Turkish Journal of Zoology Turk J Zool (2016) 40: 40-56 http://journals.tubitak.gov.tr/zoology/ © TÜBİTAK Research Article doi:10.3906/zoo-1501-38

Contribution to the knowledge of the () fauna of from northeastern Anatolia, Part I

1 2, 2 Janko KOLAROV , Saliha ÇORUH *, İrfan ÇORUH 1 Faculty of Pedagogy, University of Plovdiv, Plovdiv, 2 Department of Protection, Faculty of Agriculture, Atatürk University, , Turkey

Received: 20.01.2015 Accepted/Published Online: 05.08.2015 Final Version: 01.01.2016

Abstract: This present contribution is based upon the ichneumonids collected from the northeastern Anatolian part of Turkey in 2014. A total of 57 species belonging to subfamilies Acaenitinae, , , , Cremastinae, Cryptinae, , , Metopiinae, Orthocentrinae, Oxytorinae, , and Tryphoninae have been recorded. Among them, Agrypon clandestinum (Gravenhorst), Lissonota (Lissonota) nitida (Gravenhorst), Campoletis agilis (Holmgren), Diadegma combinatum (Holmgren), Olesicampe cavigena (Thomson),Olesicampe patellana (Thomson), Aptesis assimilis (Gravenhorst), Schreineria populnea (Giraud), Xylophrurus lancifer (Gravenhorst), Diplazon pallicoxa Manukyan, Homotropus pallipes (Gravenhorst), Woldstedtius biguttatus (Gravenhorst), and Oxytorus luridator (Gravenhorst) are new records for the Turkish fauna. New data on the distribution of 40 known species were added. Additionally, a zoogeographical characterization is given for each of the species.

Key words: Hymenoptera, Ichneumonidae, new records, fauna, zoogeographical characterization, Turkey

1. Introduction 2002; Kolarov et al., 2002a, 2002b; Yurtcan and Beyarslan, Ichneumonids belong to the division , 2002; Özbek et al., 2003; Çoruh et al., 2004, Kolarov and which includes the superfamilies , Gürbüz, 2004; Çoruh et al., 2005a, 2005b; Çoruh and Chalcidoidea, , and Proctotrupoidea (Grzimek Özbek 2005; Gürbüz, 2005; Gürbüz and Aksoylar, 2005; et al., 2004). Gürbüz and Kolarov, 2005; Yurtcan and Beyarslan, 2005; Ichneumonid are important parasites of other Gürbüz and Kolarov, 2006a, 2006b; Kolarov and Gürbüz, . There are 2679 genera and 44,385 species in the 2006; Yurtcan and Beyarslan, 2006; Yurtcan et al., 2006; world. The family Ichneumonidae has a cosmopolitan Çoruh et al., 2007; Kolarov and Gürbüz, 2007a, 2007b; distribution, with 51 generally recognized subfamilies, Okyar and Yurtcan, 2007; Yurtcan, 2007; Boncukcu, 1579 genera, and 24,281 described species (Yu et al., 2012). 2008; Çoruh, 2008; Çoruh and Kesdek, 2008; Çoruh and The distribution of Ichneumonidae is one of the most Özbek, 2008a, 2008b; Çoruh and Tozlu, 2008; Gürbüz notable exceptions to the common latitudinal gradient in and Kolarov, 2008; Kırtay, 2008; Kolarov and Çoruh, species diversity because it shows greater speciation at high 2008; Kolarov and Yurtcan, 2008; Yurtcan and Okyar, latitudes than at low latitudes (Sime and Brower, 1998). 2008; Çoruh, 2009; Gürbüz et al., 2009a, 2009b; Kolarov Most of the members of this large family are parasites of and Gürbüz, 2009; Kolarov and Yurtcan, 2009; Kolarov et holometabolous insects. A few species or groups of species al., 2009; Birol, 2010; Çoruh, 2010; Çoruh and Kolarov, parasitize spiders (egg sacs, spiderlings, or adults) or egg 2010a, 2010b; Eroğlu, 2010; Kolarov and Gürbüz, 2010; sacs of pseudoscorpions (cf. Clancy and Pierce, 1966). Riedel et al., 2010; Çoruh and Özbek, 2011; Çoruh et al., In the catalogue of Kolarov (1995), 383 species were 2011; Eroğlu et al., 2011; Gürbüz et al., 2011; Kolarov and listed from Turkey. Studies on the Ichneumonidae of Çalmaşur, 2011; Riedel et al., 2011; Tozlu and Çoruh, 2011; Turkey have gained acceleration, particularly in the last Çoruh and Kolarov, 2012a, 2012b; Çoruh and Khalaim, decade and a half (Özdemir and Kılınçer, 1990; Özdemir, 2012; Kolarov and Çoruh 2012a, 2012b; Okyar et al., 2012; 1996; Kolarov et al., 1997a, 1997b; Kolarov et al., 1999; Çoruh and Kolarov, 2013; Çoruh and Özbek, 2013; Çoruh Pekel, 1999; Yurtcan et al., 1999; Kolarov et al., 2000; Özbek et al., 2013; Çoruh et al., 2014a, 2014c; Kolarov et al., et al., 2000; Pekel et al., 2000; Özdemir, 2001; Çoruh et al., 2014a, 2014b; Özdan, 2014; Riedel et al., 2014). Kolarov et * Correspondence: [email protected] 40 KOLAROV et al. / Turk J Zool al. (2014a) and Çoruh et al. (2014c) reported 1047 species 3. Results and discussion in 286 genera for the Ichneumonidae fauna of Turkey. In this study, 57 species of 42 genera belonging to the With this paper, the number of the Ichneumonidae fauna subfamilies Acaenitinae, Anomaloninae, Banchinae, of Turkey reached 1060 species and 286 genera. Campopleginae, Cremastinae, Cryptinae, Diplazontinae, In the present paper 57 ichneumonids species from the Ichneumoninae, Metopiinae, Orthocentrinae, northeastern Anatolian part of Turkey are reported. Of Oxytorinae, Pimplinae, and Tryphoninae from the family them, 13 species are new records from Turkey. Ichneumonidae were recorded from the northeastern The aim of this study is to report the ichneumonid Anatolian part of Turkey. species and the species visited by ichneumonids, collected from different localities in the northeastern 3.1. Acaenitinae Anatolian part of Turkey in an ongoing project (Kolarov Phaenolobus fulvicornis (Gravenhorst, 1829) et al., 2014a, 2014b; Çoruh et al., 2014a, 2014b). Thus, Material examined: Rize: İkizdere, Tulumpınar, each new record will contribute to the knowledge of the 40°42′48.7″N, 40°37′6.7″E, 1170 m, 19.06.2014, 1♂. Ichneumonidae fauna in Turkey. Geographic distribution: , , , , , , and Turkey. 2. Materials and methods Habitat: Meadows area. The ichneumonids specimens were collected by net Vegetation: Achillea millefolium, Bellis perennis, from different plants in different localities in northeastern Capsella bursa-pastoris, Epilobium montanum, Geranium Anatolia (Figures 1 and 2) in June 2014. Collections of pyrenaicum, G. sylvaticum, Lapsana communis, Leontodon material are preserved in the collection of the University hispidus, Medicago lupulina, Plantago major, Prunella of Plovdiv (Bulgaria) at present. The plant samples visited vulgaris, Pteridium aquilinum, Ranunculus kochii, by ichneumonid specimens were collected by hand and Rosularia sempervivum, Rubus hirtus, Salvia verticillata, pressed, and they were deposited at the herbarium of the Tanacetum macrophyllum, Trifolium repens, Urtica dioica, Department of Plant Protection of Atatürk University thapsus. (Erzurum). All ichneumonid specimens were collected by Associated plants with P. fulvicornis: Anthriscus Dr S Çoruh and E Yıldırım. Plant specimens were identified sylvestris, Heracleum sphondylium, Salvia sylvestris. according to Davis (1965–1988) and the herbarium of Atatürk University, Faculty of Agriculture, Department of 3.2. Anomaloninae Plant Protection, by the third author. *Agrypon clandestinum (Gravenhorst, 1829) General distributions of all species and associated Material examined: Rize: Çamlıhemşin, , 40°53′230″N, plants of the some species present in the catalogue were 40°55′746″E, 1054 m, 20.06.2014, 1♂. taken from Yu et al. (2012). In addition, host plants of Geographic distribution: Holarctic and Oriental some ichneumonid species were also determined. region.

Figure 1. Map of study areas (from Google Earth).

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Figure 2. Pictures of study areas: a- Erzurum (İspir-Çayırözü), meadow area; b- Erzurum (Pazaryolu- Toprakkale), garden area; c- Trabzon (Araklı-Küçükdere), meadow area; d- Trabzon (Yomra), roadside; e- Rize (İkizdere-Tulumpınar), meadow area; f- Ordu (Perşembe-Akçaova Bridge), roadside, meadow area; g- Giresun (Eynesil), hazelnut area; h- Ordu (Fatsa), garden area.

