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University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln

Center for Systematic Entomology, Gainesville, Insecta Mundi Florida

March 2001

Preliminary checklist of the (, Mantodea, Phasmatodea,) of Texas

John A. Stidham Garland, TX

Thomas A. Stidham University of California, Berkeley, CA

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Stidham, John A. and Stidham, Thomas A., "Preliminary checklist of the orthopteroid insects (Blattodea, Mantodea, Phasmatodea,Orthoptera) of Texas" (2001). Insecta Mundi. 180. https://digitalcommons.unl.edu/insectamundi/180

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 15, No. 1, March, 2001 35

Preliminary checklist of the orthopteroid insects (Blattodea, Mantodea, Phasmatodea,Orthoptera) of Texas

John A. Stidham 301 Pebble Creek Dr., Garland, TX 75040

and

Thomas A. Stidham Department of Integrative Biology, Museum of Paleontology, and Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720,

Abstract: Texas has one of the most diverse orthopteroid assemblages of any state in the United States, reflecting the varied habitats found in the state. Three hundred and eighty-nine and 78 subspecies of orthopteroid insects (Blattodea, Mantodea, Phasmatodea, and Orthoptera) have published records for the state of Texas. This is the first such comprehensive checklist for Texas and should aid future work on these groups in this area.

Introduction (Flook and Rowell, 1997). The species are listed alphabetically within families or subfamilies, and Overall, the orthopteroid fauna of North Amer- therefore the alphabetical taxonomic listing does ica is fairly well known. However, details of the not imply any phylogenetic relationships within geographic distribution of many orthopteroid spe- the subfamily or family. The reference for the cies in the U.S. are not well documented. We have published record is given after the species name compiled the published records of Texas ortho- and authority. The alpha is still unclear pteroids, as a basis for future study. Most of these for a few species included in this list, and therefore records have been confirmed through collections additional species may be added in the future as the made by the authors throughout Texas overthe last number of taxa is clarified. This checlilist is meant 20 years. The study of Texas began in to act as an aid to collectors, collection managers, the mid-nineteenth century and continues today field workers, and other researchers working on with 389 species currently recorded in the state. Texas orthopteroids. This places the Texan orthopteroid fauna as one of the most diverse of any state. There are 197 species The Orthopteroidea of Texas of known from Texas, compared to the over 200 species from California (Strohecker et Order Blattodea al., 1968), though other orthopteroid groups are more diverse in Texas. The large number of species Family Blattelidae is most lilrely related to the great variety of habitats found in Texas, from desert in the West, to high Blattella ger~nartica(Linnaeus) (Hebard, 194313) plains in the North, and subtropical climates in the Blattella vaga Hebard (Atkinson et al., 1991) South. Combined with failure of Pleistocene gla- Cariblatta lutea lutea (Saussure and Zehntner) ciers to reach Texas, the state harbors factors (Atkinson et al., 1991) necessary for long term orthopteroid diversifica- Cariblatta lutea ~ni~zi~naHebard (Atkinson et al., tion, as seen in its speciose fauna. However, the 199 1) number of endemic species is low compared to Chorisoizeura texerzsis Saussure and Zehntner (He- California, since many of the habitats (and hence bard, 194313) species ranges) in Texas extend into the adjacent Euthlastoblatta abortiva Caudell (Hebard, 1943b) states and into Mexico. Euthlastoblatta gernlna (Hebard) (Atkinson et al., Supraspecific taxonomy follows Otte (1981, 1991) 1984) and Helfer (1987). Tridactylids are placed in Ischnoptera deropeltiformis (Brunner) (Hebard, following tradition, though recent molec- 1943b) ular data suggest their placement within 36 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Isclznoptera rufa occiderztalis Saussure (Hebard, Order Mantodea 1943b) Parcoblatta bolliana (Saussure and Zehntner) (He- Family Mantidae bard, 1943b) Parcoblatta caudelli Hebard (Hebard, 194313) Brz~~z~zeaborealis Scudder (Hebard, 194313) Parcoblatta desertae (Rehn and Hebard) (Hebard, Litalzeutria minor (Scudder) (Hebard, 1943b) 194313) Oligo~zicellamexica7zus (Saussure) (Hebard, 194310) Parcoblatta divisa (Saussure and Zehntner) (He- Oligonicella scudderi (Saussure) (Hebard, 194313) bard, 194313) Plzyllovates chlorophaea (Blanchard) (Hebard, Parcoblatta fulvesce7zs (Saussure and Zehntner) 194313) (Hebard, 194313) Stag~norna,~ztiscaliforizica