Pterosaur Cladogram 251 taxa - 183 characters - Peters 2020
Huehuecuetzpalli Macrocnemus BES SC111 96 Macrocnemus T4822 100 Macrocnemus T2472 <50 Dinocephalosaurus Amotosaurus 78 84 Jianchangnathus Fuyuansaurus 85 79 Sordes 2585 3 97 100 Tanystropheus MSNM BES SC1018 Skye Middle Jurassic pterosaur Tanystropheus T/2819 81 76 Pterorhynchus Langobardisaurus 89 Changchengopterus PMOL Tanytrachelos 78 Wukongopterus 90 Cosesaurus 95 Hongshanopterus Kyrgyzsaurus 97 98 Archaeoistiodactylus 100 Kunpengopterus Sharovipteryx <50 Darwinopterus AMNH M8802 62 Longisquama 81 97 Darwinopterus modularis ZMNH M 8782 93 97 59 Darwinopterus robustodens 41H111-0309A 94 Bergamodactylus MPUM 6009 87 Raeticodactylus 100 Darwinopterus linglongtaensis IVPP V 16049 Darwinopterus YH2000 94 Austriadactylus SMNS 56342 Seazzadactylus 97 Scaphognathus crassirostris Austriadactylus SC332466 Scaphognathus SMNS 59395 68 Scaphognathus Maxberg sp. 67 91 Preondactylus TM 13104 MCSNB 2887 83 81 79 Gmu10157 Caelestiventus 98 BM NHM 42735 92 Dimorphodon macronyx Peteinosaurus Ex3359 100 100 BSp 1986 XV 132 89 Carniadactylus ELTE V 256-Pester specimen 78 97 89 MCSNB 8950 99 B St 1936 I 50 (n30) 89 70 Dimorphodon? weintraubi 97 Cycnorhamphus 86 IVPP V13758 embryo 99 99 Moganopterus Mesadactylus holotype Feilongus
94 55 Yixianopterus Dendrorhynchoides curvidentatus Mimodactylus 99 73 77 Dendrorhynchoides mutoudengensis 100 Haopterus SMNS 81928 flathead 100 JZMP embryo 97 Discodactylus NJU-57003 88 97 Boreopterus 85 Vesperopterylus Zhenyuanopterus 72 95 100 98 Anurognathus 98 Hamipterus JPM 2012-001 Arthurdactylus 96 96 CAG IG 02-81 SMNK PAL 3854 95 PIN 2585/4 flightless anurognthid 86 60 Ikrandraco 97 Batrachognathus 98 79 98 Coloborhynchus spielbergi 100 Daohugoupterus Criorhynchus Ferrodraco Jeholopterus Nurhachius BPMC 0204 82 Eudimorphodon ranzii 100 87 74 Arcticodactylus cromptonellus 56 Nurhachius IVPP V-13288 Austriadraco BSp 1994 I51 64 Istiodactylus SMNS 1136 PAL Lonchodraco Campylognathoides HLZ 50 97 Ornithocephalus 99 100 Campylognathoides zitteli SMNS 11879 97 SOS4593 89 Brasileodactylus Campylognathoides zitteli SMNS 9787 100 SOS4006 92 Ludodactylus Nesodactylus 98 91 Guidraco Campylognathoides liasicus CM 11424 SMNS 81775 Cearadactylus ligabuei 82 89 74 Cearadactylus atrox 87 Rhamphorhynchus St/Ei 8209 (n28) Painten pteroaur 76 85 82 94 Liaoningopterus Rhamphorhynchus B St 1960 I 470A (n9) Douzhanopterus Anhanguera piscator 100 Bellubrunnus 86 82 100 Jianchangopterus 95 Rhamphorhynchus MBH unnumbered (n20) Qinglongopterus Ningchengopterus 80 86 Rhamphorhynchus BMNH 37012 (n85) 87 Rhamphorhynchus B St 1959 I 400 (n10) 90 97 82 Pterodactylus AMNH 1942 75 Rhamphorhynchus TM 6924 (n1) 82 Pterodactylus SMF 405 (n37) 100 51 94 Pterodactylus Tokyo sp. 95 Rhamphorhynchus B St 1889 XI 1 (n7) Rhamphorhynchus B St 1938 I 503a (n11) 90 Pterodactylus MBR 3655 72 Rhamphorhynchus ROM 55352 69 Pterodactylus NHMW 1975/1756 Pterodactylus scolapaciceps BSP1937 I 18 (n21) 77 R sp Lauer collection Pterodactylus Lauer collection 96 56 Pterodactylus BSP AS V 29a 98 Rhamphorhynchus HMNS 1862 69 Pterodactylus