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Home , Asselian, Bactritida, Carboniferous, Cycloceras, Geisonoceratidae, Gzhelian

Paleontological Research, vol. 1, no. 1, pp.47-54, 3 Figs., April 3D, 1997 © by the Palaeontological Society of Japan

4. Late () to early () non-ammonoid from the Taishaku Group, Southwest Japan

SHWI NIKO and TOMOWO OZAWA

Department of Environmental Studies, Faculty of Integrated Arts and Sciences, Hiroshima University, Higashihiroshima, 739 Japan Department of and Planetary Sciences, Nagoya University, Chikusa-ku, Nagoya, 464-01 Japan

Received 6 June 1996; Revised manuscript accepted 21 December 1997

Abstract The non-ammonoid fauna described here was collected from the upper Gzhelian­ lower Asselian (Upper Carboniferous to Lower Permian) hydrozoan-algal buildups in the Taishaku Lime­ stone Group, Southwest Japan. The fauna consists of four orthocerids: ? sp., Bogoslovs­ kya miharanoroensis sp. nov., Geisonocerina? sp., Lopingoceras hayasakai sp. nov.; two nautilids: Parachouteauoceras bingoense gen. et sp. nov., Parachouteauoceras? sp.; and a bactritid: Bactrites sp. Parachouteauoceras is most similar to Chouteauoceras, but unlike the latter has a lobated aperture. The occurrence of Bogoslovskya miharanoroensis represents one of the latest records of the genus in the world. This fauna is important because it adds some new data to a poorly known Late Carboniferous­ Early Permian non-ammonoid cephalopod fauna.

Key words: Asselian, , Gzhelian, , , Taishaku Limestone Group

Introduction Permian greenstone-limestone sequence occurring in the Akiyoshi Terrane in the Inner Zone of Southwest Japan. The non-ammonoid cephalopods of the Taishaku Lime­ Hase et al. (1974) conducted a detailed facies analysis of the stone Group, Southwest Japan, were first noticed by Hayasa­ Taishaku Limestone Group and interpreted that it was for­ ka and Nishikawa (1963) who reported the orthocerid species med as a table complex on the submarine volcanic cf. adrianense Gemmellaro in an oral presentation mound. In Miharanoro, Hiba-gun, Hiroshima Prefecture (Koizumi, 1975). Of the fauna, only three specimens were (Figure 1), Upper Carboniferous to Lower Permian hydrozoan­ kept back for future examination. Recent studies, as de­ algal reef complexes are well developed, forming a reef core scribed by Niko et aI. (1993), show that the specimens are facies in the Taishaku Limestone Group. The Upper Car­ referable to a bactritid and represent a new species, Aktas­ boniferous to Lower Permian sequence of the Taishaku tioceras nishikawai. An additional seven species belonging Limestone Group in Miharanoro is divided into the following to the Orthocerida, Nautilida and Bactritida are described six fusulinacean zones (Ozawa and Hirakawa, MS): in and figured herein from the upper Gzhelian to lower Asselian ascending order, these are the Triticites ozawai-Carbono­ part of the Taishaku Limestone Group. This report adds schwagerina morikawai Zone, Triticites contractus Zone, some new data to a poorly known non-ammonoid ce­ Pseudoschwagerina muongthensis Zone, Pseudoschwage­ phalopod fauna in the vicinity of the Carboniferous-Permian rina miharanoensis Zone, Pseudofusulina vulgaris Zone, and boundary, and will provide a basis for future biostratigraphic Pseudofusulina kraffti Zone. The stratigraphic horizon of and paleobiogeographic studies. Aktastioceras nishikawai belongs to the Pseudoschwagerina All specimens used for the study are housed in the miharanoensis Zone, indicating a late Asselian . On the paleontological collections of the Department of Earth and other hand, localities of the present non-ammonoid Planetary Sciences, Nagoya University (ESN). cephalopods are in the Triticites contractus Zone, which coincides with the Carbonoschwagerina minatoi-Daixina cf. Geologic setting and occurrence robusta Zone (upper Gzhelian) and the Sphaeroschwagerina fusiforrnis Zone (lower Asselian) of the Akiyoshi Limestone The Taishaku Limestone Group, located in the northeast­ Group (Ozawa and Kobayashi, 1990). ern part of Hiroshima Prefecture, is a thick Carboniferous to The cephalopods described herein were collected by the 48 Shuji Niko and Tomowo Ozawa

