Natural History Sciences. Atti Soc. it. Sci. nat. Museo civ. Stor. nat. Milano, 6 (2): 3-10, 2019 DOI: 10.4081/nhs.2019.410
Vascular flora of Milan Malpensa airport (Lombardy, Italy). Part I: checklist
Marco Martignoni1, Enrico Banfi2, Gabriele Galasso2*
Abstract - Airport areas have, among flight infrastructures, large Casorate Sempione, Ferno, Lonate Pozzolo, Samarate, Somma Lombardo green spaces which, not having production purposes, are managed e Vizzola Ticino. Le campagne di raccolta si sono svolte da agosto 2010 a through interventions of minimal disturbance on habitats, with poten- settembre 2018 e i campioni sono conservati presso l’erbario del Museo tially positive effects on the conservation of biodiversity. The checklist of di Storia Naturale di Milano (MSNM). L’elenco floristico, organizzato the vascular flora of the airport of Milan Malpensa is here presented. This seguendo l’ordinamento e la nomenclatura delle recenti checklist della airport extends for 12.44 km2 in the southwestern portion of the province flora italiana, annovera 395 taxa di rango specifico e sottospecifico (9,35% of Varese (Lombardy, Italy) in the administrative districts of Cardano al dell’intera flora lombarda), appartenenti a 247 generi e 72 famiglie. Le Campo, Casorate Sempione, Ferno, Lonate Pozzolo, Samarate, Somma entità native sono 318, quelle alloctone 77. Il rapporto autoctone/alloctone Lombardo and Vizzola Ticino. The collection campaigns were carried risulta molto simile a quello calcolato a livello regionale (4,10 vs. 4,42). out between August 2010 and September 2018. The specimens collected Il contingente endemico annovera due taxa: Centaurea nigrescens subsp. are now kept in the herbarium facility of the Natural History Museum of pinnatifida, endemica italiana, ed Euphrasia cisalpina, endemica alpica. Milan (MSNM). The floristic list was organized following the arrange- L’aeroporto di Malpensa rimane ad oggi l’unico sito confermato a livello ment and the nomenclature of the recently published Italian flora check- nazionale per Lepidium heterophyllum e, a livello regionale, per Festuca lists. It includes 395 taxa of specific and subspecific rank (9.35% of muralis ed Euphrasia cisalpina. Le caratteristiche ambientali dell’area di the entire flora of Lombardy) belonging to 247 genera and 72 families. studio e l’analisi dei dati verranno presentati in un secondo contributo. The native taxa are 318, while the alien ones are 77. The native/aliens ratio is comparable to the one calculated at the regional level (4.10 vs. Parole chiave: aeroporto, Euphrasia cisalpina, Festuca muralis, 4.42). The endemic contingent includes two taxa: Centaurea nigrescens Lepidium heterophyllum, Malpensa. subsp. pinnatifida, Italian endemic, and Euphrasia cisalpina, endemic to the Alps. The Malpensa airport is still the only site confirmed at the national level for Lepidium heterophyllum and, at the regional level, for Festuca muralis and Euphrasia cisalpina. The ecological character- INTRODUCTION istics and the data analysis will be discussed in a second contribution. Airport areas include, among their flight infrastruc- tures, large green spaces (Fig. 1c) that, not having produc- Key words: airoport, Euphrasia cisalpina, Festuca muralis, Lepi- tion purposes, are managed through minimum disturbance dium heterophyllum, Malpensa. interventions on habitats, with potential positive effects on the conservation of biodiversity (Martignoni et al., 2016). Riassunto - Flora vascolare dell’aeroporto di Milano Malpensa We hereby present the checklist of the vascular flora (Lombardia, Italia). I parte: checklist. of the green areas of Milan Malpensa airport, which in- Le aree aeroportuali presentano, tra le infrastrutture di volo, ampi cludes well preserved semi-natural habitats, like heath- spazi verdi i quali, non avendo finalità produttive, vengono gestiti lands, grasslands and meadows. These areas are managed mediante interventi di minimo disturbo sull’habitat, con effetti potenzial- mente positivi sulla conservazione della biodiversità. Viene qui presentata through periodic mowing that hinders the ecological suc- la checklist della flora vascolare del sedime aeroportuale di Malpensa, cession to the forest climax vegetation. The ecological esteso per 12,44 km2 nella porzione sud-occidentale della provincia di characteristics of the study area and the data analysis will Varese (Lombardia, Italia) nel territorio dei comuni di Cardano al Campo, be discussed in a second contribution.