Rehn and Hebard (He- Parcoblatta lata (Brunner) (Hebard, 194313) bard, 194313) Parcoblatta pennsylva7~ica (DeGeer) (Hebard, Sta.gmonzarztis carolina (Johannson) (Hebard, 194333) 1943b) Parcoblatta uirgiizica (Brunner) (Hebard, 1943b) Stag~no~narztislinzbata (Hahn) (Hebard, 1943b) Parcoblatta zebra Hebard (Hebard, 194313) Yersirziops solitarium (Scudder) (Hebard, 194313) Plectoptera picta Saussure and Zehntner (Atkin- son et al., 1991) Order Phasmatodea Pseudolnops septentriorzalis Hebard (Hebard, 194313) Family Supella lo7zgipa,lpa (Fabricius) (Hebard, 194313) Ai~isoinorphaferrugi~zea (Beauvois) (Hebard, 1943b) Family Blattidae Family Blatta lateralis (Walker) (Atkinson et al., 1991) Blatta orientalis Linnaeus (Hebard, 194313) Diaplzero~nera covillea (Rehn and Hebard) (He- iveostylopyga rholnbifolea (Stoll) (Atkinson et al., bard, 194313) 1991) Diapherolnera fe~norata(Say) (Hebard, 194313) Periplaizeta anzericana (Linnaeus) (Hebard, 1943b) Diapheronzera persi~nilisCaudell (Hebard, 194313) Periplaneta au~trala~siae(Fabricius) (Hebard, Diaphero~nera tarnauliperznis (Rehn) (Hebard, 1943b) 194313) Periplaneta bruiz~zeaBurmeister (Hebard, 194313) Diaplzeroi?zera torquata Hebard (Hebard, 194313) Peripla,rzeta fuliginosa (Serville) (Hebard, 1943b) Diaphero~neravelii euc7~e1rzisHebard (Hebard, 1943b) Family Blaberidae Diapherolnera uelii velii Walsh (Hebard, 194313) Megaphas~nadenticrus (StAl) (Hebard, 194313) Pa.7~chloi-acubensis Saussure (Hebard, 194313) coloradus (Scudder) (Hebard, 194313) Pyc7~oscelus suri~~amensis(Linnaeus) (Hebard, Pseudoser~rzylestra~ninea (Scudder) (Hebard, 194313) 194313) Pseudoser7nyle strigata (Scudder) (Hebard, 194313) Serrnyle mexicarza (Saussure) (Hebard, 194313) Family Polyphagidae Order Orthoptera Arelzivaga bollialza (Saussure) (Hebard, 194310) Arenivaga erratica Rehn (Hebard, 1943b) Suborder Caelifera Arelzivag-a gra,ta Hebard (Hebard, 1943b) Areaivag-a tonkawa Hebard (Hebard; 1943b) Family Attaphila fu7zgicola Wheeler (Hebard, 194310) Cornpsodes cz~cullatusSaussure and Zehntner (At- Areotettix fe~noratus(&udder) (Rehn and Grant, kinson et al., 1991) 1961) Co7npsodes schwai-zi (Caudell) (Atkinson et al., Areotettix ~zullisiizus(Hancocli) (Rehn and Grant, 1991) 1961) Erelnoblatta subdiaphana (Scudder) (Hebard, iVoinotettix cristatus coinpressus Morse (Rehn and 194313) Grant, 1961) INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Pamtettix astecus (Saussure) (Rehn and Grant, Mer~neriabivitatta bivitatta (Serville) (Knutson, 1961) 1940) Paratettix breviperzrzis (Hancock) (Rehn and Grant, Mermeria bivitatta inaculipe7~r~isBruner (Knut- 1961) son, 1940) Paratettixc~~cullatus(Burmeister) (Rehn and Grant, Merrneriapicta 7~eo1nexica7za(Thomas) (Otte, 1981) 1961) Mermeria picta picta (Walker) (Tinkham, 1948) Paratettix mexica7zus (Saussure) (Rehn and Grant, Mer~neriatexarza Bruner (Tinkham, 1948) 1961) obscura (Thomas) (Otte, 1981) Paratettix rugosus (Scudder) (Rehn and Grant, 07-phulella. olivacea (Morse) (Gurney, 1940) 1961) Orphulella olivacea halophila Rehn and Hebard Paratettix toltecus (Saussure) (Rehn and Grant, (Gurney, 1940) 1961) Orphulella pelidlza (Burmeister) (Otte, 1981) Tetrix are~~osaar~gusta Hancock (Rehn and Grant, 01-phulella speciosa (Scudder) (Otte, 1981) 1961) Parapolnala pallida Bruner (Otte, 1981) Tetrix are7zosa are~zosaBurmeister (Rehn and Parapomala virgata Scudder (Otte, 1981) Grant, 1961) Parapo7nala wyorni~zgerzsis(Thomas) (Otte, 1981) arinata Morse (Rehn and Grant, 1961) Phlibostro~naquadrimaculatum (Thomas) (Otte, cazieri Rehn and Grant (Rehn 1981) and Grant, 1961) Pseudopolnala brachyptera (Scudder) (Otte, 1981) Tettigidea lateralis lateralis (Say) (Rehn and Grant, delicatula delicatula (Scudder) (Rehn, 1961) 1942) Psoloessa texaaa pawnee Rehn (Rehn, 1942) Family Psoloessa texarza pusilla (Scudder) (Rehn, 1942) Psoloessa texatza texa~rzaScudder (Rehn, 1942) Subfamily Gornphocerinae Rhammatocerus uiatorius (Saussure) (Tinkham, 1948) Achururn hilliardi Gurney (Helfer, 1987) Syrbula adrnirabilis (Uhler) (Otte, 1981) Achurunz irzirzi1nipe7~7zeCaudell (Otte, 1981) Syrbula montesuma (Saussure) (Otte, 1981) Achurunz sulnichrasti (Saussure) (Otte, 1981) hirtipes hirtipes (Say) (Otte, 1981) Subfamily Acridinae variegatus Bruner (Helfer, 1987) Metaleptea brevicorlzis (Johannson) (Otte, 1981) Acroloplzitus maculiperzi~is(Scudder) (Otte, 1981) Agerzeotettix brevipen7zis (Bruner) (Otte, 1981) Subfamily Oedipodinae Agerzeotettix deorurn (Scudder) (Otte, 1981) Arnphitor~ziscoloradus (Thomas) (Otte, 1981) Arzcorzia hebardi Rehn (Otte, 1984) elliotti (Thomas) (Otte, 1981) Arphia corzspersa Scudder (Otte, 1984) Aulocara felnoraturlz (Scudder) (Otte, 1981) Arphiapecos Otte (Otte, 1984) Boopedorz aurivelztris McNeill (Otte, 1981) Arphia pseudoizietarza (Thomas) (Otte, 1984) Boopedorz gracile Rehn (Otte, 