Michelinoceras? sp. Figures 2-14-16 Description.-Smooth-surfaced with gradual shell expansion, circular cross section; largest specimen (ESN 2593) of phragmocone 3.9 mm in adoral diameter; sutures straight, transverse; central, composed of orthochoanitic necks and cylindrical connecting rings. Discussion.-This description is based on three poorly preserved phragmocones of probably immature portions. Judging from the siphuncular structure, they may represent a species of Michelinoceras. However, there is no denying the possibility that the specimens are apical shells of as­ sociated Geisonocerina ? sp. (this report) until better material is recovered. Material and QCCurrence.-ESN 2593-2595 from Locality 1.

Genus Bogoslovskya Zhuravleva, 1978

Type species.-Bogoslovskya perspicua Zhuravleva, 1978.

Bogoslovskya miharanoroensis sp. nov. Figures 2-1-9 Diagnosis.-Species of relatively large Bogoslovskya with distant, very narrow ribs; cameral ratio approximately 3-4; siphuncular position midway between center and margin in Figure 1. Part of topographic map "Tojo", 1: 25,000 adoral shell. quadrangle of Geographical Survey Institute, showing fossil Description.-Shells relatively large for genus, orthoconic localities in Miharanoro, Hiroshima Prefecture. with circular cross section, largest specimen (ESN 2590) of phragmocone reaches approximately 25 mm in diameter; angle of shell expansion moderate, 4-5 degrees; surface junior author from bioclastic in channellike ornamentation consists of distant, very narrow ribs, weakly depressions of the Triticites contractus Zone. In the chan­ sinuate, slightly toward aperture on antisiphuncular side, rib nel-fill sediments, they occurred along with ammonoids, index (number of ribs per length of corresponding shell gastropods, pelecypods, , and foraminifers. diameter) 13-17, interribs space flat, smooth; septa moder­ ately deep forming straight, transverse sutures; cameral Systematic length short for genus, cameral ratio (diameter/length) 3.2- 4.1 ; siphuncle very narrow, maximum external diameter of Cephalopoda Cuvier, 1797 septal neck/shell diameter 0.03 in holotype, siphuncular Subclass Nautiloidea Agassiz, 1847 position submarginal, but shifts midway between center and Order Orthocerida Kuhn, 1940 margin in adoral shell, minimum distance of siphuncular axis Superfamily Orthocerataceae M'Coy, 1844 from shell surface per shell diameter in dorsoventral section Family M'Coy, 1844 0.19 in immature shell (9.5 mm in diameter) of paratype (ESN Subfamily Michelinoceratinae Flower, 1945 2589), increases to 0.26 adorally (20 mm in diameter); length Genus Michelinoceras Foerste, 1932 of septal necks moderately long for genus, approximately 1.8 mm at shell diameter of 21 mm, gently tapering ortho­ Type species.-Orthoceras michelini Barrande, 1866. choanitic with weak auxiliary deposits in funnel-shaped septal foramen; connecting rings not preserved; no camer-