1 Piazza G. Matteotti 25, 21050 Lonate Ceppino (VA), Italia. MATERIALS AND METHODS E-mail: [email protected] The study area includes the entire airport of Milan 2 Sezione di Botanica, Museo di Storia Naturale di Milano, Corso Malpensa (Fig.1c) that extends for 12.44 km2 in the Venezia 55, 20121 Milano, Italia. south-western portion of the province of Varese (Lom- E-mail: [email protected] bardy, Italy) (Fig. 1a) in the administrative districts of * Corresponding author: gabriele.galasso@comune.milano.it Cardano al Campo, Casorate Sempione, Ferno, Lonate Pozzolo, Samarate, Somma Lombardo and Vizzola Ti- © 2019 Marco Martignoni, Enrico Banfi, Gabriele Galasso cino (Fig. 1b). The altitude ranges from 206.0 to 234.6 m a.s.l. The soil lays on fluvio-glacial gravelly sediments Received: 21 December 2018 Accepted for publication: 17 April 2019 (Servizio Geologico d’Italia, 1967). Online publication: 24 October 2019 Average annual rainfall is 1211.7 mm, well distributed in the vegetative season with two equinoctial maxima 4 Marco Martignoni, Enrico Banfi, Gabriele Galasso
Fig. 1 - Location of the study area at national (a: red arrow) and provincial level (b: red contour) and a simplified map of the Milan Malpensa airport (c): green color highlights the vegetated areas. / Localizzazione dell’area di studio a livello nazionale (a: freccia rossa) e provinciale (b: contorno rosso) e una mappa semplificata del sedime aeroportuale di Milano Malpensa (c): il colore verde evidenzia le aree vegetate. (Drawing / disegno: M. Martignoni). (May and October) and two minima, one in winter (Janu- specimens are kept in the herbarium facility of the Natural ary and December) the other in summer (July); climatic History Museum of Milan (MSNM; acronym according drought do not occurs. Annual average temperature is to Thiers, 2018). The floristic list was organized follow- 11.6 °C, with the hottest month in summer (July, 22.2 °C) ing the arrangement and the nomenclature of the recently and the coldest in winter (January, 1.7 °C) (Centro Nazi- published Italian vascular flora checklists (Bartolucci et onale di Meteorologia e Climatologia Aeronautica, 2008). al., 2018; Galasso et al., 2018), except for native hybrids According to Rivas-Martínez et al. (2015), the bioclimate not considered in the above-mentioned checklist. is temperate continental, the thermotype mesotemperate, For each taxon, the following information is reported: the ombrotype humid. The potential natural vegetation is accepted name, cryptogenic and alien status in Lombardy. a deciduous upland oak forest (pedunculate and/or ses- These abbreviations are used in the floristic list: A -(ar sile oakwood) (Società per azioni Esercizi Aeroportuali, chaeophyte), N (neophyte), CAS (casual), NAT (natu- 2010). ralized), INV (invasive), C (cryptogenic, i.e. doubtfully The collection campaigns were carried out in the native taxon, whose origin of occurrence in Lombardy is period between August 2010 and September 2018. The unknown). Vascular flora of Milan Malpensa airport (Lombardy, Italy). Part I: checklist 5
