Ecological Aspects of Cercocarpus Montanus Raf. Communities in Central Utah
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Brigham Young University BYU ScholarsArchive Theses and Dissertations 1974-05-01 Ecological aspects of Cercocarpus Montanus raf. communities in central Utah David Lee Anderson Brigham Young University - Provo Follow this and additional works at: https://scholarsarchive.byu.edu/etd BYU ScholarsArchive Citation Anderson, David Lee, "Ecological aspects of Cercocarpus Montanus raf. communities in central Utah" (1974). Theses and Dissertations. 8011. https://scholarsarchive.byu.edu/etd/8011 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. ECOLOGICALASPECTS OF CERCOCARPUS MONTANUSRAF. COMMUNITIESIN CENTRALUTAH A Thesis Submitted to the Department of Botany and Range Science Brigham Young University In Partial Fulfillment of the Requirements for the Degree Master of Science by David Lee Anderson May 1974 This thesis, by David Lee Anderson, is accepted in its present form by the Department of Botany and Range Science of Brigham Young University as satisfying the thesis requirement for the degree of Master of Science. Typed by Sharon Bird ii ACKNOWLEDGMENTS The author wishes to express gratitude and apprecia- tion to the Instituto Nacional de Tecnologia Agropecuaria (I.N.T.A.), Argentina, for their financial support and interest in this project. Special appreciation is expressed to Dr. Jack D. Brotherson for suggesting the investigation of this problem and for his complete cooperation and guidance throughout the study and writing. Acknowledgment is also given to the Department of Botany and Range Science of Brigham Young University for their financial support and for their aid in supplying labora- tory space and field equipment necessary for the completion of this study. Appreciation is expressed to Dr. Ben Wood for his guidance and Dr. John F. Vallentine for his critical reading of the manuscript; to Dr. Sheldon Nelson for the provision of laboratory space and equipment for soil analyses; to Dr. Robert L. Park for statistical guidance. Special appreciation is expressed to my family. iii TABLE OF CONTENTS ACKNOWLEDGMENTS. iii LIST OF TABLES . vi LIST OF FIGURES . viii INTRODUCTION 1 LITERATUREREVIEW . 4 Taxonomy . • . • • . 4 Distribution and Comrrrunity Structure ........ 4 Forage Value . • . 8 METHODS. 9 General . 9 Vegetation 9 Soil . 17 RESULTSAND DISCUSSION 18 Description of Study Areas . ...... 18 Cat Canyon (Site No. 1) ............. 18 Upper Falls (Site No. 2) .•.......•.• 30 Edgemont (Site No. 3) . • . • • . 30 Spanish Fork Canyon (Site No. 4) . • . .. 31 Monk's Hollow (Site No. 5) .••••.• 32 Whiskey Springs (Site No. 6) .•...•.... 33 Stagecoach Inn (Site No. 7) . 34 Olmstead (Site No. 8) . ••....• 35 Cascade Springs (Site No. 9) 36 Sixth Water (Site No. 10) . 37 Red Hollow (Site No. 11) . 38 Thistle School (Site No. 12) . 38 Logan Canyon (Site No. 13) . 39 Timpooneke (Site No. 14) . 40 iv Fairview Canyon (Site No. 15) . 41 Santaquin Canyon (Site No. 16) ......... 41 Sheep Creek (Site No. 17) . 42 Canyon Glen (Site No. 18) ..•...... 43 Dividend (Site No. 19) . • . 43 Wanship (Site No. 20) . 44 Vegetational Analysis . 45 Cluster Analysis with C. montanus . 45 Cluster Analysis without C. montanus ...... 49 Linear Regression. .. 55 Multiple Regression . 63 C. montanus Height-Class Ratios , . • . 69 SUMMARYAND CONCLUSIONS. • . 76 BIBLIOGRAPHY . 79 APPENDIX . 81 V LIST OF TABLES Table Page 1. Prevalent Age Class as Applied to C. rnontanus 11 2. Deer Use as Applied to C. rnontanus . 12 3. Canopy-Coverage Classes 13 4. Prevalent Species List for All C. rnontanus Sites . 15 5. Physical Factors of the Twenty Study Sites 19 6. Soil Characteristics of the Twenty Study Sites 20 7. Percent Cover Summary of General Vegetation and Physical Factors . 21 8. Summary of C. montan~ parameters 22 9. Number of Deer Pellet Groups per Acre 23 10. Percent Cover Values for All Species in Twenty Study Sites . 24 11. Variables Used in Linear and Multiple Regression. 56 12. Factors Correlating with C. montanus Percent Frequency ............... 58 13. Factors Correlating with C. montanus Density per Acre . 59 14. Factors Correlating with C. montanus Canopy Cover 60 15. Factors Correlating with C. montanus Average Height . 61 16. Correlation of Factors to C. montanus Percent Frequency . .. 65 vi Table Page 17. Correlation of Factors to C. montanus Density per Acre . 66 18. Correlation of Factors to C. rnontanus Canopy Cover . 67 19. Correlation of Factors to C. rnontanus Average Height ................. 68 vii LIST OF FIGURES Figure Page 1. Location map of twenty C. montanus sites studied . 3 2. Distribution map of Cercocarpus montanus Raf. var. montanus in United States 6 3. Site clusters (with C. montanus) . 