antiquus BSP AS I 739 (n4) 70 Rhamphorhynchus Imhof specimen 69 94 Luchibang MBR5591.1 100 Rhamphorhynchus GPIT RE 7321 (n81) 83 Pterodactylus BMMS 7 98 Rhamphorhynchus NHMW 1998z0077/0001 90 Pterodactylus longicollum SMNS 56603 Rhamphorhynchus BMNH 37002 (n82) 81 55 100 Rhamphorhynchus WDC CSG 255 Rhamphorhynchus BRI 010 69 BSt 1967 i 276 BMNH 42736 80 Rhamphorhynchus TMP2008.41.001 68 50 Rhamphorhynchus JME SOS 4785 BM aM 4072 69 100 BSt ASXIX3 Rhamphorhynchus BSPG 1880 II 8 Zittel wing 100 Rhamphorhynchus HMNS-2 Altmeuhlopterus rhamphastinus 90 92 Cuspicephalus 96 Rhamphorhynchus MTM V2008.33.1 Rhamphorhynchus WU970001 94 Germanodactylus JME MOE 12 86 Eosipterus GMV 2117 86 <50 89 Wenupteryx MOZ 3625 87 Rhamphorhynchus CM 11427 (n59) SMNK PAL 3830 98 Rhamphorhynchus YPM 1778 (n33) 88 75 Rhamphorhynchus JME SOS 4009 (n62) Germanodactylus cristatus BSt 1892 IV 1(n61) 96 Elanodactylus Rhamphorhynchus TM6920/21 (n38) 79 84 Germandactylus SMNK-PAL6592 Rhamphorhynchus SMF R 4128 (n43) 85 B St 1878 VI 1 63 Rhamphorhynchus TM 6922/6923 (n74) 83 85 92 Rhamphorhynchus GPIH MYE 13 (n75) Rhamphorhynchus B St 1929 I 69 (n52) 96 Nemicolopterus 79 Mistralazhdarcho Changchengopterus CYGB-0036 70 Shenzhoupterus 95 Noripterus 94 Orientognathus Dsungaripterus Sordes PIN 2585-25 89 87 Phobetor 65 92 Sinopterus 60 Tupandactylus 95 95 97 Tapejara 73 Huaxiapterus 54 Sordes PIN 2470-1 98 Sinopterus liui Cacibuteryx 56 Tupuxura longicristatus 55 Dorygnathus Hauff sp. 82 83 Tupuxura Goshura sp. Dorygnathus Vienna sp. Thalassodromeus 71 Dorygnathus MBR 1920.16 60 Eopteranodon 75 Sericipterus 66 <50 Aurorazhdarcho? MB.R.3531 Dorygnathus SMNS 55886 51 <50 Eoazhdarcho 76 Dorygnathus Up R 156 Aurorazhdarcho NMB Sh 110 95 57 Dorygnathus purdoni cf. Tupuxuara Elgin 2014 96 Angustinaripterus 68 97 St/E1I 81 Muzquizopteryx 94 MBR 3530 (n40) 79 Nyctosaurus FHSMVP21 100 JPM 2014-004 [not Gladocephaloides] 71 72 Nyctosaurus FMNH25026 Nyctosaurus UNSM93000 73 AMNH 5147 100 81 Gnathosaurus 56 Liaodactylus Pteranodon AMNH 4908 no skull 81 Pteranodon YPM 1179 skull Ctenochasma elegans (n45) 95 81 Pteranodon FHSM 2183 C elegans Lauer collection 69 C elegans Fossilenatlas.de204 Pteranodon UUPI R197 94 50 Pteranodon YPM 1177 59 Pteranodon KUVP 2212 skull 88 70 Gegepterus Leptostomia Gladocephaloideus IG-CAGS-08-07 74 <50 Pteranodon YPM 2594 85 Ctenochasma (n65 B) 87 Pteranodon CMC VP7203 85 <50 Allkaruen 66 Pteranodon FHSM 2851 89 D2514 [not Eosipterus] Pteranodon UNSM 12167 <50 Pteranodon NMC41 358 99 Pterodaustro <50 Pterodaustro embryo 65 Pteranodon UALVP 24238 95 Pteranodon KUVP 27821 Pteranodon FHSM VP339 86 Dorygnathus SMNS51827 Dorygnathus SMNS 50164 99 TM 10341 (n1) 99 Rhamphodactylus BSPG 2011 I 133 100 Beipiaopterus 60 74 Ardeadactylus longicollum 89 Forfexopterus Bootstrap scores are based on testing 90 99 Huanhepterus Mesadactylus BYU specimen CM 11 426 (n44) overlapping subsets 87 B St 1911 I 31 (n42) 57 85 SOS2428 (n57) SOS 2179 due to limitations in computing power. 90 Jidapterus 95 Chaoyangopterus LPM-R00076 70 Microtuban 97 Chaoyangopterus holotype 100 Zhejiangopterus Quetzalcoatlus