AMEGHINIANA Revista de la Asociación Paleontológica Argentina Tomo VII Setiembre de 1970 N~ 3

THE LOWER BIVALVE \NPPbmS[[N IN PATAGONIA AND ITS PALEOGEOGRAPHIC SIGNIFICANCE FOR CONTINENTAL DRIFT

L i5 L5 c K_O\RYa]O S mzp K5 C. \SMMK\NS9

ABSTRACT: The subgenus Q·¡tmqm /Wuyqá~»á·q~z0 Bonarelli in Bonarelli and Nágera (1921) is considered to be a junior subjective synonym of Wmoo~qxxm Etheridge (1892). Wmoo~qxxm n~zm·qxxuu (Leanza, 1963) u» redescribed, and shown to be similar to Wmoo~qxxm uzoé·émám Waterhouse 1959 from New Zealand (Aptian? or Albian?) and M. ·~owñ~~pqz»u» (Etheridge ) írcrn Australia (Albian). The age of the Patagonian species u» probably Aptian. This genus u» important paleogeographically. By restoring the three regions with Wmoo~qxxm to their probable lower Cretaceous positions, based on paleomagnetic, faunal snd structural analyses, uá is shown that Wmoo~qáxm inhabited a restricted paleolatitudinal belt, close to 50° S to 60° S. It is now found over a much wider spread of latitudes, ranging from nearly 10° S to 50" S.

RESUMEN: Se considera al subgénero Q·¡nmqm /Wuyqá~»á·q~z0 Bonarelli /uz Bonarelli y Ná- gera, 1921) como un sinónimo de Wmoo~qxxm Etheridge (1892). Se redes cribe Wmoo~qxxm 774 zm·qxxuu (Leanza, 1963), señalando su similitud con M. uzoé·ímám Waterhouse, 1959, de Nueva Zelandia (Aptiano? o Albiano?) y M. ·~owñ~~pqz»u» (Etheridge) de Australia (Albiano). La edad de la especie patagónica es probablemente aptiana. Este género es importante paleo- gecgráficamente Situando las tres regiones con Wmoo~qxxm en sus probables posiciones en el Cretácico inferior, de acuerdo con datos paleomagnéticos, faunísticos y estructurales, se observa que' Wmoo~qáxm habitó un faja paleolatitudinal restringida, ubicada entre 500 S y 60° S. En la actualidad se la encuentra en sedimentos distribuidos en una gama más amplia de latitudes, desde casi los 10° S a 50° S.

INTRODUCTION SYSTEMATIC DESCRIPTION

Family Yde_YWSNKO Ichikawa 1958 Wmoo~qxxm is a short-ranged Lower Cre- (nom. transl. Cox 1961, p . .592) taceous bivalve genus, first discovered in Australia, and later found in two other Gen-rs Etheridge 1892 parts of the Southern Hemisphere, South America and New Zealand. TYPE SPECIES: (by original designation) Kíu4 The paleogeographic implications, ta- oéxm nm·wxu Moore 1870. xonomy and systematics of the South SYNONYM:Wuyqá~»á·q~z Bonarelli 1921, type American species are discussed in this species Kíuoéxm o~·nuqz»u»Moore 1870. paper. DIAGNOSIS:Medium-sized to large acline or prosocline shells with submedium or anteriorly • Departrnent of GeoJogy, University of Toronto, Ca- placed umbones, .1ft valve moderately to strong- Iy convex, right; V\J.lveflat, or weakly convex or nada. concave, posterior'; wings obtuse, may be poorly o Departrnent of Geology, McMasler University, Ca- defined or absent from right valve, anterior left nada. ear small, right anterior ear small and thick, -- 281 - usually with narrow interior subauricular notch, word. The ornament is costate (or costellate ), left byssaI gape small to Iarge, Costae of two and the inner surface of the shell is smooth at to four orders, subspinose 'in some species, maturity, though not of course in early growth growth lines prominent. Ligament area on plate stages. In our experience costae frequently affect in left valve, on posterior rim in right valve, the entire shell in juvenile stages, and so are both with opisthocline resilifer, large anterior like small plicae. left tooth and right socket, posterior adductor large submedian in each valve, pallial line pit- ted. a a (Leanza 1963) DrSCUSSION: Wuyqá~»á·q~z with type Pl. 1, figs. 1-10 species Kíuoéxm o~·nuqz»u» Moore 1870 8C98 Q·¡tmqm /Wuyqá~»á·q~z n. subgen.) was proposed by Bonare11i (in Bonarelli cfr. o~·nuqz»u» (Moore}-Bonarelli, p. 21, and Nágera, 1921) in the course of des- pl. 2, figs. 5a-b, 6. cribing Cretaceous fossils from Patago- 8C:B Q·¡tmqm cfr. o~·nuqz»u» (Moore l-Piar- nitzky, p. 74, pl. 5, figs. 24, 25. nia. Bonarelli was apparently unaware 8C?: Wuyqá~»á·q~z n~zm·qxxuun. sp., Leanza that Etheridge (1892) had included Kíu4 p. 222. oéxm o~·nuqz»u» in his genus Wmoo~qxxm3 8C?A Wuyqá~»á·q~z n~zm·qxxuuLeanza, p. 64, based on an allied species Kíuoéxm nm·w4 pl. 1, figs. 1-6. xu Moore. The two species differ con si- derably in shape, K5 nm·wxu being much NEOTYPE: The type material of the larger with a less convex left valve and species 5W5n~zm·qxxuuis amongst that des- less incurved umbo. It may eventually be cribed by Bonarelli (in Bonarelli and Ná- decided that such incurved species form gera, 1921), if we are to give priori ty to the first designation (Leanza, 1963, p. a distinct lineage of generic or subgeneric rank. But present evidence suggests that 222; 1967b, p. 63) over later designa- such "gryphaeation" recurred several ti- tions of another specimen( Leanza, 1967b, mes, possibly from different stocks, so pp. 64,65, pl. 1, figs. 1,2). Unfortuna- that placement of a11incurved Wmoo~qxxm tely, the two specimens figured by Bona- into Wuyqá~»á·q~z would obscure rather relli seem to be lost. A neotype should than clarify phylogeny. be chosen from topotype material. We therefore designate as neotype, specimen Leanza (1963, 1967b) correctly sho- MLP 11220, here figured on pl. 1, figs. wed that affinities of Wuyqá~»á·q~z lay 1, 3. with Yóá~ym rather than Q·¡tmqm3 and Cox (1969, p. L346) and Day (1969, DIAG OS1S:Medium-sized Wmoo~qxxm3left val- p. 159) synonymized Wuyqá~»á·q~z with ve convex with incurved umbo and wide byssal Wmoo~qxxm5 notch, right valve concave, costae moderately numerous (12-14 primaries), very fine and well- The Liassic species Q·¡tmqm »á·umám spaced, secondaries inconspicuous. Resilifers high- Phillipi (in Behrendsen 1891-2, p. 394, ly opisthocline under the umbo, notch in front pl. 22, figs. 5-6) was included in Wuyq4 of right hinge obscure. á~»á·q~z by Bonarelli. The specimens are MATERIAL: Three specimens with valves pcorly known, but the right valve does conjoined and seven left valves coIlected not appear to be like that of Wmoo~qxxm5 by A. C. Riccardi (MLP 11220-11229) and a composite mould of a left valve (MLP 2831) Ncr is the Callovian species Q·¡tmqm collected by DI. J. Frenguelli, aIl kept at Museo zqn·m»oqz»u»5 Meek and Hayden 1862 (d. de La Plata, Argentina. Imlay, 1948, p. 18; 1967, p. 8]) conge- LOCALITIES:Upper part of a small creek, west neric, though a1so referred to Wuyqá~»4 of Arroyo Calafate, south-east of Bahía de la á·q~z by BonareIli. Lancha .;E S, 72° 10' W), Lago San Martín, Santa Cruz, Argentina, in the upper part of the MORPHOLOGY:Speden (1968, 1969) has des- shaly Rio Mayer Formation, with Ku~x~oq·m»m·4 cribed the shell of Wmoo~0.qxxmas "plicate". It sqzáuzéy (Bonarelli ): must be stressed that the valves of Wmoo~qxxm This locality matches the one located on the are not plica te in the normal sense of the Geological Map of Bonarelli & Nágera (1921), - 282- PLATE 1

