AMEBICAN ENTOMOLOGICAL - ~- WNSTITUTE; ~-- ·- . ~ - - -- ICHNEWS Editor: Michael J. Sharkey Biological Resources Division/CLBRR Agriculture Canada Ottawa, ON KIA OC6, Canada Alternate Editor: David P. Wahl American Entomological Institute 3005 SW 56th Ave. Gainesville. FL 32608; USA Number 14 April1994

ichnmeumon-flies, or braconid from effectively communicating our EDITOR'S wasps. I'm talking about a nice, science to the public (and ultimately COMMENTS wholesome, down to earth, simple to our administrators). Lots of other common name like butterfly, bug, less diverse, and probably less eco­ ant, beetle, etc. Here we are talking nomically important groups, get Here is another issue of k.h.:. about one of the most diverse line­ more attention by the public simply ~. not too late this time. The next ages of living organisms (conserva­ because they do have a common issue will be published in early 1995 tively estimated at 50,000 species), name (e.g. grasshoppers, butterflies, and will cover newsworthy items one of the most economically impor­ ants). Do you think Holldobler & from 1994, therefore please respond tant groups of organisms (enor­ Wilson would have won the Pulitzer to the enclosed questionnaire ~ mously valuable because of Prize for 'The Ants' if the called their the end of 1994, so that you can biological control applications), and book, The Formicidae of the World? accumulate a year's worth of news. certainly one of the more fascinating I think not. Obviously, what we need Most of the items in this issue are groups in terms of its diversity of life is a really neat common name for this taken from responses sent to me al­ history strategies. and still we do not most important group, the Ichneu­ most a year ago. Fortunately, for the have a single common name to unify monoidea. Something to capture the newsletter, what was current a year the group. Surely this is one of the attention of the public, and to stimu­ ago remains so today. great, quirky artifacts of the history late and attract students to the study There are a number of items to ofscience! At first glance it's tempt­ of these wonderful organisms. The respond to in this issue of lchnews. ing to think that the ichneumonoids International Commission of Zoo­ In my article on venation on page have been overlooked in this regard logical Nomenclature doesn't apply (??) I request feedback. I would also because oftheir generally small body to common names, so who can say like to see some responses to Scott size. But how can that be the case that we shouldn't make one up. Shaw's article concerning a common when the tiniest creatures in the Having actually given this matter name for the Ichneumonoidea. Hymenoptera, the mymarids, do considerable thought, my suggestion have a common name (fairy tlies). is "Death Wasps." Why "Death Ultimately, we the scientists who Wasps" you ask? Why not? What DEATH WASPS work with braconids and ichneu­ feature more succinctly describes the monids must take the blame for not ichneumonoids than their ability to by Scott R. Shaw addressing this problem, for if we bring death to the hapless host vic­ University of Wyoming don't deal with it. who will? tim. With only a few exceptions they Do we need a common name for are true parasitoids, eventually kill­ the lchneumonoidea? What good ing the host insect. And it is this It really bugs me that we don't will it do us? Maybe it won't make ability to cause death that makes have a decent common name for the any difference at all, but then again them valuable to us (as hiocontrol Ichneumonoidea. I'm not talking maybe it might. Consider that the agents). Certainly the public has a about a modified derivation of the absence of a simple common name real fascination with parasitoid life scientific name like ichneumons. for our group may he preventing us style. Just look at the success of the lchnews 14, April 1994

"Alien" movies. The public is fasci­ he, is Dr. Wahl has kindly passed To the chalcidologists: yes, some nated with stories of parasitism, but along a few suggestions that Bra­ chalcids do spin silken cocoons, but it's up to us to relay the information conologists can deliberate. "The with .anal silk glands. It seems fitting in a form they can understand. Fi­ oldest common name for the Bra­ somehow, where chalcidoids mani­ nally, many generic names relate to conidae known to Dr. Wahl is 'The fest a reversal, they get it ass back­ the ability of the wasp to hunt, attack, supplementary Ichneumon-flies'. wards. Please give us your or kill (e.g. Microctonus, little mur­ The reference is Sharpe, D. 1895. suggestions for the next issue of~ derer; Orion, the hunter, a reference Peripatus, Myripods, Insects (Vol. 5 ~. to the sword of Orion; Doryctes, of Cambridge Natural history Se­ hunter with a spear; Sesioctonus, ries)., p. 558. Another name is 'Gar­ moth killer; Leptodrepana, slender bage Ichneumons'." According to ANOTHER LOOK AT sword). You can probably find other Dr. Wahl this probably refers to the WING VEIN/CELL examples. propensity ofsome alysiines and opi­ NOMENCLATURE. Critics might argue that the name ines to habituate foul environments. "Death Wasps" could be applied to One could simplify these names to Michael J. Sharkey any other group of parasitoid wasps 'garbage wasps' or 'filth wasps' or Agriculture Canada as well. Why restrict its use to the 'filthy garbage wasps' and have Ichneumonoidea? Well, there is the quite a catchy name; although, per­ As many readers know, a group matter ofhistorical precedence, since sonally I would like to have one of of us, known as the Parasitic the earliest observations of parasi­ these names apply strictly to the Opi­ Hymenoptera Advanced Research toid behaviour were made with the inae and Alysiinae (and this opinion Team (PHART), have embarked on Braconidae. The first published ac­ is unrelated to the fact that I would a project to key the New World gen­ count of insect parasitism is claimed be able to say that Bob Wharton is a era of Braconidae. One problem that to be Aldrovandi's 1602 work that specialist in filth wasps). By the we have had to overcome is to agree includes an account ofthe gregarious way, if Dr. Wahl is right in thinking on wing nomenclature. Most of us parasitoid Cotesia glomerata attack­ that Ichneumonologists are content are settled on the fact that we will use ing larva of the cabbage butterfly. with the common name Ichneumon­ some version of the Comstock­ "Am I mad," you say? Perhaps. flies, perhaps Braconologists might Needham system. But I challenge any of you to come do well to consider Braconid-bugs? In the paragraphs that follow I up with a better common name for All this bantering still leaves the will outline the Comstock-Needham our group. Certainly we have all had superfamily without a common system as it has been employed, for the experience of watching some­ name; and I agree wholeheartedly example, in the "Hymenoptera Of one's eyes glaze over (Roll back, and with Dr. Shaw that coining one is a The World". Following this I will flutter) as you begin to explain what valuable aim. I'm of the opinion that introduce some modifications that you do for a living. The next time 'death wasps' is a little too dramatic we are thinking of incorporating. I you are at a party and someone asks and offer a drier suggestion, 'silk would like anyone who has criti­ you what you do, tell them that you wasps or cocoon wasps'. This name cisms or suggestions for improving study Death Wasps, and see what refers to the fact that Ichneu­ the system to get in touch with me as happens. monoidea have retained the ability, soon as possible. as larvae, to produce silk with labial The Comstock-Needham silk glands. Ichneumonoids (with the system recognizes eight major longi­ "AM I MAD?" rare loss) use labial silk to spin co­ - A RESPONSE tudinal veins, abbreviated by capital TO coons and to tie leaves. Besides the letters. Starting from the anterior SCOTT SHAW. Aculeata, the only other hymenop­ margin of the wing they are: costa terans to have functional labial silk (C), subcosta (Sc), radius (R), by Michael 1. Sharkey glands are the Aulacidae, Gasterup­ medius (M), cubitus (Cu) and 3 anal B.R.D.. Agriculture Canada tiidae and Trigonalidae. These latter ( IA, 2A, 3A). In addition, there may groups can be safely ignored, and as be a short jugal (J) vein in some In a letter to the editorial depart­ for the aculeates, well, they already Symphyta. have their own ment of this newsletter Dr. Wahl common name. When a vein is branched the Cladists took some objection to the article by might be thinking that the most anterior branch is given the lack of functional Dr. S. Shaw. Specifically he was labial silk glands subscript I and the more posterior might rather mi !Ted by the suggestion that be a synapomorphy for the branches the subscripts 2, 3 .... This remainder of the Apocrita, the name Ichneumon-tlies was ridi­ i.e. the is done for all veins except R. When Apocrita minus culed. In his letter Dr. Whal noted (-) Aculcata, Ichneu­ R branches the most anterior branch monoidea, Trigonalidae, Gasterupti­ that it is the braconologist's who lack is called R 1 but the second branch is idae a common name for their group. Be­ and Aulacidae). I believe that called the radial sector (RS). ing the magnanimous individual that they would be right. 3

Two veins may fuse for part or all of their length, appearing as one vein. The resulting vein takes the name of both component veins joined by a plus ( +) sign. For exam­ ple, Rs and M are often fused for portions of their lengths. The fused portion is called Rs+M. Veins may fuse end to end so that it is impossible to know exactly where the first one ends and the second begins. In these cases the composite veins are joined with an ampersand (&). For exam­ ple, in all Ichneumonidae and many Braconidae the first abscissa of vein RS+M is often completely lost and in these cases veins lRS and 1M cannot be distinguished from one another: Figs. la & lb the composite vein is therefore termed RS&M. Wing cells, abbreviated with capital letters, take the name of the vein lying anterior to them. If sev­ eral fused veins form the anterior boundary of a cell, the cell takes the name of the vein that is theoretically most posterior. Thus, the cell poste­ rior to C+Sc+R is the radial cell (R). If more than one cell is directly be­ hind a vein. the cells are numbered consecutively from the base of the wing, e.g. three medial cells would be 1M, 2M, and 3M. New wing veins may arise in certairr lineages. Such veins are either given a new name or are named (often misleadingly) after the vein to which they are most similar in position. I follow the latter ap­ Figs. lc & ld proach for convenience but distin­ guish the new vein from its namesake by an apostrophe ( · ). Thus veins followed with an apostro­ A vein may have several seg­ Crossveins. indicated by phe(') are not (or probably not) ho­ ments or abscissae. They are delim­ lower case letters. take the name of mologous with the same vein lacking ited by the intersection of other the veins they connect, with the an­ an apostrophe('). veins, usually crossveins. Thus a terior vein given tirst. Thus. a cross­ vein that is intersected by two other vein that connects R with M is r-m. SUGGESTED ~IODIFICA­ veins has three abscissae. numbered If there are several r-m crossveins TIONS consecutively from the base to the they take numerical values as well apex of the wing. e.g.. when Cu has e.g. I r-m. 2r-m. etc. If a crossvein One problem inherent in the sys­ three abscissae. they are ICu (the joins two hranches of the same vein tem described above is that although basal portion of Cu). 2Cu and JCu the crossvein takes the name of the veins. crossvcins and vein branches (the apical portion of Cu). Vein ah­ rn~~jor longitudinal vein. e.g. a cross­ arc potentially homologous across scissac may vary imcrspccifically. vein he tween R1 and Rs is called r. the Hymenoptera. the abscissae :.lfC thcrcfl)rc ."\-Cu of one species is not This simplification is possible due to not This is hccause the ahscissae of necessarily homologous with 3-Cu the rarity llf this type of crossvein. a particular vein ~u·e named hased on of another. the posilion of veins and crossveins Ichnews 14, April 1994

CU whereas that of He/con fulvipes (Fig. lc) has only one component of abscissa ICU. Nonetheless abscissa ICU of Figure lc is easily homolo­ gized with ICU of Figure ld. The modified nomenclatorial system is illustrated using the wings of a xiphydriid (Fig. 2a and b) and of He/con fulvipes (Fig. 3a and b). These illustrate some fine points and conventions of the system which are discussed below.

