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Entry for ramosissimum subsp ramosissimum Desf. [family ]

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Page 1 of 3 Entry for subsp ramosissimum Desf. [family EQUISETACEAE]

Herbarium South African National Biodiversity Institute, Compton Herbarium, Cape Town (SAM) Collection Swaziland and allies Resource Type Reference Sources Entry from Swaziland Ferns and Fern Allies, Vol (2003) Author: J.P. Roux Names Equisetum ramosissimum subsp ramosissimum Desf. [family EQUISETACEAE], Other names ramosissimum = many-branched; Vernacular names: Scouring rush, Horsetail; Bewerasiegras, Dronkgras, Drilgras, Lidjiesgras, Perdestert, Paardestaart (Afr.); Mohlaka-photoane (Ses.); Ishobalehashi, Isikhumukele (Zulu) Information Plants terrestrial. Rhizome subterranean, jointed, 3-6 mm in diameter, roots borne in whorls at the nodes. Stems erect, to 1.5 m tall, to 5 mm in diameter, branches borne in whorls at the nodes, up to 14 in each whorl, to 130 mm long, to 1.5 mm in diameter, main stem and branches ridged, bearing silicaceous tubercules or bands, with regular rows of paracytic sunken stomata in the sulci, in cross section with a prominent central canal and small vallecular (under sinuses) and carinal (under ridges) canals. Leaves membranous to thinly chartaceus, each with a single central vein, whorled, fused into a nodal sheath, to 11 mm long, ending in acuminate teeth, to 2 mm long, turning black with age, margins hyaline, estomate. Sporangia elongate, thin-walled, dehiscing longitudinally, borne on stalked, peltate sporangiophores forming a strobilus terminally on a main stem or branch, to 18 mm long, to 5 mm in diameter, each sporangiophore bears 6-9 sporangia. Spores spheroidal, chlorophyllose, with 4 spathulate hygroscopic elaters. Figure 4C & D. Range Distribution: The is widespread in Swaziland, occurring along most of the major draining systems at altitudes ranging from ± 150 m along the Mlawula River to 800 m on the western highveld. The species is widespread in the eastern parts of westcentral tropical Africa, east and south tropical Africa, south

Page 2 of 3 tropical and southern Africa, Macaronesia, the western Indian Ocean region, Egypt, southern and central Europe and Asia, except Malaysia. Equisetum ramosissimum Desf. subsp. debile (Roxb. ex Vaucher) Hauke occurs from India and southern China through Malaysia to New Caledonia and Fiji. Habitat Ecology: Terrestrial, generally in humid sand, gravel, mud or silt on floodplains and in stream and riverbeds. Often also invasive in cultivated fields. Exposed or in partial shade. Geophyte, mesoxerophytic; aerial stems meso-xeromorphic. Seasonal pattern apparently nonexistent, strobili possible throughout the year. Vegetative reproduction by the subterranean branched rhizome. Pyrophytic. Use Uses: Equisetum is of economic importance to the stock farmer as it is poisonous when consumed in large quantities, the toxic principle probably being the enzyme thiaminase that destroys vitamin B1. This causes livestock to appear drunk and they may harm themselves. The Sesotho use the for various ailments and rituals. Phillips (1917) reports that the rhizomes are cooked and a decoction is drunk by barren woman so that they may become pregnant. The plant is used as a charm to drive caterpillars from the maize fields. This is done by encapsulating some of the caterpillars in clay balls which are then burnt in the fields. The Zulu use the sap from the plant to relieve toothache and is also applied to the wound after tooth extraction (Gerstner 1939).

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