2 1

4

5 6 7

Wmoo~qxxm n~z~·qááu»(Lcanza ): 1,3. Externa' nnd inter na l view of neol~'pe, ~ILP 11220; left val ve ; 2,4. External and internal vi ew of left val ve ~I LP 11221 ; 5. RiglJt aspect of M LP 2831 ; 6. Right aspect of !\ILP 11222, right exterior worn ; 7. Lateral view of le(t val ve MLP 11223; 8, 10. lnl erna l ami ext.er- nal aspects of rig-ht val ve MLP 11222; 9. Right uspcct of MLP 11225 .. \1I spccimens save MLP 2831 from z part Á smal l creek west Á Ar-royo SxxÁx°6 iU ÁRxáx ~° x bxzáx6 bxÓ ix Ma rt¡n , eoll. A. C. Hiccar-di . J\ILP 2831 from probably same locality , eoll. J. Frenguelli.AlI specimens natural ixe,

- 283- irnmediately to the north of "Puesto Martín", One of the specimens described tJy Leanza in the "Serie arcilloso-m argos a ... ", close to the (1967b, pl. 1, fig. 6) carne frorn the same area contact with the 'Cretaceo". The same locality, and horizon, probably, has yielded one of the specimens DISTRIBUTION: In Patagonia this species, or here described (MLP 2831), and those described forms referred to Q·¡nmqm cf. o~·nuqz»u» by Bonarelli (in Bonarelli & Nágera, 1921 under (Moore ), has been mentioned frorn San Martín Q·¡nmqm /Wuyqá~»á·q~z n. subgen.) cfr. o~·4 Lake, in Viedma. Lake (north and east shore.. nuqz»u» (Moore ) (p. 21, pl. 2, figs., 5a-b, 6). Leanza, 1967b) and a borehole in Punta Norte, The latter specimens are missing from the col- 8 km north of Puerto Coig, Atlantíc Ocean, ap- leetion of Bonarelli and Nágera which is de- praximately 51° S (Feruglio, 1949, T. 1, p. 179). posited in the Instituto Nacional de Geología y Minería (Buenos Aires, Argentina). DIMENSIONS (in mm):

Muxtruurn °Óy Umboual Umbo fl'0111 i°zé° Leug th XzéÓá 'Yi

MLP 11222 44 4H 21 1050 14 29 MLP 1122.Q 36 43 22 1100 14 27 MLP 112'23 40 46 23 1000 15 25.5 MLP 2831 37 43 20.r¡ 1130 13 24