RESULTS ibrido; analogamente a Banfi & Galasso (2010) sono The spontaneous vascular flora of Malpensa airport state incluse in questa specie. amounts to 395 taxa of specific and subspecific rank (9.35% A CAS Narcissus ×incomparabilis Mill. nothosubsp. of the entire flora of Lombardy: Galasso et al., 2018) be- incomparabilis (= N. poëticus L. × longing to 247 genera and 72 families. The native taxa are N. pseudonarcissus L. subsp. 318 including 2 hybrids, while the alien ones are 77. The pseudonarcissus) native/aliens ratio is comparable to the one calculated at the regional level (4.10 vs. 4.42). According to Bartolucci Asparagaceae et al. (2018), these statistics does not take into account the Anthericum ramosum L. native hybrids. The endemic contingent includes Centau- Asparagus officinalis L. subsp. officinalis rea nigrescens subsp. pinnatifida, Italian endemic, and Muscari comosum (L.) Mill. Euphrasia cisalpina, endemic to the Alps. The Malpensa Ornithogalum divergens Boreau airport is still the only site confirmed at the national level Polygonatum multiflorum(L.) All. for Lepidium heterophyllum (Martignoni et al., 2016) and, at the regional level, for Festuca muralis (Martignoni et Juncaceae al., 2016) and Euphrasia cisalpina (Martignoni, 2014). Juncus articulatus L. subsp. articulatus Juncus bufonius L. Juncus effusus L. subsp. effusus CHECKLIST N INV Juncus tenuis Willd. Luzula campestris (L.) DC. subsp. campestris Ferns and fern allies Luzula multiflora(Ehrh.) Lej. subsp. multiflora Equisetaceae Equisetum arvense L. Cyperaceae Carex brizoides L. Dennstaedtiaceae Carex caryophyllea Latourr. Pteridium aquilinum (L.) Kuhn subsp. Carex divulsa Stokes aquilinum Carex hirta L. Carex leporina L. Dryopteridaceae Carex pallescens L. Dryopteris filix-mas (L.) Schott Carex spicata Huds. Cyperus esculentus L. Gymnosperms N INV Cyperus microiria Steud.
Pinaceae Poaceae N NAT Pinus rigida Mill. Agrostis gigantea Roth subsp. gigantea Pinus sylvestris L. Agrostis stolonifera L. subsp. stolonifera Aira caryophyllea L. Angiosperms Anisantha diandra (Roth) Tutin ex Tzvelev Anisantha madritensis (L.) Nevski subsp. Orchidaceae madritensis Anacamptis morio (L.) R.M.Bateman, Anisantha sterilis (L.) Nevski Pridgeon & M.W.Chase Anisantha tectorum (L.) Nevski Neotinea ustulata (L.) R.M.Bateman, Anthoxanthum odoratum L. Pridgeon & M.W.Chase Arrhenatherum elatius (L.) P.Beauv. ex Platanthera bifolia (L.) Rich. J.Presl & C.Presl subsp. elatius A NAT Arundo donax L. Asphodelaceae Avena barbata Pott ex Link N NAT Hemerocallis fulva (L.) L. Avenella flexuosa(L.) Drejer subsp. flexuosa Avenula pubescens (Huds.) Dumort. subsp. Amaryllidaceae pubescens Allium vineale L. Bothriochloa ischaemum (L.) Keng Narcissus poëticus L. Brachypodium rupestre (Host) Roem. & Schult. A CAS Narcissus pseudonarcissus L. subsp. Bromopsis erecta (Huds.) Fourr. subsp. erecta pseudonarcissus s.l. Bromopsis erecta (Huds.) Fourr. subsp. Notes: There are only double-flowered cultivars