Late Campanian Nautiloids from Deir Abu Said, North-Western Jordan

Cretaceous Research 102 (2019) 81e88

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Cretaceous Research

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Short communication Late Campanian nautiloids from Deir Abu Said, north-western Jordan

* Markus Wilmsen a, , Mohamed Fouad Aly b a Senckenberg Naturhistorische Sammlungen Dresden, Museum für Mineralogie und Geologie, Sektion Palaozoologie,€ Konigsbrücker€ Landstraße 159, 01109, Dresden, Germany b Geology Department, Faculty of Science, Cairo University, Giza, Egypt article info abstract

Article history: A small faunule of nautiloids is described from the upper Campanian Bahiya Coquina Member of the Al- Received 21 February 2019 Hisa Phosphorite Formation of north-western Jordan. It consists of Eutrephoceras sphaericum (Forbes, Received in revised form 1845), Cimomia desertora (Quaas, 1902) and a specimen assigned to Hercoglossidae indet. in open 10 April 2019 nomenclature. E. sphaericum and C. desertora are reported for the ﬁrst time from Jordan. The present Accepted in revised form 20 April 2019 records further complement the information on the temporal and spatial distribution of nautiloids at the Available online 26 April 2019 southern margin of the Neotethyan Ocean during the late part of the Cretaceous Period. © 2019 Elsevier Ltd. All rights reserved. Keywords: Late Cretaceous Nautiloidea Taxonomy Southern Tethys Palaeobiogeography

1. Introduction Campanian to lower Maastrichtian Al-Hisa Phosphorite Formation that crops out at Deir Abu Said, north-western Jordan (Fig. 1). The Cretaceous nautiloids are commonly long-ranging in their scope of the present short contribution is to enhance the knowl- stratigraphic distribution, morphologically rather limited in edge on the stratigraphic and palaeogeographic distribution of external shell characters and often poorly preserved. Furthermore, nautiloids at the southern margin of the Neotethyan Ocean during they tend to be fairly conservative in terms of evolutionary devel- the late phase of the Cretaceous Period. opment (bradytelic) compared to the contemporaneous, fast- evolving ammonites (e.g., Ward and Signor, 1983). It is also due to 2. Geological setting and stratigraphy these shortcomings that comprehensive modern systematic ac- counts on Cretaceous taxa are very rare. The study area was located at the southeastern margin of the Up to now, there are very few records of nautiloids from Jordan. Neotethyan Ocean at tropical palaeolatitudes (Fig. 1A). Several Bender (1963) and Wolfart (1968) mentioned Lower Silurian nau- compressional features formed during the CampanianeEocene tiloids from southern Jordan, east of the Hejaz railroad. In an un- throughout the eastern Mediterranean when the Afro-Arabian published report, LeBlanc (2019) reported the occurrence of some Plate collided with the intra-oceanic trench to the north and nautiloid fragments and a cephalopod jaw from the Middle Eocene east (Fig. 1A) and almost all of the major phosphorite deposits of (Lutetian) Wadi Shallalah Formation at Wadi Al-Rijla Al-Bayda, the eastern Mediterranean are associated with these compres- Eastern Desert of Jordan. Cretaceous taxa have, to the best of our sional structures (Abed, 2013). There is general agreement that knowledge, only marginally been mentioned in papers dealing the major tectonic pulse of this Syrian Arc system took place with other aspects (e.g., Wiese and Schulze, 2005, p. 944). There- during the CampanianeMaastrichtian (Bowen and Jux, 1987; fore, we document herein a small faunule of rather well-preserved Guiraud and Bosworth, 1997; Walley, 2001; Abed, 2013). nautiloids that has been collected from the lower part of the upper CampanianeMasstrichtian strata are widely distributed in central, southern and northern Jordan (Blanckenhorn, 1903; Masri, 1963; Bender, 1974; Hiyari, 1985; Abed, 2013; Khrewesh et al., 2014; Ahmed et al., 2014). * Corresponding author. ﬁ E-mail addresses: [email protected] (M. Wilmsen), mfalyaly@ Blanckenhorn (1903) was the rst to describe phosphate sci.cu.edu.eg (M.F. Aly). deposits in Jordan, 16 km northwest of Amman. The Al-Hisa https://doi.org/10.1016/j.cretres.2019.04.009 0195-6671/© 2019 Elsevier Ltd. All rights reserved. 82 M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88

