Ecologica Montenegrina 22: 152-156 (2019) This journal is available online at: www.biotaxa.org/em

To the fauna (, Pterophoridae) of Myanmar (Burma)

*PETER USTJUZHANIN1,2, VASILIY KOVTUNOVICH3, & ALEXANDER STRELTZOV4

1 Altai State University, Lenina 61. RF-656049. Barnaul, Russia. E-mail: [email protected] 2 Biological Institute, Tomsk State University, Lenina Prospekt 36, Tomsk 634050, Russia. E-mail: [email protected] 3Moscow Society of Nature Explorers, Bolshaya Nikitskaya 2, Moscow, 125009, Russia. E-mail: [email protected] 4Herzen State Pedagogical University of Russia, 48, Moika Emb., Saint-Petersburg, 191186, Russia. E-mail: [email protected] *Corresponding author

Received: 21 June 2019│ Accepted by V. Pešić: 4 September 2019 │ Published online: 8 September 2019.

The article gives a faunistic review of the Pterophoridae fauna in Myanmar. The type habitats and world distribution (with new data) is indicated. The data of the collected specimens are provided. To the 10 already known species we have added 9 previously unknown for the fauna of Myanmar.

The state Myanmar (former Burma) is located in western portion of Indochina peninsula. It is bordered by , , Thailand and Bangladesh. The climate of Myanmar is tropical and sub-equatorial. Most of the country is occupied by medium-high mountain plateaus included in the Himalayan system. The mountains are richly wooded, with a large variety of tropical and plant species, including endemics. The information on the Pterophoridae of Myanmar is very scarce. According to literary data, 10 Pterophoridae species were known for this territory (Arenberger 1995, 2002, 2010; Gielis, 2003; Li, Hao & Wang 2003; Meyrick, 1905, 1908; Walsingham, 1891). The materials on Pterophoridae collected by our colleagues in Myanmar, served us to write this work, and as a result, we have added 9 species to the fauna of Myanmar. Thus, considering the literary data, currently there are 19 Pterophoridae species known for the fauna of Myanmar. The species indicated for the Myanmar fauna for the first time are marked with an asterisk *. The examined material is deposited in the private collection of the two first authors.

List of species

Ochyrotica connexiva (Walsingham, 1891) Steganodactyla connexiva Walsingham, 1891: 242. (Type locality: Burma) Distribution. Bhutan, India, Myanmar, Thailand. Note. This species was recorded from Myanmar (Walsingham, 1891).

Deuterocopus planeta Meyrick, 1908 Deuterocopus planeta Meyrick, 1908: 473. (Type locality: India). Distribution. India, , Myanmar, , New Guinea, Palau.

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Note. This species was recorded from Myanmar (Gielis, 2003).

Deuterocopus socotranus Rebel, 1907 Deuterocopus socotranus Rebel, 1907: 115. (Type locality: Yemen, Sokotra). Distribution. Yemen, Oman, Japan, India, Sri Lanka, Thailand, Taiwan, Myanmar, Indonesia, New Guinea, Australia, Somalia, Uganda, , Nigeria, Republic of . Note. This species was recorded from Myanmar (Gielis, 2003).

Stenoptilodes taprobanes (Felder & Rogenhoffer, 1875) Amblyptilia taprobanes Felder & Rogenhofer, 1875: plate 140, fig. 54. (Type locality: Sri Lanka). Material examined: 2 ex., N. Myanmar, 21 km E. Putao, Nan Sa Bon vill, h=550m, 1-5.V.1998, S. Murzin & V. Sinyaev; 1 ex., Myanmar, Lake Inle, 1-3.XII.2009, N.Vikhrev; 1 ♀, Myanmar, Rakhine State, disturbed tropical forest, 18.6215ºN, 94.3674ºE, (Fig. 1) 05.XII.2018, V. Spitsyn leg. Distribution: , Turkey, Indonesia, Sri Lanka, Thailand, Cambodia, India, Myanmar, New Guinea, Australia, Chad, Sao Tome, Cote d’lvoire, Nigeria, , Kenya, , Seychelles, USA, Paraguay, Bolivia. Note. This species was recorded from Myanmar (Li et al., 2003).