1981) AT-phiasi~nplex Scudder (Otte, 1984) Boopedon 7zubilurn (Say) (Otte, 1981) AT-phiasulphurea (Fabricius) (Otte, 1984) argentatus Bruner (Otte, 1981) Arphia xarzthoptera (Burmeister) (Otte, 1984) Cibolacris parviceps (Walker) (Otte, 1981) Chortophaga australior Rehn and Hebard (Otte, Cibolacris salnalayucae Tinkham (Otte, 1981) 1984) crelzulata (Bruner) (Otte, 1981) Chortophaga viridifasciata (DeGeer) (Otte, 1984) (Thomas) (Otte, 1981) rabula Rehn and Hebard (Otte, 1984) Dichrolnorpha elegarzs (Morse) (Otte, 1981) Co7zosoa texalza (Bruner) (Otte, 1984) Dichromo~.phaviridis (Scudder) (Otte, 1981) Derotmerza haydenii (Thomas) (Otte, 1984) Eritettix abortivus (Bruner) (Otte, 1981) Derot~netzalaticinctum Scudder (Otte, 1984) Eritettix si~nplex(Scudder) (Otte, 1981) Dissosteira carolirza (Linnaeus) (Otte, 1984) Heliaula rufa (Scudder) (Otte, 1981) Dissosteira lorzgipe7z~zis(Thomas) (Otte, 1984) Ligurotettix pla~zuln(Bruner) (Otte, 1981) 38 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Encoptolophus sordidus costalis (Scudder) (Knut- Subfamily son, 1940) subgracilis texe~zsisBruner (Knut- elegans (Scudder) (Tinkham, 1948) son, 1940) bruneri (Caudell) (Wallace, Exirnacris superbunz (Hebard) (Otte, 1984) 1955) ~nagrzifica(Rehn) (Otte, 1984) Agroecotettix inodestzcs aristus Hebard (Hebard, (Say) (Otte, 1984) 1943a) Heliastus benjamini Caudell (Otte, 1984) Agroecotettix inodestus crypsidomus Hebard (He- Heliastus subroseus Caudell (Otte, 1984) bard, 1943a) Heliastus sulnichrasti (Saussure) (Otte, 1984) Ca~npylacanthalalnprotata Rehn and Hebard (Rehn Hippiscus ocelote (Saussure) (Otte, 1984) and Hebard, 1909) Hippopedon capito (Stgl) (Otte, 1984) Ca~npylacanthaolivacea olivacea (Scudder) (Knut- Leprzcs i7zter7nedizcs Saussure (Otte, 1984) son, 1940) Leprus wheeleri (Thomas) (Otte, 1984) Calnpylaca7ztha olivacea vivax (Scudder) (Helfer, Mestobregrna plattei corrz~gata(Scudder) (Otte, 1987) 1984) Chloropus cactocaetes Hebard (Helfer, 1987) Mestobregma terricolor Rehn (Otte, 1984) Clelnatodes larrae Scudder (Tinkham, 1948) pardalinus (Sausswe) (Otte, 1984) Dactylotz~ln bicolor pictuln (Thomas) (Knutson, Pardalophora lzalde7nanii (Scudder) (Otte, 1984) 1940) Pardalophora phoenicoptera (Burmeister) (Otte, Dactylotunz bicolor uariegatuln (Scudder) (Helfer, 1984) 1987) Pardalophora saussurei (Scudder) (Otte, 1984) Derzdrotettix quercus Packard (Helfer, 1987) Platylactista aztecus (Saussure) (Tinkham, 1948) Hesperotettix speciosus (Scudder) (Knutson, 1940) Psilzidia ainplicor~zisCaudell (Otte, 1984) Hesperotettix viridis pratensis Scudder (Knutson, Psilzidia fenestralis fenestralis (Serville) (Otte, 1984) 1940) Psirzidia fenestralis frater Rehn (Rehn, 1919) Hesperotettix viridis viridis (Thomas) (Knutson, bolli Scudder (Otte, 1984) 1940) Spharagernorz bolli i~zornatuinMorse (Tinkham, Hypochlora alba (Dodge) (Knutson, 1940) 1948) alexarzde7-i Hilliard (Hilliard, 2001) Spharagemo7z calnpestris (McNeill) (Otte, 1984) Melanoplus angularis Little (Knutson, 1940) Spkaragenzolz collare (Scudder) (Otte, 1984) Melanoplus angustipen7zis angustipen7zis (Dodge) Spharagemon cristatum (Scudder) (Otte, 1984) (Helfer, 1987) Spharage~nonequale (Say) (Otte, 1984) Melanoplus a.7zgustipennis irnpiger Scudder (Knut- Spharagetnon ~narrnoratapicta (Scudder) (Otte, son, 1940) 1984) Melanoplus aridus (Scudder) (Spooner, 1989) Trachyrhachis aspera Scudder (Otte, 1984) Mela7toplus arizorzae Scudder (Helfer, 1987) Trachyrlzachis coronata Scudder (Otte, 1984) Melanoplus bispinosus Scudder (Knutson, 1940) Traclzyrhachis kiowa fuscifrons (Stgl) (Otte, 1984) Melarzoplus bivittatus (Say) (Knutson, 1940) Traclzyrlzachis Iziowa kiowa (Thomas) (Otte, 1984) Mela7zoplus bowditchi Scudder (Rehn and Hebard, T7-imerotropis citrirza Scudder (Otte, 1984) 1909) Trilnerotropis cyaneiperznis Bruner (Otte, 1984) Melanoplus confusus Scudder (Knutson, 1940) TT-iinerotropislatifasciata Scudder (Otte, 1984) Melarzoplus dakini Hilliard (Hilliard, 2001) Triinerotropis ~nela7zopteraMcNeill (Otte, 1984) Melanoplus desultorius Rehn (Spooner, 1989) Triinerotropis pallidiperznis (Burmeister) (Otte, Melanoplus differentialis ~zigricansCockerel1 (Rob- 1984) erts, 1942) Trilnerotropis pistrinaria Saussure (Otte, 1984) Melanoplus discolor (Scudder) (Knutson, 1940) Tri~nerotropissalirza McNeill (Otte, 1984) Mela7zoplus euinera Hebard (Tinkham, 1948) saxatilis McNeill (Isely, 1935) Melanoplus femur-rubrum (DeGeer) (Knutson, Tri~nerotropisschaefferi Caudell (Otte, 1984) 1940) Tr-imerotropis strenua McNeill (Otte, 1984) Melanoplus flabellatus Scudder (Knutson, 1940) T~*opidolophusfor7nosus (Say) (Otte, 1984) Melanoplus flavidus Scudder (Knutson, 1940) Xarzthippus corallipes pa~ztherinus (Scudder) Mela7zoplus foedus fluviatilis Brunner (Knutson, (Helfer, 1987) 1940) INSECTA MUNDI, Vol. 15, No. 1, March, 2001 39

