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Entomological Society of Canada I-HE INSECTS AND ARACHNIDS OF CANADA THE INSECTS AND ARACHNIDS OFCANADA PART 21 The Weevils of Canada and Alaska: Volume 1 Coleoptera: Curculionoidea, excluding Scolytidae and Curculionidae DonaldE. Bright Centre for Land and Biological Resources Research Ottawa, Ontario ResearchBranch Agriculture Canada PublicationlSS2 1993 oMinister of Supply and Services Canada 1993 Available in Canada through Associated Bookstores and other booksellers or by mail from Canada Communication Group Publishing Ottawa, Canada KlA 0S9 - Cat. No. A42-421I99I-2|E ISBN 0-660-14433-6 Canadian Cataloguing in Publication Data Bright, Donald E. The weevils of Canada and Alaska. Vol. 1, Coleoptera:Curculionoidea except Scolytidae and Curculionidae (The insects and arachnids ofCanada. 0706-7313 : pt. 21) (Publication ; 1882) To be complete in 3 v. Includes bibliographical references and index. Cat. no. 1'42-421 l99l-2lB ISBN 0-660-14433-6 1. Beetles-Canada. 2. Beetles-A]aska. I. Canada. Agriculture Canada. Research Branch. IL Title. III. Series. VI. Series: Publication (Canada. Aqriculture Canada). English ; 1882. QL596.C9.B7 1993 595.7',68'0971 C92-099101-7 Staff editor: Frances Smith The Insects and Arachnids of Canada Part 1. Collecting, Preparing, and Preserving Insects, Mites, and Spiders, compiled by J.E.H. Martin, Biosystematics Research Institute, Ottawa, 1978.182 pp. Cat. No. A42-4211977-I. Partie 1. R6colte, pr6paration et conservation des Insectes, des Acariens et des Araign6es, compil6 par J.E.H. Martin, Institut de recherche biosyst6matique, Ottawa, 1983. 205 pp. No de cat. 442-4211977-rF. Part 2. The Bark Beetles of Canada and Alaska (Coleoptera: Scolytidae), by D.E. Bright, Jr., Biosystematics Research Institute, Ottawa, 1976. 247pp. Cat. No. A42-421L976-2. [Out of print.l Part 3. The Aradidae of Canada (Hemiptera: Aradidae), by R. Matsuda, Biosystematics Research Institute, Ottawa, 1977. 176 pp. Cat. No. 442-421L977-3. Part 4. The Anthocoridae of Canada and Alaska (Heteroptera: Anthocoridae), by L.A. Kelton, Biosystematics Research Institute, Ottawa, L977. LU pp. Cat. No. A42-4211977-4. Part 5. The Crab Spiders of Canada and Alaska (Araneae: Philodromidae and Thomisidae), by C.D. Dondale and J.H. Redner, Biosystematics Research Institute, Ottawa, L978.255 pp. Cat. No. A42-4211978-5. Part 6. The Mosquitoes of Canada (Diptera: Culicidae), by D.M. Wood, PT. Dang, and R.A. Ellis, Biosystematics Research Institute, Ottawa, 1979. 390 pp. Cat. No. A42-421t979-6. [Out of print.l Partie 7. Genera des Trichoptdres du Canada et des Etats adjacents, par E Schmid, Institut de recherches biosyst6matiques, Ottawa, l-980. 296 pp. No de cat. A42-4211980-7F (in French only). [Out of print.l Part 8. The Plant Bugs of the Prairie Provinces of Canada (Heteroptera: Miridae), by L.A. Kelton, Biosystematics Research Institute, Ottawa, 1980. 408 pp. Cat. No.442-4211980-8. Part 9. The Sac Spiders of Canada and Alaska (Araneae: Clubionidae and Anyphaenidae), by C.D. Dondale and J.H. Redner, Biosystematics Research Institute, Ottawa, 1982. L94 pp. Cat. No. A42-4211982-9E. Part 10. The Spittlebugs of Canada (Flomoptera: Cercopidae), by K. G.A. Hamilton, Biosystematics Research Institute, Ottawa, 1982. 102 pp. Cat. No. A42-4211982-10E. Part 11. The Genera of Larval Midges of Canada (Diptera: Chironomidae), by D.R. Oliver and M.E. Roussel, Biosystematics Research Institute, Ottawa, 1983. 