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6013005692.Pdf УДК 598.112.13(4/5) ББК 28.693.34(051) П16 Панов Е.Н., Зыкова Л.Ю. Горные агамы Евразии. М.: Товарищество научных изданий КМК. 2016. 289 c., библ. 302 назв., 69 табл., 39 илл., 36 цветных вкладок (на англ. яз.). Книга представляет собой первый обзор на английском языке систематики и биологии ком- пактного таксона ящериц (около 20 видов), обитающих в пересеченных аридных и высоко- горных ландшафтах Евразии и северо-восточной Африки. Эти рептилии в последнее время привлекают пристальное внимание герпетологов из-за своей заметности, обязанной сугубо дневному образу жизни и высокой численности и плотности популяций, характерной для большинства видов. До сих пор интерес герпетологов был сосредоточен главным образом на систематике и номен- клатуре горных агам, а серьезные исследования по их образу жизни, экологии и поведению крайне немногочисленны. В этой книге наряду с обзором наиболее важной опубликованной информацией по изученным представителям таксона дана развернутая картина всех сторон биологии модельного вида – кавказской агамы Paralaudakia caucasia и его взаимоотношений с другим – хорасанской агамой P. erythrogaser. На протяжении 10 лет полевых исследований прослеживались биографии многих конкретных особей и изучались тонкие детали коммуника- тивного процесса в нескольких локальных популяциях. В основу исследования положен анализ социального поведения как фактора, интегрирующего дем в некую общность системной природы. Главной задачей было рассмотреть структуру в ее трансформациях из года в год как процесс изменений в поведении и социальных ролях взрос- лых особей и детенышей, ежегодно пополняющих контингент дема. Другие важные темы, затронутые в книге — это разнообразие носителей коммуникативной информации (оптическая, химическая, тактильная) у модельного и ряда других видов, а также характер взаимоотношений между близкородственными формами в зонах их контакта. К книге прилагается цифровой диск с многочисленными видеозаписями поведения 12 видов ящериц. On the front cover: Caucasian Rock Agama Paralaudakia caucasia. SW Turkmenistan. Photo by E.N. Panov © Е.Н. Панов, Л.Ю.Зыкова, 2016 ISBN 978-5-9907838-6-7 © Товарищество научных изданий КМК, издание, 2016 CONTENTS Authors preface .................................................................................................................... 8 Objectives, goals and background .................................................................................... 8 State of our knowledge of communicative behavior in lizards ...................................... 10 The state of things as deduced from systematic fi eld research data .............................11 The interdependence between the structure of the covers, coloration and behavior of lizards ............................................................................................... 12 The nature of theoretical generalizations on the evolution of signal behavior in lizards ........................................................................................ 13 Materials and methods ..................................................................................................... 14 Acknowledgements ........................................................................................................... 17 Introduction ........................................................................................................................ 18 Taxonomic structure of the genus Laudakia s. lato ........................................................ 20 Usual sources of disagreements in classifi cations of rock agamas .................................21 Intraspecifi c geographical variability.......................................................................... 21 Erroneous description of ‘new’ species based on specimens of a hybrid origin ......... 24 Undue haste in describing new forms .......................................................................... 24 Description of new taxa as an end in itself .................................................................. 24 Modern state of the rock agamas systematic ................................................................... 27 Chapter 1. General information on Eurasian rock agamas ........................................... 29 1.1. Morphological features ............................................................................................. 33 1.1.1. General characteristic ....................................................................................... 33 1.1.2. Size and proportions .......................................................................................... 37 1.1.3. Pholidosis .......................................................................................................... 39 1.1.4. Coloration .......................................................................................................... 51 1.1.5. Molt .................................................................................................................... 56 1.1.6. Karyotypes ......................................................................................................... 56 1.2. Habitats and numbers ............................................................................... 57 1.2.1. Terrain and substrate structure ......................................................................... 57 1.2.2. Temperature and the altitudinal range .............................................................. 58 1.2.3. Vegetation .......................................................................................................... 59 1.2.4. Numbers ............................................................................................................. 61 1.3. Social organisation and demographic structure of rock agama populations ...............................................................................................62 1.3.1. Social organisation ............................................................................................ 63 1.3.2. Use of space by rock agamas ............................................................................. 64 1.3.3. Sex and age structure of demes .......................................................................... 65 1.3.4. Population renewal rate .................................................................................... 66 1.4. Seasonal and diel activity patterns ............................................................................67 1.5. Thermoregulation ......................................................................................................70 1.5.1. Body temperature of active lizards .................................................................... 70 1.5.2. Body temperature of inactive rock agamas ....................................................... 72 1.5.3. Thermoregulatory behavior ............................................................................... 74 1.6. Diet and foraging behavior ....................................................................................... 75 1.6.1. Diet .................................................................................................................... 75 1.6.2. Variability of food spectra ................................................................................. 77 1.6.3. Foraging behavior ............................................................................................. 78 1.7. Reproduction .............................................................................................................79 1.7.1. Sexual maturity .................................................................................................. 79 1.7.2. Reproductive phenology .................................................................................... 80 1.7.3. Physiological changes during reproductive season .......................................... 81 1.7.4. Fecundity ........................................................................................................... 82 1.7.5. Mating behavior................................................................................................. 85 1.7.6. Female behavior during laying ......................................................................... 86 1.8. Social behavior and communication .........................................................................87 1.8.1. Territorial behavior ........................................................................................... 87 1.8.2. Socio-sexual relations ........................................................................................ 89 1.8.3. Communicative channels used in communication ............................................. 89 1.8.4. Signalling repertoire .......................................................................................... 90 1.8.5. Signal behavior in socio-sexual interactions ..................................................... 91 1.8.6. Species specifi city of push-ups movement.......................................................... 92 1.8.7. Functional aspects of communication in rock agamas ...................................... 93 1.9. Relationships between species and their place in biotic community ........................96 1.9.1. Spatial relationships between sympatric species of rock agamas ..................... 96 1.9.2. Interspecies hybridization .................................................................................. 97 1.9.3. Enemies and parasites ......................................................................................
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