Melanoplz~sfoedus foedz~sScudder (Hebard, 1936) albolilzeata (Thomas) (Hubbell, 1960) Melarzoplus foedus iselyi Hebard. (Knutson, 1940) Schistocerca alutacea (Harris) (Hubbell, 1960) Mela7zoplus gladstoni Scudder (Helfer, 1987) Sclzistocerca arnerical~a(Drury) (Knutson, 1940) Mela~~oplusglaucipes (Scudder) (Knutson, 1940) Sclzistocerca darlzizifica (Saussure) (Knutson, 1940) Melarzoplz~sherbaceous Bruner (Rehn and Hebard, Schistocerca elnargirzata (Scudder) (Knutson, 1940) 1909) Schistocerca nitem (Thunberg) (Helfer, 1987) Melanoplus incor~spicuusCaudell (Hebard, 1917a) Schistocerca obscura (Fabricius) (Helfer, 1987) Melanoplus Keele7-i (Thomas) (Knutson, 1940) Schistocerca rubigil~osa(Harris) (Helfer, 1987) Melarzoplus lalzi~zus(Scudder) (Knutson, 1940) Schistocerca shosho7ze (Thomas) (Hubbell, 1960) Mela7zoplus inastigiphallus Strohecker (Strohecli- er, 1941) Subfamily Leptysminae Melarzoplz~smexica~zr~s (Saussure) (Helfer, 1987) Melanoplus occide7~talis(Thomas) (Helfer, 1987) Leptys rna margirzicollis (Serville) (Knutson, 1940) Melarzoplus packardii Scudder (Knutson, 1940) Stelzacris uitreiper~nis(Marschall) (Hilliard, 1982) Melarzoplus plebejus (Stil) (Helfer, 1987) Melanoplus po~zderosus(Scudder) (Knutson, 1940) Subfamily Romaleinae Melanoplus ponderosus viola (Thomas) (Hebard, 194 1) Braclzystola ~nagr~a(Girard) (Rehn and Grant, 1961) Melar~opluspurzctulatus arborez~sScudder (Rehn, Plzrynotettix 7-obz~stusoccultr~s Rehn and Grant 1946) (Rehn and Grant, 1961) Melanoplus regalis (Dodge) (Helfer, 1987) Plzry~zotettixrobustus 7-obustus (Bruner) (Rehn and Mela~zoplz~srusticus obovatipe~zrtisBlatcheley (Cop- Grant, 1961) pock, 1962) Phry7zotettix tsclzivavensis tsclzivave~zsis(Halde- Melarzoplus sa7zgr~i7zipesdefectz~s Scudder (Helfer, man) (Rehn and Grant, 1961) 1987) Plzryr~otettixtschiuave~zsis uerrucula,tus Glover Mela7zoplus sa7zgz~irzipes vulturnis Gurney and (Rehn and Grant, 1961) Broolis (Gurney and Brooks, 1959) Rolnalea guttatus (Stoll) (Rehn and Grant, 1961) Melanoplz~sscudderi latz~sMorse (Knutson, 1940) Taelziopoda equues (Burmeister) (Rehn and Grant, Melanoplzcs scudderi texensis Hart (Knutson, 1940) 1961) Melanoplus tennir~alisScudder (Scudder, 1897) Melanoplus texalzus (Scudder) (Knutson, 1940) Suborder Ensifera Mela7zoplus thornasi Scudder (Helfer, 1987) Melarzoplus tuberculatus Morse (Helfer, 1987) Family Gryllacrididae Mela~zopluswarlzeri Little (Knutson, 1940) Netrosolna ~zigropleuraScudder (Helfer, 1987) Paraidelno~zafratercula Hebard (Hebard, 1918) Subfamily Rhaphidophorinae Paraideino~zalatifurcula Hebard (Hebard, 1918) Paraidelnorza rniinica (Scudder) (Hebard, 1918) Ammobae~zetesplzrixocrzelnoides (Caudell) (Helfer, Paraidemolza nutti7zgi Yin and Smith (Yin and 1987) Smith, 1989) Ceuthophilus carlsbadensis Caudell (Hubbell, 1936) Paraidemolza olsorzi Yin and Smith (Yin and Smith, Cer~tlzophiluscorzicaudus Hubbell (Hubbell, 1936) 1989) Ceuthophilus cu7zicularis Hubbell (Hubbell, 1936) Paraide~nortapl~lzctata (Stil) (Hebard, 1918) Ceuthoplzilus divergens Scudder (Hubbell, 1936) Paratylotropidea bru7~7ze7-iScudder (Knutson, 1940) Ceuthophilus isletae Hubbell (Hubbell, 1936) Paroxya atlalztica Scudder (Knutson, 1940) Ceutlzophilus nodulosus Brunner (Hubbell, 1936) Paroxya clavz~liger(Serville) (Helfer, 1987) Ceuthophilus pallidus Thomas (Hubbell, 1936) Phaedrotettix accola (Scudder) (Hebard, 1917b) Ceuthoplzilus poli7zgi Hubbell (Hubbell, 1936) Phaedrotettix duinicola (Scudder) (Hebard, 1917b) Ceuthophilus reh~ziHubbell (Hubbell, 1936) Phaedrotettix pal~neri(Scudder) (Hebard, 1917b) Ceuthophilus secretus Scudder (Hubbell, 1936) Phaulotettix co~npl-essusScudder (Helfer, 1987) Ceuthophilus umbratilis Hubbell (Hubbell, 1936) Phaulotettix eurycercus Hebard (Helfer, 1987) Ceutkophilus z~tahelzsisThomas (Hubbell, 1936) Philocleo7~nigrouittata (Stsl) (Tinkham, 1948) Ceuthoplzilus variegatus Scudder (Hubbell, 1936) Photaliotes rzebrascensis (Thomas) (Knutson, 1940) Daihirzia brevipes Haldeman (Helfer, 1987) 40 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Daihiniella bellicosus (Hubbell) (Hubbell, 1936) Subfamily Daihiniodes haz~steriferu~n(Rehn) (Hubbell, 1936) Udeopsylla robusta (Haldeman) (Helfer, 1987) Paracyrtophyllus excelsus (Rehn and Hebard) (He- bard, 1941) Subfamily Stenopelmatinae Paracyrtoplzyllz~s7-0 bustus Caudell (Hebard, 1941) Pterophylla cainellifolia ca~nellifolia(Fabricius) Ste7zopelmatz~sfuscus Haldeman (Hebard, 1916a) (Hebard, 1941) Pterophylta cainellifolia intermedia (Caudell) (He- Family bard, 1941) Pterophylla furcata furcata (Caudell) (Hebard, 1941) Subfamily Pterophylla furcata laletica Hebard (Hebard, 1941)