*Amblyptilia fibigeri Gielis, 1999 Amblyptilia fibigeri Gielis, 1999: 85. (Type locality: Nepal). Material examined: 1 ♂, N. Myanmar, 21 km E Putao, Nan Sa Bon vill, h=550m, 1-5.V.1998, S. Murzin & V. Sinyaev. Distribution. Nepal, Bhutan, Myanmar.

*Sphenarches bilineatus Yano, 1963 (Fig. 1) Sphenarches bilineatus Yano, 1963: 868. (Type locality: Samoa). Material examined: 1 ♀, Myanmar, Rakhine State, disturbed tropical forest, 17º34’54”N, 94º37’43”E, (Fig. 1) 03.XII.2018, V. Spitsyn leg. Distribution. Samoa, Thailand, Myanmar.

Sphenarches zanclistis (Meyrick, 1905) Oxyptilus zanclistis Meyrick, 1905: 582. (Type locality: Burma). Distribution. Nepal, India, Sri Lanka, Myanmar, Indonesia, Australia. Note. This species was recorded from Myanmar (Meyrick, 1905).

Oxyptilus epidectis Meyrick, 1908 Oxyptilus epidectis Meyrick, 1908: 476. (Type locality: Burma, India & Sri Lanka). Distribution. India, Sri Lanka, Myanmar. Note. This species was recorded from Myanmar (Meyrick, 1908).

Xyroptilia siami Kovtunovich & Ustjuzhanin, 2006 Xyroptilia siami Kovtunovich & Ustjuzhanin, 2006: 262. (Type locality: Chiang Mai, NW Thailand). Distribution. NW Thailand, Myanmar, China (Yunan) Note. This species was recorded from Myanmar (Arenberger, 2010)

*Megalorhipida leucodactyla (Fabricius, 1794) Pterophorus leucodactylus Fabricius, 1794: 346. (Type locality: Virgin Islands) Material examined: 1 ♂, Myanmar, Sagaing Distr., mountain tropical forest, 24º15’15”N, 96º15’05”E, h= 331 m, 13. XI.2018, V. Spitsyn leg. Distribution. Spain, Morocco, Israel, Lebanon, Jordan, Saudi Arabia, Yemen, Dubai, Oman, , Pakistan, Somalia, Congo, Nigeria, Cote d’lvoire, Cameroon, Chad, Liberia, Sierra Leone, The Gambia, Nigeria, Islands, Kenya, Tanzania, Swaziland, Rep. S. Africa, Uganda, Madagascar, Seychelles, Reunion, Mauritius, , China, Taiwan, , Thailand, Myanmar, Malaysia, India, Sri Lanka, Maldives, New Guinea, , Australia, USA, Argentina, Chile, Paraguay, Brazil, Ecuador, Colombia, Panama, French Guyana, Mexico, Peru, Venezuela, Bahamas, Jamaica, St. Thomas, St. Vincent, Puerto

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Rico, Virgin Islands, Suriname, Galapagos Islands, Mariana Islands, Wake Island, Marshall Islands, Gilbert Islands.

* atomosa (Walsingham, 1885) Aciptilia atomosa Walsingham, 1885: 885. (Type locality: India) Material examined: 1 ♀, Myanmar, Rakhine State, disturbed tropical forest, 18.6215ºN, 94.3674ºE, (Fig. 1) 05.XII.2018, V. Spitsyn leg. Distribution. India, Nepal, Myanmar, Cambodia, Ethiopia, Tanzania, Kenya, Swaziland, Madagascar.

