The Journal of Cell Biology Manuscript content verification and license to publish

Submitted Manuscript Title: Manuscript # Authors (all names):

In recognition of the author(s)’ desire to publish the manuscript identified above (the “Manuscript”) and any supplemental material in The Journal of Cell Biology (the “Journal”), and the desire of The Rockefeller University Press (“RUP”) to publish the Manuscript as quickly as possible should it be accepted, the Manuscript will be considered for publication on the following terms:

1. Corresponding Author Designation: The Corresponding Author has been designated 7. Copyright and License: by any and all co-Authors of the Manuscript (each an “Author” and collectively the a. Copyright Ownership: Ownership of copyright in the Manuscript remains with “Authors”) to sign this Agreement on their behalf. If the Manuscript was prepared jointly, the the Authors. Corresponding Author agrees to inform any and all co-Authors of the terms of this Agreement and to obtain all co-Authors’ permission to sign on their behalf. b. Retained Rights: After the Article is published in the Journal, the Authors will have the right to distribute the Article in the form and format as published by RUP as the Authors see 2. Representations and Warranties: Each of the Authors represents and warrants that fit, so long as the Authors provide a citation to the Article and a link to the DOI associated (a) the Manuscript was created by and is original to the Authors; (b) the Authors own or with the Article. The Authors’ rights include (i) the right to publish the Article on the official have the right to license the Manuscript to RUP pursuant to this agreement; (c) if any website(s) of the institutions or organizations with which the Authors are affiliated and on person other than the Authors contributed to any prior version of the Manuscript, the the Authors’ websites; and (ii) the right to deposit the Article with any funding repository if Authors have obtained from such person(s) all permissions and consents necessary for required pursuant to a grant or funding agreement. the submission of the Manuscript to RUP and the exercise by RUP of the rights granted in c. License to RUP: The Authors hereby grant to RUP, from the date of submission of the this agreement; (d) the Manuscript does not and will not infringe upon, violate, Manuscript to RUP through the date that is six (6) months after the date on which the Article misappropriate or give rise to any adverse claim with respect to any intellectual property is published in the Journal (the “Initial Publication Period”), the exclusive right to (i) edit, right or other personal, proprietary or contractual right of any third party, or defame any revise, supplement and format the Manuscript as deemed necessary by RUP in connection person, or violate any law or regulation; (e) the Authors have the full right, power and with the creation of the Article; (ii) publish, reproduce, distribute, display and store the Article authority to enter into this agreement and to grant to RUP all rights granted in this in all media and formats whether now or later known or developed; (iii) translate the Article agreement, free and clear of any and all adverse claims, encumbrances or other interests into other languages, create adaptations, summaries and extracts of the Article and other of any third party; (f) the Authors have not previously granted any rights in the Manuscript derivative works based on the Manuscript and exercise all of the rights set forth in (i) and to any third party except as permitted pursuant to the section entitled “Prior Publication” (ii) above in such translations, adaptations, summaries, extracts and derivative works; and below; and (g) when executed and delivered, this agreement will constitute a legal, valid (iv) license others to do any or all of the above. The Authors hereby grant to RUP, and binding obligation of each Author. Each Author will indemnify and hold RUP and The beginning immediately following the expiration of the Initial Publication Period and continuing Rockefeller University harmless with respect to any claim or liability resulting from any thereafter in perpetuity, all of the right set forth in this Section 7(c) on a non-exclusive basis. breach of any of Author’s representations or warranties in this agreement or any assertion d. Distribution by RUP: that is contrary to any such representations or warranties. i. Initial Publication Period: During the Initial Publication Period, RUP will grant to 3. Author Contributions and Identification: Each of the Authors represents and warrants the public the non-exclusive right to copy, distribute or display the Article, under the that the Author has contributed significantly to the content and preparation of the Manuscript following conditions: and that the Author has seen and approved the content of the Manuscript and the manner A. Attribution: The user must attribute the Article as being copyright of the in which the Authors are identified in connection with the Manuscript (including, without Authors and published in the Journal, but not in any way that suggests that the limitation, the order in which the Authors are listed and any identification of the respective Authors or RUP endorse the user or the user’s use of the Article. contributions of the Authors). B. Noncommercial: The user must not use the Article for commercial purposes. 