Bucrates malivoleizs (Scudder) (Helfer, 1987) Subfamily breuipe7znis (Scudder) (Rehn and Hebard, 1915a) Amblycorypha floridana Rehn and Hebard (Rehn Conocephalus fasciatus (DeGeer) (Rehn and He- and Hebard, 1914c) bard, 1915a) A~nblycoryplzahausteca (Saussure) (Rehn and He- Corzocephalus lzygroplzilus Rehn and Hebard (Rehn bard, 1914c) and Hebard, 1915a) A~nblycoryplzainsolita Rehn and Hebard (Rehn Corzocephalus resacelzsis Rehn and Hebard (Rehn and Hebard, 1914c) and Hebard, 1915a) Anzblycorypha oblorzgifolia (DeGeer) (Rehn and Conocephalus saltans (Scudder) (Rehn and He- Hebard, 1914c) bard, 1915a) A~nblycoi*ypkarotuizdifoliaparvipennis St51 (Rehn Co7zocephalus spartinae (H. Fox) (Rehn and He- and Hebard, 1914c) bard, 1915a) Alnblycorypha uhleri St51 (Rehn andHebard, 1914~) Co7zocephalus strictus (Scudder) (Rehn and He- Arethaea a~nbulatorHebard (Rehn and Hebard, bard, 1915a) 1914a) Neoco7zocephalus exilisca~zorus(Davis) (Helfer, Arethaea arachrzopyga Rehn and Hebard. (Rehn 1987) and Hebard, 1914a) Neoconoceplzalus nebi~asce~~sis(Brunner) (Walker, Arethaea colzstricta co~nanclzeHebard (Hebard, 1998) 1936b) Neoconocephalus robustus crepita~zs(Scudder) Arethaea constricts co7zstricta Brunner (Hebard, (Tinkham, 1944) 193613) Neoco~zocephalustriops (Linnaeus) (Helfer, 1987) Arethaea gracilipes cerciata Hebard. (Rehn and Neoconocephalus velox Rehn and Hebard (Walker, Hebard, 1914a) 1998) Arethaea gracilipes gracilipes (Thomas) (Rehn and Orclzeli7nu7n agile (DeGeer) (Morris and Walker, Hebard, 1914a) 1976) Arethaea grallator (Scudder) (Rehn and Hebard, Orchelinzum bullaturn Rehn and Hebard (Rehn 1914a) and Hebard, 1915b) Arethaea phantas~naRehn and Hebard (Rehn and Orcheli~nu~ncalcaratuln Rehn and Hebard (Rehn Hebard, 1914a) and Hebard, 1915b) Arethaea se~nialataRehn and Hebard (Rehn and Orcheli7nuln concinnu7n concin7~uinScudder (Rehn Hebard, 1914a) and Hebard, 1915b) Dichopetala brevihastata Morse (Rehn and He- Or-clzeli7nu7ncorzcirzrzum delicatululn Bruner (Tho- bard, 1914b) mas and Alexander, 1962) Dichopetala castarzea Rehn and Hebard (Rehn and Orchelilnuln rzigripes Scudder (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala catinata Rehn and Hebard (Rehn and Orchelimurn vulgare Harris (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala emargirzata Brunner (Rehn and He- Pyrgocorypha uncui~zata(Harris) (Helfer, 198'7) bard, 1914b) INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Diclzopetala gladiator Rehn and Hebard (Rehn and Family Tridactylidae Hebard, 1914b) Dichopetala oreoeca Rehn and Hebard (Rehn and Tridactylus apicalis Say (Helfer, 1987) Hebard, 1914b) Trida.ctylus rnirzutis Scudder (Helfer, 1987) Dichopetala pollicifera. Rehn and Hebard (Rehn and Hebard, 1914b) Family Myrmecophilidae Dichopetala. seeversi Strohecker (Strohecker, 1941) Insara elegans (Scudder) (Tinkham, 1944) Myrmecophila ~zebrascensisLugger (Helfer, 1987) Microcentrum latifrorzs Spooner (Spooner, 1989) Microce7ztrunz rni~zusStrohecker (Strohecker, 1952) Family Microcerztru7n rlzo7nbifoliu7n (Saussure) (Helfer, 1987) Montezumirza ~nodesta(Brunner) (Walker, 1998) Acheta domesticus (Linnaeus) (Blatchley, 1920) Scz~dderia~urvica~uda-laticauda Brunner (Rehn Allorzemobius fasciatus (DeGees) (Hebard, 1913) and Hebard, 1914) Allo7zemobius fasciatus socius (Scudder) (Hebard, Scudderia furcata Brunner (Rehn and Hebard, 1913) 1914) Allorzernobius griseus funeralis (Hart) (Hebard, Scudderia furcata furcifera (Scudder) (Rehn and 1913) Hebard, 1914) Allonemobius sparsalus (Fulton) (Walker, 1998) Scudderia ~nexicana(Saussure) (Rehn and He- Arzaxiplza exigua (Say) (Helfer, 1987) bard, 1914) A~zurogryllusrnuticus (DeGeer) (Helfer, 1987) Scudderia texelzsis Saussure and Pictet (Rehn and Cycloptiliz~~na7ttillarurn (Redtenbacher) (Hebard, Hebard, 1914) 1916b) Cycloptiliurn cornpreherzdens cop~nprehende7zsHe- bard (Hebard, 1916b) Subfamily Decticinae Cycloptiliu~nco7nprehendens fortior Hebard (He- bard, 1916b) Anabrus simplex Haldeman (Caudell, 1907) Cycloptiliu~nhelainopuin Love and Walker (Love Caprzobotes fuligi~tosus(Thomas) (Tinkham, 1944) and Walker, 1979) Er-ernopedes covilleae Hebard (Tinkham, 1948) Cycloptiliurn pigr-urn Love and Walker (Love and Ere~nopedesscudderi Cockerel1 (Tinkham, 1948) Walker, 1979) Neobarrettia spirzosus (Caudell) (Helfer, 1987) Cycloptiliurn slossorzi Scudder (Love and Walker, Neobarrettia victoriae (Caudell) (Helfer, 1987) 1979) Pediodectes daedalus (Rehn and Hebard) (Rehn Cycloptiliurn squa~nosu~nScudder (Love and Walk- and Hebard, 1920) er, 1979) Pediodectes graztdis (Rehn) (Tinkham, 1944) Cycloptiliurn trigolzipalpu~nRehn and Hebard (Love Pediodectes haldelnanii (Girard) (Tinkham, 1944) and Walker, 1979) Pediodectes rzigro7na1-ginata (Caudell) (Rentz and Cyrtoxipha columbiana Caudell (Walker, 1969) Birchim, 1968) Eulze~nobiuscaroliltus (Scudder) (Hebard, 1913) Pediodectes pratti (Caudell) (Rentz and