Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

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Journal of Asia-Pacific Biodiversity

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Original article Annotated catalog of the series Spheciformes (Hymenoptera: Apoidea) from the Korean Peninsula

Jeong-Kyu Kim*

Korean Bioresources Research Center, Seosuwon-ro 607-303, Gwonseon-gu, Suwon, Gyeonggi-do 441-746, Republic of Korea article info abstract

Article history: For future researchers, the Korean species of the series Spheciformes are catalogued. A total of 216 Received 30 August 2014 species in seven families are listed. Of these, species filtered according to four topicsdtaxonomically Received in revised form conflicting, highly questionable in Korean occurrence, not based on actual Korean material, and known 4 October 2014 only from North Koreadare listed and discussed. Accepted 7 October 2014 Copyright Ó 2014, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). Available online 15 October 2014 Production and hosting by Elsevier. All rights reserved.

Keywords: Apoidea Catalog Korean Peninsula Spheciformes

Introduction survey is provided for future researchers. Finally, species filtered by four topicsdtaxonomically conflicting, highly questionable A catalog is essentially an index of published taxa arranged so as in Korean occurrence, not based on actual Korean material, and to provide a complete series of references for both zoological and known only from North Koreadare separately listed and discussed. nomenclatural purposes (Mayr and Ashlock, 1991). Thus, an elab- orated catalog is a valuable guide for future taxonomic research. Materials and methods The series Spheciformes forms a large part of the aculeate Hy- menoptera. However, recognition of Korean Spheciformes fauna is The author follows the higher classification system and species fragmentary and out of date. Kim (1970) provided a catalog of this names provided by Pulawski (2014). For each species and subspe- taxon based on his pilot study (Kim, 1963). Tsuneki (1974, 1982) cies, reference to the original publication of the valid name is examined North Korean specimens, put together previously provided, together with type information. The remainder of the known species, and listed 163 species/subspecies within the family synonym list includes only selected references indicating new Sphecidae that are known from the Korean Peninsula. Later, Paik taxonomic changes (new status and new combination, etc.) and (1985), Kim et al. (1994), and Lee et al. (2010) provided checklists justifiable Korean occurrences. that were probably based on simple accumulation of previous re- Acronyms for type depositories are as follows: [BMNH] British fi cords without veri cation of listed species. In terms of the fact that Museum of Natural History, London, Britain; [DEI] Deutsches a list of names deserves the name checklist only if a careful Entomologisches Institute, Eberswalde, Germany; [ETHZ] Eidge- distinction is made within it between valid names and synonyms nössische Technische Hochschule, Zürich, Switzerland; [HMS] (Mayr and Ashlock, 1991), these three Korean checklists are nothing Heude Museum, Shanghai, China; [HUM] The Hokkaido University but uncritical prerevisionary listings of nominal species. Museum, Sapporo, Hokkaido, Japan; [IRSN] Institut Royal des Sci- In the present study, an annotated catalog of the series Sphe- ence Naturelles, Brussels, Belgium; [KUF] Kyushu University, ciformes from the Korean Peninsula through careful literature Fukuoka, Japan; [KUK] Kyoto University, Kyoto, Japan; [LS] Linnean Society, London, Britain; [MNHAH] The Museum of Nature and Human Activities, Hyogo, Japan; [MNHN] Muséum National d’His- * þ Corresponding author. Tel.: 82 (0)10 8794 3670. ’ E-mail address: [email protected]. toire Naturelle, Paris, France; [MHNG] Musée d Histoire Naturelle, Peer review under responsibility of National Science Museum of Korea (NSMK) and Genève, Switzerland; [MSNV] Museo Civico di Storia Naturale, Korea National Arboretum (KNA). Venice, Italy; [NMK] National Museum, Kraków, Poland; [NMM] http://dx.doi.org/10.1016/j.japb.2014.10.003 2287-884X/Copyright Ó 2014, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). Production and hosting by Elsevier. All rights reserved. 416 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

The National Museum, Melbourne, Australia; [NHMW] Natur- Pingtung Prefecture: Kuraru (now Kenting Park) [KUF]; Kim, historisches Museum, Wien, Austria; [NHMU] Natural History 1970: 572 (diagnosis, no Korean material), 809 (in the list of Museum, Uppsala University, Sweden; [NRS] Naturalhistrriska Hymenoptera from Korea); Paik, 1985: 196 (in the list of Sphe- Riksmuseet, Stockholm, Sweden; [OMNH] the collection of Osaka cidae of Korea); Kim et al., 1994: 260 (in Check List of Insects City Museum of Natural History, Osaka, Japan; [SYKU] Seiki Yamane from Korea); Lee et al., 2010: 208. Collection, Kagoshima University, Kagoshima, Japan; [TMB] Ter- Distribution. ?Korea, Taiwan. mészettodomanyi Múzeum, Budapest, Hungary; [USNM] United Note. Kim (1970) may have recorded the occurrence of this States National Museum, Washington, USA; [UZIL] Universitets species in Korea for the first time. His record, however, was not Zoologiska Institut, Lund, Sweden; [YKU] Yasumatsu collection, based on examination of the Korean specimen. This species is Kyushu University, Fukuoka, Japan; [ZMHU] Museum für Natur- known to be endemic to Taiwan. It seems to be more reasonable to kunde, Humboldt University, Berlin, Germany; [ZMUC] Zoological omit this species from the Korean fauna. Museum, University of Copenhagen, Copenhagen, Denmark. Abbreviations for Korean provincial names from north to south: Ampulex satoi Yasumatsu, 1936 사토오는쟁이벌 [RG] Ryanggang-do; [HB] Hamgyeongbuk-do; [PN] Pyeongannam- Ampulex satoi Yasumatsu, 1936: 204, _ (holotype), \, Korea: “Sui- do; [SL] Seoul; [GG] Gyeonggi-do; [GW] Gangwon-do; [IC] Incheon gen” (Suweon) [depository uncertain]; Bohart and Menke, 1976: Metropolitan City; [CB] Chungcheongbuk-do; [CN] Chung 78 (including Korea in distribution); Tsuneki, 1982: 14 (in the list cheongnam-do; [GB] Gyeongsangbuk-do; [GN] Gyeongsangnam- of the species of the Sphecidae known from the Korean Penin- do; [JB] Jeollabuk-do; [JN] Jeollanam-do; [JJ] Jejudo. sula); Kim, 1970: 573 (diagnosis, no examination of Korean material), 809 (in the list of Hymenoptera from Korea); Paik, Annotated Catalog of the Series Spheciformes (Hymenoptera: 1985: 196 (in the list of Sphecidae of Korea); Kim et al., 1994: Apoidea) From the Korean Peninsula 260 (in Check List of Insects from Korea); Lee et al., 2010: 208. Distribution. S Korea (GG). Order Hymenoptera 벌목 구멍벌과 Superfamily Apoidea Latreille, 1802 꿀벌상과 Family Sphecidae Latreille, 1802 나나니아과 Series Spheciformes (Fossorial wasps; Sphecidae sensu lato) 구멍 Subfamily Ammophilinae André, 1886 벌류 Genus Ammophila Kirby, 1798 나나니속 Ammophila Kirby, 1798: 199. Type species: Sphex sabulosus Lin- naeus, 1758, designated by International Commission of 는쟁이벌과 Family Ampulicidae Shuckard, 1840 Zoological Nomenclature (ICZN), 1946: 571 (Opinion 180). Subfamily Ampulicinae Shuckard, 1840 는쟁이벌아과 는쟁이벌족 Tribe Ampulicini Shuckard, 1840 Ammophila campestris Latreille, 1809 애기나나니 는쟁이벌속 Genus Ampulex Jurine, 1807 Ammophila campestris Latreille, 1809: 54, sex and locality not Ampulex Jurine, 1807: 132. Type species: Chlorion compressum of indicated (holotype or syntype) [depository uncertain]; ¼ Latreille and of Fabricius [ Chlorion compressum (Fabricius, Yasumatsu, 1930:30e32 (diagnosis, N Korea: Hamgyeongbuk- ¼ 1804) Sphex compressus Fabricius, 1781, designated by do: Hoiryeng); Kim, 1963: 118 (in Catalog of Hymenoptera Audouin, 1822: 301]. from Korea); Kim et al., 1975: 48 (an open circle in map desig- nating previous record of Uchida (1930) from North Korea is 는쟁이벌 Ampulex dissector (Thunberg, 1822) A. campestris, the other closed circles are Podalonia affinis); Kim, Ichneumon dissector Thunberg, 1822: 272, sex not indicated (holo- 1980: 133 (as in Kim et al., 1975); Paik, 1985: 197 (in the list of fi type or syntypes), Japan: no speci c locality [NHMU]. Sphecidae of Korea); Kim et al., 1994: 262 (specific name Ampulex dissector (Thunberg): Roman, 1912: 251 (new combina- incorrectly campoctris, in Check List of Insects from Korea); Lee tion); Bohart and Menke, 1976: 77 (including Korea in distri- et al., 2010: 209 (in Checklist of Korean Insects); Nemkov, bution); Tsuneki, 1982: 14 (in the list of the species of the 2012b: 434 (including Korea in distribution). Sphecidae known from the Korean Peninsula); Paik, 1985:196 Distribution. Transpalearctic: N Korea (HB), Europe to China, (in the list of Sphecidae of Korea); Yamane in Yamane et al., Russian Far East. 1999: 471 (including Korea in distribution); Kim et al., 1994: 260 (in Check List of Insects from Korea); Terayama and Tano, Ammophila clavus japonica Kohl, 1906 별나나니, 홍다리나나니 2006: 12, 17 (including Korea in distribution); Yamane, 2008: Ammophila japonica Kohl, 1906b: 328, \ (lectotype, designated by 554 (including Korea in distribution); Lee et al., 2010: 208; Suda, Menke in Bohart and Menke, 1976: 151), Japan: Kofou [MNHN], 2013: 7 (including Korea in biogeographic comment). synonymized with A. clavus by Dollfuss, 2013a:419;Kim, 1970: Ampulex amoena Stål, 1857: 63, sex not indicated (holotype or 604 (diagnosis of \, no Korean material), 814 (in the list of Hy- fi syntypes), China: no speci c locality [NRS], synonymized with menoptera from Korea); Kim et al., 1994: 262 (in Check List of Ampulex dissector by Kohl, 1893: 489; Yasumatsu, 1936: 167, Insects from Korea). e 170 175 (including Korea in distribution with the personal Ammophila clavus Fabricius: Tsuneki, 1962: 25 (in key, including communication of H Doi); Yasumatsu, 1965: 297 (including Korea in distribution); Kim, 1963: 118 (specific name incorrectly Korea in distribution); Kim, 1970: pl. 91, Figure 665, 572 (diag- as clauus, in Catalog of Hymenoptera from Korea); Yasumatsu, nosis, no Korean material). 1965: Pl. 150, 299 (including Korea in distribution); Kim, 1970: fi Distribution. Korea (no speci c locality), China, Japan; India, S Pl. 60 Figure 717, 604 (diagnosis of _, Korea: Mt. Gamaksan), 814 Asia, Taiwan. (in the list of Hymenoptera from Korea); Kim, 1980: Pl. xix HYSP Note. Just based on his personal communication with H Doi, 03, 134 (diagnosis, Korea: Gyeonggi-do, Chungcheongbuk-do, Yasumatsu (1936) recorded the Korean occurrence. To date, any Chungcheongnam-do, Gyeongsangbuk-do, Gyeongsangnam-do, Korean material of this species has not been examined. Jeollanam-do); Yamane, 2008: Pl. 167, 557 (including Korea in distribution). 왜는쟁이벌 Ampulex kurarensis Yasumatsu, 1936 Ammophila atripes japonica Kohl: Bohart and Menke, 1976:151 _ \ Ampulex kurarensis Yasumatsu, 1936: 208, (holotype), , Taiwan: (new status, in Catalog of World Sphecidae); Tsuneki, 1982:14 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 417

(in the list of the species of the Sphecidae known from the Seoul, Gangwon-do, Gyeonggi-do, Chungcheongbuk-do, Korean Peninsula); Paik, 1985: 197 (in the list of Sphecidae of Chungcheongnam-do, Gyeongsangbuk-do, Jeollanam-do, Jeju Korea); Tsuneki, 1991: 199 (in Sphecidae from Korea); Kazenas Is.); Nemkov, 2009: 51 (including Korea in distribution); Lee in Nemkov et al., 1995: 386 (in key, including Korea in distri- et al., 2010: 209 (in Checklist of Korean Insects); Nemkov, bution); Yeo et al., 1998: 21 (in key), 21e22 (S Korea: Seoul, 2012b: 434 (including Korea in distribution); Dollfuss, 2013a: Gyeonggi-do, Chungcheongbuk-do, Chungcheongnam-do, 389 (in key), 394 (in key), 400 (in key), 487e488 (including Gyeongsangbuk-do, Gyengsangnam-do); Terayama and Tano, Korea in distribution). 2006: 14 and 18 (including Korea in distribution); Lee et al., Distribution. Korean Peninsula (PN, SL, GG, GW, CB, CN, GB, JN, JJ), 2010: 209 (in Checklist of Korean Insects). Mongolia, China, Russian Far East, Taiwan. Ammophila clavus japonica Kohl: Tsuneki, 1967a: 9 and 11 (in keys), 15 (new status); Tsuneki, 1974: 359 (North Korea: “Prov. South Ammophila vagabunda Smith, 1856 꼬마나나니 Pyongan: Chang-lyongsan”). Ammophila vagabunda Smith, 1856:218,\ (syntype), China: “Tei- Distribution. Korean Peninsula, NE China (PN, SL, GG, CB, CN, GB, tung: Foo-chou-foo”; Sumatra: no specific locality; Indonesia: GN), Japan. no specific locality; northern India: no specific locality [BMNH]; Dollfuss, 2013a: 389 (in key), 390 (in key), 400 (in key), 500-501 Ammophila infesta Smith, 1873 나나니 (including Korea in distribution); Dollfuss, 2013b: 586 (N Korea: Ammophila infesta Smith, 1873: 190, \ and _ (syntypes), Japan: “Kesong Province: Pagyon water-falls”). Hyogo [BMNH]; Yasumatsu, 1934: 63 (including Korea in dis- Ammophila solowiyofkae Matsumura, 1911:111,\ (syntypes), Russia tribution); Kim, 1963:117e118 (in Catalog of Hymenoptera from now Dolinsk: Solovievka, Galkinowraskoye, Sakhalin [HUM], Korea); Tsuneki, 1974: 360 (N Korea: Kaesong: Mts. Pakyon: synonymized with A. vagabunda by Dollfuss, 2013a:500;Bohart Pakyon popo); Kim, 1980: Pl. xix HYSP 04, 135 (diagnosis, S and Menke, 1976: 153 (possibly senior synonym of Ammophila Korea: Gangwon-do, Gyeonggi-do, Cheongcheongbuk-do, sabulosa nipponica). Chungcheongnam-do, Gyeongsangbuk-do, Gyeongsangnam- Ammophila sabulosa solowiyofkae Matsumura: Nemkov, 2005:141 do, Jeollabuk-do, Jeollanam-do, Jeju Is.); Tsuneki, 1982:1e2(N (including Korea in distribution); Nemkov, 2009:50e51 Korea: “Prov. Gang-won district: On-dzong: Kum-gang san: (including Korea in distribution); Nemkov, 2012b: 434 Mandzang-tae”); Paik, 1985: 197 (in the list of Sphecidae of (including Korea in distribution). Korea); Tsuneki, 1991: 199 (in Sphecidae from Korea); Kazenas Ammophila sabulosa (Linnaeus): Okamoto, 1924: 203 (Korea: Jeju in Nemkov et al., 1995: 387 (in key, including Korea in distri- Is.); Uchida, 1925: 332 (Korea: Suwon), 369 (including Korea in bution); Yeo et al., 1998: 21 (in key), 22e23 (S Korea: Seoul, distribution); Kim, 1963: 118 (in Catalog of Hymenoptera from Gangwon-do, Gyeonggi-do, Jollanam-do); Yamane in Yamane Korea); Yasumatsu, 1965: 299 (including Korea in distribution); et al., 1999:479e480 (including Korea in distribution); Kim, 1970: 604-605 (diagnosis, no Korean material), 814 (in the Terayama and Tano, 2006: 15, 18 (including Korea in distribu- list of Hymenoptera from Korea); Paik, 1985: 197 (in the list of tion); Yamane, 2008: Pl. 167, 557 (including Korea in distribu- Sphecidae of Korea); Kim et al., 1994: 262 (in Check List of In- tion); Nemkov, 2009: 49 (including Korea in distribution); sects from Korea); Yamane in Yamane et al., 1999: 479 (including Nemkov, 2012b: 434 (including Korea in distribution); Korea in distribution); Lee et al., 2010: 209 (in Checklist of Dollfuss, 2013a: 390 (in key), 400 (in key), 449e450 (including Korean Insects). Korea in distribution). Ammophila sabulosa nipponica Tsuneki, 1967a: 23, \ (holotype) _, Sphex infesta (Smith): Okamoto, 1924: 203 (new combination, Japan: Fukui Prefecture: Ohno [MNHAH], synonymized with including Korea in distribution); Uchida, 1925: 332 (Korea: Ammophila sabulosa solowiyofkae by Nemkov, 2005:141; Weonsan, Seokwangsa, Sambang, Seoul, Suwon, Hwasan, Tsuneki, 1968b: 50 (Korea: Jeju Is.: Mt. Hanna); Kim, 1970: 605 Busan), 369 (including Korea in distribution). (diagnosis, no Korean material); Tsuneki, 1974: 360 (N Korea: Ammophila sabulosa infesta (Smith): Tsuneki, 1962: 25 (in key, “Prov. Kanwon: Kumgangsan: Go-song chon; Kengia Bagyon including Korea in distribution); Yasumatsu, 1965: Pl. 150, 299 san: San-chon tong; Prov. South Pyongan: Chang-lyong san, (including Korea in distribution); Kim, 1970: Pl. 60 Figure 720, Nung-ra do; Prov. Ryang-gang: Chan-Pay Plateau: Sam-zi-yan; 605e606 (diagnosis, Korea: Ulleung Is.), 814 (in the list of Hy- Kaesong: Mts. Packyon”); Tsuneki, 1976a: 288e289 (including menoptera from Korea); Bohart and Menke, 1976: 153 (including Korea in distribution); Tsuneki, 1982: 2 (status confirmed; N Korea in distribution); Kim et al., 1994: 262 (in Check List of In- Korea: “Prov. Gang-von district: On-dzong: Kum-gang san: sects from Korea); Yamane, 2008: 557 (including Korea in dis- Mandzang-tae; N Korea: Prov. Pyong-sung: Bek-sung-li: Za-mo tribution); Lee et al., 2010: 209 (in Checklist of Korean Insects). san”; N Korea: “Prov. Ryang-gang: River karim”), 14 (in the list Distribution. Korean Peninsula (SL, GW, GG, CN, CB, GB, GN, JB, of the species of the Sphecidae known from the Korean Penin- JN, JJ), Russian Far East, Japan. sula); Paik, 1985: 197 (in the list of Sphecidae of Korea); Tsuneki, 1991: 199 (in Sphecidae from Korea); Kim et al., 1994: 262 (in Ammophila sickmanni Kohl, 1901 식크맨나나니 Check List of Insects from Korea); Kazenas in Nemkov et al., Ammophila sickmanni Kohl, 1901:151,\ (syntype), China: Tientsin: 1995: 387 (in key, as questionable synonym of A. solowiyofkae, Kalgan [NHMW]; Tsuneki, 1962: 26 (in key); Tsuneki, 1967a:10, including Korea in distribution); Yeo et al., 1998: 21 (in key), 23 11 (in key), 115 (description of _ from Korea); Tsuneki, 1974: 360 (S Korea: Seoul, Gangwon-do, Gyeonggi-do, Chungcheongbuk- (N Korea: “Prov. South Pyongan: Desangsan, Lyongak-san, do, Chungcheongnam-do, Gyeongsangbuk-do, Jeollabuk-do, Chang-lyong san; Kaesong: Mts. Packyon”); Bohart and Jeollanam-do, Jeju Is.); Terayama and Tano, 2006:15,18 Menke, 1976: 153 (including Korea in distribution); Tsuneki, (including Korea in distribution); Lee et al., 2010: 209 (in 1982: 14 (in the list of the species of the Sphecidae known Checklist of Korean Insects). from the Korean Peninsula); Paik, 1985: 197 (in the list of Distribution. Korean Peninsula (YG, PN, SL, GG, GW, CB, CN, GB, Sphecidae of Korea); Tsuneki, 1991: 199 (in Sphecidae from JB, JN, JJ), Mongolia, China, Russian Far East, Japan, India, Indonesia. Korea); Kim et al., 1994: 262 (in Check List of Insects from Ko- rea); Kazenas in Nemkov et al., 1995: 387 (in key, including Genus Hoplammophila de Beaumont, 1960 왕나나니속(신칭) Korea in distribution); Yeo et al., 1998: 21 (in key), 22 (S Korea: Hoplammophila Beaumont, 1960: 1. Type species: Ammophila 418 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 armata (Illiger, 1807) [¼ Sphex armatus Illiger, 1807], by original Sphex cyaneus Linnaeus, 1758) ¼ Pelopeus [sic] californicus de designation. Saussure, 1867], designated by Patton, 1880a: 378.

Hoplammophila aemulans (Kohl, 1901) 왕나나니 Chalybion japonicum (Gribodo, 1883) 보석나나니 Ammophila aemulans Kohl, 1901:144,\ (syntypes), SE Siberia and Pelopaeus japonicus Gribodo, 1883: 264, \ (lectotype, designated by Korea: no specific locality [NHMW]; Tsuneki, 1962: 25 (in key, Hensen, 1988: 54), _ (as tentative new species), Japan: no spe- including Korea in distribution); Kim, 1963: 118 (in Catalog of cific locality [RMNH]. Hymenoptera from Korea); Yasumatsu, 1965: Pl. 150, 229 Chalybion japonicum punctatum (Kohl): Bohart and Menke, 1976: (including Korea in distribution); Kim, 1970: Pl. 59 Figure 715, 103 (new status by Vecht, including Korea in distribution). 603 (diagnosis, Korea: Gwangleung), 814 (in the list of Hy- Chalybion japonicum (Girbodo): Bohart and Menke, 1976:102 menoptera from Korea); Kim, 1980: Pl. xviii HYSP 01, 132 (new combination); Tsuneki, 1968b: 50; Paik, 1985: 197 (in the (diagnosis, Korea: Gangwon-do, Gyeonggi-do, Gyeongsangbuk- list of Sphecidae of Korea); Hensen, 1988: 53 (examination of do). Korean material in BMNH and RMNH); Yamane in Yamane Hoplammophila aemulans (Kohl): Menke, 1966: 150 (new combi- et al., 1999:482e483 (including Korea in distribution); nation); Menke, 1974: 582 (including Korea in distribution); Kazenas in Nemkov et al., 1995: 382 (in key, including Korea in Tsuneki, 1982: 14 (in the list of the species of the Sphecidae distribution); Yeo et al., 1998: 22 (S Korea: Seoul, Gyeonggi-do, known from the Korean Peninsula); Paik, 1985: 197 (in the list of Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- Sphecidae of Korea); Kim et al., 1994: 262 (in Check List of In- do, Jeju Is.); Terayama and Tano, 2006: 16, 18 (including Korea sects from Korea); Kazenas in Nemkov et al., 1995: 385 (in key, in distribution); Nemkov, 2009: 43 (including Korea in dis- including Korea in distribution); Yeo et al., 1998: 18 (in key), 20 tribution); Nemkov, 2012b: 433 (including Korea in (S Korea: Gyeonggi-do); Yamane, 2008: Pl. 167, 557; Nemkov, distribution). 2009: 46 (including Korea in distribution); Lee et al., 2010: Sceliphron inflexum Sickmann, 1894:220,\ and _ (syntypes), 209 (in Checklist of Korean Insects); Nemkov, 2012b: 434 China: Hopei Province: Tientsing [Münster], synonymized (including Korea in distribution). with Sceliphron curvatum by Kohl, 1906a: 193, with Chalybion Distribution. S Korea (GG, GW, GB), Russian Far East, Japan, japonicum punctatum by Vecht in Bohart and Menke, 1976: Taiwan. 103, and later with Chalybion japonicum by Hensen, 1988:53: Yasumatsu, 1934: 63 (including Korea in distribution); Kim, Genus Podalonia Spinola, 1853 애기나나니속(신칭) 1963: 118 (in Catalog of Hymenoptera from Korea); Podalonia Spinola, 1853: 53. Suppressed by ICZN, 1968: 88 (Opinion Yasumatsu, 1965: Pl. 150, 230 (including Korea in distribution); 857). Tsuneki, 1968b: 50 (Korea: Jeju Is.: Jeju-City); Kim, 1970:Pl.60 Podalonia Fernald, 1927: 11. Validated by ICZN, 1968: 88 (Opinion Figure 723, 607 (diagnosis, Korea: Mt. Gayasan), 815 (in the list 857). Type species: Ammophila violaceipennis Lepeletier, 1845, of Hymenoptera from Korea); Tsuneki, 1968b: 50 (Korea: Jeju designated by ICZN, 1968: 88 (Opinion 857). Is.: Cheju-City); Tsuneki, 1976a: 293 (including Korea in dis- tribution); Kim, 1980: Pl. xix HYSP 06, 138 (diagnosis, Korea: Podalonia affinis (Kirby, 1798) 애기나나니 Gyeonggi-do, Chungcheongnam-do, Gyeongsangbuk-do, Ammophila affinis Kirby, 1798: 205, \ (syntype), Great Britain: Gyeongsangnam-do, Jeollabuk-do, Jeollanam-do, Jeju Is.); Martlesham Heath near Woodbridge [depository uncertain]; Kim et al., 1994: 262 (in Check List of Insects from van der Linden, 1827: 357 (redescription); Tsuneki, 1962: 25 (in Korea); Yamane, 2008: Pl. 167, 558 (including Korea in key, including Korea in distribution); Tsuneki, 1976a: 292 distribution). (including Korea in distribution). ? Sceliphron violaceum Fabricius: Uchida, 1925: 332 (Korea: Hwa- Podalonia affinis (Kirby): Guichard, 1938: 72 (new combination); san, Sambang), 369 (including Korea in distribution). Tsuneki, 1982: 14 (in the list of the species of the Sphecidae ? Chalybion violaceum Fabricius: Matsumura and Uchida, 1927:41 known from the Korean Peninsula); Paik, 1985: 197 (in the list of (including Korea in distribution). Sphecidae of Korea); Kazenas in Nemkov et al., 1995: 385 (in key, Distribution. Korean Peninsula (HB, SL, GG, CB, CN, GB, JJ), China, including Korea in distribution); Yeo et al., 1998: 18 (in key), 21 Russian Far East, Japan, India, SE Asia. (S Korea: Seoul, Gyeonggi-do, Chungcheongbuk-do, Note. As listed with question marks in synonymic list, there is Chungcheongnam-do, Gyeongsangbuk-do, Gyeongsangnam- currently no direct evidence that Korean occurrence records of do); Nemkov, 2009: 47 (including Korea in distribution); Sceliphron violaceum and Chalybion violaceum by Uchida (1925) and Nemkov, 2012b: 434 (including Korea in distribution). Matsumura and Uchida (1927) were the misconceptions of Sceli- Ammophila campestris Latreille: Kim, 1970: Pl. 60 Figure 716, 603 phron japonicum. Sphex violaceus Fabricius is junior primary hom- (as campoctris, diagnosis, Korea: Mt. Dobongsan), 814 (in the onym of Sphex violaceus Scopoli 1763, and is now Chalybion list of Hymenoptera from Korea), misidentification; Kim, 1980: bengalensis (Dahlbom, 1845)(Pulawski, 2014). Chalybion benga- Pl. xviii HYSP02, 133 (specific name incorrectly as campoctris, lensis is known to distribute in Central Ryukyus of Japan as the diagnosis, Korea: Gangwon-do, Gyeonggi-do, Jeollanam-do), northernmost distributional limit, South Ryukyus of Japan, Taiwan, misidentification; Paik, 1985: 197 (in the list of Sphecidae of Old World tropics, and Australia, and easily separated from Korea). C. japonicum by smaller body, longer abscissa 4 of radius and gently Distribution. Transpalearctic: S Korea (SL, GG, CB, CN, GB, GN), N curved metasomal petiole (Yamane in Yamane et al., 1999). Africa, Europe to Mongolia, China, Russian Far East, S to SE Asia. Regarding the currently known distributional range of the species, it seems more supportive that C. bengalensis is not one of the Subfamily Sceliphroninae Ashmead, 1899 보석나나니아과(신칭) Korean sphecid elements. Tribe Sceliphronini Ashmead, 1899 보석나나니족(신칭) Genus Chalybion Dahlbom, 1843 보석나나니속(신칭) Genus Sceliphron Klug, 1801 노랑점나나니속(신칭) Chalybion Dahlbom, 1843: 21. Type species: Chalybion caeruleum [¼ Sceliphron Klug, 1801: 561. Type species: Sphex spirifex Linnaeus, Sphex caeruleus of Linnaeus, 1767 ¼ Sphex caeruleus (nec Sphex 1758, designated by Bingham, 1897:235. caeruleus Linnaeus, 1758) ¼ Sphex cyaneus Fabricius, 1775 (nec JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 419

