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Self-Repair and Self-Cleaning of the Lepidopteran Proboscis
Clemson University TigerPrints All Dissertations Dissertations 8-2019 Self-Repair and Self-Cleaning of the Lepidopteran Proboscis Suellen Floyd Pometto Clemson University, [email protected] Follow this and additional works at: https://tigerprints.clemson.edu/all_dissertations Recommended Citation Pometto, Suellen Floyd, "Self-Repair and Self-Cleaning of the Lepidopteran Proboscis" (2019). All Dissertations. 2452. https://tigerprints.clemson.edu/all_dissertations/2452 This Dissertation is brought to you for free and open access by the Dissertations at TigerPrints. It has been accepted for inclusion in All Dissertations by an authorized administrator of TigerPrints. For more information, please contact [email protected]. SELF-REPAIR AND SELF-CLEANING OF THE LEPIDOPTERAN PROBOSCIS A Dissertation Presented to the Graduate School of Clemson University In Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy ENTOMOLOGY by Suellen Floyd Pometto August 2019 Accepted by: Dr. Peter H. Adler, Major Advisor and Committee Co-Chair Dr. Eric Benson, Committee Co-Chair Dr. Richard Blob Dr. Patrick Gerard i ABSTRACT The proboscis of butterflies and moths is a key innovation contributing to the high diversity of the order Lepidoptera. In addition to taking nectar from angiosperm sources, many species take up fluids from overripe or sound fruit, plant sap, animal dung, and moist soil. The proboscis is assembled after eclosion of the adult from the pupa by linking together two elongate galeae to form one tube with a single food canal. How do lepidopterans maintain the integrity and function of the proboscis while foraging from various substrates? The research questions included whether lepidopteran species are capable of total self- repair, how widespread the capability of self-repair is within the order, and whether the repaired proboscis is functional. -
Biodiverse Master
Montane, Heath and Bog Habitats MONTANE, HEATH AND BOG HABITATS CONTENTS Montane, heath and bog introduction . 66 Opportunities for action in the Cairngorms . 66 The main montane, heath and bog biodiversity issues . 68 Main threats to UK montane, heath and bog Priority species in the Cairngorms . 72 UK Priority species and Locally important species accounts . 73 Cairngorms montane, heath and bog habitat accounts: • Montane . 84 • Upland heath . 87 • Blanket bog . 97 • Raised bog . 99 ‘Key’ Cairngorms montane, heath and bog species . 100 65 The Cairngorms Local Biodiversity Action Plan MONTANE, HEATH AND BOG INTRODUCTION Around one third of the Cairngorms Partnership area is over 600-650m above sea level (above the natural woodland line, although this is variable from place to place.). This comprises the largest and highest area of montane habitat in Britain, much of which is in a relatively pristine condition. It contains the main summits and plateaux with their associated corries, rocky cliffs, crags, boulder fields, scree slopes and the higher parts of some glens and passes. The vegeta- tion is influenced by factors such as exposure, snow cover and soil type. The main zone is considered to be one of the most spectacular mountain areas in Britain and is recognised nationally and internationally for the quality of its geology, geomorphology and topographic features, and associated soils and biodiversity. c14.5% of the Cairngorms Partnership area (75,000ha) is land above 600m asl. Upland heathland is the most extensive habitat type in the Cairngorms Partnership area, covering c41% of the area, frequently in mosaics with blanket bog. -
Lepidoptera in Cheshire in 2002