New for the Turkish fauna. Geranium purpureum, G. pyrenaicum, Heracleum Habitat: Semiwet, sloping, brushwood area. sphondylium, Hypericum hirsutum, Inula vulgaris, Vegetation: Alopecurus myosuroides, Arctium minus, Lapsana communis, Lysimachia vulgaris, Matricaria Astragalus viciifolius, Atropa belladonna, Coronilla chamomilla, Melica ciliata, Mentha longifolia, Orchis orientalis, Crepis setosa, Dactylis glomerata, Dactylorhiza palustris, Phleum phleoides, Pilosella hoppeana, Prunella urvilleana, Eryngium giganteum, Euphrasia rostkoviana, vulgaris, Pteridium aquilinum, Ranunculus cappadocicus,

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R. repens, angustifolius, Rumex alpinus, Salvia Associated plants with A. bipunctoria: Alnus glutinosa, verticillata, Sambucus nigra, Sedum alpestre, Senecio Angelica sp., vulgare, Peucedanum oreoselinum, viscosus, Symphytum asperum, Tanacetum macrophyllum, Picea sp., Quercus robur. Thalictrum minus, Telekia speciosa, Trifolium pratense, T. Lissonota (Lissonota) folii Thomson, 1877 repens, Urtica dioica, . Material examined: Rize: Çamlıhemşin, Çat, Associated plants with A. clandestinum: not known. 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♂; Zilkale, cruentatum (Geoffroy, 1785) 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♀. Material examined: Erzurum: Pazaryolu, Kumaşkaya, Geographic distribution: Holarctic region. 40°31′50.523″N, 40°45′16.686″E, 1825 m, 19.06.2014, 1♂. Habitat: Semiwet, sloping, brushwood area and shrub Geographic distribution: Oriental and Palearctic area. region. Vegetation: Alopecurus myosuroides, Arctium minus, Habitat: Meadow area. Astragalus viciifolius, Atropa belladonna, Calystegia Vegetation: Achillea biebersteinii, A. millefolium, silvatica, Coronilla orientalis, Crepis setosa, Dactylis Amaranthus retroflexus, Carum carvi, Chenopodium album, glomerata, Dactylorhiza urvilleana, Daucus carota, Echium Cirsium arvense, Epilobium angustifolium, E. parviflorum, Filipendula vulgaris, Gladiolus atroviolaceus, Heracleum vulgare, Erigeron acer, Eryngium giganteum, Euphrasia pastinacifolium, Lysimachia vulgaris, neglecta, rostkoviana, Geranium purpureum, G. pyrenaicum, Pedicularis comosa, Plantago atrata, P. lanceolata, P. major, Heracleum sphondylium, Hypericum hirsutum, Inula Ranunculus grandiflorus, Rhinanthus angustifolius, Rumex vulgaris, Lapsana communis, Lysimachia vulgaris, acetosella, Silene compacta, Thalictrum minus, Trifolium Matricaria chamomilla, Melica ciliata, Mentha longifolia, pratense, T. repens. Mycelis muralis, Orchis palustris, Paspalum dilatatum, Associated plants with A. cruentatum: Anthriscus Phleum phleoides, Pilosella hoppeana, Plantago major, sylvestris, Peucedanum oreoselinum. Prunella vulgaris, Pteridium aquilinum, Ranunculus Barylypa delictor (Thunberg, 1822) cappadocicus, R. repens, Rhinanthus angustifolius, Rumex Material examined: Erzurum: Pazaryolu, Kumaşkaya, alpinus, Salvia verticillata, Sambucus nigra, Sedum 40°31′50.523″N, 40°45′16.686″E, 1825 m, 19.06.2014, 1♂. alpestre, Senecio viscosus, Symphytum asperum, Tanacetum Geographic distribution: Algeria, , Canary macrophyllum, Thalictrum minus, Telekia speciosa, Islands, Europe, , , , Morocco, Trifolium pratense, T. repens, Urtica dioica, Verbascum , and Turkey. thapsus. Habitat: Meadow area. Associated plants with L. (L.) folii: not known. Vegetation: Achillea biebersteinii, A. millefolium, *Lissonota (Lissonota) nitida (Gravenhorst, 1829) Amaranthus retroflexus, Carum carvi, Chenopodium album, Material examined: Erzurum: İspir, Çayırözü, Cirsium arvense, Epilobium angustifolium, E. parviflorum, 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂. Filipendula vulgaris, Gladiolus atroviolaceus, Heracleum Geographic distribution: Europe, Georgia, and pastinacifolium, Lysimachia vulgaris, Malva neglecta, Kazakhstan. Pedicularis comosa, Plantago atrata, P. lanceolata, P. major, New for the Turkish fauna. Ranunculus grandiflorus, Rhinanthus angustifolius, Rumex Habitat: Ovit Mountain foothills, meadow area. acetosella, Silene compacta, Thalictrum minus, Trifolium pratense, T. repens. Vegetation: Achillea millefolium, Alchemilla caucasica, Associated plants with B. delictor: Peucedanum Capsella bursa-pastoris, Centaurea glastifolia, Cephalaria oreoselinum. procera, Daucus carota, orientalis, Galium incanum, Hypericum perforatum, Lamium album, Papaver 3.3. Banchinae bracteatum, Plantago lanceolata, P. major, Poa bulbosa, Apophua bipunctoria (Thunberg, 1822) P. pratensis, Polygonum bistorta, Ranunculus kotschyi, Material examined: Ordu: Perşembe, Akçaova bridge, 40°59.940′N, 37°49.329′E, 28 m, 23.06.2014, 1♂. Reseda lutea, Rhinanthus angustifolius, Rumex acetosella, Geographic distribution: Holarctic region. R. crispus, Sanguisorba minor, Thalictrum minus, Trifolium Habitat: Roadside, meadow area. pratense, Vicia cracca. Vegetation: Anthemis tinctoria, Aristolochia pontica, Associated plants with L. (L.) nitida: Angelica Artemisia vulgaris, Calystegia sepium, Cichorium intybus, sylvestris. Convolvulus arvensis, Conyza canadensis, Daucus carota, Lissonota (Loxonota) histrio (Fabricius, 1798) Hypericum perforatum, Lactuca serriola, Medicago sativa, Material examined: Ordu: Perşembe, Alınca, Plantago major, Polygonum persicaria, Rubus discolor, 41°1.560′N, 37°48.216′E, 0 m, 23.06.2014, 1♀. Senecio vulgaris, Setaria glauca, Trifolium pratense, Geographic distribution: Holarctic region. Tussilago farfara, Urtica dioica, Xanthium spinosum. Habitat: Seaside, garden area.

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Vegetation: Artemisia vulgaris, Calystegia sepium, Associated plants with C. agilis: Peucedanum Chenopodium album, Datura stramonium, Daucus carota, oreoselinum. Lapsana communis, Mycelis muralis, Plantago major, Poa Campoletis latrator (Gravenhorst, 1829) pratensis, Polygonum persicaria, Rubus discolor, Rumex Material examined: Rize: Çamlıhemşin, Zilkale, crispus, Setaria glauca, Sonchus oleraceus, Sorghum 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♀. halepense, Tripleurospermum oreades, Urtica dioica, Geographic distribution: Europe and Turkey. , Xanthium spinosum. Habitat: Semiwet, shrub, sloping area. Associated plants with L. (L.) histrio: Chaerophyllum Vegetation: Arctium minus, Calystegia silvatica, Daucus aromaticum, Daucus carota, Pastinaca graveolens, carota, Echium vulgare, Erigeron acer, Galium aparine, Peucedanum oreoselinum, Salvia sylvestris. Geranium purpureum, Heracleum sphondylium, Inula Lissonota (Loxonota) mediterranea Seyrig, 1927 vulgaris, Lapsana communis, Mycelis muralis, Paspalum Material examined: Ordu: Perşembe, Akçaova bridge, dilatatum, Phleum phleoides, Plantago major, Prunella 40°59.940′N, 37°49.329′E, 28 m, 23.06.2014, 1♀, Alınca, vulgaris, Pteridium aquilinum, Ranunculus repens, Rumex 41°1.560′N, 37°48.216′E, 0 m, 23.06.2014, 1♂. alpinus, Salvia verticillata, Tanacetum macrophyllum, Geographic distribution: , , , Trifolium repens, Urtica dioica. former Yugoslavia, , Turkey, and . Associated plants with C. latrator: not known (Yu et Habitat: Roadside, seaside, garden and meadow area. al., 2012). Vegetation: Anthemis tinctoria, Artemisia vulgaris, *Diadegma combinatum (Holmgren, 1860) Aristolochia pontica, Artemisia vulgaris, Calystegia sepium, Material examined: Ordu: Gülyalı, Turnasuyu, Chenopodium album, Cichorium intybus, Convolvulus 40°58.624′N, 38°0.749′E, 1 m, 22.06.2014, 1♀, Fatsa, arvensis, Conyza canadensis, Datura stramonium, 41°1.934′N, 37°34.879′E, 20 m, 23.06.2014, 1♀. Daucus carota, Hypericum perforatum, Lactuca serriola, Geographic distribution: China (), Europe, Lapsana communis, Medicago sativa, Mycelis muralis, Poa Greenland, and Siberia. pratensis, Plantago major, Polygonum persicaria, Rubus New for the Turkish fauna. discolor, Rumex crispus, Senecio vulgaris, Setaria glauca, Habitat: Seaside and garden area. Sonchus oleraceus, Sorghum halepense, Trifolium pratense, Vegetation: Amaranthus retroflexus, Anthemis Tripleurospermum oreades, Tussilago farfara, Urtica dioica, tinctoria, Artemisia vulgaris, Avena fatua, Bromus Verbascum phlomoides, Xanthium spinosum. japonicas, Chenopodium album, Cichorium intybus, Cynodon dactylon, Daucus carota, Holcus lanatus, Associated plants with L. (L.) mediterranea: not Hypericum perforatum, Lapsana communis, Lolium known. perenne, Malva neglecta, Plantago lanceolata, P. major, 3.4. Campopleginae Polygonum persicaria, Rubus discolor, R. hirtus, Rumex *Campoletis agilis (Holmgren, 1860) crispus, Scrophularia scopolii, Setaria glauca, Solanum Material examined: Giresun: Eynesil, Dizgine, 41°3.41′N, nigrum, Sonchus oleraceus, Trifolium pratense, T. repens, 39°7.185′E, 49 m, 22.06.2014, 1♂, Piraziz, 40°56.987′N, Verbascum thapsus, Xanthium spinosum. 38°6.522′E, 16 m, 23.06.2014, 1♂, Tirebolu, 40°58.117′N, Associated plants with D. combinatum: Potentilla 38°44.945′E, 6 m, 24.06.2014, 3♂♂. fruticosa (Yu et al., 2012). Geographic distribution: Europe. Dusona stygia (Förster, 1868) New for the Turkish fauna. Material examined: Erzurum: Pazaryolu, Toprakkale, Habitat: Semiwet area at the water’s edge and hazelnut 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 1 ♂; area. Ordu, Fatsa, 41°1.934′N, 37°34.879′E, 20 m, 23.06.2014, Vegetation: Alopecurus myosuroides, Alyssum repens, 1♀. Anagallis arvensis, Artemisia vulgaris, Calystegia sepium, Geographic distribution: Algeria, , Europe, Chenopodium album, Conyza canadensis, Corylus Iran, Kazakhstan, Morocco, Tunisia, Turkey, and avellana, Daucus carota, Euphorbia helioscopia, Geranium . asphodeloides, Holcus lanatus, Hypericum perforatum, Habitat: Meadows, seaside and garden area. Lactuca serriola, Lolium perenne, Myosotis lazica, Poa Vegetation: Alchemilla caucasica, Allium armenum, longifolia, Polygonum persicaria, Plantago lanceolata, P. Amaranthus retroflexus, Anthemis cretica, A. tinctoria, major, Pteridium aquilinum, Rubus hirtus, Rumex crispus, Artemisia vulgaris, Avena fatua, Centaurea depressa, Salvia forskahlei, Senecio vulgaris, Setaria glauca, Sonchus Cerinthe minor, Chenopodium album, Cichorium intybus, oleraceus, Tanacetum macrophyllum, Tripleurospermum Daucus carota, Euphorbia stricta, Ferula orientalis, Galium oreades, Urtica dioica, Verbascum phlomoides, Vicia cracca, incanum, G. verum, Holcus lanatus, Hypericum perforatum, Xanthium spinosum. Juncus articulatus, Lapsana communis, Lolium perenne,