Fig. 1. Geologic and stratigraphic framework. A, Campanian palaeogeographic and plate tectonic situation (modiﬁed after Barrier and Vrielynck, 2008). The map area of Fig. 1B is outlined and the nautiloid occurrences according to the records cited in the taxonomic descriptions are indicated. B, Location map of the studied section, north-western Jordan. C, Stratigraphic section of the upper CampanianeMaastrichtian sequence at Deir Abu Said, north-western Jordan. M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88 83

Phosphorite Formation, named after the type area of Al-Hisa in et al. (1996) assigned an early Maastrichtian age whereas Central Jordan (Hiyari, 1985), is equivalent to the upper part of others suggested a late Campanian age (Bender, 1974; Pufahl Calcaires silex (Wetzel and Morton, 1959), the upper part of the et al., 2003; Powell and Moh'd, 2011; Galmed et al., 2013; Amman Formation (Masri, 1963; Parker, 1970) as well as the Abed, 2013). The Al-Hisa Phosphorite Formation was assigned upper part (unit B2b) of the B2 Silicified Limestone and a Campanian age by Wetzel and Morton (1959),and Phosphorite Formation, and the phosphorite member of Bender CampanianeMaastrichtian by Bender (1974). Reiss et al. (1985) (1974). Isolated outcrops of this formation are also known from allocated the reported ammonites Libycoceras sp. and Anaklino- the area of Deir Abu Said which is located north of Amman ceras reflexum Stephenson to the late Campanian (see also Lewy, (Fig. 1B). The Al-Hisa Phosphorite Formation includes three 1986). formal members which are, in ascending order, the Sultani Phosphorite, the Bahiya Coquina, and Qatrana Phosphorite 3. Systematic palaeontology members (Fig. 1C). Lithologically, the Al-Hisa Phosphorite For- mation is composed of alternating beds of phosphatic chert, Cretaceous nautiloids are morphologically fairly poor in diag- phosphatic limestone, phosphate, chalky limestone, micritic nostic external shell features, bradytelic in their macroevolu- limestone, marl, and cross-bedded oyster beds (coquinas). The tionary patterns, and commonly long-ranging. Furthermore, they facies suite was interpreted to be deposited in shallow marine are often poorly preserved, especally in carbonate facies, where subtidal to intertidal environments (Moumani, 2002). The their fossil record is mostly restricted to internal moulds of very collected and described nautiloids come from the Bahiya variable preservational quality (Malchyk et al., 2017). Conse- Coquina Member, which is approximately 6 m thick in the quently, the group is rather poorly known and only a few synoptic study area (Fig. 1C). modern works on their systematics and taxonomy exist (e.g., The Bahiya Coquina Member is characterized