Sceliphron deforme atripes (Morawitz, 1888) 노랑점나나니 Mt. Gayasan), 814 (in the list of Hymenoptera from Korea); Kim, Pelopoeus atripes Morawitz, 1888:271,\ (lectotype, designated by 1980: 142 (Korean localities in Note of Sphex subfuscatus Dahl- Hensen, 1987: 240), Kazakhstan: Semipalatinsk [ZIN]. bom); Kim et al., 1994: 262 (in Check List of Insects from Korea); Sceliphron deforme atripes (Morawitz): Dalla Torre, 1897: 378 (new Lee et al., 2010: 210 (in Checklist of Korean Insects). combination); Bohart and Menke, 1976: 106 (new status); Palmodes occitanicus Lepeletier et Audinet-Serville: Bohart and Hensen, 1987: 234 (in key), 240 (Korea: Gyeonsanggnam-do: Menke, 1976: 127 (new combination); Tsuneki, 1982: 14 (in Jinyang-gun: Geumgok; Gyeonggi-do: Shoyozan); Yeo et al., the list of the species of the Sphecidae known from the Korean 1998:19e20 (S Korea: Seoul, Gangwon-do, Gyeonggi-do, Peninsula); Kazenas in Nemkov et al., 1995: 384 (in key, Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- including Korea in distribution); Nemkov, 2009: 45 (including do, Gyeongsangnam-do). Korea in distribution). Sceliphron formosum Smith var. koreanum Uchida, 1925: 332, \_, Distribution. Transpalearctic: S Korea (GG, CB, GN), N Africa, Korea: Hwasan, Sambang, synonymized with Sceliphron defrome Europe to Mongolia, China, Russian Far East, south to SE Asia. atripes by Hensen, 1987:240. Sceliphron deforme koreanum Uchida: Tsuneki, 1967a:6e7 (new Genus Prionyx van der Linden, 1827 어리조롱박벌속(신칭) status only for Korean specimens, redescriptions \ and _); Prionyx van der Linden, 1827: 362. Type species: Ammophila kirbii Tsuneki, 1971: 7 (listed); Bohart and Menke, 1976:106 van der Linden, 1827, by monotypy. (including Korea in distribution); Paik, 1985: 197 (in the list of Sphecidae of Korea). Prionyx subfuscatus (Dahlbom, 1845) 어리조롱박벌 Sceliphron deforme Smith: Kim, 1963: 118 (in Catalog of Hyme- Sphex subfuscatus Dahlbom, 1845: 436, sex not indicated (holotype noptera from Korea); Yasumatsu, 1965: Pl. 150, 299 (including or syntypes), Ukraine: “Tauria” (lost); Yasumatsu, 1938a:93e99 Korea in distribution); Kim, 1970: Pl. 6 Figure 722, 606e607 (Korea: “Keikido, Heianhokudo, Zenlahokudo, Chuseinando”); (diagnosis, Korea: Mt. Gayasan), 815 (in the list of Hymenoptera Kim, 1963: 117 (in Catalog of Hymenoptera from Korea); from Korea); Tsuneki, 1976a: 292e193; Kim, 1980: Pl. xix Tsuneki, 1968b: 50 (Korea: Jeju Is.: “Kinneikutsu”); Kim, 1970: Pl. HYSP05, 137 (diagnosis, Korea: Gangwon-do, Gyeonggi-do, 59 Figure 712, 602 (diagnosis, Korea: Mt. Gayasan), 813 (in the Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- list of Hymenoptera from Korea); Kim, 1980: pl. xix HYSP 10, 142 do, Jeollabuk-do, Jeollanam-do); Kim et al., 1994: 262 (specific (diagnosis, Korea: Gyeonggi-do); Tsuneki, 1982: 14 (in the list of name incorrectly as deforine, in Check List of Insects from Ko- the species of the Sphecidae known from the Korean Peninsula); rea); Kazenas in Nemkov et al., 1995: 382 (in key, including Kim et al., 1994: 263 (in Check List of Insects from Korea); Lee Korea in distribution); Yamane in Yamane et al., 1999:481e482 et al., 2010: 210 (in Checklist of Korean Insects). (including Korea in distribution); Terayama and Tano, 2006:16, Prionyx subfuscatus (Dahlbom): Diniz, 1965: 4 (new combination); 18 (including Korea in distribution); Yamane, 2008: Pl. 167, 558 Tsuneki, 1982: 14 (in the list of the species of the Sphecidae (including Korea in distribution); Nemkov, 2009:43e44; Lee known from the Korean Peninsula); Kazenas in Nemkov et al., et al., 2010: 210 (in Checklist of Korean Insects); Nemkov, 1995: 385 (in key, including Korea in distribution); Nemkov, 2012b: 433 (including Korea in distribution). 2009: 46 (including Korea in distribution); Nemkov, 2012b: Distribution. Korean Peninsula (HB, SL, GG, GW, CB, CN, GB, GN, 434 (including Korea in distribution). JB, JN), SE Asia, China, Russian Far East, Japan. Distribution. Transpalearctic: Korean Peninsula (PB, GG, CN,GB, JB, JJ), Africa, Europe to Mongolia, China, Russian Far East, south to Sceliphron madraspatanum kohli Sickmann, 1781 애황나나니 India, SE Asia. Sceliphron kohli Sickmann, 1894:218,\ and _ (syntypes), China: Hopei Province: Tientsin [NHMW]. Tribe Sphecini Latrille, 1802 구멍벌족(신칭) Sceliphron madraspatanum kohli (Kohl): Gussakovskij, 1938: 4 (new Genus Isodontia Patton, 1880 조롱박벌속(신칭) status, China: Kiangsu Prov.); Paik, 1985: 197 (in the list of Isodontia Patton, 1880a: 380. Type species: Sphex philadelphica Sphecidae of Korea). Lepeletier, 1845, by original designation. Sceliphron madraspatanum Fabricius: Kim, 1963: 118 (in Catalog of Hymenoptera from Korea); Kim,1970: 607 (diagnosis, no Korean Isodontia harmandi (Pérez, 1905) 홍다리조롱박벌 material), 815 (in the list of Hymenoptera from Korea); Kim Sphex harmandi Pérez, 1905:151,_ (holotype or syntype; actually \ et al., 1994: 262 (in Check List of Insects from Korea); Lee by Berland, 1926: 283), presumably Tokyo area [MNHN]; Kim, et al., 2010: 210 (in Checklist of Korean Insects). 1970: Pl. 59 Figure 710, 601 (diagnosis, Korea: Mt. Cheonma- Distribution. ?Korea, China, Japan; south to Taiwan, SE Asia. san), 813 (in the list of Hymenoptera from Korea); Kim, 1980: Pl. Note. As far as I am aware, no Korean material was examined. xix HYSP 08, 140 (diagnosis, Korea: Gangwon-do, Gyeonggi-do, Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- Subfamily Sphecinae Latreille, 1802 구멍벌아과(신칭) do, Jeollabuk-do, Jeollanam-do, Jeju Is.). Tribe Prionynchini Bohart and Menke,1963 어리조롱박벌아과(신칭) Isodontia harmandi (Pérez): Lin, 1966: 244 (new combination); Genus Palmodes Kohl, 1890 홍배조롱박벌속(신칭) Tsuneki, 1982: 14 (in the list of the species of the Sphecidae Palmodes Kohl, 1890: 112. Type species: Sphex occitanicus Lepeletier known from the Korean Peninsula); Paik, 1985: 197 (in the list of et Audinet-Serville, 1828, designated by Fernald, 1906:318. Sphecidae of Korea); Kim et al., 1994: 262 (in Check List of In- sects from Korea); Terayama and Tano, 2006: 13 and 17 Palmodes occitanicus (Lepeletier et Audinet-Serville, 1825e1828) (including Korea in distribution); Lee et al., 2010: 209 (in 홍배조롱박벌 Checklist of Korean Insects). Sphex occitanicus Lepeletier et Audinet-Serville, 1828: 462, _ (syn- Distribution. S Korea (GG, GW, CB, CN, GB, JB, JN, JJ), Japan. types), France: Hérault: Montpellier area (M. Spinola collection, Torino); Uchida,1925: 332 (N Korea: Seokwangsa), 369 (including Isodontia nigella (Smith, 1856) 먹조롱박벌 Korean in distribution); Yasumatsu, 1938a:79e83 (Korea: “Zokuri, Sphex nigellus Smith, 1856: 255, \_ (syntypes), China: Shanghai Sharin, Shoyozan”); Kim, 1963: 117 (in Catalog of Hymenoptera [BMNH]; Yasumatsu, 1938a:99e103 (Korea: “Keijo” (Seoul)); from Korea); Kim, 1970: Pl. 59 Figure 712, 602 (diagnosis, Korea: Kim, 1963: 117 (in Catalog of Hymenoptera from Korea); 420 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Tsuneki, 1968b: 50 (Korea: Jeju Is.: Jeju-City); Kim, 1970: Pl. 59 list of the species of the Sphecidae known from the Korean Figure 711, 601e602 (diagnosis, Korea: Seoul), 813 (in the list of Peninsula); Paik, 1985: 197 (in the list of Sphecidae of Korea); Hymenoptera from Korea); Tsuneki, 1974: 361 (N Korea: “Prov. Terayama and Tano, 2006: 18 (including Korea in distribution). South Pyongan: Bongwa-ri”); Kim, 1980: Pl. xix HYSP 09, 141 Sphex haemorrhoidalis Fabricius: Yasumatsu, 1938a:64e69 (Korea: (diagnosis, S Korea: Gyeonggi-do, Chungcheongnam-do, “Kojo, Koryo, Yusen, Kazan, Jinsen, Eishu Islet, Keishu”), cor- Gyeongsangnam-do); Tsuneki, 1991: 199 (in the list of Spheci- rected to Sphex subtruncatus by Tsuneki, 1982: 1 (N Korea: “Prov. dae of Korea). Pyong-sung, Bek-sung-li: Za-mo san”); Kim, 1963: 117 (in Cat- Isodontia nigella (Smith): Patton, 1880a: 381 (new combination); alog of Hymenoptera from Korea). Tsuneki, 1982: 14 (in the list of the species of the Sphecidae Sphex nigripes Smith: Uchida, 1925: 332 (Korea: Hwasan, Suwon), known from the Korean Peninsula); Paik, 1985: 197 (in the list of 369, corrected as S. haemorrhoidalis by Yasumatsu, 1938a:65; Sphecidae of Korea); Kim et al., 1994: 262 (specific name Matsumura and Uchida,1927: 39 (including Korea in distribution). incorrectly given as nigellus in Check List of Insects from Korea); Sphex nigripes Smith var. muticus Kohl: Matsumura, 1911:116; Kazenas in Nemkov et al., 1995: 384 (in key, including Korea in Matsumura and Uchida, 1927: 39 (including Korea in distribution); Terayama and Tano, 2006: 13, 17 (including Korea distribution). in distribution); Nemkov, 2009: 45 (including Korea in distri- Sphex paludosus Rossi: Uchida, 1925: 332 (N Korea: Hwasan), 369, bution); Lee et al., 2010: 210 (in Checklist of Korean Insects); corrected to S. haemorrhoidalis by Yasumatsu, 1938a: 65; Kim, Nemkov, 2012b: 434 (including Korea in distribution). 1963: 117 (in Catalog of Hymenoptera from Korea); Kim, 1970: Distribution. Korean Peninsula (PN, SL, GG, CN, GN, JJ), Japan; 813 (in the list of Hymenoptera from Korea); Paik, 1985: 197 (as India, Taiwan, Indo-China, Philippines, Myanmar, Australia. pauldosa in the list of Sphecidae of Korea); Kim et al., 1994: 263 (in Check List of Insects from Korea); Lee et al., 2010: 210 (in Genus Sphex Linnaeus, 1758 구멍벌속 Checklist of Korean Insects). Sphex Linnaeus,1758: 569. Type species: Sphex flavipennis Fabricius, Distribution. Korean Peninsula (PS, GG), China, ?Japan, India, S 1793, designated by ICZN, 1946: 571 (Opinion 180), which Asia. reversed its earlier designation as the type of Sphex sabulosus Linnaeus, 1758 (ICZN, 1911: 76, Opinion 32). Family Crabronidae Latrille, 1802 은주둥이벌과 Subfamily Astatinae Lepeletier, 1845 파리구멍벌아과 Sphex argentatus fumosus Kohl, 1890 조롱박벌 Genus Astata Latreille, 1796 파리구멍벌속 Sphex umbrosus var. fumosus Kohl, 1890: 408 (as fumosa, incorrect Astatus Latreille, 1796: 114 (no species included). Type species: original termination, authorship attributed to Mocsáry), \ (ho- Tiphia abdominalis Panzer, 1798 [¼ Sphex boops Schrank, 1781], lotype or syntypes), Japan: no specific locality [TMB]. designated by Latreille, 1802b:337 (first included species). See Sphex argentatus fumosus Kohl: Tsuneki, 1962: 6 (new status); Kim, also ICZN, 1943a: 37, Opinion 139. 1970: Pl. 59 Figure 709, 601 (specific name incorrectly as argen- Astata Latreille, 1796: xiii (in Errata: “au lieu d’Astatus lisez Astata”). tata, diagnosis, Korea: Mt. Gayasan), 814 (in the list of Hymenop- tera from Korea); Kim, 1980: Pl. xix HYSP 07, 139 (author Astata boops (Schrank, 1781) 파리구멍벌 incorrectly as Mocsáry, diagnosis, Korea: Gangwon-do, Gyeonggi- Sphex boops Schrank, 1781: 384, _ (holotype or syntypes), Austria: do, Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- Wien: Prater (destroyed). do, Jeollanam-do, Jeju Is.); Tsuneki, 1982:14(inthelistofthe Astata boops (Schrank): Kohl, 1885: 434e435 (new combination); species of the Sphecidae known from the Korean Peninsula); Paik, Yasumatsu, 1939b:101 (Korea: Gyeonggi-do: Dobongsan); Kim, 1985: 196 (in the list of Sphecidae of Korea); Kim et al., 1994:262 1970: Pl. 56 Figure 668, 574 (diagnosis, Korea: Gwangleung), (specific name and author incorrectly as argentata and Mocsáry, in 809 (in the list of Hymenoptera from Korea); Kim, 1980: Pl. xviii Check List of Insects from Korea); Terayama and Tano, 2006:13,17 HYAS 01, 129 (diagnosis, Korea: Gangwon-do, Gyeonggi-do, (including Korea in distribution); Lee et al., 2010: 210 (author Gyeongsangbuk-do, Jeollabuk-do); Tsuneki, 1974: 362 (N Korea: incorrectly as Mocsáry, in Checklist of Korean Insects). “Prov. South Pyongan: Sa-gam”); Tsuneki, 1982: 15 (in the list of Sphex argentatus Fabricius: Kim, 1970: 600 (specific name given the species of the Sphecidae known from the Korean Peninsula); incorrectly as argentata, diagnosis, no Korean material), 813 (in Paik, 1985: 198 (in the list of Sphecidae of Korea); Tsuneki, 1991: the list of Hymenoptera from Korea); Paik, 1985: 197 (in the list 199 (in Sphecidae from Korea); Kim et al., 1994: 260 (in Check of Sphecidae of Korea); Kim et al., 1994: 262 (specific name List of Insects from Korea); Nemkov in Nemkov et al., 1995: given incorrectly as argentata, in Check List of Insects from Ko- 406e408 (in key, including Korea in distribution); Nemkov, rea); Lee et al., 2010: 210 (in Checklist of Korean Insects). 2009:74;Lee et al., 2010: 208 (in Checklist of Korean Insects); Sphex (Sphex) umbrosus Christ, 1791: 293, sex not indicated (holo- Nemkov, 2012b: 437 (including Korea in distribution). type or syntypes), origin not indicated [destroyed], synony- Distribution. Transpalearctic: Korean Peninsula (PN, GG, GW, GB, mized with Sphex argentatus by van der Vecht, 1961: 28, and JB), Europe to Mongolia, NE China, Russian Far East, Japan. 1973: 345; Kim, 1963: 117 (in Catalog of Hymenoptera from Korea). Subfamily Bembicinae Latreille, 1802 코벌아과 Chlorion umbrosus (Christ): Okamoto, 1924: 203 (new combination, Tribe Alyssontini Dalla Torre, 1897 알리손어리감탕벌족 new to Korea: Jeju Is.); Yasumatsu, 1938a:54e61 (Korea: Genus Alysson Panzer, 1806 알리손어리감탕벌속(신칭) “Kongosan” (Mt. Geumgangsan), “Keijo”). Alysson Panzer, 1806: 169. Type species: Pompilus spinosus Panzer, Distribution. Korean Peninsula (GG, GW, GG, CB, CN, GB, JN, JJ), 1801, designated by Morice and Durrant, 1915:406. Japan. Alysson pertheesi Gorski, 1852 홍가슴어리감탕벌 Sphex subtruncatus Dahlbom, 1843 로시조롱박벌 Alysson pertheesi Gorski, 1852: 178 (as Perthéesi, incorrect original Sphex subtruncatus Dahlbom, 1843:25,\ (lectotype, designated by capitalization and diacritic mark), \ (syntypes), Lithuania: Vil- Vecht 1973: 350), Africa (no specific locality but actually Oriental nius [lost, except one specimen in ZMHU]; Tsuneki, 1974:361(N Region according to Schulz, 1912: 94) [MZLU]; Tsuneki, 1982:1 Korea: “Prov. South Pyongan: Pyongyan”); Tsuneki, 1982: 18 (in (North Korea: “Prov. Pyong-sung, Bek-sung-li, Za-mo san”), 14 (in the list of the species of the Sphecidae known from the Korean JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 421

Peninsula); Paik, 1985: 203 (in the list of Sphecidae of Korea); 534 (including Korea in distribution); Terayama, 2006:18 Tsuneki, 1991: 199 (in Sphecidae from Korea); Kim et al., 1994: (including Korea in distribution); Lee et al., 2010: 210 (in 263 (in Check List of Insects from Korea); Nemkov in Nemkov Checklist of Korean Insects). et al., 1995: 455 (in key, including Korea in distribution); Distribution. Korean Peninsula (PN, GW, GG, GB, JB, JN), C Asia Terayama, 2006: 4, 19 (including Korea in distribution); (Kazakhstan), Mongolia, N China, Russian Far East. Nemkov, 2009: 127 (including Korea in distribution); Lee et al., 2010: 210 (in Checklist of Korean Insects); Nemkov, 2012b:445 Subtribe Exeirina Dalla Torre, 1897 큰줄나나니아족(신칭) (including Korea in distribution). Genus Argogorytes Ashmead, 1899 큰줄나나니속(신칭) Distribution. Transpalearctic: N Korea (PN), Europe to Russian Argogorytes Ashmead, 1899: 324. Type species: Gorytes carbonarius Far East, Japan. Smith, 1856, by original designation.

Alysson ratzeburgi Dahlbom, 1843 라쯔버그어리감탕벌 Argogorytes mystaceus grandis (Gussakovskij, 1932) 큰줄나나니 Alyson ratzeburgi Dahlbom, 1843:145(asRatzeburgi, incorrect Gorytes grandis Gussakovskij, 1932: 28, \ (syntypes), Russia: Pri- original capitalization), \ and _ (lectotype, designated by de morskiy Kray: Sedanka, Sutshan, and Tigrovaya [NRS]; Kim et al., Beaumont, 1953: 197), Norway: Ostre Naess Vaerdaliae 1994: 263 (in Check List of Insects from Korea). [MZLU]; Tsuneki, 1965a: 25 (in key, include N Korea in dis- Argogorytes mystaceus grandis (Gussakovskij): Tsuneki, 1963a:2 tribution); Tsuneki, 1965b: 47 (Korea: “Nansetsu-rei” (Nam- (new combination, in key), 6-7 (Korea: “Mt. Hakuto, Tonei, seolryeong, Ryanggang-do)); Kim, 1970: 612 (diagnosis, no Gykusenji”); Tsuneki, 1965a:28e29 (in key, including Korea in Korean material), 815 (in the list of Hymenoptera from Korea); distribution); Kim, 1970: Pl. 92 Figure 729, 613e614 (diagnosis, Tsuneki, 1982: 18 (in the list of the species of the Sphecidae no Korean material), 816 (in the list of Hymenoptera from known from the Korean Peninsula); Paik, 1985: 203 (in the list Korea); Tsuneki, 1982: 18 (in the list of the species of the of Sphecidae of Korea); Kim et al., 1994: 263 (in Check List of Sphecidae known from the Korean Peninsula); Paik, 1985: 202 Insects from Korea); Nemkov in Nemkov et al., 1995: 455 (in (in the list of Sphecidae of Korea); Kim et al., 1994: 263 (sub- key, including Korea in distribution); Nemkov, 2009:127 specific name incorrectly as gromdis, in Check List of Insects (including Korea in distribution); Lee et al., 2010: 210 (in from Korea); Nemkov in Nemkov et al., 1995: 459 and 460 (in Checklist of Korean Insects); Nemkov, 2012b: 445 (including key, including Korea in distribution); Terayama, 2006: 10 and Korea in distribution). 19 (including Korea in distribution); Nemkov, 2009:132 Distribution. Transpalearctic: N Korea (RG), Europe to Russian (including Korea in distribution); Lee et al., 2010: 210 (in Far East, Japan. Checklist of Korean Insects); Nemkov, 2012b: 446 (including Korea in distribution). Tribe Bembicini Latreille, 1802 코벌족 Distribution. N Korea (RG), NE China, Russian Fat East, Japan. Subtribe Bembicina Latrille, 1802 코벌아족(신칭) Genus Bembix Fabricius, 1775 코벌속 Subtribe Gorytina Lepeletier de Saint Farageau, 1845 줄나나니아 Bembix Fabricius, 1775: Characteres Generum, p. (¼ xxiii), no 족(신칭) included species. Type species: Bembex rostrata of Fabricius, Genus Eogorytes Bohalt, 1976 털보줄나나니속(신칭) 1781 (¼ Apis rostrata Linnaeus, 1758), designated by Latreille, Eogorytes Bohart in Bohart and Menke, 1976: 52 and 505. Type 1810: 438 (one of three species included in Bembyx by Fab- species: Gorytes fulvohirtus Tsuneki, 1963, by original ricius, 1775: 361). Morice and Durrant (1915: 400) gave the first designation. type species designation for the spelling of Bembix: Apis rostrata Linnaeus, 1757. Eogorytes fulvohirtus (Tsuneki, 1963) 털보줄나나니 Gorytes fulvohirtus Tsuneki, 1963a: 3 (in key), 11e12, \ (holotype), Bembix niponica picticollis (F. Morawitz, 1889) 코벌 Japan: Fukui Prefecture: Koike, and Paratype from Korea Bembex picticollis F. Morawitz, 1889:144,\_ (syntypes), China or (“Keijo”) [MNHAH]; Tsuneki, 1965a: 30 (in key, including Korea Mongolia: Hetshuen [ZIN]; Kim et al., 1994: 263 (as Bembix, in in distribution); Kim, 1970: 614 (diagnosis, no Korean material), Check List of Insects from Korea). 816 (in the list of Hymenoptera from Korea); Kim et al., 1994: Bembix niponica picticollis (F. Morawitz): Tsuneki, 1965a: 27 (new 263 (in Check List of Insects from Korea); Lee et al., 2010: 210 (in status, in key, including Korea in distribution); Kim, 1970: Pl. 61 Checklist of Korean Insects). Figure 743, 620 (diagnosis, Korea: Mt. Taebaeksan); Tsuneki, Eogorytes fulvohirtus (Tsuneki): Bohart and Menke, 1976: 505 (new 1974: 361 (N Korea: “Prov. South Pyongan: Sa-gam”); Tsuneki, combination, including Korea in distribution); Tsuneki, 1982:18 1976a: 295e296 (including Korea in distribution); Tsuneki, (listed with question mark, in the list of the species of the 1982: 11, 18 (in the list of the species of the Sphecidae known Sphecidae known from the Korean Peninsula), 19 (Korean from the Korean Peninsula); Paik, 1985: 202 (in the list of occurrence uncertain); Terayama, 2006: 13, 19 (including Korea Sphecidae of Korea); Tsuneki, 1991: 199 (in Sphecidae from in distribution). Korea); Kazenas in Nemkov et al., 1995: 466 (in key, including Gorytes sinensis Yasumatsu: Ishikawa, 1965: 301 (including Korea in Korea in distribution); Wu and Zhou, 1996: 15 (including Korea distribution), misidentification, corrected by Tsunek, 1982: 19 in distribution); Nemkov, 2009: 142 (including Korea in distri- and Pulawski, 2014, and now Ammatomus sinensis that is known bution); Lee et al., 2010: 210 (in Checklist of Korean Insects); to be endemic to China; Yamane, 2008: 559 (including Korea in Nemkov, 2012b: 446 (including Korea in distribution). distribution). Bembex nipponica Smith, 1873a: 194, _ (syntypes), Japan: Hyogo Distribution. S Korea (SL), Japan. [BMNH]; Kim, 1970: Pl. 61 Figure 742, 620 (diagnosis, Korea: Note. Tsuneki (1982: 19) commented that the Korean occurrence Naeseolak), 817 (in the list of Hymenoptera from Korea); Kim, of Eogorytes fulvohirtus was uncertain, and he (1982: 18) listed with 1980: Pl. xx HYBE 01a, 143 (specific name incorrectly as question. However, Tsuneki (1963a: 12) originally designated one niphonica, diagnosis, Korea: Gangwon-do, Gyeonggi-do, sample of Korean material as the paratype of this species, thus I Jeollabuk-do, Jeollanam-do); Kim et al., 1994: 263 (as Bembix in confirm the occurrence of this species in Korea. Check List of Insects from Korea); Yamane in Yamane et al., 1999: 422 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Genus Gorytes Latreille, 1805 줄나나니속(신칭) NMK]; Nemkov in Nemkov et al., 1995: 463 and 464 (in key, Gorytes Latreille, 1805: 180. Type species: Mellinus quinquecinctus including Korea in distribution); Terayama, 2006:10,19(including Fabricius, 1793, by monotypy. Korea in distribution); Nemkov, 2009: 136 (including Korea in dis- tribution); Nemkov, 2012b: 446 (including Korea in distribution). Gorytes laticinctus koreanus Handlirsch, 1888b 참줄나나니 Gorytes hakutozanus Tsuneki, 1963a: 4 and 6 (in key), 17e18, \ (ho- Gorytes koreanus Handlirsch, 1888b:471(asKoreanus, incorrect orig- lotype) and _, North Korea: Ryanggang-do: “Nansetsu-rei” inal capitalization), \ (holotype), Korea: no specificlocality[NMK]; [MNHAH], synonymized with Gorytes neglectus by Nemkov,1989: Tsuneki,1963a: 3 and 5 (in key, including Korea in distribution),13e 72; Tsuneki, 1965a:32e33 (in key, including Korea in distribu- 14; Tsuneki,1965a: 33 (in key, including Korea in distribution); Kim, tion); Kim, 1970: Pl. 60 Figure 732, 615 (diagnosis, Korea: Mt. 1970: 615 (diagnosis, no Korean material), 815 (in the list of Hy- Seolaksan), 816 (in the list of Hymenoptera from Korea); Bohart menoptera from Korea); Tsuneki, 1974:361(NKorea:“Prov. Ryang- and Menke, 1976: 501 (including Korea in distribution); Tsuneki, gang: Chan-Pay Plateau”); Tsuneki, 1982: 18 (in the list of the spe- 1982: 18 (in the list of the species of the Sphecidae known from cies of the Sphecidae known from the Korean Peninsula); Paik, the Korean Peninsula); Paik, 1985: 202 (in the list of Sphecidae of 1985: 202 (in the list of Sphecidae of Korea); Tsuneki, 1991:199 Korea); Kim et al., 1994: 263 (in Check List of Insects from Korea); (in Sphecidae from Korea); Kim et al., 1994: 263 (in Check List of Lee et al., 2010: 210 (in Checklist of Korean Insects). Insects from Korea); Terayama, 2006: 13, 20 (including Korea in Distribution. Transpalearctic: Korean Peninsula (RG, GW), distribution); Nemkov, 2009: 135 (including Korea in distribution); Europe to Russian Far East, Japan. Lee et al., 2010:210(inChecklistofKoreanInsects). Gorytes laticinctus koreanus Handlirschch: Nemkov, 1989: 68, 70, 71 Gorytes pieli Yasumatsu, 1943 타케우치줄나나니 (new status); Nemkov in Nemkov et al., 1995: 461 and 464 (in Gorytes pieli Yasumatsu, 1943:15,\ (holotype), China: Inner key, including Korea in distribution); Nemkov, 2012b:446 Mongolia: Chahar: Yangkianping [HMS]; Nemkov in Nemkov (including Korea in distribution). et al., 1995: 463 and 464 (in key, including Korea in distribu- Gorytes radoszkowskyi Handlirsch, 1888b:470(asRadoszkowskyi, tion); Nemkov, 2009: 136 (including Korea in distribution); incorrect original capitalization), \ (holotype), Korea: no specific Nemkov 2012: 446 (including Korea in distribution). locality [NMK], synonymized with Gorytes laticinctus koreanus by Gorytes takeuchii Tsuneki, 1963a: 5 (in key), 18e19, _ (holotype), Nemkov, 1989:71;Dalla Torre, 1897: 553 (including Korea in Korea: “Kazan” (Hwasan) [MNHAH], synonymized with Gorytes distribution); Maidl and Klima, 1939: 71 (including Korea in pieli by Nemkov, 1989:72;Tsuneki, 1965a: 32 (in key, including distribution); Tsuneki, 1963a: 3 (in key), 15 (Korea: “Daihyori, Korea in distribution); Kim, 1970: 616 (diagnosis, no Korean Shoyzan”); Tsuneki, 1965a: 31 (in key, including Korea in distri- material), 816 (in the list of Hymenoptera from Korea); Bohart bution); Kim, 1970: 616 (diagnosis, no Korean material), 816 (in and Menke, 1976: 502 (including Korea in distribution); the list of Hymenoptera from Korea); Bohart and Menke, 1976: Tsuneki, 1982: 18 (in the list of the species of the Sphecidae 501 (including Korea in distribution); Tsuneki, 1982:18(inthelist known from the Korean Peninsula); Paik, 1985: 202 (in the list of of the species of the Sphecidae known from the Korean Penin- Sphecidae of Korea); Kim et al., 1994: 263 (in Check List of In- sula); Paik,1985: 202 (in the list of Sphecidae of Korea); Kim et al., sects from Korea); Lee et al., 2010: 210 (in Checklist of Korean 1994: 263 (in Check List of Insects from Korea). Insects). Distribution. Korean Peninsula (PN, GG), Russian Far East, Japan. Distribution. N Korea (Hwasan), Mongolia, NE China, Russian Far East. Gorytes maculicornis (F.Morawitz, 1889) 황줄나나니 Hoplisus maculicornis F. Morawitz, 1889:149,\ (lectotype, desig- Gorytes quadrifasciatus (Fabricius, 1804) 네줄나나니 nated by Nemkov, 1989:72), China: Gansu: Sigu [ZIN]. Mellinus quadrifasciatus Fabricius, 1804: 298, _ (holotype), Ger- Gorytes maculicornis (F. Morawitz): Handlirsch, 1895: 922 (new many: no specific locality (as 4-fasciatus, incorrect original combination, original description copied); Nemkov, 1989: 68, 70 spelling) [ZMUC]. (including Korea in distribution); Nemkov in Nemkov et al., Gorytes quadrifasciatus (Fabricius): Spinola, 1806: 93 (new combina- 1995: 462 and 464 (in key, including Korea in distribution); tion); Tsuneki, 1963a: 3 and 6 (in key), 15 (N Korea: “Kussho”, Terayama, 2006: 10, 19 (including Korea in distribution); “Nansetsurei”); Tsuneki, 1965a:30e31 (in key, including Korea in Nemkov, 2009: 136 (including Korea in distribution); Nemkov, distribution); Kim,1970:615e616 (diagnosis, no Korean material), 2012b: 446 (including Korea in distribution). 816 (in the list of Hymenoptera from Korea); Tsuneki, 1982: 18 (in Gorytes eous Gussakovskij, 1932: 30, \ (lectotype, designated by the list of the species of the Sphecidae known from the Korean Nemkov, 1989: 72) and _, Russia: Primorskiy Kray: Sedanka Peninsula); Paik,1985: 202 (in the list of Sphecidae of Korea); Kim [OMNH], synonymized with Gorytes maculicornis by Nemkov, et al., 1994: 263 (in Check List of Insects from Korea); Nemkov in 1989:72;Tsuneki, 1963a: 4 and 5 (in key), 14e15 (N Korea: Nemkov et al., 1995: 461 and 464 (in key, including Korea in dis- “Kussho, Nanyo, Muzanrei”); Tsuneki, 1965a: 31 and 32 (in key, tribution); Nemkov, 2009:136e137 (including Korea in distribu- including Korea in distribution); Yasumatsu, 1965:301 tion); Lee et al., 2010: 210 (in Checklist of Korean Insects). (including Korea in distribution); Kim, 1970: 614 (diagnosis, no Distribution. Transpalearctic: N Korea (RG), N Africa, Europe to Korean material), 816 (in the list of Hymenoptera from Korea); Russian Far East, Japan. Tsuneki, 1982: 18 (in the list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 202 (in the list of Gorytes tricinctus (Pérez, 1905) 줄나나니 Sphecidae of Korea); Kim et al., 1994: 263 (in Check List of In- Hoplisus tricinctus Pérez, 1905:157,\ (holotype or syntypes), Japan: sects from Korea); Yamane, 2008: 559 (including Korea in dis- no specific locality but presumably Tokyo area [MNHN]. tribution); Lee et al., 2010: 210 (in Checklist of Korean Insects). Gorytes tricinctus (Pérez): Maidl and Klima, 1939: 72 (new combi- Distribution. N Korea (RG), NE China, Russian Far East, Japan. nation); Kim, 1970: Pl. 92 Figure 737, 617 (diagnosis, no Korean material), 815 (in the list of Hymenoptera from Korea); Bohart Gorytes neglectus Handlirsch, 1895 백두산줄나나니 and Menke, 1976: 502 (listed, including Korea in distribution); Gorytes neglectus Handlirsch, 1895:916,\ and _ (syntypes), Russia: Paik, 1985: 202 (in the list of Sphecidae of Korea); Nemkov, Siberia between Irkutsk and mouth of Ussuri River [NHMW or 1989: 68, 70 (including Korea in distribution); Nemkov, 1990: JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 423

682, 685 (including Korea in distribution); Kim et al., 1994: 263 (in the list of Sphecidae of Korea); Nemkov in Nemkov et al., (in Check List of Insects from Korea); Terayama, 2006: 13, 20 1995: 465 (in key, including Korea in distribution); Nemkov, (including Korea in distribution); Lee et al., 2010: 210 (in 2009: 139 (including Korea in distribution); Nemkov, 2012b: Checklist of Korean Insects). 447 (including Korea in distribution). Distribution. ? Korea, Japan. Distribution. Korean Peninsula (PN, GG), Mongolia, N China, Note. Kim (1970: 815) first reported the Korean occurrence of this Russian Far East. species with the listing “Yasumatsu, Icon. Ins. Jap. P. 1470, 1953”.In 1953, Yasumatsu was not one of authors of the book Icon. Ins. Jap Genus Lestiphorus Lepeletier, 1832 두눈줄나나니속 (Iconographia Insectorum Japonicorum). It must be Yano (1953: Lestiphorus Lepeletier, 1832: 70. Type species: Crabro bicinctus 1740). However, Gorytes tricinctus illustrated and diagnosed by Yano Rossi, 1794, junior primary homonym of Crabro bicinctus Fab- (1953: 1740) was misidentified as G. maculicornis (refer to synonym ricius, 1793 (a name suppressed by ICZN, Opinion 675, 1963: list of G. maculicornis above). Furthermore, Tsuneki (1963a) who 331), validated in the same Opinion, by monotypy. revised the Japanese and Korean Gorytini also did not include Korea in the distributional range of G. tricinctus. Although Bohart and Lestiphorus bilunulatus yamatonis Tsuneki, 1963 두눈줄나나니 Menke (1976) included Korea in the distribution once more, unfor- Lestiphorus bilunulatus yamatonis Tsuneki, 1963a: 2 (in key), 9, \ tunately the source of the citation was not traceable. It is highly (holotype), Japan: Nikko [MNHAH], including Korea in distri- possible that later records for Korean occurrences were mere cita- bution with question mark; Tsuneki, 1965a: 29 (in key, material tions of misinterpreted work by Kim (1970) or Bohart and Menke from Ussuri and Korea may belong to this subspecies); Kim, (1976). There was no evidence that Korean material was examined, 1970: 617 (subspecific name listed incorrectly as yasumatsui, thus this species should be excluded from Korean fauna. diagnosis, no Korean material), 816 (in the list of Hymenoptera from Korea); Tsuneki, 1982:18 (in the list of the species of the Genus Harpactus Shuckard, 1807 민줄나나니속(신칭) Sphecidae known from the Korean Peninsula); Paik, 1985: 202 Arpactus Jurine, 1807: 192, junior homonym of Arpactus Panzer, (in the list of Sphecidae of Korea); Kim et al., 1994: 263 (in Check 1805, and of Arpactus Panzer, 1806. Type species: Arpactus for- List of Insects from Korea); Lee et al., 2010: 210 (in Checklist of mosus Jurine, 1807, designated by Shuckard, 1837: 220. Korean Insects). Harpactus Shuckard, 1837: 221 (emendation of Arpactus Jurine, Lestiphorus bilunulatus A. Costa: Handlirsch, 1888b: 453 (including 1807). Korea in distribution); Nemkov in Nemkov et al., 1995: 465 (in key, including Korea in distribution); Terayama, 2006: 13, 20 Harpactus laevis (Latreille, 1792) 민줄나나니 (including Korea in distribution); Nemkov, 2009:139e140 Mutilla laevis Latreille, 1792: 11, sex not indicated, holotype or (including Korea in distribution); Nemkov, 2012b:447 syntypes: SW France: Angoulême region: no specific locality (including Korea in distribution). (lost). Distribution. Korea (no specific locality), Russian Far East, Japan. Harpactus laevis (Latreille): F. Smith, 1853: 65 (new combination); Nemkov in Nemkov et al., 1995: 460 (in key, including Korea in Subtribe Stizina A. Costa, 1859 대만어리코벌아족(신칭) distribution). Genus Bembecinus A. Costa, 1859 대만어리코벌속 Dienoplus laevis (Latreille): Wolf, 1959: 30 (new combination); Bembecinus A. Costa, 1859:4. Type species: Bembecinus meridionalis Tsuneki, 1963a: 2 and 5 (in key, Korean and Japanese occurrence A. Costa, 1859, by monotypy. doubtable), 9 (possibly discovered in Korea, considering the distributional range); Kim, 1970: 816 (in the list of Hymenoptera Bembecinus quadratus Tsuneki, 1968 남방어리코벌 from Korea); Paik, 1985: 202 (in the list of Sphecidae of Korea). Bembecinus pacificus Tsuneki, 1968a:17,_ (holotype) and \, Taiwan: Distribution. Transpalearctic: ?Korea, N Africa, Europe to Taitung [USNM], 25 (S Korea: “Shoyozan”), junior secondary Mongolia, China, ?Japan. homonym of Bembecinus pacificus (R. Turner, 1917); Bohart and Note. Tsuneki (1963a) commented on the possibility of a future Menke, 1976: 531 (including Korea in distribution). discovery of this species in Korea and Japan. Therefore, this species Bembecinus quadratus Tsuneki, 1976b: 434, substitute name for should be excluded from Korean fauna until real material is available. Bembecinus pacificus Tsuneki; Tsuneki, 1982: 12, 18 (in the list of the species of the Sphecidae known from the Korean Peninsula); Genus Hoplisoides Girbodo, 1884 가시줄나나니속 Paik, 1985: 202 (in the list of Sphecidae of Korea). Hoplisoides Gribodo, 1884: 276. Type species: Hoplisoides intricans Distribution. S Korea (GG), Taiwan. Gribodo, 1884, by monotypy. Bembecinus hungaricus formosanus (Sonan, 1928) 대만어리코벌 Hoplisoides gazagnairei distinguendus (Yasumatsu, 1939c) 가시줄 Stizus formosanus Sonan, 1928:262, \ (holotype) and _, Taiwan: 나나니 Boryo [J. Sonan coll.]; Tsuneki, 1965a: 14, as synonym of Bem- Gorytes distinguendus Yasumatsu, 1939c:12,\ (holotype), China: becinus hungaricus. Heilongjianag: Changchu [KUF] Bembecinus formosanus (Sonan): Lohrmann, 1943:191 (new Hoplisoides distinguendus (Yasumatsu); Tsuneki, 1963a: 2 and 4 (in combination). key), 10e11 (new combination, Korea: “Shyozan”); Tsuneki, Bembecinus hungaricus Frivaldzky: Tsuneki, 1965a:28(inkey, 1965a:29e30 (in key); Kim, 1970:617e618 (diagnosis, no including Korea in distribution); Nemkov in Nemkov et al., 1995: Korean material), 816 (in the list of Hymenoptera from Korea); 466 (in key, including Korea in distribution); Terayama, 2006:16, Kim et al., 1994: 262 (in Check List of Insects from Korea); Lee 20 (including Korea in distribution); Nemkov, 2009: 140 (including et al., 2010: 210 (in Checklist of Korean Insects). Korea in distribution); Nemkov, 2012b: 446 (including Korea in Hoplisoides gazagnairei distinguendus (Yasumatsu): Tsuneki, 1971: distribution); Nemkov, 2012c: 4 (including Korea in distribution). 11 (new status); Tsuneki, 1974: 361 (N Korea: “Prov. South Bembecinus hungaricus formosanus (Sonan): Tsuneki, 1968a: 6 (new Pyongan: Chang-lyong san”); Tsuneki, 1982: 18 (in the list of the status), 22, 25 (S Korea: Seoul); Kim, 1970: Pl. 61 Figure 741, species of Sphecidae known from the Korean Peninsula); 619e620 (diagnosis, Korea: Gwangleung), 817 (in the list of Tsuneki, 1991: 199 (in Sphecidae from Korea); Paik, 1985: 202 Hymenoptera from Korea); Kim, 1980: 143 (Korean localities in 424 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

