1972] MOORE: CHELYOCARPUS ALLIANCE 85

16. A leaf of Itaya amicorum removed from plant press for photography after folding 17. An inflorescence of Itaya amicorum. Note (Moore, Salazar & Smith 8447). split petiole base immediately at left of tape measure and bracts of another inflorescence at left. The pinnate leaf in the background composing the Chelyocarpus alliance. is from a Scheelea (Moore, Salazar & Guti- Habit, leaf, inflorescence, flowers, and errez 9509). fruit have been illustrated photographic­ ally by Allen (1953) and Bartlett (1935). and persisting as fibrous margins on The root-spines on the trunk are unique the bases of the petioles and at length among palms and the androecium is fraying into slender elongate fibers at distinctive. A possibly specialized mode the apex; petioles· elongate, unarmed, of pollination is suggested by the ar­ the bases prominently split, more or less rangement of anthers at anthesis. In long-persistent in a latticework effect general, Cryosophila seems to be more below the crown, rounded below and advanced than Chelyocarpus, less so than channelled above, basally becoming bi­ Itaya. convex and rhomboid in section distally with obtuse margins, terminating adax­ Itaya H. E. Moore, gen. nov. ially in a small, deltoid, elevated hastula Palma solitaria hermaphrodita foliis basally plicately grooved, and abaxially induplicatis flabelliformibus ad 3/4 in a narrow deltoid margin; rachis not bipartitis lateraliter in 4--7 segmenta developed; blade flat, thin, three-fourths elongato-cuneata partitis. Inflorescentiae orbicular in outline, the segments circum­ 0 interfoliatae bracteis sterilibus 5 ramis scribing an arc of about 270 , divided 6-7. Flores bisexuales sepalis 3 et peta­ about three-fourths to the base at the lis 3 ad medium connatis, staminibus middle, each half again deeply divided 18-24, carpello 1. Fructus stigmatibus into several (4--7) elongate-cuneate, 4--7­ terminalib~s, seminis endospermio homo­ ribbed segments, these very shallowly geneo embryone prope basin sito. divided apically into briefly bifid 1­ Solitary, unarmed, hermaphroditic ribbed segments. palms of moderate size, the trunks with Inflorescences several, interfoliar, smooth cortex (drying roughened), elongate, arcuate; peduncle terete, bear­ naked except for fibrous residual sheaths ing a bicarinate abaxially split prophyll below the crown. and ca. 5 chartaceous, persistent, and at Leaves numerous in a spreading crown, length marcescent sterile bracts with induplicately palmate, flabelliform; tubular bases and inflated acute apices s eaths short, split opposite the petiole split on one side; primary branches 86 PRINCIPES [Vol. 16 6-7, each subtended by a persistent and Type: Itaya amicorum H. E. Moore. at length marcescent fertile bract similar Distribution: Peru, in seasonal rain­ to the sterile bracts but progressively forest at low elevations. smaller and the uppermost scarcely tub­ Chromosome complement: unknown. ular at the base, branches more or less Anatomical features: flattened, adnate to the rachis often Central vascular bundles of the nearly to the succeeding bract, the lower petiole with two phloem strands; branches again twice- to once-branched, late metaxylem elements with the upper ones once-branched into slender simple perforation plates and slightly sinuous rachillae, each rachilla often a characteristic ligule subtended by a linear acute bract. (Larry Klotz, personal communi_ Flowers perfect, borne singly in a cation) . spiral along the rachillae, each on a very Foliar and floral anatomy-see short pedicel subtended by a small acute Uhl, 1972a & b. bract; sepals 3, connate in an acutely 3-lobed cupular calyx; petals 3, connate Itaya amicorum H. E. Moore, sp. nov. about half their length in a 3-lobed (Fig. 13A, 15-17). corolla, the lobes rounded and erect at Caulis ad 5 m. altus foliis 12-20 ad anthesis, probably valvate in bud, sta­ 1.25 m. longis 2 m. latis. Inflorescentia mens 18-24, one or two opposite each ad 1.25 m. longa. Flores 3 mm. alti. sepal, remainder opposite petals, fila­ Fructus 2.4 cm. longus. ments connate basally in a fleshy tube Trunk to 5 m. high, 9 cm. in diam., less than half their length, slightly adnate gray. to corolla basally, fleshy and more or Leaves ca. 12-20, spreading from less subulateabove, anthers oblong in ascending petioles; sheath ca. 5 dm. outline, dorsifixed at the middle, versa­ long, densely light-brown floccose-tomen· tile, laterally dehiscent by longitudinal tose; petiole ca. 2.6 m. long above sheath, slits, exserted at anthesis, bifid at apex brown-furfuraceous basally, minutely and base, carpel 1, excentrically ovoid, deciduous-floccose distally, 1.3-1.4 cm. narrowed to a slender curved style and wide, 7 mm. thick at apex; hastula ca. oblique papillose stigma, ovule hemiana­ 1 cm. high, 1.5 cm. wide; blades to 1.25 tropous, attached adaxially at the base, m. long at middle, 2 m. wide, segments the short funicle bearing a prominent 4--7 on each side, to 1.25 m. long, 9-20 oblique aril. cm. wide at apex, again divided 1-7 cm. Fruit oblong-ovoid or subglobose with deep into segments 0.5-3 cm. wide and excentrically apical stigmatic residue; bifid to 3-10 mm., upper surface epicarp minutely granular-roughened glossy when dry with numerous promi­ and irregularly beset with minute per­ nent cross-veinlets, lower surface densely appressed white-tomentose, the principal forations; exocarp ca. 0.5 mm. thick nerves conspicuously elevated, tomentose with sclerosomes; mesocarp ca. 1 mm. or glabrescent, secondary nerves few, thick, white, dry, with anastamosing not elevated, cross-veinlets evident. fibers; endocarp not discrete: seed Inflorescence to 1.25 m. long or more, oblong-ovoid; hilum ellipsoid, subbasal, creamy white with brownish bracts at raphe-branches ascending-spreading; en­ anthesis; peduncle to 2.3 cm. wide near dosperm homogeneous; embryo excentri­ base; prophyll (on a smaller inflores­ cally basal; germination not known; cence) ca. 14, cm. or more long, 2.5 cm. seedling with undivided, elliptic eophylls. wide, peduncular bracts to ca. 6 dm.