by large-scale Baudouin et al., 2016). Important contributions to post-Triassic cross-bedded, graded beds of shelly limestone with abundant nautiloids were presented by Kummel (1956, 1964), Wiedmann well-preserved oysters along with abundant phosphatic layers (1960) and Dzik (1984). Shimansky (1975) and Matsumoto et al. containing the studied nautiloids. The member also contains well- (1984) focused on Cretaceous species and their phylogenetic re- preserved bivalves (mainly oysters), gastropods and ammonoids lations. However, in spite of these important groundworks, clas- (Libycoceras spp. and Baculites sp.) as well as fish teeth and poorly sification at the family and genus levels is still controversially preserved fish fragments. The ranges of the ammonoid taxa provide discussed, especially with respect to ribbed representatives of the the basis for a twofold division of the upper Campanian Bahiya family Cymatoceratidae Spath, 1927 (e.g., see Wilmsen, 2000; Coquina Member with the biozones of Libycoceras spp. (below) and Cichowolski, 2003; Wilmsen and Yazykova, 2003; Wilmsen and Baculites sp. (above, Fig. 1C). The specimens of Libycoceras from Esser, 2004; Frank, 2010; Frank et al., 2013; Machalski and northern Jordan, originally assigned to L. ismaelis (Zittel) by Galmed Wilmsen, 2015; Lehmann et al., 2017). Herein, we follow the et al. (2013), have been re-determined as L. crossense Zaborski and classification of Shimansky (1975) with the exception that we L. afikpoense Reyment based on their sutures and shell morphol- regard the genus Deltoidonautilus Spath, 1927 as a synonym of ogies by Ifrim (2017), supporting a late Campanian age. Well-dated Angulithes de Montfort, 1808. L. ismaelis, however, are inferred to be always derived from Maas- The present paper is only a short communication, thus, the trichtian strata (Ifrim, 2017). The stratigraphic interpretation of the synonymies are limited to a minimum (first reference, regional nautiloid find layer as upper Campanian is supported by an early records, important revisions). Morphological features and terms Maastrichtian age assignment of the overlying Muwaqqar Forma- are used according to Teichert (1964). Biometric measurements of tion (Farouk et al., 2014; Ahmad et al., 2015). maximum diameter (D), whorl breadth (Wb), whorl height (Wh), The age of the phosphorite deposits of Jordan has been and size of umbilicus (U) were obtained using a Vernier Caliper and controversially discussed in the past. Authors such as Burdon are given in mm and % of maximum diameter (in brackets; Fig. 2). (1959), Hamam (1977), Abed and Ashour (1987),andCapetta The material is stored in the collections of the Geological Museum, Cairo University (repository CUGM/JCAM).

