Author(s) Year Title Publication URL Ahrens, D. 2021 Two new species of the Neoserica (sensu stricto) group from China Journal of Natural History 54(45-46):2927- 2936

Ahrens, D. 2021 New species of the genus Gynaecoserica Brenske, 1896 from Indochina Zootaxa 5005(2):218-226

Ahrens, D. & Pham, P.V. 2021 Additions to the Neoserica calva group from continental South East Asia Zootaxa 5032(3):357-378

Ahrens, D., Liu, W.G., Fabrizi, S. & Bai, M. 2021 Taxonomic review on the Trioserica Moser, 1922 species of China Zootaxa 4999(4):343-355

Allsopp, P.G. 2021 The Australian endemic genera Mesystoechus Waterhouse, 1878, Zootaxa 4965(2):363-374 Amblochilus Blanchard, 1851, and Bilobatus Machatschke, 1970 revisited

Allsopp, P.G. 2021 After the blaze: bushfire recovery survey finds a ninth species of Watkinsia Australian Entomologist 48(3):187-194 Brittion, 1995

Andújar, C. & Grebennikov, V.Y. 2021 Endogean of Madagascar: deep soil sampling and illustrated Zootaxa 4963(3):317-334 overview

Anichtchenko, A.V., Minkina, L., Vasiljeva, A. 2021 A review of the genus Rhyparus in the Philippines, with descriptions of two Acta entomologica Musei nationalis Pragae https://www.aemnp.eu/data/article- & Medina, M.N.D. new species from Mindanao 61(1):99-111 1879/1914-61_1_99.pdf

Aquino dos Santos, M.F., Mattos, I., 2021 A new passalid fossil from the Santana Formation (Crato member, Lower Cretaceous Research 118(104664):1-15 Mermudes, J.R.M., Scheffler, S.M. & Reyes- Cretaceous), Araripe Basin, NE Brazil: Paleoecological and Castillo, P. paleobiogeographic implications

Aragón-Sánchez, M., Aragón-García, A., 2021 Actualización de las especies del género Macrodactylus Dejean para el Acta Zoológica Mexicana (nueva serie) https://azm.ojs.inecol.mx/index.php/azm/articl Arce-Pérez, R., Pérez-Torres, B.C., Cuate- estado de Puebla, México 37:1-10 e/view/2403/2845 Mozo, V.A. & López-Olguín, J.F.

Arce-Pérez, R. & Reynoso-Velasco, D. 2021 Two new species of Macrodactylus from Mexico and Guatemala Zootaxa 4903(4):493-500

Avtzis, D.N., Markoudi, V., Mizerakis, V., 2021 The Aegean Archipelago as cradle: divergence of the glaphyrid genus Systematics and Biodiversity 19(4):346-358 Devalez, J., Nakas, G., Poulakakis, N. & Pygopleurus and phylogeography of P. Foina Petanidou, T.

Ballerio, A. 2021 First contribution to the knowledge of the Ceratocanthinae from the Solomon Telnov D., Barclay M.V.L. & Pauwels O.S.G. Islands, with descriptions of five new species (eds.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea 4:61-73 Barria, M.D., Quirós, D.I. & Emmen, D. 2021 Bioacoustic analysis of a compound sound with stridulation and forced air Revista Chilena de Entomologia 47(2):187- produced by the larva of Phileurus valgus (Olivier, 1789) 194

Ben Aba, W. & Bouragaoui, Z. 2021 Confirmation of the presence of the vulnerable species Calicnemis obesa Munis Entomology & Zoology 16(1):381-384 obesa Erichson, 1841 in Tunisia Bento, M. & Grossi, P.C. 2021 Two New Species of Macraspis MacLeay, 1819 from Brazil, with Neotropical Entomology 50(2):247-257 Supplementary Description on the Chromatic Variation and Endophallus of Macraspis laevicollis (Waterhouse, 1881)

Beza-Beza, C.F., Jiménez-Ferbans, L. & 2021 Historical biogeography of New World passalid beetles reveals Journal of Biogeography 48(8):2037-2052 McKenna, D.D. Mesoamerican tropical forests as a centre of origin and taxonomic diversification

Bhattacharyya, B., Bhagawati, S., 2021 Species diversity and relative abundance of scarab beetles in Jorhat district Environment 24(1):46-49 Manpoong, N.S., Das, P.P.G. & Devi, E.B. of Assam, India

Bihan, C.Z. & Zhan, Z. 2021 A new species of Lucanus Scopoli, 1763 from Tibet, China Faunitaxys 9(27):1-4 https://www.faunitaxys.fr/app/download/1204 8606997/Faunitaxys_f110.pdf?t=1632404617

Boilly, O. & Vaz-de-Mello, F.Z. 2021 Descriptions de deux nouvelles espèces de Neoathyreus et clé des groupes Bulletin de la Société entomologique de d’espèces proposés pour le sous-genre nominatif France 126(1):41-50

