Redalyc.Global Series and Stages for the Ordovician System: a Progress
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Paleontological Contributions Number 18 Efficient Ornamentation in Ordovician Anthaspidellid Sponges Stephen B. Church August 9, 2017 Lawrence, Kansas, USA ISSN 1946-0279 (online) paleo.ku.edu/contributions Ridge-and-trough ornamented outer-wall fragment of the Ordovician anthaspidellid sponge Rugocoelia eganensis Johns, 1994. Paleontological Contributions August 9, 2017 Number 18 EFFICIENT ORNAMENTATION IN ORDOVICIAN ANTHASPIDELLID SPONGES Stephen B. Church Department of Geological Sciences, Brigham Young University, Provo, Utah 84602-3300, [email protected] ABSTRACT Lithistid orchoclad sponges within the family Anthaspidellidae Ulrich in Miller, 1889 include several genera that added ornate features to their outer-wall surfaces during Early Ordovician sponge radiation. Ornamented anthaspidellid sponges commonly constructed annulated or irregularly to regularly spaced transverse ridge-and-trough features on their outer-wall surfaces without proportionately increasing the size of their internal wall or gastral surfaces. This efficient technique of modi- fying only the sponge’s outer surface without enlarging its entire skeletal frame conserved the sponge’s constructional energy while increasing outer-wall surface-to-fluid exposure for greater intake of nutrient bearing currents. Sponges with widely spaced ridge-and-trough ornament dimensions predominated in high-energy settings. Widely spaced ridges and troughs may have given the sponge hydrodynamic benefits in high wave force settings. Ornamented sponges with narrowly spaced ridge-and- trough dimensions are found in high energy paleoenvironments but also occupied moderate to low-energy settings, where their surface-to-fluid exposure per unit area exceeded that of sponges with widely spaced surface ornamentations. Keywords: lithistid sponges, Ordovician radiation, morphological variation, theoretical morphology INTRODUCTION assumed by most anthaspidellids. -
Cambrian Ordovician
Open File Report LXXVI the shale is also variously colored. Glauconite is generally abundant in the formation. The Eau Claire A Summary of the Stratigraphy of the increases in thickness southward in the Southern Peninsula of Michigan where it becomes much more Southern Peninsula of Michigan * dolomitic. by: The Dresbach sandstone is a fine to medium grained E. J. Baltrusaites, C. K. Clark, G. V. Cohee, R. P. Grant sandstone with well rounded and angular quartz grains. W. A. Kelly, K. K. Landes, G. D. Lindberg and R. B. Thin beds of argillaceous dolomite may occur locally in Newcombe of the Michigan Geological Society * the sandstone. It is about 100 feet thick in the Southern Peninsula of Michigan but is absent in Northern Indiana. The Franconia sandstone is a fine to medium grained Cambrian glauconitic and dolomitic sandstone. It is from 10 to 20 Cambrian rocks in the Southern Peninsula of Michigan feet thick where present in the Southern Peninsula. consist of sandstone, dolomite, and some shale. These * See last page rocks, Lake Superior sandstone, which are of Upper Cambrian age overlie pre-Cambrian rocks and are The Trempealeau is predominantly a buff to light brown divided into the Jacobsville sandstone overlain by the dolomite with a minor amount of sandy, glauconitic Munising. The Munising sandstone at the north is dolomite and dolomitic shale in the basal part. Zones of divided southward into the following formations in sandy dolomite are in the Trempealeau in addition to the ascending order: Mount Simon, Eau Claire, Dresbach basal part. A small amount of chert may be found in and Franconia sandstones overlain by the Trampealeau various places in the formation. -
Chapter 2 Paleozoic Stratigraphy of the Grand Canyon
