Marek Zabkar and Simon D Pollard2 A check-list of Salticidae (Arachnida: Araneae) of ,

Abstract Acheck-list ofjumping (Salticidae) from adansoni) where the number of referencesis huge, we New Zealand is presented,including synonymy, recommend the catalogue by Proszynski (2002), where a bibliography,remarks on relationshipsand distribution. complete bibliography is given.

Family Salticidae Blackwall, 1841 Keywords Salticidae,checklist, New Zealand Attidae Sundevall,1883 (Attides) Genuslttzs Walckenaer,I 805 Introduction In the past, many speciesof unclear status were included Only 49 speciesofjumping spiders(Salticidae) have been in the ltlas. Bonnet (1955), for instance,lists over describedor recorded from New Zealand.In fact, the 200 species,10 of them fromNew Zealand.Here, following (2002), numberof descriptionsis much largeras somenames were Proszynski we retain the name as provisional. synonymised.Today, fewer than l0 speciesare easily None of the specieslisted below has proper documentation recognisable- usuallyunder incorrect generic names (e.g., and their verification will be extremely difficult., if possible Salticus, Euophrys, ). The lack of type specimens, at all. poorly documentedoriginal descriptions,great intraspecifi c variationin sizeand colour and interspecificuniformity in Attus abbrevialzs Walckenaer,I 837 genitalicshucture make proper verificationoftaxa a very A t tu s abb rev i a/us Walckenaer, 183'7 :47 7 (female difficult task. In the past, the efforts to provide a proper Attus abbreviatrs: Bonnet, 1955 :793 list of New ZealandSalticidae or to verify earlierdescrip- Salticus albreviatus: Panott. 1942:90 tions were undertakenby Urquhart (1892a),Panott (1942) Type locality' : unknown. and Bryant (1935a,b), all with limited success. In 2000we begana long-termproject on New Zealand Attus aquilus Urquhart, 1886 Salticidaewith the following goals:( 1) to describe Attus aquilus Urquhart, 1886:115 (male,female) (redescribe)all the taxa availablefrom the country (2) to Salticus aquilis: Parrott, 1942:90 analysethe faunistic diversity for particular areasand Attus aquilus:Urquhart, 1892a:230.Bonnet, 1955:794 habitats,(3) to analysethe relationshipsbetween salticids Type locality: North Island: Waiwera, on rocks. ofNew Zealand and other areasofthe S-W Pacific, (4) to estimatethe influence of Gondwanan heritage and pos! Attus coo ki Walckenaer,I 837 Attus (female) Gondwananisolation on New Zealandsalticid faunaand. in cookii Walckenaer,1837 :47 8 the more distantfuture: (5) to reconstructthe Attus cooki:White & Doubleday,1843: 271. Urquhart,1897: zoogeographicalhistory of SalticidaeofAustralia, New 284. Bonnet, 1955:796 Guinea and New Zealand' Some preliminary data of the Salticus cookii: Parrott, 1942:90 project have already been presented(Zabka and Pollard Type locality: unknown. 2001, Zabka et al. 2002). Attus monticolzs The list given below is the result of a critical review of Urquhart, 1891 literature(Roewer 1954; Bonnet 1955, 1956, 1957, 1958, Attus monticolersUrquhart, I 891: 186 (female) 1959;Brignoli 1983;Platnick 1989, 1993, 199'7,2001; Attus monticolrus:Urquhart, 1892a :230 Proszl,nski2002) but also the consequenceof our own Sa lticus mo nt icul a: P anotl, 1942:90 field and laboratory research. Attus monticola:Bonnet, 1955:802 Although we are aware of the provisional nature of the Type locality: South Island: Mount Cook. majority of the genericnames, we believethe check-listwill be a good starting point for future research. Attus montinus Urquhart, I 891 (female) In the text, we eitherrefer to the papersdealing strictly Attus montinus Urquhart, 189 1: I 84 with New Zealand species,or to other works comprising Attus montinus:Urquhart, 1892a:230. Bonnet I 55:803 data on New Zealand taxa. In few cases(e. g., Hasarius Salticusmontinus : P anott, 19 42:90

'Departmentof Zoology,Academy of Podlasie,08-110 Siedlce, Poland, [email protected] 2Canterbury Museum,Rolleston Avenue, Christchurch 800 I , New Zealand,[email protected]

Recordsof CanterburyMuseum Vol l6 pp 73-82 1A

Typelocality: South Island: Mount Cook. Remark. The species is rather uncornmon, though found in number of localities and in various habitats, including leaf Attuspullus Urquhart,1890 litter, sand dunes, sea-shoreand river beds, the latter being Attuspullus Urquhart,I 890:263(female) probably the species' dispersal route. Logunov's (2000) Attuspullus: Urquhart, 1892a:230 Bonnet, 1 955 :805 suggestionthat B. compactus(Urquhart) is ajunior synonl.rn Salticuspullus: Parrott, 1942:90 of B. maculatu,s,cannot be proved as type specimensof the Typelocality: South Island: Bluff, Otago,on Leptospermum. former do not exist.

