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A Simple Generative Model of Trilobite Segmentation and Growth
paleo.peercommunityin.orG Open Access A SIMPLE GENERATIVE MODEL OF TRILOBITE Peer-ReVIEWED SEGMENTATION AND GROWTH RESEARCH ARTICLE Melanie J Hopkins1 1 Division OF Paleontology (INVERTEBRates), American Museum OF NaturAL History – NeW York, USA Cite as: Hopkins MJ (2020). A SIMPLE GENERATIVE MODEL OF TRILOBITE SEGMENTATION THIS ARTICLE HAS BEEN PEER-REVIEWED BY PCI PALEO AND GROwth. PaleorXiv VERSION 3, Read THE Editor’S RECOMMENDATION AND ReferEES’ REPORTS AT DOI:10.24072/pci.paleo.100004 peer-rEVIEWED BY PCI Paleo. DOI: 10.31233/osf.io/zt642 Published: 27 January 2020 ABSTRACT Recommender: GenerATIVE GROWTH MODELS HAVE BEEN THE BASIS FOR NUMEROUS STUDIES OF MORPHOLOGICAL DIVERSITY AND Evolution. Christian Klug Most WORK HAS FOCUSED ON MODELING ACCRETIONARY GROWTH systems, WITH MUCH LESS ATTENTION TO DISCRETE GROWTH systems. GenerATIVE GROWTH MODELS FOR MOLTING ORganisms, SUCH AS ARTHRopods, HAVE REMAINED PARTICULARLY ReVIEwers: Kenneth De Baets AND Lukáš Laibl elusive. HoWEver, OUR UNDERSTANDING OF post-embryonic GROWTH IN TRILOBITE SPECIES IS SUFFICIENTLY MATURE THAT IT IS NOW POSSIBLE TO MODEL GROWTH IN A WAY THAT INCORPORATES THE ADDITION OF NEW PARTS AS WELL AS DIFFERENTIAL Correspondence: GROWTH RATES FOR EXISTING PARTS ACROSS THE TRILOBITE BODY plan. Furthermore, BODY SIZE DATA FOR A LARGE SAMPLE OF [email protected] SPECIMENS OF THE TRILOBITE SPECIES AulacopleurA KONINCKII (Barrande, 1846) MAKE IT POSSIBLE TO GENERATE ROBUST ESTIMATES FOR MODEL PARameters. Although THE GENERATIVE MODEL DESCRIBED HERE WAS BASED ON A RELATIVELY SIMPLE SEGMENTATION schedule, A DIVERSE ARRAY OF OBSERVED BODY SIZES AND RELATIVE PROPORTIONS OF BODY REGIONS CAN BE Open Data ATTAINED BY ALTERING ONLY A FEW PARAMETERS AT A time. -
Nautiloid Shell Morphology
MEMOIR 13 Nautiloid Shell Morphology By ROUSSEAU H. FLOWER STATEBUREAUOFMINESANDMINERALRESOURCES NEWMEXICOINSTITUTEOFMININGANDTECHNOLOGY CAMPUSSTATION SOCORRO, NEWMEXICO MEMOIR 13 Nautiloid Shell Morphology By ROUSSEAU H. FLOIVER 1964 STATEBUREAUOFMINESANDMINERALRESOURCES NEWMEXICOINSTITUTEOFMININGANDTECHNOLOGY CAMPUSSTATION SOCORRO, NEWMEXICO NEW MEXICO INSTITUTE OF MINING & TECHNOLOGY E. J. Workman, President STATE BUREAU OF MINES AND MINERAL RESOURCES Alvin J. Thompson, Director THE REGENTS MEMBERS EXOFFICIO THEHONORABLEJACKM.CAMPBELL ................................ Governor of New Mexico LEONARDDELAY() ................................................... Superintendent of Public Instruction APPOINTEDMEMBERS WILLIAM G. ABBOTT ................................ ................................ ............................... Hobbs EUGENE L. COULSON, M.D ................................................................. Socorro THOMASM.CRAMER ................................ ................................ ................... Carlsbad EVA M. LARRAZOLO (Mrs. Paul F.) ................................................. Albuquerque RICHARDM.ZIMMERLY ................................ ................................ ....... Socorro Published February 1 o, 1964 For Sale by the New Mexico Bureau of Mines & Mineral Resources Campus Station, Socorro, N. Mex.—Price $2.50 Contents Page ABSTRACT ....................................................................................................................................................... 1 INTRODUCTION -
Part I. Revision of Buttsoceras. Part II. Notes on the Michelinoceratida