Birchim, Eurze~nobius carolinus neornexica7zus (Scudder) 1968) (Hebard, 1913) Pediodectes stevensoni (Thomas) (Tinkham, 1944) Falcicula hebardi Rehn (Blatchley, 1920) Pediodectes tirzkharni Hebard (Rentz and Birchim, Gryllodes sigillatus Walker (Alexander and Walker, 1968) 1962) firma Scudder (Alexander, 1968) Family Gryllotalpidae Gryllus fulto~ziAlexander (Alexander, 1968) Gryllus pennsylvarzicus Burmeister (Alexander, Gryllotalpa cultriger Uhler (Frank et al., 1995) 1968) Neocurtilla lzexadactyla (Perty) (Helfer, 1987) Gryllusperso~tatusUhler (Rehn and Hebard, 1915c) Scapteriscus borelli Giglio-Tos (Frank et al., 1995) Gryllus ruberzs (Scudder) (Walker, 1998) Scapteriscus vici~zusScudder (Frank et al., 1995) Gryllus texerzsis Cade and Otte (Cade and Otte, 2000) agitator Uhler (Helfer, 1987) 42 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Hoplosplzy7-~~1r~boreale (Scudder) (Helfer, 1987) Flook, P.K. and C.H.F. Rowell. 1997. The phy- Miogryllus lineatus (Scudder) (Hebard, 1915) logeny of the Caelifera (Insecta, Orthoptera) as Miogryllus ue7-ticalis (Serville) (Hebard, 1915) deduced from mtrRNA gene sequences. Mol. Neo7ze7nobius cube~zsis~normorzius (Scudder) (He- Phylog. Evol. 8: 89-103. bard, 1913) Frank, J.H., T.R. Fasulo, and D.E. Short. 1995. bipunctata (DeGeer) (Walker, 1963) Mcricliet Knowledgebase http://gnv.lfas.ufl.edu/ Neoxabea forinosa (F. Wallier) (Walker, 1967) -entI/mcricl.;et/. Oecantlzus a7-gentirzus Saussure (Walker, 1962) Gurney, A.B. 1940. A revision of the grasshoppers Oecanthz~scalifornicus Saussure (Walker, 1963) of the genus Orphulella fi.oin America North of Oecar~thz~scelerinictz~s T.J. Walker (Walker, 1962) Mexico. Entomologica Am.. 20: 85-157. Oecanthus exclarnationis Davis (Walker, 1963) Gurney, A.B. and A.R. Brooks. 1959. Grasshop- Oecantlzz~sfultoni T.J. Walker (Walker, 1963) pers of the Mexicanus group, genus Melano- Oecanthus leptogra~n~nusT.J. Walker (Walker, plus (Orthoptera: Acrididae). Proc. U.S. Natl. 1963) MUS. 110: 1-93. Oecanthus ~~iueus(DeGeer) (Walker, 1963) Hebard, M. 1913. A revision of the species of the Oecanthus pir~iBeutenmuller (Walker, 1962) genus Ne~nobius(Orthoptera: Gryllidae) found Oecanthus quadripurzctatus Beutenmuller (Walli- in North of the Isthmus of er, 1962) Panama. Proc. Acad. Nat. Sci. Phila. 65: 394- Oroclzaris saltator Uhler (Walker, 1969) 492. P1~yllopalpz~spulchellus (Uhler) (Walker, 1998) Hebard, M. 1915. The American species of the Trigo~~ido~nirnzcsbelfragei Caudell (Caudell, 1912) genus Miogryllus. J. New York Entomol. Soc. 23: 101-121. References Hebard, M. 1916a. A study of the species of the genus Stenopelma~tusfoundin the United States. Alexander, R.D. 1968. Life cycle origins, specia- J. New York Entomol. Soc. 24: 70-86. tion, and related phenomena in crickets. Quart. Hebard, M. 1916b. The Mogoplistinae (Ortho- Rev. Biol. 43: 1-41. ptera, Gryllidae) of the United States. Trans. Alexander, R.D. and T.J. Walker. 1962. Two Am. Entomol. Soc. 42: 135-160. introduced field crickets new to Eastern United Hebard, M. 1917a. New genera and species of the States (Orthoptera: Gryllidae). Ann. Entomol. Melanopli found within the United States and Soc. Am. 55: 90-94. Canada (Orthoptera: Acrididae). Parts X to Atkinson, T.A., P.G. Koehler, and R.S. Patter- XIV. Trans. Am. Entomol. Soc. 43:147-163. son. 1991. Catalog and atlas of the cocliroaches Hebard, M. 1917b. Notes on Mexican Melanopli () of North America North of Mexi- (Orthoptera: Acrididae). Proc. Acad. Nat. Sci. co. Misc. Publ. Entomol. Soc. Amer. 78: 85 pp. Phila. 69: 251-275. Blatchley, W.S. 1920. Orthoptera of Northeastern Hebard, M. 1918. New genera and species of America. Nature Publ. Co., Indianapolis, Indi- Melanopli found within the United States (Or- ana. thoptera: Acrididae). Trans. Am. Entomol. Soc. Cade, W.H. and D. Otte. 2000. Gryllus texensis n. 44: 141-169. sp.: a widely studied field (Orthoptera; Hebard, M. 1936a. New genera and species of the Gryllidae) from the . Melanopli found within the U.S. and Canada, Trans. Amer. Entomol. Soc. 126: 117-123. Part VIII. Trans. Am. Entomol. Soc. 62: 182- Caudell, AX 1907. The Decticinae (a group of 190. Orthoptera) of North America. Proc. U.S. Natl. Hebard, M. 193613. VI. A revision of the genus Mus. 32: 285-410. Arethaea (Tettigoniidae, Phaneropterinae). Caudell, A.N. 1912. A new genus and species of Trans. Am. Entomol. Soc. 62: 231-256. Gryllidae from Texas. Proc. Entomol. Soc. Wash. Hebard, M. 1941. The group Pterophyllae as found 14: 187-189. in the United States (Tettigoniidae: Pseudo- Coppock, S., Jr. 1962. The grasshoppers of Okla- phyllinae). Trans. Am. Entomol. Soc. 67: 197- homa (Orthoptera: Acrididae) State 219. Univ. Agric. Exp. Sta. Report P-399: 143 pp. Hebard, M. 1943a. New genera and species of Melanoplifound within the U.S., Part IV. Trans. Am. Entomol. Soc. 68: 49-66. INSECTA MUNDI, Vol. 15, No. 1, March, 2001