Narcis( - stenophylla (Link) H.Scholz & Valdés sus Group 4), which cannot be traced for sure to a partic- N INV Bromopsis inermis (Leyss.) Holub subsp. ular species or hybrid; following the suggestion of Banfi inermis & Galasso (2010), they were included in this species. Bromus hordeaceus L. subsp. hordeaceus Note: Sono presenti soltanto cultivar a fiore doppio (Narcissus Gruppo 4), che non possono essere ricon- Bromus hordeaceus L. subsp. molliformis dotte con sicurezza a una specie o a un determinato (J.Lloyd ex Billot) Maire & Weiller 6 Marco Martignoni, Enrico Banfi, Gabriele Galasso
Calamagrostis epigejos (L.) Roth subsp. Berberidaceae epigejos N NAT Mahonia aquifolium (Pursh) Nutt. N NAT Ceratochloa cathartica (Vahl) Herter Cynodon dactylon (L.) Pers. Ranunculaceae Dactylis glomerata L. subsp. glomerata Clematis vitalba L. Danthonia decumbens (L.) DC. subsp. Ficaria verna Huds. subsp. verna decumbens Ranunculus acris L. subsp. acris N NAT Dichanthelium acuminatum (Sw.) Gould & Ranunculus bulbosus L. C.A.Clark subsp. implicatum (Scribn.) Freckmann & Lelong Papaveraceae C Digitaria sanguinalis (L.) Scop. Chelidonium majus L. Echinochloa crus-galli (L.) P.Beauv. subsp. Fumaria officinalisL. subsp. wirtgenii crus-galli (W.D.J.Koch) Arcang. Elymus repens (L.) Gould subsp. repens C Papaver rhoeas L. subsp. rhoeas N INV Eragrostis pectinacea (Michx.) Nees Festuca bromoides L. Crassulaceae Festuca danthonii Asch. & Graebn. subsp. Sedum sexangulare L. danthonii Festuca filiformisPourr. Saxifragaceae Festuca lachenalii (C.C.Gmel.) Spenn. Saxifraga tridactylites L. Festuca muralis Kunth Festuca myuros L. subsp. myuros Vitaceae Festuca rubra L. subsp. juncea (Hack.) N INV Parthenocissus inserta (A.Kern.) Fritsch K.Richt. N CAS Parthenocissus tricuspidata (Siebold & Zucc.) Festuca rubra L. subsp. rubra Planch. Festuca stricta Host subsp. trachyphylla N NAT Vitis ×bacoi Ardenghi, Galasso & Banfi (Hack.) Patzke ex Pils N INV Vitis ×koberi Ardenghi, Galasso, Banfi & Holcus lanatus L. subsp. lanatus Lastrucci Holcus mollis L. subsp. mollis N NAT Vitis ×novae-angliae Fernald Hordeum murinum L. subsp. leporinum (Link) Arcang. Fabaceae Koeleria pyramidata (Lam.) P.Beauv. N CAS Albizia julibrissin Durazz. Lolium arundinaceum (Schreb.) Darbysh. Astragalus glycyphyllos L. subsp. arundinaceum Cytisus nigricans L. subsp. nigricans Lolium multiflorumLam. Cytisus scoparius (L.) Link subsp. scoparius Lolium perenne L. Ervilia hirsuta (L.) Opiz Molinia arundinacea Schrank A NAT Galega officinalis L. N INV Muhlenbergia schreberi J.F.Gmel. Genista germanica L. N INV Panicum dichotomiflorum Michx. Genista tinctoria L. N NAT Paspalum dilatatum Poir. Hippocrepis comosa L. subsp. comosa Phleum pratense L. subsp. pratense Lathyrus sylvestris L. subsp. sylvestris Phragmites australis (Cav.) Trin. ex Steud. Lotus corniculatus L. subsp. corniculatus subsp. australis A NAT Medicago sativa L. Poa angustifolia L. Onobrychis viciifolia Scop. Poa annua L. Ononis spinosa L. subsp. spinosa Poa bulbosa L. subsp. bulbosa N INV