Boilly, O. & Vaz-de-Mello, F.Z. 2021 Description d'un nouveau Dichotomius Hope, 1838 du Brésil, du Suriname, Acorep-France: Coléoptères de Guyane du Venezuela et de Guyane 13:22-24

Bordat, P. 2021 Nouvelles synonymies au sein des Aphodiinae Catharsius. La Revue 22:15-23

Bousquet, Y. & Davies, A.E. 2021 On the dating of the publications (1805–1885) of the Société Impériale des Sherbornia 7(2):5-20 Naturalistes de Moscou

Byk, A., Bunalski, M. & Trzak, P. 2021 Nowe stanowiska Melinopterus reyi (Reitter, 1892) w Polsce Wiadomości Entomologiczne 40(2):7-8 http://pte.au.poznan.pl/we/2021/7N-Byk-i- in.pdf

Calcetas, O.A., Adorada, J.L., Adorada, J.R., 2021 New Records of Scarab Insect Pests of Cacao (Theobroma cacao L.) in the Philippine Journal of Science 150(5):1197- Caoili, B.L., Rosales, A.M. & Dimapilis, E.F. Philippines 1206

Carvalho, T.G., Seidel, M. & Grossi, P.C. 2021 Taxonomic revision of the genus Oplognathus MacLeay, 1819 European Journal of Taxonomy 764:62-84 https://europeanjournaloftaxonomy.eu/index. php/ejt/article/view/1471/4885

Chamorro, W., Lopera-Toro, A. & Rossini, M. 2021 A new species and distribution records of Dichotomius Hope, 1838 in Zootaxa 4942(2):193-206 Colombia

Chandra, K., Ahrens, D., Bhunia, D., 2021 New species and records of Sericini from India Zootaxa 4951(3):492-510 Sreedevi, K. & Gupta, D.

Cheng, C.T., Jeng, M.L., Tsau, J.F., Li, C.L. 2021 Two mitochondrial genomes of Taiwanese rhinoceros beetles, Oryctes Mitochondrial DNA Part B:6(8):2260-2262 & Wu, L.W. rhinoceros and Eophileurus chinensis

Cherman, M.A., Basílio, D.S., Mise, K.M., 2021 Liogenys Guérin-Méneville, 1831 from the Chacoan Province and its Zootaxa 4938(1):1-59 Frisch, J. & Almeida, L.M. boundaries: taxonomic overview with four new species

Cherman, M.A., Basílio, D.S., Mise, K.M., 2021 Liogenys Guérin-Méneville, 1831 of northern South America and Central Zootaxa 4990(2):201-226 Frisch, J., Smith, A.B.T. & Almeida, L.M. America: taxonomic overview with four new species Chou, M.H., Tseng, W.Z., Sang Y.D., 2021 Incipient speciation and its impact on taxonomic decision: a case study using Biological Journal of the Linnean Society :1- Morgan, B., De Vivo, M., Kuan, Y.H., Wang, a sky island sister-species pair of stag beetles 15 online first L.J., Chen, W.Y. & Huang, J.P.

Clavijo-Bustos, J., Bacca, T. & Canal, N.A. 2021 Nuevos registros de Aphodiinae en Colombia Revista de la Academia Colombiana de https://raccefyn.co/index.php/raccefyn/article/ Ciencias Exactas, Físicas y Naturales view/1331/3129 45(176):738-746

Correa, C.M.A., Puker, A. & Vaz-de-Mello, 2021 Diversidade de Besouros Scarabaeinae do Ecótono Cerrado-Pantanal Ecótono Cerrado Pantanal: Meio Ambiente F.Z. e História Natural. Capítulo XI:177-189

Correia Costa, F., Cherman, M.A. & 2021 Phylogenetic relationships of Manonychus Moser among the Neotropical Zoologischer Anzeiger 292:1-13 Iannuzzi, L. Melolonthinae

Costa, L.O., Duarte, P.R.M., Iannuzzi, L. & 2021 A hidden Megasoma Kirby species under the enigmatic genus Gibboryctes Zootaxa 4964(2):390-394 Grossi, P.C. Endrödi

Costa-Silva, V., Strümpher, W.P. & Vaz-de- 2021 Review of the Brazilian species of Omorgus Erichson, 1847 Journal of Natural History 54(31-32):1993- Mello, F.Z. 2024

Coutinho, G.V., Gomes, E.S., Avila, C.J., 2021 First record of Plectris aliena Chapin, 1934 as a potential sugarcane pest in Scientia Agricola (2022) 79(2):1-7 https://www.scielo.br/pdf/sa/v79n2/1678- Silva, I.F., Costa, E.N. & Cherman, M.A. Brazil 992X-sa-79-02-e20200128.pdf

Cupello, M., Reiro-Costa, C.S. & Vaz-de- 2021 The evolution of Bolbites onitoides: its phylogenetic significance, Zoological Journal of the Linnean Society Mello, F. geographical polychromatism and the subspecies problem :1-62 online first