Figure 2. 1-9. 8ogoslovskya miharanoroensis sp. nov., 1-4: holotype, ESN 2586, 1, side view of siphuncular side, x2, 2, lateral view, siphuncular side on right, x 2, 3, septal view of apical end, x 2, 4, dorsoventral polished section, arrow indicates siphuncular position, x 3, 5, 6: paratype, ESN 2587, 5, lateral view, siphuncular side on left, x 2, 6, dorsoventral thin section, details of siphuncular structure, note auxiliary deposits, x 10, 7: paratype, ESN 2589, septal view of apical end, x 2, 8: paratype, ESN 2591, lateral view, siphuncular side on right, x 2, 9: paratype, ESN 2590, lateral view, siphuncular side on left, x2. 10-13. Geisonocerina? sp., 10-12: ESN 2596, 10, side view, x2, 11, septal view, x2, 12, longitudinal polished section, x3, 13: ESN 2597, side view, x2. 14-16. Michelinoceras? sp., ESN 2593, 14: side view, X4, 15: septal view of apical end, x 4, 16: longitudinal thin section, x 4. Gzhelian- Asselian cephalopods from Taishaku 49

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15 50 Shuji Niko and Tomowo Ozawa al deposits detected. Lopingoceras hayasakai sp. nov. Discussion.-The possession of surface ribs of Bogos/ov­ Figures 3-1-7 skya miharanoroensis sp. nov. clearly distinguishes it from all other known species of the genus. The range of Bogos/ov­ Diagnosis.-Annulated orthocones with conspicuous sur­ skya was previously restricted to the Middle to Late face lirae, annulations have subangular profile; cameral of the Urals (Zhuravleva, 1978) and the Middle Carboniferous deposits episeptal-mural, hyposeptal; endosiphuncular of the Akiyoshi Limestone Group (Niko et aI., 1995). Thus, deposits form continuous lining. this late Gzhelian to early Asselian species represents the Description.-Annulated orthocones with gradual shell youngest record of the genus. expansion, angle approximately 3-4 degrees; cross section Etymo/ogy.-The specific name is derived from the type of shell circular; largest specimen (ESN 2600) of phrag­ locality name Miharanoro. mocone reaches approximately 10 mm (reconstructed) in Material and occurrence.-Holotype, ESN 2586, incomplete diameter; annulations relatively low, subangular profile in phragmocone, 22.5 mm in length; five paratypes, ESN longitudinal section, bearing weak lateral sinus; annulations 2587-2591, incomplete phragmocones, are surely, and a and interspaces of annulations ornamented by conspicuous specimen, ESN 2592, is questionably, assigned to this lirae that run parallel with annulation, salient forms over species. All specimens from Locality 1. injured portion of shell as repaired; sutures straight, trans­ verse; septal curvature moderate, cameral ratio approxi­ Family Geisonoceratidae Zhuravleva, 1959 mately 2.2 in holotype; there are usually two annulations in Genus Geisonocerina Foerste, 1935 single ; siphuncle subcentral, shifts slightly dorsally from center, septal necks short, suborthochoanitic to weakly Type species.-Orthoceras wauwatosense Whitfield, 1882. cyrtochoanitic, 0.45 mm in length in holotype, connecting rings subcylindrical with constrictions at septal foramen and Geisonocerina? sp. weak dorsal inflations, maximum external diameter of con­ necting rings/shell diameter in each segment of holotype Figures 2-10-13 approximately 0.2; cameral deposits episeptal-mural, Description.-Orthocones with gradual shell expansion, hyposeptal; endosiphuncular deposits well developed, form circular cross section; surface ornamentation consists of thick continuous lining on siphuncular wall, thicker in venter distinct transverse lirae, adorally indicating weak sinuations ; than dorsum. largest specimen (ESN 2597) of phragmocone reaches Discussion.-Generic assignment of the species is tenta­ approximately 16 mm (reconstructed) in diameter; septa tive at present. A combination of surface annulations and thick, deeply concave, forming straight, transverse sutures; conspicuous lirae relates the form to Reticycloceras Gordon, siphuncle central; septal necks long, orthochoanitic, but 1960, but the possession of hyposeptal deposits in the slightly dilated terminally; connecting rings cylindrical. species clearly separates them. Lopingoceras Shimanskiy Discussion.-This description is based on two fragmentary in Ruzhentsev (1962) is based on an Upper Permian species phragmocones. They are not sufficiently well preserved to from Dzhul'fa, of which species has an annulated . identify more precisely, but lack of periodic thickening of the The distinguishing feature of Lopingoceras is angularity of lirae suggests this species probably belongs to Geisonocer­ outline of annulations, and the present species shears this ina rather than Geisonoceras Hyatt, 1884. morphology. Although the structure of deposits is unknown Material and occurrence.-ESN 2596, 2597. In addition, a in the generic type species, we consider that the present specimen (ESN 2598) is questionably assigned to this species probably belongs to Lopingoceras. species. All specimens from Locality 1. Shimanskiy (1954, pI. 7, figs. 6a-v) referred a species from the lower Asselian of the southern Urals to Cycloceras Superfamily Pseudorthocerataceae Flower and Caster, 1935 laevigatum M'Coy (1844, pI. 2, fig. 3), which was described Family Pseudorthoceratidae Flower and Caster, 1935 from the Lower Carboniferous of . With the excep­ Subfamily Spyroceratinae Shimizu and Obata, 1935 tion of a somewhat larger angle of shell expansion (approxi­ Genus Lopingoceras Shimanskiy in Ruzhentsev, 1962 mately 6 degrees), the Russian species is very likely closely related to Lopingoceras hayasakai sp. nov. Cycloceras was Type species.-Orthoceras /opingense Stoyanow, 1909. defined on an internal mold of a body chamber by M'Coy