MEMOIR 10 PART I Revision of Buttsoceras PART II Notes on the Michelinoceratida By ROUSSEAU H. FLOWER 1 9 6 2 STATE BUREAU OF MINES AND MINERAL RESOURCES NEW MEXICO INSTITUTE OF MINING AND TECHNOLOGY CAMPUS STATION SOCORRO, NEW MEXICO NEW MEXICO INSTITUTE OF MINING & TECHNOLOGY E. J. Workman, President STATE BUREAU OF MINES AND MINERAL RESOURCES Alvin J. Thompson, Director THE REGENTS MEMBERS Ex OFFICIO The Honorable Edwin L. Mechem ........................................ Governor of New Mexico Tom Wiley .......................................................... Superintendent of Public Instruction APPOINTED MEMBERS William G. Abbott ............................................................................................... Hobbs Holm 0. Bursum, Jr. ......................................................................................... Socorro Thomas M. Cramer ......................................................................................... Carlsbad Frank C. DiLuzio ...................................................................................... Albuquerque Eva M. Larrazolo (Mrs. Paul F.) ............................................................... Albuquerque Published October I2, 1962 For Sale by the New Mexico Bureau of Mines & Mineral Resources Campus Station, Socorro, N. Mex.—Price $2.00 Contents PART I REVISION OF BUTTSOCERAS Page ABSTRACT ....................................................................................................................... INTRODUCTION ............................................................................................................................. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant. -
Contributions in BIOLOGY and GEOLOGY
MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - --- -
Current Views on Chelicerate Phylogeny —A Tribute to Peter Weygoldt
Current views on chelicerate phylogeny —A tribute to Peter Weygoldt The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Giribet, Gonzalo. 2018. “Current Views on Chelicerate phylogeny— A Tribute to Peter Weygoldt.” Zoologischer Anzeiger 273 (March): 7– 13. doi:10.1016/j.jcz.2018.01.004. Published Version doi:10.1016/j.jcz.2018.01.004 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:37308630 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Open Access Policy Articles, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#OAP 1 Current views on chelicerate phylogeny—a tribute to Peter Weygoldt 2 3 Gonzalo Giribet 4 5 Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard 6 University, 26 Oxford Street, CamBridge, MA 02138, USA 7 8 Keywords: Arachnida, Chelicerata, Arthropoda, evolution, systematics, phylogeny 9 10 11 ABSTRACT 12 13 Peter Weygoldt pioneered studies of arachnid phylogeny by providing the first synapomorphy 14 scheme to underpin inter-ordinal relationships. Since this seminal worK, arachnid relationships 15 have been evaluated using morphological characters of extant and fossil taxa as well as multiple 16 generations of molecular sequence data. While nearly all datasets agree on the monophyly of 17 Tetrapulmonata, and modern analyses of molecules and novel morphological and genomic data 18 support Arachnopulmonata (a sister group relationship of Scorpiones to Tetrapulmonata), the 19 relationships of the apulmonate arachnid orders remain largely unresolved. -
The Evolution of Trilobite Body Patterning
ANRV309-EA35-14 ARI 20 March 2007 15:54 The Evolution of Trilobite Body Patterning Nigel C. Hughes Department of Earth Sciences, University of California, Riverside, California 92521; email: [email protected] Annu. Rev. Earth Planet. Sci. 2007. 35:401–34 Key Words First published online as a Review in Advance on Trilobita, trilobitomorph, segmentation, Cambrian, Ordovician, January 29, 2007 diversification, body plan The Annual Review of Earth and Planetary Sciences is online at earth.annualreviews.org Abstract This article’s doi: The good fossil record of trilobite exoskeletal anatomy and on- 10.1146/annurev.earth.35.031306.140258 togeny, coupled with information on their nonbiomineralized tis- Copyright c 2007 by Annual Reviews. sues, permits analysis of how the trilobite body was organized and All rights reserved developed, and the various evolutionary modifications of such pat- 0084-6597/07/0530-0401$20.00 