Order Nautilida Agassiz, 1847 Superfamily Nautiloidea Blainville, 1825 Family Nautilidae Blainville, 1825 Genus Eutrephoceras Hyatt, 1894. Type species: Nautilus dekayi Morton, 1834 (p. 33, pl. 8, fig. 4). Eutrephoceras sphaericum (Forbes, 1845) Figs. 3A, 4A *1845 Nautilus sphaericus sp. nov., Forbes, p. 98. 1891 Nautilus sphaericus Forbes e Foord, p. 298. 1956 Eutrephoceras sphaericum (Forbes) 1846 e Kummel, p. 383. 1960 Eutrephoceras sphaericum (Forbes) e Wiedmann, p. 170, pl. 18, figs. DeG, pl. 23, figs. G, M, N. 1960 Eutrephoceras sphaericum geinitzi n. ssp. e Wiedmann, p. 171. 1985 Eutrephoceras sphaericum geinitzi Wiedmann, 1960 e Klin- ger, p. 3, text-fig. 3AeF. e Fig. 2. Taxonomically important features and biometric parameters of the planispiral 2016 Eutrephoceras sphaericum (Forbes, 1845) Wilmsen, p. 62, nautiloid shell. pls 4c, 5, text-fig. 2a. 84 M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88

Material. One fully septate internal mould (CUGM/JCAM 49). 1981 Eutrephoceras desertorum (Quaas) e Barthel and Dimensions. Herrmann-Degen, p. 157. 1984 Eutrephoceras desertorum (Quaas) e Dzik, p. 180, fig. 71/5. Specimen D Wb (%) Wh (%) Wb/Wh U (%) 1995 Cimomia aff. sowerbyana (d'Orbigny, 1840) e Morris, p. 252, pl. 1, fig. 4, pl. 2. CUGM/JCAM 49 69 69 (100) 48 (70) 1.44 Occluded 2002 Eutrephoceras desertorum (Quaas) e Hewaidy and Azab, p. 213, pl. 4, figs. 1e4. Description. Globular, strongly depressed (Wb/Wh ¼ 1.44) nau- 2019 Eutrephoceras desertorum (Quaas, 1902) e Hewaidy et al., tilid with occluded umbilicus. During ontogeny, whorl breadth p. 50, text-figs. 4A, B. increases rapidly, forming a globular conch in which Wb is Material. One fully septate internal mould (CUGM/JCAM 50). approximating D. The whorl cross-section is transverse elliptical Dimensions. and the venter broadly rounded. The suture is nearly straight with only weak undulations while the distance of septa is rather low. The position of the siphuncle cannot be seen. Specimen D Wb (%) Wh (%) Wb/Wh U (%) Remarks. Eutrephoceras sphaericum is a very inflated nautilid spe- CUGM/JCAM 50 100 75 (75) 66 (66) 1.14 occluded cies with Wb/D z 1 (e.g., Forbes, 1845; Wiedmann, 1960). In association with the simple suture it can thus be well separated from Description. Slightly depressed (Wb/Wh ¼ 1.14) nautiloid with contemporaneous less inflated nautilids. Our specimen is very close occluded umbilicus. Greatest whorl breadth is just above the in sagittal section to the holotype in Forbes (1845, p. 98, see Fig. 4A). umbilical margin from where the flanks converge towards the Also the Wb/D ratio of 0.7 is very close to data given for this species broadly rounded venter without any ventrolateral shoulder, in the literature (Wiedmann, 1960, p. 170). However, it should be forming a semi-circular whorl cross-section (Fig. 4B). The su- noted that E. dekayi (Morton) is a fairly similar and contempora- ture rises from an umbilical lobe towards a narrow and neous nautilid species in which Wb/D can approach a ratio of 1, too. shallow saddle across the umbilical margin followed by a wide, Mainly known from the upper CampanianeMaastrichtian of the shallow lobe on the flanks. On the venter, a very shallow, Cretaceous of the Western Interior Seaway in North America (e.g., narrow saddle and a somewhat broader lobe are present. In- Sohl and Koch, 1983), the species has also been reported from dividual septa are rather closely spaced (ca. 18 septa in the last northern Africa by Wanner (1902, p. 142) from strata contempo- whorl). The siphuncle has a dorso-central position (Fig. 4B). raneous to the find layers in Jordan, unfortunately without illus- tration. As we did not trace Wanner's specimens, we cannot finally Remarks. The present specimen corresponds well in shell pro- conclude on the taxonomic affinity of his material, and an exami- portions, shape of suture, position of siphuncle and closely spaced nation