CHAPTER 2 PALEOZOIC STRATIGRAPHY OF THE GRAND CANYON PAIGE KERCHER INTRODUCTION The Paleozoic Era of the Phanerozoic Eon is defined as the time between 542 and 251 million years before the present (ICS 2010). The Paleozoic Era began with the evolution of most major animal phyla present today, sparked by the novel adaptation of skeletal hard parts. Organisms continued to diversify throughout the Paleozoic into increasingly adaptive and complex life forms, including the first vertebrates, terrestrial plants and animals, forests and seed plants, reptiles, and flying insects. Vast coal swamps covered much of mid- to low-latitude continental environments in the late Paleozoic as the supercontinent Pangaea began to amalgamate. The hardiest taxa survived the multiple global glaciations and mass extinctions that have come to define major time boundaries of this era. Paleozoic North America existed primarily at mid to low latitudes and experienced multiple major orogenies and continental collisions. For much of the Paleozoic, North America’s southwestern margin ran through Nevada and Arizona – California did not yet exist (Appendix B). The flat-lying Paleozoic rocks of the Grand Canyon, though incomplete, form a record of a continental margin repeatedly inundated and vacated by shallow seas (Appendix A). IMPORTANT STRATIGRAPHIC PRINCIPLES AND CONCEPTS • Principle of Original Horizontality – In most cases, depositional processes produce flat-lying sedimentary layers. Notable exceptions include blanketing ash sheets, and cross-stratification developed on sloped surfaces. • Principle of Superposition – In an undisturbed sequence, older strata lie below younger strata; a package of sedimentary layers youngs upward. • Principle of Lateral Continuity – A layer of sediment extends laterally in all directions until it naturally pinches out or abuts the walls of its confining basin. -
Tetrapod Biostratigraphy and Biochronology of the Triassic–Jurassic Transition on the Southern Colorado Plateau, USA
Palaeogeography, Palaeoclimatology, Palaeoecology 244 (2007) 242–256 www.elsevier.com/locate/palaeo Tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau, USA Spencer G. Lucas a,⁎, Lawrence H. Tanner b a New Mexico Museum of Natural History, 1801 Mountain Rd. N.W., Albuquerque, NM 87104-1375, USA b Department of Biology, Le Moyne College, 1419 Salt Springs Road, Syracuse, NY 13214, USA Received 15 March 2006; accepted 20 June 2006 Abstract Nonmarine fluvial, eolian and lacustrine strata of the Chinle and Glen Canyon groups on the southern Colorado Plateau preserve tetrapod body fossils and footprints that are one of the world's most extensive tetrapod fossil records across the Triassic– Jurassic boundary. We organize these tetrapod fossils into five, time-successive biostratigraphic assemblages (in ascending order, Owl Rock, Rock Point, Dinosaur Canyon, Whitmore Point and Kayenta) that we assign to the (ascending order) Revueltian, Apachean, Wassonian and Dawan land-vertebrate faunachrons (LVF). In doing so, we redefine the Wassonian and the Dawan LVFs. The Apachean–Wassonian boundary approximates the Triassic–Jurassic boundary. This tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau confirms that crurotarsan extinction closely corresponds to the end of the Triassic, and that a dramatic increase in dinosaur diversity, abundance and body size preceded the end of the Triassic. © 2006 Elsevier B.V. All rights reserved. Keywords: Triassic–Jurassic boundary; Colorado Plateau; Chinle Group; Glen Canyon Group; Tetrapod 1. Introduction 190 Ma. On the southern Colorado Plateau, the Triassic– Jurassic transition was a time of significant changes in the The Four Corners (common boundary of Utah, composition of the terrestrial vertebrate (tetrapod) fauna. -
The Geologic Time Scale Is the Eon
Exploring Geologic Time Poster Illustrated Teacher's Guide #35-1145 Paper #35-1146 Laminated Background Geologic Time Scale Basics The history of the Earth covers a vast expanse of time, so scientists divide it into smaller sections that are associ- ated with particular events that have occurred in the past.The approximate time range of each time span is shown on the poster.The largest time span of the geologic time scale is the eon. It is an indefinitely long period of time that contains at least two eras. Geologic time is divided into two eons.The more ancient eon is called the Precambrian, and the more recent is the Phanerozoic. Each eon is subdivided into smaller spans called eras.The Precambrian eon is divided from most ancient into the Hadean era, Archean era, and Proterozoic era. See Figure 1. Precambrian Eon Proterozoic Era 2500 - 550 million years ago Archaean Era 3800 - 2500 million years ago Hadean Era 4600 - 3800 million years ago Figure 1. Eras of the Precambrian Eon