DESCRIPTIaN: Oóáq·zmxD The shells are me- opisthocline (80°) or vertical, or sometimes pro- dium-sized for the genus, the left valve mode- socline. Behind lies resilifer, opisthocline, ge- rately convex, with incurved anteriorly placed nerally bordered by low ridges, and marked by prosogyraus umbo, the right valve gently CO!l- strang cross-ridges, with its dorsal first formed cave.Anterior byssal gape in left valve lamellose, part curving and widenrng posteriorly from very large, up to 7 mm deep from commissure, under the umbo.Remainder of the hinge plate, Iargely infillec1 when valves shut by right valve, including the tooth, lightly striated by growth which lies partly in the concavity, and has ri lines. flange-like extension. Posterior wing moderately Hinge of right valve thickened, large socket long, gently convex in left valve, varying a little enclosed by two elevated ridges, Anterior ridge in length in different specimens, and probably merges with anterior flange to help close the often broken, feebly developed or absent from byssal notch of left valve. Posterior ridge of right valve as now preserved. socket fits into gap between left tooth and Ieft Nine to fifteen, usually 12 primary costae pit, adjoins feebly depressed, posteriorly facing extend from close to umbonal tip of left valve resilifer pit of right valve. to ventral margin with narrow convex crests, Musculature feebly irnpressed. Posterior nd- not divided longitudinally, and broadly concave ductor of left valve lies near rniddle of valve, interspaces. Secondary costae arise 15-35 mm. behind ligament pit, below mid-height, close to from the umbo and tertiary costae arise on the anterior hinge plate. Posterior adductor clearly anterior and posterior flanks, and probably defined on right valve in corresponding posi- around the ventral margins of large shells. Both tion. Three-four pits exposed along pallial line. sets remain finer than the primaries. Valve crossed by low growth Iines, 3-5 per mm, OCC¡J- sionally forming long "spines" across the costae. RESEMBLANCES: This species resembles Growth steps 3-10 mm aparto Only traces of cos- \4 uzoé·ímám Waterhouse 1959 of New tellae seen on right valves frorn the Bahia de la Zealand in its elongated outline, and mo- Lancha. A better preserved specimen MLP 2831 derately incurved left val ve, but the Ar- from Lago San Martín seems to have only pri- mary costae on the right valve opposing those gentinian form is larger with a wider of the left valve. umbo and an entirely concave right valve, The shell is lamellate, and 3-4 mm thick. whereas the right valve of the New Zea- Száq·zmxD Hinge plate of left valve roofs an land species is often partly convexo Cos- umbonal chamber, extending further from the tae are finer and less spinose and more hinge with increase in size, and infilled dorsally widely spaced with more numerous prim- with callus. Byssal gape bordered by a ridge aries and fewer tertiaries. On the New in some shells, not in others; tooth lies just in front of or below the umbonal tip, varies in in- Zealand species, primary costae are fewer, clination in different specimens, being generally 8-10 usually, and coarse. Secondary costae - 284- arise sooner on the New Zealand form but this is an error by a factor of 10 to and grow stronger.Growthlines are com- 12. Speden also counted 9-11 growth Ii- parable. In the right valve the anterior nes per mm in W5 uzoé·ímám3 yet his ru4 notch in front of the tooth buttress is gure /rus5 6), admittedly with no scale less incised than in ubS5uzoé·ímám3 and the given in the caption, suggests only 2 per resilifer of the left valve curves more mm, or if the specimen u» twice natural sharply back from under the umbo. size, 4 per mm. Dr. 1. G. Speden, New Speden (1968, p. 706) claimed that a Zealand Geological Survey, has kindly re- "higher number" of "plicae", as many as examined his material for us, and writes 18-22, were present around the ventral that 4-5 lamellae per mm occur on \4 uz4 margin of a new found left valve of \4 oé·ímám (TM 3913), and 3-4 per mm in uzoé·ímám3 in alleged contrast to the report W5 o~·nuqz»u» (WM 9047, UFQ 35518; by Waterhouse (1959, p. 493) of 8-10 WM 9048-UFQ 35520) noting that the costae. But Waterhouse had reported number increases to 4-6 per mm in bands, these as primary costae, correctly in our related to seasonal changes.(Dr. 1. G. opinion, and had counted in addition se- Speden letter, january, 1970). condary and tertiary costae, to make a The anterior auricular notch is more total of about 20-24 costae around the incised in W5 o~·nuqz»u»3 and the resilifer ventral margino Dr. Speden seerns to have apparently less opisthocline. All these dif- failed to consider the likelihood that his ferences suggest that W5 o~·nuqz»u»was "high number" of so-called plica e around less adapted to an attached habito A spe- the ventral margin included secondary and cimen figured as W5 o~·nuqz»u» by Lud- perhaps tertiary as well as primary costae. brook (1966, pl. 14, fig. 25, 28) from The Patagonian species was originally South Australia is almost as incurved as compared to Wmoo~qxxm o~·nuqz»u» Moore the Patagonian formo by Bonarelli (1921), and regarded as the W5 d. o~·nuqz»u» of Skwarko (1966, contemporaneous Patagonian counterpart p. 74, pl. 6, rus»5 1-11) from late Neoco- of this Australian species bv Day (1969, mian and early Aptian beds of Northern p. 159). The two are distinct. \4 o~·4 Territory, Australia, are somewhat closer nuqz»u» is a slightly smaller species with to the Patagonian species in incurvature less convex left valve, and convex rather and number and strength of the ribs. The than con cave right valve. The posterior shells are smaller, and the pos terior wing wings are slightly better defined, with much better developed. The right valve slightly alate cardinal extremities, and the was described as straight (meaning fIat? ) byssal gape of the left valve is smaller or slightly convexo The specimens are not than in the Patagonian formo Concentric conspecific with W5 o~·nuqz»u»3but come ornament seems to be slightly more pro- closer to W5 ·~owñ~~pqz»u» Etheridge, minent on W5 o~·nuqz»u» (2-3 per mm) which Skwarko erroneously synonymised and spines possibly less evident, but this with W5 o~·nuqz»u»5 needs verification. Costae are more nu- The type specimen of Wmoo~qxxm ·~ow4 merous at 17 or so, with 3-4 more arising á~~~pqz»u» (Etheridge, 1892, p. 448, pl. in front of the posterior wing, and more 8;2 rus5 15) u» a left valve with narrower closely spaced. This number may include umbo, longer posterior wing and coarser, a few secondaries that arise close to the more close-set costae, noted by Water- umbo. Other intercalated costae rapidly house (1959 p. 496) to be close to \4 uz4 become almost as strong as the primary oé·ímám5 Specimens figured by Ludbrook costae. Costae on the right valve are ver y (1966, pl. 14, figs. 22-24, 26) from Al- numerous. bian beds of South Australia verify this Speden (1968, p. 708) reported 14-16 similarity. They are smaller with a more growth lines per mm in W5 o~·nuqz»u»3 rounded outline, but agree in having a - 285- flat or concave right va1ve, and relatively v/ AMMONOID EVIDENCE: The speci- well spaced slender costae. It thus appears mens of Wmoo~Txoáxmn~zm·qxxuu (Leanza) to be closest of the Australian forms he re described were collected in beds with though many details have not been des- Ku~x~oq·m» m·sqzáuzéy (Bonarelli) and cribed. It is one of the younger species Xq~tun~xuáq» d. »qyuomzmxuoéxmáé»(Blain- in Australia, Day (1969) listing it as ville); the beds lie above the horizons of upper A1bian. Ko~zqoq·m» sp. and 'Kzox~oq·m»' ¡mám4 M. »uy¡xqó (Moore, 1870, pl. 11, fig. ¡°x Sto11ey, and below the horizon 3) is slightly incurved with few we11 spa- of ]mzym·áuz~oq·m»¡máms~zuoéy Bonare11i. ced costae. The age of M. n~zm·qxxuuhas been con- sidered as Albian by Leanza (1963, p. 222, 1967b, p. 69) fo11owing Piatnitzky AGE OF W5 LYXK\OVVSS (1938, p. 50). This was based on the wrong assignation by Bonarelli (in Bona- m0 MACCOYELLAAFFINITIES: M. n~zm4 relli and Nágera, 1921) of his material ·qxxuuappears to be relatively "advanced" from San Martin Lake, to the Albian ge- in its gryphaeoid morphology, but this nere Mxq~zuoq·m» Parona and Bonarelli, may reflect adaptation to substratum rath- Lqépmzáuoq·m» HitzeI, and atxusqxxm Ja- er than a young age. cob. These specimens are now placed in M. n~zm·qxxuu3M. uzoé·émám and M. ·~ow4 Ku~x~oq·m» Whitehouse /or5 Whitehouse, ñ~~pqz»u» a11occur in silty or muddy se- 1926, p. 206: Casey, 1961b, p. 132) a diment, though M. d. o~·nuqz»u» d. Skwar- genus known only from South Patagonia ko (1966) is found in sandstones. Wmo4 and Australia. The lower Albian occu- o~qxxm uzoé·ímám Waterhouse from New rrence reported by Collignon, 1949, from Zealand is moderately close. \X'aterhouse Madagascar, and quoted in Arke11 qx mx5 ( 1959, 1965), preferred a Lower Creta- (1957, p. L394) has been discounted by ceous, possibly Aptian age on fauna1 evi- Casey (1961b, p. 169). dence, but noted that M. uzoé·émám was According to Day (1969, p. 159), Ku~4 relatively advanced in incurvation and or- x~oq·m» is late Aptían because of its stra- nament so that it could prove younzer. tígraphic occurrence in proximity to ]mz4 The recent discovery by Norris and Wa- ym·áuz~oq·m» s. s. at San Martín Lake, terhouse (1970), that the so-called Juras- Patagonia, and because of its association sic Hawk Crag Breccia, is in fact, Upper near Tambo, Australia, with _·~¡mqéy Albian, may require some upward revi- Sowerby which shows crioceratid coiling. sion of the Wmoo~qxxm3 to perhaps as. Day (1969), quoting Casey (1961b, p. young as Albian. 132), considered that ]mzym·áuz~oq·m» Speden ( 1968) a1so supported a Lower s. s. was restricted to the Upper Aptian. Cretaceous age for M. uzoé·émám3but sta- Day (1967, 1969) also considered the ted that "an Upper age should Australian _·~¡mqéy and Ké»á·mxuoq·m» not be completely discounted". However, Whitehouse as of late Aptian age, follow- it is difficult to see a good reason for such ing Casey's (1961a) dating of the extra- an early age, especia11y since earlier re- Australian occurrences of species with ports of supposed Wmoo~qxxm in the Aus- crioceratid coiling. tralian jurassic, questioned byWatethopse (1959) and corrected by Brunnschwéiler An exclusively late Aptian age of ]mz4 (1960), have been shown false (confir- ym·áuz~oq·m» s. s. is not firmly established. med by Dr. R. Brunnschweiler, pers. Originally, Bonarelli and Nágera (1921) comm. to J. B. W., Ietter, 1960;~letter to considered S. ¡máms~zuoéy Bonarelli as Dr. 1. G. Speden, 1966, quoted in ~peden Upper Albian-Lower Cenomanian, because 1968, p. 709). of its occurrence at San Martin Lake above - 286- 4B·------'\----t--l------;~------_I_-_l