the Note of Bembex nipponica Smith); Tsuneki, 1982: 12, 18 (in Nysson maculosus (Gmelin): Lomholdt, 1975c: 213 (new combina- the list of the species of the Sphecidae known from the Korean tion); Tsuneki, 1982: 18 (in the list of the species of the Sphe- Peninsula); Paik, 1985: 202 (in the list of Sphecidae of Korea); cidae known from the Korean Peninsula); Paik, 1985: 203 (in the Kim et al., 1994: 263 (in Check List of Insects from Korea); Lee list of Sphecidae of Korea); Nemkov in Nemkov et al.,1995: 467e et al., 2010: 210 (in Checklist of Korean Insects). 459 (in key, including Korea in distribution); Terayama, 2006:6, Distribution. Transpalearctic: S Korea (SL, GW, GG), Europe to 19 (including Korea in distribution); Nemkov, 2009:130e131 Russian Far East, Japan, Taiwan. (including Korea in distribution); Nemkov, 2012b:446 (including Korea in distribution). Genus Stizus Latreille, 1802 어리코벌속 Distribution. Transpalearctic: N Korea (HB), Europe to Mongolia, Stizus Latreille, 1802b: 344. Type species: “Stizus ruficornis” Fab- China, Russian Far East. ricius [¼ Bembix ruficornis Fabricius, 1787 (junior secondary homonym of Vespa ruficornis J. Forster, 1771) ¼ Vespa ruficornis J. Subfamily Crabroninae Latreille, 1802 은주둥이벌아과 Forster, 1771, designated by Blanchard, 1846: Pl. 121. Tribe Crabronini Latreille, 1802 참빗은주둥이벌족 Subtribe Crabronina Latreille, 1802 침빗은주둥이벌아족(신칭) Stizus perrisi Dufour, 1838 어리코벌 Genus Crabro Fabricius, 1775 참빗은주둥이벌속 Stizus perrisi Dufour, 1838: 269 (as Perrisi, incorrect original capi- Crabro Fabricius, 1775: 373, junior homonym of Crabro Geoffroi de talization), _ (lectotype, designated by Bitsch, 1996: 21), France: Saint Hilaire,1762, which was suppressed by the ICZN, 1943b:91 Landes: Mont-de-Marsan [MNHN]. (Opinion 144) and 1994: 59 (Opinion 1754). Type species: Vespa Larra pulcherrima F. Smith, 1856:348,\ (holotype or syntypes), cribraria Linnaeus, 1758, designated by the ICZN, 1943b:91 China: Shanghai [BMNH], synonymized with Stizus perrisi by (Opinion 144). Nemkov, 2012a: 59. Stizus pulcherrimus (F. Smith): Handlirsch, 1892:172 (new combi- Crabro cribrarius (Linnaeus, 1758) 빗은주둥이벌 nation); Tsuneki, 1965a: 27 (in key, including Korea in distri- Vespa cribraria Linnaeus, 1758: 573, sex not indicated, \ (holotype), bution); Yasumatsu, 1965: 301 (including Korea in distribution); Europe: no specific locality [LSL]. Kim, 1970: 618 Pl. 61 Figure 740 (diagnosis, S Korea: Mt. Crabro (Crabro) cribrarius (Linnaeus): Fabricius, 1775: 374 (new Cheonmasan), 817 (in the list of Hymenoptera from Korea); combination, redescription); Tsuneki, 1947a: 286 (N Korea: Tsuneki, 1971: 8 (including Korea in distribution); Tsuneki, “Nansetsu-rei”); Kim, 1970: 640, 819 (in the list of Hymenoptera 1976a: 296e297 (including Korea in distribution); Kim, 1980: from Korea); Tsuneki, 1982: 17 (in the list of the species of the 143 (Korean localities in the Note of Bembex nipponica Smith); Sphecidae known from the Korean Peninsula); Paik, 1985:201 Tsuneki, 1982:18 (in the list of the species of the Sphecidae (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Check known from the Korean Peninsula); Paik, 1985: 202 (in the list of List of Insects from Korea); Nemkov in Nemkov et al., 1995:444 Sphecidae of Korea); Kim et al., 1994: 263 (in Check List of In- (in key, including Korea in distribution); Nemkov, 2009:113 sects from Korea); Kazenas in Nemkov et al., 1995: 465e466 (in (including Korea in distribution); Lee et al., 2010: 210 (in key, including Korea in distribution); Terayama, 2006: 15, 20 Checklist of Korean Insects); Nemkov, 2012b: 440 (including (including Korea in distribution); Yamane, 2008: 560 (including Korea in distribution). Korea in distribution); Nemkov, 2009: 141 (including Korea in Distribution. Transpalearctic: N Korea (RG), Europe to Mongolia, distribution); Lee et al., 2010: 210 (in Checklist of Korean In- Russian Far East. sects); Nemkov, 2012b: 447 (including Korea in distribution). Distribution. S Korea (SL, GG, CB, JJ), E Mongolia, E and NE China, Crabro peltarius (Schreber, 1784) 방패은주둥이벌 Russian Far East, Japan. Sphex peltarius Schreber, 1784: 98 (as peltaria, incorrect original termination), \_(syntypes), Germany: Neuwarp, now Poland: Tribe Nyssonini Latreille, 1804 어리감탕벌족 Nowe Warpno (depository uncertain). Subtribe Nyssonina Latrille, 1804 어리감탕벌아족(신칭) Crabro peltarius (Schreber): Schrank, 1802: 336 (new combination); Genus Nysson Latreille, 1802 어리감탕벌속(신칭) Kim, 1970: 640, 819 (in the list of Hymenoptera from Korea); Nysson Latreille, (1802b: 340). Type species: Crabro spinosus Fab- Tsuneki, 1957b: 60 [S Korea: “Zokurisan” (Mt. Sokrisan)]; Tsuneki, ricius, 1775 (¼ Sphex spinosus J. Forster, 1771), designated by 1982: 17(in the listof the species of the Sphecidae known from the Shuckard, 1837: 99. Validated by the ICZN, 1979:175 (Opinion Korean Peninsula); Paik, 1985: 201 (in the list of Sphecidae of 1115). Korea); Kim et al., 1994: 264 (in Check List of Insects from Korea); Nemkov in Nemkov et al., 1995: 444 (in key, including Korea in Nysson maculosus (Gmelin, 1790) 어리감탕벌 distribution); Yeo et al., 2008: 10 (in key), 10e13 (S Korea: Seoul, Sphex maculatus Fabricius, 1787: 277 (as maculata, incorrect original Gangwon-do, Gyeonggi-do, Chungcheongnam-do, Gyeongsang- termination), sex not indicated (lectotype, designated by van nam-do, Jeollanam-do); Nemkov, 2009:114e115 (including Korea der Vecht, 1961: 47), Germany: Kiel [ZMUC], junior primary in distribution); Lee et al., 2010: 210 (in Checklist of Korean In- homonym of Sphex maculatus Drury, 1775. sects); Nemkov, 2012b: 440 (including Korea in distribution). Nysson maculatus (Fabricius): Latreille, 1805: 306 (new combina- Distribution. Transpalearctic: S Korea (SL, GW, GG, CN, CB, GN, tion); Yasumatsu, 1938c: 195 (N Korea: “Mosanrei”); Maidl and JN), Europe to Mongolia, N China, Russian Far East. Klima, 1939: 123 (including Korea in distribution); Tsuneki, 1965a: 25 (in key, including Korea in distribution); Kim, 1970: Crabro ussuriensis Gussakovskij, 1932 참빗은주둥이벌 611 (diagnosis, no Korean material), 815 (in the list of Hyme- Crabro (Crabro) ussuriensis Gussakovskij, 1932:18,\ (holotype), noptera from Korea); Kim et al., 1994: 262 (in Check List of In- Russia: Primorskiy Kray: Sutshan [OMNH]; Tsuneki, 1977b:4 sects from Korea); Lee et al., 2010: 210 (in Checklist of Korean (redescription of _, S Korea: Gyeongsangbuk-do: Mt. sudosan); Insects). Tsuneki, 1982:9(NKorea:“Prov. Pyong-gang, Hyesan: Mt. Ze- Sphex maculosus Gmelin, 1790: 2731 (as maculosa, incorrect original dang”), 17 (in the list of the species of the Sphecidae known from termination), substitute name for Sphex maculatus Fabricius, the Korean Peninsula); Paik, 1985: 201 (in the list of Sphecidae of 1787. Korea); Nemkov in Nemkov et al., 1995: 444 and 446 (in key, JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 425

including Korea in distribution); Yeo et al., 2008: 10 (in key), 13e Crossocerus assimilis collaris Tsuneki, 1974 량강은주둥이벌(신칭) 15 (Korea: Seoul, Gangwon-do, Gyeonggi-do, Chungcheongnam- Crossocerus assimilis collaris Tsuneki, 1974: 377, _ (holotype) and \, do, Gyeongsangbuk-do, Gyeongsangnam-do); Nemkov, 2009: Korea: “Prov. Ryang-gang: Chan-Pay Plateau: Samzi-yan” [TMB]; 114 (including Korea in distribution); Nemkov, 2012b:441 Bohart and Menke, 1976: 629 (including Korea in distribution); (including Korea in distribution). Tsuneki, 1982: 17 (in the list of the species of the Sphecidae Crabro koreanus Tsuneki, 1947a: 287e289, \ (holotype), South Ko- known from the Korean Peninsula); Paik, 1985: 201 (in the list of rea: “Shoyo-zan” ca 16 km S Demilitarized Zone [MNHAH], Sphecidae of Korea); Tsuneki, 1991: 202 (as new species, nomen synonymized with Crabro ussuriensis by Marshakov, 1978: 859; nudum); Leclercq, 2009: 165 (in the subgenus Ablepharipus, Leclercq, 1954: 251 (including Korea in distribution); Kim, 1970: including Korea in distribution); Tano, 2010a: 37 (including 640, 819 (in the list of Hymenoptera from Korea); Paik, 1985: Korea in distribution). 201 (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Crossocerus assimilis (F. Smith): Nemkov in Nemkov et al., 1995: 434 Check List of Insects from Korea); Lee et al., 2010: 210 (in and 437 (in key, including Korea in distribution); Nemkov, 1999: Checklist of Korean Insects). 98 (including Korea in distribution); Nemkov, 2012b:441 Distribution. Korean Peninsula (RG, SL, GW, GG, CN, GB, GN), (including Korea in distribution). China, Russian Far East. Distribution. N Korea (RG).

Genus Crossocerus Lepeletier et Brullé, 1835 꼬마은주둥이벌속 Crossocerus ceratus (Schukard, 1837) 갑옷은주둥이벌 Crossocerus Lepeletier et Brullé, 1835: 763. Type species: Crabro Crabro cetratus Shuckard, 1837: 131, _ (syntypes), Great Britain: scutatus Fabricius, 1787 (¼ Sphex palmipes Linnaeus, 1767), London area (depository unknown). designated by Westwood, 1839: 80. Crossocerus ceratus (Schukard): Tsuneki, 1974a: 378 (N Korea: “Prov. Corenocrabro Tsuneki, 1974: 374. Type species: Corenocrabro ecte- Ryang-gang: Chan-Pay Plateau”); Tsuneki, 1982b: 17 (in the list miformis Tsuneki, 1974, by original designation and monotypy. of the species of the Sphecidae known from the Korean Penin- Synonymized with Crossocerus by Marshakov, 1979a: 96 and by sula); Paik, 1985: 201 (in the list of Sphecidae of Korea); Kim Nemkov in Nemkov et al., 1995: 428 (as new synonym). et al., 1994: 264 (in Check List of Insects from Korea); Nemkov in Nemkov et al., 1995: 435 and 440 (in key, including Korea in Crossocerus amurensis (Kohl, 1892) 아무르은주중이벌 distribution); Nemkov, 2009: 101 (including Korea in distribu- Crabro amurensis Kohl, 1892: 199, \ (holotype or syntypes), Russia tion); Lee et al., 2010: 210 (in Checklist of Korean Insects); or China: Amur area: no specific locality [NMNHA]. Nemkov, 2012b: 441 (including Korea in distribution). Crossocerus amurensis (Kohl): Leclercq, 1954: 233 (new combi- Distribution. Transpalearctic: N Korea (RG), Europe to Russian nation); Tsuneki, 1954b: 62 and 71 (including Korea in dis- Far East, Japan. tribution); Tsuneki, 1957b: 61 (Korea: “Shutotsu” (Hamgyeongbuk-do: Juheul), “Berg Gongo” (Mt. Geumgang- Crossocerus cinxius (Dahlbom, 1839) 남설령은주둥이벌(신칭) san)); Tsuneki, 1959b: 69 and 75 (including Korea in distri- Crabro cinxius Dahlbom, 1839: 38, \ and _ (syntypes), Norway: near bution); Kim, 1970: 636 (diagnosis, no Korean material), 822 Dovre [MZLU]; Tsuneki, 1947a: 290 (new to Korea, N Korea: (in the list of Hymenoptera from Korea); Tsuneki, 1982:17 “Nansetsu-rei”). (specific name incorrectly as amulensis in the list of the spe- Crossocerus cinxius (Dahlbom): Kirchner, 1867: 225 (new combina- cies of the Sphecidae known from the Korean Peninsula); Paik, tion); Tsuneki, 1954b: 63 and 69 (including Korea in distribu- 1985: 201 (in the list of Sphecidae of Korea); Kim et al., 1994: tion); Tsuneki, 1959b: 70 and 74 (including Korea in 264 (in Check List of Insects from Korea); Nemkov in Nemkov distribution); Kim, 1970: 635e636 (diagnosis, no Korean mate- et al., 1995: 434 and 439 (in key, including Korea in distri- rial), 822 (in the list of Hymenoptera from Korea); Tsuneki,1982: bution); Leclercq, 2009: 163 (in the subgenus Neoblepharipus, 17 (in the list of the species of the Sphecidae known from the including Korea in distribution); Nemkov, 2009: 111 Korean Peninsula); Paik, 1985: 201 (in the list of Sphecidae of (including Korea in distribution); Lee et al., 2010:210(in Korea); Kim et al.,1994: 264 (in Check List of Insects from Korea); Checklist of Korean Insects); Nemkov, 2012b: 442 (including Nemkov in Nemkov et al., 1995: 436 and 441 (in key, including Korea in distribution). Korea in distribution); Nemkov, 2009: 102 (including Korea in Distribution. N Korea (HB, GW), China, Russian Far East, Japan. distribution); Lee et al., 2010: 210 (in Checklist of Korean In- sects); Nemkov, 2012b: 441 (including Korea in distribution). Crossocerus annulipes hokkaidoensis Tsuneki, 1954 북해도은주둥 Distribution. Transpalearctic: N Korea (RG), Europe to N China, 이벌 Russian Far East, Japan. Crossocerus ambiguus hokkaidoensis Tsuneki, 1954b: 78, \ (holo- type), Japan: Hokkaido: Sapporo [MNHAH], corrected to Cross- Crossocerus denticoxa (Bischoff, 1932) 밑마디은주둥이벌(신칭) cerus annulipes hokkaidonensis by Pulawski 2014:13;Tsuneki, Crabro denticornis Gussakovskij, 1932: 24, _ (holotype), Russia: 1974: 378 (N Korea: “Prov. South Pyongan: Pyongyan”); Primorskiy Kray: Sedanka near Vladivostok [OMNH], junior Tsuneki, 1991: 202 (in Sphecidae from Korea). primary homonym of Crabro denticornis F. Smith, 1879. Crossocerus annulipes hokkaiddoensis Tsuneki: Tsuneki, 1982: 17 (in Crossocerus denticornis (Bischoff): Coulon, 1925: 123 (new combi- the list of the species of the Sphecidae known from the Korean nation); Tsuneki, 1974: 380 (N Korea: “Prov. South Pyongan: Peninsula); Paik, 1985: 201 (in the list of Sphecidae of Korea); Pyongyan, Nung-ra do, Chang-lyong san, Guk-san-bong”); Leclercq, 2009: 164 (in the subgenus Ablepharipus, including Tsuneki, 1982: 7 (additional description, N Korea: “Prov. South Korea in distribution). Pyonggan: Nampo”), 17 (in the list of the species of the Sphe- Crossocerus annulipes Lepeletier et Brullé: Nemkov in Nemkov et al., cidae known from the Korean Peninsula); Paik, 1985: 201 (in the 1995: 436 (including Korea in distribution); Nemkov, 2009:100 list of Sphecidae of Korea); Tsuneki, 1991: 203 (in Sphecidae (including Korea in distribution); Nemkov, 2012b:441 from Korea). (including Korea in distribution). Crabro denticoxa Bischoff, 1932:44,_ (holotype), Germany: Baden- Distribution. N Korea (PN). Württemberg: Wasenweiler near Kaiserstuhl [ZMHU], synony- mized with Crossocerus denticornis Gussakovskij by Marshakov, 426 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

1980:356;Nemkov, 2012b: 442 (including Korea in 501 (including Korea in distribution); Nemkov, 2009:105 distribution). (including Korea in distribution); Lee et al., 2010: 210 (in Crossocerus denticoxa (Bischoff): Leclercq, 1954: 226 (new combi- Checklist of Korean Insects); Nemkov, 2012b: 442 (including nation); Nemkov in Nemkov et al., 1995: 431 and 438 (in key, Korea in distribution). including Korea in distribution); Nemkov, 2009: 104 (including Crabro pacificus Gussakovskij, 1932: 25, _ (holotype), Russia: Vla- Korea in distribution). divostok [ZIN], synonymized with Crabro emarginataus by Distribution. Transpalearctic: N Korea (PN), Europe to Russian Tsuneki, 1947b: 432; Tsuneki, 1947a: 290 (new to Korea, Korea: Far East, south to India. “Keijo”). Distribution. Korean Peninsula (PN, SL), Mongolia, China, Russian Crossocerus denticrus Herrich-Schaeffer, 1841 잔니은주둥이벌 Far East, Japan. Crossocerus denticrus Herrich-Schaeffer, 1841b: fasc. 181: plate 20, Germany; Tsuneki, 1954b: 64 and 73 (including Korea in dis- Crossocerus exiguus (Linden, 1829) 짧은은주둥이벌 tribution); Tsuneki, 1959b: 71 and 76 (including Korea in dis- Crabro exiguus Linden, 1829:74,\ (holotype or syntypes), Belgium: tribution); Kim, 1970: 638 (diagnosis, no Korean material), 822 Bruxelles [IRSNB]. (in the list of Hymenoptera from Korea); Tsuneki, 1974: 381e Crossocerus exiguus (Linden): Lepeletier et Brullé, 1835:791 (new 382 (N Korea: “Prov. South Pyongan: Pyongyan”); Tsuneki, combination); Tsuneki, 1974: 381 (N Korea: “Prov. South Pyon- 1982: 17 (in the list of the species of the Sphecidae known gan: Pyongyan; Kaesong: Mts. Pakyon”); Tsuneki, 1982: 17 (in from the Korean Peninsula); Paik, 1985:201(inthelistof the list of the species of the Sphecidae known from the Korean Sphecidae of Korea); Kim et al., 1994:264(inCheckListof Peninsula); Paik, 1985: 201 (in the list of Sphecidae of Korea); Insects from Korea); Nemkov in Nemkov et al., 1995:432(in Nemkov in Nemkov et al., 1995: 432 and 439 (in key, including key, including Korea in distribution); Nemkov, 2009:105 Korea in distribution); Nemkov, 2009: 106 (including Korea in (including Korea in distribution); Lee et al., 2010:210(in distribution); Nemkov, 2012b: 442 (including Korea in Checklist of Korean Insects); Nemkov, 2012b: 442 (including distribution). Korea in distribution). Distribution. Transpalearctic: N Korea (PN, GG), Europe to Crabro denticrus Herrich-Schaffer: Yasumatsu, 1942a: 90 [N Korea: Mongolia, NE China, Russian Far East. “Mozan” (Musan, Hamgyeongbuk-do)]; Tsuneki, 1947a:291e 292 (in Crabronidae fauna of Korea). Crossocerus hakusanus Tsuneki, 1954 Distribution. Transpalearctic: N Korea (HB, PN), N Africa, Europe Crossocerus hakusanus Tsuneki, 1954b:74,_ (holotype), Japan: to Mongolia, Russian Far East, Taiwan. Ishikawa Prefecture: Shiramine-mura (MNHAH); Bohart and Menke, 1976: 401 (including Korea in distribution); Yamane in Crossocerus dimidiatus sapporensis (Kohl, 1915) 삿뽀로은주둥이 Yamane et al., 1999: 501 (including Korea in distribution). 벌 Distribution. ?Korea, N China, Russian Far East, Japan, Taiwan. Crabro serripes sapporoensis Kohl, 1915:213,\ (holotype), Japan: Note. Both of two books (Bohart and Menke 1976; Yamane et al., Sapporo [NMNHA]. 1999) reporting the Korean occurrence of this species had no proof Crossocerus dimidiatus sapporoensis (Kohl): Tsuneki, 1954b: 59 and that they were based on examination of Korean material, although 68 (in key, new combination); Tsuneki, 1959b: 68 and 73 this species is known to be broadly distributed in eastern Asia. It is (including Korea in distribution); Yasumatsu, 1965: 302 currently more reasonable to exclude this species from the Korean (including Korea in distribution); Kim, 1970: 637 (diagnosis, no fauna. Korean material), 822 (in the list of Hymenoptera from Korea); Kim et al., 1994: 264 (in Check List of Insects from Korea); Crossocerus heydeni Kohl, 1880 헤이든은주둥이벌(신칭) Yamane, 2008: 554 (including Korea in distribution); Leclercq, Crossocerus heydeni Kohl, 1880: 216 (as Heydeni, incorrect original 2009: 168 (including Korea in distribution); Lee et al., 2010: capitalization), _ (syntypes), Austria: Tirol: Bozen area, now 210 (in Checklist of Korean Insects). Italy: Alto Adige: St. Gries and Kollern near Bolzano [NHMW]; Crossocerus dimidiatus Fabricius: Tsuneki, 1947: 289e290 (new to Tsuneki, 1982:8e9 (N Korea: “Prov. Ryang gang: Chan-Pay Korea, Korea: “Kodai-san”, “Keijo”, “Nanseitsu-rei”); Nemkov Plateau”), 17 (in the list of the species of the Sphecidae known in Nemkov et al., 1995: 430 and 437 (in key, including Korea in from the Korean Peninsula); Paik, 1985: 201 (in the list of distribution); Nemkov, 2009: 109 (including Korea in distri- Sphecidae of Korea); Nemkov in Nemkov et al., 1995: 435 and bution); Nemkov, 2012b: 442 (including Korea in distribution). 441 (in key, including Korea in distribution); Nemkov, 2009:102 Distribution. Korean Peninsula (HB, SL, GW), Japan. (including Korea in distribution); Nemkov, 2012b:441 (including Korea in distribution). Crossocerus emarginatus (Kohl, 1899) 우수리은주둥이벌 Distribution. Transpalearctic: N Korea (RG), Europe to Russian Crabro emarginatus Kohl, 1899: 307, _ (holotype or syntypes), Far East, Japan. Mongolia: northern Mongolia: no specific locality [NHMW]. Crossocerus emarginatus (Kohl): Tsuneki, 1952b: 69 (new combi- Crossocerus malaisei (Gussakovskij, 1932) 아이노은주둥이벌 nation); Tsuneki, 1954b: 64 and 73 (including Korea in distri- Crabro malaisei Gussakovskij, 1932:21,\ (holotype), Russia: Pri- bution); Tsuneki, 1959b: 71 and 76 (including Korea in morie Krai: Sutshan [NRS]. distribution); Kim, 1970: 638e639 (diagnosis, no Korean mate- Crossocerus malaisei (Gussakovskij): Leclercq, 1954: 241 (new rial), 822 (in the list of Hymenoptera from Korea); Tsuneki, 1974: combination); Nemkov in Nemkov et al., 1995: 430 (in key, 381 (N Korea: “Prov. South Pyongan: Lyong-ak san”); Tsuneki, including Korea in distribution); Nemkov, 2009: 100 (including 1982: 17 (in the list of the species of the Sphecidae known Korea in distribution). from the Korean Peninsula); Paik, 1985: 201 (in the list of Crabro aino Tsuneki, 1947b:413,\ (holotype), Japan: Hokkaido: Sphecidae of Korea); Tsuneki, 1991: 203 (in the list of Sphecidae Jozankei [HMHAN], synonymized with Crossocerus malaisei by of Korea); Kim et al., 1994: 264 (in Check List of Insects from Marshakov, 1979: 93; Tsuneki, 1954b: 58 and 67 (in key, Korea); Nemkov in Nemkov et al., 1995: 432 and 439 (in key, including Korea in distribution); Tsuneki, 1959b: 68 and 73 (in including Korea in distribution); Yamane in Yamane et al., 1999: key, including Korea in distribution). JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 427

Crossocerus aino (Tsuneki): Tsuneki, 1952b: 69 (new combination); Crossocerus styrius (Kohl): Leclercq, 1954: 233 (new combination). Tsuneki, 1957b: 61 (Korea: “Berg Gongo”); Kim, 1970: 634 Crabro pauxillus Gussakovskij, 1932: 22, \ (holotype), Russia: Pri- (diagnosis, no Korean material), 821 (in the list of Hymenoptera morskiy Kray: Sedanka [OMNH], synonymized with Crossocerus from Korea); Tsuneki, 1982: 17 (in the list of the species of the styrius by Bitsch and Leclercq, 1993:113. Sphecidae known from the Korean Peninsula); Paik, 1985:201 Crossocerus pauxillus (Gussakovskij): Leclercq, 1954: 235 (new (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Check combination); Tsuneki, 1968b: 54 (Korea: Jeju Is.: Mt. Hanna); List of Insects from Korea); Lee et al., 2010: 210 (in Checklist of Kim,1970: 636 (diagnosis, no Korean material), 821 (in the list of Korean Insects). Hymenoptera from Korea); Tsuneki, 1973b: 126 (three speci- Distribution. N Korea (GW), Russian Far East, Japan. mens from Korea without specific localities); Paik, 1985: 202 (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Check Crossocerus palmipes chosensis Tsuneki, 1957 마당발은주둥이벌 List of Insects from Korea); Lee et al., 2010: 210 (in Checklist of Crossocerus palmipes chosensensis Tsuneki, 1957b: 62, \ (holotype) Korean Insects). and _, central Korea: “Shinshu” [KUK]; Tsuneki, 1959b:70and Distribution. Transpalearctic: S Korea (JJ), Europe to Russian Far 76 (including Korea in distribution); Kim, 1970: 639, 822 (in East, Japan. the list of Hymenoptera from Korea); Bohart and Menke, 1976: 402 (including Korea in distribution); Tsuneki, 1982:17(inthe Crossocerus takeuchii Tsuneki, 1957 타케우치은주둥이벌 list of the species of the Sphecidae known from the Korean Crossocerus takeuchii Tsuneki, 1957: 61e62, \ (holotype), N Korea: Peninsula); Paik, 1985: 201 (in the list of Sphecidae of Korea); “Sanyodai” (Sanyangdae, Ryanggang-do) [KUK]; Tsuneki, 1959: Kim et al., 1994: 264 (in Check List of Insects from Korea); 74e75 (including Korea in distribution); Kim, 1970: 636e637 Nemkov in Nemkov et al., 1995: 431 and 438 (in key, including (diagnosis, no Korean material), 821 and 822 (in the list of Hy- Korea in distribution); Leclercq, 2009:161,178(inthesubge- menoptera from Korea); Tsuneki, 1982: 17 (in the list of the nus Crossocerus); Nemkov, 2009: 106 (including Korea in dis- species of the Sphecidae known from the Korean Peninsula); tribution); Lee et al., 2010: 210 (no subspecific recognition, in Paik, 1985: 201 (in the list of Sphecidae of Korea); Kim et al., Checklist of Korean Insects); Nemkov, 2012b: 442 (including 1994: 264 (in Check List of Insects from Korea); Nemkov in Korea in distribution). Nemkov et al., 1995: 435 and 441 (in key, including Korea in Distribution. Korea (“Shinshu), Russian Far East. distribution); Nemkov, 2009: 104 (including Korea in distribu- tion); Lee et al., 2010: 210 (in Checklist of Korean Insects); Crossocerus podagricus hokusenensis Tsuneki, 1974 유럽은주둥 Nemkov, 2012b: 441 (including Korea in distribution). 이벌 Distribution. N Korea (RG), Russian Far East, Japan. Crossocerus podagricus hokusenensis Tsuneki, 1974, \ (holotype), N Korea: “Prov. Ryang-gang: Chann-Pay Plateau: Sam-zi-yan” Crossocerus tarsatus (Shuckard, 1837) 슈카아드은주둥이벌: [TMB]: Bohart and Menke, 1976: 629 (including Korea in dis- Crabro tarsatus Shuckard, 1837: 133, _ (holotype or syntypes), tribution); Tsuneki, 1982: 17 (in the list of the species of the Britain: Highgate (depository unknown). Sphecidae known from the Korean Peninsula); Paik, 1985:201 Crossocerus tarsatus (Shuckard): Richards, 1937b: 106 (new com- (in the list of Sphecidae of Korea); Lee et al., 2010:210(in bination); Tsuneki, 1957b: 63 and 72 (including Korea in dis- Checklist of Korean Insects). tribution); Tsuneki, 1957b: 62 (N Korea: “Sanyodai”); Tsuneki, Crossocerus podagricus (Linden): Tsuneki, 1947a: 290 (new to Korea, 1959b: 70 and 76 (including Korea in distribution); Kim, 1970: N Korea: “Nanseitsu-rei”); Tsuneki, 1954b: 59 and 69 (including 638, 822 (in the list of Hymenoptera from Korea); Paik, 1985: Korea in distribution); Tsuneki, 1959b: 68 and 74 (including 201 (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Korea in distribution); Kim, 1970: 634 (diagnosis, no Korean Check List of Insects from Korea); Nemkov in Nemkov et al., material), 822 (listed as podagticus, in the list of Hymenoptera 1995: 431 (in key, including Korea in distribution); Nemkov, from Korea); Kim et al., 1994: 264 (in Check List of Insects from 2009: 107 (including Korea in distribution); Lee et al., 2010: Korea); Nemkov in Nemkov et al., 1995: 434 and 437 (in key, 210; Nemkov, 2012b: 442 (including Korea in distribution). including Korea in distribution); Nemkov, 2009: 99 (including Distribution. Holarctic: N Korea (RG), N Africa, Europe to Russian Korea in distribution). Far East, Canada, USA. Distribution. N Korea (RG, HB). Crossocerus vagabundus koreanus Tsuneki, 1957 나그네은주둥 Crossocerus pseudopalmarius (Gussakovskij, 1932) 넓적발은주 이벌 둥이벌 Crossocerus vagabundus koreanus Tsuneki, 1957b:61,_ (holotype) Crabro pseudopalmarius Gussakovskij, 1932:26,\ (holotype), and \, S Korea: “Zokurisan” [KUK]; Kim, 1970:821(inthelistof Russia: Primorskiy Kray: Sutshan [NRS]. Hymenoptera from Korea); Bohart and Menke, 1976:403 Crossocerus pseudopalmarius (Gussakovskij): Leclercq, 1954: 228 (including Korea in distribution); Tsuneki, 1982:17(inthelist (new combination); Tsuneki, 1974: 379 (new to Korea, N Korea: of the species of the Sphecidae known from the Korean “Prov. South Pyongan: Pyongyan”); Tsuneki, 1982: 17 (in the list Peninsula); Kim et al., 1994: 264 (in Check List of Insects from of the species of the Sphecidae known from the Korean Penin- Korea); Li and He, 2005: 160 (including Korea in distribution); sula); Paik, 1985: 201 (in the list of Sphecidae of Korea); Nemkov Leclercq, 2009: 187 (including Korea in distribution); Lee et al., in Nemkov et al., 1995: 430 and 436 (in key, including Korea in 2010: 210 (in Checklist of Korean Insects). distribution); Leclercq, 2009: 180 (including Korea in distribu- Croossocerus vagabudus (Panzer): Tsuneki, 1976a:78(“N Korea: tion); Nemkov, 2009: 110 (including Korea in distribution); Kan-nan: Hutempo: Taitimpyoo”); Nemkov in Nemkov et al., Nemkov, 2012b: 442 (including Korea in distribution). 1995: 434 and 437 (in key, including Korea in distribution); Distribution. N Korea (PN), NE China, Russian Far East. Nemkov, 2009: 99 (including Korea in distribution); Nemkov, 2012b: 441 (including Korea in distribution). Crossocerus styrius (Kohl, 1892) 꼬마은주둥이벌 Crabro vagabundus Panzer f. quadricinctus Tsuneki, 1947a: 289, \, Crabro styrius Kohl, 1892: 198, \ (holotype), Austria: Steiermark: Korea: “Keijo”, corrected to Crossocerus vagabundus koreanus by Admont [NHMV]. Tsuneki 1957b:61;Kim, 1970: Pl. 63, Figure 764, 635 (diagnosis, 428 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

S Korea: Mt. Taebaeksan), 821 (in the list of Hymenoptera from distribution); Lee et al., 2010: 210 (in Checklist of Korean In- Korea). sects); Nemkov, 2012b: 442 (including Korea in distribution). Crossocerus vagabundus yamamotonis Tsuneki, 1947: 412: Tsuneki, Distribution. S Korea (SL, GW), Russian Far East, Japan. 1954b: 58 (including Korea in distribution); Tsuneki, 1959b:67 and 72 (including Korea in distribution); Yasumatsu, 1965: 302 Genus Ectemnius Dahlbom, 1945 은주둥이벌 (including Korea in distribution); Kim, 1970: 635 (in Note), 821 Ectemnius Dahlbom, 1945: 389. Type species: “E. guttatus, Dahlb.” (in the list of Hymenoptera from Korea); Yamane, 2008: 562 (¼ Crabro guttatus of Dahlbom, 1845 ¼ Crabro guttatus Linden, (including Korea in distribution). 1829), designated by Ashmead, 1899:172. Corenocrabro ectemiformis Tsuneki, 1974a: 374e375, _ (holotype) and \, N Korea: Prv. Suth Pyongan: Pyongyan [TMB], synony- Ectemnius borealis (Zetterstedt, 1839) 먹은주둥이벌 mized with Crossocerus vagabundus by Marshakov, 1979a: 96; Crabro borealis Zetterstedt, 1839: 443, _ (syntypes), Norway: Bohart and Menke, 1976:629 (including Korea in distribution); Alteidet, Altengaard, and Bossekop; Finland or Sweden: Calix on Tsuneki, 1991: 202 (as new species, nomen nudum). Botnia Gulf [MZLU]. Distribution. Korean Peninsula (HN, PN, SL, GW, CB). Ectemnius borealis (Zetterstedt): Leclercq, 1974b: 279 (new combi- nation); Tsuneki,1982: 7 (N Korea: “Prov. Ryang-gang: Chan-pay Crossocerus varus Lepeletier and Brullé, 1835 별난은주둥이벌 Plateau”), 16 (in the list of the species of the Sphecidae known Crossocerus varus Lepeletier and Brullé, 1835: 775, \ and _, junior from the Korean Peninsula); Paik, 1985: 200 (in the list of secondary homonym of Crossocerus varus (Panzer, 1799), but Sphecidae of Korea); Nemkov in Nemkov et al.,1995: 447 (in key, valid under Article 23.9.1 of the Code (nomen protectum), including Korea in distribution); Leclercq, 1999: 46 (including Syntypes: France [coll. Spinola, Torino]; Tsuneki, 1974: 381 Korea in distribution); Yeo, 2009: 9 (in key), 12e13 (N Korea: (specific name listed incorrectly as varius, N Korea: “Prov. Ryang- Ryanggang-do: Mt. Baekdusan); Nemkov, 2009: 119 (including gang: Chan-pay Plateau: Sam-zi-yan”); Tsuneki, 1982: 17 (in the Korea in distribution); Nemkov, 2012b: 443 (including Korea in list of the species of the Sphecidae known from the Korean distribution). Peninsula); Paik, 1985: 201 (in the list of Sphecidae of Korea); Crabro nigrinus Herrich-Schaeffer, 1841: 15, Pl. 15, synonymized Nemkov in Nemkov et al., 1995: 431 (in key, including Korea in with Ectemnius borealis by Leclercq, 1974b: 280: Tsuneki, 1947a: distribution); Nemkov, 2009: 108 (including Korea in 284e285 (new to Korea, N Korea: “Nansetsu-rei, Daitaku, Kus- distribution). sho”); Tsuneki, 1958a: 12 (including Korea in distribution). Crossocerus pusillus Lepeletier and Brullé, 1835: 778, _ (holotype), Ectemnius nigrinus (Herrich-Schaeffer): Tsuneki, 1957b:60(NKo- France: Landes: Saint Sever [coll. Spinola, Torino], Synonymized rea: “Sanyodai”); Kim, 1970: 644 (listed with distribution), 820 with Crossocerus varus by Bitsch and Leclercq, 1993:297:Kim (in the list of Hymenoptera from Korea); Kim et al., 1994: 265 (in et al., 1994: 264 (in Check List of Insects from Korea); Lee Check List of Insects from Korea); Lee et al., 2010: 211 (in et al., 2010:210;Nemkov, 2012b: 442 (including Korea in Checklist of Korean Insects). distribution). Distribution. Holarctic: N Korea (RG), Europe to Mongolia, Distribution. Transpalearctic: N Korea (RG), N Africa, Europe to Russian Far East, Japan, Canada, USA. Mongolia, Russian Far East, Japan. Ectemnius cavifrons (Thomson, 1870) 오목이마은주둥이벌 Crossocerus wesmaeli parvicorpus Tsuneki, 1974 웨스멜은주둥이 Crabro cavifrons Thomson, 1870:173,\ and _ (syntypes), Sweden: 벌 no specific locality [MZLU]; Tsuneki, 1943a: 156 [N Korea: Crossocerus wesmaeli parvicorpus Tsuneki, 1974: 381, \ (holotype) “Hakutosan” (Mt. Baekdusan, Ryanggang-do)]; Tsuneki, 1947a: and _, North Korea: “Prov. South Pyongan: Pyongyan” [TMB]; 283 (Korea: “Hakutosan, Kodai-san, Nansetu-rei”). Bohart and Menke, 1976: 629 (including Korea in distribution); Ectemnius cavifrons (Tomson): Leclercq, 1949a: 13 (new combina- Tsuneki, 1982: 17 (in the list of the species of the Sphecidae tion); Tsuneki, 1957b: 59 [N Korea: “Mozanrei (Musanryeong, known from the Korean Peninsula); Paik, 1985: 202 (in the list of Hamgyeongbuk-do), Berg Gongo”]; Tsuneki, 1958a:13 Sphecidae of Korea). (including Korea in distribution); Kim, 1970: 644 (listed with Crossocerus wesmaeli (Linden): Nemkov in Nemkov et al., 1995: distribution), 819 (in the list of Hymenoptera from Korea); 431e432 (in key, including Korea in distribution); Nemkov, Tsuneki, 1974: 374 (N Korea: “Prov. Ryang-gang: Chan-Pay 2009: 108 (including Korea in distribution); Nemkov, 2012b: Plateau: Sam-zi-yan”); Tsuneki, 1982: 16 (in the list of the spe- 442 (including Korea in distribution). cies of the Sphecidae known from the Korean Peninsula); Paik, Distribution. N Korea (PN). 1985: 200 (in the list of Sphecidae of Korea); Tsuneki, 1991: 202 (in Sphecidae from Korea); Kim et al., 1994: 265 (specific Crossocerus yanoi (Tsuneki) 야노은주둥이벌 name incorrectly as clavifrons, in Check List of Insects from Ko- Crabro yanoi Tsuneki, 1947b: 409, _ (holotype) and _, Japan: Hok- rea); Nemkov in Nemkov et al., 1995: 450 (in key, including kaido: Sapporo [MNHAH]. Korea in distribution); Leclercq, 1999: 46 (including Korea in Crossocerus yanoi (Tsuneki): Tsuneki, 1952b: 68 (new combination); distribution); Yeo, 2009: 11 (in key), 14e17 (the Korean Penin- Tsuneki, 1954b: 59 and 67 (including Korea in distribution); sula: Ryanggang-do, Gangwon-do, Gyeonggi-do, Seoul, Incheon Tsuneki, 1957b: 61 [Korea: “Kodaisan” (Odaesan, Gangwon-do), Metropolitan City, Chungcheongbuk-do, Chungcheongnam-do, “Keijo”]; Tsuneki, 1959b: 68 and 73 (including Korea in distri- Gyeongsangbuk-do, Gyeongsangnam-do, Jeollanam-do, Jeju Is.); bution); Kim, 1970: 637 (diagnosis, no Korean material), 821 (in Nemkov, 2009: 117 (including Korea in distribution); Lee et al., the list of Hymenoptera from Korea); Tsuneki, 1982: 17 (in the 2010: 210 (in Checklist of Korean Insects); Nemkov, 2012b: list of the species of the Sphecidae known from the Korean 443 (including Korea in distribution). Peninsula); Kim et al., 1994: 264 (in Check List of Insects from Distribution. Transpalearctic: Korean Peninsula (RG, HB, GW, SL, Korea); Nemkov in Nemkov et al., 1995: 430 and 437 (in key, GG, IC, CB, CN, GB, GN, JN, JJ), Europe to Mongolia, China, Russian Far including Korea in distribution); Leclercq, 2009: 188 (including East, Japan. Korea in distribution); Nemkov, 2009: 110 (including Korea in JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 429