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Orchis mascula, Ornithogalum narbonense, Pedicularis atrata, P. lanceolata, Prunella vulgaris, Pteridium camosa, Plantago major, Poa pratensis, Polygonum aquilinum, Rubus discolor, R. hirtus, Rumex pulcher, Urtica bistorta, P. persicaria, Ranunculus kotschyi, Rhinanthus dioica. angustifolius, Rubus discolor, R. hirtus, Rumex acetosella, Associated plants with M. collector: Angelica sylvestris, R. crispus, Scrophularia scopolii, Setaria glauca, Solanum Peucedanum oreoselinum, Picea abies, Picea excelsa. nigrum, Sonchus oleraceus, Thalictrum minus, Verbascum *Olesicampe cavigena (Thomson, 1887) thapsus, Veronica gentianoides. Material examined: Rize: Çamlıhemşin, Zilkale, Associated plants with D. stygia: not known. 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♀. Dusona terebrator (Förster, 1868) Geographic distribution: Europe. New for the Turkish Material examined: Rize: Çamlıhemşin, Çat, fauna. 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 3♂♂, Habitat: Semiwet, shrub, sloping area. Zilkale, 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♂; Vegetation: Arctium minus, Calystegia silvatica, Daucus Trabzon: Araklı, Birlik, 40°51.677′N, 40°3.836′E, 118 m, carota, Echium vulgare, Erigeron acer, Galium aparine, 21.06.2014, 1♂. Geranium purpureum, Heracleum sphondylium, Inula Geographic distribution: Palearctic region. vulgaris, Lapsana communis, Mycelis muralis, Paspalum Habitat: Semiwet, shrub, sloping, meadow and dilatatum, Phleum phleoides, Plantago major, Prunella hazelnut area. vulgaris, Pteridium aquilinum, Ranunculus repens, Rumex Vegetation: Alopecurus myosuroides, Arctium minus, alpinus, Salvia verticillata, Tanacetum macrophyllum, Astragalus viciifolius, Atropa belladonna, Calystegia Trifolium repens, Urtica dioica. silvatica, Carex pallascens, Cichorium intybus, Coronilla Associated plants with O. cavigena: not known. orientalis, Corylus avellana, Crepis setosa, Dactylis *Olesicampe patellana (Thomson, 1887) glomerata, Dactylorhiza urvilleana, Daucus carota, Echium Material examined: Erzurum: Pazaryolu: Kumaşkaya, vulgare, Erigeron acer, Eryngium giganteum, Euphrasia 40°31′50.523″N, 40°45′16.686″E, 1825 m, 19.06.2014, 1♂; rostkoviana, Equisetum arvense, Geranium purpureum, Rize: İkizdere, Tulumpınar, 40°42′48.7″N, 40°37′6.7″N, G. pyrenaicum, Heracleum sphondylium, Holcus lanatus, 1170 m, 19.06.2014, 1♂, 1♀. Hypericum hirsutum, H. perforatum, Inula vulgaris, Geographic distribution: Europe. Lapsana communis, Lolium perenne, Lysimachia vulgaris, New for the Turkish fauna. Matricaria chamomilla, Melica ciliata, Mentha longifolia, Habitat: Meadow area. Mycelis muralis, Orchis palustris, Paspalum dilatatum, Vegetation: Achillea biebersteinii, A. millefolium, Phleum phleoides, Pilosella hoppeana, Plantago lanceolata, P. Amaranthus retroflexus, Bellis perennis, Capsella bursa- major, Prunella vulgaris, Pteridium aquilinum, Ranunculus pastoris, Carum carvi, Chenopodium album, Cirsium cappadocicus, R. repens, Rhinanthus angustifolius, Rumex arvense, Epilobium angustifolium, E. montanum, E. alpinus, Salvia verticillata, Sambucus nigra, Sedum alpestre, parviflorum, Filipendula vulgaris, Geranium pyrenaicum, Senecio viscosus, Sonchus oleraceus, Sorghum halepense, G. sylvaticum, Gladiolus atroviolaceus, Heracleum Rubus discolor, R. hirtus, Rumex crispus, Symphytum pastinacifolium, Lapsana communis, Leontodon hispidus, asperum, Tanacetum macrophyllum, Thalictrum minus, Lysimachia vulgaris, Malva neglecta, Medicago lupulina, Telekia speciosa, Trifolium pratense, T. repens, Urtica dioica, Pedicularis comosa, Plantago atrata, P. lanceolata, P. major, Verbascum thapsus. Prunella vulgaris, Pteridium aquilinum, Ranunculus Associated plants with D. terebrator: Cirsium grandiflorus, R. kochii, Rhinanthus angustifolius, Rosularia lanceolatum. sempervivum, Rubus hirtus, Rumex acetosella, Salvia Meloboris collector (Thunberg, 1822) Material examined: Trabzon: Araklı, Oylum, verticillata, Silene compacta, Tanacetum macrophyllum, 40°45.651′N, 40°3.498′E, 797 m, 21.06.2014, 1♀. Thalictrum minus, Trifolium pratense, T. repens, Urtica Geographic distribution: , Azerbaijan, dioica, Verbascum thapsus. Azores, , China (Qinghai, Shanxi), Europe, Associated plants with O. patellana: not known. Iceland, Israel, Madeira Islands, , South Africa, 3.5. Cremastinae and Turkey. Pristomerus armatus (Lucas, 1849) Habitat: Roadside, hazelnut area. Material examined: Erzurum: İspir, Çayırözü, Vegetation: Artemisia vulgaris, Calystegia sepium, 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂. Campanula rapunculoides, Carex pallascens, Centaurea Geographic distribution: Algeria, Europe, Georgia, hypoleuca, Chaerophyllum temulum, Conium maculatum, Iran, Kazakhstan, Kyrgyzstan, Morocco, Turkey, Corylus avellana, Daucus carota, Erigeron acer, Holcus Turkmenistan, and Siberia. lanatus, Leontodon hispidus, Medicago falcate, Plantago Habitat: Ovit Mountain foothills, meadow area.