Attusscindus Urquhart, I 890 Gents Simon, 1901 Attusscindus Urquhart, 1890:264 (female) The only speciesof true Clynotis fC. severus (L. Koch, Attusscindus: Urquhart, 1892a:230 Bonnet, 1955:806 1879)l is known fromAusfalia (Zabka 1988).Others, Salticus s c indus: Parrott, 1 9 42:9 0 including some of those below, should be transferredto a Typelocality: Scinde Island, Hawke's Bay, above the cliffs. new genus comprising a number ofAustralian and New Zealand species. Attussubfuscus Urquhart, I 887 Attussub-fuscrzs Urquhart, I 887:I 17(female) C$norts banesiHogg, 1909 Attussub-fuscus: Urquhart, 1892a:230 Bonnet, 1955:808 Clynotis barceslsHogg, 1909:176 (male, female) Salticussubfuscas: Parrott, 19 42:90 Clynotis Banesl: Dalmas, l9l7:419 Attussubfuscus: Bonnet, 1955 :808 Clynotisb arresrs: Berland, 193| a: 92, 193 I b:35 7. Roewer, Typelocality: North Island:Waiwera, on Cordylineshntbs. 1954:1255 Clynotis banesi: Berland, 1934:170 Parrott, 1942:89. Attus tenebro sasUrquhart, I 893 Bonnet,1956:1 166 Forster, 1964:102 Attustenebrosr.rs Urquhart, 1893: 188 (male) Type locality: Campbell Islands. Salticustenebrosus: Parrott, 1942:90 Attustenebroszs: Bonnet: 1 955.:808 Clynotis knoxi Forster, 1964 Typelocality: North Island: Hunua. Clynotisknoxi Forster,1964:105 (male, female) Clynotisknoxi: Brigroli, I 983.:634 Attus valentulusUrquhart, 1891 Type locality: SnaresIsland. Attusvalentulus Urquhart, I 891: 187 (female) At t u s va I en tu lus: Urquhart,| 892a:23 0 Bonnet: I 955 : 8 09 Clynotis saxatilis (Urquhart, I 886) Salticusvalentulus'. Parrott, 1942:90 Attus saxatilis Urquhart, I 886:204 (female) Typelocality: North Island:Auckland. Attus saxatil i s: Urquhart, 1892a:230 Cly n o t i s sax at i I i s: Bry ant, I 935 b : 88, Parrott, | 942:89, GenusBisnor Peckham,1885 Roewer,195 4:1255, Bonnet, 1956:1 | 67 Proszynski(2002) lists over 20 species,most ofthem from Type locality: South Island: Two Thumb Range,Lake Tekapo, theOld World.Single species are known from thePacific Canterbury Miller's Branch, ClarenceRiver. area(Berry et al. 1996;Logunov 2000). GenusCosmophasis Simon, I 90 I Bianor compaclzs(Urquhart, 1885) A large tropical genus,widely distributed in SE Asia, New Salticuscompaclas Urquhart, I 885:50(female) Guinea, N and in some W Pacific Islands Balluscompaclzs: Urquhart, 1892a:229 (Proszynski 1996; Berry et al. 1997). The records from Bianorcompaclrs: Dalmas, 1917: 419 . Bryant,I 935a:68. Africa should be confirmed. The genus has notbeen found Parrott,1942: 88. Roewer, 1954: 1231. Bonnet, I 955:885. in New Zealandand it is unlikely to occur. Logunov,2000:235 Typelocality: South Island: Lake Tekapo. Co s m op h asis ar ch eyi B erland, 193 | Remarks.Logunov (2000) suggests that the species'nameis Co s mo p h as is ar chey i B erland, I 9 3 I b :3 63 (male, female). ajunior synonymof B. maculatus(Keys.) (see below). Cosmophasisarcheyi: Roewer, 1954:1152. Bonnet, 1956:1241 Bianor maculalzs(Keyserling, I 883) Clynoti s arc heyi: Forster, 1964 : | 04 Scythropamaculata Keyserling, I 883 : I 447 Type locality: Auckland Islands. Bianormaculalus: Simon, I 90I : 638, 641 . Roewer,I 954 Remark. The presenceof Cosmophaslson subantarctic Bonnet,1955 Proszynski, 1971:385 Platnick, 1989:545; islands of New Zealand is unlikely. 1993:740; 1997 :865 . Logunov,2000:240. Typelocality: Peak Downs (Australia). 75