Ectemnius chrysites (Kohl, 1892) 노란줄은주둥이벌 Check List of Insects from Korea); Nemkov in Nemkov et al., Crabro chrysites Kohl, 1892,197,\ (holotype or syntypes), Sikkim: 1995: 451 (in key, including Korea in distribution); Yeo, 2009: no specific locality [NHMW]; Iwata, 1934: 8 [Korea: “Onsenji” 11 (in key), 19e22 (the Korean Peninsula: Ryanggang-do, (Onjeongsa, Gangwon-do), “Hosenji” (Bongseonsa, ? Gyeonggi- Gyeonggi-do); Nemkov, 2009: 119 (including Korea in distri- do), “Koryo” (Gwangleung)]; Tsuneki, 1947a: 283 (in Crabroni- bution); Lee et al., 2010: 211 (in Checklist of Korean Insects); dae fauna of Korea). Nemkov, 2012b: 443 (including Korea in distribution). Ectemnius chrysites (Kohl): Leclercq, 1954: 288 (new combination); Solenius vagus Linnaeus: Iwata, 1933: 11 (Korea: no specific local- Tsuneki, 1957b: 59 (Korea: “Gyokusenji”); Tsuneki, 1958a:15 ity), misidentification, misidentified references list in Pulawski (include Korea in distribution); Kim, 1970: 644 (listed with (2014). Sphex vagus Linnaeus is a junior synonym of Mellinus distribution), 820 (in the list of Hymenoptera from Korea); arvensis. Bohart and Menke, 1976: 425 (including Korea in distribution); Crabro vagus (Linnaeus): Tsuneki, 1943a: 156 (N Korea: “Hakuto- Nemkov in Nemkov et al., 1995: 449 (in key, including Korea in zan”) misidentification, misidentified references list in Pulawski distribution); Yeo, 2009: 10 (in key), 17e19 (the Korean Penin- (2014). Sphex vagus Linnaeus is a junior synonym of Mellinus sula: Pyeongannam-do: Pyeongyang, Gyeonggi-do, Seoul, arvensis. Chungcheongbuk-do, Gyeongsangbuk-do, Gyeongsangnam-do, Distribution. Holarctic: Korean Peninsula (RG, GW, GG, GB, JJ), Jeollanam-do, Jeju Is.); Nemkov, 2009: 121 (including Korea in Europe to Mongolia, China, Russian Far East, Japan; Canada, USA, distribution); Nemkov, 2012b: 443 (including Korea in Cuba, Guatemala, Venezuela. distribution). Ectemnius chrysites chosenensis Tsuneki, 1974:372,\ (holotype), N Ectemnius dives (Lepeletier et Brullé, 1835) 북방목대장은주둥이 Korea: Kaesong: Mount Pakyon, 27 Km NE Kaesong; N Korea: 벌, 유럽은주둥이벌 “Prov. South Pyongan: Pyongyan, Za-mo san” (paratypes) [TMB], Solenius dives Lepeletier et Brullé, 1835: 716, \ and _ (syntypes), synonymized with nominotypical subspecies by Yeo, 2009:18; France: Paris (lost), \ (neotype designated by Leclercq, 1974d: Tsuneki, 1982: 16 (in the list of the species of the Sphecidae 280), France: Versailles [MNHN]. known from the Korean Peninsula); Paik, 1985: 200 (in the list of Ectemnius dives Lepeletier et Brullé: Kirchner, 1854: 313 (new Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of Insects combination); Tsuneki, 1947: 285 (new to Korea, N Korea: from Korea); Lee et al., 2010: 211 (in Checklist of Korean Insects). “Nansetsu-rei”); Tsuneki, 1958a: 12 (including Korea in distri- Ectemnius (Metacrabro) ssp. n.: Tsuneki, 1991: 202, Korea: Mount bution); Kim, 1970: 644 (listed with distribution), 821 (in the list Pakyon 27 km NE Kaesong. of Hymenoptera from Korea); Tsuneki, 1982: 7 (N Korea: Prov. Distribution. Korean Peninsula (PN, GG, SL, CB, GB, GN, JN, JJ), Ryong-gang: River Karim), 16 (in the list of the species of the Russian Far East, Japan; Taiwan, India, South East Asia. Sphecidae known from the Korean Peninsula); Paik, 1985:200 (in the list of Sphecidae of Korea); Kim et al., 1994: 265 (in Check Ectemnius confinis (Walker, 1871) 윤은주둥이벌 List of Insects from Korea); Nemkov in Nemkov et al., 1999: 446 Crabro confinis Walker, 1871: 27, _ and \ (syntypes), Egypt: Cairo (in key, including Korea in distribution); Yeo, 2009: 9 (in key), [lost], _ (neotype) Egypt: no specific locality [MNHN], desig- 22e23 (the Korean Peninsula: Ryanggang-do, Gangwon-do); nated by Leclercq, 1974d: 276. Nemkov, 2009: 119 (including Korea in distribution); Nemkov, Ectemnius confinis (Walker): Leclercq, 1975b: 3 (new combination); 2012b: 443 (including Korea in distribution). Tsuneki, 1982: 16 (as confinus in the list of the species of the Distribution. Holarctic: Korean Peninsula (RG, GW), Europe to Sphecidae known from the Korean Peninsula); Paik, 1985:200 Russian Far East, Japan, Canada, USA. (in the list of Sphecidae of Korea); Yamane in Yamane et al., 1999: 505 (including Korea in distribution). Ectemnius fossorius manchurianus Tsuneki, 1976 코노우은주둥 Crabro laevigatus De Stefani, 1884, synonymized with Ectemnius 이벌 confinis by Leclercq, 1974b:276:Yasumatsu, 1942a: 88 [N Korea: Sphex fossorius Linnaeus, 1758: 571 (as fossoria, incorrect original “Koka-islet: Onsei-ri” (Onjeong-ri, Gangwon-do)]; Tsuneki, termination), \ (holotype), Europe: no specificlocality[LS]. 1947: 284 (in Crabro fauna of Korea); Kim, 1970: 640 (listed with Ectemnius fossorius (Linnaeus): Leclercq, 1949a: 17 (new combina- distribution). tion); Tsuneki, 1982: 16 (in the list of the species of the Sphe- Distribution. Transpalearctic: N Korea (GW), N Africa, Europe to cidae known from the Korean Peninsula); Paik, 1985: 200 (in the Russian Far East, Japan. list of Sphecidae of Korea); Nemkov in Nemkov et al., 1995:449 (in key, including Korea in distribution); Yeo, 2009: 10 (in key), Ectemnius continuus (Fabricius, 1804) 은주둥이벌 23e26 (Korea: Gangwon-do); Nemkov, 2009:121e122 Crabro continuus Fabricius, 1804:312,\ (holotype), Morocco: (including Korea in distribution); Nemkov, 2012b:443 Tanger [ZMUC]; Tsuneki, 1947a: 283e284 [N Korea: “Hakuto- (including Korea in distribution). zan”, “Daitaku” (Daetaek, Ranggang-do), “Nansetsu-rei”]. Crabro konowi Kohl, 1905: 218 (as Konowi, incorrect original capi- Ectemnius continuus (Fabricius): Pate, 1946: 10 (new combination); talization), _ (holotype or syntypes), Japan: Tokyo [NMNHA], Leclercq, 1949a: 11 (including Korea in distribution); Leclercq, synonymized with Crabro fossorius by Marshakov, 1979: 103; 1954: 269 (including Korea in distribution); Tsuneki, 1957b:60 Iwata, 1938: 81 (Korea: no specific locality); Tsuneki, 1947a: 282 (N Korea: “Hakugan, Berg Gongo”); Tsuneki, 1958a:17 (N Korea: “Hakuto-san, Nansetsu-rei, Daita-ku”). (including Korea in distribution); Kim, 1970: Pl. 64 Figure 778, Ectemnius konowii (Kohl): Leclercq, 1950: 10 (new combination); 641e642 (diagnosis, S Korea: Uidong), 820 (in the list of Hy- Tsuneki, 1957a: 77 [N Korea: “Kan-nan (Hamgyeongnam-do): menoptera from Korea); Tsuneki, 1974: 372 (N Korea: “Prov. Hutempo: Taitimpyoo”]; Tsuneki, 1957b: 59 [N Korea: “Tonai” Ryang-gang: Chan-Pay Plateau: Sam-zi yan”); Kim, 1980: Pl. xx (Donae, Ryanggnag-do)]; Tsuneki, 1958a: 15 (including Korea in HYCR 01, 147 (diagnosis, Gangwon-do, Gyeonggi-do, distribution); Yasumatsu, 1965: 302 (including Korea in distri- Gyeongsangbuk-do, Jeju Is.); Tsuneki, 1982: 7 (N Korea: “Prov. bution); Kim, 1970: Pl. 64 Figure 783, 643 (diagnosis, S korea: Ryang-gang: Chan-Pay Plateau”), 16 (in the list of the species of Naeseolak), 820 (in the list of Hymenoptera from Korea); Kim, the Sphecidae known from the Korean Peninsula); Paik, 1985: 1980: Pl. xx HYCR 02, 148 (diagnosis, Korea: Gyeonggi-do, 200 (in the list of Sphecidae of Korea); Kim et al., 1994: 265 (in Gyeongsangbuk-do); Kim et al., 1994: 265 (in Check List of 430 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Insects from Korea); Yamane, 2008: 561 (including Korea in 1982: 16 (in the list of the species of the Sphecidae known distribution); Lee et al., 2010: 211 (in Checklist of Korean Insects). from the Korean Peninsula); Paik, 1985: 200 (in the list of Ectemnius fossorius konowi (Kohl): Tsuneki, 1977b: 8 (new status); Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of In- Tsuneki, 1982: 16 (in the list of the species of the Sphecidae sects from Korea); Nemkov in Nemkov et al., 1995: 450 (in key, known from the Korean Peninsula); Paik, 1985: 200 (in the list of including Korea in distribution); Leclercq, 1999: 59 (including Sphecidae of Korea); Leclercq, 1999:54e55 (including Korea in Korea in distribution); Yeo, 2009: 11 (in key), 28e30 (new to S distribution). Korea, Korea: Ryangang-do, Gangwon-do); Nemkov, 2009:117 Ectemnius konwii manchurianus Tsuneki, 1976a: 299e300, \ (ho- (including Korea in distribution); Lee et al., 2010: 211 (in lotype), N China: Manchuria: Maershan [HMHAN] and N Korea: Checklist of Korean Insects); Nemkov, 2012b: 443 (including “Mt. Hakuto, Mt. Nanseitsu-rei, Diataku-Hills” (paratypes). Korea in distribution). Ectemnius fossorius manchurianus Tsuneki: Tsuneki, 1977b: 4 (new Distribution. Holarctic: Korean Peninsula (RG, GW), Europe to subspecific combination). Mongolia, China, Russian Far East, Japan, Canada, USA. Distribution. Korean Peninsula (RG, HN, GG, GW, GB), N China, Russian Far East. Ectemnius martjanowi (F. Morawit, 1892) 시베리아은주둥이벌 Crabro martjanowi F. Morawit, 1892: 177 (as Martjanowi, incorrect Ectemnius iridifrons (Pérez, 1905) 흰줄은주둥이벌 original capitalization), _ (holotype), Russia: Krasnoyarskiy Crabro iridifrons Pérez, 1905a: 154, \ (holotype or syntypes), Japan: Kray: Minussinsk [ZIN]; Tsuneki, 1947a: 282 (Korea: “Hakuto- no specific locality but presumably Tokyo area [MNHN]; Iwata, san, Nansetsu-rei, Daitaku”). 1934: 8 (N Korea: “Kongo-san”); Iwata, 1938: 81-82 (including Ectemnius martjanowi (F. Morawitz): Leclercq, 1954: 287 (new Korea in distribution); Tsuneki, 1947a: 282 (in Crabronidae combination); Tsuneki, 1956c:128e131 (discussion with N fauna of Korea); Kim et al., 1994: 264 (in Check List of Insects Korean material); Tsuneki, 1957a: 77 (N Korea: “Kan-nan: from Korea); Lee et al., 2010: 210 (in Checklist of Korean Insects). Hutempo: Taitimpyoo”); Tsuneki, 1957b: 59 (N Korea: “Haku- Ectemnius iridifrons (Pérez): Tsuneki, 1952b: 63 (new combination); gan”); Tsuneki, 1958a: 14 (including Korea in distribution); Kim, Leclercq, 1954: 289, 322 (including Korea in distribution); 1970: 644 (diagnosis, no Korean material), 821 (in the list of Tsuneki, 1958a: 15 (include Korea in distribution); Kim, 1970: Pl. Hymenoptera from Korea); Bohart and Menke, 1976:427 64 Figure 782, 643 (diagnosis, S korea: Mt. Gayasan), 819 (in the (including Korea in distribution); Tsuneki, 1982:6e7 (N Korea: list of Hymenoptera from Korea); Tsuneki, 1982: 16 (in the list of “Prov. Ryang-gang: Chan-Pay Plateau”), 16 (in the list of the the species of the Sphecidae known from the Korean Peninsula); species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 200 (in the list of Sphecidae of Korea); Kim et al., Paik, 1985: 200 (in the list of Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of Insects from Korea); Nemkov in 1994: 265 (in Check List of Insects from Korea); Nemkov in Nemkov et al., 1995: 449 (in key, including Korea in distribu- Nemkov et al., 1995: 448 (in key, including Korea in distribu- tion); Leclercq, 1999: 55 (including Korea in distribution); tion); Leclercq, 1999: 61 (including Korea in distribution); Yeo, Yamane in Yamane et al., 1999:504e505 (including Korea in 2009: 10 (in key), 30e32 (new to S Korea: the Korean penin- distribution); Yamane, 2008: 562 (including Korea in distribu- sula: Ryanggnag-do, Gangwon-do); Nemkov, 2009:122 tion); Yeo, 2009: 10 (in key), 26e27 (Korea: Gangwon-do, (including Korea in distribution); Lee et al., 2010: 211 (in Gyeonggi-do, Gyeongsanm-do); Nemkov, 2009: 122 (including Checklist of Korean Insects); Nemkov, 2012b: 443 (including Korea in distribution); Lee et al., 2010: 211 (in Checklist of Korea in distribution). Korean Insects); Nemkov, 2012b: 443 (including Korea in Distribution. Transpalearctic: Korean Peninsula (RG, GW), distribution). Europe to China, Russian Far East, Japan, south to India. Distribution. Korean Peninsula (GW, GG, GN), Russian Far East, Japan. Ectemnius nigritarsus (Herrich-Schaeffer, 1841) 검은발목은주 둥이벌 Ectemnius furuichii (Iwata, 1934) 후루이치은주둥이벌 Crabro nigritarsus Herrich-Schaeffer, 1841: 25, \ (holotype or Crabro furuichii Iwata, 1934:17,\ (holotype), Japan: Honshu: syntypes), Germany: no specific locality (depository un- Todoromi: Mount Hachifuse [KUK]; Tsuneki, 1947a: 283 (new to known); Tsuneki, 1947a: 283 (new to Korea, Korea: “Shoyo- Korea, Korea: “Shoyo-zan”). zan”). Ectemnius furuichii (Iwata): Leclercq, 1954: 284 (new combination); Ectemnius nigritarsus (Herrich-Schaeffer): Leclercq, 1949b:180 Tsuneki, 1960a: 241 (examination of one female Korean spec- (new combination); Tsuneki, 1958a: 14 (including Korea in imen); Tsuneki, 1982: 16 (in the list of the species of the Sphe- distribution); Kim, 1970: 640, 819 (in the list of Hymenoptera cidae known from the Korean Peninsula); Paik, 1985:200(inthe from Korea); Paik, 1985: 200 (in the list of Sphecidae of Korea); list of Sphecidae of Korea); Leclercq, 1999: 55 (including Korea in Kim et al., 1994: 265 (in Check List of Insects from Korea); Yeo, distribution); Lee et al., 2010: 211 (in Checklist of Korean Insects). 2009: 10 (in key), 32e34 (Korea: Seoul, Gyeonggi-do, Gyeong- Ectemnius furuichii formosanus Tsuneki: Kim, 1970: 642 (diagnosis, sangbuk-do, Gyeongsangnam-do); Nemkov, 2009:116 no Korean material), 821 (in the list of Hymenoptera from Ko- (including Korea in distribution); Lee et al., 2010: 211 (in rea); Kim et al., 1994: 265 (in Check List of Insects from Korea). Checklist of Korean Insects); Nemkov, 2012b: 442 (including Distribution. S Korea (GG), Japan. Korea in distribution). Distribution. Transpalearctic: S Korea (SL, GG, GB, GN), Europe to Ectemnius lapidaries (Panzer, 1803) 북극은주둥이벌 China, Russian Far East, Japan. Crabro lapidarius Panzer, 1803 (Heft 90): 12, _ (holotype or syn- types), Germany: no specific locality (depository uncertain). Ectemnius nitobei (Matsumura, 1912) 니토베은주둥이벌 Ectemnius lapidaries (Panzer): Pate, 1946: 2 (new combination); Crabro nitobei Matsumura, 1912:173,\ (holotype or syntypes, Tsuneki, 1957b: 59 (N Korea: “Tonai”); Tsuneki, 1958a:12 actually _), Japan: Aomori Prefecture (Hokkaido University): (including Korea in distribution); Kim, 1970: 644, 820 (in the list Kim, 1970: 645 (diagnosis, no Korean material), 823 (in the list of of Hymenoptera from Korea); Tsuneki, 1974: 374 (N Korea: Hymenoptera from Korea); Paik, 1985: 201 (in the list of Sphe- “Prov. Ryang-gang: Chan-Pay Plateau: Sam-zi-yan”); Tsuneki, cidae of Korea). JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 431

Ectemnius nitobei (Matsumura): Tsuneki, 1958b:156e160 (new Sweden: Muonioniska at 6757’N2342’E (depository combination; redescription of _, and description of \). uncertain). Lestica nitobei (Matsumura): Kim et al., 1994: 265 (in Check List of Ectemnius ruficornis (Zetterstedet): Leclercq, 1974d: 281 (new Insects from Korea); Lee et al., 2010: 211 (in Checklist of Korean combination); Tsuneki, 1982: 10, 16 (in the list of the species of Insects). the Sphecidae known from the Korean Peninsula); Paik, 1985: Distribution. ?Korea, Japan. 200 (in the list of Sphecidae of Korea); Nemkov in Nemkov et al., Note. Kim (1970) first recorded this species in Korea, but no 1995: 450 (in key, including Korea in distribution); Nemkov, Korean material was examined. His inclusion of this species in the 2009:117e118 (including Korea in distribution); Nemkov, Korean fauna was groundless. 2012b: 443 (including Korea in distribution). Ectemnius planifrons (Thomson), 1807: 173 (as Crabro), synony- Ectemnius pedicellaris (F. Morawit, 1889) 중국은주둥이벌 mized with Ectemnius nigrifrons by Pate, 1946c: 2: Tsuneki, Crabro pedicellaris F. Morawit,1889: 160, \ (lectotype, designated by 1947a: 283 (new to Korea, Korea: “Hakoto-san, Kodai-san, Marshakov, 1979: 105), China: Gansu: “Monasterium Dshoni” Nansetus-rei”); Tsuneki, 1958a: 13 (including Korea in distri- [ZIN]. bution); Yasumatsu, 1965: 302 (include Korea in distribution); Ectemnius pedicellaris (F. Morawitz): Marshakov, 1979: 105 (new Kim,1970: 641 (diagnosis, no Korean material), 820 (in the list of combination); Nemkov in Nemkov et al., 1995: 450 (in key, Hymenoptera from Korea); Kim et al., 1994: 265 (specific name including Korea in distribution); Yeo, 2009: 11 (in key), 34e35 incorrectly as flamifrons, in Check List of Insects from Korea); (N Korea: Pyeongannam-do: Pyeongyang: Mt. Changryongsan); Yamane, 2008: 554 (including Korea in distribution); Lee et al., Nemkov, 2009: 120 (including Korea in distribution); Nemkov, 2010: 211 (in Checklist of Korean Insects). 2012b: 443 (including Korea in distribution). Ectemnius nigrifrons (Cresson), synonymized with Ectemnius rufi- Crabro nielseni Kohl, 1915: 74 (as Nielseni, incorrect original capi- cornis by Leclercq, 1974b: 281, 260: Tsuneki, 1957b: 59 (N Korea: talization), \ (syntypes), China: Harning [BMNH] and Tsingtau “Tonai”); Kim, 1970: 644 (listed with Korean locality), 820 (in [ZMHU], synonymized with Crabro pedicellaris by Marshakov, the list of Hymenoptera from Korea). 1979: 105; Iwata, 1934: 10 [Korea: “Seiryo-ri” (Cheongryang-ri, Distribution. Holarctic: Korean Peninsula (RG, GW), Europe to Seoul)]; Tsuneki, 1947a: 284 (Korea: “Shoyozan”); Tsuneki, Mongolia, Russian Far East, Japan, Canada, USA. 1958a: 17 (including Korea in distribution). Ectemnius nielseni (Kohl): Leclercq, 1954: 271 (new combination); Ectemnius schlettereri schlettereri (Kohl, 1888) 쉴레테레은주둥 Tsuneki, 1957b: 60 (Korea: “Zokurisan”), “Berg Gaya”; Kim,1970: 이벌 641e642 (diagnosis, no Korean material), 820 (in the list of Crabro schlettereri Kohl, 1888a: 135 (as Schlettereri,incorrect Hymenoptera from Korea); Tsuneki, 1974: 374 (N Korea: “Prov. original capitalization), \ (holotype or syntypes), Austria: South Pyongan: Chang-lyong san, Bongwa-ri”); Bohart and Südirol: St. Pauls near Bozen, now Italy: Alto Adige: St. Pauls Menke, 1976: 427 (including Korea in distribution); Tsuneki, near Bolzano [NHMW]; Tsuneki, 1947a: 284 (new to Korea, 1982: 16 (in the list of the species of the Sphecidae known Korea: “Soyo-zan”, “Temma-san” (Mt. Cheonmasan, Gyeonggi- from the Korean Peninsula); Paik, 1985: 200 (in the list of do), “Kodai-san”). Sphecidae of Korea); Tsuneki, 1991: 202 (in Sphecidae from Ectemnius schlettereri (Kohl): Iwata, 1937: 130 (new combination); Korea); Kim et al., 1994: 265 (in Check List of Insects from Ko- Tsuneki, 1957b: 60 (Korea: Shutosu, Berg Kaya, Zokurisan); rea); Lee et al., 2010: 211 (in Checklist of Korean Insects). Tsuneki, 1958a: 17 (include Korea in distribution); Kim, 1970: Distribution. Korean Peninsula (PN, SL, CB, GB), Mongolia, NE 642 (diagnosis and line drawings, no Korean material), 820 (in China, Russian Far East. the list of Hymenoptera from Korea); Tsuneki, 1974:372(N Korea: “Prov. South Pyongan: Lyong-ak san”); Tsuneki, 1976a: Ectemnius radiatus (Pérez, 1905) 방사은주둥이벌 304 (including Korea in distribution); Tsuneki, 1982: 16 (in the Crabro radiatus Pérez, 1905: 155, _ (Holotype or syntypes), Japan: list of the species of the Sphecidae known from the Korean no specific locality but presumably Tokyo area [MNHN]. Peninsula); Paik, 1985: 200 (in the list of Sphecidae of Korea); Ectemnius radiates (Pérez):Leclercq, 1954: 289 (new combination); Tsuneki, 1991: 202; Kim et al., 1994: 265 (in Check List of Insects Tsuneki, 1958a: 14 (including Korea in distribution); Kim, 1970: from Korea); Nemkov in Nemkov et al., 1995: 451 (in key, 644 (listed with distribution), 819 (in the list of Hymenoptera including Korea in distribution); Yamane in Yamane et al., 1999: from Korea); Leclercq, 1982: 153 (Korea: no specific locality); 505 (including Korea in distribution); Yeo, 2009: 11 (in key), 36e Tsuneki, 1982: 16 (in the list of the species of the Sphecidae 38 (the Korean Peninsula: Pyeongannam-do, Seoul, Gangwon- known from the Korean Peninsula); Paik, 1985: 200 (in the list of do, Gyeonggi-do, Chungcheongbuk-do, Chungcheongnam-do, Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of In- Gyeongsangbuk-do, Jeollanam-do, Jeji Is.); Nemkov, 2009:121 sects from Korea); Yamane in Yamane et al.,1999: 506 (including (including Korea in distribution); Lee et al., 2010: 211 (in Korea in distribution); Nemkov, 2009: 116 (including Korea in Checklist of Korean Insects); Nemkov, 2012b: 443 (including distribution); Lee et al., 2010: 211 (in Checklist of Korean In- Korea in distribution). sects); Nemkov, 2012b: 443 (including Korea in distribution). Distribution. Transpalearctic: Korean Peninsula (PN, SL, GG, GW, Ectemnius mizuho Tsuneki, 1952b: 63, _ (holotype) and \, Japan, CB, CN, GB, JN, JJ), Europe to China, Russian Far East, Japan, south to Hokkaido: Sounkyo [HMHAN], synonymized with Ectemnius India. radiatus by Tsuneki, 1959c:7:Tsuneki, 1957b: 59 (N Korea: “Berg Gongo”). Ectemnius schlettereri horvatovichi (Kohl, 1974) 호르바토빗츠은 Ectemnius uadiatus Pérez: Kim et al., 1994: 265 (in Check List of 주둥이벌 Insects from Korea), maybe misspelled in specific name of Ectemnius horvatovichi Tsuneki, 1974:373,_ (holotype) and \,N radiatus. Korea: “Prov. South Pyongan: Pyongyan”; N Korea: “Prov. South Distribution. N Korea (GW), Russian Far East, Japan. Pyongan: Bongwa-ri” (paratype and others) [TMB]; Bohart and Menke, 1976: 629 (including Korea in distribution); Tsuneki, Ectemnius ruficornis (Zetterstedet, 1838) 따라지은주둥이벌 1982: 16 (in the list of the species of the Sphecidae known Crabro ruficornis Zetterstedt, 1838: 443, _ (holotype or syntypes), from the Korean Peninsula); Paik, 1985: 200 (in the list of 432 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Sphecidae of Korea); Tsuneki, 1991: 202 (as new species, nomen Lestica alata basakis F. Smith: Kim et al., 1994: 265 (in Check List of nudum). Insects from Korea). Ectemnius schlettereri horvatovichi Tsuneki: Tsuneki, 1977a: 10 (new Distribution. Transpalearctic: Korean Peninsula (RG, PN, GG, GW, status); Leclercq, 1999: 72 (including Korea in distribution). IC, JN), Europe to Mongolia, China, Russian Far East, Japan. Distribution. N Korea (PN). Lestica camelus (Eversmann, 1849) 목대장은주둥이벌 Ectemnius shimoyamai Tsuneki, 1958 시모야마은주둥이벌 Crabro camelus Eversmann, 1849: 424, _ (syntypes), Russia: Ural Ectemnius shimoyamai Tsuneki, 1958b:198e199, \ (holotype), foothills: no specific locality [ZIN]; Tsuneki,1943a: 156 (N Korea: Japan: Honshu: Aomori Prefecture: Towada [HMHAN]; Kim, “Hakuto-san”); Tsuneki, 1947a: 285 (N Korea: “Hakuto-san, 1970: Pl. 92 Figure 784, 643e644 (diagnosis, no Korean mate- Nansetsu-rei”). rial), 821 (in the list of Hymenoptera from Korea); Paik, 1985: Lestica camelus (Eversmann): Leclercq, 1954: 295 (new combina- 200 (in the list of Sphecidae of Korea); Paik, 1985: 200 (in the list tion); Tsuneki, 1957a: 77 (N Korea: “Kan-nan: Hutempo: Tai- of Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of timpyoo”); Tsuneki, 1957b: 60 (N Korea: “Sanyodai”); Kim, 1970: Insects from Korea); Lee et al., 2010: 211 (in Checklist of Korean Pl. 92 Figure 788, 646 (diagnosis, no Korean material), 823 (in Insects). the list of Hymenoptera from Korea); Tsuneki, 1974: 377 (N Distribution. ? Korea, Japan. Korea: “Prov. Ryang-gang: Chan-Pay Plateau: Sam-zi-yan”); Note. As far as I am aware, no Korean material has been exam- Tsuneki, 1976a: 304 (including Korea in distribution); Tsuneki, ined so far. Korean occurrence records listed herein are groundless. 1982: 17 (in the list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 201 (in the list of Ectemnius spinipes (A. Morawitz) 가시은주둥이벌 Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of In- Crabro spinipes A. Morawitz, 1866: 265, \ and _ (syntypes), Russia: sects from Korea); Nemkov in Nemkov et al., 1995: 451 and 453 Amur area, Spask, and Kazan [ZIN]; Tsuneki, 1947a: 282 (Korea: (in key, including Korea in distribution); Yeo et al., 2008: 35 (in “Hakuto-san, Nansetsu-rei, Shoyo-zan”). key), 48e49 (N Korea: Ryanggang-do: Samjiyeon: Mt. Baekdu- Ectemnius spinipes (A. Morawitz): Leclercq, 1954: 288 (new com- san); Nemkov, 2009: 124 (including Korea in distribution); Lee bination); Tsuneki, 1957a: 77 (N Korea: “Kan-nan: Hutempo: et al., 2010: 211 (in Checklist of Korean Insects); Nemkov, Taitimpyoo”); Tsuneki, 1957b: 59 (N Korea: “Tonai”); Tsuneki, 2012b: 444 (including Korea in distribution). 1958a: 15 (including Korea in distribution); Kim, 1970:644e Distribution. Transpalearctic: N Korea (RG), Europe to NE China, 645, 821 (in the list of Hymenoptera from Korea); Tsuneki, 1982: Russian Far East, Japan. 6 (N Korea: “Prov. Pyong-sung: Bek-sung-li: Za-mo san”), 16 (in the list of the species of the Sphecidae known from the Korean Lestica collaris (Matsumura) 목도리목대장은주둥이벌 Peninsula); Paik, 1985: 200 (in the list of Sphecidae of Korea); Crabro collaris Matsumura, 1912: 174, 175, \ (holotype or syntypes), Kim et al., 1994: 265 (in Check List of Insects from Korea); Japan: Hokkaido: Sapporo [HUM]. Nemkov in Nemkov et al., 1995: 449 (in key, including Korea in Lestica collaris (Matsumura): Leclercq, 1954: 291 (new combina- distribution); Leclercq, 1999: 73 (including Korea in distribu- tion); Tsuneki, 1957b: 60 (S Korea: “Berg Kaya”); Kim, 1970: tion); Yeo, 2009: 10 (in key), 38e40 (the Korean Peninsula: 646e647 (diagnosis, no Korean material), 823 (in the list of Pyeongannam-do, Gangwon-do, Gyeonggi-do, Seoul, Hymenoptera from Korea); Tsuneki, 1982: 17 (in the list of the Chungcheongnam-do, Gyeongsangnam-do, Jeollabuk-do); species of the Sphecidae known from the Korean Peninsula); Nemkov, 2009: 122 (including Korea in distribution); Lee Paik, 1985: 201 (in the list of Sphecidae of Korea); Kim et al., et al., 2010: 211 (in Checklist of Korean Insects); Nemkov, 1994: 265 (in Check List of Insects from Korea); Nemkov in 2012b: 443 (including Korea in distribution). Nemkov et al., 1995: 452 and 453 (in key, including Korea in Distribution. Transpalearctic: Korean Peninsula (RG, HN, PN, SL, distribution); Yeo et al., 2008: 36 (in key), 39e40 (S Korea: GG, GW, CN, GN, JB), Europe to China, Russian Far East, Japan. Gangwon-do, Gyeonggi-do, Incheon Metropolitan City, Chungcheongbuk-do, Chungcheongnam-do, Gyeongsangbuk- Genus Lestica Billberg 목대장은주둥이벌속 do, Gyeongsangnam-do); Nemkov, 2009: 124 (including Korea Lestica Billberg, 1820: 107. Type species: Crabro subterraneus Fab- in distribution); Lee et al., 2010: 211 (in Checklist of Korean In- ricius, 1775, designated by Rohwer, 1911: 154. sects); Nemkov, 2012b: 444 (including Korea in distribution). Distribution. S Korea (IC, GW, GG, CB, CN, GB, GN), China, Russian Lestica alata (Paner) 황다리은주둥이벌 Fat East, Japan. Crabro alatus Panzer, 1797 (Heft 46): 6, _ (holotype or syntypes), Germany: no specific locality (depository uncertain); Iwata, Lestica heros (Kohl) 여신은주둥이벌 1938: 84 (N Korea: no specific locality); Tsuneki, 1943a: 156 (N Crabro heros Kohl, 1915: 120, \ (holotype or syntypes), Korea: no Korea: “Hakuto-san”); Tsuneki, 1947a: 285e286 (Korea: “Keijo, specific locality [NHMW]; Iwata, 1938: 84 (description of _ with Hakoto-san, Shoyo-san, Kodai-san”). Japanese material); Tsuneki, 1947a: 286 (in Crabronidae fauna of Lestica alata (Paner): Billberg, 1820: 107 (new combination); Korea); Kim, 1970: 640 (listed with distribution), 822 (in the list Leclercq, 1954: 292, 324; Kim, 1970: Pl. 64 Figure 786, 645 (as of Hymenoptera from Korea); Lee et al., 2010: 210 (in Checklist alatus, diagnosis, Korea: Paldang), 822 (listed as alatus); Tsuneki, of Korean Insects). 1982: 9 (N. Korea: “Prov. South Pyongan: Nam-po”), 17 (in the Lestica heros (Kohl): Tsuneki, 1952b: 66 (new combination); list of the species of the Sphecidae known from the Korean Leclercq, 1954: 293 (including Korea in distribution); Tsuneki, Peninsula); Paik, 1985: 200 (in the list of Sphecidae of Korea); 1982: 17 (in the list of the species of the Sphecidae known Nemkov in Nemkov et al., 1995: 451 and 453 (in key, including from the Korean Peninsula); Paik, 1985: 200 (in the list of Korea in distribution); Yeo et al., 2008: 36 (in key), 36e37 (S Sphecidae of Korea); Nemkov in Nemkov et al., 1995: 452 and Korea: Gangwon-do, Gyeonggi-do, Incheon Metropolitan City, 453 (in key, including Korea in distribution); Yeo et al., 2008:35 Jeollanam-do); Nemkov, 2009: 123 (including Korea in distri- (in key), 43e44 (new to S Korea: Gyeonggi-do); Nemkov, 2009: bution); Lee et al., 2010: 211 (in Checklist of Korean Insects); 124 (including Korea in distribution); Nemkov, 2012b:444 Nemkov, 2012b: 444 (including Korea in distribution). (including Korea in distribution). JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 433