Lepidoptera in Cheshire in 2002 A Report on the Micro-Moths, Butterflies and Macro-Moths of VC58 S.H. Hind, S. McWilliam, B.T. Shaw, S. Farrell and A. Wander Lancashire & Cheshire Entomological Society November 2003 1 1. Introduction Welcome to the 2002 report on lepidoptera in VC58 (Cheshire). This is the second report to appear in 2003 and follows on from the release of the 2001 version earlier this year. Hopefully we are now on course to return to an annual report, with the 2003 report planned for the middle of next year. Plans for the ‘Atlas of Lepidoptera in VC58’ continue apace. We had hoped to produce a further update to the Atlas but this report is already quite a large document. We will, therefore produce a supplementary report on the Pug Moths recorded in VC58 sometime in early 2004, hopefully in time to be sent out with the next newsletter. As usual, we have produced a combined report covering micro-moths, macro- moths and butterflies, rather than separate reports on all three groups. Doubtless observers will turn first to the group they are most interested in, but please take the time to read the other sections. Hopefully you will find something of interest. Many thanks to all recorders who have already submitted records for 2002. Without your efforts this report would not be possible. Please keep the records coming! This request also most definitely applies to recorders who have not sent in records for 2002 or even earlier. It is never too late to send in historic records as they will all be included within the above-mentioned Atlas when this is produced. -
T1)E Bedford,1)Ire Naturaii,T 45
T1)e Bedford,1)ire NaturaIi,t 45 Journal for the year 1990 Bedfordshire Natural History Society 1991 'ISSN 0951 8959 I BEDFORDSHffiE NATURAL HISTORY SOCIETY 1991 Chairman: Mr D. Anderson, 88 Eastmoor Park, Harpenden, Herts ALS 1BP Honorary Secretary: Mr M.C. Williams, 2 Ive! Close, Barton-le-Clay, Bedford MK4S 4NT Honorary Treasurer: MrJ.D. Burchmore, 91 Sundon Road, Harlington, Dunstable, Beds LUS 6LW Honorary Editor (Bedfordshire Naturalist): Mr C.R. Boon, 7 Duck End Lane, Maulden, Bedford MK4S 2DL Honorary Membership Secretary: Mrs M.]. Sheridan, 28 Chestnut Hill, Linslade, Leighton Buzzard, Beds LU7 7TR Honorary Scientific Committee Secretary: Miss R.A. Brind, 46 Mallard Hill, Bedford MK41 7QS Council (in addition to the above): Dr A. Aldhous MrS. Cham DrP. Hyman DrD. Allen MsJ. Childs Dr P. Madgett MrC. Baker Mr W. Drayton MrP. Soper Honorary Editor (Muntjac): Ms C. Aldridge, 9 Cowper Court, Markyate, Herts AL3 8HR Committees appointed by Council: Finance: Mr]. Burchmore (Sec.), MrD. Anderson, Miss R. Brind, Mrs M. Sheridan, Mr P. Wilkinson, Mr M. Williams. Scientific: Miss R. Brind (Sec.), Mr C. Boon, Dr G. Bellamy, Mr S. Cham, Miss A. Day, DrP. Hyman, MrJ. Knowles, MrD. Kramer, DrB. Nau, MrE. Newman, Mr A. Outen, MrP. Trodd. Development: Mrs A. Adams (Sec.), MrJ. Adams (Chairman), Ms C. Aldridge (Deputy Chairman), Mrs B. Chandler, Mr M. Chandler, Ms]. Childs, Mr A. Dickens, MrsJ. Dickens, Mr P. Soper. Programme: MrJ. Adams, Mr C. Baker, MrD. Green, MrD. Rands, Mrs M. Sheridan. Trustees (appointed under Rule 13): Mr M. Chandler, Mr D. Green, Mrs B. -
Mardon Skipper Site Management Plans
Mardon Skipper (Polites mardon) Site Management Plans Gifford Pinchot National Forest Service Cowlitz Valley Ranger District Prepared by John Jakubowski North Zone Wildlife Biologist Reviewed by Rich Hatfield, The Xerces Society for Invertebrate Conservation October 2, 2015 Cowlitz Valley Ranger District Mardon Skipper Sites Group Meadow Longitude Latitude Area (acres) Elevation (ft.) Midway Midway 121 32.0 46 21.2 8 4,313 Midway PCT 121 31.1 46 21.1 2 4,530 Midway 115 Spur 121 30.9 46 21.0 3 4,494 Midway Grapefern 121 30.9 46 21.5 3 4,722 Midway 7A North 121 31.4 46 21.5 2 4,657 Midway 7A South 121 31.5 46 21.4 2 4,625 Midway 7A 121 31.1 46 21.4 7 4,676 Muddy Muddy 121 32.2 46.18.5 4 4,450 Muddy Lupine 121 31.8 46 18.7 3 4,398 Spring Cr Spring Cr. 121 33.5 46 20.4 unknown 3,900 Goal of the Management Plans Maintain and improve grassland/forb habitat at known occupancy meadow sites to ensure continued occupancy by mardon skipper butterfly as well as other important pollinator species such as western bumble bee. 1 Introduction On the Gifford Pinchot National Forest (GPNF), mardon skippers were first detected on the Mt. Adams Ranger District (MTA) in 2000 and on Cowlitz Valley Ranger District (CVRD) in 2002. Mardon skippers are known to inhabit ten, upland dry grassy meadows on the CVRD. Portions of the meadows are mesic and are unsuitable mardon skipper habitat. -