45 KOLAROV et al. / Turk J Zool

Vegetation: Achillea millefolium, Alchemilla caucasica, Geranium purpureum, G. pyrenaicum, Heracleum Capsella bursa-pastoris, Centaurea glastifolia, Cephalaria sphondylium, Hypericum hirsutum, Inula vulgaris, procera, Daucus carota, Ferula orientalis, Galium Lapsana communis, Lysimachia vulgaris, Matricaria incanum, Hypericum perforatum, Lamium album, Papaver chamomilla, Melica ciliata, Mentha longifolia, Orchis bracteatum, Plantago lanceolata, P. major, Poa bulbosa, palustris, Phleum phleoides, Pilosella hoppeana, Prunella P. pratensis, Polygonum bistorta, Ranunculus kotschyi, vulgaris, Pteridium aquilinum, Ranunculus cappadocicus, Reseda lutea, Rhinanthus angustifolius, Rumex acetosella, R. repens, Rhinanthus angustifolius, Rumex alpinus, Salvia R. crispus, Sanguisorba minor, Thalictrum minus, Trifolium verticillata, Sambucus nigra, Sedum alpestre, Senecio pratense, Vicia cracca. viscosus, Symphytum asperum, Tanacetum macrophyllum, Associated plants with P. armatus: not known. Thalictrum minus, Telekia speciosa, Trifolium pratense, T. Temelucha genalis (Szépligeti, 1899) repens, Urtica dioica, Verbascum thapsus. Material examined: Erzurum: İspir, Çayırözü, Associated plants with A. senicula: not known. 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♀. Buathra laborator (Thunberg, 1822) Geographic distribution: Armenia, Azerbaijan, Material examined: Erzurum: Pazaryolu, Kumaşkaya, Europe, and Turkey. 40°31′50.523″N, 40°45′16.686″E, 1825 m, 19.06.2014, Habitat: Ovit Mountain foothills, meadow area. 1♀, Toprakkale, 40°16′41.47″N, 40°59′56″E, 2246 m, Vegetation: Achillea millefolium, Alchemilla caucasica, 19.06.2014, 1♀. Capsella bursa-pastoris, Centaurea glastifolia, Cephalaria Geographic distribution: Holarctic region. procera, Daucus carota, Ferula orientalis, Galium Habitat: Meadow area. incanum, Hypericum perforatum, Lamium album, Papaver Vegetation: Achillea biebersteinii, A. millefolium, bracteatum, Plantago lanceolata, P. major, Poa bulbosa, Alchemilla caucasica, Allium armenum, Amaranthus P. pratensis, Polygonum bistorta, Ranunculus kotschyi, retroflexus, Anthemis cretica, Carum carvi, Centaurea Reseda lutea, Rhinanthus angustifolius, Rumex acetosella, depressa, Cerinthe minor, Chenopodium album, Cirsium R. crispus, Sanguisorba minor, Thalictrum minus, Trifolium arvense, Epilobium angustifolium, E. parviflorum, pratense, Vicia cracca. Euphorbia stricta, Ferula orientalis, Filipendula vulgaris, Associated plants with T. genalis: not known Galium incanum, G. verum, Gladiolus atroviolaceus, 3.6. Cryptinae Heracleum pastinacifolium, Juncus articulatus, Lapsana *Aptesis assimilis (Gravenhorst, 1829) communis, Lysimachia vulgaris, Malva neglecta, Orchis Material examined: Erzurum: Pazaryolu, Toprakkale, mascula, Ornithogalum narbonense, Pedicularis camosa, 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 7♂♂. Plantago atrata, P. lanceolata, P. major, Poa pratensis, Geographic distribution: Europe. Polygonum bistorta, Ranunculus grandiflorus, R. kotschyi, New for the Turkish fauna. Rhinanthus angustifolius, Rumex acetosella, Silene Habitat: Meadow area. compacta, Thalictrum minus, Trifolium pratense, T. repens, Vegetation: Alchemilla caucasica, Allium armenum, Veronica gentianoides. Anthemis cretica, Centaurea depressa, Cerinthe minor, Associated plants with B. laborator: Peucedanum Euphorbia stricta, Ferula orientalis, Galium incanum, G. oreoselinum. verum, Juncus articulatus, Lapsana communis, Orchis Cryptus viduatorius Fabricius, 1804 mascula, Ornithogalum narbonense, Pedicularis camosa, Material examined: Erzurum: İspir, Çayırözü, Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 2♂♂, Rhinanthus angustifolius, Rumex acetosella, Thalictrum 1♀, Pazaryolu, Toprakkale, 40°16′41.47″N, 40°59′56″E, minus, Veronica gentianoides. 2246 m, 19.06.2014, 4♂♂, 1♀. Associated plants with A. assimilis: Listera ovata, Geographic distribution: Azerbaijan, , Georgia, Pastinoca sativa. Europe, Siberia, and Turkey. Aptesis senicula (Kriechbaumer, 1893) Habitat: Ovit Mountain foothills, meadow area. Material examined: Rize: Çamlıhemşin, Çat, Vegetation: Achillea millefolium, Alchemilla caucasica, 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♂. Allium armenum, Anthemis cretica, Capsella bursa- Geographic distribution: Azerbaijan, Bulgaria, pastoris, Centaurea depressa, C. glastifolia, Cephalaria , Moldova, , and Turkey. procera, Cerinthe minor, Daucus carota, Euphorbia stricta, Habitat: Semiwet, sloping, brushwood area. Ferula orientalis, Galium incanum, G. verum, Hypericum Vegetation: Alopecurus myosuroides, Arctium minus, perforatum, Juncus articulatus, Lamium album, Lapsana Astragalus viciifolius, Atropa belladonna, Coronilla communis, Orchis mascula, Ornithogalum narbonense, orientalis, Crepis setosa, Dactylis glomerata, Dactylorhiza Papaver bracteatum, Pedicularis camosa, Plantago urvilleana, Eryngium giganteum, Euphrasia rostkoviana, lanceolata, P. major, Poa bulbosa, P. pratensis, Polygonum

46 KOLAROV et al. / Turk J Zool bistorta, Ranunculus kotschyi, Reseda lutea, Rhinanthus Associated plants with G. profligator: Aegopodium sp., angustifolius, Rumex acetosella, R. crispus, Sanguisorba Alisma sp., Alnus glutinosa, Angelica sp., Chaerophyllum minor, Thalictrum minus, Trifolium pratense, Veronica sp., Euphorbia nicaeensis, Euphorbia seguieriana, gentianoides, Vicia cracca. Euphorbia virgata, Heracleum sp., Heracleum sphondylium, Associated plants with C. viduatorius: Anethum Peucedanum oreoselinum, Poa pratensis, Quercus sp., Tilia graveolens, Angelica sylvestris, Daucus carota, Daucus cordata. carota sativus, Euphorbia sp., Euphorbia nicaeensis, Glyphicnemis vagabunda (Gravenhorst, 1829) Euphorbia virgata, Ferula communis, Heracleum Material examined: Erzurum: İspir, Çayırözü, sphondylium, Medicago sativa, Peucedanum oreoselinum 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂; (Çoruh and Çoruh, 2008). Pazaryolu, Toprakkale, 40°16′41.47″N, 40°59′56″E, Remark: This species was collected on Ferula orientalis 2246 m, 19.06.2014, 1♂; Kumaşkaya, 40°31′50.523″N, while feeding. 40°45′16.686″E, 1825 m, 19.06.2014, 1♂. Endasys plagiator (Gravenhorst, 1829) Geographic distribution: Azerbaijan, Europe, Georgia, Material examined: Erzurum: Pazaryolu, Toprakkale, Iran, and Turkey. 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 1 ♂. Habitat: Ovit Mountain foothills, meadow area. Geographic distribution: Europe and Turkey. Vegetation: Achillea biebersteinii, A. millefolium, Habitat: Meadow area. Alchemilla caucasica, Allium armenum, Amaranthus Vegetation: Alchemilla caucasica, Allium armenum, retroflexus, Anthemis cretica, Capsella bursa-pastoris, Anthemis cretica, Centaurea depressa, Cerinthe minor, Carum carvi, Centaurea depressa, C. glastifolia, Cephalaria Euphorbia stricta, Ferula orientalis, Galium incanum, G. procera, Cerinthe minor, Chenopodium album, Cirsium verum, Juncus articulatus, Lapsana communis, Orchis arvense, Daucus carota, Epilobium angustifolium, mascula, Ornithogalum narbonense, Pedicularis camosa, E. parviflorum, Euphorbia stricta, Ferula orientalis, Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, Filipendula vulgaris, Galium incanum, G. verum, Gladiolus Rhinanthus angustifolius, Rumex acetosella, Thalictrum atroviolaceus, Heracleum pastinacifolium, Hypericum minus, Veronica gentianoides. perforatum, Juncus articulatus, Lamium album, Lapsana Associated plants with E. plagiator: not known. communis, Lysimachia vulgaris, Malva neglecta, Orchis agilis (Fabricius, 1775) mascula, Ornithogalum narbonense, Papaver bracteatum, Material examined: Trabzon: Araklı, Birlik, Pedicularis camosa, Plantago atrata, P. lanceolata, P. major, 40°51.677′N, 40°3.836′E, 118 m, 21.06.2014, 1♂. Poa bulbosa, P. pratensis, Polygonum bistorta, Ranunculus Geographic distribution: Palearctic region. grandiflorus, R. kotschyi, Reseda lutea, Rhinanthus Habitat: Roadside, hazelnut area. angustifolius, Rumex acetosella, R. crispus, Sanguisorba Vegetation: Carex pallescens, Cichorium intybus, minor, Silene compacta, Thalictrum minus, Trifolium Corylus avellana, Daucus carota, Erigeron acer, Equisetum pratense, T. repens, Veronica gentianoides, Vicia cracca. arvense, Holcus lanatus, Hypericum perforatum, Lapsana Associated plants with G. vagabunda: Angelica communis, Lolium perenne, Paspalum dilatatum, Plantago sp., Carum carvi, Daucus carota, Euphorbia nicaeensis, lanceolata, P. major, Trifolium pratense, Salvia verticillata, Euphorbia virgata, Fraxinus excelsior, Heracleum sp., Sonchus oleraceus, Sorghum halepense, Rubus discolor, R. Heracleum sibiricum, Heracleum sphondylium, Listera hirtus, Rumex crispus. ovata, Quercus sp., Seseli libanotis, Seseli sp. (Çoruh and Associated plants with G. agilis: Lonicera sp., Prunus Çoruh, 2008). sp., Mentha longifolia, Quercus robur, Picea excelsa, Salix Remark: This species was collected onCarum carvi sp. while feeding. Glyphicnemis profligator (Fabricius, 1775) Ischnus alternator (Gravenhorst, 1829) Material examined: Trabzon: Araklı, Birlik, Material examined: Ordu: Perşembe, Kaleyaka, 40°51.677′N, 40°3.836′E, 118 m, 21.06.2014, 2♂♂. 41°6.077′N, 37°47.013′E, 51 m, 23.06.2014, 1♀. Geographic distribution: Palearctic region. Geographic distribution: Azerbaijan, Europe, Iran, Habitat: Roadside, hazelnut area. and Turkey. Vegetation: Carex pallescens, Cichorium intybus, Habitat: Roadside and garden area. Corylus avellana, Daucus carota, Erigeron acer, Equisetum Vegetation: Bromus japonicus, Calystegia sepium, arvense, Holcus lanatus, Hypericum perforatum, Lapsana Cichorium intybus, Conyza bonariensis, Daucus carota, communis, Lolium perenne, Paspalum dilatatum, Plantago Epilobium angustifolium, Lactuca serriola, Lolium perenne, lanceolata, P. major, Trifolium pratense, Salvia verticillata, Poa pratensis, Plantago major, Rubus discolor, Rumex Sonchus oleraceus, Sorghum halepense, Rubus discolor, R. crispus, Setaria glauca, Sorghum halepense, Trifolium hirtus, Rumex crispus. pratense, Tripleurospermum oreades.