GenusHasarius Simon, 1871 l90l:595,602,608;1909: I 98 Peckham, I 894:124; Rain- In the past,a number of unrelatedspecies have wrongly bow,I 9 I 1:29 1 ; Dalmas,I 9 I 7; Berland,1924: 163, 249.; been placed within Hasarius. In fact, it seemsto be rather Petrunkevitch,1928 :206 ; Roewer,1 9 5 4 :l 257; Bonnet, small genus of Oriental origin, found in the tropics of S and 1957:2228; Mascord, 1970:28; Proszynski, 197 l:418; SEAsia, .Australia and Pacific islands.One I 984:5 1 ; Forster,197 7 a: 52; Davies & Zabka,1 9 89 :262; species(below) has cosmohopicaldistribution. Zabka,199 1 a:200; I 99I b:3 3 Typelocality: Sydney,Peak Downs (Australia). Hasarius adansozi (Audouin, I 826) Remarks.The species tnhabits Eucalyptzs tree trunks inAus- AttusAdansonl Audouin, 1826: 169 tralia.Despite Dalmas' data (1917), it is probablymissing HasariusAdansoni: Simon. 1871 : 1903 fromNew Zealand. Hasariusadansoni: O. P.-Cambridge,1881. Roewer, 1954: Bonnet,1957 Holoplatysappresszs (Powell, 1873), comb. nov. Type locality: Unknown. Salticusappressus Powell 1873:281 (male) syn. nov. Remarks.A cosmotropical species,known from North Island Sa I t i cus apre s sus: Urquhart, | 892a:229; Bonnet, 1 95 8 : 3 909 (oral information by Dr Peter Maddison). For complete Holoplatyssenills Dalmas, l9l7:416 (male,female) syn. bibliography see Proszynski (2002), Holoplatyssenills: Myers, 1927 :132.134; GenusHelpis Simon, I 901 Parrott,1942:89; Roewer,1954:1258; Bonnet, 1957 :2228; Proszynski, Thanksto excellentrevision by Wanless(1988) and later 1911 : 41 8 Zabka,1 99 1 a:l 82 papersby Gardzinska(1996) and Zabka(2002), eight ; Salticusadpressus: Parrott, 1942:90 speciesof the genus are known fromAustralia, adjacent Typelocality: for Salticusappressus'. South Island: on islandsand New Guinea,one is also found in New Zealand. palingsin andaround Christchurch. For senilis thelocality was not given. Helpis minitabunda (L. Koch, 1880) Remarks.The speciesis mostlyfound Astia minitabundaL. Koch I 880:I 16 (male) underbark of manuka(Leptospermum scoparium) and kanuka (Kunzea Astia respersaL. Koch, I 880:I 163 (male,female) ericoides),in termsofbark structure,both are Helpis minitabunda'. Simon, 1901 :43 | -432, 436, 438 the local equivalentof AustralianEucalyptus. Interestingly, Rainbow,I 9 I :280 Petrunkevitch,1928: 1 84; Roewer, the specimenshave also been found under bark ofintroduced 19 5 49 69 ; Bonnet, 19 57 :21 50. Proszl.nski,1 97 1: 417; Pinusradiata andon driftwood,which suggestspossibility Preston-Mafh am & Preston-Mafham, 19 84 :'72 ; | 99 6 : 107 . of raftingfrom Australia. Hallas& Jackson,1986b:883, 889-892. Jackson, 1986:1 194, 1204; 1987 :2-4, 8. Wanless,1988:94; Davies & Gen.Hypoblemzrz Peckham et Peckham, 1886 Zabka,| 989 :2 12 ; Gardzinska,| 996 :299:. Harland& genus Jackson,2001:449 The is knownfrom 2 Australianspecies, 1L albovittatum(Keys.) and H. vilosum(Keys.) (Davies & Helpis respersa:Simon, 1901:436;Rainbow,191 l:280; Zabka1989), the formeris alsofound Roewer,19 5 4 :969 Bonnet, 19 57 :21 5 | Proszynski, in New Zealand ; ; (Zabka 197l:417 & Pollardin print). Type locality: Australia: Sydney:Shelleys Flats. Hypoblem um albovittatum (Keyserling,1 882) Remarks.The speciesis of Australian origin and is mostly Habrocestumalbovittatum Keyserling, 1882: 1407 (male) found on North Island and in northernpart ofSouth Island. Hypoblemumalbovittatum: Simon, I 903:703, 708; Rain- Its presencein New Zealand seemsthe result of aero- or bow,1 9 1I r 299;1920: 267. Bonnet, 1957 : 2255. anthropodispersal(Patoleta & Zabka 1999). Proszynski,197 | :422 Zabka& Pollard,2002. Euophrys parvula: F orster Forster, -62; GenusHoloplalys Simon, 1885 & 1972:5 | 1976 :21 1 4-21 l7 1999 : | 19,| 27-129. Forster, 197 7 a:53 The genuscomprises about 40 describedspecies, mostly ; ; 1982b:1 64, 1 66-1 69, 17 5, 203 I 985:25 3. Hallas& known from Australia (including ),with single ; Jackson,1886b: 883-884,889-891. Jackson, 1986b:1 194; representativesreported from Papua New Guinea, Lord 1987:2-7;1988:348; 1992: 100. Jackson & Hallas,1990: . Howe Islandand (Zabka 1991 a; Patoleta Jackson& Tarsitano,1993:106-108. Jackson & Willey, 2002). DespiteForster and Forster(1999), the number of 1995:1-6.Harland & 2001:447-448 New Zealandspecies is very limited. Jackson. Typelocality: Peak Downs (Australia). Remarks.The original description byBryant (1935b) Holoplatys planissima (L. Koch, I 879) suggeststhat true E. p. representsTrite minax f. Marptusaplanissima L. Koch, I 879: I 100(male, female) f: planicepsf-groupand is neitherrelated to Euophrysnor H ol opI atys p Ianiss ima: Simon, I 8 85:4 45 ; 1898:27 5 ; Hypoblemum.The speciesis oneof the mostcommon 76