Garden Moth Scheme Report 2016
Garden Moth Scheme Report 2016 Heather Young – April 2017 1 GMS Report 2016 CONTENTS PAGE Introduction 2 Top 30 species 2016 3 Population trends (?) of our commonest garden moths 5 Autumn Moths 12 Winter GMS 2016-17 14 Antler Moth infestations 16 GMS Annual Conference 2017 19 GMS Sponsors 20 Links & Acknowledgements 21 Cover photograph: Fan-foot (R. Young) Introduction The Garden Moth Scheme (GMS) welcomes participants from all parts of the United Kingdom and Ireland, and in 2016 received 341 completed recording forms, slightly fewer than last year (355). The scheme is divided into 12 regions, monitoring 233 species of moth in every part of the UK and Ireland (the ‘Core Species’), along with a variable number of ‘Regional Species’. For each of the last seven years, we have had records from over 300 sites across the UK and Ireland, and later in the report there are a series of charts representing the population trends (or fluctuations) of our most abundant species over this period. The database has records dating back to 2003 when the scheme began in the West Midlands, and now contains over 1 ¼ million records, providing a very valuable resource to researchers. Scientists and statisticians from Birmingham and Manchester Universities are amongst those interested in using our data, as well as the ongoing research being undertaken by the GMS’s own John Wilson. There is an interesting follow-up article by Evan Lynn on the Quarter 4 GMS newsletter piece by Duncan Brown on Antler Moth infestations, and a report on the very successful 2017 Annual Conference in Apperley Village Hall, near Tewkesbury. -
January Review of Butterfly, Moth and Other Natural History Sightings 2019
Review of butterfly, moth and other natural history sightings 2019 January January started dry and settled but mostly cloudy with high pressure dominant, and it remained generally dry and often mild during the first half of the month. The second half became markedly cooler with overnight frosts and the last week saw a little precipitation, some which was occasionally wintry. With the mild weather continuing from December 2018 there were a small number of migrant moths noted in January, comprising a Dark Sword-grass at Seabrook on the 5th, a Silver Y there on the 13th and 2 Plutella xylostella (Diamond-back Moths) there on the 15th, whilst a very unseasonal Dark Arches at Hythe on the 4th may have been of immigrant origin. Dark Sword-grass at Seabrook (Paul Howe) Dark Arches at Hythe (Ian Roberts) More typical species involved Epiphyas postvittana (Light Brown Apple Moth), Satellite, Mottled Umber, Winter Moth, Chestnut, Spring Usher and Early Moth. Early Moth at Seabrook (Paul Howe) Spring Usher at Seabrook (Paul Howe) The only butterfly noted was a Red Admiral at Nickolls Quarry on the 1st but the mild weather encouraged single Buff-tailed Bumblebees to appear at Seabrook on the 7th and Mill Point on the 8th, whilst a Minotaur Beetle was attracted to light at Seabrook on the 6th. A Common Seal and two Grey Seals were noted regularly off Folkestone, whilst at Hare was seen near Botolph’s Bridge on the 1st and a Mink was noted there on the 17th. February After a cold start to the month it was generally mild from the 5th onwards. -
Two New Records for the Appalachian Grizzled Skipper (Pyrgus Wyandot)
Banisteria, Number 24, 2004 © 2004 by the Virginia Natural History Society Status of the Appalachian Grizzled Skipper (Pyrgus centaureae wyandot) in Virginia Anne C. Chazal, Steven M. Roble, Christopher S. Hobson, and Katharine L. Derge1 Virginia Department of Conservation and Recreation Division of Natural Heritage 217 Governor Street Richmond, Virginia 23219 ABSTRACT The Appalachian grizzled skipper (Pyrgus centaureae wyandot) was documented historically (primarily from shale barren habitats) in 11 counties in Virginia. Between 1992 and 2002, staff of the Virginia Department of Conservation and Recreation, Division of Natural Heritage, conducted 175 surveys for P. c. wyandot at 75 sites in 12 counties. The species was observed at only six sites during these surveys, representing two new county records. All