Single-celled and simple multicelled organisms first developed during the Precambrian eon. There are many fos- sils from this time because the sea-dwelling creatures were trapped in sediments and preserved. The Phanerozoic eon is subdivided into three eras – the Paleozoic era, Mesozoic era, and Cenozoic era. An era is often divided into several smaller time spans called periods. For example, the Paleozoic era is divided into the Cambrian, Ordovician, Silurian, Devonian, Carboniferous,and Permian periods. Paleozoic Era Permian Period 300 - 250 million years ago Carboniferous Period 350 - 300 million years ago Devonian Period 400 - 350 million years ago Silurian Period 450 - 400 million years ago Ordovician Period 500 - 450 million years ago Cambrian Period 550 - 500 million years ago Figure 2. -
Chordates (Phylum Chordata)
A short story Leathem Mehaffey, III, Fall 201993 The First Chordates (Phylum Chordata) • Chordates (our phylum) first appeared in the Cambrian, 525MYA. 94 Invertebrates, Chordates and Vertebrates • Invertebrates are all animals not chordates • Generally invertebrates, if they have hearts, have dorsal hearts; if they have a nervous system it is usually ventral. • All vertebrates are chordates, but not all chordates are vertebrates. • Chordates: • Dorsal notochord • Dorsal nerve chord • Ventral heart • Post-anal tail • Vertebrates: Amphioxus: archetypal chordate • Dorsal spinal column (articulated) and skeleton 95 Origin of the Chordates 96 Haikouichthys Myllokunmingia Note the rounded extension to Possibly the oldest the head bearing sensory vertebrate: showed gill organs bars and primitive vertebral elements Early and primitive agnathan vertebrates of the Early Cambrian (530MYA) Pikaia Note: these organisms were less Primitive chordate, than an inch long. similar to Amphioxus 97 The Cambrian/Ordovician Extinction • Somewhere around 488 million years ago something happened to cause a change in the fauna of the earth, heralding the beginning of the Ordovician Period. • Rather than one catastrophe, the late-Cambrian extinction seems to be a series of smaller extinction events. • Historically the change in fauna (mostly trilobites as the index species) was thought to be due to excessive warmth and low oxygen. • But some current findings point to an oxygen spike due perhaps to continental drift into the tropics, driving rapid speciation and consequent replacement of old with new organisms. 98 Welcome to the Ordovician YOU ARE HERE 99 The Ordovician Sea, 488 million years 100 ago The Ordovician Period lasted almost 45 million years, from 489 to 444 MYA. -
Palynology of the Middle Ordovician Hawaz Formation in the Murzuq Basin, South-West Libya
This is a repository copy of Palynology of the Middle Ordovician Hawaz Formation in the Murzuq Basin, south-west Libya. White Rose Research Online URL for this paper: http://eprints.whiterose.ac.uk/125997/ Version: Accepted Version Article: Abuhmida, F.H. and Wellman, C.H. (2017) Palynology of the Middle Ordovician Hawaz Formation in the Murzuq Basin, south-west Libya. Palynology, 41. pp. 31-56. ISSN 0191-6122 https://doi.org/10.1080/01916122.2017.1356393 Reuse Items deposited in White Rose Research Online are protected by copyright, with all rights reserved unless indicated otherwise. They may be downloaded and/or printed for private study, or other acts as permitted by national copyright laws. The publisher or other rights holders may allow further reproduction and re-use of the full text version. This is indicated by the licence information on the White Rose Research Online record for the item. Takedown If you consider content in White Rose Research Online to be in breach of UK law, please notify us by emailing [email protected] including the URL of the record and the reason for the withdrawal request. [email protected] https://eprints.whiterose.ac.uk/ Palynology of the Middle Ordovician Hawaz Formation in the Murzuq Basin, southwest Libya Faisal H. Abuhmidaa*, Charles H. Wellmanb aLibyan Petroleum Institute, Tripoli, Libya P.O. Box 6431, bUniversity of Sheffield, Department of Animal and Plant Sciences, Alfred Denny Building, Western Bank, Sheffield, S10 2TN, UK Twenty nine core and seven cuttings samples were collected from two boreholes penetrating the Middle Ordovician Hawaz Formation in the Murzuq Basin, southwest Libya. -