Pueraria lobata (Willd.) Ohwi Poa compressa L. N INV Robinia pseudoacacia L. Poa pratensis L. subsp. pratensis N INV Robinia viscosa Vent. Poa trivialis L. Securigera varia (L.) Lassen N NAT Setaria italica (L.) P.Beauv. subsp. pycnocoma Trifolium arvense L. subsp. arvense (Steud.) de Wet Trifolium campestre Schreb. C Setaria italica (L.) P.Beauv. subsp. viridis (L.) Trifolium pratense L. subsp. pratense Thell. Trifolium repens L. Setaria pumila (Poir.) Roem. & Schult. Trigonella alba (Medik.) Coulot & Rabaute A INV Sorghum halepense (L.) Pers. Trigonella officinalis(L.) Coulot & Rabaute N NAT Sporobolus neglectus Nash Vicia angustifolia L. N INV Sporobolus vaginiflorus (Torr. ex A.Gray) Vicia cracca L. Alph.Wood Vicia villosa Roth Tragus racemosus (L.) All. A CAS Triticum aestivum L. subsp. aestivum Polygalaceae Polygala vulgaris L. subsp. vulgaris Vascular flora of Milan Malpensa airport (Lombardy, Italy). Part I: checklist 7
Rosaceae Carpinus betulus L. Agrimonia eupatoria L. subsp. eupatoria Corylus avellana L. N CAS Chaenomeles speciosa (Sweet) Nakai Drymocallis rupestris (L.) Soják Celastraceae Filipendula vulgaris Moench Euonymus europaeus L. Fragaria moschata Weston Fragaria vesca L. subsp. vesca Oxalidaceae Potentilla argentea L. N NAT Oxalis articulata Savigny Notes: Some plants could correspond to the hybrid N INV Oxalis dillenii Jacq. with P. intermedia. Note: Sono presenti alcune piante che potrebbero corrispondere all’ibrido con P. intermedia. Violaceae Potentilla argentea L. × Potentilla verna L. Viola alba Besser subsp. dehnhardtii (Ten.) Potentilla erecta (L.) Raeusch. W.Becker Potentilla inclinata Vill. Viola arvensis Murray subsp. arvensis N INV Potentilla indica (Andrews) Th.Wolf Viola canina L. subsp. canina N CAS Potentilla intermedia L. Viola odorata L. Potentilla pusilla Host Potentilla recta L. subsp. recta Salicaceae Potentilla reptans L. Populus alba L. Poterium sanguisorba L. subsp. sanguisorba Populus canescens (Aiton) Sm. A NAT Prunus cerasifera Ehrh. Populus nigra L. subsp. nigra A NAT Prunus domestica L. Populus tremula L. A CAS Prunus persica (L.) Batsch Salix alba L. N INV Prunus serotina Ehrh. Salix caprea L. Prunus spinosa L. subsp. spinosa Salix purpurea L. subsp. purpurea Rubus caesius L. Rubus praecox Bertol. Hypericaceae Rubus sect. Corylifolii Lindl. N CAS Hypericum calycinum L. N NAT Spiraea salicifolia L. Hypericum humifusum L. Hypericum perforatum L. subsp. veronense (Schrank) Ces. Rhamnaceae Frangula alnus Mill. subsp. alnus Euphorbiaceae Euphorbia cyparissias L. Ulmaceae Euphorbia esula L. subsp. esula Ulmus minor Mill. subsp. minor Euphorbia helioscopia L. subsp. helioscopia A NAT Euphorbia lathyris L. Cannabaceae N INV Euphorbia maculata L. N INV Humulus japonicus Siebold & Zucc. Humulus lupulus L. Geraniaceae Erodium cicutarium (L.) L’Hér. Moraceae Geranium columbinum L. Ficus carica L. Geranium molle L. A NAT Morus alba L. Onagraceae Chamaenerion dodonaei (Vill.) Schur ex Fuss Urticaceae N NAT Oenothera biennis L. Parietaria judaica L. N NAT Oenothera chicaginensis de Vries ex Renner Urtica dioica L. subsp. dioica & Cleland Urtica urens L. N INV Oenothera stucchii Soldano
