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WR Anderson. 2006. Eight Segregates from the Neotropical Excerptfrom: W. R.Anderson. 2006. EightSegregates from the NeotropicalGenus Mascagnia (Malpighiaceae) Novon16: 168-204. fChristianella on pages190-191J 190 Novon VI. Christianella W. R. Anderson, gen. nov. TYPE: rarely 1, narrow; ventral areole circular or broadlv ChristianeLLume.\oemerica.nrt (W. R. Anderson) ovate. W. R. Anderson. It gives me great pleasure to name this genus in honor of Christiane Eva Seidenschnur Antlersor-r(b. Lianae lignosaevel frutit:es;lamina eglanclulosavel in 1944). Christiane's excellent revisions in the Malprgh- r,limidio proximali aliquot glandulis parvis rnarginalibus instnrcta: stipulae parvae prope llasim petioli portatae; iaceae and her achievements as eclitor of Systemotir: bracteaelrracrteolaeque eglanclulosae vel glanclulisparvis Boturry Monograph,s have made her an imporlant {igure rnarginalibusinstnr<r tae; lrrac'teolae apicre perd unculi poltatae; in our generation of plant taxonomists. sepalaaliquot glandulis parv.isrnarginalillus instructa, his ClristiuneLLc is distinguished by the small glan<.ls sessilibus,sulrsessilibus, vel stipitatis;petala hrtea, abax- ialiter lomentosa;filarnenta valcle heteromor'plu;samara {bund on the margin of the sepals, and in most species membrarnacea,alis lateraliltusapice clistincrtis, basi conLinuis on the margin of the bracteoles and sometimes the v"l usqrreurl nrr,'"m inrisis. brtrcts. Such rnarginal glands are founcl in a I'ervother genera of Malpighiaceae, Lrut not in any o1'the other Woody vines, or slirubby in open habitats. Petiole grolqrs inclLrded in Mrtscugnia by NiedenzLr. This eglandular or l.rearing2-10 large to small glands in 2 group of species is also notable {br thc marginal lea{' ro\,\rs;larnina eglanclular or Lrearing several snrall glands, the yellow abaxially tomentose petals, ttre sessile glands on rnargin on proximal hall; stipr-rles heteromorphic lilaments, ancl the membrulnous sa- very small, triangular, borne on petiole near base. maras rvitl'rthe lateral wings cleft to the nut at the apex Inllorescence with {lowers decusstrte to irregularly and continuous or cleft at the base. The sti{I'hairs on insertcrl in short to elongated pseucloracemes,telnri- the samara rvings are usually inserted at characteristic nal or lateral, single or (usually) grouperlin panicles; dark spots. ChristianeLLohas not been inclucled in any bracts and bracteoles eglandular or bearing several published molecular study. sul.rsessileto long-stalketl, clavate or capitate lnal-- ChristiuneLLa,consists o1'five species of Central and ginal glancls; floriferous peduncle rvell developed: Soutl'rAmerica. lrracteolesborne at apex o1'peduncle.Sepals elongat- etl but separating to expose petals during enlargement Kr,r' ro l'HnSpECTES t*' Cnntsrntr:t.t"t ol'bucl, appressedto spreaclingin anthesis,thc lateral 4 bearing large pairerd abaxial glancls, all 5 bearing 1a. Larninasericeous [o soonglabrate. the hairssessile and straight. a rorv of small sessile or subsessile to long-stalkecl, 2a. Bractsand bracteoles2-3 mm long, eglandu- clavateor capitate marginal glands; corolla bilaterally lar or with severaltinv sessileor subsessile symmetrical, the posterior petal r,vith a longer anrl rnarginal glands; marginal glands oI sepals thicker clarv than the lateral 4; petals vellow, sessileor subsessileisouthrvestern Brazil. abaxially clcnselv tomentose, aclaxially glabror-rs; ltoludicolu zl,. eru"t, o',i r,,r"",""r",i-i -; l;"t;. androecium bilaterally symmetrical; stamens 10, all rnany long-stalked marginal glnncls; marginal Iertile, glabrous; lilaments connate at base, straight, glancls of sepals long-stalkecl. stronglv heteromorpl'ric, longest and mr"rchthickenerl 3a. Bracts caducous; petiole of larger leaves opposite the 2 posterior-lati:ral petals, r-rsLrallylong 11-I5 rnrn long; lamina of larger leaves bLrt slender opposite the anterior sepal, slrortest 5-7.2 cn rvitlel samaras orhic:trlar. 