of a potential specific concordance of E. dekayi and septa to Cimomia desertora (Quaas,1902) as described and illustrated E. sphaericum is beyond the scope of the paper. in the literature (e.g., Wiedmann, 1960). In several papers, the spe- On the basis of a single specimen from the Turonian of Saxony cies has also been assigned to the genus Eutrephoceras (starting with (Germany), Wiedmann (1960, p. 171) introduced a new subspecies, Kummel, 1956) but the rather sinuous suture, the numerous (i.e., Eutrephoceras sphaericum geinitzi n. ssp. However, the differences to densely spaced) septa and the rather dorsal position of the siphuncle the nominate subspecies, E. sphaericum sphaericum, are very subtle suggest a closer affinity with Cimomia (see also Wiedmann, 1960; (shallow external lobe vs. shallow external saddle) and prone to be Shimansky, 1975). According to Kummel (1956, 1964), Cimomia is influenced by taphonomic processes (Wilmsen, 2016). Further- transitional between Eutrephoceras and Hercoglossa. A fairly similar more, we feel that the separation of subspecies in morphologically and broadly contemporaneous taxon has been described as Cimomia poorly differentiated and long-ranging Cretaceous nautiloids is aff. sowerbyana (d'Orbigny, 1840)byMorris (1995, p. 252, pl. 1, fig. 4, pointless in general. Thus, we refrain from separating any subspe- pl. 2) from the lower Maastrichtian Simsima Formation of the United cies in E. sphaericum. Arab EmirateseOman border region. It mainly differs from C. desertora as described herein by means of the more central po- Occurrence. The species ranges from the Cenomanian to the sition of the siphuncle. However, the position of the siphuncle in Maastrichtian (Wiedmann, 1960, fig. 26) and has so far been re- C. desertora changes ontogenetically from dorso-central to ventro- ported from India, Europe, South Africa and possibly Madagascar central (e.g., Quaas, 1902, p. 300) and the specimen from the Sim- (Fig. 1A). The present study is the first documentation of Eutre- sima Formation is larger (D ca. 150 mm). Thus, we regard it as phoceras sphaericum (Forbes) from the upper Campanian of Jordan. conspecifictoC. desertora. Furthermore, Angulithes sowerbyanus (d'Orbigny, 1840) is a Turonian species that is much more com- Family Hercoglossidae Spath, 1927 pressed and has an open umbilicus (Tintant and Gauthier, 2006, pl. Genus Cimomia Conrad, 1866 4, fig. 2a, b). Cimomia imperalis (Sowerby, 1812), mainly known from the Eocene of Europe, has a more sinuous suture but is otherwise Type species: Nautilus burtini Galeotti, 1837, p. 140. very similar shell proportions. Furthermore, a number of Eutrepho- Cimomia desertora (Quaas, 1902) ceras species such as E. bouchardianum (d'Orbigny, 1840)and Figs. 3B, 4B E. dorbignyanum (Forbes, 1846) share similarities to C. desertora as described herein. However, their sutures are simpler and the dis- *1902 Nautilus desertorum Zitt. (in manu.), Quaas, p. 299, pl. 29, tance of the septa is larger (e.g., Wilmsen, 2000, 2016; Nielsen and fig. 1; pl. 33, figs. 29, 30. Salazar, 2011; Machalski and Wilmsen, 2015). 1915 Nautilus desertorum Zitt. e Greco, p. 229, pl. 22, figs. 5, 6. 1956 Eutrephoceras desertorum (Quaas) e Kummel, p. 381. Occurrence. Cimomia desertora has so far been reported from the 1960 Angulithes (Cimomia) desertorum (Zittel) in Quaas 1902 e Santonian of Spain (Wiedmann, 1960), the Maastrichtian to Paleo- Wiedmann, p. 179, pl. 19, figs. J, K, M, ?N; pl. 23, fig. I; pl. cene of Egypt (Western Desert: Quaas, 1902; Barthel and Herrmann- 24, fig. 5. [with synonymies] Degen, 1981; Hewaidy et al., 2019; Eastern Desert: Greco, 1915; ?1975 Cimomia ?desertorus (Zittel in Quaas, 1902) e Shimansky, southwest Aswan: Hewaidy and Azab, 2002), and, as Cimomia aff. p. 135, pl. 30, fig. 1. sowerbyana, from the lower Maastrichtian of the United Arab M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88 85