Fagaceae Lythraceae Castanea sativa Mill. Lythrum salicaria L. Quercus robur L. subsp. robur Quercus rubra L. Sapindaceae Acer campestre L. Juglandaceae Acer pseudoplatanus L. C Juglans regia L. Simaroubaceae Betulaceae N INV Ailanthus altissima (Mill.) Swingle Betula pendula Roth 8 Marco Martignoni, Enrico Banfi, Gabriele Galasso
Cistaceae Atocion armeria (L.) Raf. Helianthemum nummularium (L.) Mill. Atocion rupestre (L.) Oxelman subsp. obscurum (Čelak.) Holub Cerastium arvense L. subsp. arvense Cerastium glomeratum Thuill. Malvaceae Cerastium holosteoides Fr. Malva alcea L. Cerastium ligusticum Viv. Malva moschata L. Cerastium semidecandrum L. Malva sylvestris L. Dianthus armeria L. subsp. armeria Petrorhagia prolifera (L.) P.W.Ball & Resedaceae Heywood Reseda lutea L. subsp. lutea Petrorhagia saxifraga (L.) Link subsp. saxifraga Brassicaceae Psammophiliella muralis (L.) Ikonn. Alliaria petiolata (M.Bieb.) Cavara & Grande Sabulina tenuifolia (L.) Rchb. subsp. Arabidopsis thaliana (L.) Heynh. tenuifolia Barbarea vulgaris R.Br. Sagina apetala Ard. subsp. apetala A NAT Brassica napus L. Saponaria officinalis L. Capsella bursa-pastoris (L.) Medik. subsp. Scleranthus perennis L. subsp. perennis bursa-pastoris Scleranthus polycarpos L. Cardamine hirsuta L. Silene gallica L. Diplotaxis tenuifolia (L.) DC. Silene latifolia Poir. Draba verna L. s.l. Silene nutans L. subsp. insubrica (Gaudin) Notes: The observed individuals are not attributable Soldano for sure to any of the subspecies reported by Barto- Silene vulgaris (Moench) Garcke subsp. lucci et al. ������������������������������������������(2018), which we believe are devoid of sy- stematic significance. vulgaris Note: Gli individui presenti non sono attribuibili con Spergula arvensis L. sicurezza a nessuna delle sottospecie riportate da Bar- Stellaria media (L.) Vill. subsp. media tolucci et al. (2018), che secondo noi sono prive di si- gnificato sistematico. Amaranthaceae Erucastrum nasturtiifolium (Poir.) N INV Amaranthus hybridus L. subsp. hybridus O.E.Schulz subsp. nasturtiifolium N NAT Lepidium heterophyllum Benth. Chenopodiaceae N INV Lepidium virginicum L. subsp. virginicum Chenopodium album L. subsp. album Microthlaspi perfoliatum (L.) F.K.Mey. Dysphania botrys (L.) Mosyakin & Clemants Raphanus raphanistrum L. subsp. N NAT Dysphania pumilio (R.Br.) Mosyakin & raphanistrum Clemants N NAT Rorippa armoracioides (Tausch) Fuss N NAT Rorippa austriaca (Crantz) Besser Phytolaccaceae Rorippa sylvestris (L.) Besser subsp. sylvestris N INV Phytolacca americana L. Teesdalia nudicaulis (L.) R.Br. Turritis glabra L. Portulacaceae C Portulaca nitida (Danin & H.G.Baker) Ricceri Santalaceae & Arrigoni Thesium linophyllon L. Primulaceae Polygonaceae Lysimachia arvensis (L.) U.Manns & Anderb. Fallopia convolvulus (L.) Á.Löve subsp. arvensis Persicaria lapathifolia (L.) Delarbre subsp. lapathifolia Ericaceae