"house salticids in New Zealand and is known as hopper" Type locality: North Island: Stratford. from number of localities throughout the country. As it is predominantly found in and around human habitations, it Marpissa leucophaea Urquhart, I 888 seemlikely to have arrived in New Zealand vlc MarpissaleucophaeumUrquhart, 1888: I2I (male,female) anthropodispersalfrom Austraha (Zabka & Pollard in M arpiss a leucophaeum: Urquhart, | 892a:229 print). The mating behaviour of Hypoblemum albovittatum Marpissa leucophaina: Parrott, 1942:89 (asEuophrys parwla) was studiedby Jackson& Willey Marpissaleucophaea'. Bonnet, 1957:2722 ( 1995). Type locality: North Island: Port Waikato. Genus"/afzsL. Koch, 1881 The genus is known from Australia (Davies & Zabka 1989) Marpissa marina (Goyen, I 892) and its presencein New Zealand needs confirmation. Marptusa marina Goyen, 1892:253 (female) Marptusa marina'.Merian, 1913:57 Jotus ravus (Urquhart, 1893) Marpissa marina: Parrott, 1942:89. Bonnet, 1957:27 22 Attus ravas Urquhart, 1893: 1 85 (male). Forster,19'7'7a:54. Jackson, 1986b: I 194;1987:2-7; Jotus ravus: Bryant, 1935:67.Roewer, 1954:1094.Bonnet, 1992:1 00. Jackson& Brassington,1987:229.Jackson et al. 1957:2328 1990:25-3 8. Jackson& Cooper, 1990:40; 1991:78-82 Type locality: North Island: Wellington. Jackson& Tarsitano,1993:108-108. Forster'& Forster, 1999:129.Harland & Jackson,2001:448-449 GenusLaufeia Simon, 1889 Type locality: South Island: coastsofOtago. A small genus, known from E and SE Asia. Despite Bryant Remarks. A well known shore-dweller, representing (1935b),its presencein New Zealandseems unlikely. Euophryinae subfamily, not related to the genus Marpissa. Some aspectsof its biology were studied by Jacksonet al. Laufeia serihirta (Urquhart, 1888) (1990) and Jacksonand Cooper (1991). M. marinaisprobably Marpissa aeri- hirta Urquhart, I 888: I 24 (female) a junior synonym of Salticus atrallrs Powell, however we do M arp iss a aeri hirt a: Urquhart, | 892:229 not proposeany formal changesuntil the generic statusofthe Laufeia aerihirta: Bryant, I 935b:80 former is revised. Laufeiaaerihirta: Bonnet, 1957:2384 Type locality: North Island: Te Karaka. M arpissa n emoralis Urquhart, 1892 M arp iss a nemor al i s Ur quhart, I 892b :2 50 (male, female) GenusMarpissa C. L. Koch, 1846 Marpissa nemoralis: Parrott, 1942:89. Bonnet, 1957:2726 This widespread genus consists of nearly 50 nominal Type locality: North Island: forests near Stratford. species(Proszynski 2002), many ofthem to be excludedas non-congeneric.No representativesoftrue Marpissa are GenusOcrisiaza Simon, 1901 known from New Zealandand all nominal specieslisted The genusincludes about 10 species,mostly from Australia below have to be transferredto other genera. (Zabka 1990) and is probably missing from New Zealand.

Marpissa arenaria Urquhart, 1888 cinerea (Koch, I 879) Marpissa arenaria Urquhart, I 888: 123 (female Marptusa cinereaKoch, I 879:1 I 12 (female) Marpissa arenaria: Urquhart, 1892a:229.Bonnet, Ocrisionacinerea'. Dalmas, 1917 :419. Parrott, 1942:89; 1957:2721 Bonnet, 1958:3124 Evophrysarenaria: Parrott, 1942:88 Remarks.The specieshas originally been describedfrom Type locality: North Island: Port Waikato, sandhills. New Zealand (with no type locality) but should be excluded from Ocrisiona and transferred either to M arpissa arm ifera Urquhart, I 892 Holoplatys or minax-grouP. Marpissa armifera Urquhart, 1892b:248 (male, female) Marpissa armifera: Bryant 1935a:69.Parrott, 1942:89. Ocrisiona leucocomis (L. Koch, I 879) Roewer 19541261.Bonnet, 195'7:27 21 Marptusa leucocomisL.Koch, 1879: 1096(male, female) Type locality: South Island: Albury, on limestone rocks. Ocrisiona leucocomis: Simon, I 90 1: 5 96,602, 608-609; 1909:200.Dalmas, 1917:418 Parrott, 1942:90. Bonnet, Marpissa cineraceaUrquhart, I 891 | 95 8 :3 | 24. Proszynski,197 1 : 4 43 . Zabka, 199 0 :30. Marpissa cineracea Urquhart, 189 1: I 88 (female) Type locality: Sydney, Bowen, Rockhampton, Peak Downs Marpissa cineracea'.Urquhart, 1892a:229. Parrott, 1942:89 (Australia). Bonnet.1957:2722 Remarks.The dataon species'presence in New Zealand(Koch M arpis s a cineracca: Bonnet, 1957 :27 22 I 879) hasto be confirmed. 77

GenusPlexipprs C. L. Koch, I 846 Attus curwts; Urquhart, 1892a:229.Parrott, 1942:90 Bonnet(1958) lists nearly 70 nominal species;only some Type locality: North Island:Whangarei Harbour. of them representtrue Plexippus. Two cosmotropical speciesrelated to human habitations (P. paykulli andP. Salticusfumosus Powell, 1873 petersi) are likely to occur in North Island, but none has SalticusfumosasPowell, 1873:284 (male, female) beenrecorded so far. Salticusfumasas: Urquhart, 1892a229.Parrott, 1942:90. Bonnet, 1958:3912 Plevippus sylvarusUrquhart,lS92 Type locality: South Island: Christchurch, Riccarton Bush, Plexippus sylvanrs Urquhart, 1892b:252(male) dead leavesclothing the trunk ofthe cabbage-trees Plexippus sylvarus: Parrott, 1942:88 (Cordyline sp.). Powell's colour plate (1873) suggeststhat Plexippus silvarzs: Bonnet, 1958:3721 this might be Trite auricoma,but we retain this specieshere Type locality: North Island: forest near Stratford. until the generic statusof T. auricoma is cleared.(see Remarks.The generic afliliation of the speciesis unknown. below).

