zoological nomenclature. Australasian Museum of Comparative Zoology and United States National Journal of Herpetology 25:14-38. Museum. Bull. Mus. Comp. Zool. Harvard 101(2):305-430. Hoser, R. T. 2015c. Best Practices in herpetology: Hinrich Kaiser’s Macleay, W. 1877. 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GENUS AND SPECIES LIST Species formerly included in Lepidodactylus Fitzinger, 1843, Luperosaurus Gray, 1845 and Pseudogekko Taylor, 1922.

GENUS LEPIDODACTYLUS FITZINGER, 1843 Allengreercolotes paulwoolfi sp. nov. Lepidodactylus lugubris (Duméril and Bibron, 1836) Allengreercolotes shebae (Brown and Tanner, 1949) (type species) Allengreercolotes tepukapili (Zug, Waitling, Alefaio, Lepidodactylus aureolineatus Taylor, 1915 Alefaio and Ludescher, 2003) Lepidodactylus herrei Taylor, 1923 Allengreercolotes vanuatuensis (Ota, Fisher, Ineich, Lepidodactylus moestus (Peters, 1867) Case, Radtkey and Zug, 1998) Lepidodactylus planicaudus Stejneger, 1905 GENUS BORNEOCOLOTES GEN. NOV. Lepidodactylus woodfordi Boulenger, 1887 Borneocolotes ranauensis (Ota and Hikida, 1988) (type Lepidodactylus labialis (Peters, 1867) species) SUBGENUS BOREALISCOLOTES SUBGEN. NOV. GENUS ROSSSADLIERCOLOTES GEN. NOV. Lepidodactylus (Borealiscolotes) balioburius Ota and Rosssadliercolotes paurolepis (Ota, Fisher, Ineich and Crombie, 1989 (type species) Case, 1995) (type species) Lepidodactylus (Borealiscolotes) christiani Taylor, Rosssadliercolotes pantai (Stubbs, Karin, Arifin, 1917 Iskandar, Arida, Reilly, Bloch, Kusnadi and McGuire, Lepidodactylus (Borealiscolotes) yami Ota, 1987 2017) GENUS SHIREENHOSERGECKO GEN. NOV. GENUS LUPEROSAURUS GRAY, 1845 Shireenhosergecko shireenhoserae sp. nov. (type Luperosaurus cumingii Gray, 1845 (type species) species) Luperosaurus angliit Brown, Diesmos and Oliveros, Shireenhosergecko browni (Pernatta and Black, 1983) 2011 Shireenhosergecko dalegibbonsi sp. nov. Luperosaurus corfieldi Gaulke, Rösler and Brown, Shireenhosergecko jarradbinghami sp. nov. 2007 Shireenhosergecko mutahi (Brown and Parker, 1977) Luperosaurus kubli Brown, Diesmos and Duya, 2007 Shireenhosergecko oortii (Kopstein, 1926) Luperosaurus macgregori Stejneger, 1907 Shireenhosergecko orientalis (Brown and Parker, Luperosaurus palawanensis Brown and Alcala, 1978 1977) Luperosaurus sorok Das, Lakim and Kandaung, 2008 Shireenhosergecko petewhybrowi sp. nov. GENUS CHARLESPIERSONCOLOTES GEN. NOV. Shireenhosergecko robjealousi sp. nov. Charlespiersoncolotes joloensis (Taylor, 1918) (type GENUS JACKYHOSERGECKO GEN. NOV. species) Jackyhosergecko jackyhoserae sp. nov. (type species) Charlespiersoncolotes yasumai (Ota, Sengoku and Jackyhosergecko euaensis (Gibbons and Brown, Hikida, 1996) 1988) GENUS SCELOTRETUS FITZINGER, 1843 Jackyhosergecko manni (Schmidt, 1923) Scelotretus (Scelotretus) vittatus (Houttuyn, 1782) SUBGENUS SOLOMONCOLOTES SUBGEN. NOV. (type species) Jackyhosergecko (Solomoncolotes) flaviocularis