Distribution. S Korea (GG), China, Russian Far East, Japan. Distribution. Holarctic: N Korea (RG), Europe to Russian Far East, Lestica reiteri (Kohl) 이와타목대장은주둥이벌 Canada, USA. Crabro reiteri Kohl, 1915: 119 (as Reiteri, incorrect original capitali- zation), \ (holotype or syntypes), Japan: Tokyo [NHMW]. Rhopalum coarctatum chosenense Tsuneki, 1974 일본호리은주둥 Lestica reiteri (Kohl): Tsuneki, 1952b: 66 (new combination); 이벌 Leclercq, 1954: 295 (including Korea in distribution); Tsuneki, Rhopalum nipponicum chosenense Tsuneki, 1974: 382, _ (holotype) 1982: 9 (N Korea: “Prov. Pyong-Sung: Bek-sung-li: Za-mo san”), and \, N Korea: “South Phenan Province” (Pyeongannam-do): 17 (in the list of the species of the Sphecidae known from the “Sa-gam, 30 km N Pyongyan” [TMB]; Bohart and Menke,1976: Korean Peninsula); Paik, 1985: 201 (in the list of Sphecidae of 630 (including Korea in distribution); Tsuneki, 1982: 17 (in the Korea); Nemkov in Nemkov et al., 1995: 452 and 453 (in key, list of the species of the Sphecidae known from the Korean including Korea in distribution); Nemkov, 2008: 23 (including Peninsula); Paik, 1985: 202 (in the list of Sphecidae of Korea); Korea in distribution); Yeo et al., 2008: 35 (in key), 44e47 (S Tsuneki, 1991: 203 (as new subspecies, nomen nudum). Korea: Gangwon-do, Gyeonggi-do, Seoul, Chungcheongnam-do, Rhopalumcoarctatum chosenense Tsuneki: Leclercq, 2011: 9 (new Gyeongsangbuk-do, Gyeongsangnam-do); Nemkov, 2009:124 subspecific combination). (including Korea in distribution); Nemkov, 2012b: 444 (including Distribution. N Korea (PN). Korea in distribution). Crabro reiteri kuramaensis Iwata, 1938:83e84, \ (holotype), Japan: Rhopalum coarctatum koreense Tsuneki, 1974 고려호리은주둥이벌 Hokkaido: Jozankei (Hokkaido University), synonymized with Rhopalum coarctatum koreense Tsuneki, 1974: 382, _ (holotype), N Lestica reiteri by Tsuneki, 1952b:66;Tsuneki, 1947a: 285 (new to Korea: “Prov. Ryang-gang, Plateau Chann-Pay: Sam-zi-yan” Korea, N Korea: “Hakoto-san: Jimmunjo”); Kim, 1970:646 [TMB]; Bohart and Menke, 1976: 630 (including Korea in dis- (diagnosis, Korea: Mt. Gayasan), 823 (in the list of Hymenoptera tribution); Tsuneki, 1982: 17 (in the list of the species of the from Korea); Kim et al., 1994: 265 (in Check List of Insects from Sphecidae known from the Korean Peninsula); Paik, 1985: 202 Korea). (in the list of Sphecidae of Korea); Yeo et al., 2008: 17 (in key), Distribution. Korean Peninsula (RG, PN, SL, GW, GG, CN, GB, GN), 17e19 (S Korea: Gyeonggi-do, Seoul, Jeju Is.); Leclercq, 2011:9 Russian Far East, Japan. (include N Korea in distribution). Rhopalum coarcitatum [sic] koreense Tsuneki, 1991: 203 (as new Genus Lindenius Lepeletier et Brullé, 1835 흰입술은주둥이벌속 subspecies, nomen nudum). Lindenius Lepeletier et Brullé, 1835: 791. Type species: Crabro albi- Distribution. Korean Peninsula (RG, GG, SL, JJ). labris Fabricius, 1793, designated by Westwood, 1839: 80. Rhopalum gracile Wesmael, 1849 호리은주둥이벌(신칭) Lindenius albilabris Fabricius, 1793 흰입술은주둥이벌 Rhopalum nigrinum Kiesenwetter, 1849:91,\ and _ (syntypes), Crabro albilabris Fabricius, 1793: 302, sex not indicated. \ (holo- Germany: Sachsen-Anstalt: Eisleben (depository uncertain), type), Germany: Halae Saxonum, now Sachsen-Anstalt: Halle junior secondary homonym of Crabro nigrinus Herrich- [ZMUC]. Schaeffer, 1841; Tsuneki, 1974: 382e383 (N Korea: “Prov. Lindenius albilabris (Fabricius): Lepeletier et Brullé, 1835: 795 (new South Pyongan: Bongha-ri, Sa-gam, Man-yong-dae, Pyongyan”); combination); Tsuneki, 1957b: 62 (N Korea: “Kainei”); Kim, Tsuneki, 1991: 203 (in the list of Sphecidae of Korea); Leclecq, 1970: 647 (listed), 823 (in the list of Hymenoptera from Ko- 2011: 11 (citing Tsuneki, 1974). rea); Tsuneki, 1982: 17 (in the list of the species of the Sphecidae Rhopalum gracile Wesmael, 1852: 592 (as Gracile, incorrect original known from the Korean Peninsula); Bohart and Menke, 1976: capitalization), \ (holotype), Switzerland: Genève area (Brux- 384 (including Korea in distribution); Paik, 1985: 202 (in the list elles), synonymized with Rhopalum nigrinum Kiesenwetter by of Sphecidae of Korea); Kim et al., 1994: 265 (in Check List of Leclercq, 1954: 189, but valid with homonymy of Rhopalum Insects from Korea); Nemkov in Nemkov et al., 1995: 426 (in key, nigrinum Kisenwetter; Tsuneki, 1982: 17 (in the list of the spe- including Korea in distribution); Nemkov, 2009:93e94 cies of the Sphecidae known from the Korean Peninsula); Paik, (including Korea in distribution); Lee et al., 2010: 211 (in 1985: 202 (in the list of Sphecidae of Korea); Nemkov in Checklist of Korean Insects); Nemkov, 2012b: 444 (including Nemkov et al., 1995: 428 (in key, including Korea in distribu- Korea in distribution). tion); Yeo et al., 2008: 17 (in key), 19e22 (Korean Peninsula: Distribution. Transpalearctic: N Korea (“Kainei”), Europe to Pyeongannam-do, Gangwon-do, Incheon Metropolitan City, Mongolia, NE China, Russian Far East, Taiwan, SE Asia. Gyeongsangbuk-do); Nemkov, 2009: 96 (including Korea in distribution); Nemkov, 2012b: 444 (including Korea in Genus Rhophalum Stephen, 1829 호리은주둥이벌속 distribution). Rhopalum Stephens, 1829: 366. Type species: Crabro rufiventris Distribution. Holarctic: Korean Peninsula (PN, GW, IC, GB), Panzer, 1799 (¼Sphex clavipes Linnaeus, 1758), designated by J. Europe to Far East Russia, Japan, Canada, USA. Curtis, 1837: 656. Rhopalum jessonicum (Bischoff, 1921) 다듬이호리주둥이벌 Rhopalum clavipes (Linnaeus, 1758) 몽둥이호리은주둥이벌 Crabro jessonicum Bischoff, 1921: 7, \ (holotype), Japan: Hokkaido: Sphex clavipes Linnaeus, 1758: 569, sex not indicated, \ (lectotype), Akke shi [ZMHU]. Europe: no specific locality [LS], designated by Day, 1979: 59. Rhopalum jessonicum (Bischoff): Tsuneki, 1952a: 112 and 115 (new Rhopalum clavipes (Linnaeus): Dahlbom, 1839: 43 and 1839e1840: combination); Yeo et al., 2008: 17 (in key), 22e23 (new to Korea: 43 (new combination); Tsuneki, 1982: 10 (North Korea: “Prov. Gyeongsangbuk-do: Uljin-gun). Ryang-gang, Chan-Pay Plateau: Sam-Zi-yan”), 17 (in the list of Distribution: S Korea (GB), Russian Far East, Japan. the species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 202 (in the list of Sphecidae of Korea); Nemkov in Rhopalum kuwayamai Tsuneki, 1952 쿠와야마호리은주둥이 Nemkov et al., 1995: 428 (in key, including Korea in distribu- 벌(신칭) tion); Nemkov, 2012b: 445 (including Korea in distribution). Rhopalum kuwayamai Tsuneki, 1952a:121,\ (holotype), Russian Far East: Kuril Islands: Etorofu [HUM]; Yeo et al., 2008:16(in 434 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

key), 24e25 (new to Korea: Gangwon-do; Gyeongsangbuk- Rhopalum venustum Tsuneki, 1955 쓰네기호리은주둥이벌 do). Rhopalum venustum Tsuneki, 1955: 107, \ (holotype), Japan: Fukui Distribution. S Korea (GW, GB), Russian Far East, Japan. Prefecture: Koike [HMNHA]: Kim et al., 1994: 265 (in Check List of Insects from Korea); Lee et al., 2010: 211 (in Checklist of Rhopalum laticorne (Tsuneki, 1947) 뿔호리은주둥이벌 Korean Insects). Crabro laticornis Tsuneki, 1947a: 292, _ (holotype) and \, N Korea: Distribution. ? Korea, Japan. “Nanetsurei” [MNHAH], and Daitaku (paratype). Note. Korean occurrence of this species is very doubtable. Both Rhopalum laticorne (Tsuneki): Tsuneki, 1952a: 111 and 114 (including records by Kim et al. (1994) and Lee et al. (2010) were just checklists Korea in distribution), 119 (new combination, N Korea: “Daitaku, that were not based on examination of Korean material. The most Nansetsu-rei”); Leclercq, 1954: 189 (including Korea in distribu- recent work on Korean Crabroninae by Yeo et al. (2008) also did not tion); Tsuneki, 1957a:78(Korea:“Kongo-san”); Tsuneki, 1957b:62 include this species in Korean fauna. (N Korea: “Mozanrei”); Tsuneki, 1960b: 57 and 60 (including Ko- rea in distribution); Kim, 1970: Pl. 64, Figure 792, 647e648 Tribe Larrini latreille, 1810 어리구멍벌족(신칭) (diagnosis, S Korea: Hyangrobong), 823 (in the list of Hymenop- Subtribe Gastrosericina André, 1886 어리구멍벌아족(신칭) tera from Korea); Bohart and Menke,1976: 389 (including Korea in Genus Tachyspex Kohl, 1883 어리은줄구멍벌속 distribution); Tsuneki, 1982: 17 (in the list of the species of the Tachysphex Kohl, 1883:166. Type species: Tachysphex filicornis Kohl, Sphecidae known from the Korean Peninsula); Paik,1985: 202 (in 1883 (¼ Tachytes fugax Radoszkowski, 1877), designated by the list of Sphecidae of Korea); Kim et al.,1994: 265 (specific name Bingham, 1897: 192. incorrectly iaticorne in Euplilis, in Check List of Insects from Korea); Kim et al.,1994: 265 (in Check List of Insects from Korea); Nemkov Tachysphex nigricolor (Dalla Torre, 1897) 검정구멍벌, 검은어리 in Nemkov et al., 1995: 426 (in key, including Korea in distribu- 구멍벌 tion); Nemkov, 2008: 21 (including Korea in distribution); Yeo Larrada nigricans F. Smith, 1873:192,\ (lectotype, designated by et al., 2008: 16 (in key), 25e28 (S Korea: Gangwon-do, Pulawski, 1975: 310), Japan: Nagasaki [BMNH], junior primary Gyeonggi-do, Gyeongsangnam-do); Nemkov, 2009:96 homonym of Larrada nigricans Walker, 1871. (including Korea in distribution); Lee et al., 2010: 211 (in Checklist Larra nigricolor Dalla Torre, 1897: 670, substitute name for Larrada of Korean Insects); Leclercq, 2011: 13 (including Korea in distri- nigricans F. Smith, 1873. bution); Nemkov, 2012b: 444 (including Korea in distribution). Tachysphex nigricolor (Dalla Torre): Pulawski, 1975: 310 (new Rhopalum (Latrorhopalum) latronum Kohl: Kim, 1970: Pl. 65, combination); Bohart and Menke, 1976: 275 (including Korea in Figure 793a, b, 648 (diagnosis, S Korea: Mt. Taebaeksan), 823 distribution); Tsuneki, 1982: 15 (in the list of the species of the (new to Korea, in the list of Hymenoptera from Korea), Sphecidae known from the Korean Peninsula); Paik, 1985:199 misidentification corrected by Yeo et al., 2008: 28; Paik, 1985: (in list of Sphecidae of Korea); Kim et al., 1994: 261 (in Check List 202 (in the list of Sphecidae of Korea); Kim et al., 1994: 265 (in of Insects from Korea); Nemkov in Nemkov et al., 1995: 410 (in Check List of Insects from Korea); Lee et al., 2010: 211 (in key, including Korea in distribution); Yamane in Yamane et al., Checklist of Korean Insects). 1999:517;Nemkov, 2008: 19 (including Korea in distribution); Distribution. Korean Peninsula (RG, HB, GW, GG, GN), NE China, Nemkov, 2009: 79 (including Korea in distribution); Lee et al., Russian Far East. 2010: 208 (in Checklist of Korean Insects); Nemkov, 2012b: 438 (including Korea in distribution). Rhopalum pygidiale Bohart, 1976 검정호리은주둥이벌 Tachysphex japonicus Iwata, 1933b:47,\ (holotype) and _, Japan: Crabro calceatus Tsuneki, 1947b: 430, _ (holotype) and \, Japan: Honshu: Ikeda, Settsu [KUK], synonymized with Tachysphex Hokkaido: Jozankei [MNHAH], junior primary homonym of nigricolor by Pulawski, 1975:311;Tsuneki, 1964b: 63 (in key, Crabro calceatus Rossi, 1794. new to Korea, no Korean material). Rhopalum calceatum (Tsuneki): Tsuneki, 1952a: 113 and 114 (in Tachytes begalensis japonicus Iwata: Tsuneki, 1967b:49e53 (new key), 123 (new combination); Tsuneki, 1974: 383 (new to Korea, combination); Pulawski, 1971: 162, corrected to Tachysphex N Korea: “Kaesong: Mts. Pakyon: Pakyon popo”); Tsuneki, 1982: japonicus. 17 (in the list of the species of the Sphecidae known from the Tachysphex bengalensis Cameron: Kim, 1970: Pl. 57 Figure 677, 581 Korean Peninsula); Paik, 1985: 202 (in the list of Sphecidae of (possibly follow interpretation of Tsuneki 1967b but without Korea); Tsuneki, 1991: 203 (in the list of Sphecidae of Korea). recognition of subspecies, diagnosis, Korea: Mt. Gayasan), 810 (in Rhopalum pygidiale Bohart in Bohart and Menke, 1976: 390, sub- the list of Hymenoptera from Korea); Paik, 1985: 199; Kim et al., stitute name for Crabro calceatus Tsuneki: Kim et al., 1994: 265 1994:261;Lee et al., 2010: 208 (in Checklist of Korean Insects). (in Check List of Insects from Korea); Nemkov in Nemkov et al., Distribution. S Korea (GN), Russian Far East, Japan. 1995: 426 (in key, specific name incorrectly as pigidiale, including Korea in distribution); Yeo et al., 2008: 16 (in key), Tachysphex pompiliformis (Panzer, 1804) 대륙어리구멍벌 28e29 (S Korea: Gangwon-do, Jeju Is.); Nemkov, 2009:95 Larra pompiliformis Panzer, 1804 (Heft 89): 13, \ (holotype), Ger- (including Korea in distribution); Lee et al., 2010: 211 (in many: no specific locality [ZSBS]. Checklist of Korean Insects); Nemkov, 2012b: 444 (including Tachysphex pompiliformis (Panzer): Kohl, 1885: 388 (new combi- Korea in distribution). nation, as synonym of T. pectinipes of Kohl); Tsuneki, 1982:15(N Distribution. Korean Peninsula (GG, GW, JJ), Russian Far East, Japan. Korea: “Hakugan: Baek-am”, in the list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985:199 Rhopalum watanabei Tsuneki, 1952 방태산호리은주중이벌 (in the list of Sphecidae of Korea); Nemkov in Nemkov et al., Rhopalum watanaibei Tsuneki, 1952: 124, _ (holotype), Japan: 1995: 410 (in key, including Korea in distribution); Nemkov, Hokkaido, Jozankei [MNHAH]; Yeo et al., 2008: 16 (in key), 28e 2009: 79 (including Korea in distribution); Nemkov, 2012b: 29 (new to Korea: Gangwon-do: Mt. Bangtaesan, Mt. 438 (including Korea in distribution). Cheongoksan). Tachytes nigripennis Spinola, 1808: 260, \ (holotype or syntypes), Distribution. S Korea (GW), Russian Far East, Japan; Taiwan. Italy: “prope Genuam” (lost), synonymized with Tachysphex pompiliformis by Pulawski, 1971: 62. JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 435

Tachysphex nigripennis (Spinola): Kohl, 1883e: 671 (new combina- distribution); Lee et al., 2010: 208 (in Checklist of Korean In- tion); Tsuneki, 1964b: 63 (in key, including Korea in distribution, sects); Baik and Kim, 2014: 302 (in key), 303e304 (S Korea: as Tachyspex sp. with comment “very similar to T. nigripennis”). Gangwon-do, Incheon Metropolitan City, Gyeonggi-do, Chung- Distribution. Transpalearctic: N Korea (“Hakugan”), N Africa, cheongnam-do, Gyeongsangnam-do, Jeollanam-do). Europe to Mongolia, China, Russian Far East. Distribution. Korean Peninsula (PN, SL, IC, GG, GW, CN, GN, JN), China, Japan, India, Thailand, Taiwan. Genus Tachytes Panzer, 1806 참구멍벌속 Tachytes Panzer, 1806: 129. Type species: Pompilus tricolor of Fab- Tachytes gyusanus Baik et Kim, 2014 규산구멍벌(신칭) ricius, 1798 (¼ Sphex tricolor Fabricius, 1793, junior primary Tachytes gyusanus Baik et Kim, 2014: 302 (in key), 303e307, \ homonym of Sphex tricolor Schrank, 1781), by monotypy. (holotype) and _, Korea: Chungcheongnam-do: Seosan-si: Haemi-myeon: Daegok-ri; Gangwon-do, Incheon Metropolitan Note. Tachytes monticola Dalla Torre (1897: 681) was substitute City, Gyeonggi-do, Chungcheongbuk-do [NIBR]. name for Tachytes montanus (Radoszkowski, 1886: 31) that was Distribution. S Korea (IC, GG, CB, CN). junior secondary homonym of Tachysphex montanus (Cresson, 1865), and now is Tachyspex brullii F. Smith 1856, which is broadly Tachytes modestus Smith, 1856 은줄구멍벌 distributed in Europe (Pulawski, 2014). Previous inclusion of out- Tachytes modestus Smith, 1856: 299, \ (lectotype, designated by of-date name Tachytes monticola to Korean fauna by Kim et al. Pulawski, 1975: 316), India: Punjab [BMNH]; Tsuneki, 1964a:2e (1994: 261) and Lee et al. (2010: 208) were unfounded. 3 (in key) and 5 (including Korea in distribution); Tsuneki, 1964b: 63 (in key, including Korea in distribution); Kim, 1970: Tachytes etruscus sibiricus Gussakovskij, 1932 닙폰구멍벌 Pl. 57 Figure 674, 579 (diagnosis, S Korea: Dnguleung), 809 (in Tachytes sibiricus Gussakovskij, 1932: 14 (as sibirica, incorrect the list of Hymenoptera from Korea); Tsuneki 1974: 362 (N original termination), \ (holotype), Russia: Primorskiy: Kha- Korea: “Prov. South Pyongan: De-sang san, Lyong-ak san”); Kim, barovsk [ZIN]. 1980: Pl. xviii HYLA 01, 130 (specific name incorrectly as mod- Tachytes etruscus sibiricus Gussakovskij: Nemkov in Nemkov et al., esta, diagnosis, Korea: Gyeonggi-do, Chungcheongnam-do, 1995: 409 (in key, new status, including Korea in distribution); Gyeongsangbuk-do); Tsuneki, 1982: 4, 15 (in the list of the Terayama, 2008: 14 and 20 (including Korea in distribution); Li species of the Sphecidae known from the Korean Peninsula); et al., 2008:154e155 (including Korea in distribution); Nemkov, Paik, 1985:199in Yamane et al., 1999: 515 (including Korea in 2009: 77 (including Korea in distribution); Nemkov, 2012b:437 distribution); Li et al., 2008:154e155 (in key); Terayama, 2008: (including Korea in distribution); Baik and Kim, 2014: 302 (in 12e14 and 20 (including Korea in distribution); Lee et al., 2010: key), 302-302 (Korea: Gyeonggi-do, Gyeongsangnam-do, Jeol- 208 (in Checklist of Korean Insects); Baik and Kim, 2014: 302 (in lanam-do). key), 308e309 (S Korea: Gangwon-do, Gyeonggi-do, Tachytes nipponicus Tsuneki, 1964a:2e3 (in key), 6e8. _ (holotype), Gyeongsangnam-do, Jeollanam-do). Japan: Fukui Prefecture: no specific locality [MNHAH], synony- Distribution. Korean Peninsula (PN, SL, GG, GW, CN, GB, GN, JN), mized with Tachytes etruscus sibiricus by Nemkov in Nemkov China, Japan, India, Thailand, Taiwan. et al., 1995: 409; Tsuneki, 1964b: 63 (in key, including Korea in distribution); Kim, 1970: Pl. 57 Figure 675, 580 (diagnosis, S Tachytes sinensis F. Smith, 1856 참구멍벌 Korea: Donguleung), 809 (in the list of Hymenoptera from Ko- Tachytes sinensis F. Smith, 1856: 299 (as Sinensis, incorrect original rea); Tsuneki, 1974: 362 (N Korea: Prov. South Pyongan: Za-mo capitalization), \ (holotype or syntypes), China: Shanghai san); Tsuneki, 1982: 4 (N Korea: “Prov. Pyong Sung: Bek-sung- [BMNH]; Tsuneki, 1964a: 2 (in key) and 3e5 (including Korea in li: Za-mo san”), 15 (in the list of the species of the Sphecidae distribution); Yasumatsu, 1965: 298 (including Korea in distri- known from the Korean Peninsula); Paik, 1985: 199 (in the list of bution); Kim, 1970: Pl. 57 Figure 675, 581 (diagnosis, S Korea: Mt. Sphecidae of Korea); Tsuneki, 1991: 199 (in the list of Sphecidae Gayasan), 809 (in the list of Hymenoptera from Korea); Tsuneki, of Korea); Kim et al., 1994: 261 (in Check List of Insects from 1974:362(NKorea:“Prov. South Pyongan: Za-mo san”); Bohart Korea); Lee et al., 2010: 208 (in Checklist of Korean Insects). and Menke, 1976: 267 (including Korea in distribution); Kim, Tachytes etruscus (Rossi): Terayama, 2008: 14 (including Korea in 1980: Pl. xviii HYLA 02, 131 (diagnosis, S Korea: Gangwon-do, distribution); Kim et al., 1994: 261 (specific name incorrectly as Gyeonggi-do, Gyeongsangbuk-do); Tsuneki, 1982: 15 (in the list etruscoides, in Check List of Insects from Korea). of the species of the Sphecidae known from the Korean Penin- Distribution. Korean Peninsula (PN, SL, GG, GN, JN), N China, sula); Paik, 1985: 199 (in the list of Sphecidae of Korea); Tsuneki Russian Far East, Japan. 1991: 199 (in the list of Sphecidae of Korea); Yamane in Yamane et al., 1999: 516 (including Korea in distribution); Yamane, Tachytes fruticis Tsuneki, 1964 구리구멍벌 2008: 554 (including Korea in distribution); Li et al., 2008: Tachytes fruticis Tsuneki, 1964a:8,_ (holotype) and \, Japan: Fukui 154e155 (in key); Terayama, 2008:14e16 and 20 (including Prefecture: Hatogayu [MNHAH]; Tsuneki, 1964b: 63 (in key, Korea in distribution); Lee et al., 2010: 208 (in Checklist of Korean including Korea in distribution); Kim, 1970: Pl. 56 Figure 673, Insects); Baik and Kim, 2014: 302 (in key), 308 (S Korea: 579 (diagnosis, S Korea: Mt. Gayasan), 809 (in the list of Hy- Gangwon-do, Incheon Metropolitan City, Gyeonggi-do, Chung- menoptera from Korea); Tsuneki, 1974: 363 (N Korea: “Prov. cheongnam-do, Gyeongsangnam-do, Jeollanam-do). South Pyongan: De-sang san, Za-mo san”); Bohart and Menke, Distribution. Korean Peninsula (PN, IC, GG, CN, GW, GB, GN, JN), 1976: 265 (including Korea in distribution); Kim, 1980:131 China, Japan, India, Thailand, Taiwan. (Korean localities in Note of Tachytes sinesis Smith); Tsuneki 1982: 15 (in the list of the species of the Sphecidae known Subtribe Larrina Latreille, 1810 홍띠구멍벌아족(신칭) from the Korean Peninsula); Paik, 1985: 199 (in the list of Genus Larra Fabricius, 1793 홍띠구멍벌속 Sphecidae of Korea); Tsuneki, 1991: 200 (in the list of Sphecidae Larra Fabricius, 1793: 220. Type species: Larra ichneumoniformis of Korea); Kim et al., 1994: 261 (in Check List of Insects from Fabricius, 1793 (¼ Sphex anathema Rossi, 1790), designated by Korea); Li et al., 2008:154e155 (including Korea in distribu- Latreille, 1810: 438. tion); Terayama, 2008: 12 and 20 (including Korea in 436 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Larra amplipennis (F. Smith, 1873) 홍띠구멍벌 (including Korea in distribution); Lee et al., 2010: 208 (in Larrada amplipennis F. Smith, 1873: 193, \ and _ (holotype or syn- Checklist of Korean Insects). types), Japan: Hyogo [BMNH]. Distribution. ?Korea, China, Japan, SE Asia, Taiwan. Larra amplipennis (F. Smith): Kohl, 1885: 241 (new combination), Note. All the publications including Korean occurrences listed 258; Kim, 1970: Pl. 56 Figure 669, 577 (diagnosis, S Korea: herein are not based on Korean material, and thus are groundless. Gwangleng), 809 (in the list of Hymenoptera from Korea); Tsuneki, 1982: 18 (in the list of the species of the Sphecidae Tribe Miscophini W. Fox, 1894 어리윤구멍벌족 known from the Korean Peninsula); Paik, 1985: 198 (in the list of Genus Lyroda Say, 1837 일본구멍벌속 Sphecidae of Korea); Kim et al., 1994: 260 (in Check List of In- Lyroda Say, 1837: 372. Type species: Lyroda subita Say 1837, desig- sects from Korea); Terayama, 2008: 7, 19 (including Korea in nated by Patton, 1880b: 386. distribution); Lee et al., 2010: 208 (in Checklist of Korean Insects). Lyroda nigra japonica Iwata, 1933 일본구멍벌 Distribution. S Korea (GG), Japan, Taiwan, Philippines, Thailand. Odontolarra nigra Cameron, 1904:12,\ (holotype or syntypes), In- dia: Sikkim: no specific locality [BMNH]. Larra carbonaria erebus (F. Smith, 1873) 먹윤구멍벌 Lyroda nigra (Cameron): Bohart and Menke, 1976: 299 (new com- Larrada erebus F. Smith, 1873: 193, \ (holotype or syntypes), Japan: bination); Tsuneki, 1983: 57 (reexamination of type); Yamane in Hyogo [BMNH]. Yamane et al., 1999: 519 (including Korea in distribution); Larra erebus (F. Smith): Kohl, 1885: 243 (new combination). Nemkov, 2008: 19 (including Korea in distribution); Terayama, Larra carbonaria (F. Smith): Tsuneki, 1967b:8e9 and 13 (in key, 2008: 19 and 20 (including Korea in distribution). including Korea in distribution); Kim et al., 1994: 261 (in Check Lyroda japonica Iwata, 1933: 8, \ and _ (holotype), Japan: Honshu: List of Insects from Korea); Yamane in Yamane et al., 1999: 509 Settsu: Ikeda (Kyoto University); Yasumatsu, 1965:297 (including Korea in distribution); Terayama, 2008:7,19 (including Korea in distribution); Kim, 1970: Pl. 56 Figure 672, (including Korea in distribution); Lee et al., 2010: 208 (in 578e579 (diagnosis, Korea: Baekgyo), 809 (in the list of Hy- Checklist of Korean Insects). menoptera from Korea); Tsuneki, 1974: 363 (N Korea: “Prov. Larra carbonaria erebus (F. Smith): Tsuneki, 1964g: 221 (new status, South Pyongan: Sa-gam”); Tsuneki, 1982: 15 (S Korea: “Shoyo- syntype male of Larrada docilis Smith,1873 ¼ carbonaria erebus); zan”, in the list of the species of the Sphecidae known from the Kim, 1970: 578 (diagnosis, no Korean material), 810 (in the list of Korean Peninsula); Paik, 1985: 199 (in Sphecidae from Korea); Hymenoptera from Korea); Tsuneki, 1982: 15 (S Korea: Mt. Tsuneki, 1991: 200 (in Sphecidae from Korea); Kim et al., 1994: Soyosan); Paik, 1985: 198 (in the list of Sphecidae of Korea). 261 (in Check List of Insects from Korea); Nemkov in Nemkov Distribution. S Korea (GG), Japan, SE Asia. et al., 1995: 413 (in key, including Korea in distribution); Nemkov, 2009: 81 (including Korea in distribution); Lee et al., Genus Liris Fabricius, 1804 귀뚜라미벌속 2010: 208 (in Checklist of Korean Insects). Liris Fabricius, 1804: 227. Type species: Sphex auratus Fabricius, Lyroda nigra japonica Iwata: Tsuneki, 1983: 58 (new status), 59; 1787, junior primary homonym of Sphex auratus Linnaeus, 1758 Terayama, 2008: 18 and 20 (including Korea in distribution); (¼ Sphex aurulentus Fabricius, 1787), designated by Patton, Yamane, 2008: 554 (including Korea in distribution); Nemkov, 1880b: 386. 2012b: 439 (including Korea in distribution). Distribution. Korean Peninsula (PN, GG), NE China, Japan. Liris festinans japonicus (Kohl, 1884) 귀뚜라미벌 Notogonia japonica Kohl, 1884:357,\ (lectotype, designated by de Genus Miscophus Jurine, 1807 두색구멍벌속 Beaumont, 1961: 237), Kyoto, Japan [NHMW]. Miscophus Jurine, 1807: 206. Type species: Miscophus bicolor Jurine, Liris japonica (Kohl): Beaumont, 1961ba: 237 (new combination); 1807, by monotypy. Tsuneki, 1964b: 62 (including Korea in distribution); Kim, 1970: 578 (diagnosis, no Korean material), 810 (in the list of Hyme- Miscophus bicolor Jurine, 1807 두색구멍벌 noptera from Korea); Tsuneki, 1974: 363 (N Korea: “Prov. South Miscophus bicolor Jurine, 1807: Pl. 11 and Pl. 11, Figure 25, _ (holo- Pyongan, Hotel garden; Kaesong, Mts. Pakyon, Pakyon popo”); type or syntypes), Europe: no specific locality, perhaps Tsuneki, 1991: 200 (in Sphecidae from Korea); Kim et al., 1994: Switzerland [MHNG]; Kim, 1970: Pl. 91 Figure 679, 583 (diag- 261 (in Check List of Insects from Korea); Lee et al., 2010: 208 (in nosis, no Korean material), 810 (in the list of Hymenoptera from Checklist of Korean Insects). Korea); Kim et al., 1994: 261 (in Check List of Insects from Ko- Liris festinans japonica (Kohl): Vecht in Bohart and Menke,1976: 245 rea); Nemkov in Nemkov et al., 1995: 413 (in key, including (new status); Tsuneki, 1982: 15 (in the list of the species of the Korea in distribution); Nemkov, 2009:81e82 (including Korea Sphecidae known from the Korean Peninsula); Paik, 1985:199 in distribution); Lee et al., 2010: 208 (in Checklist of Korean (in Sphecidae from Korea); Yamane in Tamane et al., 1999: 514. Insects). Liris festinans japonicus (Kohl): Li et al., 2007: 419 (in key), 422 (in Miscophus tsunekii de Andrade, 1960:97,\ (holotype) and _, Korea: key); Terayama, 2008: 8, 19 (including Korea in distribution); Li “Keijo” (University of Coimbra), synonymized with Miscophus et al., 2009: 117 and 118 (including Korea in distribution in key bicolor by Nemkov in Nemkov et al., 1995:413;Tsuneki, 1964g: to the species of Liris from Palearctic region). 62 (in key, including Korea in distribution); Kim, 1970: 583 Distribution. N Korea (PN, GG), China, Japan, S Asia, Taiwan. (diagnosis, no Korean material), 810 (in the list of Hymenoptera from Korea); Tsuneki, 1974: 363 (N Korea: Prov. South Pyongan: Liris subtessellatus (F. Smith, 1856) 귀뚜라미벌 Bongwa-ri); Bohart and Menke, 1976: 319 (including Korea in Larrada subtessellata F. Smith, 1856: 277, \ (syntypes), India, distribution); Tsuneki, 1982: 15 (in the list of the species of the Sumatra, Java [BMNH]. Sphecidae known from the Korean Peninsula); Paik, 1985:199 Liris subtessellata (F. Smith): de Beaumont, 1961a: 4 (new (in the list of Sphecidae of Korea); Tsuneki, 1991: 200 (in combination). Sphecidae from Korea); Kim et al., 1994: 261 (in Check List of Liris subtesellatus (F. Smith): Kim et al., 1994: 261 (in Check List of Insects from Korea); Lee et al., 2010: 208 (in Checklist of Korean Insects from Korea); Yamane in Yamane et al., 1999:513 Insects). JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 437

Distribution. Transpalearctic: Korean Peninsula (PN, SL), N Africa, nudum); Kim et al.,1994: 265 (in Check List of Insects from Korea); Europe to Mongolia, Russian Far East, Japan. Lee et al., 2010: 211 (in Checklist of Korean Insects). Distribution. N Korea (PN). Tribe Oxybelini Leach, 1815 등가시구멍벌족 Genus Oxybelus Latreille, 1796 등가시구멍벌속 Oxybelus strandi Yasumatsu, 1935 스트랜드등가시구멍벌, 야쓰마쓰 Oxybelus Latreille, 1796: 129, no included species. Type species: 등가시구멍벌 Crabro uniglumis of Fabricius, 1775 (¼ Vespa uniglumis Linnaeus, Oxybelus strandi Yasumatsu, 1935b: 39, _ (holotype) and \, Japan: 1758), designated by Latreille, 1802b: 343 (first included Honshu: Shinano: Kamikôchi [KUF]; Tsuneki, 1974: 384 (N Ko- species). rea: “Prov. South Pyongan: Pyongyan, Bongwa-ri”); Tsuneki, 1982: 10, 16 (in the list of the species of the Sphecidae known Oxybelus bipunctatus Olivier, 1812 쌍점등가시구멍벌 from the Korean Peninsula); Paik, 1985: 200 (in the list of Oxybelus bipunctatus Olivier, 1812:597,\ (holotype or syntypes), Sphecidae of Korea); Tsuneki, 1991: 203 (in the list of Sphecidae France: Paris area (lost): Tsuneki, 1982:10e11 (new to Korea, N of Korea); Kim et al., 1994: 265 (in Check List of Insects from Korea: Prov. Ryang-gang, River Karim), 16 (in the list of the Korea); Lee et al., 2010: 211 (in Checklist of Korean Insects). species of the Sphecidae known from the Korean Peninsula); Distribution. N Korea (PN), Japan. Paik, 1985: 200 (in the list of Sphecidae of Korea); Nemkov in Nemkov et al., 1995: 423 (in key, including Korea in distribu- Tribe Palarini 북지구멍벌족(신칭) tion); Nemkov, 2009: 90 (including Korea in distribution); Genus Palarus Latreille, 1802 북지구멍벌속 Nemkov, 2012b: 440 (including Korea in distribution). Palarus Latreille, 1802b: 336. Type species: Tiphia flavipes Fabricius, Distribution. Holarctic: N Korea (RG), N Africa, Europe to Russian 1793 (¼ Palarus rufipes Latreille, 1812), junior secondary hom- Far East, USA. onym of Palarus flavipes (Fabricius, 1781), by monotypy.