Figure 3. 1-7. Lopingoceras hayasakai sp. nov., 1-4: holotype, ESN 2599, 1, ventral view, X2, 2, lateral view, venter on right, x 2, 3: dorsoventral thin section, venter on left, note subangular profile of annulations, x 4, 4, details of siphuncular structure, note thick continuous lining of endosiphuncular deposits, x10, 5-7: paratype, ESN 2600, 5, lateral view, venter on right, x2, 6, dorsal view, x2, 7, septal view of apical end, venter down, x2. 8-11. Bactrites sp., ESN 2607, 8: ventral view, x 2, 9: lateral view, venter on right, x 2, 10: dorsal view, x 2, 11 : septal view of apical end, venter down, x 2. 12- 15. Parachouteauoceras bingoense gen. et sp. nov., 12-14: holotype, ESN 2601, 12, lateral view, note color markings through ammonium chloride coating, x3, 13, ventral view, x3, 14, septal view of apical end, venter up, x3, 15: paratype, ESN 2602, lateral view with complete embryonic shell, x3. 16-18. Parachouteauoceras? sp., ESN 2604, 16: lateral view, x3, 17: ventral view, x 3, 18: septal view of apical end, venter up, x 3. Gzhelian- Asselian cephalopods from Taishaku SI

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13 17 52 Shuji Niko and Tomowo Ozawa