Genus,SoltlcasLatreille, 1804 Salticus fumus Urquhart, 1885 In the past, many unrelated specieswere placed within Salticusfurwts Urquhart, 1885:47 (female) Salticus,including I I New Zealandrepresentatives. The Salticusfurvus:Parrott, 1942:90.Bonnet, 1958r3914 genusis mostly given from Palaearcticand Nearctic Regions Type locality: North Island: Karaka, Auckland. and is unlikely to occur in New Zealand. Salticus tabinus Urquhart, 1885 Salticusatbipalpis Urquhart, 1885 SalticustabinusUrquhart,1885:46(female) Salticus albopalprs Urquhart, 1885:49 (female) Attus tabinus'.Urquhart, 1892a:229 Attus albo-palpls: Urquhart, 1892a:299 ,Salticustabinus: Parrott, 1942:90 Bonnet, 1958:3925 Salticusalbipalprs: Bonnet, 1958:3908 Type locality: North Island: Scoriawalls, North Shore, Salticus albo-palpis: Panott,1942:90 Auckland. Type locality: South Island:mountains above the Otira Gorge, Westland. Salticus tenebricas Urquhart, 1885 Salticustenebrlcus Urquhart, 1885:51(male) Salticus albobarbatus Powell, 1873 Attus tenebricus: Urquhart, 1892a:229 SalticusalbobarbatusPowell,l8T3.286 (male, female) Salticustenebricus:Parrott,1942:90 Bonnet, 1958:3925 Salticusalbo-barbatus'. Urquhart, 1892:229.Parrott, Type locality: SouthIsland: shingleslopes, mountains near 1942:90.Bonnet, 1958:3908 Lake Tekapo,Canterbury. Type locality: South Island:Castle Hill, shingleslides. Salticus v-notatus Powell, 1873 Salticus alpinus Urquhart, 1885 SalticusV-notatusPowell, 1873:284 (male) Salticus alpinu.sUrquhart, 1885:48 (male, female) Salticus V-notatus:Urquhart 1892a:229.Parrott, 1942:90. ' Attus alpinus:Urquhart, 1892a:229 Bonnet,1958:3926 Salticusalpinu.s: Parrott, 1942:90. Bonnet, 1958:3908. Type locality: South Island: Christchurch,Oxford Terrace. Type locality: South Island: solid rocks ofCanterbury and Westland. Salticus zanthrofrontafis Urquhart, 1885 Salticus zanthofrontalis Urquhart, I 885:45 (male) SalticusatratusPowell,l8T3 Attuszanthofrontalis:Urquhaft,1886:203;1892a:230 Salticusatrutus Powell, 1873:283 (male) Attus zanthofrontalisvar. hirta: Urquhart, 1886 :204 Salticus atratus: Urquhart 1892a:229.Parrott, 194290. Salticusxanthofrontalis: Parrott, 1942:90. Bonnet, Bonnet,1958:2909 1958:3926 Type locality: South Island: Sumner, on rocks. Type locality: North Island: Karaka, Auckland. Remarks. It probably is the same species as"Marpissa marina" (see above) but we do not propose any formal Genus Trile Simon, 1885 changesuntil the statusofNew Zealand"Marpissa" is The list given below includesat least2 groupsofspecies: revised. Trite minax {: planicepsl representingtrue Trite and T. auricoma- which shouldbe excludedas not related.Except Salticus czruzs Urquhart, 1885 from New Zealand, true Trite is known from the W Pacific Salticus curwtslJrquharq 1885:46 (male) islands (Zabka 1988; Berry et al. 1997, Patoleta 2002). T\e Salticuscurws'. Bonnet, 19583912 numberof speciesand greatinter- and intraspecific 78