Oxybelus eximius Sickmann, 1894 어리등가시구멍벌 Palarus variegatus varius Sickmann, 1894 북지구멍벌 Oxybelus eximius Sickmann, 1894: 205, \ and _ (syntypes), China: Palarus flavipes var. varius Sickmann, 1894:215,\ and _ (syntypes), Hopei Province: Tientsin [Münster]; Yasumatsu, 1935b:39 China: Hopei Province: Tientsin [Münster]. (Korea: “Taiden”); Kim, 1970: 649 (diagnosis, no Korean mate- Palarus variegates varius Sickmann: Tsuneki, 1964g: 61 (new sta- rial), 823 (in the list of Hymenoptera from Korea); Tsuneki, 1982: tus); Kim, 1970: 581 (diagnosis, no Korean material), 814 (in the 18 (in the list of the species of the Sphecidae known from the list of Hymenoptera from Korea); Tsuneki, 1982: 15 (in the list of Korean Peninsula); Paik, 1985: 200 (in the list of Sphecidae of the species of the Sphecidae known from the Korean Peninsula); Korea); Kim et al., 1994: 265 (in Check List of Insects from Ko- Paik, 1985: 199 (in the list of Sphecidae of Korea); Kim et al., rea); Lee et al., 2010: 211 (in Checklist of Korean Insects). 1994: 261 (in Check List of Insects from Korea); Nemkov in Distribution. Korea (“Taiden”), China. Nemkov et al., 1995: 413 (in key, including Korea in distribu- tion); Nemkov, 2009: 81 (including Korea in distribution); Oxybelus haemorrhoidalis Olivier, 1812 등가시구멍벌 Nemkov, 2012b: 437 (including Korea in distribution). Oxybelus haemorrhoidalis Olivier, 1812: 597, sex not indicated (ho- Palarus variegatus (Fabricius): Lee et al., 2010: 208 (in Checklist of lotype or syntypes), France: Paris area (lost); Nemkov, 2009: Korean Insects). 90e91 (including Korea in distribution); Nemkov, 2012b:440 Palarus saishiuensis Okamoto, 1924:202,\ (holotype), South Korea: (including Korea in distribution). Quelpart Island (now Cheju Island): no specific locality (de- Oxybelus victor Lepeletier, 1845:218,\ (holotype or syntypes), pository uncertain), synonymized with Palarus variegatus varius France: Paris area (lost?), synonymized with Oxybelus haemor- by Tsuneki, 1971:34:Kim et al.,1994: 265 (in Check List of Insects rhoidalis by Bitsch and Leclercq, 1993: 253; Tsuneki, 1982: 19 (in from Korea); Lee et al., 2010: 211 (in Checklist of Korean Insects). the list of the species of the Sphecidae known from the Korean Distribution. S Korea (JJ), China, Russian Far East. Peninsula); Paik, 1985: 200 (in the list of Sphecidae of Korea); Nemkov, Kazenas, Budrys, and Antropov, 1995: 424e425 Tribe Trypoxylini Lepeletier, 1845 어리나나니족(신칭) (including Korea in distribution). Genus Pison Jurine, 1808 윤어리나나니속 Oxybelus melancholicus Chevrier, 1868: 394, \ and _ (syntypes), Genus Pison Jurine in Spinola, 1808: 255. Type species: Pison jurini Switzerland: Lake Léman area: no specific locality [MHNG], Spinola, 1808 (correctly jurinei ¼ Alyson ater Spinola, 1808), by synonymized with Oxybelus victor by Blüthgen, 1949: 87; monotypy. Yasumatsu, 1935b: 39 (Korea: “Kogendo: Kongosan”); Kim, 1970: Pl. 65 Figure 795, 650 (diagnosis, Korea: Gwangleung), Pison insigne Sickmann 윤어리나나니 823 (in the list of Hymenoptera from Korea); Kim et al., 1994: Pison insigne Sickmann, 1894:210,_ (syntypes), China: Chu-Yung- 265 (in Check List of Insects from Korea); Lee et al., 2010: 211 (in guan in Nankou Pass [Münster]; Tsuneki, 1964f: 49 (in key, Checklist of Korean Insects). Korea: “Shoyozan”); Kim,1970: Pl. 58 Figure 688, 588 (diagnosis, Distribution. Transpalearctic: Korean Peninsula (GW, GG), N Korea: Mt. Sokrisan), 810 (in the list of Hymenoptera from Ko- Africa, Europe to Mongolia, N China, Russian Far East, Japan, south to rea); Tsuneki, 1976a: 297 (including Korea in distribution); SE Asia. Tsuneki, 1982: 16 (in the list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 199 (in the list of Oxybelus koreanus Tsuneki, 1974 서북등가시구멍벌, 참등가시구 Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of In- 멍벌 sects from Korea); Wu and Zhou, 1996: 100 (including Korea in Oxybelus koreanus Tsuneki, 1974:383,_ (holotype) and \,NKorea: distribution); Nemkov, 2009: 83 (including Korea in distribu- “Prov. South Pyongyan: Sa-gam, 45 Km N of Pyongyan” [TMB]; tion); Terayama and Nambu, 2009: 23, 34 (including Korea in Bohart and Menke, 1976: 629 (including Korea in distribution); distribution); Lee et al., 2010: 209 (in Checklist of Korean In- Tsuneki, 1982: 16 (in the list of the species of the Sphecidae known sects); Li and Li, 2011: 62 (including Korea in distribution); from the Korean Peninsula); Paik, 1985: 200 (in the list of Sphe- Nemkov, 2012b: 439 (including Korea in distribution). cidae of Korea); Tsuneki, 1991: 203 (as new species, nomen 438 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Distribution. S Korea (GG, CB), China, Russian Far East, Japan. Trypoxylon chosenense Tsuneki, 1956 조선어리나나니 Trypoxylon chosenense Tsuneki, 1956a: 5 and 10 (in key), 25, \ Pison koreense (Radoszkowski, 1887) 참어리나나니 (holotype) and _, Korea: “Shoyozan” [MNHAH]; Kim, 1970: Pl. Paraceramius koreensis Radoszkowski, 1887: 433 (as Koreensis, 57 Figure 68, 584e585 (diagnosis, Korea: Uidong), 811 (in the incorrect original capitalization), \ (holotype), Korea: no specific list of Hymenoptera from Korea); Bohart and Menke, 1976: 345 locality [NMK, now lost], \ (neotype, designated by Antropov, (including Korea in distribution); Tsuneki, 1981: 3 (in key), 13e 1994: 125, Russia: Primorskiy Kray, 32 km SE Ussuriysk 15 (based on types); Paik, 1985: 199 (in the list of Sphecidae of [ZMMU]. Korea); Kim et al., 1994: 261 (in Check List of Insects from Ko- Ceramius koreensis (Radoszkowski): Dalla Torre, 1894: 3 (new rea); Lee et al., 2010: 209 (in Checklist of Korean Insects). combination). Distribution. S Korea (GG), Japan. Pison koreense (Radoszkowski): Kohl in Dalla Torre, 1897: 712 (as koreensis, new combination); Yasumatsu, 1935c: 228 (as kore- Trypoxylon exiguum exiguum Tsuneki, 1956 제주꼬마어리나나 ensis, in key), 229e230 (including Korea in distribution); 니(신칭) Yasumatsu, 1939a: 82, 83 (including Korea in distribution); Trypoxylon clavicerum exiguum Tsuneki, 1956a: 6 and 12 (in key), Tsuneki, 1964f: 48 and 49 (in key, including Korea in distribu- 37e38, \ (holotype), Japan: Fukui Prefecture [MNHAH]; Tsuneki tion); Kim, 1970: 588 (as koreensis, diagnosis, no Korean mate- 1968b: 50 (Korea: Jeju Is.: Mt. Hanna); Kim et al., 1994: 261 (in rial), 810 (in the list of Hymenoptera from Korea); Bohart and Check List of Insects from Korea); Lee et al., 2010: 209 (in Menke, 1976: 337 (including Korea in distribution); Tsuneki, Checklist of Korean Insects). 1982: 16 (in the list of the species of the Sphecidae known Trypoxylon exigum exigum Tsuneki: Antropov, 1986: 627 (new from the Korean Peninsula); Paik, 1985: 199 (in the list of status). Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of In- Trypoxylon exigum Tsuneki: Terayama and Nambu, 2009: 25, 35 sects from Korea); Antropov, 1994: 121 (in key), 125 (including (including Korea in distribution). Korea in distribution); Antropov in Nemkov et al., 1995: 415 (in Distribution. S Korea (JJ), Japan. key, including Korea in distribution); Terayama and Nambu, 2009: 13 and 34 (including Korea in distribution); Nemkov, Trypoxylon exiguum gussakovskiji Tsuneki, 1974 꼬마어리나나니 2009: 83 (including Korea in distribution); Lee et al., 2010: Trypoxylon pygmaeum Gussakovskij, 1932:13,\ (lectotype, 209 (in Checklist of Korean Insects); Li and Li, 2011:62 designated by Tsuneki, 1981: 43), Russia: Primorskiy Kray: (including Korea in distribution); Nemkov, 2012b: 439 Sedanka [OMNH], junior primary homonym of Trypoxylon (including Korea in distribution). pygmaeum Cameron; Kim et al., 1994: 261 (in Check List of Distribution. Korea (no specific locality), E China, Russian Far Insects from Korea); Lee et al., 2010: 209 (in Checklist of East, Japan, USA. Korean Insects). Trypoxylon clavicerum pygmaeum Gussakovskij: Tsuneki, 1956a:6 Pison punctifrons Shuckard, 1838 곰보어리나나니 and 12 (in key), 36e37 (new status). Pison punctifrons Shuckard, 1838:77,\ and _ (holotype or syn- Trypoxylon clavicerum gussakovskiji Tsuneki, 1974: 365 (North Ko- types), India or St. Helena: no specific locality [OXUM?]; rea: Kaesong: Mts. Pakyon: Pakyon popo), substitute name for Tsuneki, 1964f: 49 (in key, Korea: “Kodaisan”); Kim, 1970: Pl. 58 Trypoxylon clavicerum pygmaeum Gussakovskij, 1933; Tsuneki, Figure 690, 589 (diagnosis, Korea: Mt. Cheonmasan), 810 (in the 1981: 5 and 6 (in key), 42 (redescription and designation of list of Hymenoptera from Korea); Tsuneki, 1982: 16 (in the list of lectotype \); Tsuneki, 1982: 16 (in the list of the species of the the species of the Sphecidae known from the Korean Peninsula); Sphecidae known from the Korean Peninsula); Paik, 1985:199 Paik, 1985: 199 (in the list of Sphecidae of Korea); Kim et al., (in the list of Sphecidae of Korea). 1994: 261 (in Check List of Insects from Korea); Yamane in Trypoxylon exiguum gussakovskiji Tsuneki: Antropov, 1986:627 Yamane et al., 1999: 529 (including Korea in distribution); (new subspecific combination); Antropov in Nemkov et al., Terayama and Nambu, 2009: 23, 34 (including Korea in distri- 1995: 420 and 422 (in key, including Korea in distribution); bution); Lee et al., 2010: 209 (in Checklist of Korean Insects); Li Nemkov, 2009: 85 (including Korea in distribution); Nemkov, and Li, 2011: 62 (including Korea in distribution). 2012b: 439 (including Korea in distribution). Distribution. S Korea (GG, GW), China, Japan, India, Myanmar, Distribution. N Korea (GG), Russian Far East. Indo-China, Taiwan. Trypoxylon frigidum cornutum Gussakovskij, 1932 총각어리나나니 Pison strandi Yasumatsu, 1935 야쓰마츠어리나나니 Trypoxylon cornutum Gussakovskij, 1932:11,\ and _ (syntypes), Pison strandi Yasumatsu, 1935c: 228 (in key), 231, \ (holotype), Russia: Siberia: Kamchatka: Elisovo, and Primorskiy Kray: Japan: Honshu, Settsu Mount Kinri [Kyushu University, now Sedanka [NRS], also Khabarovsk [ZIN]. lost]; Tsuneki, 1964f: 48 and 49 (in key, including Korea in dis- Trypoxylon frigidum cornutum Gussakovskij: Tsuneki, 1956a: 5 and tribution, Korea: Seoul, “Shoyozan”); Tsuneki, 1982: 16 (in the 10 (in key), 24 (new status); Tsuneki, 1981: 4 (in key, including list of the species of the Sphecidae known from the Korean Korea in distribution), 22 (synonymy of Trypoxylon frigidum Peninsula); Paik, 1985: 199 (in the list of Sphecidae of Korea); chongar as T. frigidum chongar); Tsuneki, 1982: 16 (in the list of Kim et al., 1994: 261 (in Check List of Insects from Korea); the species of the Sphecidae known from the Korean Peninsula); Terayama and Nambu, 2009: 24, 34 (including Korea in distri- Paik, 1985: 199 (in the list of Sphecidae of Korea); Antropov in bution); Nemkov, 2009: 84 (including Korea in distribution); Nemkov et al., 1995: 419 (in key, including Korea in distribu- Nemkov, 2012b: 439 (including Korea in distribution). tion); Nemkov, 2009: 85 (including Korea in distribution); Distribution. S Korea (GG), Japan. Nemkov, 2012b: 439 (including Korea in distribution). Trypoxylon frigidum chongar Tsuneki, 1956a: 25, _ (holotype), N Genus Trypoxylon Latreille, 1796 어리나나니속 Korea: “Nansetsu-rei” [USNM], synonymized with Trypoxylon Trypoxylon Latreille, 1796: 121, no included species. Type species: frigidum cornutum Gusakovskij by Tsuneki, 1981: 22; Kim, 1970: Sphex figulus Linnaeus, 1758, designated by Latreille, 1802b: 339 585 (diagnosis, no Korean material), 811 (in the list of Hyme- (first included species). noptera from Korea); Bohart and Menke, 1976: 346 (including JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 439

Korea in distribution); Kim et al., 1994: 261 (in Check List of In- Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of In- sects from Korea); Lee et al., 2010: 209 (in Checklist of Korean sects from Korea); Antropov in Nemkov et al., 1995: 420 and 422 Insects). (in key, including Korea in distribution); Nemkov, 2009:87 Distribution. N Korea (RG), Russian Far East, Japan. (including Korea in distribution): Lee et al., 2010: 209 (in Checklist of Korean Insects); Nemkov, 2012b: 439 (including Trypoxylon fronticorne japonense Tsuneki, 1956 일본어리나나니 Korea in distribution). Trypoxylon pennsylvanicum japonense Tsuneki, 1956a: 6 and 9 (in Distribution. S Korea (SL), Russian Far East, Japan. key), 29e30, \ (holotype) and _, Japan: Fukui Prefecture, and Korea: “Keijo” [MNHAH]; Kim, 1970: 587 (diagnosis, no Korean Trypoxylon malaisei Gussakovskij, 1932 어리나나니 material), 811 (in the list of Hymenoptera from Korea); Tsuneki, Trypoxylon malaisei Gussakovskij, 1932:10,\ (syntypes), Russia: 1976a: 297 (including Korea in distribution); Kim et al., 1994: Primorskiy Kray: Sutshan, Sedanka near Vladivostok, and 261 (in Check List of Insects from Korea); Lee et al., 2010: 209 (in Tigrovaya [NRS]; Tsuneki, 1956a:3e4 and 7 (in key), 14 (rede- Checklist of Korean Insects). scription); Kim, 1970: Pl. 58 Figure 685, 587 (diagnosis, Korea: Trypoxylon fronticorne japonense Tsuneki, 1979: 58 (new subspecific Mt. Gayasan), 811 (in the list of Hymenoptera from Korea); combination); Tsuneki, 1981: 28 (Korea: Seoul); Paik, 1985:199 Tsuneki, 1973a: 33 and 36 (including Korea in distribution); (in the list of Sphecidae of Korea); Antropov in Nemkov et al., Bohart and Menke, 1976: 347 (including Korea in distribution); 1995: 419 (including Korea in distribution); Terayama and Kim, 1980: Pl. xx HYTR 01, 146 (diagnosis, Korea: Gyeonggi-do, Nambu, 2009: 26 and 35 (including Korea in distribution); Chungcheongbuk-do, Gyeongsangbuk-do); Tsuneki, 1981:7 Nemkov, 2009: 86 (including Korea in distribution); Nemkov, and 8 (in key), 73e75 (Central Korea: Mt. Temma, Mt. Shŏyŏ); 2012b: 439 (including Korea in distribution). Tsuneki, 1982: 16 (in the list of the species of the Sphecidae Distribution. S Korea (SL), N China (questioned by Tsuneki, 1981), known from the Korean Peninsula); Paik, 1985:199(inthelist Russian Far East, Japan. of Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of Insects from Korea); Antropov in Nemkov et al., 1995: 416 (in Trypoxylon fronticorne seoulense Tsuneki, 1981 서울어리나나 key, including Korea in distribution); Yamane in Yamane et al., 니(신칭) 1999:526e527 (including Korea in distribution); Terayama and Trypoxylon fronticorne seoulense Tsuneki, 1981: 4 (in key), 30, \ Nambu, 2009: 29, 36 (including Korea in distribution); Nemkov, (holotype), South Korea: northern suburb of Seoul [MNHAH]; 2009: 87 (including Korea in distribution); Lee et al., 2010:209 Tsuneki, 1982: 16 (specific name incorrectly listed as seulense,in (in Checklist of Korean Insects); Nemkov, 2012b: 439 (including the list of the species of the Sphecidae known from the Korean Korea in distribution). Peninsula); Paik, 1985: 199 (in the list of Sphecidae of Korea). Distribution. S Korea (GG, CB, GB), NE Myanmar, China, Russian Distribution. S Korea (SL). Far East, Japan. Note. This subspecies is sympatric with subspecies T. fronticorne japonense in Korea, which means that these two subspecies are self- Trypoxylon pacificum Gussakovskij, 1932 먹어리나나니 contradictory on their subspecific status. This subspecies was Trypoxylon pacificum Gussakovskij, 1932:12,\ and _ (syntypes), described as a sole female specimen, thus more specimens are Russia: Primorskiy Kray: Tigrovaya, Sedanka, Putyatin’s Island needed for final taxonomic confirmation. Considering the struc- [NRS]; Tsuneki, 1956a: 6, 7, and 11 (in key), 33 (redescription; tural distinctiveness from nominotypical subspecies (Tsuneki Korea: “Shoyozan”); Yasumatsu, 1965: 298 (including Korea in 1981), full specific status is highly possible. distribution); Tsuneki, 1968b: 50 (Korea: Jeju Is.: Mt. Hanna); Kim, 1970: 587 (diagnosis, no Korean material), 811 (in the list of Trypoxylon koma Tsuneki, 1956 고려어리나나니 Hymenoptera from Korea); Tsuneki, 1973: 35 and 37 (including Trypoxylon figulus koma Tsuneki, 1956a: 5 and 9 (in key), 28, \ (ho- Korea in distribution); Bohart and Menke, 1976: 347 (including lotype), Korea: “Mt. Kodai, near 38N” [MNHAH]; Kim, 1970:585 Korea in distribution); Tsuneki, 1981: 7 and 8 (in key), 73; (diagnosis, no Korean material), 811 (in the list of Hymenoptera Tsuneki, 1982: 16 (in the list of the species of the Sphecidae from Korea); Bohart and Menke, 1976: 346 (including Korea in known from the Korean Peninsula); Paik, 1985: 199 (in the list of distribution); Tsuneki, 1981: 3 (in key), 20 (redescription); Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of In- Tsuneki 1982: 6, 16 (in the list of the species of the Sphecidae sects from Korea); Antropov in Nemkov et al., 1995: 420 and 421 known from the Korean Peninsula); Pulawski, 1984: 132 (as (in key, including Korea in distribution); Yamane, 2008: 556 synonym of Trypoxylon figulus); Paik, 1985: 199 (in the list of (including Korea in distribution); Terayama and Nambu, 2009: Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of Insects 30e31, 36 (including Korea in distribution); Nemkov, 2009:89 from Korea); Lee et al., 2010: 208 (in Checklist of Korean Insects). (including Korea in distribution); Lee et al., 2010: 209 (in Trypoxylon koma Tsuneki: Antropov, 1987: 58 (full species status); Checklist of Korean Insects); Nemkov, 2012b: 440 (including Antropov in Nemkov et al., 1995: 419 (in key, including Korea in Korea in distribution). distribution); Nemkov, 2009: 87 (including Korea in distribu- Distribution. S Korea (GG, JJ), Russian Far East, Japan. tion); Nemkov, 2012b: 439 (including Korea in distribution). Distribution. S Korea (Odaesan Mountain, middle part of the Trypoxylon petiolatum F. Smith, 1858 곰보어리나나니 Korean Peninsula), Russian Far East. Trypoxylon petiolatum F. Smith, 1858: 105, \ (lectotype, designated by Tsuneki, 1978: 6), Malaysia: Sarawak [OXUM]; Tsuneki, 1982: Trypoxylon koreanum Tsuneki, 1956 장안어리나나니 16 (in the list of the species of the Sphecidae known from the Trypoxylon koreanum Tsuneki, 1956a: 11 (in key), 32, _ (holotype), Korean Peninsula); Paik, 1985: 199 (in the list of Sphecidae of Korea: “Keijo” [USNM]; Kim, 1970: 585 (diagnosis, no Korean Korea); Terayama and Nambu, 2009: 31 and 36 (including Korea material), 811 (in the list of Hymenoptera from Korea); Bohart in distribution). and Menke, 1976: 347 (including Korea in distribution); Trypoxylon obsonator F. Smith, 1873: 194, \_(syntypes), Japan: Tsuneki, 1981:38e40 (supplement description based on types); Hyogo [BMNH], synonymized with Trypoxylon petiolatum by Tsuneki, 1982: 16 (in the list of the species of the Sphecidae Tsuneki, 1981:64;Tsuneki, 1968b: 50 (Korea: Jeju Is.: city of known from the Korean Peninsula); Paik, 1985: 199 (in the list of Jeju); Kim et al., 1994: 261 (in Check List of Insects from Korea). 440 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Trypoxylon bicolor obsonator Smith: Lee et al., 2010: 209 (in type); Tsuneki, 1982: 16 (in the list of the species of the Sphe- Checklist of Korean Insects). cidae known from the Korean Peninsula, erroneously placed in Distribution. S Korea (JJ), N China, Japan; SE Asia. Pison); Paik, 1985: 199 (subspecific name listed incorrectly as nastus); Tsuneki, 1991: 200 [as new subspecies (nomen nudum) Trypoxylon regium Gussakovskij, 1932 왕어리나나니(신칭) in Sphecidae from Korea]. Trypoxylon regium Gussakovskij, 1932:11,\ (holotype), Russia: Distribution. N Korea (GG), China, Russian Far East, Japan. Primorskiy Kray: Sutshan [ZIN]; Antropov in Nemkov et al., 1995: 419 (in key, as regium regium, including Korea in distri- Subfamily Mellinae Latreille, 1802 파리잡이벌아과 bution); Nemkov, 2009: 88 (including Korea in distribution); Tribe Mellinini Latreille, 1802 파리잡이벌족 Nemkov, 2012b: 440 (including Korea in distribution). Genus Mellinus Fabricius, 1790 파리잡이벌속 Distribution. ?Korea, Russian Far East, Japan, Taiwan. Mellinus Fabricius, 1790: 226. Type species: Vespa arvensis Linnaeus, Note. As far as I am aware, no Korean specimen was examined or 1758, designated by Curtis, 1836: text for Pl. 580. designated in taxonomic or related publications. Mellinus crabroneus (Thunberg, 1787) 홍다리파리잡이벌 Trypoxylon rufimaculatum Antropov, 1987 붉은어리나나니(신칭) Sphex sabulosus Fabricius, 1787: 296, sex not indicated, \ (lectotype, Trypoxylon varipes rufimaculatum Antropov, 1987:57,\ (holotype), designated by van der Vecht, 1961: 45), Denmark: Hafniae, now Russia: Primorskiy Kray: 40 km SE Ussuriysk [ZMMU]. Copenhagen [ZMK], junior secondary homonym of Sphex sabu- Trypoxylon rufimaculatum Antropov: Antropov, 1995: 124 (full losus Linnaeus, 1758, now in Ammophila (Article 59.3: a junior species status, faunal records from Japan and Russian Far East); secondary homonym replaced before 1961 is permanently Terayama and Nambu, 2009: 32 (including Korea in invalid). distribution). Mellinus sabulosus (Fabricius): Tsuneki, 1965a: 26 (including Korea Distribution. ?Korea, China, Russian Far East, Japan. in distribution); Kim, 1970: 610 (diagnosis, no Korean material), Note. As far as I am aware, no Korean specimen was examined or 815 (in the list of Hymenoptera from Korea); Kim et al., 1994: designated in taxonomic or related publications. Nemkov (2009, 262 (in Check List of Insects from Korea); Lee et al., 2010: 210 (in 2012b) did not included Korea in distribution. Checklist of Korean Insects). Sphex crabronea Thunberg, 1791: 126, substitute name for Sphex Trypoxylon sapporoense Tsuneki, 1960 삿뽀로어리나나니(신칭) sabulosus (Fabricius, 1787). Trypoxylon sapporoense Tsuneki, 1960a:241,_ (holotype), Japan: Mellinus crabroneus (Thunberg): Siri and Bohart, 1974: 170 (new Hokkaido, Sapporo [MNHAH]; Tsuneki, 1981: 3 (in key), 9e12; combination), 174; Tsuneki, 1982: 18 (in the list of the species of Tsuneki, 1982: 16 (in the list of the species of the Sphecidae the Sphecidae known from the Korean Peninsula); Paik, 1985: known from the Korean Peninsula); Paik, 1985: 199 (in the list of 203 (in the list of Sphecidae of Korea); Nemkov in Nemkov et al., Sphecidae of Korea); Terayama and Nambu, 2009: 32, 36 1995: 455 (in key, including Korea in distribution); Nemkov, (including Korea in distribution). 2009: 126 (including Korea in distribution); Nemkov, 2012b: Trypoxylon pappi Tsuneki, 1974: 363, \ (holotype), N Korea: 20 km 445 (including Korea in distribution). NE Kaesong, Mts. Pakyon [TMB], synonymized with Trypoxylon Distribution. Transpalearctic: Korea (no specific locality), Europe sapporoense by Tsuneki, 1981:9;Bohart and Menke, 1976: 630 to Mongolia, NE China, Russian Far East, Japan. (including Korea in distribution). Distribution. N Korea (GG), Japan. Mellinus obscures Handlirschch, 1888 파리잡이벌 Mellinus obscurus Handlirsch, 1888a: 289, \ (holotype), Korea: no Trypoxylon ussuriense Kazenas, 1980 우수리어리나나니(신칭) specific locality [NMK]; Dalla Torre, 1897: 561 (including Korea Trypoxylon ussuriense Kazenas, 1980:91:\ (holotype), Russia: Pri- in distribution); Maidl and Klima, 1939: 39 and 43 (including morskiy Kray: Anisimovka (ZIL); Antropov, 1986a: 87 (descrip- Korea in distribution); Yasumatsu, 1943: 2 (new description of tion of _); Antropov in Nemkov et al., 1995: 416 (in key, _), 3 (in key, Korean material as nominotypical subspecies); including Korea in distribution). Tsuneki, 1965a: 26 (including Korea in distribution); Kim, 1970: Distribution. ?Korea, Russian Far East. 610 (diagnosis, no Korean material), 815 (in the list of Hyme- Note. Nemkov et al. (1995) included Korea in distribution of this noptera from Korea); Siri and Bohart, 1974: 170, 174; Bohart and species. Although correction on distributional information pro- Menke, 1976: 449 (including Korea in distribution); Tsuneki, vided by Nemkov et al. (1995) was not stated, later Nemkov (1999, 1982: 18 (specific name incorrectly as obscrus in the list of the 2012b) clearly regarded this species to be endemic to the Russian species of the Sphecidae known from the Korean Peninsula); Far East (Primorie Krai). It is more reasonable for me to exclude this Paik, 1985: 202 (in the list of Sphecidae of Korea); Kim et al., species from the Korean fauna. 1994: 262 (in Check List of Insects from Korea); Gupta, Gayubo and Pulawski, 2008: 210 (full species status restored, diag- Trypoxylon varipes Pérez, 1905 갈색어리나나니 nosis); Nemkov, 2009: 126 (including Korea in distribution); Trypoxylon varipes Pérez, 1905a: 157, \ (syntypes), Japan: no Tano, 2010b: 46 (full species status); Nemkov, 2012: 445 specific locality but presumably Tokyo area [MNHN]; Tsuneki, (including Korea in distribution). 1973: 34 and 37e38 (including Korea in distribution); Mellinus arvensis obscures Handlirschch: Nemkov in Nemkov et al., Yamane in Yamane et al., 1999: 528 (including Korea in dis- 1995: 455 (in key, new status, including Korea in distribution); tribution); Terayama and Nambu, 2009: 34, 36 (including Lee et al., 2010: 210 (in Checklist of Korean Insects). Korea in distribution); Nemkov, 2009: 90 (including Korea in Distribution. Korea (no specific locality), Russian Far East, China, distribution); Nemkov, 2012b: 440 (including Korea in Japan. distribution). Trypoxylon varipes nasutum Tsuneki, 1974: 365, \ (holotype), Korea: Subfamily Pemphredoninae Dahlbom, 1835 진딧물벌아과 27 km NE Kaesong, Mts. Pakyon [TMB], synonymized with Tribe Pemphredonini Dahlbom, 1835 진딧물벌족 Trypoxylon varipes by Antropov, 1995: 124; Bohart and Menke, Subtribe Pemphredonina Dahlbom, 1835 진딧물벌아족 1976: 630 (listed); Tsuneki, 1981: 4 (in key), 38 (based on JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 441

Genus Diodontus Curtis, 1834 조선꼬마구멍벌속 monilicornis collectively corrected to Passaloecus insignis Diodontus Curtis, 1834 (text for plate 496). Type species: Pem- (Linden) by Yarrow, 1970: 178. phredon tristis Linden, 1829, designated by the ICZN, Opinion Passaloecus insignis (Linden): Shuckard, 1837: 189 (new combina- 844, 1968: 10. tion); Kim, 1970: 813 (in the list of Hymenoptera from Korea); Kim et al.,1994: 261 (in Check List of Insects from Korea); Budrys Diodontus chosenensis Tsuneki, 1974 조선꼬마구멍벌 in Nemkov et al., 1995: 403 (in key, including Korea in distri- Diodontus chosenensis Tsuneki, 1974: 370, \ (holotype), North Ko- bution); Kim and Yang, 2010: 146 (in the checklist of valid rea: “Prov. Ryang-gang: Hyesan” [TMB]; Bohart and Menke, Korean species of the Subfamily Pemphredoninae); Lee et al., 1976: 629 (including Korea in distribution); Tsuneki, 1982:14 2010: 209 (in Checklist of Korean Insects); Terayama, 2012:11, (in the list of the species of the Sphecidae known from the 18 (including Korea in distribution); Nemkov, 2012b:437 Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of (including Korea in distribution). Korea); Tsuneki, 1991: 201 (as new species, nomen nudum); Kim Distribution. Transpalearctic: N Korea (GW), Europe to NE China, et al., 1994: 261 (in Check List of Insects from Korea); Budrys in Russian Far East, Japan. Nemkov et al., 1995: 398 (including Korea in distribution); Kim and Yang, 2010: 148 (in the checklist of valid Korean species of Passaloecus koreanus Tsuneki, 1974 한국꼬마구멍벌 the Subfamily Pemphredoninae); Lee et al., 2010: 209 (in Passaloecus annulatus koreanus Tsuneki, 1974:368,_ (holotype) and Checklist of Korean Insects). \, Korea: Prov. “Ryang-gang: Hyesan” [TMB]; Tsuneki, 1991:201 Distribution. N Korea (RG). (as new subspecies, nomen nudum); Kim et al., 1994: 261 (in Check List of Insects from Korea); Lee et al., 2010: 209 (in Diodontus minutus orientalis (Tsuneki, 1974) 동양꼬마구멍벌 Checklist of Korean Insects). Diodontus minutus orientalis Tsuneki, 1974: 369, \ (holotype), North Passaloecus koreanus Tsuneki, 1982: 15 (full species status); Paik, Korea: “Prov. Ryang-gang: Hyesan” [TMB]; Bohart and Menke, 1985: 198 (in the list of Sphecidae of Korea); Budrys in 1976: 629 (including Korea in distribution); Tsuneki, 1982:14 Nemkov et al., 1995: 404 (in key, including Korea in distribu- (in the list of the species of the Sphecidae known from the tion); Nemkov, 2009: 69; Kim and Yang, 2010: 146 (in the Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of checklist of valid Korean species of the Subfamily Pem- Korea); Lee et al., 2010: 209 (in Checklist of Korean Insects). phredoninae); Terayama, 2012:11e12, 18 (including Korea in Diodontus minutes (Fabricius): Budrys in Nemkov et al., 1995: 399 distribution); Nemkov, 2012b: 437 (including Korea in (in key, including Korea in distribution); Kim et al., 1994: 261 (in distribution). Check List of Insects from Korea); Nemkov, 2009:62e63 Distribution. N Korea (RG), Russian Far East, Japan. (including Korea in distribution); Kim and Yang, 2010: 148 (in the checklist of valid Korean species of the Subfamily Pem- Passaloecus monilicornis Dahlbom, 1842 염주꼬마구멍벌 phredoninae); Nemkov, 2012b: 436 (including Korea in Passaloecus monilicornis Dahlbom, 1842:12,\ and _ (lectotype, distribution). designated by Yarrow, 1970: 169), Sweden: Helsingland: Kår- Distribution. N Korea (RG). böle: Skansberget [MZLU]; Tsuneki, 1955: 3 and 5 (in key, including Korea in distribution); Kim, 1970: 598 (diagnosis, no Genus Passaloecus Shuckard, 1837 꼬마구멍벌속 Korean material), 813 (in the list of Hymenoptera from Korea); Passaloecus Shuckard, 1837: 188. Type species: Pemphredon insignis Tsuneki, 1982: 3 (N Korea: “Prov. South Pyongan: Nan-po”), 14 Linden, 1829, by original designation. (in list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of Korea); Passaloecus clypealis Faester, 1947 이마방패꼬마구멍벌 Kim et al.,1994: 261 (in Check List of Insects from Korea); Budrys Passaloecus clypealis Faester, 1947:204,_ (holotype) and \, in Nemkov et al., 1995: 403 (in key, including Korea in distri- Denmark: Basnaes [ZMUC]; Tsuneki, 1974: 368 (new to Korea, N bution); Nemkov, 2009: 70 (including Korea in distribution); Korea: “Prov. South Pyongan: Chang-lyong san”); Tsuneki, 1982: Kim and Yang, 2010: 146 (in the checklist of valid Korean species 14 (in the list of the species of the Sphecidae known from the of the Subfamily Pemphredoninae); Lee et al., 2010: 209 (in Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of Checklist of Korean Insects); Nemkov, 2012b: 437 (including Korea); Tsuneki, 1991:201;Kim et al., 1994: 261 (in Check List of Korea in distribution). Insects from Korea); Burdrys in Nemkov et al., 1995: 403 (in key, Distribution. Transpalearctic: N Korea (PN), Europe to NE China, including Korea in distribution); Nemkov, 2009: 68 (including Russian Far East, Japan. Korea in distribution); Kim and Yang, 2010: 146 (in the checklist Note. According to Pulawski (2014), this species may be a syn- of valid Korean species of the Subfamily Pemphredoninae); Lee onym or a northern subspecies of the species P. insignis. Also, he et al., 2010: 209 (in Checklist of Korean Insects); Terayama, supposed that specimens recorded prior to Yarrow (1970) were 2012: 10, 18 (including Korea in distribution); Nemkov, 2012b: mostly referred to as Passaloecus insignis. 436 (including Korea in distribution). Distribution. Transpalearctic: N Korea (PN), Europe to Russian Genus Pemphredon Latreille, 1796 진딧물벌속 Far East, Japan. Pemphredon Latreille, 1796: 128 (no species included). Type species: Pemphredon lugubris (Fabricius, 1804)(¼ Crabro lugubris Fab- Passaloecus insignis (Linden, 1829) 꼬마구멍벌 ricius, 1793), designated by Shuckard, 1837: 193 (one of the two Pemphredon insignis Linden, 1829:81,\ and _ (syntypes, and species first included by Latreille, 1802a: 342). _ ¼ Passaloecus gracilis), Belgium: Bruxelles [Inst. Roy. Sci. Nat. Belgique, Bruxelles]. Pemphredon diervillae Iwata 애진딧물벌, 애기진딧물벌 Passaloecus shuckardi Yasumatsu, 1934a: 36, \, Korea: “Kôgendo” Pemphredon diervillae Iwata, 1933b: 45, \ (holotype) _, Japan: (gangwon-do): “Kongôsan” (Mt. Geumgangsan) (depository Honshu: Settsu: Katioji [KUK]; Kim, 1970: 592 (diagnosis, no uncertain), synonymized with Passaloecus monilicornis var. examination of Korean material, but stating Mt. Yongmunsan as dahlbomi by Yasumatsu (1934b: 113) and Passaloecus collecting locality); Paik, 1985: 197 (specific name incorrectly 442 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

listed as dievillae); Kim et al., 1994: 262 (in Check List of Insects Distribution. S Korea (JJ), Nepal, Russian Far East, Japan. from Korea); Lee et al., 2010: 209 (in Checklist of Korean Insects). Distribution. ?Korea (no specific locality), Japan. Pemphredon koreana Tsuneki, 1951 고려진딧물벌 Pemphredon koreana Tsuneki, 1951: 171 (in key), 183e184 (as Pemphredon flavistigma Thomson, 1874 황무늬진딧물벌 koreanus, incorrect original termination), \ (holotype), Korea: Pemphredon flavistigma Thomson, 1874: 192, \ (holotype), Sweden: “Shoyozan” [MNHAH]; Tsuneki, 1964d: 28 and 29 (as koreanus, Småland: Sällsynt [MZLU]; Tsuneki, 1951:174e176 (Korea: “Mt. in key, including Korea in distribution); Kim, 1970: 593 (specific Hakto”); Tsuneki, 1964d: 28 and 29 (in key, including Korea in name listed incorrectly as koreanus, diagnosis, no Korean ma- distribution); Kim, 1970: 593 (diagnosis, no Korean material); terial); Bohart and Menke, 1976: 181 (including Korea in dis- Paik, 1985: 197 (in the list of Sphecidae of Korea); Tsuneki, 1982: tribution); Tsuneki, 1982: 14 (specific name incorrectly listed as 14 (in the list of the species of the Sphecidae known from the koreanus, including Korea in distribution); Paik, 1985:197 Korean Peninsula); Kim et al., 1994: 261 (in Check List of Insects (specific name incorrectly listed as koreanus, in the list of from Korea); Nemkov, 2009: 64 (including Korea in distribu- Sphecidae of Korea); Kim et al., 1994: 261 (in Check List of In- tion); Kim and Yang, 2010: 147 (in the checklist of valid Korean sects from Korea); Dollfuss, 1995: 975 (as possible variety of species of the Subfamily Pemphredoninae); Lee et al., 2010: 209 podagrica); Budrys in Nemkov et al., 1995: 400 (in key, specific (in Checklist of Korean Insects); Terayama, 2012:13,19 name incorrectly listed as koreanus, including Korea in distri- (including Korea in distribution); Nemkov, 2012b:437 bution); Kim and Yang, 2010: 147 (in the checklist of valid (including Korea in distribution). Korean species of the Subfamily Pemphredoninae); Lee et al., Distribution. Transpalearctic: N Korea (RG), Europe to Russian 2010: 209 (in Checklist of Korean Insects). Far East, Japan. Distribution. S Korea (GG).