46 4. Frequency and density of C. montanus; percent clay and percent bare soil plotted against cluster groups .........•..•.... 47 5. Percent sand plotted against cluster groups . 47 6. Percent slope, total brush cover, total grass cover, and total litter cover plotted against cluster groups ....•............ 47 7. Soluble salts and pH plotted against cluster groups . 47 8. Total vegetation cover, total annual cover, per- cent silt, average height and number of stems of C. montanus plotted against cluster groups . 48 9. Total forb cover and total surface rock cover plotted against cluster groups ........• 48 10. Site clusters (without C. montanus) . 50 11. Total vegetation cover, percent litter and percent slope plotted against cluster groups .... 51 12. Deer pellet groups plotted against cluster groups 51 13. Shrub cover plotted against cluster groups .... 51 viii Figure Page 14. Total soluble salts and soil depth plotted against cluster groups .. 51 15. Total grass cover plotted against cluster groups . 52 16. pH plotted against cluster groups 52 17. Surface rock plotted against cluster groups 52 18. Percent clay plotted against cluster groups 52 19. Percent silt plotted against cluster groups 53 20. Percent sand plotted against cluster groups 53 21. Relation of C. montanus stands to elevation, exposure and percent slope ..... 64 22 (a-t). Height-class density relationships of C. montanus---- . 70 ix INTRODUCTION Statement of Problem Native forage shrubs common to the Intermountain Region contribute much feed to livestock and wildlife resources of Utah. As the human population increases and demands for meat products and hunting resources become more critical, precise knowledge of the ecological requirements of forage plants will be necessary in order to apply correct management policies for insuring long-lasting yields. Propa- gation of woody shrub species for other purposes (i.e., the revegetation of roadsides, strip-mined areas, etc.) also demands an exacting knowledge of minimum site characteristics. Such information and knowledge can best be gained through ecological studies directed toward ecosystems where these plants appear as dominants. Cercocarpus montanus Raf. is one such native shrub and is recognized as a forage plant useful to livestock and wildlife. Present literature reveals a paucity of information on the structure, dynamics and habitat requirements of C. montanus Raf. and its communities in Utah. The prime 1 2 objective of this study was to evaluate certain community characteristics of twenty sites in north central Utah (Fig. 1) where C. montanus is a principal component, thus providing a more complete understanding of this and its associated species. 3 .,-·-·-·-·-· -·-. - ·-·-· UTAH Scale: 1 inch= 45 miles .·· ·······..·~.. :. .... ....... --\. ·-·-·-·-·-·1 I ·•...... -··· i I I ' ! \I i i I i I i i I \ +' ·-·-·-·-·-·-·-·--·-·-·-·-·--·-·-·-·-·-· ·-·-·-·-·-·-·-·-·--·-·-·-·-·+ Fig. 1.--Location of twenty C. montanus sites. LITERATUREREVIEW Taxonomy Martin (1950) revised the genus Cercocarpus and delimited eight varieties of C. montanus Raf. The entity referred to in this paper is C. montanus Raf. var. montanus. Martin described this variety and listed several synonyms. Common names for C. montanus are varied. Members of the genus were called mountain mahogany by early pioneers in this area. However, since that time, the name mahogany has been given to the genus Sweitenia and the Federal Trade Com- mission has ruled that the name mahogany should not be used to designate any other genus. Following this rule, the new U.S. Forest Service check list approves "cercocarpus" as the genus' common name (Hayes and Garrison, 1960). Nevertheless, there are several other common names firmly entrenched in the literature (e.g., true mountain mahogany, birchleaf mahogany, and alder-leaf mountain mahogany). Distribution and Community Structure CercocarQus montanus Raf. var. montanus is a widely distributed browse species in the western United States, 4 5 ranging from South Dakota, Nebraska and Oklahoma, through Colorado, Wyoming and New Mexico to Utah, Arizona and Nevada (Fig. 2). The main habitat type is found in the eastern Great Basin and Rocky Mountains where it occupies rocky bluffs and mountain slopes between 3500 and 10,000 feet elevation (Medin, 1960; Martin, 1950; Pyrah, 1964). Utah exhibits one of the most widespread distribu- tions of this species. The Soil Conservation Service (1971) reports that C. montanus, found in all resource areas of Utah, grows within the 10-23 inch rainbelt on sites having a June deficiency of moisture. Some of the more important associated species reported for Utah and Colorado