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A New Species of Acmaeodera (Coleoptera: Buprestidae) from Big Bend National Park, Texas, with Synonymy for Other Species Occurring in the United States

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A New Species of Acmaeodera (Coleoptera: Buprestidae) from Big Bend National Park, Texas, with Synonymy for Other Species Occurring in the United States University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 1998 A new species of Acmaeodera (Coleoptera: Buprestidae) from Big Bend National Park, Texas, with synonymy for other species occurring in the United States Richard L. Westcott Oregon Department of Agriculture, Salem, Oregon William F. Barr University of Idaho, Moscow, ID Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Westcott, Richard L. and Barr, William F., "A new species of Acmaeodera (Coleoptera: Buprestidae) from Big Bend National Park, Texas, with synonymy for other species occurring in the United States" (1998). Insecta Mundi. 354. https://digitalcommons.unl.edu/insectamundi/354 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 53 A new species of Acrnaeodera (Coleoptera: Buprestidae) from Big Bend National Park, Texas, with synonymy for other species occurring in the United States Richard L. Westcott Oregon Department of Agriculture Salem, OR 97310-0110, U.S.A. and William F. Barr Professor of Entomology Emeritus University of Idaho, Moscow, ID 83843, U.S.A. Abstract: Acrnaeodera tiquilia Westcott and Barr, new species, from Big Bend National Park, Texas is described, figured, and discussed in considerable detail, particularly in relation to the similar and partially sympatric A. recticollis Fall. A neotype is designated for A. quatuordecirnspilota Obenberger and that species is synonymized with A. orrzata (Fabricius). Additionally, A. gibbula gila Knull is synonymized with A. gibbula LeConte; A. rzautica Van Dyke is syn- onymized with A. sirnulata Van Dyke. Introduction ally, deeply depressed transversely above clypeus which is broadly, shallowly emarginate; surface Discovery of a new species of Acrnaeodera in densely, coarsely, rather shallowly punctate; vesti- Big Bend National Park, Texas, requires its de- ture long, mostly light brownish above, white be- scription for use in faunal lists and more general low; antennae serrate from segment 5, reaching works on the Buprestidae of that region. This de- slightly beyond middle of pronotum. Pronotum scription, plus synonymy in the genus, is also pro- 1.9X wider than long, widest at middle, strongly vided for inclusion in catalogues on Buprestidae convex, with a shallow basal median depression that are in preparation by colleagues. Collection which is distinctly deeper in front of sutural base codens (in parentheses) follow Arnett, et al. (1993) and with a vague, narrow median depression in or are as given in the acknowledgments. In verba- front of middle; front margin broadly, but slightly tim label data, "01)"denotes hand written, "(p)"or lobed medially; front angles not produced, nar- no indication means printed, "I" indicates a sepa- rowly rounded; hind margin subtruncate, deflexed rate label. from in front of umbones; lateral margins entire, abruptly constricted at hind angles, slightly re- Acmaeodera tiquilia Westcott and Barr, new flexed on anterior 112, visible from above only at species front angles; sides strongly arcuate, swollen on (Figures 1, 4, 6 and 7) posterior 112 above lateral margin; surface with rather dense, medium and shallow punctations Holotype, female: 9.6 mm long, 3.5 mm wide; medially, becoming coarsely, deeply reticulate- elongate-robust, elytra strongly flattened above, punctate laterally, anterior 112 almost entirely sides almost vertical; head, pronotum, underside with light brown setae, posterior 112 with long, and appendages shining black, vaguely brassy- dense white setae at sides and along front margin. green in certain lights; elytra black, each with 4 Elytra 2.1X longer than wide, widest along basal light yellow discal markings with thin brown bor- 114, equal to greatest pronotal width, extreme base ders (Figure l), first and second reaching lateral slightly constricted and equal to width of pronotal margin, third broadly interrupted and fourth base; sides strongly sloping almost vertical api- eclipsed laterally by a light red band that extends cally from second discal marking, subparallel on from anterior border of third marhng to apex, pale basal 112 then gradually convergent to narrowly yellow humeral spot present. rounded apex; humeral angle rather strongly, Head convex above, slightly flattened medi- acutely produced lateroventrally; umbone very 54 Vol. 12, Nos. 1 & 2, March-June, 1998, INSECTA MUNDI prominent; lateral margin broadly, shallowly "Bio.Note No.31 80, A. Hook" (1, no host). Para- emarginate near basal 114, finely and somewhat types deposited in BBNP, CLBC, DSVC, FSCA, sparsely serrate along apical 114; sutural area flat- GCWC, GHNC, NMPC, RLWE, TCMC, TTUC, tened along