Hebard, M. 194313. The Dermaptera and Ortho- Rehn, J.A.G. 1942. On the locust genus Psoloessa pterous families Blattidae, Mantidae, and Phas- (Orthoptera: Acrididae: Acridinae). Trans. Am. midae of Texas. Trans. Am. Entomol. Soc. 48: Entomol. Soc. 68: 167-237. 239-313. Rehn, J.A.G. 1946. On the Pu~~ctulatusspecies- Helfer, J.R. 1987. How to know the grasshoppers, group of the genus Mela~~oplus(Orthoptera, crickets, cockroaches and their allies. Dover Acrididae, Cyrtacanthamidinae). Proc. Acad. Publications, Inc., New York. Nat. Sci. 98: 241-269. Hilliard, J.R., Jr. 1982. Endophytic oviposition by Rehn, J.A.G. and H.J. Grant Jr. 1961. A mono- Leptyslna. ~nargi~zicollis~nargi~zicollis and Ste- graph of the Orthoptera of North America (North 7~a.crisvitreipe~z~tis (Orthoptera: Acrididae: Lep- of Mexico), Vol. 1.Monographs of the Acad. Nat. tysminae) with life history notes. Trans. Am. Sci. Phila., 12: 257 pp. Entomol. Soc. 108: 153-180. Rehn, J.A.G. and M. Hebard. 1909. An ortho- Hilliard, J.R., Jr. 2001. Two new pterological reconnaisance of the Southwest- species in the Texanus group of the genus ern United States. Part 11: New Mexico and Mela~~oplr~s(Orthoptera: Acrididae: Melano- Western Texas. Proc. Acad. Nat. Sci. Phila. 61: plinae) with biological notes on the group. Trans. 111-175. Am. Entomol. Soc. 127: 31-68. Rehn, J.X.G. and M. Hebard. 1914a. Arevision of Hubbell, T.H. 1936. A monographic revision of the the orthopterous group Insarae (Tettigoniidae, genus Ceuthophilus (Orthoptera, Gryllacrid- Phaneropterinae). Trans. Am. Entomol. Soc. idae, Rhaphidophorinae). Univ. Fla. Pub. Biol. 40: 37-184. Sci. Series 2: 1-551. Rehn, J.A.G. and M. Hebard. 1914b. A study of Hubbell, T.H. 1960. The sibling species of the the species of the genus Dichopetala (Ortho- Alutacea group of the bird-locust genus Schis- ptera: Tettigoniidae). Proc. Acad. Nat. Sci. Phi- tocerca (Orthoptera, Acrididae, Cyrtacan- la. 66: 64-160. thacridinae). Misc. Publ. Mus. Zool., Univ. Mich. Rehn, J.A.G.and M. Hebard. 1914~.Studies in 116: 1-91. American Tettigoniidae (Orthoptera) I and 11. Isely, F.B. 1935. Acridianresearches within North- Trans. Am. Entomol. Soc. 40: 271-344. eastern Texas (Orthoptera). Entomol. News 46: Rehn, J.A.G. and M. Hebard. 1915a. A synopsis 37-75. of the species of the genus Co~~ocepha,lus Knutson, H. 1940. A key to the Acrididae (Ortho- (Xiphidiu~~zof authors) found in North America ptera) of Northeastern Texas with ecological North of Mexico. Trans. Am. Entomol. Soc. 41: notes. Field and Laboratory 8: 33-58. 155-224. Love, R.E. and T.J. Walker. 1979. Systematics Rehn, J.A.G. and M. Hebard. 1915b. A synopsis and acoustic behavior of scaly crickets (Ortho- of the species of the genus Orcheli~?zurlz.Trans. ptera: Gryllidae: Mogoplistinae) of Eastern Am. Entomol. Soc. 41: 11-83. United States. Trans. Am. Entomol. Soc. 105: 1- Rehn, J.A.G. and M. Hebard. 1915c. The genus 66. Gryllus as found in America. Proc. Acad. Nat. Morris, G.K. and T.J. Walker. 1976. Calling Sci. Phila. 77: 293-322. songs of Orcheli~nummeadow katydids (Tet- Rehn, J.A.G. and M. Hebard. 1920. Description tigoniidae), 1. mechanism, terminology, and of new genera and species of North American geographic distribution. Can. Entomol. 108: Decticinae (Orthoptera: Tettigoniidae). Trans. 785-800. Am. Entomol. Soc. 46: 225-265. Otte, D. 1981. The North American grasshoppers, Rentz, D.C. and J.D. Birchim. 1968. Revisionary Volume I Acrididae, , and Acrid- studies in the Nearctic Decticinae. Mem. Pacif- inae. Harvard Univ. Press, Cambridge, Massa- ic Coast Entomol. Soc. 3: 1-173. chusetts. Roberts, H.R. 1942. Two subspecies of Mela~~o- Otte, D. 1984. The North American grasshoppers, plus differe~~tialisand related new subspecies Volume I1 Oedipodinae. Harvard Univ. Press, from Mexico with discussion of their variation. Cambridge, Massachusetts. Trans. Amer. Entomol. Soc. 68: 151-166. Rehn, J.A.G. 1919. Descriptions of new and criti- Scudder, S.H. 1897. Revision of the orthopteran cal notes upon previously known forms of North group Melanopli (Acrididae), with special refer- American Oedipodinae (Orthoptera:Acrididae). ence to North American forms. Proc. U.S. Natl. Trans. Am. Entomol. Soc. 45: 229-255. Mus. 20: 293-294. 44 Volume 15, No. 1, March, 2001, INSECTA MUNDI