4. Prior Publication: Each of the Authors represents and warrants that neither the C. Share Alike: If the user shares, alters, transforms, or builds upon the Article, Manuscript nor any paper or article containing content similar to the Manuscript has the user may distribute the resulting work only under the same license the user previously been published in any article listed in a public citation database or in a book received from RUP. identified with an ISBN. The Authors represent and warrant that the Manuscript is not D. No Mirror Sites. The user may not create, compile, publish, host, enable, currently under consideration for publication by any person or entity other than RUP and or otherwise make available a mirror site of the RUP web site, any of its agree not to submit the Manuscript for publication elsewhere. If the Manuscript is accepted constituent journal web sites (The Journal of Cell Biology, The Journal of by RUP, the Authors will not submit or permit the Manuscript to be submitted to any other Experimental Medicine, and The Journal of General Physiology), or any subset publisher. If the Manuscript or a prior version of the Manuscript has been published or of the RUP or journal web sites. A “mirror site” is a web site that contains the made available via a “preprint server,” the Authors shall notify RUP in writing of the same same content as a parent web site but is located at a different geographic at the time of submission of the Manuscript to RUP and each of the Authors represents location from the parent site. This prohibition applies regardless of the and warrants that neither the Authors nor any third party has granted any license or commercial or non-commercial nature of the mirror site. authorization in connection with the same that conflicts with any of the terms of this The full legal text of the license is available at http://rupress.org/rup-license_v1 agreement or the rights granted to RUP in this agreement. Upon publication by RUP of the ii. After Initial Publication Period: The Authors acknowledge that RUP will make the edited and formatted version of the Manuscript (such edited and formatted version, including Article freely available to the public on RUP’s website after expiration of the Initial all forms of supplemental material and any images used for cover art or for any other Publication Period, and that RUP intends to submit the Article to PubMed Central in journal feature, the “Article”), the Authors shall request that the preprint server acknowledge accordance with PubMed Central’s requirements, where the Article will be released to publication of the Article and provide a citation to the Article and a link to the DOI the public after expiration of the Initial Publication Period. After the Initial Publication associated with the Article. Period, RUP will grant to the public the non-exclusive right to copy, distribute, or display 5. No Competing Financial Interests: Each of the Authors represents and warrants that the Article under a Creative Commons Attribution-Noncommercial-Share Alike 4.0 all commercial affiliations or arrangements, stock or equity interests or patent-licensing International license, as described at https://creativecommons.org/licenses/by-nc- arrangements that could be considered to pose a conflict of interest regarding the submitted sa/4.0/legalcode, or updates thereof. article have been disclosed in writing to RUP. e. Special Notice for Governmental Works and Works for Hire: RUP acknowledges 6. Permissions: The Authors are responsible for obtaining permission for the use of that if the Manuscript was prepared under a United States Government contract or by any copyrighted material that is owned by any third party. If the Manuscript contains employees of a foreign or domestic government and is subject to government publication any copyrighted material that is owned by a third party, the Authors shall at the time of rights, then the above license is made subject to those rights. If an Author is unable to submission of the Manuscript notify RUP in writing of the same and submit to RUP copies retain copyright in a Manuscript because the Author is an employee under a “work for of all such permissions. If the Manuscript includes any material provided by others that is in hire” arrangement, the Author is responsible for ensuring that the Author’s employer the public domain or otherwise not subject to copyright protection, the Authors shall at the grants to RUP the above license rights and has authorized the Author to sign this time of submission of the Manuscript notify RUP in writing of the same and submit to RUP license. copies of all permission letters or other agreements relating to such material. In cases where a manuscript is NOT accepted, the portions of this agreement regarding licensing shall be null and void.  At least one author of this Manuscript was employed by the U.S. National Institutes of Health at the time the research was performed. Checking this box serves as submission and acknowledgement of the NIH Cover Sheet. Select one of the following:  None of these apply.  This manuscript was created exclusively by U.S. Government employees within the scope of their employment. This Manuscript is not subject to U.S. copyright protection.  This Manuscript was created by one or more employees of the Australian, Canadian, and/or UK Governments (the “Crown”) within the scope of their employment. The Crown owns the copyright interests of authors who are government employees.  This is a work made for hire, and my employer owns the copyright. If signing as an authorized representative and on behalf of an employer copyright holder, please provide the full name of the organization as it should appear in the copyright notice: ______

Corresponding Author: Signature: Date: ______ I affirm that I have read and agree to the above.