terning within the group. In several respects trilobite development and form appears comparable with that which may have charac- terized the ancestor of most or all euarthropods, giving studies of trilobite body organization special relevance in the light of recent advances in the understanding of arthropod evolution and devel- opment. The Cambrian diversification of trilobites displayed mod- Annu. Rev. Earth Planet. Sci. 2007.35:401-434. Downloaded from arjournals.annualreviews.org ifications in the patterning of the trunk region comparable with by UNIVERSITY OF CALIFORNIA - RIVERSIDE LIBRARY on 05/02/07. For personal use only. those seen among the closest relatives of Trilobita. In contrast, the Ordovician diversification of trilobites, although contributing greatly to the overall diversity within the clade, did so within a nar- rower range of trunk conditions. -
1540 Evans.Vp
An early Silurian (Aeronian) cephalopod fauna from Kopet-Dagh, north-eastern Iran: including the earliest records of non-orthocerid cephalopods from the Silurian of Northern Gondwana DAVID H. EVANS, MANSOUREH GHOBADI POUR, LEONID E. POPOV & HADI JAHANGIR The cephalopod fauna from the Aeronian Qarebil Limestone of north-eastern Iran comprises the first comprehensive re- cord of early Silurian cephalopods from peri-Gondwana. Although consisting of relatively few taxa, the assemblage in- cludes members of the orders Oncocerida, Discosorida, Barrandeocerida and Orthocerida. Coeval records of several of these taxa are from low palaeolatitude locations that include Laurentia, Siberia, Baltica and Avalonia, and are otherwise unknown from peri-Gondwana until later in the Silurian. The cephalopod assemblage occurs in cephalopod limestones that currently represent the oldest record of such limestones in the Silurian of the peri-Gondwana margin. The appear- ance of these cephalopods, together with the development of cephalopod limestones may be attributed to the relatively low latitudinal position of central Iran and Kopet-Dagh compared with that of the west Mediterranean sector during the Aeronian. This, combined with the continued post-glacial warming after the Hirnantian glaciation, facilitated the initia- tion of carbonate deposition and conditions suitable for the development of the cephalopod limestones whilst permitting the migration of cephalopod taxa, many of which were previously restricted to lower latitudes. • Key words: Llandovery, Aeronian, Cephalopoda, palaebiogeography, peri-Gondwana. EVANS, D.H., GHOBADI POUR, M., POPOV,L.E.&JAHANGIR, H. 2015. An Early Silurian (Aeronian) cephalopod fauna from Kopet-Dagh, north-eastern Iran: including the earliest records of non-orthocerid cephalopods from the Silurian of Northern Gondwana. -
The PALEONTOLOGY of KENTUCKY
The PALEONTOLOGY of KENTUCKY I The GEOLOGICAL SUCCESSION of LIFE In KENTUCKY By ROY LEE MOODIE THE GEOLOGICAL SUCCESSION OF LIFE IN KENTUCKY By ROY L. MOODIE INTRODUCTION ORIGIN OF THE EARTH The origin of the earth, from the viewpoint of geology and astronomy, is best accounted for by the Planetesimal Hypothesis, which is to the effect that, the earth, instead of originally being a molten mass, grew to its present form by the accretion of cold bodies from outer space. When the earth had attained about two- thirds its present size it is held that there was enough attractive force to retain a warm blanket of atmosphere. The presence of living things then became possible; simple plants and animals may then have begun to live. AGE OF THE EARTH The age of the earth is usually expressed in terms of millions of years, but such statements are largely meaningless since none of us can comprehend the vast stretches of time indicated by even a million years. It will help somewhat to understand the meanings of geological time if we say that the Mammoth Cave, in Edmonson County, had been in existence for many thousands of years when the most