Spooner, J.D. 1989. Microce7~trumlatifror~s, a lidae: Oecanthinae). 1. The Genus ATeoxabea new species of angular-winged katydid from and the Ariveus and Varicor~~isGroups of the Southwestern United States (Orthoptera, Tet- Genus Oecanthus. Ann. Entomol. Soc. Am. 55: tigoniidae, Phaneropterinae). Trans. Am. En- 303-322. tomol. Soc. 114: 311-317. Walker, T. J. 1963. The taxonomy and calling songs Strohecker, H.F. 1941. Two new species of Ortho- of the United States tree crickets (Orthoptera: ptera from Texas. Ann. Entomol. Soc. Am. 34: Gryllidae: Oecanthinae). 11. The Nigricorr~is 539-542. Group of the Genus Oecal~tkus.Ann. Entomol. Strohecker, H.F. 1952. Descriptions of new spe- Soc. Am. 56: 772-789. cies and notes on North American Orthoptera. Walker, T.J. 1967. Revision of the Oecanthinae Am. Midland Naturalist 48: 683-688. (Gryllidae: Orthoptera) of America South of the Strohecker, H.F., W.W. Middlekauff, and D.C. United States. Ann. Entomol. Soc. Am. 60: 784- Rentz. 1968. The grasshoppers of California 796. (Orthoptera: Acridoidea). Bull. California In- Walker, T.J. 1969. Systematics and acoustic be- sect Survey 10: 177pp. havior of United States crickets of the genus Thomas, E.S. and R.D. Alexander. 1962. Sys- Oroclzaris (Orthoptera: Gryllidae). Ann. Ento- tematic and behavioral studies on the meadow mol. Soc. Am. 62: 752-762. grasshoppers of the Orchelirrzuin cor~ci7~izum Walker, T.J. 1969. Systematics and acoustic be- group (Orthoptera: Tettigoniidae). Occ. Papers havior of United States crickets of the genus Mus. Zool., Univ. Mich. 626: 1-30. Cyrtoxiplza (Orthoptera: Gryllidae). Ann. En- Tinkharn, E.R. 1944. Biological, taxonomic and tomol. Soc. Am. 62: 945-952. faunistic studies on the shield-back katydids of Walker, T.J. 1998. Handbook of katydids and the North American deserts. Am. Midland Nat- crickets of America North of Mexico. http:// uralist 31: 257-328. csssrvr.entnem.ufl.edu/-walker/handbook/ Tinkham, E.R. 1948. Faunistic and ecological hdbkhmpg.htm. studies on the Orthoptera of the Big Bend Wallace, H.S. 1955. Revision of the genus Aeol- Region of Trans-Pecos Texas, with especial oplides (Orthoptera, Acrididae). Ann. Entomol. reference to the orthopteran zones and faunae Soc. Am. 48: 453-480. of Midwestern North America. Am. Midland Yin, X. and R.L. Smith. 1989. Three new grass- Naturalist 40: 521-663. hoppers from the Western United States (Or- Walker, T.J. 1962. The taxonomy and calling songs thoptera: Acrididae). Pan-Pacific Entomol. 65: of United States tree crickets (Orthoptera: Gryl- 166-171.