variation suggestNew Zealand is the centre oforigin and 581,583,585,590. Jackson & Willey, 1994:80.Pollard et distributionofTrite. al., 1987:137. Taylor, 1997:341-343.Forster & Forstel 1999:39, 119, l2l, 125-126 Trite auricoma (Urquhart, 1886) Triteminax: Bryant 1935a:67. Bonnet, 1959:4695 Attus auricomus Urquhart, 1886:202 (male, female) Type locality: for T. minax'. South Island: Riccarton Bush, P lexippu s cap i I I atu s Ur quhart, 18 90 :265 (female) Govemor Bay, in dead leavesclothing the trunks ofcabbage Attus auricomus: Urquhart, 1892al.230 tree (Cordyline sp.). For T. planiceps'. South Island: French Attussuffuscus Urquhart, 1893: 1 86 (female) Pass,Riccarton Bush. Attus kirkii Urquhart, I 893:1 87 (male) Remarks.A well known (as T. planiceps) representativeof Attus adustusUrquhart, I 893: I 89 (female) the genus,found on flax(). Trite vafra Dalmas, 1917:424 (male) Triteauricoma: Bryant, 1935:64.Roewer, 1954:103 1. Trite mustilina (Powell, 1873),comb. nov. Bonnet, 1959:469 4. Proszynski,197 1 :481. Forster, Salticusmustihnzs Powell, I 873:285(male) 197 7 a: 52-5 4, 56-57, 60 -61; 197 7b:43 5 -443; 1977 c:3 5 -36; Attus bimaculoszs Urquhart, I 887: I I 6 (female) | 9 82a:1 69 ; 19 82b: 169 - | 75, I 80- I 82, 20 4-205, 207 -208, Attus bimaculosrzs:Urquhart, I 888: I 25, 1892a:230 2ll; 1982c:181Jackson, 1986a: I 10; 1986b:I 196,1204; Salticus mustillnus: Urquhart , 1892a:229.Parrott, 1942:90. I 988:348; I 992:1 00; 1987:2-7 . Jackson& Hallas, Bonnet,1958:3914 1986:492. Hallas& Jackson,19 86a: 5 44;'I 986b:8 83, 88 8- Tritebinotata Simon, 1899:431(male) 892. Jackson& Brassington,1987:229. Zabka,1988474. Tritebinotata: Simon, 1903:826.Dalmas, 1917:423. Jackson& Cooper, 1990:39-40.Jackson & Hallas.Jackson Roewer,I 954:I 03 I & Tarsitano,1993:106-106. Forster & Forster,1999:1 19- Trite bimaculosa:Bryant, 1935:91.Parrot, 194:87.Roewer, 123, 125 - 126, 132. Harland & Jackson,200 1 : 447 -448 1954: 1 03 1. Bonnet, 1959:4695. Proszynski, 197 | :481. Type locality: North Island: Te Karaka, Auckland, Pirongia Forster,1982a:166; 1982b:162. Jackson, 1 986b: 1 I 96 Mountain. Type locality: for Salticus mustilinus: South Island: Remarks.It is one of the most cofirmonsalticids in New Christchurch, Riccarton Bush, on shrubs. Zealand,representing a large group ofspecies, none ofthem For Attus bimaculosus:North Island: Whangarei Harbour, relatedto the genusTrite andall shouldprobablybe placed Waiwera, StephensIsland. in a new genus.,Softiczs fumosus Powell seemsa junior synonym of T. auricoma, however, we retain the name of Remarks.The speciesbelongs to Trite minaxf= 7. the latteruntil a new genericname for it is proposed. p lanicepsl-group of species.