Tucu Tucu

SANTA

:r: ,.; ./

u /' CRUZ 4D LOCATION MAP AREA / / / \ 50.+-~ ---rl,.------tJ+------j ( ) \

V1éÓ8;8 7 Locality x Á fxxÓéx6 showing bAÓe ix I\'I~l'tíD ammonites referred to Albian genera. In Cenomanian while Arkell qá mx5 (1957) 1924, Spath (p. 74) mentioned that and Leanza (1967c) gave an Upper Ap- S. á·méá»qt~xpu Sinzow belongs to the tian-Lower Albian age. • Upper Aptian Aschiltaensis Subzone. The Two subgenera of ]mzym·áuz~qq·m» are same age was accepted by Whitehouse Lower Aptian: S. /_nqsmzqoq·m» 0 White- (1926, p. 197) for ]mzym·áuz~oq·m»3 who house is known from the Weissi Zone therefore considered Ku~x~qq·m» White- of northern Germany (Casey, 1961b, p. house as Aptian. Although most authors 132) and the type species of S. /]uz~íum0 have placed ]mzym·áuz~oq·m» in the Upper Sazonova, Kyy~zuáq» á·méá»ot~xpu Sinzow, Aptian since 1924, Maync (1949), still comes from the Lower Aptian of Russia reported the Patagonian occurrence as ° oi5 Casey, 1961b, p. 133). Upper Aptian -- 287 - is suggested by the report of S. ¡é»uxxéy Vqotuáq» Nowak, a genus of U. Albian-L. Cenomanian age (d. Arkell et al., 1957, m; zyxcR3 Jn~n Á[ Id ,1935 p. 17, fig. 16) from Greenland, since Rm4 p. t ~ig\ ., tle S ~hd~j~.Y77W7 yuáq» is of late Aptían-Iate Albian age was mentioned with unidentifiable T 3 (Arkell qx mx51957, p. L 216). Of ]mz4 ··uxuáupmqfrom Argentino Lake (Leanza, ym·áuz~oq·m» s. S., neither S. s·~qzxmzpu4 1967c, p. 173) suggest at least a Middle oéy Rosenkrantz nor 'Qm·zuq·um' ¡é»uxxm Albian age far the assemblage. Hawever, Ravn [probably a synonym of s·~qzxmz4 in his latest paper Leanza (1970, p. 199, puoéy according to Spath (1946, o. 7) 252) considered ]mzym·áuz~oq·m» Bona- and Casey (1961b, p. 131)] from Green- relli as U. Aptian, which would mean, on land, were collected uz »uáé (Ravn, 1911, his identifications, a downward extension p. 493; Rosenkrantz, 1934, p. 20). The of the stratigraphical range of Vqotuáq» assigned Upper Aptian age was based on and the _é··uxuáupmq as given in the Trea- that supposed for the genus, but as poi n- tise (1957). ted out by Maync (1949, p. 261) the pre- Another important point is the trans- sence of the Lower Aptian Nq»tmq»uáq» fer of ']uxq»uáq»' pq»y~oq·má~upq» Stolley, d. D. pq»nm»»u (Leymarie) and D. n~qs4 from San Martin Lake, to Zm·m»uxq»uáq» ímpu Rosenkrantz in the same are a could Imlay by Leanza (1967 a). This genus is suggest Lower Aptian for S. s·~qzxmzpu4 known from the Albian of Alaska and oéy5 Sardinia (Imlay, 1959, 1960; Wiedmann In Australia there seems to be no di- and Dieni, 1968). However, different spe- rect support for an Upper Aptian age of cies are involved. Leanza (1970, p. 251) ]mzym·áuz~oq·m» s. s. (d. Day, 1969, p. reported the patagonian species from le- 159). vels also containing ]mzym·áuz~oq·m» ¡m4 In Alexander Land and South Georgia, áms~zuoéy of U. Aptian age in San Mar- Antarctica, S. ¡máms~zuoéy was considered tin Lake, and associated with (?U.) Al- Upper Aptian (Wilckens, 1947; Howarth, bian ammonoids in Estancia La Vega (p. 1958) because this age was generally 252). This would indicate an U.Aptian- accepted for the genus. ( ?U.) Albian range for this species. Doubtful references to ]mzym·áuz~oq4 ]mzym·áuz~oq·m» Bonarelli, therefore, ·m» include 1) the ]mzym·áuz~oq·m» /]50 can be considered as Aptian, while any sp. (Casey, 1954), later called ]5I /]uz4 restrictions and/or extensions of its ran- ~íumI0 sp. nov. (Casey, 1961b), from ge are doubtfu1. The stratigraphic proxi- the Lower Albian Tardefurcata Zone of mity u~ ]5 uu¡°x to Ku~x~oq·m» u3 England; 2) the ]mzym·áuz~oq·m» n. sp. sqzáuzéy (Bonarelli) and Wmoo~qxxm n~4 mentioned by Breistroffer (1947) from zm·qxxuu(Leanza ) does not necessarily in- the Upper Aptian "Clansayesian" of Fran- dicate a late Aptian age for the last species. ce: 3) the dubious reports of this genus Casey's (1960) demonstration of a from imprecisely dated beds in Venezue- general phyletic trend in the ancylocera- la (Rod and Maync, 1954) and Papua tids from ancyloceratid to crioceratid coi- (Australasian Petr. Co., 1961). ling during the Aptian, agreeing with the Some dubious evidence suggests an Al- evolution of the heteromorphs as propo- bian age for ]mzym·áuz~oq·m»D Leanza sed by Wiedmann (1969), was given (1967c, p. 167) reported S. ¡máms~zuoéy as evidence by Day (1967, 1969) for a together with Pq·ésxu~oq·m» ¡umázuáwu late Aptian age for Ku~x~oq·m»5 At San Leanza. The statement refers to the San Martín Lake, 'M·u~oq·m»' [= _·~¡mqéy i Martín Lake area (Leanza, 1967c, p. 171, pqqowqu Favre with crioceratid coiling 173; 1970, p. 251). In the latter paper occurs below Ku~x~oq·m»d / . Feruglio, both species were reported together with 1949, p. 173). This would indicate an Vqotuáq» uyxmu Leanza. The presence of age not older than late Aptian for Ku~x~4 - 288- AL MAP SCALE GEOLOGIC ANCHA AREA 500 O 250 &>o 750 1000m OF BAHIA DE ~:NL MARTIN LAGO Z ARGENTINA ~

[3J iéó° ~ e Fosstutercvs Lccotrty IV 00 DAlluvlum pR ~ seno, Bovlders ond Grave!

CJ $ondy S.11 i~ Grcve! ~TIII

~ Posados Basalt ] OLlGOCENE ] ~ Rto Carbón U°° EOCENE TERTIARY

mrnII Puesto Nuevo nodesue ] SANTONIAN

~ Kochol~e Formonon J AP TIAN J CRETACEOUS_ u ~ R(o Mayer Formonon 2 8 $P"09h," Formonon J "NEOCOMIAN' I ~ • El Quemado Complex 1 '" •• • JURA5SIC ~ Arroyo de lo Mino Conglomero le I