47 KOLAROV et al. / Turk J Zool

Associated plants with I. alternator: Peucedanum Vegetation: Achillea millefolium, Alchemilla caucasica, oreoselinum. Capsella bursa-pastoris, Centaurea glastifolia, Cephalaria Meringopus calescens calescens (Gravenhorst, 1829) procera, Daucus carota, Ferula orientalis, Galium Material examined: Erzurum: Pazaryolu, Toprakkale, incanum, Hypericum perforatum, Lamium album, Papaver 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 3♂♂. bracteatum, Plantago lanceolata, P. major, Poa bulbosa, Geographic distribution: Holarctic and Oriental P. pratensis, Polygonum bistorta, Ranunculus kotschyi, region. Reseda lutea, Rhinanthus angustifolius, Rumex acetosella, Habitat: Meadow area. R. crispus, Sanguisorba minor, Thalictrum minus, Trifolium Vegetation: Alchemilla caucasica, Allium armenum, pratense, Vicia cracca. Anthemis cretica, Centaurea depressa, Cerinthe minor, Associated plants with M. titillator: Carum carvi, Euphorbia stricta, Ferula orientalis, Galium incanum, G. Euphorbia nicaeensis, Euphorbia virgata, Fraxinus excelsior, verum, Juncus articulatus, Lapsana communis, Orchis Seseli libanotis, Spiraea sp., , Chaerophyllum mascula, Ornithogalum narbonense, Pedicularis camosa, bulbosum (Çoruh and Çoruh, 2008). Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, Remark: This species was collected on Ferula orientalis Rhinanthus angustifolius, Rumex acetosella, Thalictrum while feeding. minus, Veronica gentianoides. Mesostenus albinotatus Gravenhorst, 1829 Associated plants with M. calescens calescens: Carum Material examined: Erzurum: Pazaryolu, Toprakkale, carvi (Çoruh and Çoruh, 2008). 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 1♂. Remark: This species was collected on Ferula orientalis Geographic distribution: Holarctic region. while feeding. Habitat: Meadow area. Meringopus calescens persicus Heinrich, 1937 Vegetation: Alchemilla caucasica, Allium armenum, Anthemis cretica, Centaurea depressa, Cerinthe minor, Material examined: Erzurum: İspir, Çayırözü, Euphorbia stricta, Ferula orientalis, Galium incanum, G. 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂, verum, Juncus articulatus, Lapsana communis, Orchis Pazaryolu, Toprakkale, 40°16′41.47″N, 40°59′56″E, 2246 mascula, Ornithogalum narbonense, Pedicularis camosa, m, 19.06.2014, 2♂♂. Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, Geographic distribution: Azerbaijan, Iran, Kyrgyzstan, Rhinanthus angustifolius, Rumex acetosella, Thalictrum Mongolia, and Turkey. minus, Veronica gentianoides. Habitat: Ovit Mountain foothills, meadows area. Associated plants with M. albinotatus: Daucus carota, Vegetation: Achillea millefolium, Alchemilla caucasica, Daucus carota sativus, Euphorbia seguieriana, Heracleum Allium armenum, Anthemis cretica, Capsella bursa- sphondylium, Peucedanum oreoselinum, Pimpinella pastoris, Centaurea depressa, C. glastifolia, Cephalaria tragium (Çoruh and Çoruh, 2008). procera, Cerinthe minor, Daucus carota, Euphorbia stricta, Remark: This species was collected on Euphorbia Ferula orientalis, Galium incanum, G. verum, Hypericum stricta while feeding. perforatum, Juncus articulatus, Lamium album, Lapsana Rhembobius quadrispinus (Gravenhorst, 1829) communis, Orchis mascula, Ornithogalum narbonense, Material examined: Giresun: Piraziz, 40°56.987′N, Papaver bracteatum, Pedicularis camosa, Plantago 38°6.522′E, 16 m, 23.06.2014, 2♂♂. lanceolata, P. major, Poa bulbosa, P. pratensis, Polygonum Geographic distribution: Europe and Turkey. bistorta, Ranunculus kotschyi, Reseda lutea, Rhinanthus Habitat: Hazelnut area. angustifolius, Rumex acetosella, R. crispus, Sanguisorba Vegetation: Alyssum repens, Calystegia sepium, minor, Thalictrum minus, Trifolium pratense, Veronica Chenopodium album, Corylus avellana, Lactuca serriola, gentianoides, Vicia cracca. Lolium perenne, Myosotis lazica, Polygonum persicaria, Associated plants with M. calescens persicus: not Pteridium aquilinum, Rumex crispus, Senecio vulgaris, known. Setaria glauca, Tanacetum macrophyllum, Urtica dioica, Meringopus titillator (Linnaeus, 1758) Vicia cracca, Xanthium spinosum. Material examined: Erzurum: İspir, Çayırözü, Associated plants with R. quadrispinus: Deschampsia 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂. cespitosa (Yu et al., 2012). Geographic distribution: Afghanistan, Algeria, *Schreineria populnea (Giraud, 1872) Armenia, Azerbaijan, China (Qinghai), Cyprus, Egypt, Material examined: Rize: Çamlıhemşin, Çat, Europe, Iran, Israel, Jordan, Kyrgyzstan, Morocco, 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♀. Palestine, Siberia, , Tajikistan, Tunisia, Turkmenistan, Geographic distribution: Palearctic region. New for Turkey, and . the Turkish fauna. Habitat: Ovit Mountain foothills, meadows area. Habitat: Semiwet, sloping, brushwood area.

48 KOLAROV et al. / Turk J Zool

Vegetation: Alopecurus myosuroides, Arctium minus, Leontodon hispidus, Lysimachia vulgaris, Matricaria Astragalus viciifolius, Atropa belladonna, Coronilla chamomilla, Medicago lupulina, Melica ciliata, Mentha orientalis, Crepis setosa, Dactylis glomerata, Dactylorhiza longifolia, Mycelis muralis, Orchis palustris, Paspalum urvilleana, Eryngium giganteum, Euphrasia rostkoviana, dilatatum, Phleum phleoides, Pilosella hoppeana, Plantago Geranium purpureum, G. pyrenaicum, Heracleum major, Prunella vulgaris, Pteridium aquilinum, Ranunculus sphondylium, Hypericum hirsutum, Inula vulgaris, cappadocicus, R. kochii, R. repens, Rhinanthus angustifolius, Lapsana communis, Lysimachia vulgaris, Matricaria Rosularia sempervivum, Rubus hirtus, Rumex alpinus, chamomilla, Melica ciliata, Mentha longifolia, Orchis Tanacetum macrophyllum, Salvia verticillata, Sambucus palustris, Phleum phleoides, Pilosella hoppeana, Prunella nigra, Sedum alpestre, Senecio viscosus, Symphytum vulgaris, Pteridium aquilinum, Ranunculus cappadocicus, asperum, Tanacetum macrophyllum, Thalictrum minus, R. repens, Rhinanthus angustifolius, Rumex alpinus, Salvia Telekia speciosa, Trifolium pratense, T. repens, Urtica dioica, verticillata, Sambucus nigra, Sedum alpestre, Senecio Verbascum thapsus. viscosus, Symphytum asperum, Tanacetum macrophyllum, Associated plants with D. pallicoxa: not known. Thalictrum minus, Telekia speciosa, Trifolium pratense, T. *Homotropus pallipes (Gravenhorst, 1829) repens, Urtica dioica, Verbascum thapsus. Material examined: Trabzon: Vakfıkebir, 41°2.997′N, Associated plants with S. populnea: Carpinus betulus, 39°16.907′E, 5 m, 24.06.2014, 2♀♀. Euonymus europaeus, Rhamnus frangula (Yu et al., 2012). Geographic distribution: Holarctic region and Mexico. *Xylophrurus lancifer (Gravenhorst, 1829) New for the Turkish fauna. Material examined: Erzurum: Pazaryolu, Kumaşkaya, Habitat: Seaside, garden area. 40°31′50.523N, 40°45′16.686″E, 1825 m, 19.06.2014, 2♂♂. Vegetation: Alopecurus myosuroides, Anagallis Geographic distribution: Palearctic region. arvensis, Artemisia vulgaris, Cichorium intybus, Conyza New for the Turkish fauna. bonariensis, Holcus lanatus, Lactuca serriola, Lapsana Habitat: Meadow area. communis, Medicago sativa, Melilotus alba, Poa pratensis, Vegetation: Achillea biebersteinii, A. millefolium, Rubus idaeus, Sonchus asper, Trifolium pratense, Vicia Amaranthus retroflexus, Carum carvi, Chenopodium album, cracca. Cirsium arvense, Epilobium angustifolium, E. parviflorum, Associated plants with H. pallipes: Poa pratensis, Filipendula vulgaris, Gladiolus atroviolaceus, Heracleum Tsuga heterophylla. pastinacifolium, Lysimachia vulgaris, Malva neglecta, Syrphoctonus tarsatorius (Panzer, 1809) Pedicularis comosa, Plantago atrata, P. lanceolata, P. major, Material examined: Giresun: Tirebolu, Balçıkbelen, Ranunculus grandiflorus, Rhinanthus angustifolius, Rumex 40°58.386′N, 38°52.576′E, 71 m, 22.06.2014, 1♀. acetosella, Silene compacta, Thalictrum minus, Trifolium Geographic distribution: Holarctic and Oriental pratense, T. repens. region. Associated plants with X. lancifer persicus: not known. Habitat: Hazelnut area. 3.7. Diplazontinae Vegetation: Anthemis tinctoria, Artemisia vulgaris, *Diplazon pallicoxa Manukyan, 1987 Calystegia sepium, Carex echinata, Corylus avellana, Material examined: Rize: Çamlıhemşin, Zilkale, Daucus carota, Lapsana communis, Lolium perenne, 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♂, Çat, Medicago sativa, Poa longifolia, Plantago lanceolata, P. 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♀, major, Pteridium aquilinum, Rubus hirtus, Rumex crispus, İkizdere, Tulumpınar, 40°42′48.7″N, 40°37′6.7″E, 1170 m, Sonchus oleraceus, Tanacetum macrophyllum, Trifolium 19.06.2014, 1♂. pretense. Geographic distribution: Palearctic region. Associated plants with S. tarsatorius: not known. New for the Turkish fauna. Syrphophilus bizonarius (Gravenhorst, 1829) Habitat: Semiwet, shrub, sloping area and meadow Material examined: Giresun: Eynesil, 41°3.394′N, area. 39°8.021′E, 0 m, 22.06.2014, 1♀; Trabzon: Vakfıkebir, Vegetation: Achillea millefolium, Alopecurus 41°2.997′N, 39°16.907′E, 5 m, 24.06.2014, 1♀. myosuroides, Arctium minus, Astragalus viciifolius, Atropa Geographic distribution: Holarctic and Oriental belladonna, Bellis perennis, Calystegia silvatica, Capsella region. bursa-pastoris, Coronilla orientalis, Crepis setosa, Dactylis Habitat: Seaside and hazelnut area. glomerata, Dactylorhiza urvilleana, Daucus carota, Echium Vegetation: Alopecurus myosuroides, Anagallis vulgare, Epilobium montanum, Erigeron acer, Eryngium arvensis, Artemisia vulgaris, Calystegia sepium, Cichorium giganteum, Euphrasia rostkoviana, Geranium purpureum, intybus, Conyza bonariensis, Corylus avellana, Daucus G. pyrenaicum, G. sylvaticum, Heracleum sphondylium, carota, Euphorbia helioscopia, Holcus lanatus, Hypericum Hypericum hirsutum, Inula vulgaris, Lapsana communis, perforatum, Lactuca serriola, Lapsana communis, Medicago