basal 112, becoming strongly elevated USNM, WFBC, ZMAS. apically; surface rather discretely, yet densely and Variation. The elytral maculation is quite deeply punctate-striate throughout, first 3 striae uniform in appearance, though some difference in especially distinct and impressed apically, with the markings occurs as follows: The yellow ones punctations smaller and more elongate; laterally are variably shaped, more or less discrete, the and basally, other stria1 punctations becoming thrd and fourth sometimes longitudinally con- equal to or greater than size of lateral pronotal nected. Laterally there may be considerable coa- punctations; first 5 rows of punctations (excluding lescence of markings, especially apically. The api- short, basal sutural row) separated by more than colateral red marlng varies in hue and may ex- the width of a punctation, discrete basally; rows 5- tend to the hind margin of the second yellow 8 separated by width of a punctation or less; rows marking andlor be interrupted apically. In a sam- 8-10 generally more widely spaced, especially by ple of 40 specimens, the humeral elytral spot, the prominently convex ninth interval; remaining which is located below the umbone, is present on intervals flattened except for a portion of the both sides on 37 and coalesces with the first elytral eighth shortly before it ends apically; punctations marking on 22. It is present only on one side of one of intervals fine throughout; vestiture considerably specimen and is absent on two specimens. less dense than on pronotum, mostly consisting of Sixteen specimens have some arrangement of long, erect, light brownish setae on disc, setae on submarginal yellow to yellowish-brown lateral sides shorter, recumbent to subrecumbent, mostly pronotal spots, at most (seen on 2) consisting of a whitish with a few brown ones apically. Ventral smaller spot on each side near apical angle and a surface with long, moderately dense setae which larger (usually) postmedian-subbasal spot on each are distinctly more dense at sides of first 3 visible side. The most common condition (7) is the latter abdominal sterna; prosternum slightly convex on pair only. Even the largest spots are scarcely dis- middle, front margin slightly retracted from front cernible to the naked eye. In addition, 2 specimens angles of pronotum, with a weakly developed (1 also having all 4 submarginal spots) bear a mi- rounded prominence on either side of shallowly nute median spot at the apical 115. emarginate middle; abdomen with medium-fine The front margin of the prosternum varies punctations moderately placed on middle, which from being weakly to strongly produced on each become denser and slightly larger at sides, except side of middle, which is broadly truncate to weakly first visible sternum coarsely and densely punctate emarginate. The subapical structure of the last laterally; last visible sternum subtriangular, nar- visible sternum varies from a rather vaguely de- rowly rounded apically with a distinct thickened fined ridge, usually with a distinct triangularly to triangular submarginal ridge. Ungual tooth with broadly rounded apex, or more rarely a well de- apex angularly rounded. Ovipositor as in Figure 4. fined broadly to narrowly rounded plate. It is of Material examined. Holotype (CASC) la- interest to compare this with a general comment beled: "Lower Pine Cyn., Big Bend N.P., TEX VII- on the genus made by Fall (1899:3): "It may be 16-84lon flowers of Tiquilia caltescelts n7ir.F. Barr thick or thin, broadly or narrowly rounded, trun- CollectorlHolotype Aclnaeodera tiquilia Westcott & cate or angulate, with regular or irregular edge, Barr" (h; red card). Paratypes all from Texas but I have never in the hundreds of specimens ex- [Brewster Co.], Big Bend National Park, on flow- amined discovered any variation in type within ers of Tiquilia greggii, 16117-VII-84, with excep- specific limits." tions noted: same data as holotype (2, except 1 on Length: R=8.0-9.9 mm, mean 9.1 mm; females, T. canescens); 6 mi SE Panther Jct. (8); 2.4 mi W N=16, Rz8.3-9.9 mm, mean 9.4 mm; males, N=24, Panther Jct. (I), all W. F. Barr; Pine Canyon Road, R=8.0-9.9 mm, mean 8.9 mm. 29"15'13", 103"11'30", 1160-1190 m (1); Glenn Comparison. This species appears most Springs Road, 29"15'48", 103"08'50", 945 m (15); closely related to A. recticollis Fall (Figure 2), with 1.6 mi W Basin Jct. (5) (the last 3 localities corre- which it is partially sympatric, and A. recticolloi- spond to the first 3); 10.3 mi W Basin Jct., 960 m des Westcott, a species restricted to extreme (6), all R. L. Westcott; Glenn Springs Road, 15-VII- southern California and northern Baja California. 86 (10); Lower Pine Canyon, 14-VII-86 (lo), all G. It is closer to the former, differing as follows: A. H. Nelson; Rio Grande Village, 16-VII-80, recticollis averages larger, with vestiture longer INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 55 and denser, particularly underneath.
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