Pemphredon inornata Say, 1824 슈카아드진딧물벌 Pemphredon lethifer (Shuckard, 1837) 북방진딧물벌 Pemphredon inornata Say, 1824: 339 (as inornatus, incorrect original Cemonus lethifer Shuckard,1837:201,\ and _ (syntypes), Britain: no termination), sex not indicated (holotype or syntype), USA: specific locality [W. Shuckard coll.]. Pennsylvania (destroyed); Tsuneki, 1982: 3 (as inornatus,NKo- Pemphredon lethifer (Shuckard): Schenck, 1867a: 353 (new combi- rea: “Prov. South Pyongan: Pyongan; Prov. Pyong-sung: Bek- nation); Tsuneki, 1974: 367e368 (N Korea: “Prov. South Pyngan: sung-li”), 14 (in the list of the species of Sphecidae known from Bongwa-ri, Chang-lyong san”); Tsuneki, 1976a: 298 (including the Korean Peninsula); Budrys in Nemkov et al., 1995: 401 (in Korea in distribution); Tsuneki, 1982: 14 (in the list of the spe- key, including Korea in distribution); Nemkov, 2009:64e65 cies of the Sphecidae known from the Korean Peninsula); Paik, (including Korea in distribution); Kim and Yang, 2010: 147 (in 1985: 197 (in the list of Sphecidae of Korea); Tsuneki, 1991: the checklist of valid Korean species of the Subfamily Pem- 201 (listed); Kim et al., 1994: 262 (in Check List of Insects from phredoninae); Terayama, 2012: 13, 19 (including Korea in dis- Korea); Budrys in Nemkov et al., 1995: 401 (in key, including tribution); Nemkov, 2012b: 437 (including Korea in Korea in distribution); Nemkov, 2009: 66 (including Korea in distribution). distribution); Kim and Yang, 2010: 147 (in the checklist of valid Cemonus shuckardi A. Morawitz, 1864: 460 (as Shuckardi, incorrect Korean species of the Subfamily Pemphredoninae); Terayama, original capitalization), \ and _ (syntypes), Russia: St. Peters- 2012:13e14, 19 (including Korea in distribution); Lee et al., burg (lost), synonymized with Pemphredon inornata Say by 2010: 209 (in Checklist of Korean Insects); Nemkov, 2012b: Lomholdt, 1975: 99. 437 (including Korea in distribution). Pemphredon shuckardi (A. Morawitz): Kohl, 1890a: 56 (new com- Pemphredon lethifer fabricii (Müller): Tsuneki, 1951:198e204 (Ko- bination); Tsuneki, 1951:190e194 (Korea: “Keijo, Hakuto-zan”); rea: “Keijo”), corrected Pemphredon lethifer (Shuckard) by Tsuneki, 1964d: 28 and 29 (in key, including Korea in distribu- Tsuneki, 1974: 367. tion); Kim, 1970: 592 (diagnosis, no Korean material); Tsuneki, Distribution. Holarctic: Korean Peninsula (PN, SL, GG), Europe to 1974: 368 (N Korea: “Prov. South Pyngan: Pyongyan”); NE China, Russian Far East, Japan, Canada, USA. Tsuneki, 1976a: 298 (including Korea in distribution); Paik, 1985: 197 (in the list of Sphecidae of Korea); Tsuneki, 1991: Pemphredon lugubris (Fabricius, 1793) 우수리진딧물벌 201 (in Sphecidae from Korea); Kim et al., 1994: 261 (in Check Crabro lugubris Fabricius, 1793: 302, \ (lectotype, designated by van List of Insects from Korea); Lee et al., 2010: 209 (in Checklist of der Vecht, 1961: 28), Germany: Halae Saxonum, now Sachsen- Korean Insects). Anstalt: Halle [ZMK]. Distribution. Holarctic: Korean Peninsula (RG, PN, SL), Europe to Pemphredon lugubris (Fabricius): Fabricius, 1804: 315 (new combi- NE China, Russian Far East, Japan, Canada, USA. nation, redescription); Budrys in Nemkov et al., 1995: 400 (in key, including Korea in distribution); Nemkov, 2009:66 Pemphredon japonica Matsumura, 1912 왕진딧물벌 (including Korea in distribution); Kim and Yang, 2010: 147 (in Pemphredon japonica Matsumura, 1912: 179 (as japonicum, incor- the checklist of valid Korean species of the Subfamily Pem- rect original termination), \ (holotype or syntypes), Japan: phredoninae); Terayama, 2012: 14, 19 (including Korea in dis- Honshu: Kyoto (lost); Tsuneki, 1968b: 50 (specific name incor- tribution); Nemkov, 2012b: 437 (including Korea in rectly listed as japonicus, Korea: Jeju Is.: Mt. Hanna); Paik, 1985: distribution). 197 (specific name incorrectly listed as japonicas in the list of Pemphredon pacifica Gussakovskij, 1932:8(aspacificus,incorrect Sphecidae of Korea); Kim et al., 1994: 261 (specific name original termination), \ and _ (syntypes), Russia: Kamchatka: incorrectly listed as japonicus, in Check List of Insects from Ko- Elisovo [OMNH] and Primorskiy Kray: Khabarovsk [ZIN], synon- rea); Nemkov, 2009: 65 (including Korea in distribution); Kim ymized with Pemphredon lugubris by Lomholdt, 1975:89; and Yang, 2010: 147 (in the checklist of valid Korean species of Tsuneki, 1951:179e181 (specific name incorrectly listed as the Subfamily Pemphredoninae); Lee et al., 2010: 209 (in pacificus,Korea:“Hakotozan”); Lee et al., 2010: 209 (specificname Checklist of Korean Insects); Nemkov, 2012b: 437 (including incorrectly listed as pacificus, in Checklist of Korean Insects). Korea in distribution). Pemphredon lugubris pacificus Gussakovskij: Tsuneki, 1964d: 28 and 29 (in key, new status, including Korea in distribution); Kim, JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 443

1970: 593 (diagnosis, no Korean material); Tsuneki, 1974: 367 (N species of the Subfamily Pemphredoninae); Terayama, 2012: Korea: “Prov. Ryang-gang: Hyesan; Prov. South Pyongan: Lyong- 14e15 and 19 (including Korea in distribution); Nemkov, 2012b: ak san”); Tsuneki, 1982: 14 (in the list of the species of the 437 (including Korea in distribution). Sphecidae known from the Korean Peninsula); Paik, 1985:197 Pemphredon wesmaeli (Morawitz): Lee et al., 2010: 209 (in Checklist (in the list of Sphecidae of Korea); Kim et al., 1994: 261 (in Check of Korean Insects). List of Insects from Korea); Tsuneki, 1991: 201 (in Sphecidae Pemphredon rugifer wesmaeli (A. Morawitz): Beaumont, 1951: 34 (as from Korea); Kim and Yang, 2010: 147 (in the checklist of valid rugifer f. wesmaeli, Switzerland: Neuchâtel area); Tsuneki, 1974: Korean species of the Subfamily Pemphredoninae). 367 (N Korea: “Prov. Ryang-gang: Hyesan”); Tsuneki, 1982:14 Distribution. Holarctic: N Korea (RG, PN), Europe to Russian Far (in the list of the species of the Sphecidae known from the East, Japan, Canada, USA. Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of Korea); Kim et al., 1994: 262 (specific name incorrectly listed as Pemphredon montana Dahlbom, 1845 진딧물벌 westmaeli, in Check List of Insects from Korea); Tsuneki, 1991: Pemphredon montana Dahlbom, 1845:262(asmontanus,incorrect 201 (in Sphecidae from Korea). original termination), \ (lectotype, designated by Dollfuss, 1995: Pemphredon unicolor wesmaeli A. Morawitz: Tsuneki, 1951:194e 994) and _, Sweden: Faxälven [MZLU]; Tsuneki, 1951:181e183 198 (Korea: “Keijo, Shyozan, Nansetsurei, Daitaku”); Tsuneki, (specific name incorrectly listed as montanus, NKorea:“Nanset- 1964d: 28 and 29 (in key, including Korea in distribution); surei”); Tsuneki, 1964d: 28 (specific name incorrectly listed as Kim, 1970: 592 (diagnosis, no Korean material). montanus, in key, including Korea in distribution); Yasumatsu, Pemphredon mortifer Valkeila in Valkeila and Leclercq, 1972: 697, \ 1965: 298 (specific name incorrectly listed as montanus, (holotype) and _, Russia: Karelian Isthmus: Metsäpirtti, now including Korea in distribution); Kim, 1970: Pl. 58 Figure 698, 594 Zaporozhskoye 75 km NW St. Petersburg [ZMUT], synonymized (specific name listed incorrectly as montanus, diagnosis; Korea: with Pemphredon rugifer by Dollfuss, 1991: 62; Bohart and Mt. Yongmunsan); Tsuneki, 1982:14(specificnameincorrectly Menke, 1976: 182 (including Korea in distribution); Paik, 1985: listed as montanus in the list of the species of the Sphecidae 198 (in the list of Sphecidae of Korea); Budrys in Nemkov et al., known from the Korean Peninsula); Paik, 1985: 197 (specific 1995: 402 (in key, including Korea in distribution); Kim and name incorrectly listed as montanus, in the list of the Sphecidae of Yang, 2010:147e148 (in the checklist of valid Korean species Korea); Kim et al., 1994: 261 (specific name incorrectly listed as of the Subfamily Pemphredoninae). montanus, in Check List of Insects from Korea); Budrys in Nemkov Distribution. Holarctic: Korean Peninsula (RG, SL, GG), Europe to et al.,1999: 400 (in key, including Korea in distribution); Nemkov, Russian Far East, Japan, USA. 2009:66e67 (including Korea in distribution); Kim and Yang, Note. Pemphredon wesmaeli (Morawitz, 1864) is controversial in 2010: 147 (in the checklist of valid Korean species of the Sub- its specific validity. It was a synonym of Pemphredon rugifer (Doll- family Pemphredoninae); Terayama, 2012: 14 and 19 (including fuss, 1991), or a valid species (Schmidt and Schmid-Egger, 1997). Korea in distribution); Lee et al., 2010:209(specificname Therefore, at least the subspecific combination of Pemphredn rugifer incorrectly listed as montanus in Checklist of Korean Insects); wesmaeli (Dahlbom), as often used in the Far East, is invalid. Nemkov, 2012b: 437 (including Korea in distribution). Distribution. Holarctic: Korean Peninsula (RG, GG), Europe to Genus Polemistus Saussure, 1892 병신꼬마구멍벌속(신칭) Russian Far East, Japan, Canada, USA. Polemistus Saussure, 1892: 565. Type species: Polemistus macilentus Saussure, 1892, designated by Pate, 1937: 52. Pemphredon morio Linden, 1829 뜨장이진딧물벌 Pemphredon morio Linden 1829: 84, \ (holotype), Belgium: Brux- Polemistus abnormis (Kohl, 1888) 병신꼬마구멍벌 elles (lost); Paik, 1985: 197 (in the list of Sphecidae of Korea); Passaloecus abnormis Kohl, 1888b: 726, \, Slovenia: Vipava Kim et al., 1994: 261 (in Check List of Insects from Korea); Lee [NHMW]; Tsuneki, 1955: 3 and 4 (in key), 12 (including Korea in et al., 2010: 209 (in Checklist of Korean Insects); Terayama, distribution); Kim, 1970: 597 (diagnosis, no Korean material), 2012: 14, 19 (including Korea in distribution). 813 (in the list of Hymenoptera from Korea); Lee et al., 2010: 209 Pemphredon intermedius Tsuneki 1951a: 186 (as intermedius, (in Checklist of Korean Insects). incorrect original termination), \ (holotype) and _, Japan: Sap- Polemistus abnormis (Kohl): Bohart and Menke, 1976: 185 (new poro [USNM], synonymized with Pemphredon morio by Bohart in combination); Tsuneki, 1982: 14 (in the list of the species of the Bohart and Menke, 1976: 182; Tsuneki, 1964d: 27 (in key, Sphecidae known from the Korean Peninsula); Paik, 1985:198 including Korea in distribution); Kim, 1970: 593 (diagnosis, no (in the list of Sphecidae of Korea); Kim et al., 1994: 261 (in Check Korean material), 812 (in the list of Hymenoptera from Korea). List of Insects from Korea); Budrys in Nemkov et al., 1995: 404 Distribution. Transpalearctic: Korea (no specific locality), Europe (in key, including Korea in distribution); Nemkov, 2009:70 to Russian Far East, Japan. (including Korea in distribution); Kim and Yang, 2010: 147 (in the checklist of valid Korean species of the Subfamily Pem- Pemphredon rugifer (Dahlbom, 1884) 단색진딧물벌 phredoninae); Terayama, 2012: 10 and 19 (including Korea in Sphex unicolor Panzer, 1798: 24 (nom. Praeocc, as Crabro unicolor distribution); Nemkov, 2012b: 437 (including Korea in only in the legend for illustration), [\] (holotype), Germany: distribution). Norimbergae, now Nürnberg (depository uncertain), junior Distribution. Transpalearctic: Korea (no specific locality), Europe primary homonym of Sphex unicolor Fabricius, 1787; synony- to Russian Far East, Korea (no specific locality), Japan. mized with Pemphredon lugubris (Fabricius, 1793) by Panzer, 1806: 187 and with Pemphredon rugifer by Dahlbom, 1844: 256. Subtribe Stigmina Bohart and Menke, 1976 호리꼬마구멍벌아족 Cemonus rugifer Dahlbom, 1844: 256, _ (lectotype, designated by Genus Carinostigmus Tsuneki, 1954 용골꼬마구멍벌속 Blüthgen, 1931: 128) and \, Germany: Silesia: Glogavia, now Carinostigmus Tsuneki, 1954a: 3. Type species: Stigmus congruus Poland: Glogów [MZLU]. Walker, 1860, by original designation. Pemphredon rugifer (Dahlbom): Schenck, 1867a: 353 (new combi- nation): Nemkov, 2009:67e68 (including Korea in distribu- Carinostigmus filippovi (Gussakovskij, 1934) 호리꼬마구멍벌 tion); Kim and Yang, 2010: 148 (in the checklist of valid Korean Stigmus filippovi Gussakovskij, 1934b: 83, \ (holotype), Japan: 444 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Honshu: Takao-san (60 km of Tokyo) [ZIN]; Kim, 1970: 598 Mimesa lutaria (Fabricius, 1787) 큰꼬마구멍벌 (diagnosis, no Korean material), 813 (in the list of Hymenoptera Sphex lutarius Fabricius, 1787: 273 (as lutaria, incorrect original from Korea); Paik, 1985: 198 (in the list of Sphecidae of Korea); termination), \ (lectotype, designated by van der Vecht, 1961: Kim et al., 1994: 262 (in Check List of Insects from Korea); Lee 27), Kiliae (now Kiel), Germany [ZMK]. et al., 2010: 209 (in Checklist of Korean Insects). Mimesa lutaria (Fabricius): Dahlbom, 1843: 4 (new combination); Carinostigmus filippovi (Gussakovskij): Finnamore, 1995: 225 (new Budrys in Nemkov et al., 1995: 389e390 (in key, including Korea combination); Kim and Yang, 2009: 466e467 (new to Korea: in distribution); Kim and Yang, 2010: 144 (in the checklist of Gyeonggi-do, Gangwn-do). valid Korean species of the Subfamily Pemphredoninae). Distribution. S Korea (GG, GW), Japan. Distribution. Transpalearctic: Korea (no specific locality), Europe Genus Stigmus Panzer, 1804 호리꼬마구멍벌속 to Russian Far East, Japan. Stigmus Panzer, 1804: 86. Type species: Stigmus pendulus Panzer, Note. Although Nemkov et al. (1995) included Korea in the dis- 1804, by monotypy. tribution, later Nemkov (2012b) did not include Korea in the dis- tribution. Korean occurrence of this species had not been directly Stigmus convergens Tsuneki, 1954 evidenced with a Korean specimen, thus this species should be Stigmus convergens Tsuneki, 1954a: 33, \ (holotype), Japan: Hok- excluded from the Korean fauna. kaido: Akagawa near Kucchian [HMHAN]; Terayama, 2012:16 (including Korea in distribution). Genus Mimumesa Malloch, 1933 어리꼬마구멍벌속 Distribution. ?Korea, Russian Far East, Japan. Mimumesa Malloch, 1933: 16. Type species: Psen niger Packard, Note. Terayama’s (2012) record of Korean occurrence of this 1867, by original designation. species was groundless, probably was merely a mistake. Mimumesa atratina (F. Morawitz, 1891) 어리꼬마구멍벌 Stigmus japonicus Tsuneki, 1954 왜꼬마구멍벌 Mimesa atratina F. Morawitz, 1891:206,_ (holotype or syntypes), Stigmus japonicus Tsuneki, 1954a: 29, \ (holotype) and _ (¼ Stigmus Kazakhstan: Bogdosan [ZIN]. quadriceps), Japan: Hokkaido: Kyogoku-mura near Kucchian Mimumesa atratina (F. Morawitz): Lomholdt, 1975: 158 (new com- [MNHAH]; Kim and Yang, 2009: 468e469 (Korea: Gangwn-do: bination); Budrys in Nemkov et al., 1995: 390 (in key, including Baekdam); Kim and Yang, 2010: 146 (in the checklist of valid Korea in distribution); Nemkov, 2009:54e55 (including Korea Korean species of the Subfamily Pemphredoninae); Terayama, in distribution); Kim and Yang, 2010: 145 (in the checklist of 2012: 16, 20 (including Korea in distribution). valid Korean species of the Subfamily Pemphredoninae); Distribution. S Korea (GW), Russian Far East, Japan. Nemkov, 2012b: 435 (including Korea in distribution). Mimesa longula Gussakovskij, 1932:5,_ (holotype), Russia: Pri- GenusTzustigmus Finnamore, 1995 무소꼬마구멍벌속 moskiy Kray: Sutshan [OMNH], synonymized with Mimumesa Tzustigmus Finnamore, 1995: 211. Type species: Tzustigmus syam atratina by de Beaumont, 1941a: 328 and Budrys, 1990: 944. Finnamore, 1995, original designation. Psen atratinus longulus (Gussakovskij): Tsuneki, 1964c: 10 (new subspecific combination); Tsuneki, 1974: 366 (new to Korea, N Tzustigmus rhinoceros (Budrys, 1987) 무소꼬마구멍벌(신칭) Korea: “Prov. South Pyongan: Pyongyang”); Tsuneki, 1982: 15 (in Carinostigmus rginocerus Budrys, 1987: 54, \ (holotype) and _, the list of the species of the Sphecidae known from the Korean Russia: Primorskiy Krai: Kabarovsk, Khekhtsir Range [ZIN]. Peninsula); Paik, 1985: 198 (in the list of Sphecidae of Korea); Tzustigmus rhinoceros (Budrys): Finnamore, 1995: 212 (new com- Tsuneki, 1991: 200 (in Sphecidae from Korea); Kim et al., 1994: bination); Kim and Yang, 2009:467e468 (new to Korea: 262 (in Check List of Insects from Korea); Lee et al., 2010: 209 (in Gangwon-do: Hongcheon-gun); Kim and Yang, 2010: 146 (in the Checklist of Korean Insects). checklist of valid Korean species of the Subfamily Distribution. Holarctic: N Korea (PN), Europe to Mongolia, Pemphredoninae). Russian Far East, Japan, Canada. Distribution. S Korea (GW), Russian Far East. Mimumesa dahlbomi (Wesmael, 1852) 달봄먹꼬마구멍벌 Tribe Psenini A Costa, 1858 먹꼬마구멍벌족 Mimesa dahlbomi Wesmael, 1852: 271 (as Dahlbomi, incorrect Genus Mimesa Shuckard 큰꼬마구멍벌속 original capitalization), \ and _ (lectotype, designated by Lith, Mimesa Shuckard, 1937: 228. Type species: “Psen equestris F.” (¼ 1949: 139), Belgium: Bruxelles area (Inst. Roy. Sci. Nat. Psen equestris of Latreille, 1819 ¼ Trypoxylon equestre Fabricius, Bruxelles). 1804), by original designation. Pesn dahlbomi (Wesmael): Tsuneki, 1974: 366 (new to Korea, N Korea: “Prov. Ryang-gang; Chan-Pay Plateau”); Tsuneki, 1982:15 Mimesa equestris (Fabricius, 1804) 붉은띠먹꼬마구멍벌 (in the list of the species of the Sphecidae known from the Trypoxylon equestre Fabricius, 1804: 182, sex not indicated, _ (ho- Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of lotype), no specific locality of Germany [ZMK]. Korea); Tsuneki, 1991: 200 (in Sphecidae from Korea); Kim et al., Psen equestris (Fabricius): Panzer, 1806: 110 (new combination); 1994: 262 (in Check List of Insects from Korea); Lee et al., 2010: Tsuneki, 1982: 19 (in the list of the species of the Sphecidae 209 (in Checklist of Korean Insects). known from the Korean Peninsula); Paik, 1985: 198 (in the list of Mimumesa dahlbomi (Wesmael): Lomholdt, 1975c: 160 (new com- Sphecidae of Korea). bination); Budrys in Nemkov et al., 1995: 390 (in key, including Mimesa equestris (Fabricius): Wesmael, 1852: 272 (new combina- Korea in distribution); Nemkov, 2009: 55 (including Korea in tion); Budrys in Nemkov et al., 1995: 388 (in key, including Korea distribution); Nemkov, 2012b: 435 (including Korea in distri- in distribution); Kim and Yang, 2010: 144 (in the checklist of bution); Kim and Yang, 2010: 145 (in the checklist of valid valid Korean species of the Subfamily Pemphredoninae). Korean species of the Subfamily Pemphredoninae). Distribution. Transpalearctic: Korea, Europe to Mongolia, Distribution. Transpalearctic: N Korea (RG), Europe to Russian Russian Far East. Far East, Japan. JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 445

Mimumesa littoralis (Bondroit, 1934) 해변어리꼬마구멍벌 (new to Korea, N Korea: Kaesong: Mts. Pakyon); Tsuneki, 1982: Mimesa littoralis Bondroit, 1934: 61, 64, 65, \ and _ (lectotype, 15 (in the list of the species of the Sphecidae known from the designated by Leclercq, 1974a: 194), Belgium: Ostende [IRSN]. Korean Peninsula); Tsuneki, 1991: 200 (in Sphecidae from Ko- Psen littoralis (Bondroit): Tsuneki, 1974: 366 (N Korea: South rea); Kim et al., 1994: 262 (in Check List of Insects from Korea); Phennan: Pyeongyan, De-sang san); Tsuneki, 1982:15 (in the list Nemkov, 2009: 58 (including Korea in distribution); Kim and of the species of the Sphecidae known from the Korean Penin- Yang, 2010: 145 (in the checklist of valid Korean species of the sula); Paik, 1985: 198 (in the list of Sphecidae of Korea); Tsuneki, Subfamily Pemphredoninae); Lee et al., 2010: 209 (in Checklist 1991: 200 (in Sphecidae from Korea); Kim et al., 1994: 262 (in of Korean Insects); Nemkov, 2012b: 435 (including Korea in Check List of Insects from Korea); Lee et al., 2010: 209 (in distribution). Checklist of Korean Insects). Psen caocinnus Tsuneki 1973c: 113 (invalid: Article 59.4), substitute Mimumesa littoralis (Bondroit): Lomholdt, 1975c: 157 (new com- name for Psen aurifrons Tsuneki, 1959, junior secondary hom- bination); Budrys in Nemkov et al., 1995: 391 (in key, including onym of Psen aurifrons (Taschenberg, 1875), now in Pseneo; Korea in distribution); Nemkov, 2009: 55 (including Korea in Tsuneki, 1982: 15 (in the list of the species of the Sphecidae distribution); Kim and Yang, 2010: 145 (in the checklist of valid known from the Korean Peninsula); Paik, 1985: 198 (in the list of Korean species of the Subfamily Pemphredoninae). Sphecidae of Korea); Yamane in Yamane et al., 1999:489e490 Distribution. Transpalearctic: N Korea (PN), Europe to Mongolia, (including Korea in distribution). Russian Far East, Japan. Distribution. N Korea (GG), Russian Far East, Japan.

Genus Psen Latreille, 1976 먹꼬마구멍벌속 Psen koreanus Tsuneki, 1959 고려꼬마구멍벌 Psen Latreille, 1796: 122 (no included species). Type species: Sphex Psen koreanus Tsuneki, 1959a:73e74, \ (holotype), Korea: “Keijo” ater Fabricius, 1794 (¼ Crabro ater Olivier, 1792), designated by [USNM]; Kim, 1970; 596e597 (diagnosis, no Korean material), Latreille, 1802a: 338 (first included species). 812 (in the list of Hymenoptera from Korea); Bohart and Menke, 1976: 166 (listed); Tsuneki, 1982: 15 (in the list of the species of Psen affinis Gussakovskij, 1937 구사코브스키꼬마구멍벌 the Sphecidae known from the Korean Peninsula); Kim et al., Psen affinis Gussakovskij, 1937: 652, \ (holotype), Russia: Primor- 1994: 262 (in Check List of Insects from Korea); Budrys in skiy Kray: Maikhinsk forestry station (depository uncertain); Nemkov et al., 1995: 393 (in key, including Korea in dis- Tsuneki, 1959a:65e66 (N Korea: “Daitaku, Nansetsurei”); Kim, tribution);Yamane in Yamane et al., 1999: 490 (including Korea 1970: 595e596 (diagnosis, no Korean material), 812 (in the in distribution); Nemkov, 2009: 58 (including Korea in distri- list of Hymenoptera from Korea); Budrys, 1986: 142 and 145 bution); Kim and Yang, 2010: 145 (in the checklist of valid (including Korea in distribution); Kim et al., 1994: 262 (in Check Korean species of the Subfamily Pemphredoninae); Lee et al., List of Insects from Korea); Budrys in Nemkov et al., 1995: 394 2010: 209 (in Checklist of Korean Insects); Nemkov, 2012b: (in key, including Korea in distribution); Yamane in Yamane 436 (including Korea in distribution). et al., 1999: 490 (including Korea in distribution); Budrys, Distribution. S Korea (SL), Russian Far East, Japan. 2000: 64 (including Korea in distribution); Nemkov, 2009:57; Kim and Yang, 2010: 144 (in the checklist of valid Korean species Psen seminitidus Lith, 1934 광채꼬마구멍벌 of the Subfamily Pemphredoninae); Lee et al., 2010: 209 (in the Mimesa kohli Gussakovskij, 1934a:7,\ (holotype), China: Tibet: checklist of Korean Insects); Nemkov, 2012b: 435 (including Affluent of Mekong, River Chok-Chyu [ZIN], junior secondary Korea in distribution). homonym of Psen kohli Fox, 1898. Distribution. N Korea (RG), NE China, Russian Far East, Japan. Psen seminitidus Lith, 1965: 40, substitute name for Psen kohli (Gussakovskij, 1934); Budrys in Nemkov et al., 1995: 393 (in key, Psen ater (Olivier, 1792) 먹꼬마구멍벌 including Korea in distribution); Nemkov, 2009:58e59 Crabro ater Olivier, 1792:517,\ (neotype, designated by Leclercq, (including Korea in distribution). 1974b: 259), France: Yvelines, Feucherolles [MNHN]. Psen takanensis Tsuneki, 1978c: 81, substitute name for Psen alticola Psen ater (Oliver): Spinola, 1806: 94 (new combination); Kim, 1970; Tsuneki (1977c: 370) from Japan (Ishikawa Prefecture: Mount Pl. 58 Figure 703, 596 (author incorrectly listed as Fabricius, haku), which is junior secondary homonym of Mimesa alticola diagnosis; Korea: Mt. Cheonmasan), 812 (in the list of Hyme- Viereck, 1903, and synonymized with Psen seminitidus Lith by noptera from Korea); Tsuneki, 1976a: 298 (including Korea in Burdrys, 1986: 145; Tsuneki, 1982:3e4 (N Korea: “Prov. Ryang distribution); Tsuneki, 1982: 3 (N Korea: “Porv. Pyong gang: gang: Chan-Pay Plateau”), 15 (in the list of the species of the Hyesan: Mt. Ze-dong”), 15 (in the list of the species of the Sphecidae known from the Korean Peninsula); Paik, 1985:198 Sphecidae known from the Korean Peninsula); Paik, 1985:198 (in the list of Sphecidae of Korea); Kim and Yang, 2010: 145 (in (in the list of Sphecidae of Korea); Kim et al., 1994: 262 (in Check the checklist of valid Korean species of the Subfamily Pem- List of Insects from Korea); Budrys in Nemkov et al., 1995: 392 phredoninae); Nemkov, 2012b: 436 (including Korea in (in key, including Korea in distribution); Nemkov, 2009:57 distribution). (including Korea in distribution); Kim and Yang, 2010:144e145 Distribution. N Korea (RG), NE China, Russian Far East, Japan. (in the checklist of valid Korean species of the Subfamily Pem- phredoninae); Lee et al., 2010: 209 (in Checklist of Korean In- Psen ussuriensis Lith, 1932 우수리꼬마구멍벌 sects); Nemkov, 2012b: 435 (including Korea in distribution). Mimesa orientalis Gussakovskij, 1932:5e6, _ (syntype), Russia: Distribution. Transpalearctic: Korean Peninsula (PN, GG), Europe Primorskiy Krai: Sutshan and Tigrovaya [OMNH], junior sec- to Mongolia, NE China, Russian Far East, Japan. ondary homonym of Psen orientalis Cameron, 1890. Psen ussuriensis Lith, 1959: 59, substitute name for Psen orientalis Psen aurifrons Tsuneki, 1959 누런얼굴꼬마구멍벌 (Gussakovskij); Kim and Yang, 2010:141e142 (new to Korea; Psen aurifrons Tsuneki, 1959a:63e65, \ (holotype) and _, Japan: Korea: Gyeonggi-do, Gangwon-do, Chungcheongbuk-do, Jella- Fukui Prefecture, Sabaé Prefecture [MNHAH]; Tsuneki, 1974:366 nam-do). Distribution. S Korea (GG, GW, CB, JN), Russian Far East, Japan. 446 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Genus Pseneo Malloch, 1933 러시아꼬마구멍벌속 distribution); Kim and Yang, 2010: 144 (in the checklist of valid Pseneo Malloch, 1933: 7. Type species: Psen kohlii Fox, 1898,by Korean species of the Subfamily Pemphredoninae); Lee et al., original designation. 2010: 209 (in Checklist of Korean Insects); Nemkov, 2012b: 436 (including Korea in distribution). Pseneo exaratus (Eversmann, 1849) 러시아꼬마구멍벌 Distribution. S Korea (SL), Japan. Mimesa exarata Eversmann, 1849: 361, sex not indicated (holotype or syntypes), Russia: Kazan area, Russia [ZIN]. Psenulus laevigatus (Schenck, 1857) 검정다리꼬마구멍벌 Psen exaratus (Eversmann): Beaumont, 1937: 44 (new combina- Psen laevigatus Schenck, 1857:215,\ (holotype), Germany: Weil- tion); Yasumatsu, 1942b: 94; Tsuneki, 1959a: 49 and 52 (in key), burg; Hessen (depository uncertain). 69e71 (Korea: “Shoyozan”); Kim, 1970: 596 (diagnosis, no Psenulus laevigatus (Schenck): Schulz, 1911: 136 (new combination). Korean material), 812 (in the list of Hymenoptera from Korea); Psenulus fuscipes Tsuneki, 1959g: 33, \ (holotype) and _, Japan: Tsuneki, 1974: 366 (N Korea: “Prov. South Pyongan: Mts. Fukui Prefecture: Sakatani [MNHAH], synonymized with Pse- Guksan-bong, Za-mo san”); Tsuneki, 1982: 15 (in the list of the nulus laevigatus by Budrys in Nemkov et al., 1995: 393; Tsuneki, species of the Sphecidae known from the Korean Peninsula); 1974: 367 (new to Korea, N Korea: “Prov. Ryang-gang: Hyesan”); Tsuneki, 1991: 200 (in Sphecidae from Korea); Kim et al., 1994: Tsuneki, 1982:15 (in the list of the species of the Sphecidae 262 (in Check List of Insects from Korea); Yamane in Yamane known from the Korean Peninsula); Paik, 1985: 198 (in the list of et al., 1999: 489 (including Korea in distribution); Lee et al., Sphecidae of Korea); Tsuneki, 1991: 200 (in Sphecidae from 2010: 209 (in Checklist of Korean Insects). Korea); Kim et al., 1994: 262 (in Check List of Insects from Ko- Pseneo exaratus (Eversmann): Budrys, 1988:108(newcombi- rea); Nemkov, 2009: 60 (including Korea in distribution); Lee nation); Budrys in Nemkov et al., 1995: 391 (in key, including et al., 2010: 209 (in Checklist of Korean Insects); Kim and Korea in distribution); Nemkov, 2009: 57 (including Korea in Yang, 2010: 144 (in the checklist of valid Korean species of the distribution); Kim and Yang, 2010:145e146 (in the checklist Subfamily Pemphredoninae); Nemkov, 2012b: 436 (including of valid Korean species of the Subfamily Pemphredoninae); Korea in distribution). Nemkov, 2012b: 436 (including Korea in distribution). Distribution. Transpalearctic: N Korea (RG), Europe to Russian Distribution. Transpalearctic: Korean Peninsula (PN, GG), Europe Far East, Japan. to Russian Far East, Japan, south to India, Taiwan. Psenulus nipponensis Yasumatsu, 1942 일본꼬마구멍벌 Genus Psenulus Kohl, 1897 꼬리먹꼬마구멍벌속 Psenulus nipponensis Yasumatsu, 1942b: 96, \ (holotype), Japan: Psenulus Kohl, 1897: 293. Type species: Mimesa fuscipennis Dahl- Honshu: Minoo in Settu Province, and Paratype also from North bom (¼ Psen fuscipennis Dahlbom, 1843, designated by Korea: no specific locality [KUF]; Tsuneki, 1959a: 28 (in key, Ashmead, 1899: 224. including Korea in distribution), 35e36 (including Korea in distribution); Tsuneke, 1963b: 52 (in key, including Korea in Psenulus anomoneurae (Yasumatsu, 1938b) 북방꼬리먹꼬마구멍벌 distribution); Kim, 1970: 594 (diagnosis, no Korean material), Nipponopsen anomoneurae Yasumatsu, 1938b: 84, _ (holotype) and 812 (in the list of Hymenoptera from Korea); Tsuneki, 1982:15 \, Japan: Honshu: Yashirodani [KUF]. (in the list of the species of the Sphecidae known from the Psenulus anomoneurae (Yasumatsu): Tsuneki, 1963b: 52 and 53 (in Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae of key, new combination); Dollfuss, 2004: 111 (N Korea: “Hyan- Korea); Kim et al., 1994: 262 (in Check List of Insects from Ko- gean: Mychyang Mts.”); Nemkov, 2009: 59 (including Korea in rea); Lee et al., 2010: 209 (in Checklist of Korean Insects); Kim distribution); Kim and Yang, 2010: 144 (in the checklist of valid and Yang, 2010: 144 (in the checklist of valid Korean species of Korean species of the Subfamily Pemphredoninae). the Subfamily Pemphredoninae). Distribution. N Korea (“Hyangean: Mychyang Mts.”), Russian Far Distribution. N Korea (no specific locality), Japan. East, Japan. Psenulus pallipes gussakovskij Lith, 1973 극동꼬마구멍벌 Psenulus carinifrons iwatai (Gussakovskij, 1934) 호리꼬리먹꼬마 Psenulus puncticeps Gussakovskij, 1932:6,\ (syntypes), Russia: 구멍벌 Kamchatka or Ussuri area: Kursk Tomsk [OMNH], junior sec- Eopsenulus iwatai Gussakovskij, 1934: 84e86, \ (holotype), Japan: ondary homonym of Psenulus puncticeps Cameron, 1907. Osaka: Ikeda [OMNH]. Psenulus gussakovskij Lith, 1973: 116, substitute name for Psenulus Psenulus iwatai (Gussakovskij): Tsuneki, 1956b: 9 (new puncticeps Gussakovskij, 1932, and synonymized with Psenulus combination). pallipes by Haneda et al., 2005: 45; Kim et al., 1994: 262 (in Psenulus carinifrons iwatai (Gussakovskij): Bohart and Menke, 1976: Check List of Insects from Korea); Lee et al., 2010: 209. 172 (new status): Yang and Kim, 2010:361e363 (new to Korea, S Psenulus pallipes (Panzer): Nemkov, 2009: 61 (including Korea in Korea: Incheon Metropolitan City, Gyeonggi-do). distribution); Nemkov, 2012b: 436 (including Korea in distribution). Psenulus fuscipennis japonicus Tsuneki, 1959 꼬리꼬마구멍벌 Psenulus pallipes puncticeps Gussakovskij: Tsuneki, 1959a:27e28 Psenulus fuscipennis japonicus Tsuneki, 1959a: 38, \ and _ (syn- and 29 (in key, including Korea in distribution), 41e42 (new types), Korea: “Keijo”; Japan: Hokkaido: Sapporo: Jozankei; status; Korea: “Hakugan, Keijo”); Kim, 1970: 595 (diagnosis, no Aomori Prefecture: Towada; Fukui and Ishikava Prefectures Korean material), 812 (in the list of Hymenoptera from Korea). [MNHAH]; Tsuneki, 1963b: 51 and 52 (in key, including Korea in Psenulus pallipes gussakovskiji (Lith): Lith, 1979a: 151 (new status, distribution); Kim, 1970: Pl. 58 Figure 699, 594 (diagnosis, Ko- emendation); Tsuneki, 1974: 367 (N Korea: “Prov. South Pyon- rea: Uidong), 812 (in the list of Hymenoptera from Korea); gan: Pyongyan”); Tsuneki, 1982: 15 (in the list of the species of Bohart and Menke, 1976: 173 (including Korea in distribution); the Sphecidae known from the Korean Peninsula); Paik, 1985: Tsuneki, 1982: 15 (in the list of the species of the Sphecidae 198 (in the list of Sphecidae of Korea); Tsuneki, 1991: 200 (in known from the Korean Peninsula); Paik, 1985: 198 (in the list of Sphecidae from Korea); Kim and Yang, 2010: 144 (in the Sphecidae of Korea); Kim et al., 1994: 262 (in Check List of In- checklist of valid Korean species of the Subfamily sects from Korea); Nemkov, 2009: 59 (including Korea in Pemphredoninae). JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 447