a)When two longitudinal veins meet they are presumed to merge unless there is evidence to the con­ trary. For example veins IRS and 1M meet to become RS+M. When Fig. 2a two longitudinal veins meet they do not adopt the abscissae letters and numbers of their constituent veins, rather they are numbered inde­ pendently. So in the case of the xiphydriid fore wing (Fig. 2a) the correct name is RS+M not IRS+1M. In Helcon fulvipes (Fig. 3a) the RS+M vein is intersected from be­ low by crossvein m-cu therefore there are two sections to RS+M. i.e., (RS+M)a and (RS+M)b. The brack­ ets ( ) are used to imply that the sections a and b refer to the compos­ ite vein RS+M and not to the M component of RS+M which might be inferred if the vein was named RS+Ma. Notice that (by convention) C+SC both terminate at or before the Fig. 2b stigma ofthe fore wing or. in the case of the hind wing, at or before the point where R splits into R 1 and RS. that intersect h2.th anteriorly and To solve this problem the ab­ When two fused longitudinal posteriorly with the vein in question. scissa are numbered based only on veins diverge. the new abscissae of To see how this can be a problem intersection with veins or crossveins the constituent veins take on an extra refer to Figure Ia (Helconfulvipes). that meet anteriorly. When cross ve­ numerical value. Thus in the case of Note that there are three abscissae of ins meet the longitudinal vein in the xiphydriid (Fig. 2a), vein IRS vein CU. Now refer to the fore wing question from the posterior side a fuses with M to become RS+M and of He/con pedal is tFig. lb). Here letter is added to the name ofthe vein. when they separate they give rise to leu-a is positioned tO\vards the apex For example the abscissae of vein Cu 2RS and 2M tin this case 2M is di­ of the wing such that it intersects of Helconjidvipes are named accord­ vided into 2M a and 2Mb). ICU instead of heing inrerstitial with ing to this system in Figure l c and I would appreciate your com­ M ti have exaggerated the position those of He/con pedalis are named in ments. for case of illustration). In this case Figure I d. In ~omparing tigures I c there is an extra ahs~issa of CU and and Id not icc that all abscissae ofCU none of the ahs~issac l)f CU of the arc homologous. The only differ­ two wings tFigs Ia and lh) arc ho­ ence in the two wings is that the fore llllllogous. wing 1lf Ht'lcon pt>dalis (Fig. J) has lwol-·omponcnls lo abscissa I of vein 5

when the first tergite is strongly petiolate. laterope (singular/plural): pit­ like depression situated in the glymma in many Braconidae, close to the base of the frrst metasomal tergite laterally. precoxal sulcus: (not that new term, e.g. Richard, 1956). Common longitudinal depression of the mesopleuron in Braconidae. Run­ ning nearly horizontal and situated above the sternaulus (well known in the Ichneumonidae). The different nature of both can be observed in some Opiinae where both depres­ Fig. 3a sions are present. The sternaulus is clearly bent downwards anteriorly and situated more ventrad.

c

Fig. 3b

New Morphological terms: subposterior depression that may be present e.g. in Agathidinae, because by C. van Achterberg of different topology and sculpture National Nawurhisrorisclz A1useum (editor's note: I believe the depres­ Netherlands sion found in many Agathidinae to be homologous with those found in anterior subalar depression: other Braconidae e.g. most Helcon­ depression above subalar promi­ ini. nence (Richard. 1977); may be (medial) pronope (singular/plu­ smooth. crenulate or with a distinct ral): round or transverse depression transverse carina. Not to be confused medio-dorsally on the pronotum. with the subalar pit (Richard. 1977). lateral pronope (or sub­ Med io- posterior depression of pronope): round depression in the Cardiochilinae scutellum: a posterinr depression on sides of the pronotum anteriorly. the scutellum. whidl has a median dorsope (singular/plural): pit­ carina or (more nften) fine crenula­ like depression just behind the bases tion. Not to he ~onfused with rugos­ of the dorsal carin•te of the first me­ ity or other sculpture that can he tasomal tergite. The dorso.1l carinae present in ~ydostome l)r microgas­ may he absent and sometimes the troid groups. Also different from a dllrsopc hccomes hlterally situated 6

CALLING ALL GENERA by David Wahl American Entomological Institute

The American Entomological Institute strives to maintain as complete a collection ofIchneumonidae as possible. This is especially so for genera and subgenera. Of the 1500 +genera and subgenera that are considered valid, the AEI lacks 121; these missing genera are listed below. Ifany Ichnews reader has specimens ofthese genera and is willing to trade, please contact me. The AEI has an extensive set ofduplicates that is sure to delight even the most jaded tastes.

ACAENTINAE: Coleocentrini: Eremocinetus Viktorov, 1964; Palaearctic. Acaenitini: Boloderma Morley, 1913; Oriental.

ANOMALONINAE: Aubeniana Viktorov, 1970; Palaearctic. Clypeocampulum Gauld, 1976; Palaearctic. Gravenhorstia (Ribasia Ceballos, 1921 ); Palaearctic. Neohabronyx Dasch, 1979; Nearctic. Porlzonopteron Meyer, 1931; Palaearctic. Sphaeromanus Aubert, 1979; Palaearctic. Stangepelma Porter, 1976; Neotropical. Vernanomalon Gauld, 1976; Ethiopian.

BANCHINAE: Atrophini: Tossinolodes Aubert, 1984; Palaearctic. Banchini: Geraldus Fitton, 1987; Neotropical. Neoexetastes Graf, 1984; Neotropical.

CAMPOPLEGINAE: Allotheca Cameron, 1906; Ethiopian. Breviterebra Kusigemati, 1982; Palaearctic. Cymodusa (Diverdusa) Dbar, 1985; Palaearctic. Diadegma (Auma) Dbar, 1984; Palaearctic. Neolophron Gau1d, 1984; Australia.

CREMASTINAE: Eucremastoids Ko1arov, 1980; Palaearctic. Ricrena Cameron, 1906; Ethiopian.

CTENOPELIVIATINAE: Ctenopelmatini: Rhorodes Aubert, 1970; Palaearctic. Scolobatini: Dictyopheltes Gauld, 1984; Australia. Hypopheltes Cushman, 1924; Australia. Pergaphaga Gauld, 1984; Australia. Mesoleiini: /skarus Kolarov, 1987; Palaearctic. Semimesoleius Ozols, 1963; Palaearctic. Ichnews 14, April 1994

Euryproctini: Denticeria Gauld, 1984; Australia.

DIPLAZONTINAE Episemura Kasparyan & Manukyan, 1987; Palaearctic. Eurytyloides Nakanishi, 1978; Palaearctic. Schachticraspedon Diller, 1984; Neotropical.

ICHNEUMONINAE Ichneumonini: Amblysmenus Heinrich, 1965; Oriental. Archboldiella Heinrich, 1934; Oriental. Auritus Constantineanu, 1969; Palaearctic. Bonthainiella Heinrich, 1934; Oriental. Caspipina Cameron, 1903; Oriental. Celebichneumon Heinrich, 1934; Oriental. Celebijoppa Heinrich, 1934; Oriental. Clypeocava Heinrich, 1934; Oriental. Coelojoppa Cameron, 1904; Oriental. Cornutiplisus Heinrich, 1957; Palaearctic. Cryptoplites Heinrich, 1938; Ethiopian. Dammermanlella Heinrich, 1934; Oriental. Dammermaniellops Heinrich, 1974; Oriental. Dentichasmiops Heinrich, 1969; Ethiopian. Deuterolabops Heinrich, 1975; Oriental. Gibbobystra Heinrich, 1969; Oriental. H emibystrops Heinrich, 1969; Oriental. Hyemnura Townes, 1965; Palaearctic. Hytophatnus Cameron, 1907; Oriental. Larischia Heinrich, 1969; Ethiopian. Leptomalasia Heinrich, 1965; Oriental. A1adagasichneumon Heinrich, 1938; Oriental. _ Malaisichneumon Heinrich, 1965; Oriental. Marlisia Heinrich, 1975; Oriental. Monodontichneumon Heinrich, 1969; Ethiopian. Nonpropodeum Heinrich, 1934; Oriental. Pectinorex Graf, 1976; Neotropical. Phaisurella Heinrich, 1938; Ethiopian. Pseudevirochoma Heinrich, 1969; Ethiopian. Rimbusia Heinrich, 1980; Oriental. Serratosculum Heinrich, 1969; Ethiopian. Stenichneumonopsis Heinrich, 1934; Oriental. Stenogynaia Heinrich, 1965; Oriental. Tlzaumatocephalus Heinrich, 1965; Oriental. Thaumatoplitops Heinrich, 1969; Ethiopian. Togeella Heinrich, 1980; Oriental. Heresiarchini( =Protichneuonini): Clzasmopyglum Heinrich, 1967; Ethiopian. Gathetus Cameron, 1901; Oriental. Lissophadnus Cameron, 1907; Oriental. Lllteocoelius Heinrich, 1968; Oriental. Matimmgarches Heinrich. 1934; Oriental. Neoheresiarches Uchida. 1937; Oriental. Pseudocillimops Heinrich, 1969; Oriental. Punctileptops Heinrich, 1967; Ethiopian. Stenapatetor Heinrich. 1938; Ethiopian. Trogichneumon Heinrich. 1968; Oriental. Trogopyga Heinrich. 1969; Oriental. 8

Platylabini: Acantholabus Heinrich, 1974; Oriental. Car/sonia Heinrich, 1973; Nearctic. Clypeolabus Heinrich, 1974; Oriental. Cratolabus Heinrich, 1974; Oriental. Ectopoides Heinrich, 1951; Oriental. Hoploplatystylus Schmiedeknecht, 1912; Palaearctic. Lissolaboides Heinrich, 1974; Oriental. Neeurylabia Heinrich, 1967; Ethiopian. Platybirmania Heinrich, 1974; Oriental. Ctenocalini: Listrocalus Heinrich, 1978; Palaearctic. Compsophorini: Poecilodromus Heinrich, 1975; Oriental. Pyramidamblys Heinrich, 1967; Ethiopian. Listrodromini: Poecilodromops Heinrich, 1975; Oriental.

ORTHOCENTRINAE Atabulus van Rossem, 1988; Palaearctic. Epitropus van Rossem, 1990; Palaearctic. Fetialis van Rossem, 1990; Palaearctic. Kentrotryphon Strobl, 1902; Palaearctic. Pantomima van Rossem, 1990; Palaearctic. Phosphoriana van Rossem, 1987; Palaearctic.

PAXYLOMMATINAE Eurypterna Foerster, 1962; Palaearctic. Ghilaromma Tobias, 1988; Palaearctic.

PHRUDINAE /caromimus Seyrig, 1932; Ethiopian.

PHYGADEUONTINAE Phygadeuontini: Apotemnus Cushman, 1940; Nearctic. Ceplzalobaris Kryger, 1915; Palaearctic. Diaglyptellana Horstmann, 1976; Palaearctic. Dolichomastax Ceballos, 1924; Ethiopian. Notostilbus Townes, 1983; Holarctic. Mesostenini: Cyanodolius Seyrig, 1952; Ethiopian. Hackerocryptus Gauld, 1984; Australia. Jonathania Gupta, 1987; Oriental. Piambia Seyrig, 1952; Ethiopian. Pseudotricapus Jonathan, 1987; Oriental. Savolia Seyrig, 1952 Ethiopian Tretobasis Porter, 1973; Neotropical Xylacis Porter, 1987; Neotropical

PIMPLINAE Polysphinctini: Afrosphincta Benoit, 1953; Ethiopian

TERSILOCHINAE Diaparsis (Pseudaneuclis Horstmann, 1971 ); Palaearctic Ichnews 14, April 1994

Horstmannolochus Gauld, 1984. Australia

TRYPHONINAE Phytodietini: Brevitubula Wang, 1983; Oriental Tryphonini: Parablastus M. Constantineau, 1973; Paleartic Exenterini: Eremodolius Kasparyan, 1985; Palaearctic Schelocentrus Kasparyan, 1976; Palaearctic