Trite herbigrada (Urquhart, I 889) Trile pamula (Bryant, 1935) comb. nov. PlexippusherbigradusUrquhart, I 889:150 (male, female). Evophrysparvula Bryant, 1835b:90(male), syn. nov. Plexippus herbigradus: Urquhart, 1892a:229 Evophrysparvula: Bonnet, 1956: 1885. Trilelucida Simon, 1902:395 Type locality: South Island: Christchurch. Trit e lucida: Simon, 1903 : 8 26, 827,829. Dalmas,19 l7 :423 Remarks.Since its original description, the specieshas not Trite herbigrada: Bryant, 1935a:66.Roewer, 195:1032. been studied. In the last decades,its name has wrongly been Bonnet, 1959:4695.Proszynski, 197 1 :481 used for other unrelated species(Hypoblemum albovittatum Type locality: North Island: summit of Te Aroha up the - seeabove). WairongomaiGorge, on ferns. Trite uwillei (Dalmas, 1917) Trite mintx (Powell, 1873) Holoplatys urvillei Dalmas, 1917 :417 (female). Salticusminax Powell 1873:282 (male) Holoplatys urvillei: P arrott, 1942:89. Roewer, 1954:1254. P lexippus minax: Urquhart, | 892a:229 Bonnet, 1957:2228 Triteplaniceps Simon, 1899:430 (female) Trite urvill ei: Zabka, 199 1a: 17 2 Triteplaniceps: Dalmas 1917:442. Roewer, 1954:1032. Type locality: North Island: Taumaranui. Proszynski,197 | :481.Forster, 197'7 a: 52-53, 5 6; 1979 :79 - Remarks. A common species found inside rolled-up flax 93; I 982b:I 65, 170-173. I 982b:I 6 l, 176-182, 195, 202- (Phormium tenax) leaves, wrongly considered to represent 205,21 1 ; 1982c:179 -1 83 ; 1985.251, 253-260,264-265, the genus Holoplatys. Its biology (as Holoplatys sp.) was 267-268.Hallas & Jackson,1986b:883, 890-892. Jackson, studiedby Jacksonand Harding ( I 982). I 886:1 I 96, | 198, 1202, 1204; 1987: 2-7; 1992:100 Jackson& Hallas, 1986:492. Jackson& Brassington, Acknowledgements 1987:229. Zabka, 1988:473. Jackson& Tarsitano, The researchwas possibledue to the 1993:106-108. Jackson & van Olphen,l99l:577-578, 580- Grant 6PO4C01115 79 provided for MZ by the Polish State Committee of Douladoure,Toulouse, 2( I ): I -91 8 ScientificResearch (KBN). MZ is also gateful to the BonnetP (1956)BibliographiaAraneorum. Les Frdres Drrector and staff of the Canterbury Museum (Christch- Douladoure,Toulouse, 2(2):9 19 -1 926 urch, New Zealand), and to the fiiends ofthe Invertebrate BonnetP (1957) BibliographiaAraneorum. Les Frdres Zoology Department who were most kind and co-operative Douladoure,Toulouse, 2(3): 1927 -3026 during his 3-month researchvisit. Mathew Anstey is BonnetP (1958)BibliographiaAraneorum. Les Frdres acknowledgedfor his enthusiasm,help and professional Douladoure,Toulouse, 2(4):3027 -4230 skills. Specialthanks go to Brian Patrick, Erena Barker and BonnetP (1959)BibliogaphiaAraneorum. Les Frdres Simon Wylie (OtagoMuseum, Dunedin), John Early Douladoure,Toulouse, 2(5):423 | -5058 (AucklandMuseum EntomologyCollection), Phil Sirvid BrignoliPM ( 1983)A Catalogueof theAraneae described (Museum of New Zealand, Wellington), Trevor Crosby between1940 and 1981.Manchester University Press, (New ZealandArthropod Collection,Auckland) and Cor Manchester Vink (Entomology ResearchMuseum, Lincoln University, BryantE (1935a)Notes on someUrquhart's Species of Lincoln) - the curatorsofarachnological collections, who Spiders.Rec Cant Mus 4;54-70 were most co-operative in providing material for study. The BryantE (1935b)Some New andLittle Known Species of project has also been supportedby Mike Gray (Australian NewZealand Spiders. Rec Cant Mus 4:71-94 Museum,Sydney), Robert Raven (QueenslandMuseum, DalmasR de( I 9 17) Araneign6es de Nouvelle Z6lande. Ann Brisbane), Robert Jacksonand his students (Canterbury Socent Fr 86:3I 7-430 University, Christchurch), Cor Vink (Lincoln University, DaviesTV, ZabkaM (1989)Illustrated keys to thegenera Lincoln) and Jerzy Proszynski (Museum and Institute of ofjumpingspiders (Araneae:Satticidae). NZ J Zool4:51- Zoology,Polish Academy of Sciences,Warsaw). 62 The researchwould not havebeen possible without decades ForsterLM (1977a)Aqualitative analysis ofhunting ofarachnological researchcarried out by the late Ray behaviourin jumpingspiders (Araneae: Salticidae). NZ J Forster (Otago Museum, Dunedin) whose impressive Zool4:51-62 collections from throughout New Zealand contributed ForsterLM (1977b)Some factors affecting feeding enormouslyto the project. behaviourin youngTriteauricoma spiderlings (Araneae: Salticidae).NZ J Zool4:435-443. Bibliography ForsterLM (1977c)Mating behaviour in Triteauricoma, a NewZealand jumping . Peckhamia | :35-36. Audouin V (1826)Explication sommaire des Planches ForsterLM (1979)Visual mechanisms of huntingbehaviour (4): -339 d'Arachnidesde I'Egypte et de la Syrie 1 1 in Triteplaniceps, ajumping spider (Araneae: Salticidae). Berland L ( I 93 I a) Les Araign6esdes iles avoislant la NZ I Zool 6:79-93 Nouvelle Zelandeet les relationsentre I'Australie et ForsterLM (1982a)Vision and Prey-Catching Strategies in I'Ameriquedu Sud.C R Socbiogeogr 1930,60:90-94 JumpingSpiders. Amer Sci 70: 165-175 BerlandL ( 193 1b) Araign6esdes iles Auckland et Campbell. ForsterLM (1982b)Visual communication injumping RecCant Mus 3:357-365 spiders(Salticidae). In: Witt PN,Rovner JS (eds) Spider (1 In: BerlandL 934) LesAraign6es du Pacifique. Contribu- Communication:Mechanisms and Ecological Signifi- tion d 1'6tudedu peuplementzoologique et botaniquedes cance.Princeton Univ. Press, Princeton, NJ ppl6l-212. ilesdu Pacifique.Publ Socbiogeogr 4: 155-180 ForsterLM (1982b)Visual communication in jumping (1996) Beny JW, Beatty JA, ProszynskiJ Salticidaeof the spiders(Salticidae). In: Witt PN, RovnerJS (eds) Spider genera, PacificIslands. I. Distribution of twelve with Communication:Mechanisms and Ecological Sigrifi- descriptionsofeighteen new species.J Arachnol 24:214- cance.Princeton Univ. Press, Princeton, NJ ppl6l-212 253 ForsterLM ( I 982c)Non-visual prey-capture in Trite BerryJW, Beatty JA, Proszynski J (1997) Salticidae of the planiceps,ajumping spider (Araneae, Salticidae). J genera, PacificIslands. II. Distribution of nine with Arachnol10:179-183 descriptionsofeleven new species.J Arachnol 25:109- ForsterLM (1986)Target discrimination in jumping 136 spiders.Proceedings ofthe Ninth InternationalCongress Blackwall J ( 1841 ) The Difference in the Number of Eyes ofArachnology.Smithsonian Institution Press, p3 I 3 proposed with which Spiders are provided as the basis ForsterRR (l 964)The Araneae and Opiliones of the of their Distribution into Tribes; with Descriptions of subantarcticislands of New Zealand.Pac Isl Monogt newly discovered Speciesand the Charactersof a new 7:58-ll5 Family and three new Genera of Spiders. Trans Linn Soc ForsterRR, ForsterLM (1973)New Zealandspiders: an Londl8:601-670 introduction.Collins, Auckland BonnetP (1955) BibliographiaAraneorum.Les Frdres ForsterRR, ForsterLM (1976)Jumping spiders. New 80