(1844), thus the diagnosis of the genus calls for nothing Ireland. All occurrences are in the Lower Carboniferous. further than an annulated orthocone with low angles of Parachouteauoceras can be distinguished from expansion. Sweet (1964) properly stated that "no species Chouteauoceras by its peristome shape. The growth lines other than the type species should be referred to Cycloceras of Chouteauoceras indicate a transverse peristome with until its type is better known". This problem is still unsolved. weak lateral saddles. Etymology.-The specific name refers to the late Dr. Ichiro Apogonoceras remotum Ruzhentsev and Shimanskiy Hayasaka, a pioneer in the study of the (1954, pI. 11, figs. 8a, 8b; only known species and the type of Japan. species of the genus) from the of the southern Urals has a shell shape generally similar to Para­ Material and occurrence.-Holotype, ESN 2599, incomplete chouteauoceras bingoense, but has nearly straight sutures in phragmocone, 20.1 mm in length; paratype, incomplete the adoral shell and a somewhat subtriangular whorl section. phragmocone, ESN 2600. Both from Locality 2. The peristome shape of Apogonoceras is unknown. Etymology.-The specific name is derived from Bingo, which is the historic province name of the region in which Order Nautilida Agassiz, 1847 the type locality lies. Superfamily Trigonocerataceae Hyatt, 1884 Material and occurrence.-Holotype, ESN 2601, incomplete Family Trigonoceratidae Hyatt, 1884 phragmocone with apical body chamber, 25.5 mm in length; Genus Parachouteauoceras gen. nov. two paratypes, ESN 2602, 2603, apical phragmocones. All specimens from Locality 2. Type species.-Parachouteauoceras bingoense sp. nov. Diagnosis.-Like Chouteauoceras but differs in possession of lobed peristome with linguiform ventral sinus, ventrolateral Parachouteauoceras? sp. saddle, dorsolateral sinus, dorsal saddle. Figures 3-16-18 Etymology.-The generic name is derived from the Greek para, meaning near, and Chouteauoceras. Description.-Loosely coiled juvenile phragmocones losing contact; whorl section compressed, ovoid, width/height Parachouteauoceras bingoense sp. nov. ratio 0.90; shell height reaches 4.9 mm in largest specimen ESN 2604; surface ornamentation consists of weak longitu­ Figures 3-12-15 dinal ridges and growth lines indicating ventral sinus; sutures nearly transverse; siphuncie subcentral. Diagnosis.-As for the genus. Discussion.-This species is represented by two fragmen­ Description.-Loosely coiled, gyrocones losing contact; tary phragmocones of juvenile portion. The specimens whorl section weakly compressed, width/height ratio ranges differ from Parachouteauoceras bingoense gen. et sp. nov. from 0.90 to 0.97 in holotype, venter and flanks slightly (this report) in smaller shell size in the corresponding shell convex with subangular ventral shoulders, dorsum broadly portion. The lack of adult shells of this species leaves rounded; adoral end of body chamber of holotype reaches doubt upon assignment to Parachouteauoceras. 8.6 mm in height; embryonic shell weakly inflated in dor­ Material and QCCurrence.-ESN 2604, 2605 from Locality 1. soventral direction, with cone-shaped primary shell; most inflated portion of embryonic shell 5.3 mm in height, 4.7 mm in width, giving a ratio of 0.89; surface ornamentation Subclass Bactritoidea Shimanskiy, 1951 consists of numerous longitudinal ridges and growth lines; Order Bactritida Shimanskiy, 1951 growth lines nearly transverse, producing a latticework in Family Bactritidae Hyatt, 1884 embryonic shell; later growth lines indicate lobed peristome Genus Bactrites Sandberger, 1843 of deep linguiform ventral sinus, rounded ventrolateral sad­ dle, shallow dorsolateral sinus, broadly rounded dorsal sad­ Type species.-Bactrites subconicus Sandberger, 1843. die; sutures transverse with broadly rounded lateral lobes, dorsal and ventral saddles; siphuncle small, subcentral, Bactrites sp. slightly shifts to venter; preserved color markings consist of Figures 3-8-11 7 or 8 relatively wide bands, which run parallel with peris­ tome, in ventral shoulder to ventrolateral position on Description.-Orthocones with gradual shell expansion, holotype. angle approximately 3.5 degrees, circular cross section; Discussion.-The gyroceraconic shells with a compressed largest specimen (ESN 2607) of phragmocone reaches 11.5 whorl section, the longitudinal surface ridges, and rounded mm in diameter; transverse lirae of surface ornamentation lateral lobes in the sutures of Parachouteauoceras bingoense slightly sinuate, very fine, closely spaced; sutures straight, gen. et sp. nov. suggest a relationship to the trigonoceratid transverse with ventral lobe, siphuncle relatively large, ventral genus Chouteauoceras Miller and Garner, 1953. margin in position. Chouteauoceras is a rare genus which embraces the type Discussion.-This species resembles most closely species C. americanum (Miller and Furnish, 1938, pI. 48, figs. Bactrites costatus Mapes (1979, pI. 31, figs. 7-9, 13) from the 4-6) from Missouri and some other species questionably upper Virgilian of Texas in having closely spaced surface assigned to the genus from , Belgium and lirae. Bactrites costatus differs from the present species in Gzhelian-Asselian cephalopods from Taishaku 53

