Taxonomic Checklist of Chameleons (Squamata: Chamaeleonidae)
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65 (2): 167 – 246 © Senckenberg Gesellschaft für Naturforschung, 2015. 23.6.2015 Taxonomic checklist of chameleons (Squamata: Chamaeleonidae) compiled by Frank Glaw at the request of the Nomenclature Specialist of the CITES Animals Committee and the German Federal Agency for Nature Conservation (BfN) Funded by the German Federal Ministry of the Environment, Nature Conservation, Building and Nuclear Safety (BMUB) 2015 65 (2): 167 – 283246 © Senckenberg Gesellschaft für Naturforschung, 2015. 23.6.2015 Taxonomic checklist of chameleons (Squamata: Chamaeleonidae) Frank Glaw Zoologische Staatssammlung München, Münchhausenstraße 21, 81247 München, Germany; [email protected] Accepted 26.5.2015. Published online at www.senckenberg.de / vertebrate-zoology on 5.6.2015. Contents Abstract ............................................................................................................................................................. 174 Introduction ............................................................................................................................................................. 174 Collection acronyms ................................................................................................................................................. 174 Familiy Chamaeleonidae GRAY, 1825 ...................................................................................................... 175 Subfamily Brookesiinae ANGEL, 1942 .......................................................................................................... 176 Genus Brookesia GRAY, 1865 .................................................................................................................. 176 1. Brookesia ambreensis RAXWORTHY & NUSSBAUM, 1995 ................................................ 176 2. Brookesia antakarana RAXWORTHY & NUSSBAUM, 1995 ............................................... 176 3. Brookesia bekolosy RAXWORTHY & NUSSBAUM, 1995 .................................................... 176 4. Brookesia betschi BRYGOO, BLANC & DOMERGUE, 1974 ................................................ 176 5. Brookesia bonsi RAMANANTSOA, 1980 ........................................................................... 177 6. Brookesia brunoi CROTTINI, MIRALLES, GLAW, HARRIS, LIMA & VENCES, 2012 .............. 177 7. Brookesia brygooi RAXWORTHY & NUSSBAUM, 1995 ..................................................... 177 8. Brookesia confidens GLAW, KÖHLER, TOWNSEND & VENCES, 2012 ................................. 177 9. Brookesia decaryi ANGEL, 1939 .................................................................................... 177 10. Brookesia dentata MOCQUARD, 1900 ............................................................................. 177 11. Brookesia desperata GLAW, KÖHLER, TOWNSEND & VENCES, 2012 ................................ 177 12. Brookesia ebenaui (BOETTGER, 1880) ............................................................................ 178 13. Brookesia exarmata SCHIMMENTI & JESU, 1996 ............................................................. 178 14. Brookesia griveaudi BRYGOO, BLANC & DOMERGUE, 1974 ............................................ 178 15. Brookesia karchei BRYGOO, BLANC & DOMERGUE, 1970 ................................................ 178 16. Brookesia lambertoni BRYGOO & DOMERGUE, 1970 ...................................................... 178 17. Brookesia lineata RAXWORTHY & NUSSBAUM, 1995 ....................................................... 179 18. Brookesia micra GLAW, KÖHLER, TOWNSEND & VENCES, 2012 ...................................... 179 19. Brookesia minima BOETTGER, 1893 ............................................................................... 179 20. Brookesia perarmata (ANGEL, 1933) ............................................................................ 179 ISSN 1864-5755 169 Glaw, F.: Taxonomic checklist of Chamaeleonidae 21. Brookesia peyrierasi BRYGOO & DOMERGUE, 1975 ........................................................ 179 22. Brookesia ramanantsoai BRYGOO & DOMERGUE, 1975 .................................................. 179 23. Brookesia stumpffi BOETTGER, 1894 .............................................................................. 180 24. Brookesia superciliaris (KUHL, 1820) ........................................................................... 180 25. Brookesia therezieni BRYGOO & DOMERGUE, 1970 ........................................................ 180 26. Brookesia thieli BRYGOO & DOMERGUE, 1969 ................................................................ 180 27. Brookesia tristis GLAW, KÖHLER, TOWNSEND & VENCES, 2012 ....................................... 