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: Vegetative Organs

Leaves: Stem: Support BI 103: Gas exchange Transport Light absorption Storage An examination of leaves Chapter 43 cont. : Anchorage Storage Form = Function Transport Absorption

Introduction Adapted for Photosynthesis • Other functions of leaves: • Leaves are usually thin – Wastes from metabolic processes accumulate in leaves and are disposed of – High surface area-to-volume when leaves are shed. ratio – Promotes diffusion of carbon – Play major role in movement of water dioxide in, oxygen out absorbed by roots • occurs when water evaporates • Leaves are arranged to from surface. capture sunlight • Guttation - pressure forces water out hydathodes at tips of leaf veins in some . – Are held perpendicular to rays of sun – Arranged so they don’t shade one another

Common Leaf Forms Internal Anatomy of Leaves

Specialized structures: DICOT MONOCOT • Veins axillary – surrounded by bundle sheath blade • Mesophyll node • Stomata– openings for gas exchange blade sheath

node

1 leaf blade Leaf Vein (one ) cuticle : Cuticle leaf vein Upper Epidermis

Palisade • Waxy cuticle secreted by epidermis cells Mesophyll

stem • Protective layer against disease

Spongy • Reduced water loss from cells Water, dissolved Mesophyll mineral ions from roots and stems move into leaf Lower vein (blue arrow) Epidermis 50m cuticle-coated cell of lower epidermis Photosynthetic products (pink one (opening arrow) enter across epidermia) vein, will be Carbon transported Oxygen and water vapor dioxide in throughout outside air plant body diffuse out of leaf at enters leaf at stomata. stomata. Fig. 29-14, p.501

Guard Cells Dermal • Epidermis - Single layer of cells covering the entire surface of the leaf – Devoid of – Coated with cuticle – Functions to protect tissues inside leaves – Waste materials may accumulate in epidermal cells. – Different types of glands may also be present in the epidermis.

Vascular plants: have stomata : pores bordered by 2 guard cells • Bryophytes like moss and liverworts lack stomata • Have pores that are always open

Cross section of liverwort thallus (Conocephalum)

Open when water is abundant. Close when water is scarce.

2 Epidermis: Trichomes

Glandular of Stinging Nettle Urtica dioica

Trichomes on an Arabidopsis leaf. Trichomes are extensions from the plant's epidermis and they occur in numerous shapes and sizes in various plants. Trichomes may provide defense against insects

Mesophyll= Epidermal tissue: Prickles Most photosynthesis takes Prickles: epidermal place in the mesophyll between the cells- larger tichomes two epidermal layers. Examples: Rosa spp. I. Palisade Mesophyll – Compactly stacked, barrel-shaped parenchyma cells, commonly in two rows

– Contains most of leaf’s chloroplasts II. Spongy Mesophyll

– Loosely arranged parenchyma cells with abundant air spaces

Photosynthesis: The Components What are the inputs? ? ?

? What are the outputs?

? ?

3 Light is a wave of energy: Photosynthesis: The Components colors are different wavelengths

6CO2 + 6H2O + Light energy  C6H12O6 + 6O2

What are the inputs? - light - water - carbon dioxide

What are the outputs? - glucose - oxygen

Photosynthesis: Input - Light Photosynthesis: Light Absorption

The color you see Intensity

Plants are green because… • Reflect green light. • Absorb red, blue and purple light. Energy (wavelength) absorbed

Photosynthesis: Light Absorption Leaf Veins: Vascular Bundles Plant pigments absorb light….

• Xylem and phloem; often

strengthened with fibers

• In dicots, veins are netlike

• In monocots, they are parallel Absorb red, blue and purple light Reflect green light

4 Specialized Leaves Specialized Leaves: Carnivorous Carnivorous leaves Sundews • Grow in swampy areas • Have round to oval leaves covered with glandular and bogs hairs that have a sticky • Nitrogen and other fluid of digestive enzymes elements are deficient in at tip soil. Sundew • Specialized leaves trap Venus’s Flytraps and digest insects. • Only in North Carolina and South Carolina – Pitcher Plants • Blade halves trap insects. • Insects trapped and digested inside cone- Pitcher plant shaped leaves. Venus’s Flytraps

Specialized Leaves Specialized Leaves: • Floral Leaves (bracts) Example: Flowering dogwood, Cornus nutallii – At bases of or stalks – Poinsettia - Flowers do not have , instead brightly colored bracts surround flowers. – Clary’s sage - Colorful bracts are at top of flowering stalks above flowers.

Poinsettia Clary’s sage

Specialized Leaves Specialized Leaves • Leaves of Arid Regions • Leaves of Arid Regions – Conifers Tsuga canadensis – Sunken stomata – Arid regions have limited availability of water,

– wide temperature ranges, and high light intensities. – Leaves reduce loss of water by: • Thick, leathery leaves • Fewer stomata or sunken stomata • Succulent, water-retaining leaves, or no leaves • Dense, hairy coverings

Picea

5 Specialized Leaves Specialized Leaves

• Leaves of Aquatic Areas Tendrils – Less xylem and phloem – Mesophyll not differentiated into palisade and spongy layers. – Large air spaces Water lily leaf Tendrils Elodea leaf Spines Thorn

Thorns - Modified stems arising in the axils of leaves of woody plants

Spine

Autumnal Changes in Leaf Color

• Chloroplasts of mature leaves contain several groups of pigments: – - Green – Carotenoids - Yellows – In fall, chlorophylls break down and other colors are revealed. • Water soluble anthocyanins (red or blue) and betacyanins (red) may also be present in the .

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