Bahío dero Loncha Fo-mot.on ]LOWER ]8U UPPER OEVQNIAN ..J..(5 m

Fig. 2. - Geological map of Bahla de la Lancha a rea, Lago San Martín, A rr ow indicates Wmoo~qxxm occurreuce oq·m» m·sqzáuzéy and consequently for ( Archangelsky, 196 7 ). According to Dr. ]mzym·áuz~oq·m» ¡máms~zuoéy5 However, Archangelsky this flora is dominated by a Wiedmann (1969) also reported a recoi- new species of Mxm»»~¡~xxu»an3 d conside- ling trend in the crioceratíds in the Hau- ring the absence of angiosperms, the ho- terivian-; the material of 'M·u~4 rizon cannot be younger than early Albian oq·m»' pqqowqu Favre, therefore, needs to (pers. comm. 1970). be re-examined in order to establish its In summary, it seems reasonable to position with regard to the íterative evo- accept an Aptían age for the Ku~x~oq·m» lution of the heteromorphs during the assemblage wíth Xq~tun~xuáq» d. N. »qyu4 early Cretaceous. Moreover, during the om·umxuoéxmáé»(Blainville) and W moo~qxxm change from ancyloceratid to crioceratid n~zm·qxxuu (Leanza). The association of coiling of the ancyloceratids, different de- W5 n~zm·qxxuuwíth Rmáotq·uoq·m» Stanton grees of uncoiling probably coexisted as reported by Leanza (1967b, p. 69) does illustrated by the Lower Aptían _·~¡mqéy not contradicr this conclusion, since the n~u~q·nmzxquSowerby as figured by Casey Rmáotq·uoq·m» occurrence above Ku~x~oq4 (1960, text-fig. 7, pl. IV, fig. 1-3). Crio- ·m» at Posadas Lake does not establish ceratíd coiling, therefore, occurred also an Albian age for Rmáotq·uoq·m» as pos- in the Lower Aptian and ít seems rather tulated by Leanza (1963, 1967c). Day difficult to draw a line between the Lo- (1969) agreed with Woods' datíng of wer and Upper Aptían based on the de- Rmáotq·uoq·m» in Australia as Neocomian gree of coiling of the ancyloceratids. /rupq op. cit.). Day (1967) suggested that an Upper However, there exist the possibility Aptian age for Ké»á·mxuoq·m»was indicated of an extension of the stratigraphical ran- by its association wíth Vuátmzooxé» mé»4 ge of Wmoo~qxxm n~zm·qxxuuupwards, ac- á·mxu»Day; but the late Aptian age of cording to the report of this species with L. mé»á·mxu»is uncertain. Vuánmzoíoxé» ran- ( ?U.) Albian ammonoids in Estancia La ges from Barrernian to the Low~r Aptian, Vega (Leanza, 1970, p. 252). according to Casey (1960); Upper Ap- tian, therefore, seem unlikely in the ab- sence of strong supporting evidence. The PALEOGEOGRAPHY alleged similarity to two Californian spe- cies noted by Day (1967) is not convin- Wmoo~qxxm in Australia, New Zealand cing evidence for Upper Aptían; the 'Rq4 and Patagonia is one of many faunal and yunmoéxuáq»' yu·mnuxu» figured by Ander- floral links between these countries.Auth- son (1938) appears to be of middle to ors have particularly stressed the increa- late Aptían age (Popenoe qá mx531960) to se of common genera in the upper Jurassic judge from the associated fossils (Mur- (Howarth, 1958) and Cretaceous (Casey phy, 1956); the 'Rqyunmoéxuáq»' zqxqé» 1960; Fleming, 1962; Stevens 1965; Day is associated, according to Anderson 1969), coupled in at least New Zealand (1938), with 'Ko·~áqéátu»' mn~·usuzmxu» with weakening Cretaceous affinities to of Hauterivian-Barremian age (Srevens faunas from south Asia-Mediterranean 1965, p. 166; Murphy 1956, Text fig. 6; (Fleming, 1967). This latter phenome- Popenoe qá mx51960, chart 10 e), and non may have been related to Lower Cre- with 'Zm·mn~¡xuá~upq»'d. 'Z' oq··m»qz4 taceous cooling, which emphasized clima- »u» of late Aptían age (Popenoe qá mx53 tic zonation, discovered through oxygen 1960, p. 1508). isotope work on samples from Europe by The microflora, particularly the spores, Bowen (see 1966) and Australia and New occurring together with and above ]mz4 Zealand (Dorman and Gill, 1959; Clayton ym·áuz~oq·m» at San Martín Lake is very and Stevens, 1965). This lower Creta- similar to that in the Baqueró Formation ceous cool episode is reinforced by ana- - 29D - lysis of faunas in Queensland (Day, position. Both these are debatable, and 1969). Paleogeographers have thus be en overlook the fact that can cross encouraged to look for landlinks or sha- most barriers, given time (Waterhouse, llow sea-links between these regions, in 1969). the belief that similarity of life betokens One writer (Stevens, 1965) proposed either geographic proximity, or ready ac- that migrations along shallow ma- cess, either along continental bridges (Fle- rine shelves were obstructed periodically ming 1962), now foundered, or along by submarine canyons. He overlooked the pathways now displaced and disorganized fact that such canyons must have been by continental drift (Melville, 1966).Re- bordered by water shallow enough for constructions by continental drift also life to have slipped around. Prevention show New Zeáland, Tasmania and eastern of such migration must have been due Australia in close juxtaposition ( Carey to additional factors, such as temperature. 1958, Waterhouse and Vella, 1965, Wa- In our view, similarity of fauna mean terhouse, 1967a) shortening the migration similarity of living conditions, including distance for Wmoo~qxxm by 1000 miles. temperature, and similar rernperature is Oceanographic surveys have suggested at least on a world scale, likely to have that New Zealand is linked to Antarctic reflected similar paleolatitude. At present by oceanic ridges (Brodie 1964), Cullen the three major occurrences of fossil Wm4 ( 1968), Summerhayes (1967), W right oo~qxxm are within 12°_30°' S in north ( 1966), and van der Linden (1968) have Australia, South Australia, Queensland; proposed that New Zealand in the Meso- 38° -S. in New Zealand, and 49° S-51 ° S zoic lay close to the Antarctic, off Marie in Patagonia, this is a comparatively wide Byrd land, or the Ross Sea, this in turn scatter for a discontinuously distributed being a short geological step from the and locally restricted sessile genus. As a Malvinas Islands and on to Patagonia. general hypothesis geographically restric- Halpern ( 1968) has shown that Cretaceous ted and short-ranged animals inhabited si- granites extend in a chain from South milar latitudes. It is possible to test this America into Western Antarctica, and hypothesis from the distribution of Wm4 on into New Zealand (Aronson, 1965). oo~qxxm5 This view must be counterposed with Paleomagnetic data in Irving (1964), other evidence, that New Zealand was Creer (1967), show that Queensland lay linked with south-east Asia, through the across the parallels 50° and 60° S in the Indonesian and Melanesian ares (Carey, Cretaceous. Faunal ' support for this posi- 1958; Waterhouse, 1967a). To us, it tion is given- by Day (1969). Paleomag- appears feasible that Australia formed netic data (Irving, 1964, Creer, 1967; part of Gondwana, and that the South Valencio, 1969; Valencio and Vílas, Asian, Indonesian-Melanesian-New Zea- 1969); show that South America and the- land geological belt lay to the north and refore Patagonia has not changed its lati- east and continued through western An- tude much since Mesozoic (Irving, 1964; tarctica, Graham Land and Scotia are Creer et al, 1969; Francheteau and Scla- into South America. More faunal and pa- ter, 1969; Vilas and Valencio 1970a, b ). leomagnetic analyses are needed to show This is to be expected, in view of its the exact relative positions of New Zea- position between the South Pacific and land along this belt. South Atlantic centres of sea-flor spread- The significant aspect of the similari- ing oriented longitudinally. ties between east Australian, New Zea- Thus Patagonia probably lay at much land and South American faunas does not the same Cretaceous latitude as Queens- lie only in the possibility of interconnec- land. tion of shallow seas, or geographic juxta- New Zealand is now in an intermediate

- 291- Oegrees of mchnauon af growth line commissure from preceeding growth line 90