*Exelastis pumilio (Zeller, 1873) Mimeseoptilus pumilio Zeller, 1873: 324. (Type locality: Texas, USA) Material examined: 1 ♀, Myanmar, Sagaing Distr., mountain tropical forest, 24º14’26”N, 96º14’45”E, h =334 m, 14. XI. 2018, V. Spitsyn leg. Distribution. Nepal, Cambodia, Myanmar, , New Guinea, Chad, The Gambia, Tanzania, Mali, Zimbabwe, Swaziland, Rep. S. Africa, Seychelles, Reunion Island, Madagascar, Nigeria, Kenya, USA., Argentina, Brazil, Ecuador, Bolivia, Colombia, Surinam, Costa Rica, Cuba, Puerto Rico, Guadeloupe, Mexico, Jamaica, Virgin Islands, Fatu Hiva, Guam, Palau.

(?) *Exelastis sp. Material examined: 1 ♀, Myanmar, Rakhine State, disturbed tropical forest, 18.6215ºN, 94.3674ºE, (Fig. 1) 05.XII.2018, V. Spitsyn leg. Note. This species was impossible to determine, it requires additional materials.

Tabulaephorus sesamitis (Meyrick, 1905) Alucita sesamitis Meyrick, 1905: 582. (Type locality: Burma). Distribution. Turkmenistan, Afghanistan, India, Myanmar. Note. This species was recorded from Myanmar (Meyrick, 1905).

* microdactyla (Hubner, [1813]) Alucita microdactyla Hubner, [1813]: pl. 5, figs. 26, 27. (Type locality: Europe). Material examined: 6 ex., N. Myanmar, 40 km N Myitkyina, Chane Kand vill., h=235m, 23-24.IV.1998; N. Myanmar, 21 km E Putao, Nan Sa Bon Vill, h=550m, 1-5.V.1998, S. Murzin & V. Sinyaev; 1 ex., Myanmar, Lake Inle, 1-3.XII.2009, N. Vikhrev. Distribution. Europe, Russia (European part), Turkey, Israel, Iran, Georgia, Nepal, China, Japan, Madagascar, Vietnam, Myanmar, Indonesia, Philippines, Taiwan, New Guinea, Solomon Islands.

*Pterophorus elaeopus (Meyrick, 1908) Alucita elaeopa Meyrick, 1908: 490. (Type locality: India). Material examined: 1 ♂, 1 ♀, N. Myanmar, 40 km N Myitkyina, Chane Kand vill., h=235m, 23-24.IV.1998, S. Murzin & V. Sinyaev. Distribution. India, Myanmar, Thailand, Malaysia, Indonesia.

Pterophorus lacteipennis (Walker, 1864) Aciptilus lacteipennis Walker, 1864: 949. (Type locality: India). Distribution. India, Myanmar, Malaysia, Singapore, Sarawak, Taiwan, Indonesia, New Guinea, Australia. Note. This species was recorded from Myanmar (Gielis, 2003).

*Pterophorus leucadactylus (Walker, 1864) Aciptilus leucadactylus Walker, 1864: 949. (Type locality: Sri Lanka). Material examined: 1 ♀, N. Myanmar, 50 km NW Putao, Wa Sa Dam vill., h=950m, 17.V.1998, S. Murzin & V. Sinyaev; 1 ♂, Myanmar, Rakhine State, disturbed tropical forest, 18.6215ºN, 94.3674ºE, (Fig. 1) 05.XII.2018, V. Spitsyn leg. Distribution. India, Sri Lanka, Myanmar, Thailand, Vietnam, Indonesia.

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Figure1. Biotope: Rakhine State, disturbed tropical forest. Foto V.Spitsyn.

Cosmoclostis leucomochla Fletcher, 1947 Cosmoclostis leucomochla Fletcher, 1947: 46. (Type locality: Sri Lanka). Distribution. Sri Lanka, Myanmar, Philippines. Note. This species was recorded from Myanmar (Gielis, 2003).

As a result of examining the material on Pterophoridae, 19 species have been found for Myanmar, nine of them are new to the fauna of the country. There is no doubt that further research on plume of Myanmar will significantly increase this list.

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Acknowledgments The authors express their gratitude to their colleagues: V. Spitsyn, (Arkhangelsk, Russia), S. Murzin, V. Sinyaev, N. Vikhrev, (Mosсow, Russia) for providing material for examination.

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