The Adventures of ICHMAN

MeANW'HIL.E', TN£HY11EN HAVE .S'IF, /NTHIIIR CLIJTCNE$. 10

THE PUBLICATION DATES OF HEINRICH'S "SYNOPSIS AND RECLASSIFICATION OF THE ICHNEUMONINAE STENOPNEUSTICAE OF SOUTH AFRICA AND THE SAHARA"

by David Wahl ( 1987) in which Paraphyscoteles American Entomological Heinrich is cited as being described Institute in 1968 but with the bibliography giving an unrelated publication for Heinrich's 5-volume magnum that year. opus on Ethiopian ichneumonines is Literature Cited commonly cited as 1967-1968 (He­ Gauld, I.D. 1984. An introduc­ inrich (1977)) or 1967 (Gauld (1984) tion to the Ichneumonidae of Austra­ and Gupta (1987)). Townes and lia. British Museum (Natural Townes (1973) cite the volumes as History), London. 413 pp. covering the span 1967-1969. Henry Gupta, V.K. 1987. The Ichneu­ Townes' copy of volume V in the monidae of the Indo-Australian area American Entomological Institute li­ (Hymenoptera). Memoirs of the brary has the dates he received his American Entomological Institute copies written adjacent to the printed 42: 1-1210. dates of issue on page 1258. They Heinrich, G. 1977. Ichneumoni­ are as follows (the printed dates of nae of Florida and neighboring states issue are in brackets): (Hymenoptera: Ichneumonidae, sub­ volume I - December 20, 1967 family Ichneumoninae ). Arthropods [April 3, 1967] of Florida and neighboring areas 9: volume II - December 20, 1967 1-350. Dirrhopinae [June 28, 1967] volume III - June 18, 1968 [De­ cember 21, 1967] volume IV - February 18, 1969 [June 20, 1968] ICHNEUMONIDAE GENERA AND SPECIES IN THE volume V- August 8, 1969 [No­ UNIVERSITY OF MINNESOTA COLLECTION vember 10, 1968] Henry Townes wrote the follow­ by John Luhman Campopleginae (using new clas­ ing passage below the dates; it is Minnesota Dept. ofAgriculture. sification): 4 Tribes/30 Genera/31 reproduced here in full: "In August, spp., about 1000 mostly Helictini un­ 1969, I asked Hilda Heinrich why the The University ofMinnesota In­ det. above dates are so much earlier than sect Collection, St.Paul, is estimated Ctenopelmatinae: 5 Tribes/45 the dates on which I received copies. to be the 8th largest in the U.S. for Genera/61 spp. (includes 9 spp. of Her reply indicates that the above the number of identified ichneu­ Perilissus). dates are the ones on which the books monids. It has over 1,200 species, Microleptinae Cylloceria 113. Diplazontinae: 9/41. were~ in Germany. After this, and 40,000 specimens, are deter­ a copy was sent air mail to Gerd mined to at least genus. Those need­ Ichneumoninae (still working, Heinrich & the rest came by ship, ing specimens for revision will find with 85% identified): Phaeogenini- through customs, and to Farmington most Nearctic, and some Neotropical 5/5, and about 600 undet. to genus; State College. She believes that I am material for most taxa. Recently Gyrodonmini- 311137; Ichneumon­ among the first subscribers to receive much Neotropical material from ini (=Protichneumonini:Heinrich)- copies. This means that the publica­ Costa Rica has been added. I have 3/24; Listrodromini- 2/2; Platy­ tion dates are 7-10 days prior to the identified all of the undetermined labini- 4113; Trogini- 6/9 + 9 undet. dates that I received my copies. H. ichneumons in the following sub­ tropical species; Acanthojoppini- Townes. Aug. 1969." families, with diversity shown by 3/ I,+ 4 undet. tropical species. Sum­ Given this information, the dates numbers (by tribe) genus, and spe­ mery: 7 Tribes/54 Genera/191 spp., of publication should be regarded as cies (Genus/species). Those wishing + 13 undet. tropical species. those listed by Townes (and used in loans should write to the curator, Dr. Labeninae: 2/2, + 4 undet. tropi­ Townes & Townes, 1973). Sys­ Phil Clausen, Dept. of Entomology, cal Labena tematists and users of catalogues Hodson Hall, Univ. of Minnesota, Pimplinae: Pimplini- 14/48; should take these into account, St. Paul, MN 55108, USA. Polysphinctini-9/22; Delomeristini- avoiding such mixups as Gupta 3/7; Siacritini- 1/1; Echthromorphini Ichnews 14, April 1994

(=Ephialtini)- 7/33; Therionini-2/4; this work: 1) it was put together by a Pimplini from the United States Poemenini (=Neoxoridini)-3/6; non-systematist without any input gives species of both Pimp/a and Rhyssini- 4/12; Sum= 8 Tribes/43 from specialists; 2) it uses a species Coccygomimus. Any competent Genera/133 spp., + several undet. nomenclature that is not in accord­ specialist knows that these are one tropical species. ance with the International Code of and the same genus, with the name Xoridinae: 3/21. Zoological Nomenclature (hence­ preference depending upon whether All other subfamilies are also forth referred to as "the Code"), and one follows the ICZN and its Opin­ represented except Agriotypinae and 3) it is riddled with errors and incon­ ions. As far as I can make out, if a Phrudinae. sistencies. I shall deal with each of species was originally described in these in tum. Pimpla, it is kept there even if treated 1) Decisions are made and spe­ as Coccygomimus by later authors; if REVIEW cies assigned according by naive in­ described in Coccygomimus but terpretation of recent literature, never placed in Pimpla, it remains in rather than by the author's judge­ Coccygomimus. Another example is FORWARD INTO THE ment or informed scholarly opinion in Ichneumon. The numerous West­ PAST by advisors who are more cognizant em Palaearctic species have never revised and a [Review of TAXA, Dicky Yu, of the situation. When two nomen­ been authoritatively of various ichneumonine (see enclosed notice) clatures or taxonomic interpretations wide array are at odds, Yuhas followed the most genera (or even non-ichneumonines) under the name. I printed out a by David Wahl recently published paper, rather than lurk of American Entomological Institute querying specialists (TAXA work­ list of 271 species and subspecies After checking Gainesville, Florida book, p. 1). He is " ... not trying to German Ichneumon. create a 'catalogue' which would be a variety of references that dealt with a declaration of what I think the tax­ the genus as it is perceived today, I In terms ofaccessibility and ease onomy of the group should be, but a found that the true affiliation of 169 of study, ichneumonids have the system that reflects the published in­ names is simply unknown; these in­ odds stacked against them. The formation available. If this means clude many species by Christ, sheer size of the family, with at least the system may yo-yo as taxonomists Schrank, and other early authors 60,000 species worldwide, has dis­ argue back and forth through their which have not been dealt with since couraged many potential workers. publications, so be it." (Yu. pers. their description. The most prudent Then there are the various historical comm.). This database is a de facto course of action would been to create factors: the 1869 Forster names; the catalogue, however, although the a separate category of "Ichneumon proliferation of names in the late author may not chose to call it such, incertae sed is" for all names not 19th - early 20th centuries that cre­ especially for other hymenopterists dealt with in modern publications. ated a situation in which it was al­ and applied workers. I know of no One may be sure that non-specialists most impossible to identify taxonomic listing of similar breadth will interpret these species as all be­ extra-Holarctic species; the Opin­ and aims that has not been the work longing in the genus and thus pro­ ions of the International Commission of a specialist and has similarly re­ duce conflated and misleading on Zoological Nomenclature (ICZN) flected the compiler's specialized figures for various purposes. on certain crucial type species; knowledge. For biological control 2. Perhaps the single most damn­ Townes' use of a idiosyncratic fam­ specialists and other scientists with ing aspect of this database is Yu's ily-group nomenclature at odds with similar needs, this is one of the big­ new system ofspecies nomenclature, that used by other zoologists; and gest strikes against TAXA. Applied which he discusses at length in Yu finally the lack of a catalogue for the workers need consistent and reliable ( 1993). He rightly points out that the important fauna of the Western Pa­ nomenclature. As identifiers. we in recombinations of species under dif­ laearctic. One would expect that a the systematics profession strive to­ ferent genera, coupled with the. ne­ database of the world ichneumonid wards this goal by examining the cessity to conform with Latin fauna would be a tremendously use­ arguments for the application of a grammar, creates homonymy that ful tool for organizing information name and then sticking by them until will interfere with computerized in­ for systematists and applied workers. such time as new evidence is brought formation retrieval systems. He is Unfortunately, the database re­ forth. This concern is absent in also right in stating that the original viewed here is instead a step back­ TAXA and the use of this database by binomen is the only stable anchor ward in ichneumonid studies and non-systematists is against their own that can correctly associate name must be placed with the hindrances best interests. variations. Unfortunately, he pro­ list\!d above. It is one of the most This deliberate blindness toward poses that the rules of nomenclature confusing and destabilizing taxo­ argumentation and evidence has led be changed so that the original bino­ nomic creations that I've examined. to egregious blunders. Two TAXA men becomes a permanent part ofthe There arc three major strikes against examples will suffice. A listing of scientific name. leading to such un- 12

wieldy monstrosities as Stirexepha­ they are not available (Article 45, tion ofGupta ( 1987) who placed it in nes signatus Togea formosana e-f). I randomly came across these the Acaenitini. Cursory examina­ Uchida, 1926. Even worse, the sys­ examples while checking taxa ofper­ tion of specimens reveals Gupta's tem has been implemented in the da­ sonal interest and no doubt many placement to be a mistake. Such an tabase as if this was accepted more exist. anomalous record should have been procedure. This is all needless. I've c) Ninety-six holotypes of the referred to a specialist for clarifica­ discussed the situation with various American Entomological Institute tion if the compiler was not qualified systematists who are putting together are erroneously listed as belonging to to investigate; instead, error is per­ large taxonomic databases and they the Smithsonian Institution. The petuated. point out that changing the Code is publication discussing the situation g) ANOMALONINAE. Theri­ unnecessary: relational database pro­ (Townes & Townes (1983)) was not ini Viereck, 1918 is not listed as a grams can incorporate the original examined. synonym of Gravenhorstiini (treated bin omen as background keys that are d) Carlson (1979: 738-739) as a subfamily in TAXA). invisible to the searcher but yet will listed 21 Nearctic species as un­ h) BANCHINAE. Lissonotini, associate homonyms. TAXA, how­ placed to subfamily or genus, 15 of which is used here, is a junior syno­ ever, is not a relational database and which had the type missing. Yuhas nym of Atrophini (Gauld 1984). Ar­ the unwieldy association of current uncritically assigned the latter to choprotus porteri Brethes, whose combinations with original bino­ various genera. For example, Lim­ type is lost, was treated by Townes mens is Yu's solution. Basically, the neria illepida Cresson is placed in & Townes (1966) as a possible atro­ zoological community is being asked Olesicampe since Limneria is a jun­ phine. Yuhas placed it, with no ex­ to make unnecessary nomenclatural ior synonym ofthat genus. Asymmi­ planation, in the Banchini. changes due to deficient program­ ctus nigrofrons Davis is placed in i) CAMPOPLEGINAE. This ming. Alloplasta on similar grounds. It is subfamily has more than its share of 3. a) The endings ofjunior syno­ imprudent to place species in this mistakes. Similar to the case of nyms are changed, as an automatic manner and they should have been Nearctic Pimp/a mentioned above, function of TAXA, to the gender of treated in a manner similar to Nearctic Dusona has been split into the genus under which they are cur­ Carlson's. Dusona and Delopia: species from rently placed. This is not required by e) Townes ( 1971) listed 11 fos­ this area are in Dusona (if one ac­ the Code nor followed in any other sil ichneumonid genera, three of cepts Gauld ( 1984) on the matter), work of this nature. Valid names are which were assigned to subfamilies. but not in both! Even though Gupta not exempt from this either, as Ben­ Since then, six more genera have ( 1987) rejected the record of Micro­ jaminia maurus is changed to Ben­ been described, placed either in their charops taitica from Tahiti as an er­ jaminia maura even though the name own subfamilies (Tanychorinae and ror, it is listed here as from that was put forth as a noun in apposition Pherhombinae) or in the tryphonine island. It is with the tribes, however, (Wahl, 1989). While on the topic of tribes Tryphonini, Oedemopsini, and that massive mistakes are made. A specific names, I note that misspell­ ldiogrammatini. With the exception maximum of five tribes are usually ings of specific epithets are rather of Pherhombus, Yuhas placed them recognized (Townesian names in common. all in the Ephialtitidae. which is pe­ brackets): Nesomesochorini, Hell­ b) Species names proposed as culiar considering that he cites no wigiini, Cymodusini, Campoplegini "variety" or "form" are not listed as publications that would justify such [= Porizontini], and Limneriini [= such in synonymies. An example is action (and there are none to my Macrini]. Yuhas managed to use six Neotheronia lineata var. concolor knowledge). The fossil ichneu­ tribes, using both Campoplegini and Krieger, 1905, which is listed simply monid subfamily Tanychorinae Porizontini. Campoplegini is used in as Neotheronia lineata concolor. (Rasnitsyn, 1980) is not even listed. the traditional sense (consisting of Some names proposed as varieties Ephialtitidae and Ichneumonomidae Campoplex and related genera; after 1961, such as Alomya debella­ are placed in the lchneumonoidea, Gupta ( 1987) places Neolophron and tor var. flavotibiata Constantineanu, even though Rasnitsyn ( 1988) ex­ Eucaphila here, whereas Yu places 1965, or A. d. var. nigrita Constan­ cluded these taxa from the superfa­ them in his Porizontini). Porizontini tineanu, 1965, are listed as subspe­ mily. is used as well, however, and the cies. Neither one is available under (The following sections list er­ contents of Limneriini are split be­ the Code (Article 45. e-g). In the rors discovered while randomly tween the two tribes! Related genera Acaen i ti nae, Lissonota ornatus checking subfamilies and taxa of are placed in different tribes (Me/a­ Hellen, 1937 and trochanterus personal interest. The reader should boris and Nepiera, various genera of Hellen, 1937 are listed as junior keep in mind that many more un­ the Hyposoter complex, Diadegma synonyms of Coleocemrus excita­ doubtedly exist) and Enytus) contrary to all previous tor: as hoth were proposed as aberra­ t) ACAENITINAE. All authors usage. tions (sec Horstmann. 1990: 183 for have considered Leptacoenites to be j) ICHNEUMONINAE. The a uiscussion ofHellt~n 's aberrations). in the Coleoccntrini, with the cxccp- mistakes in the Campoplcginae tind Ichnews 14, April 1994