170 la

20 15 10 O __ HEIGHT-

Fig. 3. - Cornpar ison of curvatures of Jeft val ves of various Wmoo~qxxm3 based moclelled OD Waterhouse (1 g59, fiig. 4) posrnon, There is no published paleo- much similarity. Oxygen-isotope analyses magnetic data for its Cretaceous position. on Patagonian samples by Bowen (1961) The summary of faunas by indicates high temperatures of 23.7" e Waterhouse (1967; 1969,1970, in press) and 25.7" e for Hauterivian-Barrernian showed that eastern New Zealand lay at faunas, as interpreted by us (though assig- a paleolatitude of approximately 55" S, ned to the Aptian by Leanza 1963, probably north of Queensland and South 1967c). Another determination (Bowen, Australia, and higher than the paleolati- 1963) based probably on Xq~tun~xuáq» d. tude of Western Australia. Western New »qyuomzmxuoéxmáé»(Aptian ) yielded tern- Zealand formed part of Gondwana, being peratures of 30.7" to 32.7" C. Both ages similar in rocks and faunas to Tasmania. and identifications lack certainty, because Cretaceous faunas of New Zealand have of inattention to precise age or precise not been analysed for diversity or paleo- identification. latitude. It is our impression that the Such temperatures are considerably faunas are not quite as diverse as those of higher than those of Australia. From Lake either Australia or Patagonia. Although Eyre, central Australia, specimens of Zq4 authorities have implied that rhis might ·má~nqxé» gave average values of 13.8° e be due to isolation, we suggest it may and 16.5" e, comparable to temperatures reflect a more southerly, higher latitud e, found at about 40° south, and accepted by 5° to 10° than present, i. e. 45-55-600S. as reasonable by Day (1969). There is General faunal assemblages, and absen- no result for this part of the column in ce of dolomite and reefs or salt suggest New Zealand. Such a wide difference be- that it has occupied temperate latitudes tween these results would suggest very since the Mesozoic and Tertiary (Well- low latitudes for Patagonia, in contrast to man, 1959, p. 126). high latitudes for Australia, in conflict Paleotemperature work does not suggest with reconstructions based on paleomag- - 292- netic and paleontologic data. Most authors Nqéá»on5 sq~x5 Qq»qxx»onmvá15, 1891, 43: 369- agree that the Patagonian results seem 420, pIs. 22-25; II, 1892, 44: 1-42, pls. 1-4. BONARELLI,G. & NÁGERA,J. J., 1921. Obser- anomolously high (Bowen, 1966), and vaciones Geológicas en las inmediaciones del note must be taken of criticisms of poor Lago San Martín (Territorio de Santa Cruz). procedures in analyses and reliability of Nu·5 M·mx5Wuzm»3L~x5327 B, 39 pp., 6 pIs. the samples, possibly applicable to the BOWEN, R., 1961. Paleotemperature analyses of Patagonian results (Stahl and Jordan Belemnoidea and jurassic Paleoclimatology. g~é·zmx Mq~x~s3 69: 309-320, 2 figs. 196 9; Longinelli 196 9 ). -1963. 01'/0" Paleotemperature measurements on mesozoic Belemnoidea from Neuquen and Santa Cruz Provinces, Argentina. g~é·5 Zmxq~zá3 ACKNOWLEDGEMENTS 37: 714-718, 2 figs, 48C??5 Zmxq~áqy¡q·máé·qKzmx»u»5EIsevier Publ. Co., Amsterdam. 265 pp. We thank Dr. R. W. Day, Australian BREISTROFFER,M., 1947. Sur les zones d'Am- National University, Canberra, and Dr. monites dans l'Albien de France et d'Angle- 1. G. Speden, New Zealand Geological terre. _·mí5 Vmn5 Mq~x5Zmo~ ]ou5azuí5 Q·qz~4 Survey, Lower Hutt, for information as nxq3 26: 17-104. acknowledged in the text, and gift of du- BRODIE,J. W., 1964. Bathymetry of the New plicates. We also acknowledge helpful ex- Zealand region. N. Z. Nq¡á5 ]ou5 Szpé»5 \q»53 161: 1-55. change of information about the ammoni- BRUNNSCHWEILER,R. O., 1960. Marine fossils te ages with Dr. M. R. A. Thomson, Bri- from the Upper Jurassic and Lower Cretaceous tish Antarctic Survey. Photographs are of Dampier Península, Western Australia. Léxx5 by Mr: B. O'Donovan, and two text fi- Lé·5 Wuzq·5 \q»~é·o5 Qq~x5 Qq~¡n»5 Ké»á·mx53 gures by Mr. F. Jurgenheit, Department 59, 46 pp., illustr. CAREY, S. W., 1958. A tectonic Approach to of Geology, University of Taronto. In the Continental Drift Continent. N·uuá ]y¡~»5 text, systematics is principally by J. B. Mq~x5 Nq¡á5 azuí5 _m»ymzumD8AA4:<<5 Waterhouse, the ammonite ages by A. CASEY, R., 1954. Pmxouvq·mxxm3a new genus of C. Riccardi and the paleogeography by the Gault Ammonites, with a review of the fa- both of uso mily Aconeceratidae in the British Cretaceous. Z·~o5 Qq~x5 K»»~o5 V~zp53 65: 262-277, pl. 7, figs. 1-3. REFERENCES -1960. The Amrnonoidea of the Lower Green- sand. Part 1. Zmxmq~zá~s·m¡nuomx ]~ouqá3 113: I-XXXVI; 1-44, pIs. 1-10. ANDERSON,F. M., 1938. Lower Cretaceous De- posits in California and Oregon. Qq~x5 ]~o~ 48C?8 u4 The Amrnonoidea of the Lower Green- Ky5 ]¡qo5 Zm¡q·3 16: x + 339 pp., 84 pIs. sand. P¡¡rt lI. Zmxmq~zá~s·m¡nuomx]~ouqá3 88;D 45-118, pIs. 11-25. ARCHANGELSKY,S., 1967. Estudio de la For- mación Baqueró, Cretácico Inferior de Santa 48C?8 n5The Arnrnonoidea of the Lower Green- Cruz, Argentina. \qí5 Wé»5 Vm Zxmám(3n. s.}, sand. Part lII. Zmxmq~zá~s·m¡nuomx ]~ouqá3 Zmxq~zá535: 63-171, 1 mapa, 13 cuadros, Iáms. 115: 119-216, pIs. 26-35. 1-8. CLAYTON,R. N. & STEVENS,G. R., 1965. Pa- ARKELL,W. J., KUMMEL, B. & WRIGHT,C. W., Ieotemperatures of New Zealand Belemnites. 1957. Mesozoic . In R.C. MOORE ]ámnxq S»~á~¡q»uz Yoqmz~s·m¡tuo ]áépuq» mzp (Ed.). _·qmáu»q ~z Szíq·áqn·máq Zmxq~zá~x~s5 Zmxq~áqy¡q·máé·q»3 Pisa. 1-6. L. W~xxé»omD80-437. Univ. Kansas Press. COLLIGNON,M., 1949. Recherches sur les faunes ARONSON,J. L., 1965. Reconnaissance Rubidium- albiennes de Madagascar. 1, L'Albien d'Am- Strontium Geochronology of New Zealand barimaninga. Kzz5 Qq~x5 ]q·í5 Wuzq» Wmpm4 Plutonic and Metamorphic Rocks.N.Z. g~é·5 sm»qm·316; 128 pp., 22 pls. Qq~x5 Qq~¡n»53 8: 401-23. Cox, L. R., 1961. New genera and subgenera of AUTRALASIANPETROLEUMCo., 1961. Geologi- Mesozoic Bivalvia. Zmxmq~zá~x~s3 4 (4): 592- cal Results of Petroleum exploration in West- 598. ern Papua. 