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Über Die Puppen Der Mittel- Und Westeuropäischen Gracillariinae (Lepidoptera: Gracillariidae)

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Über Die Puppen Der Mittel- Und Westeuropäischen Gracillariinae (Lepidoptera: Gracillariidae) Über die Puppen der mittel- und westeuropäischen Gracillariinae (Lepidoptera: Gracillariidae) J. Patoöka PATOCKA, J., 2001. ON THE PUPAE OF THE CENTRAL AND WESTERN EUROPEAN GRACILLARIINAE (LEPIDOPTERA: GRACILLARIIDAE). - ENT. BER., AMST. 61 (11): 157-164. Abstract: Pupae of seven species of Central and Western European Gracillariinae (Lepidoptera: Gracillariidae) are de¬ scribed and illustrated. Diagnostic characteristics of the species are compared with those of similar taxa of this subfami¬ ly. Some biological data are added. Institut für Waldökologie der SAW, Stürova 2, 960 53 Zvolen, Republik Slowakei. Die vorliegende Arbeit stellt einen Nachtrag Raupen an Asteraceae spp. In Mitteleuropa zu den Arbeiten von Patocka (1992a, 1992b) mit einer, in Westeuropa mit keiner Art ver¬ und Patocka & Zach (1995) vor. Sie behandelt treten. die Puppen der sieben von dem Autor bisher Bemerkung: Bei der Bestimmung dieser nicht untersuchten Arten der mittel- und Gattung mit Hilfe der Gattungstabelle der westeuropäischen Gracillariinae. Eine Cha¬ Gracillariidae in Patocka (1992 a) kommt man rakteristik der Lepidopteren-Puppen und Be¬ zum Punkt 11, Gattungen: Acrocercops Wal- stimmungstabelle deren mitteleuropäischer lengren und Parectopa Clemens. Von den Überfamilien und Familien anhand der Pup¬ beiden unterscheidet sich Aristaea durch die penmerkmale bringt Patocka (1999). Das frontale Richtung des Kopffortsatzes und System und die Nomenklatur folgt Karsholt & dadurch, daß der Kopf kaudal davon nur Razowski (1996) und berücksichtigt auch schwach gewölbt ist (Abb. 2-4). Von Parec¬ Leraut (1997) und Lastuvka (1998). Das topa ferner durch die Abwesenheit je einer untersuchte Material stammt aus der Samm¬ starken Kopfborste (Abb. 3), von Acrocercops lung des Verfassers, aus dem Museum für Na¬ und Dialectica Clemens durch das in der turkunde Berlin und der Zoologischen Staats¬ Mitte nur mittelstark verjüngte Pronotum sammlung München. (Abb. 4). Gattung Aristaea Meyrick Aristaea pavoniella (Zeller) (Abb. 1-9) Puppen mittelklein und mittelschlank (Abb. 1). Kopf läuft in einem spitzen, praktisch Untersuchtes Material: 5 6, 5 9, Slowakei frontal gerichteten Fortsatz aus, dorsal davon und Mähren. ist er nur schwach gewölbt. Eine große Cly- Puppe 3,8-4,5 x 0,7-0,9 mm, orange braun, pealborste nicht vorhanden (Abb. 2-4). Exuvie heller. Kopf und Thorax fein, stellen¬ Fabium überall deutlich begrenzt. Proboscis weise etwas runzelig skulptiert. 2.-8. Abdomi¬ länger als die Vorderbeine (Abb. 1). Pronotum nalsegment dorsal mit Mikrodömchen besetzt, in der Mitte mittelstark (etwa 3 x) verkürzt nur je ein Kaudalstreifen fast glatt. Borsten am (verschmälert, Abb. 4). Dörnchen an mittleren Metanotum, dorsal und subdorsal am 1-8. Ab¬ Abdominalsegmenten winzig (Abb. 6). Kre¬ dominalsegment ziemlich groß, sonst kaum master nicht vorhanden. 10. Abdominalseg¬ unterscheidbar. Labrum abgerundet. Mandi¬ ment kaudal verjüngt und am Ende konkav bulae länglich, schräg gestellt. Oculi deutlich, mit drei Paaren von Spitzen (Abb. 7-9). relativ groß. Palpi maxillares nicht oder kaum Ent. Ber., Amst. 61 (2001) 158 Abb. 1-11. 