Fig. 3. Nautiloids from the upper Campanian of Jordan; all ﬁgures in natural size. A, Eutrephoceras sphaericum (Forbes, 1845), specimen CUGM/JCAM 49 in lateral (A1), apertural (A2) and ventral (A3) views. B, Cimomia desertora (Quaas, 1902), specimen CUGM/JCAM 50 in lateral (B1, B3), and apertural (B2) views. C, Hercoglossidae indet., specimen CUGM/JCAM 51 in lateral (C1), apertural (C2) and ventral (C3) views. 86 M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88

Fig. 4. Whorl cross-sections and external sutures of nautiloids from the upper Campanian of Jordan (except D). A, Eutrephoceras sphaericum (Forbes, 1845); in grey cross-section of Nautilus sphaericus in Forbes (1845, p. 98). B, Cimomia desertora (Quaas, 1902). C, Hercoglossidae indet. D, Cimomia jordani (from Wanner, 1902, pl. 19, ﬁgs. 21, 21a).

Emirates (Morris, 1995)(seeFig. 1A). A questionable record is from a flexed suture and the siphuncle is close to the dorsum, and the the uppermost Cretaceous of Crimea (Shimansky, 1975). Herein, triangular whorl section is very similar to specimen CUGM/JCAM Cimomia desertora is firstly recorded from the upper Campanian of 51, albeit somewhat more broadly rounded (Fig. 4D). Another Jordan. similar and broadly contemporaneous taxon with triangular whorl section has been described as Deltoidonautilus salisfilius sp. Hercoglossidae indet. nov. by Morris (1995,p.251,pl.1,figs. 1e3)fromthebaseofthe Figs. 3C, 4C Simsima Formation (probably lower Maastrichtian) of the United Material. One internal mould (CUGM/JCAM 51). Arab EmirateseOman border region. It shares the flexed suture Dimensions. and dorsal position of the siphuncle with C. jordani but seems to be more compressed. However, our specimen cannot be safely identified and the exact taxonomic relationship between Specimen D Wb (%) Wh (%) Wb/Wh U (%) C. jordani and (sic) Angulithes salisfilius (Morris) needs to be CUGM/JCAM 51 55 35 (64) 35 (64) 1.00 occluded evaluated elsewhere. We thus keep specimen CUGM/JCAM 51in open nomenclature. Description. The small specimen (D ¼ 55 mm) is convolute. Occurrence. The specimen is from the Al-Hisa Phosphorite Forma- Greatest whorl breadth is directly at the umbilical margin from tion, Bahiya Coquina Member, upper Campanian. where the flat flanks converge to a slender, rounded venter, resulting in a triangular whorl cross-section with Wb z Wh. 4. Discussion and concluding remarks Due to the poor state of preservation of the studied specimen, the suture lines along with the position of the siphuncle cannot Cretaceous nautiloids from Jordan have not been dealt with in be seen. However, faint growth lines at the venter (see Fig. 3C3) any detail so far. In order to minimize this dark spot in this show the presence of a rather deep hyponomic sinus. palaeobiogeographically important region of the Middle East, we Remarks. The imperfect preservation impedes a specific and even report and describe a small faunule of nautiloids from the upper a safe generic identification of specimen CUGM/JCAM 51. The Campanian Bahiya Coquina Member of the upper Campanian to characteristic triangular whorl shape suggests that it may be a lower Maastrichtian Al-Hisa Phosphorite Formation of north- representative of the widespread genus Angulithes which ranges western Jordan. from the Upper Cretaceous into the Eocene. However, Wanner The Bahiya Coquina Member at Deir Abu Said, north-northwest (1902, p. 143, pl. 19, figs. 12, 21a) introduced a new species, Nau- of Amman, is composed of large-scale cross-bedded bioclastic tilus jordani, from the Maastrichtian of western Egypt that has limestone with abundant well-preserved oysters and layers of later been placed in Cimomia by Kummel (1956) and Shimansky phosphatic clasts. It contained the studied nautiloids and was (1975),andinAngulithes (Cimomia)byWiedmann (1960).Ithas deposited in an agitated, shallow-marine environment as shown by M. Wilmsen, M.F. Aly / Cretaceous Research 102 (2019) 81e88 87 the coarse-grain size and fabric of the strata as well as the