49 KOLAROV et al. / Turk J Zool sativa, Melilotus alba, M. officinalis, Poa pratensis, Plantago Vegetation: Alopecurus myosuroides, Alyssum repens, major, Polygonum persicaria, Prunella vulgaris, Pteridium Amaranthus retroflexus, Anagallis arvensis, Anthemis aquilinum, Raphanus raphanistrum, Rorippa sylvestre, tinctoria, Artemisia vulgaris, Avena fatua, Bromus japonicas, Rubus discolor, R. idaeus, Rumex crispus, Senecio vulgaris, Calystegia sepium, Chenopodium album, Cichorium Solanum nigrum, Sonchus asper, S. oleraceus, Tanacetum intybus, Conyza bonariensis, Corylus avellana, Cynodon macrophyllum, Trifolium pratense, Vicia cracca. dactylon, Daucus carota, Epilobium angustifolium, Holcus Associated plants with S. bizonarius: not known. lanatus, Hypericum perforatum, Lactuca serriola, Lapsana *Woldstedtius biguttatus (Gravenhorst, 1829) communis, Lolium perenne, Malva neglecta, Medicago Material examined: Trabzon: Vakfıkebir, 41°2.997′N, sativa, Melilotus alba, Myosotis lazica, Phragmites austrialis, 39°16.907′E, 5 m, 24.06.2014, 1♀. Plantago lanceolata, P. major, Poa pratensis, Polygonum Geographic distribution: Palearctic region. New for persicaria, Pteridium aquilinum, Rubus discolor, R. hirtus, the Turkish fauna. R. idaeus, Rumex crispus, Scrophularia scopolii, Senecio Habitat: Seaside, garden area. vulgaris, Setaria glauca, Solanum nigrum, Sonchus asper, Vegetation: Alopecurus myosuroides, Anagallis S. oleraceus, Tanacetum macrophyllum, Trifolium pratense, arvensis, Artemisia vulgaris, Cichorium intybus, Conyza T. repens, Tripleurospermum oreades, Urtica dioica, bonariensis, Holcus lanatus, Lactuca serriola, Lapsana Verbascum thapsus, Vicia cracca, Xanthium spinosum. communis, Medicago sativa, Melilotus alba, Poa pratensis, Associated plants with I. proletarius: Anethum Rubus idaeus, Sonchus asper, Trifolium pratense, Vicia graveolens, Angelica sylvestris, Anthriscus sylvestris, cracca. Chaerophyllum aromaticum, Daucus carota, Euphorbia Associated plants with W. biguttatus: not known. virgata, Ferulago sylvatica, Heracleum sphondylium, 3.8. Ichneumoninae Medicago sativa, Pastinaca graveolens, Pastinaca sp., Cratichneumon viator (Scopoli, 1763) Peucedanum oreoselinum, Rubus fruticosus. Material examined: Rize: İkizdere, Sivrikaya, Remark: This species was collected on Hypericum 40°40′37.4″N, 40°42′30.6″E, 1871 m, 19.06.2014, 1♂; perforatum while feeding. Ordu: Perşembe, Çaytepe, 41°7.697′N, 37°41.092′E, 10 m, Obtusodonta equitatoria (Panzer, 1786) 23.06.2014, 1♂. Material examined: Erzurum: Pazaryolu, Toprakkale, Geographic distribution: Holarctic region. 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 1♀. Habitat: Roadside, meadow, seaside, and garden area. Geographic distribution: Palearctic region. Vegetation: Alchemilla caucasica, Anthemis tinctoria, Habitat: Meadow area. Artemisia vulgaris, Bellis perennis, Bromus japonicas, Vegetation: Alchemilla caucasica, Allium armenum, Campanula rapunculoides, Cichorium intybus, Coronilla Anthemis cretica, Centaurea depressa, Cerinthe minor, orientalis, Daucus carota, Echium vulgare, Equisetum Euphorbia stricta, Ferula orientalis, Galium incanum, G. arvense, Heracleum platytaenium, Hypericum perforatum, verum, Juncus articulatus, Lapsana communis, Orchis Lactuca serriola, Neslia paniculata, Orchis palustris, mascula, Ornithogalum narbonense, Pedicularis camosa, Papaver lateritium, Pilosella hoppeana, Plantago lanceolata, Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, P. major, P. me di a , Polygonum persicaria, Pteridium Rhinanthus angustifolius, Rumex acetosella, Thalictrum aquilinum, Rosularia sempervivum, Rubus discolor, R. minus, Veronica gentianoides. hirtus, Rumex alpinus, R. crispus, Salvia verticillata, Silene Associated plants with O. equitatoria: Chaerophyllum compacta, Sonchus asper, Tanacetum macrophyllum, aromaticum, Daucus carota, Heracleum sphondylium, Trifolium pratense, T. repens, Tripleurospermum oreades, Medicago sativa, Melilotus indica, Oryza sativa, Vicia cracca. Peucedanum oreoselinum, Pinus sylvestris. Associated plants with C. viator: not known. Virgichneumon callicerus (Gravenhorst, 1820) Ichneumon proletarius Wesmael, 1848 Material examined: Trabzon: Araklı, Küçükdere, Material examined: Giresun: Görele, Çavuşlu, 40°54.395′N, 40°3.361′E, 33 m, 21.06.2014, 1♂. 41°2.449′N, 39°4.403′E, 0 m, 22.06.2014, 1♂, Piraziz, Geographic distribution: Azerbaijan, China (Qinghai 40°56.987′N, 38°6.522′E, 16 m, 23.06.2014, 8♂♂; Ordu: and Xinjiang), Europe, Iran, Kazakhstan, Siberia, and Turnasuyu, 40°58.624′N, 38°0.749′E, 1 m, 22.06.2014, Turkey. 8♂♂, 1♀; Fatsa, 41°1.934′N, 37°34.879′E, 20 m, 23.06.2014, Habitat: Meadows area. 3♂♂; Trabzon: Vakfıkebir, 41°2.997′N, 39°16.907′E, 5 m, Vegetation: Artemisia vulgaris, Briza maxima, 24.06.2014, 2♂♂. Bromus japonicus, Calystegia sepium, Cichorium intybus, Geographic distribution: Algeria, Azerbaijan, Europe, Conyza bonariensis, Dactylis glomerata, Daucus carota, Georgia, Iran, and Turkey. Erigeron acer, Holcus lanatus, Hypericum perforatum, Habitat: Seaside, garden, and hazelnut area. Lolium perenne, Medicago lupulina, M. sativa, Paspalum