Reinterpretation of the Enigmatic Ordovician Genus Bolboporites (Echinodermata)
Reinterpretation of the enigmatic Ordovician genus Bolboporites (Echinodermata). Emeric Gillet, Bertrand Lefebvre, Véronique Gardien, Emilie Steimetz, Christophe Durlet, Frédéric Marin To cite this version: Emeric Gillet, Bertrand Lefebvre, Véronique Gardien, Emilie Steimetz, Christophe Durlet, et al.. Reinterpretation of the enigmatic Ordovician genus Bolboporites (Echinodermata).. Zoosymposia, Magnolia Press, 2019, 15 (1), pp.44-70. 10.11646/zoosymposia.15.1.7. hal-02333918 HAL Id: hal-02333918 https://hal.archives-ouvertes.fr/hal-02333918 Submitted on 13 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. 1 Reinterpretation of the Enigmatic Ordovician Genus Bolboporites 2 (Echinodermata) 3 4 EMERIC GILLET1, BERTRAND LEFEBVRE1,3, VERONIQUE GARDIEN1, EMILIE 5 STEIMETZ2, CHRISTOPHE DURLET2 & FREDERIC MARIN2 6 7 1 Université de Lyon, UCBL, ENSL, CNRS, UMR 5276 LGL-TPE, 2 rue Raphaël Dubois, F- 8 69622 Villeurbanne, France 9 2 Université de Bourgogne - Franche Comté, CNRS, UMR 6282 Biogéosciences, 6 boulevard 10 Gabriel, F-2100 Dijon, France 11 3 Corresponding author, E-mail: [email protected] 12 13 Abstract 14 Bolboporites is an enigmatic Ordovician cone-shaped fossil, the precise nature and systematic affinities of 15 which have been controversial over almost two centuries. -
New Radiometric 40Ar–39Ar Dates and Faunistic Analyses Refine
www.nature.com/scientificreports OPEN New radiometric 40Ar–39Ar dates and faunistic analyses refne evolutionary dynamics of Neogene vertebrate assemblages in southern South America Francisco J. Prevosti1,2*, Cristo O. Romano2,3, Analía M. Forasiepi2,3, Sidney Hemming4, Ricardo Bonini2,5, Adriana M. Candela2,6, Esperanza Cerdeño2,3, M. Carolina Madozzo Jaén2,7,8, Pablo E. Ortiz2,7, François Pujos2,3, Luciano Rasia2,6, Gabriela I. Schmidt2,9, Matias Taglioretti10,11,12, Ross D. E. MacPhee13 & Ulyses F. J. Pardiñas2,14,15 The vertebrate fossil record of the Pampean Region of Argentina occupies an important place in South American vertebrate paleontology. An abundance of localities has long been the main basis for constructing the chronostratigraphical/geochronological scale for the late Neogene–Quaternary of South America, as well as for understanding major patterns of vertebrate evolution, including the Great American Biotic Interchange. However, few independently-derived dates are available for constraining this record. In this contribution, we present new 40Ar/39Ar dates on escorias (likely the product of meteoric impacts) from the Argentinean Atlantic coast and statistically-based biochronological analyses that help to calibrate Late Miocene–Pliocene Pampean faunal successions. For the type areas of the Montehermosan and Chapadmalalan Ages/Stages, our results delimit their age ranges to 4.7–3.7 Ma and ca. 3.74–3.04 Ma, respectively. Additionally, from Buenos Aires Province, dates of 5.17 Ma and 4.33 Ma were recovered for “Huayquerian” and Montehermosan faunas. This information helps to better calibrate important frst appearances of allochthonous taxa in South America, including one of the oldest records for procyonids (7.24–5.95 Ma), cricetids (6.95– 5.46 Ma), and tayassuids (> 3.74 Ma, oldest high-confdence record). -
RESEARCH New Biostratigraphic Evidence of Late Permian to Late
RESEARCH New biostratigraphic evidence of Late Permian to Late Triassic deposits from Central Tibet and their paleogeographic implications Gui-chun Wu1,*, Zhan-sheng Ji2, Wei-hua Liao3, and Jian-xin Yao1 1KEY LABORATORY OF STRATIGRAPHY AND PALAEONTOLOGY, MINISTRY OF LAND AND RESOURCES, INSTITUTE OF GEOLOGY, CHINESE ACADEMY OF GEOLOGICAL SCIENCES, BEIJING 100037, CHINA 2CHINESE ACADEMY OF GEOLOGICAL SCIENCES, BEIJING 100037, CHINA 3NANJING INSTITUTE OF GEOLOGY AND PALAEONTOLOGY, CHINESE ACADEMY OF SCIENCES, NANJING, 210008, CHINA ABSTRACT Triassic deposits in the Bangong-Nujiang Suture Zone are important for understanding its tectonic nature and evolutionary history, but have not been systematically studied due to a lack of biostratigraphic data. For a long time, the Upper Triassic Quehala Group featuring clasolite has been regarded as the only rocky unit. In recent years, the silicite-dominated Gajia Formation that bears radiolarian fossils was