observations since 1992 combined account for <80 individuals. Due to forest succession and threats from gypsy moth control measures, all recent sites for P. c. wyandot in Virginia may be degrading in overall habitat quality. Key words: Lepidoptera, Pyrgus centaureae wyandot, conservation, shale barrens, Virginia. INTRODUCTION wyandot) in Virginia. Parshall (2002) provides a comprehensive review of the nomenclature and The Appalachian grizzled skipper (Pyrgus taxonomy of P. c. wyandot. Most authors classify this centaureae wyandot) has a rather fragmented range, skipper as a subspecies of the Holarctic Pyrgus occurring in northern Michigan as well as portions of centaureae (e.g., Opler & Krizek, 1984; Iftner et al., Ohio, Pennsylvania, Maryland, West Virginia, and 1992; Shuey, 1994; Allen, 1997; Opler, 1998; Virginia; isolated historical records are known from Glassberg, 1999; Parshall, 2002), although some Kentucky, New York, New Jersey, North Carolina, and lepidopterists treat it as a full species (Shapiro, 1974; the District of Columbia (Opler, 1998; NatureServe, Schweitzer, 1989; Gochfeld & Burger, 1997). -
The Consequences of a Management Strategy for the Endangered Karner Blue Butterfly
THE CONSEQUENCES OF A MANAGEMENT STRATEGY FOR THE ENDANGERED KARNER BLUE BUTTERFLY Bradley A. Pickens A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE August 2006 Committee: Karen V. Root, Advisor Helen J. Michaels Juan L. Bouzat © 2006 Bradley A. Pickens All Rights Reserved iii ABSTRACT Karen V. Root, Advisor The effects of management on threatened and endangered species are difficult to discern, and yet, are vitally important for implementing adaptive management. The federally endangered Karner blue butterfly (Karner blue), Lycaeides melissa samuelis inhabits oak savanna or pine barrens, is a specialist on its host-plant, wild blue lupine, Lupinus perennis, and has two broods per year. The Karner blue was reintroduced into the globally rare black oak/lupine savannas of Ohio, USA in 1998. Current management practices involve burning 1/3, mowing 1/3, and leaving 1/3 of the lupine stems unmanaged at each site. Prescribed burning generally kills any Karner blue eggs present, so a trade-off exists between burning to maintain the habitat and Karner blue mortality. The objective of my research was to quantify the effects of this management strategy on the Karner blue. In the first part of my study, I examined several environmental factors, which influenced the nutritional quality (nitrogen and water content) of lupine to the Karner blue. My results showed management did not affect lupine nutrition for either brood. For the second brood, I found that vegetation density best predicted lupine nutritional quality, but canopy cover and aspect had an impact as well. -
Grizzled Skipper
Species: Grizzled Skipper (Pyrgus wyandot) Global Rank: G1G2Q State Rank: S1 State Wildlife Action Plan: Immediate Concern Responsibility Species Climate Change Vulnerability: Highly Vulnerable Confidence: Very High Note: This assessment is expected to be similar for other butterflies of specialized or moderately specialized forest clearing habitats, with specific food plants, habitats exposed to gypsy moth spray; and lacking a fire resistant dormant stage (larval and/or pupal). Some examples: - Frosted Elfin (Callophrys irus); Global Rank G3, State Rank S2; Caterpillar hostplant Wild Indigo (Baptisia tinctoria); Habitat typically grassy (Andropogon spp.) openings in oak habitats on sandy rocky soils; sometimes found in disturbed areas with hostplant such as powerline right-of-ways. - Persius Duskywing (Erynnis persius); Global Rank G5T1T3, State Rank S1; Caterpillar hostplant Wild Indigo (Baptisia tinctoria); Habitats include pitch pine- scrub oak barrens, scrubby ridgetops, or powerline right-of-ways within such settings with sandy-gravelly soils. - Northern Metalmark (Calephelis borealis), Global Rank G3G4, State Rank S2; Caterpillar hostplant Round-leaved Ragwort (Senecio obovatus); Habitats are openings within forested or wooded areas such as natural outcrops, shale or limestone barrens, glades or powerline right-of-ways. Habitat (adapted from NatureServe 2008 and Schweitzer 1989): The Grizzled Skipper butterfly is an Appalachian Mountain habitat specialist that requires shale barren habitats with abundant exposed crumbly rock or soil. Shale barrens are semi-open shale slopes with sparse herbaceous vegetation and tend to be surrounded by scrubby oak or oak-hickory woodlands, often with a component of Virginia Pine (Pinus virginiana). A natural area which meets the habitat requirements should maintain itself as a shale barren. -