their match in this subfamily. 1) Gyrodontina. In particular, Steno­ ance in a paper dealing with Oxytori­ Family-group names are incorrect. barichneumon, Vulgichneumon, nae in a less restricted sense (van IfAlomya is to be placed with Phaeo­ Richtichneumon, Virgichneumon, Rossem, 1990). genes, the tribal name is Alomyini, Carinodes and Paraditremops form m) PHYGADEUONTINAE. as Townes & Townes (1951: 276) a well-defined complex centred The unfortunate decision was made made a first-revisor decision in be­ about Melanichneumon, which was to follow the few authors (Gupta, tween the two. Yu uses Diller's correctly placed in Gyrodontina. Er­ 1970, 1987; Jonathan & Gupta, (1981) subtribes and leaves out the rors of this type can be found all over 1973) who elevated the tribes to subtribe Phaeogenina, leaving a -- Afrotrogus and Pepsijoppa are separate subfamilies (viz., Phy­ large number of genera floating placed in the Trogina, instead of the gadeuontinae s.s., Hemigastrinae, without subtribal affiliation. Gyro­ Callajoppina where Heinrich ( 1967) Mesosteninae). As pointed out by dontina should take precedence over placed them. The most inexplicable Gauld (1984: 101), the tribes are Cratichneumonina. Similarly, Her­ action, however, is the placement of closely related and no other authors esiarchini is the correct name for 19 Neotropical genera in the Ichneu­ recognize this action. Protichneumonini if Heresiarches monina in the absence of any publi­ n) PIMPLINAE. Eremochila is and its relatives are kept within this cation. As Heinrich ( 1967) did not placed with the Pimplini (sensu Fit­ tribe (as Yu does). 2) I was surprised propose subtribes in Ichneumonini ton & Gauld (1976) -- due, I sup­ to find Amblyteles recorded from until after Townes & Townes ( 1966), pose, to confusion over the North America, thinking that the ge­ the Neotropical ichneumonin fauna conflicting tribal nomenclature in the nus contained only one species, the was not assigned to subtribes. Need­ subfamily. Palaearctic armatorius Forster. The less to say, the same errors afflicting o) TATOGASTRINAE. Wahl Nearctic species in question, Am­ the Melanichneumon complex men­ ( 1990) is cited as providing a larval blyteles crudosus Cresson, was tioned earlier apply to this fauna as figure for Tatogaster. As stated in placed in the genus by Townes well. 5) In the Alomyini, both the paper, the larva is unknown. (1945) although Carlson (1979) had Phaeogenes and Tycherus are used: p) TRYPHONINAE. Gupta deliberately left it as unplaced in researchers who use Tycherus do so ( 1988) restricted the Eclytini to Amblytelina. Ichneumonine taxon­ in the sense of Diller (1981), restrict­ Eclytus, and this was maintained in omy has advanced considerably ing Phaeogenes to semivulpinus Townes et al (1992). Thymaridini is since 1945 and unpublished notes of Gravenhorst and its relatives. As a junior synonym of Oedemopsini Townes in the AEI indicate that it is used in TAXA, Phaeogenes contains (Carlson, 1979). Yet in TAXA, oede­ a species of Diphyus. A search of numerous species that should be mopsin genera are randomly distrib­ TAXA found I ll species were re­ placed in Tycherus. Even if one does uted between Thymaridini and corded in Amblyteles. These turned not agree with the separation of Ty­ Oedemopsini. Acaenitellus is in out to be mostly species described in cherus, the situation here is guaran­ Thymaridini, even though Gupta the late 19th - early 20th century by teed to please no one. 6) The (1988) explicitly put it in Oede­ Rudow, Berthoumieu, Tischbein, publication dates of Heinrich's mopsini. Eclytus is also in Thy­ and other European authors when monographic series on Ethiopian maridini. As far as I can make out, Amhlyteles was a considerably more ichneumonines are incorrect (see the genera in Oedemopsini are those diffuse concept than today (the treat­ note on this subject, this issue of mentioned in Townes et al (1992); ment of Amblyteles by Constan­ lchnews); those given in Townes & thymaridin genera are those not men­ tineanu ( 1959) and Perkins ( 1960) is Townes ( 1973) should have been fol­ tioned (with the exception of Thy­ in the old style). As in Ichneumon, lowed. maris which apparently stayed with prudence should have dictated Yu' s k) LABENINAE. Labiini is the tribal name. actions. 3) Acanthojoppini and Jop­ used instead of Groteini, even Some final comments relating to pocryptini are kept separate, al­ though it is a junior synonym of the appearance and utility of TAXA: The though Heinrich ( 1977: 268) clearly latter. The Clasini are retained in this presentation of data on screen is indicated that Acanthojoppini was subfamily, even though Gauld rather primitive, below the standards the junior synonym. 4) Heinrich ( 1983) demonstrated that they do not of other taxonomic databases that ( 1977) and Carlson ( 1979) assigned belong here and should be regarded I've examined. In particular, syno­ the Nearctic Ichneumonini to sub­ as a tribe of Phygadeuontinae. nyms are presented in an unhelpful tribes. With no revisionary work on I) OXYTORINAE. Oxytorinae, format. Arrangement is by the cate­ this fauna since these publications, although otherwise seeming to con­ gories "synonym" and "junior syno­ why was the subtribal association so form to the restricted definition in nym" and then within each category badly mishandled? Homotherus, Wahl ( 1990), includes Epitropus, numerically by taxon number. Anisopygus, Limonethe, Ectopimor­ Fetialis, and Palllomima. These Worst ofall, past synonyms are listed pha. Stenobaridmeumon and a host clearly belong in the Orthocentrinae even though those names have been nf other genera are in the lchneu­ (Helictinae of authors); Yu was ap­ transferred out of the taxon under monina; they should he placed in parently confused by their appear- examination. Using Campoplex as 14

an example, Sinophorus (with its (I wish to thank the following __. 1987. A revision of the junior synonym Eulimneria) and persons for input during the prepara­ genus Microcharops (Hymenoptera: Sesioplex are listed as synonyms tion of the review: John Barron, Ian Ichneumonidae). Contributions of (even though they are recognized Gauld, Gary Gibson, Klaus the American Entomological Insti­ elsewhere as valid genera), along Horstmann, Charles Porter, Mike tute 23: 1-42. with Phaedroctonus (a junior syno­ Sharkey, Larry Spears, and Chris ___. 1988. Relationships of nym of Porizon and listed as such Thompson.) the tryphonine tribe Oedemopsini under that genus). This convoluted and a revision of Acaenitellus Mor­ practice serves only to confuse the Literature Cited: ley. In V .K. Gupta (ed. ), Advances in unwary who lack literature re­ parasitic Hymenoptera research. sources. Finally, about 7 megabytes Carlson, R.W. 1979. Family Ich­ E.J. Brill, Leiden/New York. 546 pp. of scanned digitized images are in­ neumonidae. In K. V. Krombein, cluded. After installation of TAXA, P.D. Hurd, D.R. Smith, & B.D. Heinrich, G .H. 1967. Synopsis the first priority of the user should be Burks (eds.), Catalog of the and reclassification of the /chneu­ to dump the image files. The images Hymenoptera in America North of moninae Stenopneusticae of Africa are so crude that no really usable Mexico, Vol. 1. Smithsonian Institu­ south of the Sahara. 1. Farmington information is presented. I have a tion Press, Washington, D.C. University Press, Maine. 250 pp. VGA screen and about all I could say Constantineanu, M.I. 1959. Fa­ __. 1977. Ichneumoninae of was "well, yes, this looks like a cam­ milia Ichneumonidae, tribus Ichneu­ Florida and neighbouring states poplegine ... " moninae Stenopneusticae. Fauna (Hymenoptera: lchneumonidae, sub­ After all the errors of commis­ Republicii Populare Romine 9( 4 ): 1- family Ichneumoninae). Arthropods sion and omission listed above, and 1248. of Florida and Neighbouring Land the reader should remember that Diller, E. 1981. Bemerkungen Areas 9: 1-350. these were found after only cursory zur Systematik der Phaeogenini mit Jonathan, J.K. & V.K. Gupta. examination, the question arises einem vorHiufigen Katalog der Gat­ 1973. Ichneumonologia Orientalis. whether TAXA is of any utility. In­ tungen (Hymenoptera, Ichneumoni­ Part III. The Goryphus-complex deed it is, but to only about 15-20 dae). Entomofauna 2: 93-109. (Hymenoptera: Ichneumonidae). people worldwide. These are ich­ Fitton, M.G. & I.D. Gauld. 1976. Orientai Insects Monographs 3: 1- neumonid specialists who will find The family-group names of the Ich­ 203. the list of literature exceedingly use­ neumonidae (excluding Ichneumon­ Perkins, J.F. 1960. Ichneumoni­ ful (which it is!) or will generate lists inae) (Hymenoptera). Systematic dae, Ichneumoninae II, Alomyinae, of species for various taxa or areas. Zoology 1: 247-258. Agriotypinae and Lycorininae. I found it useful for finding over­ Gauld, I.D. 1983. The classifica­ Handbooks for the Identification of looked species in various groups of tion, evolution and distribution ofthe British Insects 7 (2aii): 117-213. my current research interest. West­ Labeninae, an ancient southern Rasnitsyn, A.P. 1980. [Origin ern European workers will appreci­ group oflchneumonidae (Hymenop­ and evolution of Hymenoptera.] ate it as a crude substitute for a tera). Systematic Entomology 8: 167- Transactions of the Paleontological catalog of that area. BUT: only these 178. Institute ofthe Academy ofSciences people will have the expertise and __. 1984. An introduction to ofthe USSR 147: 1-134. collection/literature resources to fil­ the /chneumonidae of Australia. __. 1988. An outline of evo­ ter out the errors that affect this data­ British Museum (Natural History), lution of the hymenopterous insects base. The benefits of TAXA to a few London. 413 pp. (Order Vespida). Oriental Insects are far outweighed by its potential __. 1991. The Ichneumoni­ 22: 115-145. harm to many. TAXA cannot be used dae of Costa Rica, I. Memoirs ofthe van Rossem, G. 1990. Key to the as a reliable guide to current ichneu­ American Entomological Institute genera ofthe Palaearctic Oxytorinae, monid taxonomy. Unless Yu puts 47: l-589. with the description of three new the species nomenclature into ac­ Gauld, I.D. & B. Bolton. 1988. genera (Hymenoptera: lchneumoni­ cordance with current practice and The Hymenoptera. British Museum dae). Zoologische Mededelingen 62: brings an advisory panel of special­ (Natural History) & Oxford Univer­ 309-323. ists on board to compensate for his sity Press, Oxford. 332 pp. Townes, H., V.K. Gupta, & M. unfamiliarity with ichneumonids, Gupta, V.K. 1970. Ichneumon Townes. 1992. The ichneumon-flies TAXA will remain a bountiful fount hunting in India. Delhi. I09 pp. of America north of Mexico. 11. of bad scholarship to plague ichneu­ __. 1987. The Ichneumoni­ Tribes Oedemopsini, Tryphonini, monid systematics for years to come. dae of the Indo-Australian area and ldiogrammatini (Hymenoptera: I cannot recommend its purchase to (Hymenoptera). Memoirs of the Ichneumonidae: Tryphoninae). hymenoptcrists or the general ento­ American Entomological Institute Memoirs of the American Entomo­ mological community. 41: 1-1210. logical Institute 50: 1-296. Ichnews 14, April 1994