50 KOLAROV et al. / Turk J Zool dilatatum, Phleum pratense, Plantago lanceolata, P. major, 3.11. Oxytorinae Rubus discolor, Sonchus oleraceus, Trifolium pratense. *Oxytorus luridator (Gravenhorst, 1820) Associated plants with V. callicerus: not known. Material examined: Rize: Çamlıhemşin, Zilkale, 3.9. Metopiinae 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 1♂, ′ ″ ′ ″ Exochus mitratus Gravenhorst, 1829 İkizdere, Tulumpınar, 40°42 48.7 N, 40°37 6.7 E, 1170 m, ♂ Material examined: Ordu: Fatsa, 41°1.934′N, 37°34.879′E, 19.06.2014, 1 . Geographic distribution: Europe and Georgia. 20 m, 22.06.2014, 1♀. New for the Turkish fauna. Geographic distribution: Holarctic region. Habitat: Semiwet, shrub, sloping and meadow area. Habitat: Seaside, garden area. Vegetation: Achillea millefolium, Arctium minus, Bellis Vegetation: Amaranthus retroflexus, Anthemis perennis, Calystegia silvatica, Capsella bursa-pastoris, tinctoria, Artemisia vulgaris, Avena fatua, Chenopodium Daucus carota, Echium vulgare, Epilobium montanum, album, Cichorium intybus, Daucus carota, Holcus lanatus, Erigeron acer, Galium aparine, Geranium purpureum, Hypericum perforatum, Lapsana communis, Lolium G. pyrenaicum, G. sylvaticum, Heracleum sphondylium, perenne, Plantago major, Polygonum persicaria, Rubus Inula vulgaris, Lapsana communis, Leontodon hispidus, discolor, Rumex crispus, Scrophularia scopolii, Setaria Medicago lupulina, Mycelis muralis, Paspalum dilatatum, glauca, Solanum nigrum, Sonchus oleraceus, Verbascum Phleum phleoides, Plantago major, Prunella vulgaris, thapsus. Pteridium aquilinum, Ranunculus kochii, R. repens, Associated plants with E. mitratus: Peucedanum Rosularia sempervivum, Rubus hirtus, Rumex alpinus, oreoselinum, Quercus lusitanica. Salvia verticillata, Tanacetum macrophyllum, Trifolium Exochus suborbitalis Schmiedeknecht, 1924 repens, Urtica dioica, Verbascum thapsus. Material examined: Giresun: Tirebolu, Balçıkbelen, Associated plants with O. luridator: not known. 40°58.386′N, 38°52.576′E, 71 m, 22.06.2014, 1♀; Ordu: 3.12. Pimplinae Fatsa, 41°1.934′N, 37°34.879′E, 20 m, 23.06.2014, 1♀. Exeristes roborator (Fabricius, 1793) Geographic distribution: Palearctic region. Material examined: Erzurum: İspir, Çayırözü, Habitat: Hazelnut and seaside, garden area. 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♂,1♀. Vegetation: Amaranthus retroflexus, Anthemis tinctoria, Geographic distribution: Afrotropical region, Artemisia vulgaris, Avena fatua, Calystegia sepium, Carex Oriental, and introduced in North America and the echinata, Chenopodium album, Cichorium intybus, Corylus Palearctic. avellana, Daucus carota, Holcus lanatus, Hypericum Habitat: Ovit Mountain foothills, meadow area. perforatum, Lapsana communis, Lolium perenne, Medicago Vegetation: Achillea millefolium, Alchemilla caucasica, sativa, Poa longifolia, Plantago lanceolata, P. major, Capsella bursa-pastoris, Centaurea glastifolia, Cephalaria Polygonum persicaria, Pteridium aquilinum, Rubus discolor, procera, Daucus carota, Ferula orientalis, Galium R. hirtus, Rumex crispus, Scrophularia scopolii, Setaria incanum, Hypericum perforatum, Lamium album, Papaver glauca, Solanum nigrum, Sonchus oleraceus, Tanacetum bracteatum, Plantago lanceolata, P. major, Poa bulbosa, macrophyllum, Trifolium pratense, Verbascum thapsus. P. pratensis, Polygonum bistorta, Ranunculus kotschyi, Associated plants with E. suborbitalis: Peucedanum Reseda lutea, Rhinanthus angustifolius, Rumex acetosella, oreoselinum. R. crispus, Sanguisorba minor, Thalictrum minus, Trifolium pratense, Vicia cracca. 3.10. Orthocentrinae Associated plants with E. roborator: Anethum Megastylus (Megastylus) flavopictus (Gravenhorst, 1829) graveolens, Daucus carota, Euphorbia virgata, Fraxinus Material examined: Ordu: Perşembe, Kaleyaka, excelsior, Heracleum sphondylium, Pastinaca graveolens, 41°6.077′N, 37°47.013′E, 51 m, 23.06.2014, 1♀. Peucedanum oreoselinum, Quercus sessiliflora, Salvia Geographic distribution: Holarctic region. sylvestris, Seseli tortuosum, Tamarix sp. Habitat: Roadside and garden area. Remark: This species was collected on Daucus carota Vegetation: Bromus japonicus, Calystegia sepium, while feeding. Cichorium intybus, Conyza bonariensis, Daucus carota, Gregopimpla inquisitor (Scopoli, 1763) Epilobium angustifolium, Lactuca serriola, Lolium perenne, Material examined: Rize: Çamlıhemşin, Çat, Poa pratensis, Plantago major, Rubus discolor, Rumex 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♀. crispus, Setaria glauca, Sorghum halepense, Trifolium Geographic distribution: Holarctic region. pratense, Tripleurospermum oreades. Habitat: Semiwet, sloping, brushwood area. Associated plants with M. (M.) flavopictus: not Vegetation: Alopecurus myosuroides, Arctium minus, known. Astragalus viciifolius, Atropa belladonna, Coronilla

51 KOLAROV et al. / Turk J Zool orientalis, Crepis setosa, Dactylis glomerata, Dactylorhiza Habitat: Meadow area. urvilleana, Eryngium giganteum, Euphrasia rostkoviana, Vegetation: Achillea biebersteinii, A. millefolium, Geranium purpureum, G. pyrenaicum, Heracleum Amaranthus retroflexus, Carum carvi, Chenopodium album, sphondylium, Hypericum hirsutum, Inula vulgaris, Cirsium arvense, Epilobium angustifolium, E. parviflorum, Lapsana communis, Lysimachia vulgaris, Matricaria Filipendula vulgaris, Gladiolus atroviolaceus, Heracleum chamomilla, Melica ciliata, Mentha longifolia, Orchis pastinacifolium, Lysimachia vulgaris, Malva neglecta, palustris, Phleum phleoides, Pilosella hoppeana, Prunella Pedicularis comosa, Plantago atrata, P. lanceolata, P. major, vulgaris, Pteridium aquilinum, Ranunculus cappadocicus, Ranunculus grandiflorus, Rhinanthus angustifolius, Rumex R. repens, Rhinanthus angustifolius, Rumex alpinus, Salvia acetosella, Silene compacta, Thalictrum minus, Trifolium verticillata, Sambucus nigra, Sedum alpestre, Senecio pratense, T. repens. viscosus, Symphytum asperum, Tanacetum macrophyllum, Associated plants with P. gnathaulax: not known. Thalictrum minus, Telekia speciosa, Trifolium pratense, T. Remark: This species was collected on Heracleum repens, Urtica dioica, Verbascum thapsus. pastinacifolium while feeding. Associated plants with G. inquisitor: Alnus glutinosa, Pimpla spuria Gravenhorst, 1829 Daucus carota sativus, Epilobium angustifolium, Heracleum Material examined: Giresun: Görele, Çavuşlu, sphondylium, Impatiens parviflora. 41°2.449′N, 39°4.403′E, 0 m, 22.06.2014, 1♂, Piraziz, Remark: This species was collected on Hypericum 40°56.987′N, 38°6.522′E, 16 m, 23.06.2014, 3♂♂; Ordu: hirsutum while feeding. Turnasuyu, 40°58.624′N, 38°0.749′E, 1 m, 23.06.2014, ♂. Itoplectis maculator (Fabricius, 1775) Geographic distribution: Oriental and Palearctic Material examined: Erzurum: İspir, Çayırözü, region. 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♀; Habitat: Seaside and hazelnut area. Trabzon: Yomra, 40°56.058′N, 39°52.526′E, 401 m, Vegetation: Alyssum repens, Amaranthus retroflexus, 21.06.2014, 1♀; Araklı, Birlik, 40°51.677′N, 40°03.836′E, Anthemis tinctoria, Artemisia vulgaris, Avena fatua, 118 m, 21.06.2014, 1♀. Bromus japonicas, Calystegia sepium, Chenopodium album, Geographic distribution: Armenia, Europe, Iran, Cichorium intybus, Conyza bonariensis, Corylus avellana, Mongolia, Morocco, Siberia, Tunisia, and Turkey. Cynodon dactylon, Daucus carota, Epilobium angustifolium, Habitat: Ovit Mountain foothills, meadows, roadside Holcus lanatus, Hypericum perforatum, Lactuca serriola, and hazelnuts area. Lapsana communis, Lolium perenne, Malva neglecta, Vegetation: Achillea millefolium, Alchemilla caucasica, Myosotis lazica, Phragmites austrialis, Plantago lanceolata, Artemisia vulgaris, Calystegia sepium, Capsella bursa- P. major, Polygonum persicaria, Pteridium aquilinum, pastoris, Carex pallescens, Centaurea glastifolia, Cephalaria Rubus discolor, R. hirtus, Rumex crispus, Scrophularia procera, Chenopodium album, Cichorium intybus, Conyza scopolii, Senecio vulgaris, Setaria glauca, Solanum nigrum, bonariensis, Corylus avellana, Daucus carota, Erigeron Sonchus oleraceus, Tanacetum macrophyllum, Trifolium acer, Equisetum arvense, Euphorbia helioscopia, Ferula pratense, T. repens, Tripleurospermum oreades, Urtica orientalis, Galium incanum, Holcus lanatus, Hypericum dioica, Verbascum thapsus, Vicia cracca, Xanthium perforatum, Lactuca serriola, Lamium album, Lapsana spinosum. communis, Lolium perenne, Medicago lupulina, Melilotus Associated plants with P. spuria: Acer campestre, officinalis, Paspalum dilatatum, Papaver bracteatum, Anethum graveolens, Daucus carota, Daucus carota sativus, Plantago lanceolata, P. major, Poa bulbosa, P. pratensis, Euphorbia nicaeensis, Heracleum sphondylium, Tamarix sp. Polygonum bistorta, Ranunculus kotschyi, Reseda lutea, Remark: This species was collected on Hypericum Rhinanthus angustifolius, Rumex acetosella, R. crispus, perforatum while feeding. Salvia verticillata, Sanguisorba minor, Seseli libanotis, 3.13. Tryphoninae Sonchus asper, S. oleraceus, Sorghum halepense, Rubus Aderaeon hamatum Kasparyan, 1971 discolor, R. hirtus, Rumex crispus, Thalictrum minus, Material examined: Erzurum: İspir, Çayırözü, Trifolium pratense, T. repens, Tripleuspermum oreades, 40°33′53.1″N, 40°53′24.2″E, 2123 m, 19.06.2014, 1♀; Vicia cracca. Pazaryolu, Toprakkale, 40°16′41.47″N, 40°59′56″E, 2246 Associated plants with I. maculator: not known. m, 19.06.2014, 2♂♂. Remark: This species was collected on Daucus carota Geographic distribution: Armenia, Azerbaijan, while feeding. Bulgaria, Georgia, Iran, Russia (), Turkey, Paraperithous gnathaulax (Thomson, 1877) and Ukraine. Material examined: Erzurum: Pazaryolu, Kumaşkaya, Habitat: Ovit Mountain foothills and meadow area. 40°31′50.523″N, 40°45′16.686″N, 1825 m, 19.06.2014, 1♀. Vegetation: Achillea millefolium, Alchemilla caucasica, Geographic distribution: Palearctic region. Allium armenum, Anthemis cretica, Capsella bursa-