Distribution. Transpalearctic: Korean Peninsula (PN, SL), Russian Mt. Gongo”); Kim, 1970: Pl. 61 Figure 746, 622 (diagnosis, Korea: Far East, Japan. Mt. Sokrisan), 819 (in the list of Hymenoptera from Korea); Kim, 1980: Pl. xx HYCE 01, 144 (diagnosis, Korea: Gangwon-do, Subfamily Philanthinae Latreille, 1802 노래기벌아과 Gyeonggi-do); Tsuneki, 1982: 18 (in the list of the species of Tribe Cercerini Lepeletier, 1845 노래기벌족 the Sphecidae known from the Korean Peninsula); Paik, 1985: Genus Cerceris Latreille, 1802 노래기벌속 203 (in the list of Sphecidae of Korea); Kim et al., 1994: 263 (in Cerceris Latreille, 1802: 367. Type species: Philanthus ornatus Fab- Check List of Insects from Korea); Kazenas in Nemkov et al., ricius, 1790 (¼ Sphex rybyensis Linnaeus, 1771), designated by 1995: 473 (in key, including Korea in distribution); Yamane in Latreille, 1810: 438. Yamane et al., 1999: 545 (including Korea in distribution); Terayama and Tano, 2005: 6 and 15 (including Korea in distri- Cerceris adelpha Kohl, 1887 아델파노래기벌 bution); Yamane, 2008: 554 (including Korea in distribution); Cerceris adelpha Kohl in Schletterer, 1887:447,\ (lectotype, desig- Lee et al., 2008: 64 (in the checklist of Korean species of Phi- nated by Kohl 1916: 107), Korea: no specific locality [NMK]; lanthidae); Nemkov, 2009: 145 (including Korea in distribu- Schletterer, 1887: 485 (including Korea in distribution); Tsuneki, tion); Lee et al., 2010: 211 (in Checklist of Korean Insects); 1961: 12 and 18 (in key, including Korea in distribution), 42; Nemkov, 2012b: 447 (including Korea in distribution). Kim, 1970: 622 (diagnosis, no Korean material), 817 (in the list of Distribution. Transpalearctic: Korean Peninsula (SL, GG, GW, CB), Hymenoptera from Korea); Bohart and Menke, 1976:576 North Africa through Europe to NE China, Russian Far East, Japan. (including Korea in distribution); Kazenas, 1979: 85 (including Korea in distribution); Tsuneki, 1982:12e13 (N Korea: “Prov. Cerceris bicincta Klug, 1835 네점박이노래기벌 Pyong sung: Bek-sung-li: Za-mo san”), 18 (in the list of the Cerceris bicincta Klug in Waltl, 1835:100,_ (holotype), Spain: species of the Sphecidae known from the Korean Peninsula); Andalusia [ZMHU]; Tsuneki, 1961: 10 and 16 (in key, including Paik, 1985: 203 (in the list of Sphecidae of Korea); Kim et al., Korea in distribution), 41 (Korea: “Shoyozan”); Kim, 1970: 623 1994: 263 (in Check List of Insects from Korea); Kazenas in (diagnosis, no Korean material), 817 (in the list of Hymenoptera Nemkov et al., 1995: 473 (in key, including Korea in distribu- from Korea); Tsuneki, 1982: 18 (in the list of the species of the tion); Lee et al., 2008: 64 (in the checklist of Korean species of Sphecidae known from the Korean Peninsula); Paik, 1985: 203 Philanthidae); Nemkov, 2009:145e146 (including Korea in (in the list of Sphecidae of Korea); Kim et al., 1994: 263 (in Check distribution); Lee et al., 2010: 211 (in Checklist of Korean In- List of Insects from Korea); Kazenas in Nemkov et al., 1995:472 sects); Nemkov, 2012b: 447 (including Korea in distribution). (in key, including Korea in distribution); Lee et al., 2008: 64 (in Cerceris hokkanzana Tsuneki, 1961: 17 (in key), 43, _ (nec \), Korea: the checklist of Korean species of Philanthidae); Nemkov, 2009: “Soyozan, Kainei” [MNHAH], synonymized _ with C. adelpha by 146 (including Korea in distribution); Lee et al., 2010: 211 (in Tsuneki, 1982:13;Tsuneki, 1976a: 293 (including Korea in dis- Checklist of Korean Insects); Nemkov, 2012b: 447 (including tribution); Paik, 1985: 203 (in the list of Sphecidae of Korea); Korea in distribution). Kim et al., 1994:264. Distribution. Transpalearctic: Korean Peninsula (GG), Europe Distribution. Korean Peninsula (PN, GG), Mongolia, NE China, through central Asia to Mongolia, N China, Russian Far East. Russian Far East. Cerceris carinalis Pérez, 1905 Cerceris albofasciata (Rossi, 1790) 흰띠노래기벌 Cerceris carinalis Pérez, 1905, \ (holotype or syntypes), Japan: Vespa albofasciata Rossi, 1790:87,\ (holotype), Italy: “Etruria” (now Yokohama [MNNH]: Terayama and Tano, 2005: 6, 15 (including Toscana) (lost). Korea in distribution). Cerceris albofasciata (Rossi): Rossi, 1792: 138 (new combination); Distribution. ?Korea, Japan. Tsuneki, 1961: 10, 17, 25, 26, and 41 (Korea: “Keijo, Shyozan”); Note. The record of Korean occurrence by Terayama and Tano Yasumatsu, 1965: 302 (including Korea in distribution); Kim, (2005) was groundless. 1970: Pl. 61 Figure 745, 622 (diagnosis, Korea: Naeseolak), 818 (in the list of Hymenoptera from Korea); Kim, 1980: 145 (Korean Cerceris coreensis Tsuneki, 1961 참노래기벌 localities in Note of Cerceris hortivaga Kohl); Tsuneki, 1982:18 Cerceris coreensis Tsuneki, 1961: 11 and 17e18 (in key), 45, \ (ho- (in the list of the species of the Sphecidae known from the lotype) and _, Korea: “Shoyozan” [USNM]; Kim, 1970: Pl. 62 Korean Peninsula); Paik, 1985: 203 (in the list of Sphecidae of Figure 748, 623 (diagnosis, S Korea: Naeseolak), 817 (in the list of Korea); Kim et al., 1994: 263 (in Check List of Insects from Ko- Hymenoptera from Korea); Bohart and Menke, 1976:579 rea); Kazenas in Nemkov et al., 1995: 472 (in key, including (including Korea in distribution); Tsuneki, 1982: 19 (in the list of Korea in distribution); Yamane in Yamane et al., 1999: 546e547 the species of the Sphecidae known from the Korean Peninsula); (including Korea in distribution); Terayama and Tano, 2005:6, Paik, 1985: 203 (in the list of Sphecidae of Korea); Kim et al., 15 (including Korea in distribution); Yamane, 2008: 554 1994: 263 (in Check List of Insects from Korea); Kazenas in (including Korea in distribution); Lee et al., 2008: 64 (in the Nemkov et al., 1995: 472 (in key, including Korea in distribu- checklist of Korean species of Philanthidae); Nemkov, 2009:145 tion); Lee et al., 2008: 64 (in the checklist of Korean species of (including Korea in distribution); Lee et al., 2010: 211 (in Philanthidae); Nemkov, 2009: 147 (including Korea in distri- Checklist of Korean Insects); Nemkov, 2012b: 447 (including bution); Lee et al., 2010: 211 (in Checklist of Korean Insects); Korea in distribution). Nemkov, 2012b: 447 (including Korea in distribution). Distribution. Transpalearctic: S Korea (SL, GG, GW), Europe to E Distribution. S Korea (GG, GW), Russian Far East. Mongolia, NE China, Russian Far East, Japan. Cerceris hortivaga Kohl, 1880 노래기벌 Cerceris arenaria (Linnaeus, 1758) 띠노래기벌 Cerceris hortivaga Kohl, 1880: 223, \ and _ (syntypes), Austria: Sphex arenaria Linnaeus, 1758: 571, \ (holotype), Sweden: Fahuln Südtirol: Levico (now Italy: Alto Adige: Levico) [NHMW]; [LS]. Radoszkowski, 1890: 231 (Korea: no specific locality); Tsuneki, Cerceris arenaria (Linnaeus): Billberg, 1829: 107 (new combination); 1961: 9, 15, and 24e25 (in key, including Korea in distribution), Tsuneki, 1961: 19 (in key), 41 (Korea: “Shyozan, Kodaisan, Keijo, 36 (Korea: “Shoyzan, Kodaisan, Temmasan, Shu-otsu, Konggosan, 448 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Gyokyusenji, Kayasan, Utsuryo Is.”); Yasumatsu, 1965: 302 Korean species of Philanthidae); Lee et al., 2010: 211 (in (including Korea in distribution); Kim, 1970: Pl. 62 Figure 750, Checklist of Korean Insects). 624 (diagnosis, S Korea: Ulleung Is.), 819 (in the list of Hyme- Distribution. S Korea (JJ), Japan. noptera from Korea); Tsuneki, 1974: 371 (N Korea: “Prov. South Pyngan: Pyongyan”); Tsuneki, 1976a: 294 (including Korea in Cerceris pedetes Kohl, 1887 홍다리노래기벌 distribution); Kim, 1980: Pl. xx HYCE 02, 145 (diagnosis, Korea: Cerceris bicornuta F. Smith, 1856: 455, \ (holotype or syntypes), Gangwon-do, Gyeonggi-do, Chungcheongnam-do, Gyeong- North China: no specific locality [BMNH], junior primary hom- sangbuk-do, Gyeongsangnam-do, Jeollabuk-do, Jeollanam-do, onym of Cerceris bicornuta Guérin-Méneville 1844, and synon- Jeju Is.); Tsuneki, 1982: 18 (in the list of the species of the Sphe- ymized with Cerceris pedetes by R. Turner, 1912:814. cidae known from the Korean Peninsula); Paik, 1985: 203 (in the Cerceris smithii Dalla Torre, 1897: 199, substitute name for Cerceris list of Sphecidae of Korea); Tsuneki,1991: 201 (in Sphecidae from bicornuta F. Smith, 1856; Paik, 1985: 203 (in the list of Sphecidae Korea); Kim et al.,1994: 264 (in Check List of Insects from Korea); of Korea). Kazenas in Nemkov et al., 1995: 471 (in key, including Korea in Cerceris pedetes Kohl in Schletterer, 1887: 449, 498, \ (holotype or distribution); Yamane and Tano, 1995: 14 (including Korea in syntypes), Korea: no specific locality [NMK]; Tsuneki, 1961:13 distribution); Yamane in Yamane et al., 1999: 545 (including and 25 (in key), 42 (including Korea in distribution); Kim, 1970: Korea in distribution); Terayama and Tano, 2005: 7 and 15 Pl. 62 Figure 754, 625 (diagnosis, Korea: no specific locality), 817 (including Korea in distribution); Yamane, 2008: 554 (including (in the list of Hymenoptera from Korea); Tsuneki, 1982: 18 (in Korea in distribution); Lee et al., 2008: 64 (in the checklist of the list of the species of the Sphecidae known from the Korean Korean species of Philanthidae); Nemkov, 2009: 148 (including Peninsula); Paik, 1985: 203 (in the list of Sphecidae of Korea); Korea in distribution); Lee et al., 2010: 211 (in Checklist of Kim et al., 1994: 264 (in Check List of Insects from Korea); Korean Insects); Nemkov, 2012b: 448 (including Korea in Kazenas in Nemkov et al., 1995: 474 (in key, including Korea in distribution). distribution); Terayama and Tano, 2005: 8, 15 (including Korea Distribution. Transpalearctic: Korean Peninsula (PN, SL, GG, GW, in distribution); Lee et al., 2008: 65 (in the checklist of Korean CN, GB, GN, JB, JN, JJ), N Africa, Europe to Mongolia, N China, Russian species of Philanthidae); Nemkov, 2009:149;Lee et al., 2010: Far East, Japan. 211 (in Checklist of Korean Insects); Neov, 2012b: 448 (including Korea in distribution). Cerceris koma Tsuneki, 1961 고려노래기벌 Distribution. Korean Peninsula (PN, GG), NE China, Russian Far Cerceris koma Tsuneki, 1961: 8 (in key), 40, \ (holotype), Korea: East, Japan. “Keijo: Hokkanzan” [USNM]; Kim, 1970: 624 (diagnosis, no Korean material), 818 (in the list of Hymenoptera from Korea); Cerceris quadrifasciata (Panzer, 1799) 네줄노래기벌 Bohart and Menke, 1976: 583 (including Korea in distribution); Philanthus quadrifasciatus Panzer, 1799, Heft 63: 14, \ (holotype or Tsuneki, 1982: 18 (in the list of the species of the Sphecidae syntypes), Austria: Vienna (depository uncertain). known from the Korean Peninsula); Paik, 1985: 203 (in the list of Cerceris quadrifasciata (Panzer): Latreille, 1805: 316 (new combi- Sphecidae of Korea); Kim et al., 1994: 264 (in Check List of In- nation); Tsuneki, 1961: 12 and 18 (in key), 44 (Korea: “Shoyo- sects from Korea); Kazenas in Nemkov et al., 1995: 470 (in key, zan”); Kim, 1970: Pl. 62 Figure 755, 626 (diagnosis, no Korean including Korea in distribution); Lee et al., 2008: 65 (in the material), 818 (in the list of Hymenoptera from Korea); Tsuneki, checklist of Korean species of Philanthidae); Lee et al., 2010:211 1982: 18 (in the list of the species of the Sphecidae known from (in Checklist of Korean Insects). the Korean Peninsula); Paik, 1985: 203 (in the list of Sphecidae Distribution. S Korea (SL). of Korea); Kim et al., 1994: 264 (in Check List of Insects from Korea); Kazenas in Nemkov et al., 1995: 473 (in key, including Cerceris koryo Tsuneki, 1961 광릉노래기벌 Korea in distribution); Lee et al., 2008: 65 (in the checklist of Cerceris koryo Tsuneki, 1961: 18 (in key), 43, _ (holotype), Korea: Korean species of Philanthidae); Nemkov, 2009: 150 (including “Koryo” [USNM]; Kim, 1970: 625 (diagnosis, no Korean mate- Korea in distribution); Lee et al., 2010: 211 (in Checklist of rial), 818 (in the list of Hymenoptera from Korea); Bohart and Korean Insects); Nemkov, 2012b: 448 (including Korea in Menke, 1976: 583 (including Korea in distribution); Tsuneki, distribution). 1982: 18 (in the list of the species of the Sphecidae known Distribution. Transpalearctic: S Korea (GG), Europe to NE China, from the Korean Peninsula); Paik, 1985: 203 (in the list of Russian Far East. Sphecidae of Korea); Kim et al., 1994: 264 (in Check List of In- sects from Korea); Kazenas in Nemkov et al., 1995: 479 (in key, Cerceris quinquefasciata seoulensis Tsuneki, 1961 서울네줄노래 including Korea in distribution); Lee et al., 2008: 65 (in the 기벌 checklist of Korean species of Philanthidae); Lee et al., 2010:211 Cerceris quinquefasciata seoulensis Tsuneki, 1961:12and18(in (in Checklist of Korean Insects). keys), 44, \ (holotype), _, Korea: Seoul [USNM]; Kim, 1970: 626 Distribution. S Korea (GG). (as C. quadrifasciata seoulensis, diagnosis, no Korean material), 818 (in the list of Hymenoptera from Korea); Bohart and Cerceris nipponensis Tsuneki, 1961 니폰노래기벌 Menke, 1976: 586 (including Korea in distribution); Tsuneki, Cerceris nipponensis Tsuneki, 1961:31,\ (holotype) and _, Japan: 1982: 18 (in the list of the species of the Sphecidae known Ishikawa: Mont Haku [MNHAH]; Tsuneki, 1968b: 51 (Korea: Jeju from the Korean Peninsula); Paik, 1985: 203 (in the list of Is.: Mt. Hanna); Tsuneki, 1968c: 59 (Korea: Jeju Is.: Mt Hanna); Sphecidae of Korea); Kim et al., 1994: 264 (in Check List of Kim, 1970: Pl. 62 Figure 753, 625 (diagnosis, S Korea: Mt. Insects from Korea); Kazenas in Nemkov et al., 1995: 473 (in Gayasan), 817 (in the list of Hymenoptera from Korea); Paik, key, including Korea in distribution); Lee et al., 2008:65(inthe 1985: 203 (in the list of Sphecidae of Korea); Kim et al., 1994: checklist of Korean species of Philanthidae); Nemkov, 2009: 264 (in Check List of Insects from Korea); Yamane and Tano, 151 (including Korea in distribution); Lee et al., 2010:211(in 1995: 17; Yamane in Yamane et al., 1999: 547 (including Korea Checklist of Korean Insects). in distribution); Terayama and Tano, 2005: 8, 15 (including Distribution. S Korea (SL), NE China, Russian Far East. Korea in distribution); Lee et al., 2008: 65 (in the checklist of JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 449

Cerceris rubida (Jurine, 1807) 두줄노래기벌 Distribution. S Korea (SL, GG, GB), Central and NE China, Russian Philanthus rubidus Jurine, 1807: 202, \ (holotype), Europe: no spe- Far East. cific locality, perhaps France [MHNG]. Cerceris rubidus (Jurine): A. Costa, 1867c: 154 (new combination); Cerceris saishuensis Tsuneki, 1968 제주노래기벌 Lee et al., 2008:59e61 (new to Korea, S Korea: Gyeonggi-do, Cerceris saishuensis Tsuneki, 1968b: 53, _ (holotype), Korea: Quel- Gangwon-do, Chungcheongbuk-do, Chungcheongnam-do, quepart Island (now Jeju Island): Mt. Hanna [MNHAH]; Kim, Gyeongsangbuk-do, Jeollabuk-do, Jeollanam-do); Lee et al., 1970: Pl. 63 Figure 759, 627 (diagnosis, S Korea: Jejudo: Chuja 2010: 211 (in Checklist of Korean Insects). Is.), 818 (in the list of Hymenoptera from Korea); Bohart and Distribution. Transpalearctic: S Korea (GG, GW, CB, CN, GB, JB, Menke, 1976: 587 (including Korea in distribution); Paik, 1985: JN), South Europe to W Asia, Asia Minor, NE China, Russian Far East, 203 (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Japan. Check List of Insects from Korea); Lee et al., 2008: 66 (in the checklist of Korean species of Philanthidae); Lee et al., 2010:211 Cerceris ruficornis (Fabricius, 1793) 백두산노래기벌 (in Checklist of Korean Insects). Philanthus ruficornis Fabricius, 1793: 292, _ (lectotype, designated Distribution. S Korea (JJ). by van der Vecht, 1961: 65), Italy: no specific locality [ZMCU]. Cerceris ruficornis (Fabricius): van der Vecht, 1961: 65 (new com- Cerceris semilunata Radoszkowski, 1870 북한산노래기벌 bination); Tsuneki, 1961: 13 and 17 (in keys), 44 (N Korea: “Mt. Cerceris semilunata Radoszkowski, 1870: 105, \ (holotype), Russia: Hakuto: Jimmjo”); Kim, 1970: 626, 817 (in the list of Hyme- Amur area (type depository uncertain); Kazenas in Nemkov noptera from Korea); Tsuneki, 1976a: 293 (include N Korea in et al., 1995: 474 (in key, including Korea in distribution), 478 distribution); Tsuneki, 1982: 18 (in the list of the species of the (in key); Lee et al., 2010: 211 (in Checklist of Korean Insects). Sphecidae known from the Korean Peninsula); Paik, 1985: 203 Cerceris hokkanzana Tsuneki, 1961: 13 and 17 (in key), 42, \ (holo- (in the list of Sphecidae of Korea); Kim et al., 1994: 264 (in Check type, nec _, see synonym list under C. adelpha), Korea: “Keijo: List of Insects from Korea); Kazenas in Nemkov et al., 1995:474 Hokkanzan” [USNM], synonymized with C. semilunata by (in key, including Korea in distribution); Lee et al., 2008: 65 (in Kazenas in Nemkov et al., 1995:474;Kim, 1970: 624 (diagnosis, the checklist of Korean species of Philanthidae); Nemkov, 2009: no Korean material), 818 (in the list of Hymenoptera from Ko- 151 (including Korea in distribution); Lee et al., 2010: 211 (in rea); Tsuneki, 1974: 372 (N Korea: Kaesong: Mts. Pakyon); Checklist of Korean Insects); Nemkov, 2012b: 448 (including Bohart and Menke, 1976: 582 (including Korea in distribution); Korea in distribution). Tsuneki, 1982: 18 (in the list of the species of the Sphecidae Distribution. Transpalearctic: N Korea (RG), Europe to N China, known from the Korean Peninsula); Tsuneki, 1991: 201 (listed); Russian Far East. Kim et al., 1994: 264 (in Check List of Insects from Korea); Lee et al., 2008: 66 (in the checklist of Korean species of Philan- Cerceris rybyensis (Linnaeus, 1771) 둥근무늬노래기벌 thidae); Nemkov, 2009: 152 (including Korea in distribution); Sphex rybyensis Linnaeus, 1771: 88, \ (lectotype, designated by Day, Nemkov, 2012b: 448 (including Korea in distribution). 1979: 71), Sweden: Söermanland [LSL]. Distribution. Korean Peninsula (SL, GG), Mongolia, N China, Cerceris rybyensis (Linnaeus): Kohl, 1880: 144 (new combination); Russian Far East. Tsuneki, 1974: 371 (N Korea: “Prov. South Pyongan: Mang-yong- dae, Chang-ryong san, Bongwa-ri”); Tsuneki, 1982: 18 (in the list Cerceris shirozui Tsuneki, 1968 시즈로노래기벌 of the species of the Sphecidae known from the Korean Penin- Cerceris shirozui Tsuneki, 1968b:51,\ (holotype), Korea: Quel- sula); Paik, 1985: 203 (in the list of Sphecidae of Korea); Tsuneki, quepart Island: Mt. Hanna [MNHAH]; Kim, 1970: 627, 818 (in the 1991: 201 (in Sphecidae from Korea); Kim et al., 1994: 264 (in list of Hymenoptera from Korea); Bohart and Menke, 1976: 587 Check List of Insects from Korea); Kazenas in Nemkov et al., (including Korea in distribution); Kim et al., 1994: 264 (in Check 1995: 471 and 476e477 (in key, including Korea in distribu- List of Insects from Korea); Lee et al., 2008: 66 (in the checklist of tion); Lee et al., 2008: 65 (in the checklist of Korean species of Korean species of Philanthidae); Lee et al., 2010: 211 (in Philanthidae); Nemkov, 2009: 152 (including Korea in distri- Checklist of Korean Insects). bution); Lee et al., 2010: 211 (in Checklist of Korean Insects); Distribution. S Korea (JJ). Nemkov, 2012b: 448 (including Korea in distribution). Distribution. Transpalearctic: N Korea (PN), N Africa through Cerceris sobo Yasumatsu and Okabe, 1936 소보노래기벌 Europe Russian Far East, Japan. Cerceris sobo Yasumatsu and Okabe, 1936:497e499, \ (holotype), Japan: Kyushu: Kôaru [KUF]; Tsuneki, 1961: 7, 15, and 23e24 (in Cerceris sabulosa subgibbosa Yasumatsu, 1935 사불로사노래기벌 key), 39 (description of _ from Korea: “Shoyozan, Gyokusenji”); Cerceris subgibbosa Yasumatsu, 1935a: 15, 25, \ (holotype), China: Kim, 1970: Pl. 63 Figure 760, 628 (diagnosis, S Korea: Mt. Hopeh: Cheng-teh (depository uncertain). Gayasan), 818 (in the list of Hymenoptera from Korea); Tsuneki, Cerceris sabulosa subgibbosa Yasumatsu: Tsuneki, 1961: 10 and 16 1974: 372 (N Korea: “South Pyongan: De-sang san, Chang-ryong (in key), 36 (new status); Kim, 1970: Pl. 62 Figure 757, 627 san, Za-mo san”); Tsuneki, 1982: 18 (in the list of the species of (diagnosis, Korea: Mt. Taebaeksan), 818 (in the list of Hyme- the Sphecidae known from the Korean Peninsula); Paik, 1985: noptera from Korea); Kazenas in Nemkov et al., 1979: 83 and 87 203 (in the list of Sphecidae of Korea); Tsuneki, 1991:201;Kim (including Korea in distribution); Tsuneki, 1982: 13, 18 (in the et al., 1994: 264 (in Check List of Insects from Korea); Kazenas in list of the species of the Sphecidae known from the Korean Nemkov et al., 1995: 468 and 476 (in key, including Korea in Peninsula); Paik, 1985: 203 (in the list of Sphecidae of Korea); distribution); Terayama and Tano, 2005: 8 and 15 (including Kim et al., 1994: 264 (in Check List of Insects from Korea); Korea in distribution); Lee et al., 2008: 66 (in the checklist of Kazenas in Nemkov et al., 1995: 471 and 477 (in key, including Korean species of Philanthidae); Lee et al., 2010: 212 (in Korea in distribution); Lee et al., 2008: 66 (in the checklist of Checklist of Korean Insects). Korean species of Philanthidae); Lee et al., 2010: 211 (in Distribution. Korean Peninsula (PN, GG, GN), Japan. Checklist of Korean Insects). 450 JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456

Cerceris supraconica Tsuneki, 1961 코주부노래기벌 Fabricius, 1790 (¼ Sphex coronata Thunberg, 1784), designated Cerceris supraconica Tsuneki, 1961: 8 and 15 (in key), 38, \ (holo- by Shuckard, 1837: 246; Lee et al., 2008: 66 (listed). type) and _, Korea: Mt. Kaya [USNM]; Kim, 1970: Pl. 63 Figure 761, 629 (diagnosis, S Korea: Seoul), 818 (in the list of Philanthus coronatus (Thunberg, 1784) 금관노래기벌 Hymenoptera from Korea); Bohart and Menke, 1976: 588 Sphex coronata Thunberg, 1784: 25 (as coronata, incorrect original (including Korea in distribution); Tsuneki, 1982: 18 (in the list of termination) _ (holotype or syntypes), France: Parisiis, now the species of the Sphecidae known from the Korean Peninsula); Paris (depository uncertain). Paik, 1985: 203 (in the list of Sphecidae of Korea); Kim et al., Philanthus coronatus (Thunberg): Thunberg, 1815a: 128 (new 1994: 264 (in Check List of Insects from Korea); Kazenas in combination); Tsuneki, 1943b:33e36 (nesting habitat observed Nemkov et al., 1995: 470 and 476 (in key, including Korea in in Seoul, Korea); Tsuneki, 1982: 18 (in the list of the species of distribution); Lee et al., 2008: 66 (in the checklist of Korean the Sphecidae known from the Korean Peninsula); Paik, 1985: species of Philanthidae); Lee et al., 2010: 212 (in Checklist of 203 (in the list of Sphecidae of Korea); Lee et al., 2010: 211 (in Korean Insects). Checklist of Korean Insects). Distribution. S Korea (SL, GN). Distribution. Transpalearctic: Korea (SL) and Europe through Russia to NE China (Wu and Zhou, 1996: 160). Cerceris tuberculata evecta Shestakov, 1918 왕노래기벌 Cerceris evecta Shestakov, 1918b: 3, \ (Syntypes) and _, China: Discussion Tientsin, also spelled Tianjin [ZIN]; Lee et al., 2010: 211 (in Checklist of Korean Insects). To comprehend Korean Spheciformes biodiversity, the following Cerceris tuberculata evecta Shestakov: Bohart and Menke, 1976: 588 species separately grouped into four topics should be more criti- (new subspecific combination); Paik, 1985: 203 (in the list of cally studied in the future. Sphecidae of Korea). 1) The following three paired conspecific subspecies are sym- Cerceris tuberculata var. bicornutum Smith: Tsuneki, 1961: 53, cor- patric, thus their subspecific status is self-contradictory. They were rected to Cerceris rufipes evecta by Tsuneki; Kim, 1970: 629 established by means of a single specimen. Future taxonomic (listed with brief comment on previous collecting record from clarification is required with more sufficient material. Pyeongyang of North Korea); Kim et al., 1994: 264 (in Check List of Insects from Korea). Ectemnius schlettereris chlettereri (Kohl 1988) and Ectemnius Cerceris fufipes evecta Shestakov: Yasumatsu, 1942b: 107 (new sta- schlettereri horvatovichi (Tsuneki 1974). tus); Kim, 1970: 629 (listed with brief comment on previous Rhopalum coartatum chosenense (Tsuneki 1974) and Rhopalum collecting record from Pyeongyang of North Korea in text); Kim coarctatum koreense (Tsuneki 1974). et al., 1994: 264 (in Check List of Insects from Korea). Trypoxylon fronticorne japonense (Tsuneki 1956) and Trypoxylon Distribution. ? N Korea (PN), Mongolia, E China. fronticorne seoulense (Tsuneki 1981). Note. Unfounded listing by Kim (1970: 629) was the first record of Korean occurrence of this species. This species is known to be 2) It is highly recommended that the following species should distributed in Mongolia and E China. Thus, I omit this species from be excluded from the Korean fauna (refer to Note in each species). the Korean fauna. Their exclusions from the Korean fauna could be documented through the currently known distributional range or confirmation Cerceris unifasciata Smith, 1856 외줄노래기벌 of the misconceptional citation of the previous record. Cerceris unifasciata Smith, 1856: 456, \ (holotype or syntypes), North China: no specific locality [BMNH]; Uchida, 1925a:331(N Ampulex kurarensis (Yasumatsu 1936) Korea: Hwasan); Uchida,1925b: 369 (in table, including Korea in Gorytes tricinctus (Pérez 1905) distribution); Lee et al., 2008: 66 (in the checklist of Korean Crossocerus hakusanus (Tsuneki 1954) species of Philanthidae); Lee et al., 2010: 212 (in Checklist of Rhopalum venustum (Tsuneki 1955) Korean Insects). Tachytes monticola (Dalla Torre 1897) [sic] [¼ Tachyspex brullii Distribution. N Korea (Hwasan), N China. (Smith 1856)] Remarks. Based on the misconception of Matsumura (1911: 115), Trypoxylon ussuriense (Kazenas 1980) Far Eastern Cerceris hortivaga was very often misidentified as this Stigmus convergens (Tsuneki 1954) species in the early 1920s (refer to Tsuneki, 1961). Thus, future Cerceris carinalis (Pérez 1905) confirmation of the Korean occurrence of this species is needed. Cerceris tuberculata evecta (Shestakov 1918)

Cerceris verhoeffi Tsuneki, 1961 민노래기벌 3) As far as I am aware, Korean occurrences of the following Cerceris verhoeffi Tsuneki, 1961: 55, \ (holotype) and _, China: species were commented on without examination and designation Heilungkiang: Harbin, [RMNH]; Lee and Kim, 2008: 61e63 (new of Korean material in any taxonomic publications. Unfortunately, to Korea: Gyeonggi-do, Gangwon-do, Chungcheongbuk-do, there is no way currently to confirm whether these records were Chungcheongnam-do, Gyeongsangbuk-do, Gyengsangnam-do, based on circumstantial evidence or examination of Korean mate- Jeollabuk-do); Lee et al., 2010: 211 (in Checklist of Korean rial although they were not designated therein. Insects). Distribution. S Korea (GG, GW, CB, CN, GB, GN, JB), NE China, Ampulex dissector (Thunberg 1822) Russian Far East. Sceliphron madraspatanum kohli (Sickmann 1781) Ectemnius nitobei (Matsumura 1912) Tribe Philanthini Latreille, 1802 황노래기벌족 Ectemnius shimoyamai (Tsuneki 1958) Subtribe Philanthina Latreille, 1802 황노래기벌아족(신칭) Rhopalum venustum (Tsuneki 1955) Genus Philanthus Fabricius, 1790 황노래기벌속 Ectemnius nitobei (Matsumura 1912) Philanthus Fabricius, 1790: 224. Type species: Philanthus coronatus Ectemnius shimoyamai (Tsuneki 1958) JK Kim / Journal of Asia-Pacific Biodiversity 7 (2014) 415e456 451

Liris festinans japonica (Kohl 1884) Diodontus chosenensis Tsuneki, 1974 Liris subtesellatus (Smith 1856) Diodontus minutus orientalis (Tsuneki 1974) Oxybelus bipunctatus (Olivier 1812) Passaloecus clypealis (Faester 1947) Trypoxylon regium (Gussakovskij 1932) Passaloecus insignis (Linden 1829) Trypoxylon rufimaculatum (Antropov 1987) Passaloecus koreanus (Tsuneki 1982) Mellinus sabulosus (Fabricius 1787) Passaloecus monilicornis (Dahlbom 1842) Pemphredon diervillae (Iwata 1933) Pemphredon flavistigma (Thomson 1874) Pemphredon morio (Linden 1829) Pemphredon lugubris (Fabricius 1793) Polemistus abnormis (Kohl 1888) Mimumesa atratina (Morawitz 1891) Mimesa equestris (Fabricius 1804) Mimumesa dahlbomi (Wesmael 1852) Mimesa lutaris (Fabricius, 1787) Mimumesa littoralis (Bondroit 1934) Psen aurifrons (Tsuneki 1959) 4) When stating the distribution related to the Korean Penin- Psen seminitidus (Lith 1965) sula, many authors used inclusively “Korea” (even “Choseon” in Psenulus anomoneurae (Yasumatsu 1938) some earlier Japanese articles) without distinction between North Psenulus laevigatus (Schenck 1857) Korea and South Korea. It is obviously true that the climate of the Psenulus nipponensis (Yasumatsu 1942) northernmost mountain regions of the Korean Peninsula (espe- Cerceris ruficornis (Fabricius 1793) cially RG and HB) is different from that of other regions of the Cerceris rybyensis (Linnaeus 1771) Korean Peninsula. The fauna of the former regions are likely to be Cerceris unifasciata (Smith 1856) more closely related to either NE China or the Russian Far East than one of the other Korean regions. Not a few Korean occurrence re- References cords were made with the specimens from those regions. For sound biodiversity evaluation of Korean fauna, the distributional range in Andre Ed. 1886e1891. Species des Hyménoptères d’Europe & d’Algérie. Vol. 3. Les the Korean Peninsula of the following species that were known Sphégiens. Chez l’auteur, Beaune (Côte d’Or). 340 þ 30 pp., XV pls. (Fasc. 24e26: e e e only from North Korea must be ascertained. 1 104, 1 June 1886; Fasc. 27: 105 168, pls. I IV, VI, VII, 1 Apr 1888; Fasc. 30: 169e248, 1888; Fasc. 33: 249e320, 1889; Fasc. 38: 25e30, explanations for plates XIeXV, 1891). Alysson pertheesi (Gorski 1852) Antropov AV. 1986. K poznaniyu royuschchikh os roda Trypoxylon (Hymenoptera, e Alyson ratzeburgi (Dahlbom 1843) Sphecidae) palearkticheskoy fauny. Zoologicheskiy Zhurnal 65:624 628. Antropov AV. 1987. K faune royuschchikh os roda Trypoxylon Latreille (Hymenoptera, Argogorytes mystaceus grandis (Gussakovskij 1932) Sphecidae) vostochnoy Palearktiki. In: Lehr PA, Storozheva NA, editors. Novye Gorytes maculicornis (F. Morawitz, 1889) dannye po sistematike nasekomykh Dal’nego Vostoka. Vladivostok: Dal’nevos- Gorytes quadrifasciatus (Fabricius 1804) tochnoe Otdelenie Akademii Nauk SSSR, Biologo-Pochvennyi Institut. pp. 57e61. Antropov AV. 1994. A review of the agile species group of Pison (Hymenoptera: Harpactus laevis (Latreille 1792) Sphecidae: Trypoxylini). Journal of Hymenoptera Research 3:119e132. Nysson maculosus (Gmelin, 1790) Antropov AV. 1995. Four new species of the digger wasps genus Trypoxylon Latreille Crabro cribrarius (Linnaeus 1758) (Hymenoptera, Sphecidae) of the Palaearctic and Oriental Regions, with taxo- nomic notes on some others previously described. Russian Entomological Journal Crossocerus amurensis (Kohl, 1892) 3:123e133. Crossocerus annulipes hokkaiddoensis (Tsuneki 1954) Audouin I. 1822. Ammophile and Ampulex. In: Vincent Bory J de Saint, editor. Crossocerus assimilis collaris (Tsuneki 1974) Dictionnaire classique d’histoire naturelle. Tome premier. Paris: Rey et Gravier and e Crossocerus ceratus (Schukard 1837) Boudouin Frères. pp. 280 281. 301. Ashmead WH. 1899. Classification of the entomophilous wasps, or the superfamily Crossocerus cinxius (Dahlbom 1839) Sphegoidea. The Canadian Entomologist 31. 145e155, 161e174, 212e225, 238e Crossocerus denticornis (Bischoff 1932) 251, 291e300, 322e330, 345e357. Crossocerus heydeni (Kohl 1880) Baik YB, Kim JK. 2014. 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