Zealandnatural heritage, pp21 14-21 17 andprior residencyon the outcomeof male-male Forster RR, Forster LM (1999) Spiders of New Zealmd interactions of Marpissa marina, a New Zealand and their Worldwide Kin. University of Otago press, (Araneae, Salticidae). Ethol Ecol Evol Dunedin 3:78-82 GardzinskaJ (1996) New speciesand recordsofAstieae JacksonRR, Hallas SEA (1986) Predatoryversatility and (Araneae:Salticidae) from Australia and PapuaNew inhaspecific interactions of spartaeinejumping spiders Guinea.Mem Qd Mus 39:297-305 (Araneae:Salticidae): Brettus adonis, B. cingulatus, GoyenP (1892) On New ZealandAraneae. Trans proc N Cyrba algerina, and Pheacius sp. indet. NZ J Zool ZealInst 24:253-257 13:491-520 Hallas SEA, JacksonRR (1986a)A comparativestudy of JacksonRR, Harding DP (1982) Intraspecificinteractions Old and New World Lyssomanines(Araneae: Salticidae): of Holoplatys sp. indet., a New Zealand jumping spider utilisationof silk and predatorybehaviour of Asemonea (Araneae:Salticidae). NZ J Zool 9:487-510 tenuipes and Lyssomanesviridis. N Zeal J Zool 13.543- JacksonRR, Pollard SD, MacnabAM, CooperKJ (1990) 551 The complex communicatory behaviour of Marpissa HallasSEA, JacksonRR ( 1986b)Prey-holding abilities of marina, a New Zealandjumping spider(Araneae: the nestsand webs ofjumping spiders(Araneae, Salticidae).NZ J Zool 17:25-38 Salticidae).J Nat Hist 20:881-894 JacksonRR, TarsitanoMS (1993) ResponseSofjumping HarlandDP, JacksonRR (2001) Prey classificationby spidersto motionlessprey. Bull Br arachnolSoc 9: 105- Portiafimbriala, a salticid spiderthat specializesat 109 preying on other salticids:species that elicit crlptic JacksonRR, Willey MB (1994) The comparativestudy of stalking.J Zool Lond 255:445-460 the predatory behaviour of Myrmarachne, ant-like Hogg HR ( I 909) Spidersand Opiliones from the jumping spiders(Araneae: Salticidae). Zool J Linn Soc SubantarcticIslands ofNew Zealand.In: The 110:77-102. SubantarcticIslands of New Zealand I : I 55- I 8 I JacksonRR, Willey MB (1995) Display and mating http://research.org/entomology/spiders/catalog81-87l behaviourof Euophrysparwla,aNew Zealandjumping INTROl.html spider(Araneae: Salticidae). NZ I Zool22:l-16 JacksonR.R, van Olphen A (1991) Prey capturetechniques KeyserlingE (1882) Die ArachnidenAustraliens, nach der and prey preferencesof Corythalia canosa andPystira Natur beschriebenund abgebildet.Niimberg, 2:1325- orb icul ata, ant eatingjumping spiders (Araneae, 1420 -591 Salticidae).I Zool 23:577 KeyserlingE (1882) Die ArachnidenAustraliens, nach der JacksonRR (1886b) Cohabitationofmales andjuvenile Natur beschriebenund abgebildet.Nrirnberg, 2: 1421 - females: a prevalent mating tactic of spiders. J Nat Hist 1489 20:1193-1210 Koch CL ( 1846) Die Arachniden.Niirnberg, 13: 1-23 4 JacksonRR ( I 986a)The biology of Phyacescomosus Koch L (1879) Die ArachnidenAustraliens, nach der Natur (Araneae:Salticidae), predatory behaviour, antipredatory beschriebenund abgebildet.Nrimberg, 2:1045-1 156 adaptationsand silk utilization. Bull Br Mus nat Hist Koch L (1880) Die ArachnidenAustraliens, nach der Natur (Zool)50: I 09-1I 6 beschriebenund abgebildet.Niimberg, 2: I | 57-1 21 2 JacksonRR (1987) Comparativestudy ofreleaser Koch L ( I 88I ) Die ArachnidenAushaliens, nach der Natur pheromonesassociated with the silk ofjumping spiders beschriebenund abgebildet.Ntirnberg, 2:1213-127 | (Araneae,Salticidae). NZeal JZool l4:1-10 LatreillePA (1804) Tableaum6thodiques des Insectes.N JacksonRR (1988) The biology of Taualalepidus, a Dic hist nat24:129-200 jumping spider(Araneae: Salticdae) from : LogunovDV (2000)Aredefinition of the generaBianor display and predatorybehaviour. NZ J Zool l5:34i-364 Peckham& Peckham,1885 andHarmochirusSimon, JacksonRR (l 992) Conditional strategiesand 1885, with the establishmentof a new genusSibianor interpopulationvariation in the behaviourofjumping gen.n. (Aranei: Salticidae).Arthrop Sel9:221-286 spiders.NZ I Zool 19:99-11l MascordR (1970) AustralianSpiders in Colour.AH and JacksonRR, BrassingtonRJ ( I 987) The biology of Pholcus AW Reed Pty Ltd., Sydney phalangioides (Araneae,Pholcidae): predatory versatil- Merian P ( I 9 I 3) Les Araigndes de la Terre de Feu et de la ity, araneophagyand aggressivemimicry. J Zool Lond Patagonie,comme point de ddpart de comparaisons 211:227-238 gdographiquesentre couchesfaunistiques. Rev Mus La JacksonRR, CooperKJ (1990) Variability in the responses Plata20:7-100 ofjumping spiders(Araneae: Salticidae) to sex Myers JG (1927) EthologicalNotes on someNew Zealand pheromones.NZ J Zool l7:39-42 Spiders.Second Contribution. NZ J Sci 9:129-136 JacksonRR, CooperKJ (1991) The influenceofbody size Ocrisiona Simon 1901 . Rec Austral Mus 42:27 -43 81

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For almost all species listed in this paper neither the type specimensnor the type localities have been formally "type designated,thus locality''here means the localities taken fiom the original papers.