its larger angle (5-7 degrees) of shell expansion. Miller, A.K. and Garner, H.F., 1953: Lower Material and occurrence.-Based on two phragmocones: cephalopods of Michigan. Part II. Coiled nautiloids. ESN 2606 from Locality 1, and ESN 2607 from Locality 2. Contributions from the Museum of Paleontology, Uni­ versity of Michigan, vol. 11, p.111-151, pis. 1-4. Niko, S., Nishida, T. and Hamada, T., 1993: Aktastioceras Acknowledgments nishikawai n. sp., a first Permian bactritoid cephalopod from Japan. Journal of Paleontology, vol. 67, p. 314- The second author is deeply indebted to Mitsuaki Hira­ 316. kawa and Akihiko Shioe, former post-graduate students of Niko, S., Nishida, T. and Kyuma, Y., 1995: A new Carbonif­ the Hyogo University of Teacher Education, for their kind erous cephalopod Bogos/ovskya akiyoshiensis from help in sampling. Southwest Japan. Transactions and Proceedings of the Paleontological Society of Japan, New , no. References cited 179, p. 193-195. Ozawa, T. and Hirakawa, M., MS: Foraminiferal biostratigra­ Barrande, J., 1866: Systeme silurien du centre de la Bo­ phy and sedimentary facies of the Taishaku Limestone heme, Premiere Partie: Recherches paleontologiques, Group in the Akiyoshi Terrane, southwest Japan. 14 p., vol. 2, Classe des Mollusqes, Ordre des Cephalopodes. 3 pis. Part 7, pis. 108-244, Privately published, Prague and Ozawa, T. and Kobayashi, F., 1990: Carboniferous to Per­ Paris. mian Akiyoshi Limestone Group. Guidebook for Field Flower, RH., 1945: Classification of Devonian nautiloids. Trip No.4, Benthos '90, the Fourth International Sympo­ The American Midland Naturalist, vol. 33, p. 675-724. sium on Benthic , Sendai, p. E1-E31, pis. 1- Flower, RH. and Caster, K.E., 1935: The stratigraphy and 13. paleontology of northwestern Pennsylvania. Part II : Ruzhentsev, V.E., 1962: Fundamentals of Paleontology Paleontology. Section A: The cephalopod fauna of (Osnovy paleontologil). Vol. V. -Cephalopoda 1. the Conewango Series of the Upper Devonian in New 425 p., 32 pis. Izdatel'stvo Akademii Nauk SSSR, York and Pennsylvania. Bulletins of American Moskva. (translated from RUSSian, Israel Program for Paleontology, vol. 22, p.199-271. Scientific Translations, Jerusalem, 1974) Foerste, A.F., 1932: Black River and other cephalopods from Ruzhentsev, V.E. and Shimanskiy, V.N., 1954: Nizhneperm­ Minnesota, Wisconsin, Michigan, and Ontario (Part 1). skie svernutie i sognutie nautiloidei yuzhnogo Urala Denison University Bulletin, Journal of the Scientific (Lower Permian coiled and curved nautiloids of the Laboratories, vol. 27, p.47-136, pis. 7-37. southern Urals). Akademiia Nauk SSSR, Trudy Foerste, A.F., 1935: Big Horn and related cephalopods. Paleontologicheskogo Instituta, vol. 50, p.1-152, pis. 1- Denison University Bulletin, Journal of the Scientific 15. (in Russian) Laboratories, vol. 30, p.1-96, pis. 1-22. Sandberger, G., 1843: Schilderung der palaontologischen