35- 60 mnl rvir,le,the lateral rving continuous op1rositethe posterior petal; anthers subeqrral or at lrase, t:left to nut at aliex; weslern largcr on longer lilaments; gynoecium bilaterally Arnazonia . I. C. glarululy'bru synurtetrical; carpels very shortl,v connatt' in ovary; 3b. Bracts persistent or decidrrotrsin fruit; anterior style straight anrl shorter tl'ran the others, petiole of larger leaves 2-B mm long; the posterior 2 straight <ir bowed {r'om the base; lamina of larger leaves 2.5-5 cm rvicle; samaras butterfly-shaped, 22-42 mm apex ol' styles dorsally truncate to short-hooked, wide, the lateral wing cleft to nut at base the stigrna internal. Fruit clry, breaking apart into and apex; southeastern Mexir:o to Pana- samaras separating {rom a short obscurc pyramiclal rna 2. C. ntesoutneriL:rLnu torus; samara suborbicular to butter-lly-shaped, the lb. Lamina clensely and persistentlv tomentose. velu- tinous, or rvooll,v, especially abaxially, the hairs lateral wing(s) dominant, membranous with reticr.rlate- stalked. atrastornosingvenaticln, cleli to nut at apex, t,r-rntintr- 4a. Pedunr:le anrj pedicel very deeply rvoolly. l- ous at base or cle{i to nrrt, bearing still', r-rsualli' 1.5 rnm or rlol'e tliarn. inclucling vesture; hairs basi{ixetl or sub-basi{ixed I'rairs insertctl at dark so clenseon abaxial leaf strrfac:eas to make it spots, the margin undulate or coarsely tlerrtate;rlorsal difficult to distinguish the rrrossJriet:eof any given hail bracteoles abaxiallv very densely rving srnall, extendecl {brwarcl at apex through gap spreacling-w<xllly,aclaxially tornentose,at least rrr latera,l rving, clistinct at base ol conlltrent with on the distal half; sepals abaxially tlccpli lateral rving; intermediate winglets mostly lacking, spreading-rvoolly; petals al.raxially densely Volume16, Number 2 Anderson 191 2006 Segregatesfrom Mascagnia rvoolly; southern Bolivra, northem Paraguay, mrLLtigLarulllLosovar. surinumensis Kostermans, trncl southern l3razil . 3. C. mtLlt,iglun.tluLoso Mecletl. Bot. Mus. HerLr. Rijksuniv. Utrecht 25: 4b. Per.luncle and pedicel clensely tolrentose or (Kostcrmans) veiutinous but only 0.4-0.i rnnr diarn. in- 5. 1936. Mascugniu surinumen.\i.i r.rluding vesture: leal hairs mur:h less rlense. W. R. Anclerson, Contr. Univ. Michigan Herb. tlre crosspieces of inclividual hairs eersr- 17: 53. 1990. TYPE: Suriname. Brownsberg,23 l,v distinguished; bracteoles abaxrall,v ap- Sep. 1931, W. C. u. Emden (holotype, U; ist-rtypes, presserl-tontentose. adaxially glabrous or to- nlentose only at verv apexl sepals alraxially K, MICH). Figure B. appressecl-tomentose or srrbseric,eous;petals This species is rvidespreacl in Amazonian South abaxially rnoderately appressed-tornentose: Arnazonian Sorrth America . 5. C. suri,nu,rnensis America, including Cuyana and Suriname (Anderson 1990b:53). l. Christianella glandulifera (Cutrtrecasas) W. R. Anclerson, comb. nov. Basionym: Mascognin gLuntluLifbnr Cuatrecasas, Webbia 13: 365. 1958. TYPE: Colombia. Amazonas-Vaup6s: Rfo Apaporis, R. E. SchuLt,es& L Cubreru l.l209 (holotype,US; isotypes,GH, NY). 2. Christianella rnesoarnericana (W. R. Anderson) W. R. Anderson, comb. nov. Basiorrym: Moscrtg- niu m,esoumcriconuW. R. Anderson. Contr. Univ. Michigan Herb. t6: 105. 1987. TYPE: Guate- mala. Retall-ruleu: Between Retalhuleu and Nueva Linda, P. C. StaruLLey87301 (holotype, F: isotype, US). This species is known Ii'orn stlrtheasteln Mexico, Cuatemala,antl Panama(Antlerson, 1987: 105),and has rec:entlybeen founcl in Costa Rica (,lcostaB0J, INB). 3. Christianella rnultiglandulosa (Nieclenzu in Chodat & Hassler) W. R. Anclerson,comb. nov. Basionym: M asr:agn,inmuLtigLrtntLulo.lo Niedenzu in Choclat& Flassler,Bull. Herb. Boissier,56r. 2, 7:284.1907. TYPE: Paraguay.Near Valenzuela, E. HossLer6998 (lectotype, clesignateclhere, C; isotypes,A, C, K, MO, NY). Tfre frcrlotl,pe sheet ol' HassLer 6998 vtas destroyed at B. There ar€r{our isotypes at C; the lectotype is the isotype that was annotated lectotype by W. R. Anclersonin 1993. This specics is knolvn fi'om Paraguay and southern Brazil (Anderson 1990b: 53) and southern Bolivia (Cutiirrez e.toL. I170, MICH). 4. Christianella paludicola (W. R. Anclerson) W. R. Anderson, comb. nov. Basionym: Mascugnirt paLutlicoLuW. R. Anderson, Contr. Univ. Michi- gan Herb. 16: 106. 1987. TYPE: Brazil. Amaztrnas: Rio Purfrs. Rio Ituxi. C. T. Prunce ct uL.l.l9lB (holotype,INPA; isotypes,K, MICH, I{O, NY, U, US,VEN). 5. Christianella surinarnensis (Kostermans) W. R. Antlerson, comb. nov. Basionym: Muscugrtio.
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