263 pp. Cat. No. A42-4211983-11E. Pafi, 12. The Families and Subfamilies of Canadian Chalcidoid Wasps (Hymenoptera: Chalcidoidea), by C.M. Yoshimoto, Biosystematics Research Institute, Ottawa, L984.149 pp. Cat. No. A42-4211983-12E. Part 13. The Carrion Beetles of Canada and Alaska (Coleoptera: Silphidae and Agyrtidae), by R.S. Anderson and S.B. Peck, Biosystematics Research Institute, Ottawa, 1985. 121- pp. Cat. No. A42-4211985-13E. Part 14. The Grasshoppers, Crickets, and Related Insects of Canada and Adjacent Regions (lllonata: Dermaptera, Cheleutoptera, Notoptera, Dictuoptera, Grylloptera, and Orthoptera), by V.R. Vickery and D.KMcE. Kevan, Biosystematics Research Institute, Ottawa, 1986. 918 pp. Cat. N o. 4.42-4211985-148. [Out of print.l Part .l-5. The Metallic Wood-boring Beetles of Canada and Alaska (Coleoptera: Buprestidae), by D.E. Bright, Biosystematics Research Centre, Ottawa, 1987. 335 pp. Cat. No. A42-4211986-158. Part 16. The Horse Flies and Deer Flies of Canada and Alaska (Diptera: TabarLidae), by H.J. Teskey, Biosystematics Research Centre, Ottawa, 1990. 381pp. Cat. No. A42-4211990-16E. Part |.7. The Wolf Spiders, Nurseryweb Spiders, and Lynx Spiders of Canacla and Alaska (Araneae: Lycosidae, Pisauridae, and Oxyopidae), by C.D. Dondale and J.H. Redner, Biosystematics Research Centre, Ottau'a, 1990. 383 pp. Cat. No. A42-4211990-17E. Part .[8. The Flower Flies of the Subfamily Syrphinae of Canada, Alaska, and Greenland (Diptera: Syrphidae), by J.R. Vockeroth, Centre for Land and Biological Resources Research, Ottawa, t992.456 pp. Cat. No. 442-4211991-18E. Part .l-9. The Ground Spiders of Canada and Alaska (Araneae: Gnaphosidae), by N.I. Platnick and C.D. Dondale, Centre for Land and BioloE;ical Resources Research, Ottawa, 1992. 297 pp. Cat. No. p'42-4217991-19E. Part 20. The Genera and Subgenera of the Sawflies of Canada and Alaska (Hymenoptera: Symphyta), by H. Goulet, Centre for Land and Biological Resources Research, Ottawa, 1992. 235 pp. Cat. No. 442-42ft992-208. Contents Abstract Acknowledgments . 8 Introduction ......::: ::: :::::::::::::... 8 Anatomy 10 Methods t2 Classification.... 12 Key to families of Curculionoidea in Canada and Alaska 13 Cl6 des familles des Curculionoidea du Canada et de I'Alaska . 15 Family Anthribidae t7 Key to genera of Anthribidae in Canada and Alaska . t7 Cl6 des genres des Anthribidae du Canada et de l'Alaska . 18 Genus Araecerus Schoenherr 19 Genus Euxenus LeConte 2L Genus Euparius Schoenherr 22 Genus Tropideres Schoenherr 25 Key to species of Tropideres in Canada 26 Cl6 des espdces du genre Tropideres du Canada 26 Genus Allandrus LeConte 33 Key to species of Allandrus in Canada and Alaska 34 Cl6 des espdces du genre Allandrus du Canada et de I'Alaska 34 Genus Trigonorhinzs Wollaston . 37 Key to species of Trigonorhinus in Canada 38 Cl6 des espdces du genre Trigonorhiruzs du Canada . 39 Genus Ormiscus Waterhouse 47 Key to species of Ormiscus in Canada 48 Cl6 des espbces du genre Ormiscus du Canada 49 Family Nemonychidae . 51 Key to genera of Nemonychidae in Canada 52 CI6 des genres des Nemonychidae du Canada 53 Genus Lecontellus Kuschel 53 Genus Cim.beris des Gozis 55 Key to species of Cimberis in Canada 56 Cl6 des espdces du genre Cimberis du Canada 56 Genus Acromacer Kuschel 64 Genus Pityornacer Kuschel 65 Family Brentidae 66 Genus Arrheno de s Schoenherr 67 Family Apionidae 70 Key to genera of Apionidae in Canada and Alaska 70 Cl6 des genres des Apionidae du Canada et de l'Alaska . 