1937-1961. g~é·5 Qq~x5]~o~ Ké»4 -1969. Family Oxytomidae Ichikawa 1958. Sz á·mxum38 (1): 1-133. R. C. MOORE(Ed.) _·qmáu»q ~z Sz~q·áqn·máq BEHRENDSEN, O., 1891-92. Zur Geologie des Zmxq~zá~x~s3 Part N, vol. 1, W~xxé»om 6 Lu4 Ostabhanges der argentischen Cordillere. fqé5 ímxíum3344-346, fig. e 70. Univ. Kansas Press. - 293 CREER, K. M., 1967. A synthesis of World-wide ICHIKAWA,K., 1958. Zur Taxionomie und Phy- palaeomagnetic data, Sz RUNCORN,S. K. (Ed.) logenie der Triadischen "Pteriidae" (Lamelli- Wmzáxq» 78 ánq Om·át mzp _q··q»á·umx Zxmzqá»5 branch.). Zmxmq~zá~s·m¡tuomlII, Abh. A: 131- 351-382, 10 figs., 10 tables. 212, pls. 21-24. CREER,K. M., EMBLETON,B. J. J. & VALENCIO, IMLAY, R. W., 1948. Characteristic Marine ju- D. A., 1969. Comparison between the Upper rassic Fossils from the \'V'estern Interior of the Paleozoic and Mesozoic Palaeomagnetic poles United States. h4 S. Qq~x5 ]é·í5 Z·~65 Zm¡53 for South America, Africa and Australia. Om·át 214 B: 13-33, pls, 4-9. Zxmzqá5 ]ou5 Vqááq·»3 7: 288-292, 5 figs., 3 -1959. New genera of early Cretaceous (AI- tables. bian ) ammonites from Alaska. g~é·5 Zmxq~zá53 CULLEN, D. J., 1968. Movement of Sialic Blocks 33: 179-185, pls. 29-30. oblique to the Direction of sea-floor spread- -1960. Early Cretaceous (Albian) Ammonites ing. Om·át Zxmzqá5]ou5 Vqááq·»35: 123-126. from the Chitina Valley and Talkeetna Moun- DAY,R. W., 1967. Vuátmzoxé» mé»á·mxu» sp. nov. tains, Alaska. h4 ]5 Qq~x5]é·í5 Z·~r5 Zm¡qx.3 A new ammonite from the Aptian of Queens- 354-D: 87-114, pIs. 11-19, text-figs. 21-24. land. T~é·5 Qq~x5 ]~o~ Ké»á·mxum314 (1): 19- -1967. Twin Creek Limestone (Jurassic) in the 22, pIs. 2. Western Interior of the United States. a5 ]5 --1969. The Lower Cretaceous of the Great Ar- Qq~x5]é·~5 Z·~r5 Zm¡q·3 540: 1-105, pIs. 1-16, tesian Basin. Sz K. S. W. CAMPBELI. (Ed.) text-figs. 1-18. ]á·máus·m¡t mzp Zmxq~zá~x~s O»»m»3 uz R~4 lRVING, E., 1964. Zmxq~ymszqáu»y mzp uám¡¡x» u4 z~é· ~r N~·~át V°ññ2140-173, figs. 28-35. omáu~z á~ Qq~x~suomx mzp Qq~¡n»uomx ¡·~nxqy»5 Austral. Nat. Univ. Press. John Wiley and Sons, New York, 399 pp. DORMAN, F. H. & GILL, E. D., 1959. Oxygen LEANZA, A. F., 1963. Zmáms~zuoq·m»gen. nov. Isotope Paleotemperature measurements on (Binneyitidae) y otros ammonites del Cretáci- Australian fossils. Z·~o5 \~5 ]~o~ buoá~·um3 co superior de Chile meridional, con notas 71: 73-98. acerca de su posición estratigráfica. L~x5 Komp5 Xmo5 M»5 /Mnm50343: 203-225, 4 pIs. ETHERIDGE, R. (Jr.), 1892. Sz ]ACK, R. L., ETHERIDGE, R., 1892. _tq Qq~x~s mzp Zm4 48C?Am5 Descripción de la fauna de Zxmoqzáu4 xmq~zá~x~s ~r [éqqz»xmzp mzp Xqñ Qéuzqm5 oq·m» del Cretácico superior de Patagonia Aus- tral con consideraciones acerca de su posición Government Printer, Brisbane, Dulau & Co., London, 768 pp., 44 pls. estratigráfica. L~x5 Komp5Xmo5 M»5 /Mnm5046 3 ( 1): 5-28, pIs. 1-10. FERUGLIO, E., 1949. Nq»o·u¡ouz Qq~xsuom pq 48C?A n5Descripción de la especie tipo de Wu4 xm5Zmáms~zum51: I-XV, 1-334, 60 láms., 78 figs. yqá~»á·q~z Bonarelli, 1921,Emend. Leanza, Dir, Gral. Yac. Petr. Fisc., Buenos Aires. 1963 (Moll., Pel.) del Cretácico de Santa FLEMING, C. A., 1962. New Zealand Biogeopra- Cruz (Patagonia Austral). L~x5 Komp5 XmoM»5 5 phy. A Paleontologist's Approach. Témám·m /Mnm50 346 (1): 61-70, 1 pl. 10 (2): 53-108. ' 48C?Ax4 Anotaciones sobre los fósiles Jurásico- -1967. Biogeographic Change related to Meso- Cretácicos de Patagonia Austral (Colección Fe- zoic orogenic history in the south-west Paci- ruglio) conservados en la Universidad de Bo- fic. _qoá~z~¡n»uo»3 4 (4-6): 419-427. logna. Koám Qq~x5 Vuxx~mzm39: 121-187, pIs. FRANCHETEAU,J. & SCLATER,J. G., 1969. Pa- 1-3. leomagnetism of the Southern Continents and -1970. Arnmonites nuevos o poco conocidos del Plate Tectonics. Om·án Zxmzqá5]ou5 Vqááq·»3?D Aptiano, Albiano y Cenomaniano de los An- 93-106, 4 figs. des Australes con notas acerca de su posición estratigráfica. \qí5 K»~o5 Qq~x5 K·s5325: 197- FREBOLDH.,, 1935. Marines Aptien von der Kol- 261, figs. 1-42. dewey Insel (nordliches Ostgronland ). Wqpp5 U}ñ42 95: 1-112, 8 pIs., 20 figs. LONGINELLI,A., 1969. Oxygen-18 variations in Bclernnite Guards. Om·án Zxmzqá5]ou5 Vqááq·»3 GREGORYJ,. W. & SMITH, F., 1902. A new Arn- 7 (2): 209-212. monite from the Cretaceous rocks of Queens- LUDBROOK,N. H., 1966. Cretaceous Biostrati- Iand. Z·~o5 \~5 ]~o~ buoá~·um3(2): 15 n. S. graphy of the Great Artesian Basin in South 141-144, pIs. 22. Australia. Qq~x5 ]é·í5 ]~éát Ké»á·mxumLéxu53 HALPERN, M., 1968. Ages of Antarctic and Ar- 40: 1-223, illustr. gentine Rocks bearing on Continental Drift. MAYNC,W., 1949. The Cretaceous Beds between Om·át Zxmzqá5]ou5 Vqááq·»35: 159-167. Kohn Island and Cape Franklin (Gauss Pe- HOWARTH, M. K., 1958. Upper Jurassic and ninsula), Northern East Greenland. Wqpp5 Cretaceous Ammonite Faunas of Alexander x3}2 133 (3): 1-291, 4 pls, Land and Graham Land. Zmxw5 S»5 Nq¡]é·í55 MEEK, F. B. & HAYDEN, F. V., 1861. Descrip- ]ou5 \q¡53 21: 1-16, pIs. 1-5. tions of new lower , (Primorrlial ), ju- - 294- rassic, Cretaceous, and Tertiary Fossils, col- SPEDEN, 1. G., 1968. Additional details of the lected in Nebraska by the Exploring Expedi- morphology of Wmoo~qxxm uzoé·ímámWater- tion under the command of capto Wm. F. house and a reconsideration of the age of the RAYNOLDS,U. S. Top. Engrs., with some re- genus Wmoo~qxxm5 Xqñ fqmxmzpg~é·5 Qq~x5 marks on the rocks from which were obtain- Qq~¡n»53 11 (3): 706-710. ed. Z·~o5 Komp5 Xmáé·mx ]ou5Ztuxm¡qx¡num3 -1969. Lower Cretaceous Marine fossils, in- (publ. 1862): 415-447. cluding Wmoo~qxám sp. from the Whatarangi -1864. Invertebrates, Sz Palaeontology of the Formation, east side of Palliser Bay, New Zea- Upper Missouri. A report upon collections land. _·mz»5 \~5 ]~o~ Xqñ fqmxmzp Om·át made principally by