suggested to represent Ladinian to Carnian deposits. The Upper Permian and Lower Triassic rocks have never been excavated and thus are considered to be absent. This research, however, reveals that fossils aged from the Late Permian to Anisian of the Middle Trias- sic and Norian of the Late Triassic have been preserved in the central Bangong-Nujiang Suture Zone, which provides evidence of Upper Permian to early Middle Triassic deposits and provides new insights on the Upper Triassic strata as well. A new Triassic strata succes- sion is thus proposed for the Bangong-Nujiang Suture Zone, and it demonstrates great similarities with those from Lhasa to the south and Qiangtang to the north. Therefore, we deduce that the Bangong-Nujiang Suture Zone was under a similar depositional setting as its two adjacent terranes, and it was likely a carbonate platform background because limestones were predominant across the Triassic. -
Late Silurian Trilobite Palaeobiology And
LATE SILURIAN TRILOBITE PALAEOBIOLOGY AND BIODIVERSITY by ANDREW JAMES STOREY A thesis submitted to the University of Birmingham for the degree of DOCTOR OF PHILOSOPHY School of Geography, Earth and Environmental Sciences University of Birmingham February 2012 University of Birmingham Research Archive e-theses repository This unpublished thesis/dissertation is copyright of the author and/or third parties. The intellectual property rights of the author or third parties in respect of this work are as defined by The Copyright Designs and Patents Act 1988 or as modified by any successor legislation. Any use made of information contained in this thesis/dissertation must be in accordance with that legislation and must be properly acknowledged. Further distribution or reproduction in any format is prohibited without the permission of the copyright holder. ABSTRACT Trilobites from the Ludlow and Přídolí of England and Wales are described. A total of 15 families; 36 genera and 53 species are documented herein, including a new genus and seventeen new species; fourteen of which remain under open nomenclature. Most of the trilobites in the British late Silurian are restricted to the shelf, and predominantly occur in the Elton, Bringewood, Leintwardine, and Whitcliffe groups of Wales and the Welsh Borderland. The Elton to Whitcliffe groups represent a shallowing upwards sequence overall; each is characterised by a distinct lithofacies and fauna. The trilobites and brachiopods of the Coldwell Formation of the Lake District Basin are documented, and are comparable with faunas in the Swedish Colonus Shale and the Mottled Mudstones of North Wales. Ludlow trilobite associations, containing commonly co-occurring trilobite taxa, are defined for each palaeoenvironment. -
Lower Paleozoic
Paleozoic Time Scale and Sea-Level History Sponsored, in part, by: Time ScaLe R Creator Updated by James G. Ogg (Purdue University) and Gabi Ogg to: GEOLOGIC TIME SCALE 2004 Cen Mesozoic (Gradstein, F.M., Ogg, J.G., Smith, A.G., et al.2004) Paleozoic ICS Precambrian and The Concise Geologic Time Scale (Ogg, J.G., Ogg, G. & Gradstein, F.M., 2008) Paleozoic Sequences -- Haq and Schutter (Science, 2008) Geo- magnetic Standard Chronostratigraphy Mega- Sequences Coastal Onlap and Mean Sea- Paleozoic Long- and Silurian-Ordovician Ordovician Trilobite Zones and Polarity Sequences (nomenclature by Level (intermediate term) Short-Term Sea Level Sea Level Sealevel Events major Cambrian events Conodont Zonation Graptolite Zonation Age Period Epoch Stage Primary of Sloss J. Ogg) 0 100 200m 0 100 200m 0 100 200m (Baltoscandia) South China 416.1 Monograptus bouceki - Oulodus elegans detortus transgrediens - perneri Silur-8 Monograptus branikensis - Pri-1 417.5 lochkovensis 418.02 Ozarkodina remscheidensis Pridoli Interval Zone Monograptus parultimus - ultimus 418.7 Lud-3 419.0 Ozarkodina crispa Monograptus formosus 419.75 Silur-7 Ozarkodina snajdri Interval Zone Neocucullograptus kozlowskii - 420 Ludfordian Ludlow-Pridoli Polonograptus podoliensis Lud-2 420.5 420.7 Polygnathoides siluricus mixed-polarity interval 421.0 Saetograptus leintwardinensis 421.3 Lud-1 Ancoradella ploeckensis Ludlow Lobograptus scanicus Gorstian 422.0 Silur-6 [ not zoned ] 422.9 Kockelella stauros Neodiversograptus nilssoni Gor-1 423.0 Colonograptus ludensis Colonograptus praedeubeli