large- Ahmed, F., Ifrim, C., Farouk, S., 2015. The last ammonoids of Jordan e implications € scale cross-bedding. Despite the unfavorable taphonomic condi- from the Outhriate section. Neues Jahrbuch für Geologie und Palaontologie, Abhandlungen 278, 245e256. tions prone to physical destruction of organic remains, the pres- Barrier, E., Vrielynck, B., 2008. Map 7: Early Campanian (80.6e83.5 Ma). In: ervation of the nautiloid specimens is moderate to good. The small Barrier, E., Vrielynck, B. (Eds.), Palaeotectonic maps of the Middle East e tec- e faunule (three specimens) consists of Eutrephoceras sphaericum tono-sedimentary palinsspastic maps from the Late Norian to Pliocene. Com- mission for the Geological Map of the World. CGMW/CCGM), Paris. (Forbes, 1845), Cimomia desertora (Quaas, 1902) and a specimen Barthel, K.W., Herrmann-Degen, W., 1981. Late Cretaceous and Early Tertiary stra- assigned in open nomenclature to the family Hercoglossidae. Both tigraphy in the Great Sand Sea and its SE margins (Farafra and Dakhla oases), specifically identified taxa, i.e., E. sphaericum and C. desertora,are SW Desert, Egypt. Mitteilungen der Bayerischen Staatssammlung für € e fi Palaontologie und historische Geologie 21, 141 182. reported for the rst time from Jordan. E. sphaericum is a very Baudouin, C., Delanoy, G., Moreno-Bedmar, J.A., Pictet, A., Vermeulen, J., Conte, G., inflated species (Figs. 3A, 4A) that, in association with its simple Gonnet, R., Boselli, P., Boselli, M., 2016. Revision of the Early Cretaceous genera suture, can be readily separated from contemporaneous nautilids. It Heminautilus Spath, 1927, and Josanautilus Martínez & Grauges, 2006 (Nautilida, Cenoceratidae). Carnets de Geologie 16, 61e212. ranges throughout the Upper Cretaceous and has so far been re- Bender, F., 1963. Geology of the Arabian Peninsula. Jordan. U.S. Geological Survey ported from India, Europe, South Africa and possibly Madagascar Professional Paper 560-I, ievi, I1eI36. (see Fig. 1A, e.g., Wiedmann, 1960). C. desertora is a less depressed Bender, F., 1974. Geology of Jordan, 196pp., Berlin (Borntraeger). e species with more sinuous suture, closely spaced septa and a rather de Blainville, H.M.D., 1825 27. Manuel de malacologie et de conchyliologie, 664pp. (1825), 87 pls. (1827). Levrault, Paris and Strasbourg. dorsal position of the siphuncle (Figs. 3B, 4B) that has often been Blanckenhorn, M., 1903. Die Mineralschatze€ Palastinas.€ Mitteilungen und Nach- placed in the genus Eutrephoceras by other authors (e.g., Kummel, richten des Deutsch-Palastinensischen€ Vereins 1902, 65e70. 1956; Hewaidy et al., 2019). However, the above mentioned Bowen, R., Jux, U., 1987. Afro-Arabian Geology. Chapman and Hall, London, 295 pp. Burdon, D.J., 1959. Handbook of the geology of Jordan. Government of Jordan, morphological characters suggest a close affinity with the genus Amman, 82 pp. Cimomia Conrad, 1866 (see genus definition in Kummel, 1964: Capetta, H., Pfeil, F., Schmidt-Kittler, N., Martini, E., 1996. New biostratigraphic data K456). C. desertora has so far been reported from the Santonian to on the marine Upper Cretaceous and Paleogene of Jordan. Unpublished report. Jordan Phosphate Mining Company, Amman. Paleocene of Spain, Egypt, the United Arab Emirates and possibly Cichowolski, M., 2003. The nautiloid genus Cymatoceras from the Cretaceous of the Crimea (Fig. 1A). The third compressed nautiloid specimen can only Neuquen and Austral basins, Argentina. Cretaceous Research 24, 375e390. be assigned to the family Hercoglossidae but its characteristic Conrad, T.A., 1866. Observations on recent and fossil shells, with proposed new genera and species. American Journal of Conchology 2, 101e103. triangular whorl shape (Figs. 3C, 4C) suggests that it may be a Dzik, J., 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica 45, 1e220. representative of the widespread Late Cretaceous to Eocene genus Farouk, S., Marzouk, A.M., Ahmad, F., 2014. The Cretaceous/Paleogene boundary in Angulithes. 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