1 A molecular phylogeny of Cochylina, with confirmation of its relationship to Euliina 2 (Lepidoptera: Tortricidae) 3 4 John W. Brown*1, Leif Aarvik2, Maria Heikkilä3, Richard Brown4, and Marko Mutanen5 5 6 1 National Museum of Natural History, Smithsonian Institution, Washington, DC, USA, e-mail: 7
[email protected] 8 2 Natural History Museum, University of Oslo, Norway, e-mail:
[email protected] 9 3 Finnish Museum of Natural History, LUOMUS, University of Helsinki, Helsinki, 00014, 10 Finland, e-mail:
[email protected] 11 4 Mississippi Entomological Museum, Mississippi State, MS 39762, USA, e-mail: 12
[email protected] 13 5 Ecology and Genetics Research Unit, PO Box 3000, 90014, University of Oulu, Finland, e- 14 mail:
[email protected] 15 *corresponding author 16 17 This is the peer reviewed version of the following article: Brown, J.W., Aarvik, L., Heikkilä, M., 18 Brown, R. and Mutanen, M. (2020), A molecular phylogeny of Cochylina, with confirmation of its 19 relationship to Euliina (Lepidoptera: Tortricidae). Syst Entomol, 45: 160-174., which has been 20 published in final form at https://doi.org/10.1111/syen.12385. 21 1 22 Abstract. We conducted a multiple-gene phylogenetic analysis of 70 species representing 24 23 genera of Cochylina and eight species representing eight genera of Euliina, and a maximum 24 likelihood analysis based on 293 barcodes representing over 220 species of Cochylina. The 25 results confirm the hypothesis that Cochylina is a monophyletic group embedded within a 26 paraphyletic Euliina. We recognize and define six major monophyletic lineages within 27 Cochylina: a Phtheochroa Group, a Henricus Group, an Aethes Group, a Saphenista Group, a 28 Phalonidia Group, and a Cochylis Group.