1-9, Aristaea pavoniella; 10-11. Caloptilia falconipennella. 1, Puppe ventral; 2, Frontalteil der Puppe lateral; 3, 10, ventral; 4, 11, dorsal; 5, Metanotum, Abdomenbasis (linke Hälfte); 6, 5. Abdominalsegment (linke Hälfte); 7, Abdomenende dorsal; 8, ventral; 9, lateral. Abkürzungen: A = Antenna; An = Analnaht; As = abdominales Spiraculum; Cl = Clypeus; F = Frons; S Gf = <? Genitalfeld; S Gn = S Genitalnaht; Hb = Hinterbein; Hfl = Hinterflügel; Kf = Kopffortsatz; Fb = Labium; Lbr = Fabrum; Mb = Mittelbein; Md = Mandibula; Msn = Mesonotum; Mtn = Metanotum; O = Oculus; Pn = Pronotum; V = Vertex; Vb = Vorderbein; Vfl = Vorderflügel; VS = Vorderschenkel; 1-10 = 1.-10. Abdominalsegment. unterscheidbar. Vorderschenkel etwas länger lappen. Das männliche Genitalfeld groß, als das Labium, Vorderbeine kürzer, Mittel¬ Analfeld klein, am Abdomenende liegend, beine viel länger als die Proboscis. Antennae dieses dort konkav. 10. Abdominalsegment und die langen Enden der Hinterbeine etwa kaudal verjüngt und mit drei Paaren spitzer zum Abdomenende hin ragend. Kopffortsatz Dörnchen versehen. ziemlich breit und spitz, der Kopf dorsal Raupe miniert in Aster, insbesondere A. davon nur mäßig gewölbt. Frons dorsal amellus Linnaeus. In Süden Mitteleuropas an stumpfwinklig auslaufend. Pronotum in der trockenwarmen Standorten, lokal. Mitte kaum zu 1/3 seiner größten Länge verkürzt, seine Seiten fein vertieft und Gattung Caloptilia Hübner gerunzelt, dorsal davon je ein Grübchen. Me¬ tanotum etwa zu 1/3 seiner Länge abgerundet Charakteristik der Puppen dieser Gattung in ausgeschnitten mit starker Dorsalborste und Patocka & Zach (1995). einigen schrägen Runzeln auf den Frontal¬ Ent. Ber., Amst. 61 (2001) 159 Caloptilia falconipennella (Hübner) bedeckt (Abb. 22, 23). Borsten sehr lang (Abb. 10-17) (Abb. 18, 23). Kopffortsatz sehr groß, lang und spitz (Abb. 19-21). Proboscis viel länger Untersuchtes Material: 1 S, Deutschland. als die Vorderbeine, Antennae überragen das Puppe 6,5 X 0,8 mm, rotbraun, Rücken Abdomen nicht (Abb. 18). Pronotum in der dunkler, Exuvie heller. Kopf und Thorax fast Mitte breit unterbrochen (Abb. 20). 10. Abdo¬ glatt und ziemlich glänzend. 1. Abdominal¬ minalsegment mit zwei langen, divergieren¬ segment mit dünnen Papillen, 2.-7. Segment den Ausläufern, außerdem mit Dörnchen dorsal mit deutlichen dunklen Dörnchen bewehrt, kein Kremaster (Abb. 24-26). besetzt, aber die Kaudalstreifen dieser Seg¬ Bemerkung: Diese Gattung ist von allen mente und das 8.-10. Segment sind nur etwas Gattungen der Gracillariinae durch den rauh skulptiert. Borsten am Thorax und 1.-9. großen Kopffortsatz (Abb. 18-21), die langen Abdominalsegment groß. Kopf ventral und Ausläufer am Abdomenende (Abb. 18, 24- lateral stumpf abgerundet ohne große Borsten. 26), und so weiter, leicht zu unterscheiden. Sie Kopffortsatz stumpf, höckerartig, ventral ist der Gattung Phyllocnistis Zeller (Gracilla- gerichtet. Frons dorsal am Ende stumpf¬ riidae: Phyllocnistinae) ähnlich, unterscheidet winklig. Pronotum in der Mitte kaum zu 1/3 sich zum Beispiel durch die längere Proboscis seiner größten Länge verkürzt, lateral und im und Antennae (Abb. 18), sowie durch die Mittelteil mit je einer Vertiefung. Metanotum perianalen und dorsalen Dörnchen am 10. Ab¬ kaum zu 1/3 seiner größten Länge und abge¬ dominalsegment (Abb. 24-26). rundet stumpfwinklig ausgeschnitten. Dorsal Raupen an Oleaceae, auch an Aquifolia- am 8. Abdominalsegment je ein Höcker mit je ceae spp. Nur eine Art in Süd- und Südwest¬ zwei Zähnchen an der Spitze. 