E-News Spring 2019
Spring e-newsletter March 2019 Welcome to Spring! Green Hairstreak - Iain Leach Orange-tip - Iain Leach Garden Tiger caterpillar - Roy Leverton INSIDE THIS ISSUE: Contributions to our newsletters Dates for your Diary……………………….2 The Wish List…………………….…13-14 are always welcome. Borders News...........................................3 Our Conservation Strategy…….….15-20 Please use the contact details Munching Caterpillars Scotland.…………4 Carrion Beetles…………….……….21-22 below to get in touch! Peatlands for People………..…………….5 Moth Equipment - for sale...............23 If you do not wish to receive our Recording butterflies using Apps………...6 SW Branch Events 2019……….....24-25 newsletter in the future, simply Adopt a Transect………………………......7 Highland Branch Events 2019..….26-29 reply to this message with the Rare migrant on Islay!..............................8 East Branch Events 2019..….…...30-34 word ’unsubscribe’ in the title - Coul Links Update……………………….9-10 thank you. Northern Brown Argus, Kincraig………11-12 Contact Details: Butterfly Conservation Scotland t: 01786 447753 Balallan House e: [email protected] Allan Park w: www.butterfly-conservation.org/scotland Stirling FK8 2QG Dates for your Diary Wildlife Recorders’ Gathering - Saturday 30th March 10.30 - 4.30pm - Dumfries An informal day of talks, presentations, networking and displays covering the wonderful wildlife of SW Scotland. Contact SWSEIC at [email protected] for more details. Highland Branch AGM - Saturday, 13th April 2019 Our Highlands & Island Branch will be holding their AGM on Saturday, 13th April at the Kingsview Christian Centre, Balnafettack Road, Inverness, IV3 8TF. See Highland Branch Events (Page 27) for more info. South & West Branch Members’ Day/AGM - Saturday, 27th April 2019 Our Glasgow & Southwest Branch will be holding their Members’ Day/AGM on Saturday, 27th April at Chatelherault Country Park. -
Scottish Macro-Moth List, 2015
Notes on the Scottish Macro-moth List, 2015 This list aims to include every species of macro-moth reliably recorded in Scotland, with an assessment of its Scottish status, as guidance for observers contributing to the National Moth Recording Scheme (NMRS). It updates and amends the previous lists of 2009, 2011, 2012 & 2014. The requirement for inclusion on this checklist is a minimum of one record that is beyond reasonable doubt. Plausible but unproven species are relegated to an appendix, awaiting confirmation or further records. Unlikely species and known errors are omitted altogether, even if published records exist. Note that inclusion in the Scottish Invertebrate Records Index (SIRI) does not imply credibility. At one time or another, virtually every macro-moth on the British list has been reported from Scotland. Many of these claims are almost certainly misidentifications or other errors, including name confusion. However, because the County Moth Recorder (CMR) has the final say, dubious Scottish records for some unlikely species appear in the NMRS dataset. A modern complication involves the unwitting transportation of moths inside the traps of visiting lepidopterists. Then on the first night of their stay they record a species never seen before or afterwards by the local observers. Various such instances are known or suspected, including three for my own vice-county of Banffshire. Surprising species found in visitors’ traps the first time they are used here should always be regarded with caution. Clerical slips – the wrong scientific name scribbled in a notebook – have long caused confusion. An even greater modern problem involves errors when computerising the data.