Scelotretus (Scelotretus) daranini sp. nov. (Brown, McCoy and Rodda, 1992) (type species) Scelotretus (Scelotretus) haroldcoggeri sp. nov. GENUS BOBBOTTOMCOLOTES GEN. NOV. Scelotretus (Scelotretus) jenandersonae sp. nov. Bobbottomcolotes bobbottomi sp. nov. (type species) Scelotretus (Scelotretus) remotus (Rösler, Ineich, Bobbottomcolotes crusmaculosus sp. nov. Wilms and Bo¨hme, 2012). Bobbottomcolotes magnus (Brown and Parker, 1977) SUBGENUS HAROLDCOGGERCOLOTES SUBGEN. Bobbottomcolotes pumilus (Boulenger, 1885) NOV. Bobbottomcolotes potens sp. nov. Scelotretus (Haroldcoggercolotes) iskandari (Brown, GENUS MARTINEKCOLOTES GEN. NOV. Supriatna and Ota, 2000) (type species) Martinekcolotes listeri (Boulenger, 1889) Scelotretus (Haroldcoggercolotes) gulat (Brown, GENUS ADELYNHOSERGECKO GEN. NOV. Diesmos, Duya, Garcia and Rico, 2010) Adelynhosergecko novaeguineae (Brown and Parker, GENUS GEORGEMARIOLISCOLOTES GEN. NOV. 1977) (type species) Georgemarioliscolotes brooksii (Boulenger, 1920) Adelynhosergecko adelynhoserae sp. nov. (type species) Adelynhosergecko brettbarnetti sp. nov. Georgemarioliscolotes browni (Russell, 1979) Adelynhosergecko buleli (Ineich, 2008) GENUS PSEUDOGEKKO TAYLOR, 1922 Adelynhosergecko haydnmcphiei sp. nov. Pseudogekko compresicorpus (Taylor, 1915) (type Adelynhosergecko huonensis sp. nov. sp. nov. species) Adelynhosergecko judyfergusonae sp. nov. Pseudogekko chavacano Siler, Welton, David, Watters, Adelynhosergecko lachlanmcpheei sp. nov. Davey, Diesmos, Diesmos and Brown, 2014 Adelynhosergecko lucybennettae sp. nov. Pseudogekko ditoy Siler, Welton, David, Watters, Adelynhosergecko madangensis sp. nov. Davey, Diesmos, Diesmos and Brown, 2014 Adelynhosergecko matteoae sp. nov. Pseudogekko isapa Siler, Davis, Diesmos, Guinto, Adelynhosergecko oligoporus (Buden, 2007) Whitsett and Brown, 2016 Adelynhosergecko pulcher (Boulenger, 1885) Pseudogekko pungkaypinit Siler, Welton, David, Adelynhosergecko sloppi sp. nov. Watters, Davey, Diesmos, Diesmos and Brown, 2014 Adelynhosergecko stevebennetti sp. nov. Pseudogekko smaragdinus (Taylor, 1922) GENUS ALLENGREERCOLOTES GEN. NOV. Pseudogekko sumiklab Siler, Davis, Watters, Freitas, Allengreercolotes guppyi (Boulenger, 1884) (type sp.) Griffith, Binday, Lobos, Amarga and Brown, 2017 Allengreercolotes allengreeri sp. nov. SUBGENUS ROBWATSONCOLOTES SUBGEN. NOV. Allengreercolotes gardineri (Boulenger, 1897) Pseudogekko (Robwatsoncolotes) brevipes (Boettger, Allengreercolotes intermedius (Darevsky, 1964) 1867) (type species) Allengreercolotes lombocensis (Mertens, 1929) Pseudogekko (Robwatsoncolotes) atiorum Davis, Allengreercolotes pauldarwini sp. nov. Watters, Köhler, Whitsett, Huron, Brown, Diesmos and Hoser 2018 - Australasian Journal of Herpetology 38:32-64. Available online at www.herp.netSiler, 2015. AustralasianCopyright- KotabiJournal Publishing of Herpetology - All rights reserved Issue 38 was published in hard copy on 10 August 2018.