Townes, H. 1945. A catalogue Current projects: Work on Me­ and reclassification of the Nearctic PERSONAL NOTES topiinae of the Palaearctic. Ichneumonidae. Memoirs of the Collection sites: Mediterranean American Entomological Society 11: area. 478-925. (Na­ __. 1971. The genera of Ich­ . Kees van Achterberg tional Natuurhistorisch Museum· Alexandre P. Aguiar (Pur­ neumonidae, part 4. Memoirs of the Postbox 9517, 2300 RA Leiden' due University; 1158 Entomology American Entomological Institute NETHERLANDS). ' Hall West Lafayette, IN, 47907- 17: 1-372. Current projects: 1. Revision of 1158, USA). Townes, H. & M. Townes. 1951. the genera of the Betylobraconinae Current projects: Revision of the Family Ichneumonidae. In C.F.W. and related groups. 2. Revision of Genus Hemistephanus Enderlein, Muesebeck, K.V. Krombein, & H.K. the genus Aleiodes Wesmael of the 1906 (Hymenoptera: Stephanidae). Townes, Hymenoptera of America West Palaearctic region, with M.R. . Request for help and coopera­ North of Mexico: Synoptic Catalog. Shaw). 3. Revision of the Australian .t.wn: I have been looking for the USD~ Agricultural Monograph 2, Betylobraconinae, with A. Austin. types of Hemistephanus peruanus Washmgton, D.C. 1420 pp. 4. Revision of the Indo-Australian and H. pehlkei (Enderlein, 1906) --· 1966. A catalogue and re­ and Afro tropical Macrocentrinae. 5. [Stephanidae], but after writing to classification of Neotropic Ichneu­ Revision of the genus Tropobracon. several museums in Europe, USA monidae. Memoirs of the American 6. Illustrated key to the subfamilies and Canada, I am still unable to lo­ Entomological Institute 8: 1-367. ofthe Braconidae. 7. Revision ofthe cate them. The species were de­ __. 1973. A catalogue and re­ Northwest European Opiinae and a scribed from just one specimen each, classification of Ethiopian Ichneu­ key to the genera of the Palaearctic and now I have evidence that they monidae. Memoirs of the American Opiinae, with J.W. van Zuijlen. 8. could be synonymous with previous Entomological Institute 19: 1-416. Several smaller papers on Opiinae described species of Cameron and ---· 1983. A description of and Alysiinae (Asobara, Alysia, Westwood. Any information re­ the Townes collection of Hymenop­ Aphaereta), Ecnomiinae etc.. garding the location of these types tera. Contributions of the American Museum notes: The staff of the will be highly appreciated. Entomological Institute 20: 15-30. National Natuurhistorisch Museum, Museum notes: The entomologi­ Wahl,D.B.1990.Areviewofthe Lei den is working hard to get accept­ cal collection at Purdue University mature larvae ofDiplazontinae, with able plans for the new museum has nearly 1,500,000 mounted in­ notes on larvae of Acaenitinae and building to be opened in 1997. Fi­ sects, most from USA, but also from Orthocentrinae and proposal of two nally the museum will have a large countries in South America and Af­ new subfamilies (Insecta: Hymenop­ permanent exposition. rica. There is no estimate ofthe num­ tera, Ichneumonidae). Journal of New literature: Achterberg C. ber of Ichneumonoidea, but most of Natural History 24: 27-52. van. 1993. Illustrated key to the sub­ the Hymenoptera are non-identified Yu, D.S. 1993. A proposed sys­ families of the Braconidae. Zoolo­ ichneumonids. This collection may t~m f?r stabilizing the names of spe­ gische Verhandelingen. 283:1-189. not be known by many professionals Cies, Illustrated with reference to the Achterberg C. van. 1993. Revi­ since Purdue's major emphasis is not Ichneumonidae (Hymenoptera). sion of the subfamily Macrocentri­ on systematic research. Bulletin ofZoological Nomenclature nae from the Palaearctic region. 50: 7-12. Zoologische Verhandelingen. 286: 1- Rafael Ala yo S. (lnstituto de 110. Ecologia y Sistematica; Carretera de Achterberg C. van and C. Varona Km 3 l/2; Capdevila, Boy­ O'Toole, 1993. Annotated Cata­ eros, Apartado Postal 801 0; 10800 logue of the types of Braconidae in Ciudad de Ia Habana, CUBA). the Oxford University Museum. Current projects: Systematics Zoologische Verhandelingen. 287:1- and ecology of Cuban Braconidae. 48. Collection sites: for 1993 Andy Austin (Department of (Sept./Oct.) field research was Crop Protection; Waite Campus; planned on Sumbawa and the Sula University of Adelaide; P.O. Glen Archipelago (Indonesia). Osmond, S.A 5064. AUSTRALIA). New literature· Achterberg, C. Aeschlimann (CSIRO; Bio­ van & A.D. Austin ( 1992). Revision control Unit; 335 Av. Parguel; 34090 of the genera of the subfamily Sigal­ Montpellier. FRANCE). phinae (Hymenoptera: Braconidae), Cheloninae including a revision of the Australian 16 species. Zoologische Verhandelin­ Current projects: Sergey hopes Tamaulipas, Mexico. We are going gen 280: 1-44. to finish his revision of the Pa­ to collect different pests too. I con­ Austin, A.D. & P.C. Dangerfield laearctic species of Spathius, for this tinue my collecting of Ichneu­ (1992). Synopsis ofAustralasian Mi­ reason he would like to borrow any monoidea and other Hymenoptera in crogastrinae (Hymenoptera:Bra­ material from the southern Pa­ different areas of Mexico." conidae), with a key to genera and laearctic including the south-eastern description of new taxa. Invertebrate and south-western parts. He contin­ Miroslav Capek (Boettin­ Taxonomy 6: 1-76. ues to work on a key to the braconid grova 38; 636 00 Brno 36; CZECH Austin, A.D. & R.A. Wharton species of the Russian Far East. Ser­ REPUBLIC). (1992). New records of subfamilies, gey reports an excellent and very im­ New Literature: apek, M. & van tribes and genera of Braconidae (In­ portant visit to Ottawa Canada and Achterberg, C., 1992: A revision of secta: Hymenoptera) from Australia, College Station USA. (He was here the genus Microtypus Ratzeburg with description of seven new spe­ for three weeks on a Canacoll grant (Hym.:Braconidae). Zool. Med. cies. Transactions of the Royal Soci­ and in Texas with a grant from funds 66:323-338. ety of South Australia 116: 41-63. available to Bob Wharton). Current projects: Braconids as Austin, A.D., R.A. Wharton & parasitoids of economic pests. P.C. Dangerfield (1993). Revision of Miloje Brajkovi (Institute of Request for help: Reprints of pa­ the endemic Australian subfamily Zoology, Faculty of Science, Uni­ pers on Braconidae wanted. Trachypetinae Schulz s.l. (including versity of Belgrade, 16, Akademski Cercobarconinae Tobias) trg, Belgrade 11000, Yugoslavia) James M. Carpenter (Hymenoptera:Braconidae). Sys­ Current projects: Study of the (Dept. of Entomology; American tematic Entomology 18: 97-119. Braconidae of Yugoslavia. Museum of Natural History; Central Current projects· 1. Biology of Request for help: Identification Park West at 79th St.; New York,NY Mesostoa kerri. 2. Revision ofBety­ of species of the genera Bracon, 10024, USA). lobraconinae. Apanteles and Opius. Dr. Brajkovic Current projects: Phylogeny of Other items of interest: Andy also would like to have literature on Aculeate families and various groups spent 3.5 months on study leave in braconids. therein, with collaborators too nu­ November 1992-January 1993, merous to mention "What do you working at collections around Aus­ Ahmet Beyarslan (Trakya want, at least it's the sister-group of tralia. He spent 2 months at ANIC Universitesi; Fen-Ed. Faliiltesi Biy­ the Ichneumonoidea --- maybe." (Australian National Insect Collec­ oloji BOliimti; 22030 Edirne, TUR­ Request for help or cooperation: tion) and 3 weeks doing field work KEY). "Now that you mention it, ethanol­ in Northern Territory. Results were Current projects: Taxonomic re­ preserved specimens of Vespidae s.l. mediocre due to the prolonged dry search on the Braconidae fauna ofthe would be useful, and I know all of season. Eastern Marmara (Turkey). you followers ofHenry Townes have been diligent ... " Nancy Beckage (University Santiago Bordera (Depar­ Museum notes: The American of California; Riverside, California tamento de Ciencias Ambientales y Museum of Natural History recently 92521-0314; Assistant Professor of Recursos Naturales Facultad de brought its collection back out of Entomology; Department of Ento­ Ciencias; Universidad de Alicante; storage, where it had been for several mology; 4435 Boyce Hall; USA). Ap. Correos 99; 03080 Alicante, years during compactor construc­ Current projects: Assessment of SPAIN). tions. the role of polyclivaviruses in caus­ Current projects: Study of the CoJJection sites: During a recent ing alterations in host physiology. fuana of Cryptinae (Hym., Ichneu­ (Sept. 1992) trip, Jim drove 3000km Isolation of venom components in monidae) from the Iberian Penin­ through pine and eucalyptus planta­ species that paralyse their host. Re­ sula. tions in Misiones, Argentina. quests for help or cooperation: Need assistance in collecting wasps: Amo­ Eurigie Ruiz Cancino Ronald D. Cave (Depto. de phila, eumenid species, ectoparasites (Uni versidad Autonoma de Proteccion Vegetal; Escuela with paralysing venoms. (She will Tamaulipas; Fac. de Agronomia; Agricola Panamericana; Apdo. 93, pay for collecting activities)! Centro de Inv. y Desarrollo El Zamorano, Tequciqalpa, HON­ Agropecuario Forestal y de Ia Fauna, DURAS). Sergey Belokobylskij MEXICO). Current projects: Massing-rear­ (Zoological Institute, Dept. of Ento­ Current projects: "I am now in ing and release for establishment of mology, Russian Academy of Sci­ the beginning of a research project Cotesia plutellae and Diadromus ences, 199164 St. Petersburg, B-164, about Hymenoptera and other natu­ collaris. Inventories of natural ene­ RUSSIA) ral enemies on pinyon pines in mies of key pests and general inven- Ichnews 14, April 1994