181 28. Brookesia tuberculata MOCQUARD, 1894 ....................................................................... 181 29. Brookesia vadoni BRYGOO & DOMERGUE, 1968 ............................................................. 181 30. Brookesia valerieae RAXWORTHY, 1991 ......................................................................... 181 Genus Palleon GLAW, HAWLITSCHEK & RUTHENSTEINER, 2013 .............................................................. 181 31. Palleon lolontany (RAXWORTHY & NUSSBAUM, 1995) .................................................... 182 32. Palleon nasus (BOULENGER, 1887) ................................................................................ 182 Palleon nasus nasus (BOULENGER, 1887) ............................................................... 182 Palleon nasus pauliani (BRYGOO, BLANC & DOMERGUE, 1972) .............................. 182 Subfamily Chamaeleoninae GRAY, 1825 ...................................................................................................... 183 Genus Archaius GRAY, 1865 ................................................................................................................... 183 33. Archaius tigris (KUHL, 1820) ........................................................................................ 183 Genus Bradypodion FITZINGER, 1843 ..................................................................................................... 183 34. Bradypodion atromontanum BRANCH, TOLLEY & TILBURY, 2006 .................................. 184 35. Bradypodion caeruleogula RAW & BROTHERS, 2008 ..................................................... 184 36. Bradypodion caffer (BOETTGER, 1889) .......................................................................... 184 37. Bradypodion damaranum (BOULENGER, 1887) .............................................................. 184 38. Bradypodion dracomontanum RAW, 1976 .................................................................... 184 39. Bradypodion gutturale (SMITH, 1849) ........................................................................... 185 40. Bradypodion kentanicum (HEWITT, 1935) ..................................................................... 185 41. Bradypodion melanocephalum (GRAY, 1865) ............................................................... 185 42. Bradypodion nemorale RAW, 1978 ............................................................................... 185 43. Bradypodion ngomeense TILBURY & TOLLEY, 2009 ...................................................... 185 44. Bradypodion occidentale (HEWITT, 1935) ..................................................................... 186 45. Bradypodion pumilum (GMELIN, 1789) ......................................................................... 186 46. Bradypodion setaroi RAW, 1976 ................................................................................... 186 47. Bradypodion taeniabronchum (SMITH, 1831) ............................................................... 186 48. Bradypodion thamnobates RAW, 1976 .......................................................................... 187 49. Bradypodion transvaalense (FITZSIMONS, 1930) ........................................................... 187 50. Bradypodion ventrale (GRAY, 1845) ............................................................................. 187 Genus Calumma GRAY, 1865 .................................................................................................................. 188 51. Calumma amber RAXWORTHY & NUSSBAUM, 2006 ........................................................ 188 52. Calumma ambreense (RAMANANTSOA, 1974) ................................................................ 188 53. Calumma andringitraense (BRYGOO, BLANC & DOMERGUE, 1972) ................................ 188 54. Calumma boettgeri (BOULENGER, 1888) ........................................................................ 188 55. Calumma brevicorne (GÜNTHER, 1879) ......................................................................... 189 56. Calumma capuroni (BRYGOO, BLANC & DOMERGUE, 1972) ........................................... 189 57. Calumma crypticum RAXWORTHY & NUSSBAUM, 2006 .................................................. 189 58. Calumma cucullatum (GRAY, 1831) .............................................................................. 189 59. Calumma fallax (MOCQUARD, 1900) .............................................................................. 190 60. Calumma furcifer (VAILLANT & GRANDIDIER, 1880) ...................................................... 190 61. Calumma gallus (GÜNTHER, 1877) ................................................................................ 190 62. Calumma gastrotaenia (BOULENGER, 1888) .................................................................