7l Genus Nanophyes Schoenherr 7t Genusdpioz Herbst 74 Key to subgenera and species of Apion in Canada and Alaska /D Cl6 des sous-genres et des espdces des Apion du Canada etdel'Alaska.... 82 Genus Podapion Riley . t40 Family Rhynchitidae . L42 Key to genera of Rhynchitidae in Canada L43 Cl6 des genres des Rhynchitidae du Canada t44 Genus Pterocolus S.y . t45 Genus Eugnamptus Schoenherr . 148 Genus Pselaphorhynchites Schilsky L52 Key to species of Pselaphorhynchites in Canada r52 Cl6 des espdces du genre Pselnpharhyrrch#es du Canada ' ' . 153 Genus Merhynchites Sharp 155 Key to species of Merhynchites in Canada 156 Cl6 des espdces du genre Merhynchites du Canada . ' . t57 Genus Auletobius Desbrochers . 162 Key to species of Auletobius in Canada t62 Cl6 des espdces du genre Azletobius du Canada 163 Genus Deporaus Samouelle L67 G enus H ap lor hynclzif es Voss 168 Family Ithyceridae t7t Genus lthycerus Schoenherr t7r Family Attelabidae 175 Key to genera of Attelabidae in Canada . 175 Cl6 des genres des Attelabidae du Canada 175 Genus Hinratolabus Jekel r76 Genus Homoeola.bus Jekel r78 Genus Attelabus Linnaeus . 180 Family Platypodidae ... 184 Genus Platypus Herbst 184 Glossarv 198 References .....'.201 Index ......212 Abstract This monograph includes all species in the families Anthribidae (18), Apionidae (47), Attelabidae (3), Brentidae (L), Ithyceridae (1), Nemonychidae (B), Platypodidae (1), and Rhynchitidae (11) that occur or may occur in Canada and Alaska. The work includes a key to all the families of Curculionoidea in North America, as well as keys to all genera and species of the families included. Each genus is described. Species entries include a list of synonyms, a description, data on distribution, information on host plants and on other aspects of biolory (if known), and diagnostic features. Distribution maps and illustrations of taxonomic characters for a number of species are included, as well as habitus illustrations of representative species. Acknowledgments Several individuals contributed to the preparation of this volume. To all those listed below I express my thanks. The excellent habitus illustrations were prepared by Mr. Claude Paquette and the line drawings were completed by Mr. Go Sato, both staff artists, Centre for Land and Biological Resources Research. Jason R. Bright helped to complete the distribution maps. Drs. E. C. Becker, Y. Bousquet, PT. Dang, and A. Smetana, Centre for Land and Biological Resources Research, reviewed an early draft of the manuscript. The final manuscript was reviewed by Drs. R.A. Anderson and H.R. Burke, both at the Department of Entomolory, Texas A & M University, College Station, Tex., and by Dr. WE. Clark, Department of Entomolory, Auburn University, Auburn, Ala. Mr. Robert Skidmore assisted in many ways and his help is gratefully acknowledged. The manuscript was greatly improved by their comments. Ms. Frances Smith, Research Program Service, Agriculture Canada, edited and prepared this work for publication. Her skillful editing and her patient, capable, and amiable way of dealing with an obstinate author are much appreciated. During the course of my studies on Canadian weevils, I examined numerous specimens from various regional laboratories of the Canadian Forestry Service and Agriculture Canada.
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