52 KOLAROV et al. / Turk J Zool pastoris, Centaurea depressa, C. glastifolia, Cephalaria Habitat: Meadow area. procera, Cerinthe minor, Daucus carota, Euphorbia stricta, Vegetation: Alchemilla caucasica, Allium armenum, Ferula orientalis, Galium incanum, G. verum, Hypericum Anthemis cretica, Centaurea depressa, Cerinthe minor, perforatum, Juncus articulatus, Lamium album, Lapsana Euphorbia stricta, Ferula orientalis, Galium incanum, G. communis, Orchis mascula, Ornithogalum narbonense, verum, Juncus articulatus, Lapsana communis, Orchis Papaver bracteatum, Pedicularis camosa, Plantago mascula, Ornithogalum narbonense, Pedicularis camosa, lanceolata, P. major, Poa bulbosa, P. pratensis, Polygonum Poa pratensis, Polygonum bistorta, Ranunculus kotschyi, bistorta, Ranunculus kotschyi, Reseda lutea, Rhinanthus Rhinanthus angustifolius, Rumex acetosella, Thalictrum angustifolius, Rumex acetosella, R. crispus, Sanguisorba minus, Veronica gentianoides. minor, Thalictrum minus, Trifolium pratense, Veronica Associated plants with T. (T.) rutilator: not known. gentianoides, Vicia cracca. Tryphon (Tryphon) thomsoni Roman, 1939 Associated plants with A. hamatum: not known. Material examined: Giresun: Tirebolu, Balçıkbelen, Ctenochira angulata (Thomson, 1883) 40°58.386′N, 38°52.576′E, 71 m, 22.06.2014, 1♂. Material examined: Rize: Çamlıhemşin Çat, Geographic distribution: Armenia, Azerbaijan, 40°53.230′N, 40°55.746′E, 1054 m, 20.06.2014, 1♂. Europe, Georgia, Iran, Tajikistan, Siberia, and Turkey. Geographic distribution: Palearctic region. Habitat: Hazelnut area. Habitat: Semiwet, sloping, brushwood area. Vegetation: Anthemis tinctoria, Artemisia vulgaris, Vegetation: Alopecurus myosuroides, Arctium minus, Calystegia sepium, Carex echinata, Corylus avellana, Astragalus viciifolius, Atropa belladonna, Coronilla Daucus carota, Lapsana communis, Lolium perenne, orientalis, Crepis setosa, Dactylis glomerata, Dactylorhiza Medicago sativa, Poa longifolia, Plantago lanceolata, P. urvilleana, Eryngium giganteum, Euphrasia rostkoviana, major, Pteridium aquilinum, Rubus hirtus, Rumex crispus, Geranium purpureum, G. pyrenaicum, Heracleum Sonchus oleraceus, Tanacetum macrophyllum, Trifolium sphondylium, Hypericum hirsutum, Inula vulgaris, pratense. Lapsana communis, Lysimachia vulgaris, Matricaria Associated plants with T. (T.) thomsoni: Peucedanum chamomilla, Melica ciliata, Mentha longifolia, Orchis oreoselinum. palustris, Phleum phleoides, Pilosella hoppeana, Prunella vulgaris, Pteridium aquilinum, Ranunculus cappadocicus, 3.14. Zoogeographic characterization R. repens, Rhinanthus angustifolius, Rumex alpinus, Salvia The zoogeographical characterization mainly follows the verticillata, Sambucus nigra, Sedum alpestre, Senecio chorotype classification of the Near East fauna proposed viscosus, Symphytum asperum, Tanacetum macrophyllum, by Taglianti et al. (1999). After investigation of the recent Thalictrum minus, Telekia speciosa, Trifolium pratense, T. geographic distribution of the species listed above, they repens, Urtica dioica, Verbascum thapsus. can be divided into the following groups: Associated plants with C. angulata: not known. 1. Species with multiregional distribution – here belongs Polyblastus (Polyblastus) cothurnatus (Gravenhorst, Exeristes roborator. 1829) 2. Species distributed in two zoogeographical regions Material examined: Rize: Çamlıhemşin, Zilkale, (Holarctic/Palearctic and Oriental/Neotropical). 40°55.322′N, 40°57.477′E, 764 m, 20.06.2014, 2♂♂. Holarctic and Oriental ranges have the following Geographic distribution: Palearctic region. species: Agrypon clandestinum, Meringopus calescens Habitat: Semiwet, shrub, sloping area. calescens, Syrphoctonus tarsatorius, and Syrphophilus Vegetation: Arctium minus, Calystegia silvatica, Daucus bizonarius; Palearctic and Oriental ranges have the carota, Echium vulgare, Erigeron acer, Galium aparine, species Anomalon cruentatum and Pimpla spuria; Geranium purpureum, Heracleum sphondylium, Inula Palearctic (Asiatic-European) and Afrotropical vulgaris, Lapsana communis, Mycelis muralis, Paspalum ranges have the species Meloboris collector. dilatatum, Phleum phleoides, Plantago major, Prunella 3. With Holarctic ranges are the following species: vulgaris, Pteridium aquilinum, Ranunculus repens, Rumex Apophua bipunctoria, Lissonota (Lissonota) folii, L. alpinus, Salvia verticillata, Tanacetum macrophyllum, (Loxonota) histrio, Buathra laborator, Mesostenus Trifolium repens, Urtica dioica. albinotatus, Cratichneumon viator, Exochus mitratus, Associated plants with P. (P.) cothurnatus: not known. Megastylus (Megastylus) flavopictus, and Gregopimpla Tryphon (Tryphon) rutilator (Linnaeus, 1761) inquisitor. Material examined: Erzurum: Pazaryolu, Toprakkale, 4. Most numerous are the species with Palearctic ranges. 40°16′41.47″N, 40°59′56″E, 2246 m, 19.06.2014, 1♀. Here belong 13 species – Dusona terebrator, Gelis Geographic distribution: Armenia, Azerbaijan, agilis, Glyphicnemis profligator, Schreineria populnea, Europe, Georgia, Iran, Siberia, and Turkey. Xylophrurus lancifer, Diplazon pallicoxa, Woldstedtius

53 KOLAROV et al. / Turk J Zool

biguttatus, Obtusodonta equitatoria, Exochus 10. Turano-European distribution is seen for the suborbitalis, Itoplectis maculator, Paraperithous species Glyphicnemis vagabunda, Ischnus alternator, gnathaulax, Ctenochira angulata, and Polyblastus and Aderaeon hamatum (Turano-East European (Polyblastus) cothurnatus. distribution). 5. Asiatic European distribution has the species 11. European ranges contain 10 species: Campoletis Pristomerus armatus, Meringopus titillator, and agilis, C. latrator, Olesicampe cavigena, O. patellana, Virgichneumon callicerus. Temelucha genalis, Aptesis assimilis, A. senicula, 6. Four species have a Sibero-European range: Diadegma Endasys plagiator, Rhembobius quadrispinus, and combinatum, Cryptus viduatorius, Tryphon (Tryphon) Oxytorus luridator. rutilator, and T. (T.) thomsoni. 12. One species, Lissonota (Loxonota) mediterranea, has 7. With Central Asiatic-European ranges are the species a Mediterranean range. Barylypa delictor and Lissonota (Lissonota) nitida. 8. Central Asiatic distribution is seen for the species Acknowledgments Meringopus calescens persicus. The preparation of this article was supported by Project 9. With Turano-Europeo-Mediterranean range are the Nr. BAP – 2012/234 (Atatürk University) and we are very species Phaenolobus fulvicornis, Dusona stygia, and thankful for it. We are also indebted to Dr Erol Yıldırım Ichneumon proletarius. (Atatürk University, Erzurum) for collecting specimens.

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