10. Abdominal¬ europa. segment plötzlich quer abgeschnitten, dort mit einem Kranz von etwa fünf oder sechs Paar Metriochroa latifoliella (Minière) Dörnchen gesäumt. Am Kaudalende der Pup¬ (Abb. 18-26) pe liegt das mittelkleine Analfeld. Bemerkung: Bei der Bestimmung der Untersuchtes Material: 2 ?, Kroatien. ! Puppe dieser Art mit Hilfe der Bestimmungs¬ Puppe 3,8-4,1 X 0,75-0,85 mm, ockergelb, tabelle der Caloptilia-Äxten in Patocka & Exuvie heller. Skulptur am Kopf und Thorax Zach (1995) kommt man zum Punkt 11. Von fein, an den Vorderflügeln papillös, am 1., 9. den dort erwähnten Arten, Caloptilia alchi- und 10. Abdominalsegment aus winzigen, am miella (Scopoli) und C. robustella (Jäckh) 2.-8. Segment dorsal überall aus etwas größe¬ unterscheidet sich C. falconipennella durch ren Mikrodörnchem. Borsten am Clypeus, die gewölbt abgerundete Kopfform ( Abb. 10, Meso-, Metanotum und 1.-9. Abdominalseg¬ 12); Durch diese ist sie C. elongella (Lin¬ ment sehr lang. Kopffortsatz etwa halb so lang naeus) am Ähnlichsten, es fehlen bei ihr aber wie der Clypeus, anfangs parallelrandig, dann Idie großen Kopfborsten und sie unterscheidet schlank zugespitzt, frontal gerichtet, in sich durch die gezähnten Höcker dorsal am 8. Lateralansicht etwas ventral geneigt. Labrum Abdominalsegment (Abb. 16, 17). abgerundet. Labium überall deutlich begrenzt, Raupe an Ainus sp., miniert anfangs, später kürzer als die Vorderschenkel, Proboscis viel skelettiert sie unter einem umgeschlagenen länger als die Vorderbeine. Antennae länger Blattrand. Überwinterung als Falter; Gewäs¬ als die Hinterbeine, doch das Abdomen nicht serufer und Erlenbrüche, nicht häufig. überragend. Frons dorsal kurz und hinten eher konkav, Vertex groß und kaudal spitz. Pro¬ notum zweiteilig, in der Mitte breit unter¬ Gattung Metriochroa Busck brochen. Metanotum mäßig ausgeschnitten. Puppen mittelklein, mittelschlank (Abb. 18). Abdomen zum Ende stärker verjüngt. 10 2.-8. Abdominalsegment mit Mikrodörnchen Segment mit je einem Paar perianaler und 160 Ent. Ber., Amst. 61 (2001) Abb. 12-25. 12-17, Caloptilia falconipennella\ 18-25, Metriochroa latifoliella. 12, 19, Frontalteil der Puppe lateral; 13, 22, Metanotum, Abdomenbasis (linke Hälfte); 14, 23, 5. Abdominalsegment (linke Hälfte); 15, 24, Abdomenende ventral; 16, 25, lateral; 17, dorsal; 18, Puppe ventral; 20, Frontalteil der Puppe dorsal; 21, ventral. dorsaler Dörnchen und großer schlanker und 27). Vertex relativ kurz. Pronotum im Mittel¬ divergierender Ausläufer am Ende. teil extrem stark verkürzt (verschmälert, Abb. Raupe miniert an Oleaceae spp., auch an 30). 10. Abdominalsegment mit vier Paaren Ilex ciquifolium Linnaeus (von A. Lastüvka von Dörnchen. Kaudal ist er kegelförmig beobachtet). In Südwest- und Südeuropa ver¬ verjüngt, ein kurzer Kremaster ist vorhanden breitet. (Abb. 33, 34). Bemerkung: Bei der Bestimmung der Pup¬ pen dieser Gattung mit Hilfe der Bestim¬ Gattung Dialectica Walsingham
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