tory of parasitic Hymenoptera of Request for help: Requests co­ Requests for help or cooperation: Honduras. operation on taxonomic revisions of anybody interested in seeing and Request for help or cooperation: chinese Braconidae and adaptation identifying Hymenoptera from Indo­ Ronald has requested Malaise traps of indigenous parasitoids to exotic nesia. and in return he will supply speci­ immigrants. Requests reprints of pa­ Collection sites: Sumatra, Bor­ mens of interest collected in the pers on Braconidae especially neo. traps. Aphidiinae. Museum notes: The EAP collec­ Clement E. Dasch (160 tion currently holds approximately Chen, Xuexin (Institute of Montgomery Blvd.; New Concord, 3000 specimens of Braconidae (52 Applied Entomology; Zhejiang Ag­ Ohio 43762, USA). genera, 23 species identified) and ricultural University; Hangzhou New Literature: Nearctic Mi­ 2000 specimens of Ichneumonidae 310029; CHINA). croleptinae, He1ictinae, Cyllocerii­ (104 genera and 28 species identi­ New Literature: The Chinese nae, & Oxytorinae. Mem. Amer. Ent. fied). Homolobus species (Chen, 1991; lost. 52: 1-4 70, 1992. Collection sites: Newly pre­ Chen, He & Ma, 1991); the Chinese Current projects: Working on my served areas (national parks, biologi­ Aleiodes species with pale antenna! private collection, especially Ichneu­ cal reserves) in Honduras want to segments (Chen & He, 1991); the moninae. know more about their fauna; Ronald Chinese Acampsis species (Chen & can arrange collecting trips to these He, 1992); new records of Aleiodes Luis De Santis (Facultad de areas for anyone wanting to sample species from China, i & ii ( Chen & Ciencias Naturales y Museo; Paseo the entomodiversity. He, 1992); the Chinese Fomicia spe­ del Bosque; 1900 La Plata, REPffi­ cies (Chen, He & Ma, 1993); a new LICA ARGENTINA). Bjorn Cederberg (Mora species of Agriotypus from Hubbi, Current projects. Dr. De Santis Folkhogskola; Box 423 S-79227 China (He & Chen, 1991 ); the Chi­ continues to work on the Braconidae Mora, SWEDEN). nese Sigalphus species (He & Chen, and Ichneumonidae of Argentina. Current project: Revision of 1993). North European Ophioninae. Current projects: Revision of the Hugo Delfin (Universidad subfamilies Sigalphinae and Meteo­ Autonoma de Yucatan, Facultad de Chao, Hsiu-fu (Biological ridiinae ( 1992-1993 ), Rogadinae and Medicina Veterinaria y Zootecnia, Control Research Institute; Fujian Macrocentrinae (1992-1994) from Merida, Yucatan, MEXICO) Agricultural College: Fuzhou, Fu­ China (with He Junhua). Current projects: Hugo has re­ Request jian 350002; P.R. CHINA). for help or cooperation: cently started a Ph.D. on the sys­ Current projects: Chinese Agri­ Literature on the taxonomy and biol­ tematics of the Braconidae. His area ogy of Braconidae, otypidae, with descriptions of a new especially on the ofresearch is still not exactly defined subfamilies genus and three new species (Ichneu­ indicated above. but he is thinking of working on monidae) (in press). Streblocera Collection sites: Chebaling and some aspect of the faunistics of the from Fujian province. On the late Heishiding natural reserves in Braconidae of the Yucatan. Dr. H. Townes' collection of Xan­ Guangdong Province (vi.1992); Gu­ Request for help: Hugo would thopimpla Saussure from Indo-Ma­ tian Shan natural reserve in Zhejiang like to have literature on Neotropical laysia. Studies on Chinese Spathiini Province (vii.1992). Braconidae. and Xanthopimpla (lchneumoni­ dae). Paul C. Dangerfield (Uni­ Erich Diller (Zoologische Museum notes: We have a col­ versity of Adelaide; Waite Campus; Staatssammlung; Munchhausenstr. lection of about 260.000 specimens Waite Rd.; Glen Osmond, South 21; D-8000 Munchen 60; Tel-Nr. of parasitic Hymenoptera in our Bio­ Australia 5064. AUSTRALIA). (089) 8107-0). logical Control Research Institute, Current projects: Taxonomy and Current projects: Systematics of Fujian Agricultural College. phylogeny of the Indo-Australian & Phaeogenini (Ichneumoninae). Re­ Pacific species of Cardiochiles. This vision of Stenodontina (lchneumon­ Chen, Jia Hua (Beneficial project is likely to be expanded to a inae, Phaeogenini). Insects Lab.; Dept. of Plant Protec­ generic revision of the subfamily tion: Fujian Agricultural University; Cardiochilinae later in 1993. Fernando Fernandez C. Fuzhou. Fujian P.R. 350002, (Apartado Aereo, 77038: Santa Fe CHINA). D. Christopher Darling De Bogota 2 D.C.. COLUMBIA). Current projects: Prof. Chen con­ (Entomology - Royal Ontario Mu­ Current projects: Curation of tinues research on taxonomy of chi­ seum: 100 Queen's Park. Toronto, Hymenoptera at the Museo De His­ nese Aphidiinac and other Ontario. CANADA). toria Natural. Universidad Nacional Braconidae as well as biocontrol of Current projects: mouth parts of De Colombia. Santa Fe De Bogota. aphids. braconid subfamilies. 18

Parasitica and Aculeata survey of Current projects: Emendation to son), Hymenoptera of Costa Rica, Hamacatacu National Park (Ama­ the revision of the Opiinae ofthe Old with chapters by Scott Shaw, Jim zonas), La Mac arena Reserve (Meta) World (Braconidae). Opiinae-fauna Carpenter, Dave Smith, Lynn Kim­ and Zambrano Region (Magdalena). of Finland (together with M. Kopo­ sey, Denis Brothers, the innumerable Requests for help or cooperation: nen). Opiinae-fauna of Turkey (per­ New World chalcidologists, Mas­ Identification, at least to genus, of haps together with A. Beyarslan). simo Olmi, Mary-Jane West-Eber­ ichneumonid, braconid, and other Revisionary studies on Old World hard and others. parasitica of the museum and my col­ Alysiinae. Revisionary studies on lection trips. Calyptus and related groups by Peter Milka Glavendekic (For­ Collection sites: Colombia: Stadlhofer. estry Faculty ofBelgrade University; Magdelana: Zambrano - dry forest Requests for help or cooperation: Kneza Viseslava 1; YU-11030 Beo­ on Magdelena rivers. Colombia: Dr. Fischer would like to obtain in­ grad, YUGOSLAVIA). Meta: La Macarena - gallery forest, formation on collections of Old Current projects: Fauna ofYugo­ savannas and agro-ecosystems in World Alysiinae (identified or not). slavian Ichneumonidae. Fauna of north eastern Macarena Reserve. Presumably he would like to borrow Durmitor Mountain. Amazonas: Pan Hamacayaru - rain them for his revisionary studies. Museum notes: I visited Zoolo­ forest near Amazonas River. Colom­ Dr. Fischer would also like to gischen Staatssammlung Munchen bia: Choco: Riosurio - very humid know if anyone has any ideas (pub­ in Feb.-March 1993. rain forest near Atrato River. lished or not) for the function and Collection sites: Montenegro: evolution of the cyclostome condi­ Durmitor, Adriatic Sea. Serbia: Na­ Dominga Carolina Berta tion found in many Braconidae. tional Park "Djerdap", Kopaonik De Fernandez (Departamento Also the function of the carapace Mountain, Raska. Zoologia, Entomologia; Fundacton found in many and diverse groups of Miguel Lillo; Miguel Lillo 251; Braconidae. Vladimir E. Gokhman 4000 San Miguel de Tucuman; AR- Museum notes: Dr. Fischer has (Plant Protection Division; Botani­ indicated that his museum may have GENTINA). cal Garden; Moscow State Univer­ limited funds to purchase collections Current projects: Ms. Berta has sity; Moscow 119899, RUSSIA). of Braconidae. Please let him know finished her doctoral thesis on Neot­ New literature: Gokhman, V.E. if you have anything to offer. ropical Cremnops and Zacremnops. 1992. On the origin ofendoparastism in the subfamily Ichneumoninae Eduardo Portuondo Fer­ Mike Fitton (Dept. of Ento­ (Hymenoptera, Ichneumonidae). mology; The Natural History Mu­ Zhurnal Obshchei Biologii 53: 600- rer (Centro De Ciencias Naturales; seum; London, SW7 5BD; 608. Ave. Manduley 308 esq. 13; Vista ENGLAND). Current projects: Investigation of Alegre. Stgo. de cuba, CUBA cp Current projects: Phylogeny of the phylogenetic affinities ofthe sub­ 90400). Ichneumonid subfamilies (with family Inchneumoninae (together Current projects: Systematics Donald Quicke). Hymenopterous with M.G. Fitton and D.L.J. Quicke). and Ecology of Cuban Ichneumoni­ parasitoids of British/European but­ Study of ichneumonids of the tribe dae, especially Ichneumoninae. terflies (with Mark Shaw). Phaeogenini from the Russian Far Museum notes: My department East. Karyological study of Russian bas interesting material of the fol­ Ulf Gardenfors (Dept. of Ichneumoninae. lowing taxa: Chalcidoidea, Ichneu­ Wildlife Ecology; Box 7002, S-750 Request for help or cooperation: monoidea, Proctotrupoidea and 07 Uppsala, SWEDEN). Specimens of Phaeogenini from the Cynipoidea. collected in mountains Current projects: "I now have a Palaearctic (especially from the Far of eastern Cuba. Anyone interested position at the Threatened Species East) for loan/exchange. Any living in studying the material is welcome. Unit (working with Red-Listed in­ Ichneumoninae (preimaginal Request: I am interested in ob­ vertebrates), but I still try to work a stages/adult females). taining literature on Neotropical Ich­ bit with the aphidiines." Museum notes: April 1993 - neumonidae, especially study for a few days of the collection Icbneumoninae. of Phaeogenini and some other ich­ Other items of interest: Sys­ Ian Gauld (The Natural His­ neumonids at the British Museum of tematics and ecology of Braconinae tory Museum: Cromwell RA: Lon­ Natural History. and Rogadinae. don SW 7 5BD: U.K.). Current projects: Ichneumonidae Collection site: Summer 1992 - of Costa Rica: subfamilies collected Ichneumoninae in the Mos­ Maximilian Fischer (Di­ Anomaloninae, Typhoninae, cow and Volgograd regions. rektor Ocr 2 Zoolog.ABT. AM; Ctcnopclmatinae, Lycorninae, Naturhistorischen Museum: A-1 014 Phrudinac and Pimplinae (addi­ Wicn. Burgring 7, AUSTRIA). tions). Book (with co-editor P. Han- Ichnews 14, April 1994