Persicaria lapathifolia (L.) Delarbre subsp. Calluna vulgaris (L.) Hull pallida (With.) Á.Löve Polygonum arenastrum Boreau Rubiaceae N INV Reynoutria bohemica Chrtek & Chrtková Cruciata glabra (L.) C.Bauhin ex Opiz Rumex acetosa L. subsp. acetosa Cruciata pedemontana (Bellardi) Ehrend. Rumex acetosella L. subsp. acetosella Galium aparine L. Rumex acetosella L. subsp. acetoselloides Galium mollugo L. (Balansa) Den Nijs Galium parisiense L. Rumex obtusifolius L. subsp. obtusifolius Galium rubrum L. Galium verum L. subsp. verum Caryophyllaceae Sherardia arvensis L. Arenaria serpyllifolia L. subsp. serpyllifolia Vascular flora of Milan Malpensa airport (Lombardy, Italy). Part I: checklist 9
Gentianaceae Mentha spicata L. Centaurium erythraea Rafn subsp. erythraea Prunella laciniata (L.) L. Prunella vulgaris L. subsp. vulgaris Apocynaceae Salvia pratensis L. subsp. pratensis N NAT Asclepias syriaca L. Teucrium scorodonia L. Vinca minor L. Thymus oenipontanus Heinr.Braun ex Borbás Vincetoxicum hirundinaria Medik. subsp. Ziziphora acinos (L.) Melnikov subsp. acinos hirundinaria Orobanchaceae Convolvulaceae Euphrasia cisalpina Pugsley Convolvulus arvensis L. Convolvulus sepium L. Verbenaceae N INV Cuscuta campestris Yunck. Verbena officinalisL.
Solanaceae Campanulaceae Solanum nigrum L. Campanula rapunculoides L. subsp. rapunculoides Boraginaceae Campanula rapunculus L. Echium vulgare L. subsp. vulgare Myosotis ramosissima Rochel subsp. Asteraceae ramosissima Achillea millefolium L. s.l. Notes: Populations are difficult to ascribe; according Oleaceae to Valentino Casolo (in litt.), they would partially N CAS Forsythia viridissima Lindl. correspond to A. collina (Becker ex Wirtg.) Heimerl, Fraxinus excelsior L. subsp. excelsior A. millefolium subsp. millefolium, A. collina × Fraxinus ornus L. subsp. ornus A. millefolium, and A. collina × A. setacea. Note: Le popolazioni sono difficilmente attribuibili; N INV Ligustrum sinense Lour. secondo Valentino Casolo (in litt.) sarebbero parzialmente corrispondenti ad A. collina (Becker ex Plantaginaceae Wirtg.) Heimerl, A. millefolium subsp. millefolium, A. Anarrhinum bellidifolium (L.) Willd. collina × A. millefolium e A. collina × A. setacea. Linaria angustissima (Loisel.) Borbás Achillea setacea Waldst. & Kit. Linaria pelisseriana (L.) Mill. Notes: According to Valentino Casolo (in litt.), some Plantago lanceolata L. plants could correspond to hybrids with A. roseoalba Ehrend. and with A. collina (Becker ex Wirtg.) Heimerl. Plantago major L. Note: Secondo Valentino Casolo (in litt.) alcune piante Veronica arvensis L. potrebbero corrispondere a ibridi con A. roseoalba N INV Veronica filiformis Sm. Ehrend. e con A. collina (Becker ex Wirtg.) Heimerl. Veronica hederifolia L. N INV Ambrosia artemisiifolia L. Veronica officinalisL. Anthemis arvensis L. subsp. arvensis Arctium minus (Hill) Bernh. Scrophulariaceae N INV Artemisia annua L. N INV Buddleja davidii Franch. N INV Artemisia verlotiorum Lamotte Scrophularia canina L. Artemisia vulgaris L. Scrophularia