Gordon, M.Jr., 1960: Some American Midcontinent Carboni­ Verhaltnisse der alteren Formationen Nassaus. Ver­ ferous cephalopods. Journal of Paleontology, vol. 34, p. sammlung Deutscher Naturforscher und Aerzte Mainz, 133-151, pis. 27, 28. Bericht 20, p.154-160. Hase, A., Okimura, Y. and Yokoyama, T., 1974: The Upper Shimanskiy, V.N., 1954: Pryamye nautiloidei i baktritoidei Paleozoic formations in and around Taishaku-dai, sakmarskogo i artinskogo yarusov yuzhnogo Urala Chugoku Massif, Southwest Japan; with special refer­ (Straight nautiloids and bactritoids from the ence to the sedimentary facies of limestones. Geol­ and Artinskian stages of the Southern Urals). ogical Report of the Hiroshima University, no. 19, p.1-39, Akademiia Nauk SSSR, Trudy Paleonto/ogicheskogo pis. 1-8. (in Japanese with English abstract) Instituta, vol. 44, p. 1-156, pis. 1-12. (in Russian) Hayasaka, I. and Nishikawa, I., 1963: On some Permian Shimizu, S. and Obata, T., 1935: New genera of Gotlandian macrofossils from Hiroshima Prefecture, Japan. (A pre­ and nautiloids. Journal of the Shanghai liminary note). (Palaeontological Society of Science Institute, Section 2, Geology, Paleontology, Japan), no. 6, p.27. (in Japanese) Mineralogy, and Petrology, vol. 2, p.1-1O. Hyatt, A., 1883-1884: Genera of fossil cephalopods. Pro­ Stoyanow, A.A., 1909: On the character of the boundary of ceedings of the Boston Society of Natural History, vol. Paleozoic and near Djulfa. The Diary of the 22, p. 253-338. Xllth Congress of Russian Naturalists and Physicians in KOizumi, H., 1975: Paleozoic cephalopods of Japan. 149 p. , no. 4, p. 142. (not seen) Privately published, Chiba. (in Japanese) Sweet, W.C., 1964: Nautiloidea-Orthocerida. In, Moore, Mapes, RH., 1979: Carboniferous and Permian Bactritoidea RC. ed., Treatise on Invertebrate Paleontology. p. K216- (Cephalopoda) in North America. The University of K261. The Geological Society of America and the Paleontological Contributions, Article 64, p.1- University of Kansas Press. 75, pis. 1-41. Whitfield, RP., 1882: Palaeontology. Geology of Wiscon­ M'Coy, F., 1844: A synopsis of the characters of the sin, vol. 4, p.161-363, pis. 1-17. (not seen) fossils of Ireland. 274 p. Zhuravleva, FA, 1959: 0 semeistve Michelinoceratidae Privately published. (reissued by Williams and Norgate, Flower, 1945 (On the family Michelinoceratidae Flower, London, 1862) 1945). Materialy k "Osnovam paleontologii", part 3, p. Miller, A.K. and Furnish, W.M., 1938: Lower Mississippian 47-48. (in Russian) cephalopods of Missouri. The University of Zhuravleva, FA, 1978: Devonskiye ortocerody, nadotryad Missouri Studies, vol. 13, p.149-178, pis. 38-48. Orthoceratoidea (Devonian orthocerids, superorder 54 Shuji Niko and Tomowo Ozawa

Orthoceratoidea). Akademiia Nauk SSSR, Trudy Russian) Pa/eont%gicheskogo /nstituta, vol. 168, p.1-223. (in

Bingo frm~, Hiba-gun Jt~w, Miharanoro =.W-lIlf§, Taishaku'i'iI;fR, Tojo *~