the expeditions under com- ]ou536 (12): 143-153, pl. 1. mand of Lieut. G. K. WARREN, U. S. Top. STAHL, W. & JORDAN,R., 1969. General Consi- Engrs. in 1855 & 1856, Part 1, ]yuát»~zumz derations on Isotopic Paleotemperature deter- M~zá·unéáu~z» á~ Uz~ñxqpsq172: I-IX, 1-135, mina tions and analyses on J urassic ammonites. pIs. I-IV. Om·án Zxmzqá5 ]ouV5mááq·»36: 173-178. MELvILLE, R., 1966. Continental Drift, Mesa- STEVENS,G. R., 1965. The jurassic and Creta- zoic Continents, and the migrations of the ceous Belemnites of New Zealand and a review Angiosperms. Xmáé·q 211: 116-120. of the Jurassic and Cretaceous Belemnites of MooRE, c., 1870. Australian Mesozoic Geology the Indo-Pacific Region. Xqñ fqmxmzp Qq~x5 and Palaeontology. [ém·á5 g~é·5 Qq~x5 ]~o~ ]é·í5 Zmxq~zá5Léxx5336: 1-283, pls. 1-25. V~W5326: 226-61, pls, 10-18. SUMMERHAYES,C. P., 1967. New Zealand re- gion volcanism and structure. Xmáé·q3 215: MURPHY,M. A., 1956. Lower Cretaceous strati- 610-611. graphic units of northern California. Ky5 K»»~o5 Zqá·mx5 Mq~x5Léxá5340: 2098-2119, 6 VALENCIO,D. A, 1969. El Paleomagnetismo de figs, algunas magmatitas del Triásico superior, Gru- po Cacheuta, Prov. de Mendoza, Rep. Argen- NORRIS, G. & WATERHOUSE,J B., 1970. Age tina. \qí5 Kyo5 Mq~x5 K·s5324 (3): 191-198, of the Hawks Crag Breccia. _·mz»5 ·~~ ]~o~ 3 figs. Xqñ fqmxmzp Om·át ]ou537 (13): 241-250. VALENCIO,D. A., VILAS, J F., 1969. Age of PIATNITZKY, A., 1938. Observaciones geológi- the Separation of South AmerÍca and Africa. cas en el oeste de Santa Cruz (Patagonia). Xmáé·q3 223: 1353-1354, 2 figs. L~x5 Szx5 Zqá·53165: 45-85, láms. 1-10. VAN DER LINDEN, W. J M., 1968. Rotation of POPE 'OE, W P., IMLAY, R. W., & MURPHY, M. the Melanesian Complex and of West Antarc- A., 1960. Correlation of the Cretaceous For- tica - a key to the configuration of Gondwana? mations of the Pacific Coast (United States Zmxq~sq~s53 Zmxq~oxuy53 Zmxq~qo~x536: 37-44. and Northwestern Mexico). Qq~x5 ]~o~ Ky5 VILAS, J F. Y VALENCIO,D. A., 1970 u4 Paleo- M~··qxmáu~z Mnm·á» r~· X~·át Kyq·uom130e., geographic reconstructions of the Gondwanic 50 pp., 1 pl., 5 figs. Continents based on Palaeomagnetic and Sea- RAVN,J P. J., 1911. On jurassic and Cretaceous fIoor Spreading Data. Om·át Zxmzqá5 ]ou5Vqá4 Fossils from North-East Greenland. Wqpp5 áq·»37: 397-405, 3 figs. o4q3 45 (10): 437-500, pls. 32-38. 48CA7 n5The Recurrent Mesozoic Drift of South ROD, E. & MAYNC,W., 1954.Revision of Lower America and Africa. Om·át Zxmzqá3]ou5Vqááq·»3 Cretaceous Stratigraphy of Venezuela. Léxx5 7: 441-444, 1 fig. Ky5 K»»~o5Zqá·5 Mq~x5338, (2): 193-283. WATERHOUSE,J. B., 1959. A New Species of ROSENKRANTZA.,, 1934. Beschreibung der von Wmoo~qxámfrom Raukumara Peninsula, with a R. Bo gvad wahrend der Jahre 1929-30 in Revision of W5 ymszmámMarwick. Xqñ fqm4 Ostgr 2Inland Gesammelten unterkretacischen xmzp g~é·5 Qq~x5 Qq~¡n»3 2: 489-500. Fossilien und die sich aus diesen ergebenden -1965. A Historical Survey of the Pre-Creta- Stratigraphischen schlussfolgerungen. Wqpp5 ceous Geology of New Zealand, Part 1. Xqñ M·7zx53 93 (1): 17-28, pls. 1-5. fqmxmzp g~é·5 Qq~x5 Qq~¡n»53 8 (6): 931- SKWARKO,S. K., 1966. Cretaceous Stratigraphy 998. and Paleontology of the Northern Territory. 48C?Am5 The Continental Displacement of New Léxx5 Lé·5 Wuzq·5 \q»~é·5 Qq~x5 Qq~¡n»5 Zealand. ASyrnposiurn on Continental Drift Ké»á·mx5373: 1-133, pls. 1-15. emphasizing the History of the South Atlan- tic Area. azq»o~ W~záqíupq~ ]y¡~»uéy 657 SPATH, F., 1924. On the ammonites of the Spee- M~záuzqzámxN·uxá5 ton Clay and the subdivisions of the Neoco- mian. Qq~x5 Wms5361: 73-89. 48C?A n5A new Permian fauna from New Ca- ledonia, and its relationships to Gondwana and SPATH, L. F., 1946. Preliminary Notes on the the Tethys. S]uy¡~»u~ Vzáq·z5 O»á·máusZ5m4 Cretaceous Ammonite Faunas of East Green- xq~zá5 Q~zpéumzm3 Léqz~» Ku·q» 4 K·sqzáuzm3 land. Wqpp5 o4q3 132 (4). 1-11. p. 249-266. - 295 -1969. The Paleoclimatic Significance of Per- WHITEHOUSE, F. W., 1926. The Cretaceous mian Praductacea from Queensland. Sz K. S. Arnmonoidea of eastern Australia. Wqy5 [p5 W. CAMPBELL,(ed.) ]á·máus·m¡t mzp Zmxq~z4 Wé»538: 195-242, pls. 34-41. á~x~s O»»m»3 uz R~z~é· 78 N~·~án Ruxx3 1. WIEDMANN, J., 1969. The Heteromorphs and 226-235, figs. 40-43. Aust. Nat. Univ. Press. Ammonoid Extinction. Lu~x5\qí53 44: 563-602, -1970. A new Permian fauna from New Cale- 23 figs., pls, 1-3. donia, and its relationships to Gondwana and WIEDMANN, J. & DIENI, J., 1968. Die Kreide the Tethys. azq»o~ Q~zp5 ]á·má5 SaM] ]y¡»5 Sardiniens und ihre CephaIopoden. Zmxq~zá~s·5 249-272. Vámxuom364 (n. S. 34): 1-171, pls. 1-18. WATERHOUSE,J. B. & VELLA, P., 1965. Permian WILCKENS, O., 1947. Palaontologische und Geo- fossils from Para para Peak, north-west Nelson. logische Ergebnisse der Reise von KohI-Larsen _·mz»5 \~5 ]~o~ Xqñ fqmxmzp Mq~x533 (5): (1928-29) nach Süd-Georgien. Knt5 ]qzow5 57-84, pls. 1-4. Xmá5 Qq»53474: 1-66, 9 pls, WELLMAN, H. W., 1959. Divisions of the New WRIGHT, J. B., 1966. Convection and Continen- Zealand Cretaceous. _·mz»5 \~5 ]~o~ Xqñ tal Drift in the south-west Pacific. _ qoá~z~4 fqmxmzp3 87: 99-163. ¡n»uo»3 3 (2): 69-81.

NOTA DEL EDITOR

En el trabajo de C. S. Churcher, "OBSERVACIONES SOBRE EL STATUS TAXONÓ- MICO DE O¡uqé·oq·~» AMEGHINO 1889 y sus ESPECIES E. á·ézoé» y E. ¡·~óuyé»'3 publicado en el número 1O del tomo IV de AMEGHINIANA, con respecto al pá- rrafo de la página 353 donde dice: ... "Más aún: describió una especie nueva, E. ¡·~óuyé»3 fundada en un buen ejemplar de muda de cornamenta recogida por el señor Edgardo Rohn, en 1944, en la playa de las barrancas derechas del Río Paraná ... ", la Dra. Pierina Pasotti nos envió una nota donde aclara que 'qx quqy4 ¡xm pq E. ¡·~óuyé» réq qónéymp~ ¡~· qx »q~· Opsm·p~ \~nz3 qz 1944, pq xm nm··mzom pq xm ·unq·m pq·qotm pqx ·~ Zm·mz'3

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