nodosa L. Bellis perennis L. Verbascum densiflorum Bertol. N INV Bidens frondosa L. Verbascum lychnitis L. Carduus nutans L. subsp. leiophyllus Verbascum nigrum L. (Petrovič) Stoj. & Stef. Verbascum phlomoides L. Centaurea deusta Ten. Centaurea jacea L. subsp. gaudinii (Boiss. & Lamiaceae Reut.) Gremli Ajuga genevensis L. Centaurea nigrescens Willd. subsp. Ajuga reptans L. pinnatifida(Fiori) Dostál Betonica officinalisL. Chondrilla juncea L. Clinopodium nepeta (L.) Kuntze subsp. Cichorium intybus L. nepeta Cirsium arvense (L.) Scop. Clinopodium vulgare L. subsp. vulgare Cirsium vulgare (Savi) Ten. subsp. vulgare Glechoma hederacea L. Crepis capillaris (L.) Wallr. Lamium maculatum L. Crepis foetida L. subsp. foetida Lamium purpureum L. Crepis setosa Haller f. Mentha ×piperita L. nothosubsp. piperita N INV Erigeron annuus (L.) Desf. (= M. aquatica L. subsp. aquatica × M. N INV Erigeron canadensis L. spicata L.) Eupatorium cannabinum L. subsp. cannabinum 10 Marco Martignoni, Enrico Banfi, Gabriele Galasso
N INV Helianthus tuberosus L. ticular Davide Pisoni and Elena Portone. For the identifi- Hieracium umbellatum L. subsp. umbellatum cation of the critical groups are indebted to Nicola M.G. Hypochaeris maculata L. Ardenghi (Festuca, Vitis), Fabrizio Bartolucci (Thymus), Hypochaeris radicata L. Valentino Casolo (Achillea), Günter Gottschlich (Pilosel- Lactuca sativa L. subsp. serriola (L.) Galasso, la), Thomas Gregor (Potentilla p.p.), and Adriano Soldano Banfi, Bartolucci & Ardenghi (Oenothera). We are also grateful to Enzo Meneguzzo for Leontodon hispidus L. subsp. hispidus reporting Anisantha diandra that he noticed from a shuttle Leucanthemum ircutianum DC. subsp. bus while boarding a plane. ircutianum Logfia minima(Sm.) Dumort. Matricaria chamomilla L. References Picris hieracioides L. subsp. hieracioides Banfi E. & Galasso G. (eds.), 2010 – La flora esotica lombarda. Museo di Storia Naturale di Milano, Milano. (+ Cd-rom). Pilosella officinarumVaill. Bartolucci F., Peruzzi L., Galasso G., Albano A., Alessandrini A., Pilosella piloselloides (Vill.) Soják subsp. Ardenghi N.M.G., Astuti G., Bacchetta G., Ballelli S., Banfi piloselloides E., Barberis G., Bernardo L., Bouvet D., Bovio M., Cecchi L., Pilosella velutina (Hegetschw.) F.W.Schultz & Di Pietro R., Domina G., Fascetti S., Fenu G., Festi F., Foggi Sch.Bip. B., Gallo L., Gottschlich G., Gubellini L., Iamonico D., Iberite M., Jiménez-Mejías P., Lattanzi E., Marchetti D., Martinetto E., Scorzoneroides autumnalis (L.) Moench Masin R.R., Medagli P., Passalacqua N.G., Peccenini S., Pennesi N INV Senecio inaequidens DC. R., Pierini B., Poldini L., Prosser F., Raimondo F.M., Roma- Senecio vulgaris L. subsp. vulgaris Marzio F., Rosati L., Santangelo A., Scoppola A., Scortegagna S., N INV Solidago gigantea Aiton Selvaggi A., Selvi F., Soldano A., Stinca A., Wagensommer R.P., Sonchus asper (L.) Hill subsp. asper Wilhalm T. & Conti F., 2018 – An updated checklist of the vascular flora native to Italy. Plant Biosystems, 152 (2): 179-303.