V.K. Gupta (Entomology literature for me - being not em­ New Literature: Klunker, R. & Dept.; University ofFlorida; Gaines­ ployed in a museum or an institute Fabritius, K. (1992): Bibliographie ville, Florida 32611-0620, USA). with a large library". (Editor's note: uber Pupariumparasitoide synan­ Current projects: Taxonomy of If enough readers have the same throper Fliegen.-Beitr. Entomol. Ichneumonidae of Florida - Hand­ opinion we will reintroduce a recent Berlin 42(2):331-429. books. Taxonomy of Oriental Ich­ literature section. My feeling is that neumonidae. Revisions of genera. most readers don't need the service. J ank Kolarov (Department Database of ichneumonid literature. Please let us know bow you feel ofZoology; Biological Faculty; Uni­ about this when you return the 1994 versity of Sofia; 8 Dragan Zankov He, Junhua (Institute of Ap­ questionnaire.) Boul., 1421 Sofia, BULGARIA). plied Entomology; Zhejiang Agri­ Current prQjects: Conclusion of New Literature: Kolarov, J., cultural University; Hangzhou the revisions of Western Palaearctic 1992. A catalogue ofIchneumonidae 310029, CHINA). Tersilochinae. Revisions of several from Albania Entomofauna 13(16): New Literature: A supplemen­ genera of Western Palaearctic Cam­ 261-268. Kolarov, J., 1992. On the tary catalogue of Ichneumonidae popleginae and Cryptinae-Phy­ Hibernation and spreading of some from China (HE, 1992); the Chinese gadeuontini. Search for types and ichneumonids. Acta Zool. Bulg, Sigalphus species (HE & CHEN, type revisions ofWestern Palaearctic 45,104-106. 1993); the Chinese Homolobus spe­ Ichneumonidae. Current projects: Catalogues of cies (CHEN, HE & MA, 1991); the Museum notes: The types of lch­ Balkan countries, Turkey and Cy­ Chinese Acampsis species (CHEN & neumonidae described by Pic and prus (lchneumonidae ), Taxonomy of HE, 1992); the Chinese Fomicia Berthoumiae present in the Paris Cremastinae of these regions. species (CHEN, HE & MA, 1993). Museum are now labelled and listed. Requests for help or cooperation: Current projects: Revision of the This done, more than 95% of types Materials and literature from the re­ Chinese members of the subfamilies of Westt!rn Palaearctic Ichneumoni­ gion. Sigalphinae, Meteoridiinae ( 1992- dae present in the Museum National CoJJection sites: Dr. Kolarov has 1993), Rogadinae and Macrocentri­ d' His to ire N aturelle Paris are now been collecting in the Balkan coun­ nae (1992-1994) (with Chen recognized and labelled, and most of tries and Cyprus. Xuexin). them are placed into separate boxes. Request for help or cooperation: Dolly M. Lanfranco Literature on the taxonomy and biol­ A. B. Jakimavicius (Insti­ Leverton (Institute de Silvicul­ ogy of Braconidae, especially on the tute of Ecology; Akademijos 2; Vil­ tura; Universidad Austral de Chile; subfamilies indicated above. nius 2600, LITHUANIA). Casilla 567; Valdivia, CHILE). Collection sites: Chebaling and Current projects: Generalised Current projects: Biological Heishiding Natural Reserve in research on the Braconidae of Lithu­ control by parasitoids in Rhyacionia Guangdong Province (vi.l992); Gu­ ania. buoliana (pine shoot moth) and Sirex tian Shan Natural Reserve in Zhe­ nortilio (woodwasp) a potential pest jiang Province (vi.l992). Jim Johnson (Dept. of Ento­ for Chile. mology; Michigan State University; Erasmus Haeselbarth 243 Nat. Sci. Bid 8; East Lansing, MI John Luhman (Minn DepL (Zoologische Staatssammlung, 48824-l I 15, USA). of Ag.-PPD; 90 W. Plato Blvd.; St. Milnchbausenstr. 21, d-81247 Current projects: Revision of the Paul, MN, USA 551 07). Milnchen, GERMANY). species of American Aphidius north Current projects: Revision of Current projects: Dr. haeselbarth of Mexico is ongoing. Population subtribe Stilpnina (Phygadeuonti­ is retired but cominues his studies on genetics and morphometric analysis nae, Ichneumonidae); Identifying European, /chneutes and some of native and exotic aphid parasitoids undetermined Ichneumonids in Uni­ Eupborinae (Perilitus, Microctonus, - This project is investigating parasi­ versity of Minnesota collection. and Syntretus and allied genera) toids of the Russian wheat aphid D. Other items of interest: "Ichneu­ noxia, both from a biochemical and moninae workers! I have a complete Klaus Horstmann morphological basis. Primary groups list with data of all the Enderlein (Lebrstubl Zoologie Ill; Biozen­ of interest are native and exotic types and Heinrich types in the col­ trum; Am Hub land; W -8700 Wur­ Diaeretiella rapae and Aphidius spp. lection of the Institute of Zoology, zburg, GERMANY). Polish Academy of Sciences New Literature: "I regret that you Ronald Klunker (Bundes­ (=PAN), Warsaw. The former is in do not further include lists of litera­ gesundheitsamt Laboratorium f.ge­ press the latter, in preparation. ture in the ICHNEWS. I understand s undhe itsscbadl. Arthropoden; the reasons for not doing it. but these Thielallee 88-92. PF 33 00 13; D- lists were an important source for 1000 Berlin 33. GERMANY). 20

Paul M. Marsh ( P.O. Box Queiroz" da Universidade de Sao Work on the increasingly large genus 384 North Newton, Kansas 67117; Paulo, Brazil" studying braconid Yelicones is progressing though USA). parasitoids of fruit flies in Brazil. slower than hoped, as are revisions Current projects: Continuing Requests help or cooperation: I of Indo-Australian Physaraia, Tri­ study of the North American species need literature on the biology of Do­ gastrotheca and Cosmophorus (with of the genus Aleiodes with Scott ryctobracon, Opius and other gen­ C. van Achterberg). Other members Shaw. Probably about 80 species in­ era. of the group have also been busy. cluding new species. Professor You Lanshao from Hunan, Other items of interest: On Sep­ Angelica Maria Pen­ China, bas been visiting for six tember 30, 1993 I retired from the teado-Dias (Universidade Federal months trying to get to grips with the Systematic Entomology Laboratory de Sao Carlos; Centro de Ciencias genera of Braconinae and also de­ after nearly 30 years of enjoyable Biologicas e da Saude; Depar­ scribing a few new Chinese species. service. I have moved to Kansas tamento de Ecologia e Biologia Evo­ Jamil Chishti is nearing completion where I plan to continue work on lutiva; Via Washington Luiz, km 235 of his Ph.D. and the revision oflndo­ unfinished projects, albeit at a more - Caixa Postal676; CEP 13565-905, Australian Stenobracon. Whilst leisurely pace. (Editor's note: Paul Sao Carlos, SP, BRAZIL). new Ph.D. students, Md Ismail Miah has also stepped down as a co-editor Current project: Dr. Penteado­ and Azidah Abdul Aziz are coming of lchnews.) Dias is working on the biology and to terms with various ichneumonids. taxonomy of Braconidae from Bra­ Kaoru Maeto (Forestry and zil. My Ph.D. student from Turkey, Forest products research Institute, Request for help and coopera­ Hasan Basibuyuk who is working in Hokkaido Research Centre, Hitsuji­ .ti.Qn: I would like to obtain literature the Entomology Department of the gaoka 7, Toyohira, Sapporo, JAPAN on Neotropical Braconidae. I hope Natural History Museum, London, is 062. to help in the production of the key now trying to integrate his studies of Current projects: 1. Intergeneric to the genera of Braconidae of the antenna! and antenna cleaner mor­ variations in the male genitalia of the New World. phology with grooming behaviour Microgastrinae. 2. Japanese species Collection sites: (1992) collect­ and he would be particularly inter­ of Microgastrinae. 3. Braconid ing trips were in Brazil areas of "cer­ ested in hearing from anyone who fauna of the Pasoh Forest Reserve, rado" and forest. might be able to help him obtain Peninsular Malaysia. living material of any of the follow­ ing hymenopterans: Pamphiliidae, Donald Quicke (Dept. of Bi­ Siricidae, Stephanidae, Orussidae, Bob Matthews (413 Biologi­ ology, Imperial College at Silwood Trigonalyidae, Aulacidae, Pelecini­ cal Sciences Building, Dept. of En­ Park, Ascot, Berks SL5 7PY, ENG­ dae, lbaliidae and Vanhorniidae. He tomology, University of Georgia; LAND. Email address can Athens, Georgia, 30602 USA). d.quicke@ ic.ac.uk) be contacted by phone (071 938 9148) or e-mail c/o Requests for help or cooperation: Current Projects: Revision of [email protected]. Postage and I would like to obtain any live Melit­ world species of Yelicones. (North packing costs would be reimbursed tobia (Eulopbidae) taken from bum­ American species are almost com­ of course. ble bee nests. pleted Oriental species are next.) The Ichneumonoid Research Mike Fitton, David Notton and I are Group (IRG) an informal group in Sergio M. Ovruski (Centro now nearing completion of a mega­ the NHM's Department of Entomol­ De Investigaciones Para La Regu­ analysis of icbneumonid relation­ ogy, within the framework of the lacion De Poblaciones De Organis­ ships including lots of new internal museum's science programme/re­ moso Nocivos; C.C. 90-4000 San characters search group system. The group Miguel de Tucuman, ARGEN­ "What's New at Silwood? Fol­ comprises 14 people co-led by Mike TINA). lowing my move to Silwood Park Fitton, Donald Quicke and Ian Gauld Current projects: "I am working (part of Imperial College) under the and includes four graduate students. on my Doctoral Thesis on the biol­ NERC's Initiative in Taxonomy last Research projects are centred around ogy, taxonomy and economic impor­ October. there have been a number hymenopteran and especially icb­ tance of parasitoid Hymenoptera of new developments on the research neumonoid taxonomy. Most of the attacking fruit tlies in Argentina. I front. Of course, ovipositor. sperm, members work at the Natural History am especially concentrating on the venom gland and alpha-taxonomic Museum, but a few are based at Sit­ biology of Doryctobracon brasilien­ work still feature high but work is wood Park, the out-of-town half of sis and D. tucumanus. In January also now under way on a number of the Imperial College Biology De­ 1993. I spent one week in the Depart­ molecular projects including the re­ partment. The research group meets ment of Entomology of "Escola Su­ lationships between icbneumonoid approximately every month for re­ perior de Agricultura" "Luiz de subfamilies and between the genera search talks or seminars nn such top- of various hraconid subfamilies. Ichnews 14, April 1994

ics as wing venation, cladistics and Azhar Saeed (Department of Costa Rica. As a member of the preparing illustrations for publica­ Crop Protection; Waite Campus, Parasitic Hymenoptera Advanced tion." University of Adelaide; Glen Os­ Research Team, I have a number of Other items of interest: ARTY mond. SA-5064, AUSTRALIA). responsibilities. I will be writing a WASPS AND WASPERS? "As a bit Current prQjects: I am a post­ key to the subfamilies of the Bra­ of fun I'm trying to compile a list of graduate student from Pakistan, conidae of the New World - keys to works of art in museums that include sponsored by the Rotary Foundation the genera of the Agathidinae, Hel­ parasitic wasps. I know there are not and I am working with Andy Austin coninae s.l. and several smaller sub­ many but my experience of walking towards a Ph.D. My research project families. As well I am responsible around galleries has shown that a is on the systematics and phylogeny for creating an interactive key for all surprising number of Old Master of the genus Diolcogaster Ashmead of the genera of Braconidae of the still-lifes have a parasitoid posed on and will include a revision of the New World. I have completed the some fruit or suchlike. The postures Australian species. genera of the Ichneutinae in this fash­ of the beasties illustrated are also Request for help or cooperation: ion, and anyone wishing a copy need informative, some were almost cer­ Although I have borrowed most Di­ only send me a floppy disc (any size). tainly seen alive but others have that olcogaster specimens from the major Presently I am using the software characteristic window ledge look!" world collections, I will be very in­ DELTA but I am also looking at terested in receiving any more mate­ some other programs before I com­ Alexandr P. Rasnitsyn rial collected from the mit myself. If anyone has any sug­ (The Arthropoda Laboratory, Pa­ Indo-Australian region, particularly gestions please let me know. leontological Institute, Russian if it has associated host data. Request: I would like to borrow Academy of Sciences, Prof­ any Agathidinae from the east Pa­ soyuznaya Str. 123, Moscow Martin Schwarz (Univer­ laearctic especially Korea and main­ 117647, RUSSIA) sitat Salzburg, Zoologisches Institut, land China. Current Projects: Dr. Rasnitsyn Hellbrunnerstrasse 34, A-5020 reports that due to the dire economic Saltzburg, AUSTRIA) Mark R. Shaw (Royal Mu­ conditions in Russia and especially Current Projects: Dr. Schwarz is seum of Scotland; Chambers Street; in his laboratory he has no progress revising the wingless species of Edinburgh EH9 2LU; Scotland, in his science to report to the reader­ Gelis of the Western Palaearctic re­ UK). ship of Ichnews. gion as well as various genera of Current projects: Interests: Biol­ Other items of interest: Dr. Ras­ Cryptini of the Western Palaearctic. ogy and taxonomy of Ichneumoni­ nitsyn had a few comments on the dae especially those parasitising article by David Wahl and myself in 1\'lichael J. Sharkey (BRD, Lepidoptera and of Spiders. Host the first volume of the Journal of CLBRR, CEF, Agriculture Canada, range/community structure of Hymenoptera Research. Ottawa, Ontario, K 1 A OC6 CAN­ parasitoids. Currently working on Sharkey and Wahl, Pg. 19: "We ADA. Email address: European Aleiodes with Kees van are unaware of any Apocrita with sharkeym@ ncccot2.agr.ca) Achterberg. two r-m crossveins, with the excep­ Recent Literature: Sharkey, M.J. Request for help or cooperation: tion of some braconids." Lep­ ( 1992) Cladistics and tribal classifi­ I would like to see reared W. Pa­ tephialtites gigas A. Rasnitsyn cation of the Agathidinae. The Jour­ laearctic Aleiodes s.lat. (Ephialtitidae, cf. Rasnitsyn 1975, nal of Natural History 26:425-447. hymenoptera Apocrita of Mesozoic: Sharkey, M.J. (1993) Braconidae. Scott R. Shaw (University of Fig.19) had two r-m crossveins (I pp.362-395 in Hymenoptera of the Wyoming; Department of Plant, believe due to a reversion and not as World, Goulet, H. and J.T. Huber Soil, and Insect Sciences; College of a symplesiomorphy with sym­ eds.. Agriculture Canada Publication Agriculture; P.O. Box 3354, phytans). 1894/E pp. 688. Laramie, WY. USA). Sharkey and Wahl, Pg. 20: "Ma­ Current Projects: A revision of Current projects: Book chapter son ( 198 l) was the tirst to examine the world genera of Ichneutinae, in on Costa Rican braconids is due for Hybrizon· s relationship to Ichneu­ press in The Journal of Natural His­ publication soon. Revision of North monidae and braconidae from a phy­ ~. (with R. Wharton). Revision of American species of Aleiodes (with logenetic perspective." Rasnitsyn the Japanese species of Agathidinae Paul Marsh). Phylogeny of ( 19XO:pg81) did this a year before (I'm almost tinished. honestly). Re­ Aleiodes. Chapters on Rogadini. with the same result. vision of the Agathidinae of the Rus­ Neoneurinae. Meteorinae, Euphori­ (Euitor's note: We welcome cor­ sian Far East. Revision of the nae. and Cheloninae for the Identifi­ rections. comments. and reviews of Agathidinae of Korea. Revision of cation Manual to Braconid Genera of puhl ishcd articles.) the genera of Cremnoptini and Diso­ the New World. A new genus of phrini (Agathidinae). Description of Euphorinae (sister-group to Cryp­ a new species of Sigalplws from toxilos). Behaviour of Nconeurus. 22

