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0 20 40 60 80 100 Time (Ma) Papilionidae Hesperiidae Pieridae Hamearis lucina Cigaritis acamas Lycaena dimorpha 44.88 12.92 Lycaena phlaeas 8.55 Lycaena bleusei Lycaena virgaureae 108.3 Lycaena ottomana 11.11 Lycaena tityrus 88.16 Lycaena helle 8.2 Lycaena alciphron Lycaena dispar Lycaena thetis Lycaena thersamon Lycaena hippothoe Lycaena candens Laeosopis roboris 107.07 Thecla betulae 54.91 22.53 35.56 Favonius quercus Tomares ballus 30.98 5.55 Tomares nogelii Tomares callimachus Callophrys avis 22.62 3.33Callophrys rubi Callophrys suaveola Callophrys chalybeitincta 16.66 Neolycaena rhymnus 6.58 Satyrium pruni 10.42 Satyrium w-album Satyrium acaciae 8.09 Satyrium spini 50.51 Satyrium ledereri Satyrium ilicis Satyrium esculi Azanus ubaldus 101.63 18.35 Azanus jesous Zizeeria knysna Zizeeria karsandra 21.14 Tarucus theophrastus Tarucus balkanicus 25.18 Cacyreus marshalli 15.07 Lampides boeticus 22.43 Celastrina argiolus Phengaris alcon 35.18 5.12 Phengaris arion 23.36 Phengaris teleius Phengaris nausithous 15.56 Scolitantides orion 10.07 Praephilotes anthracias 11.7 Iolana iolas Iolana debilitata 10.1 Glaucopsyche melanops Glaucopsyche alexis 13.49 33.59 Glaucopsyche paphos Turanana taygetica 12.26 Pseudophilotes bavius Pseudophilotes barbagiae 8.12 Pseudophilotes abencerragus Pseudophilotes panoptes Pseudophilotes baton Pseudophilotes vicrama Cyclyrius webbianus 8.12 Leptotes pirithous Tongeia fischeri Cupido argiades 12.82 27.13 Cupido decoloratus Cupido alcetas 6.58 Cupido osiris Cupido minimus 23.5 Cupido lorquinii Luthrodes galba 97.93 Freyeria trochylus Plebejus argus 17.21 6.3 Plebejus idas Plebejus bellieri Plebejus argyrognomon 14.55 Plebejidea loewii 6.72 Eumedonia eumedon 8.16 Kretania eurypilus Kretania hesperica Kretania trappi 10.79 Kretania sephirus 8.72 Kretania pylaon Cyaniris semiargus Agriades optilete Kretania psylorita Agriades orbitulus 6.57 Agriades dardanus Agriades pyrenaicus Agriades zullichi 9.84 Agriades aquilo Agriades glandon Glabroculus cyane Aricia morronensis 8.92 Aricia anteros Aricia cramera 5.82 Aricia nicias Aricia artaxerxes 9.43 Aricia montensis Aricia agestis Neolysandra coelestina Lysandra bellargus Lysandra corydonius 8.34 Lysandra coridon Lysandra caelestissima Lysandra hispana 7.66 Lysandra albicans Polyommatus escheri Polyommatus thersites 6.34 Polyommatus amandus Polyommatus daphnis 6.11 Polyommatus icarus Polyommatus eros Polyommatus celina 5.81 Polyommatus nivescens Polyommatus dorylas 5.57 Polyommatus golgus Polyommatus damon Polyommatus iphigenia Polyommatus damocles Polyommatus admetus Polyommatus nephohiptamenos Polyommatus ripartii 3.9 Polyommatus damone Polyommatus violetae Polyommatus fulgens Polyommatus fabressei Polyommatus dolus Polyommatus humedasae Polyommatus orphicus Polyommatus aroaniensis Polyommatus timfristos Nymphalidae 100 8 0 6 0 4 0 2 0 0 Time (Ma).
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