ancient Egyptians were first entering the Nile Valley, long before any of the pyramids had been thought of, thousands of years before the city of Babylon was built. Huge mountain chains had arisen and were worn down long before the waters of the earth 4 THE PALEONTOLOGY OF KENTUCKY had dissolved the first tiny crevice of the colossal cavern. When we realize that solid rock blocks creep down a slope at the rate of about one inch a century, or that a thousand years may bring about almost imperceptible changes in earth's feature, then we gain some concept of the meaning of geological time. -
Introduction to the Trilobites: Morphology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Macroevolution and More By Michelle M. Casey1 and Bruce S. Lieberman1,2, 1Biodiversity Institute and 2Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Undergraduate laboratory exercise for sophomore/junior level paleontology course Learning Goals and Pedagogy This lab is intended for an upper level paleontology course containing sophomores and juniors who have already taken historical geology or its equivalent; however it may be suitable for introductory biology or geology students familiar with geological time, phylogenies, and trace fossils. This lab will be particularly helpful to those institutions that lack a large teaching collection by providing color photographs of museum specimens. Students may find previous exposure to phylogenetic methods and terminology helpful in completing this laboratory exercise. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens, not just diagrammatic representations; 3) to highlight major events or trends in the evolutionary history and ecology of Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other solely extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They were exclusively marine but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter. -
(Late Ordovician) of Porkuni, Estonia
Concentrations of juvenile and small adult cephalopods in the Hirnantian cherts (Late Ordovician) of Porkuni, Estonia BJÖRN KRÖGER Kröger, B. 2007. Concentrations of juvenile and small adult cephalopods in the Hirnantian cherts (Late Ordovician) of Porkuni, Estonia. Acta Palaeontologica Polonica 52 (3): 591–608. The quarry in the north Estonian village of Porkuni provides a succession of shallow−water limestones and cherts span− ning the Ashgillian Normalograptus? extraordinarius graptolite Biozone. This interval comprises the initial pulse of the end−Ordovician extinction. The succession of Porkuni contains abundant and extraordinarily well−preserved fossils. 71 cephalopod specimens were extracted from these strata at Porkuni. Many of these specimens are fragments of juvenile shells or small adults. The embryonic shells of the cephalopods are usually preserved and provide insight into their early ontogeny. The faunal composition is considered as autochthonous and reflects a “palaeo−nursery” in a Hirnantian reef en− vironment. The collected specimens represent twelve genera and four orders. Small oncoceridans and orthoceridans dom− inate the association. The rate of endemism is very high, since only two genera found in Porkuni, are known from outside Baltoscandia. The new genera Parvihebetoceras, Pomerantsoceras, Porkunioceras, and the new species Parvihebeto− ceras wahli, Pomerantsoceras tibia, Porkunioceras tuba, and Strandoceras orvikui are erected. Key words: Cephalopoda, Nautiloidea, mode of life, end−Ordovician extinction, Ashgillian. Björn Kröger [[email protected]], Institut für Paläontologie, Museum für Naturkunde, D−10115 Berlin, Invaliden− strasse 43, Germany. Introduction lar pattern for bryozoans at the Ordovician–Silurian boundary interval. The Porkuni quarry, which yields this fauna is there− The name Porkuni was given to the youngest regional stage of fore a unique archive of a potential survival and recovery the Ordovician of Baltoscandia.