Other items of interest: As of the als about aphids, their parasitoids, Sharkey, Scott Shaw, Bob Wharton fall of 1993 Scott has had two new hyperparasitoids.' and Jim Whitfield. Together, under graduate students: "Joseph Fortier is the direction of Bob and Paul Marsh, working on the cladistics ofAleiodes Petr Star (Institute of Ento­ the team is preparing an illustrated funded by a three year grant from the mology; Czech Academy of Sci­ key to the genera of New World Bra­ National Science Foundation. Nina ences; Branisovska 31; 370 05 eske conidae. This will be in a similar Zitani is currently working on a new Budjovice; CZECH REPUBLIC). format to the key produced by species ofM eteorus from Costa Rica Current projects: 1992-93: FAO Marsh, Shaw and Wharton which (that apparently has a defense against consultancy - Diuraphis noxial dealt with the braconid fauna of leaf-cutter ants). Scott's under­ Mordv./in Chile. Biocontrol. Parasi­ America north of Mexico except that graduate student Jeri Wright recently toids of D. noxia - world. Neotropi­ there will be a subfamily key in­ won a honourable mention at the cal Aphidiidae, esp. Chile. cluded and more in the way of text to ESA meeting in Indianapolis for her Aphidiidae associated with Notho­ outline the diversity and biology of work on "anomalous diversity" and fagus-aphids. the Braconidae. According to com­ Costa Rican Braconidae. More im­ mitments made to NSF the project is portantly, Jeri was recently awarded David Wahl (American Ento­ to be completed in August of 1996 a Rhodes Scholarship and will attend mological Institute; 3005 SW 56th and one can reasonably expect a pub­ Oxford University next fall. She is Ave.; Gainesville, FL 32608-5047; lished product a year thereafter. As interested in tri-trophic interactions USA). part of the NSF funding Dr. Wharton of plants/leafminers/and parasitoids Current Projects: Labenine and is committed to presenting several (especially Mirax). mesochorine generic revisions have courses on the identification of the Requests for help and coopera­ been published or are in press, vari­ Braconidae. In August he travelled .ti.Qn: We are already swamped in ous small projects are under way, and to a research station near Chamela Aleiodes specimens from various I am nearing the end (hopefully) of which is several miles from the Pa­ collections, but we are always inter­ the generic revision of Alomyini cific Ocean in the state of Jalisco, ested in seeing more material if it is (Phaeogenini of authors). Mexico (along with yours truly, reared (send the mummies). Willing Museum Notes: Most of my time MJS) to give a course to a group of to exchange identified New World this year has been spent on curational Mexican students and professionals. Aleiodes for identified Aleiodes duties at the AEI. The collection has The research Station is surrounded from other regions. finally broken the one million mark. by seasonally dry deciduous forest Collection sites: Travelled to There are approximately 589,000 and is a great place to collect. There Brazil (August 1992) and Costa Rica ichneumon ids and 161,000 bra­ are inexpensive and high quality ac­ (March 1993). con ids (other holdings include commodations and meals to be had 18,000 "Symphyta", 19,000 misc. right at the reserve. For information Rajendra Singh (Aphid­ small parasitoid families, 77,000 the interested reader can write to Fe­ Biocontrol Laboratory; Department "microhymenoptera", and 184,000 lipe A. Noguera, Estaci6n de Bi­ of Zoology; University of Aculeata). The ichneumonids have ologia, UNAM, Apartado Postal 21, Gorakhpur; Gorakhpur-273009; IN­ been completely recurated and a ma­ San Patricio, Jalisco 48980, Mexico. DIA). jor effort is underway to update the Besides his responsibilities with Current projects: "Faunistic sur­ AEI literature resources. It should PHART, Bob is continuing some vey of aphids and their aphidiine be mentioned that the ichneumonid studies on Australian Braconidae parasitoids in terai belt of N.E. Uttar habitus drawings used in the 4 vol­ and remains busy with his non-bra­ Pradesh to explore the possible bio­ umes of Genera of Ichneumonidae conid research on endangered birds logical control agent against the have been sorted out and filed away, and fruit tlies. aphids of economic importance. and are available for use (with certain Factors affecting life-table ofand sex restrictions). Zettel, Herbert (Naturhis­ allocation by an aphid parasitoid torischen Museum; A-1014 Wien, Lysiphlebus delhiensis (Subba Roa Robert (Bob) E. Wharton Burgring 7, Austria). & Sharma). (Editor's note: In some (Department of Entomology. Texas Recent Literature: 1992. Revi­ of the responses that I received I A&M University, College Station, sion der Phanerotoma-Arten Norda­ came across the name Aphidiidae; I Texas 77843, USA) merikas (Hym., Braconidae. have exercised my editorial powers Current Projects: Bob received a Cheloninae).- Linzer bioi. Beitr. and changed these to Aphidiinae) NSF grant to fund the efforts of the 24/l :275-330. Special interests: Biology of Parasitic Hymenoptera Advanced aphid parasitoids and their intertro­ Research Team, which is composed phic relationships. of 7 principle investigators: Kces Cooperation: Dr. Singh would van Achterherg. Jim Johnson. Paul like to. 'exchange ideas and materi- Marsh. Donald Quickc. Mike Ichnews 14, April 1994

are available in addition to singles. HYIVIENOPTERA WORK­ The cost will be £60.00 inclusive of SHOP"V" bed and breakfast on 1Oth and 11th and packed lunches for Saturday and The Canacoll Foundation in col­ Sunday. Participants requiring ac­ laboration with the Centre for Land commodation for shorter periods and Biological Resources Research will pay proportionately less. Sil­ will present for the fifth time a course wood Park is easily reached by train aimed towards identification of from London and the journey takes Hymenoptera to family and subfam­ approximately 50 mins. Further de­ ily level. This will take place June tails from: Donald Quicke, Dept. of 22-23, 1994 at the K.W. Neatby Biology. Imperial College at Sil­ Building, C~ntral Experimental wood Park. Ascot. Berks SL5 7PY, Farm, Ottawa, Canada. The course England. Email address fee is $400.00 (Cdn.) and limited to [email protected] 14 participants. Further infonnation from: Mike Sarazin, CLBRR. K.W. Neatby Bldg. CEF, Ottawa, ON. KIA OC6. FAX: (613) 995-1823. TEL: (613) 996-1665.

Cheloninae

ANOUNCEl\'IENTS

INSECT PARASITOIDS SIG FIELD WEEKEND

Following the success of last year's lst Insect P:uasitoids SIG tield weekend. it has been decided to organise a lnd for a weekend in early The location will be June this year. Euphorinae the same: the 24.5 J.cres of diverse habitats which surrounds the Impe­ rial College Department of Biology at Silwootl Park nt!:lr .-\scot in South­ TAXONOL\'IY AND ern England. L~t year. the meeting BIOLOGY OF PARA­ was attended by some 20 RES mem­ SITIC HYl\'IENOPTERA bers and several interesting discover­ ies were made including the discovery of living Histeromerus A short residential course run Ento­ Homolobinae which has even led to J. publication. jointly by the Department of In addition. novices in the tield (both mology. Tht: Natural History Mu­ senses) were given some informal seum, London and the Department of tuition in how best to nt!t tor other­ Biology. Imperial College. Univer­ wise) their prey. This yt!ar it has been sity of London. The cost will be £550 all per rarti~ipant. with dates Je~.:iJeJ to cxtenJ the invit~uion to ****** I 1994, members ,lf the Imernatinnal So~.:iety set for 24 .-\pril w M~1y at nf Hymen,lpterists in the hope nf fur­ Imperial College. Silwood P:uk. Ichnews was assembled using further details pro­ ther in~.:reasing ~ommuni~.:ation be­ London EnglanJ. For \VPerfcct. Corel. and Ventura tween people with similar inlerests. please ~.:onla~t: Dr. Mike Fitton, grams. with the new found cxpcrtise The Natur~tl Eri~ :\~~omnwdation will ~e tn dean and Dept nf Entonwlogy. of Rickey. RnaJ. lilly (though lhll plush) student History Museum. Cromwell S\V7 5B D. EnglanJ. 1"01111\S Ill\ SI!C. s,,ltl~' ,l,,ublc 1'001\\S Lllndun. 24 TAXA

1) TAXA, a biosystematic data management system, and a Ichneumonidae data set.

TAXA is a software system that manages records of taxonomic names. It operates on any ffiM compatible per­ sonal computer with a hard disk memory. All published taxonomic and biological information can be entered into the system, for example, the author's name, the year of original description, current and past combinations, current and past junior synonyms, description, distribution, revision, catalogues, biology, hosts etc. Each entry includes a source reference. The taxa are arranged in a taxonomic hierarchical tree with kingdom as the highest possible level and subspecies and synonyms as the lowest level.

TAXA can create various lists of species depending on the conditions set for the search procedure, e.g., a list of taxa parasitizing a certain host, a list of taxa occurring in Canada or a list of types in a museum. TAXA can also gen­ erate various statistics, e.g. the regional distribution or the average body size of any taxonomic group. The Ichneu­ monidae data set includes 35,173 names, 485,883 records of referenced information and 921 digitized line drawings taken from 4,351 references covering the period from 1900 to 1990. The program and the data set occupy 19 mega­ bytes of memory.

2) REFIND, a references management system.

REFIND is an ffiM compatible software system that manages references for fast retrieval. Each reference is composed of an identification number, the year of publication, the name or names of the author, the title of the refer­ ence, the source from which the reference was found and comments. REFIND is capable of creating bibliography and importing and exporting references.

ORDER FORM

Send order to: From:

Dicky S. Yu Name: P.O. Box 273 Address: Lethbridge Alberta TlJ 3Y7 CANADA

Please send me:

- TAXA and the lchneumonidae data set. (Fourteen 3 112 " HD disks) (TAXA)US$300.00/Can$375.00 (REFIND)US$30.00/Can$37.50

Note: Canadian order please add 7% G.S.T. No postage will be added to prepaid order. TAXA: A biosystematic data management system: Insecta Hymenoptera Ichneumonidae, 1900-1990 and REFIND: a references management system. Dicky S. Yu ($330 [USA] for both; available from author: P.O. Box 273; Lethbridge, Alberta; Canada TlJ 3Y7)