Daniel, G.M., Sole, C.L., Scholtz, C.H. & 2021 Historical diversification and biogeography of the endemic southern African Biological Journal of the Linnean Society Davis, A.L.V. dung genus, Epirinus 133(3):751-765

Delatour, T., Dupuis, F. & Sudre, J. 2021 A field survey in the Western Cape Province of South Africa. Part 2: Entomologia Africana 26(1):39-48 Coleoptera, , Dynastinae

Dendi, D., Ajong, S.N., Amori, G. & Luiselli, 2021 Decline of the Commercially Attractive White Morph in Goliath Beetle Diversity 13(8)(388):1-5 https://www.mdpi.com/1424- L. Polymorphic Populations 2818/13/8/388/pdf

Dupuis, F. 2021 Nouvelles espèces et synonymies dans le genre Cyclocephala Dejean, 1821 Coléoptères 27(2):7-18 http://www.coleopteres.fr/Coleopteres27(2).p df

Egorov, L.V., Ruchin, A.B., Semionenkov, 2021 Some data on the Coleoptera Fauna of the Mordovia State Nature Reserve. Proceedings of the Mordovia State Nature O.I., Semishin, G.B. & Esin, M.N. Report 10 Reserve 26:96-128

El-Shafie, H.A.F. 2021 The date palm borers of the genus Oryctes:bionomics, economic impact and Insect Environment 24(1):12-17 possible management measures

Fabbriciani, F. & Patacchiola, D. 2021 Prima segnalazione accertata di Trox (Granulitrox) granulipennis Fairmaire, Giornale Italiano di Entomologia 16(66):127- 1852 per l'Italia 132

Fabrizi, S., Liu, W.G., Bai, M., Yang, X. & 2021 A monograph of the genus Maladera Mulsant & Rey, 1871 of China Zootaxa 4922(1):1-400 Ahrens, D. Ferreira, A., Vaz-de-Mello, F. & Bravo, F. 2021 A new species of Pelidnota MacLeay from Southeast Brazil Papéis Avulsos de Zoologia https://www.revistas.usp.br/paz/article/view/1 61(e20216139):1-9 78518/170187

Fery, H., Dellacasa, G. & Dellacasa, M. 2021 Notes on Aphodius coenobita Gravenhorst, 1807, a forgotten name Zootaxa 4990(3):553-562

Filipović, I., Hereward, J.P., Rašić, G., 2021 The complete mitochondrial genome sequence of Oryctes rhinoceros based PeerJ e10552:1-16 https://peerj.com/articles/10552.pdf Devine, G.J., Furlong, M.J. & Etebari, K. on long-read nanopore sequencing

Frolov, A.V. & Akhmetova, L.A. 2021 A new species of the Aegidium Arrow from Colombia Zootaxa 4927(4):583–586

Frolov, A.V. & Akhmetova, L.A. 2021 Taxonomic revision of the Afrotropical scarab beetle genus Cerhomalus European Journal of Taxonomy 739:36-50 https://europeanjournaloftaxonomy.eu/index. Quedenfeldt, 1884 php/ejt/article/view/1265/3745

Frolov, A.V., Vishnevskaya, M.S. & 2021 Identification of larvae of two Aphodius Helwig, 1798 species using Zootaxa 5047(2):153-164 Akhmetova, L. morphology and DNA barcode

Fuhrmann, J. 2021 Zagurya, a new genus from Brazil with notes on claw movement in Zootaxa 5016(3):395-406 Macrodactylini

Fuhrmann, J. & Smith, A.B.T. 2021 Compsodactylus Fuhrmann, 2012: distributional notes and a new species Zootaxa 4990(2):387-393 https://www.mapress.com/zt/article/view/zoot axa.4990.2.12/44376

Gámez, J., Acconcia, R. & Noriega, J.A. 2021 Nuevos registros de distribución de Digitonthophagus gazella (Fabricius, Entomotropica 36:46-50 1787) en Venezuela

Gao, C.B, Bai, M., Fang, H. & Li, J. 2021 Miridiba hani Gao, a new species from Hainan, China, with a new Zootaxa 4996(2):331-342 combination from Vietnam

Gao, C.B. & Coca-Abia, M.M. 2021 Revision of the genus Miridiba Reitter, 1902: genital morphotypes and new European Journal of Taxonomy 749:1-94 https://europeanjournaloftaxonomy.eu/index. taxonomic data php/ejt/article/view/1355/4215

Giraldo-Mendoza, A.E. 2021 A new species of Golofa Hope, 1837 from Peruvian Andes Studies and Reports Taxonomical Series 17(2):259-264

Gomes, E.S., Ávila, C.J. & Cherman, M.A. 2021 Biological and morphological aspects of Plectris aliena in sugarcane in Brazil Papéis Avulsos de Zoologia https://www.revistas.usp.br/paz/article/view/1 61(e20216113):1-7 74122/168358

González-Alvaredo, A. & Vaz-de-Mello, F.Z. 2021 Towards a comprehensive taxonomic revision of the Neotropical dung beetle PLos One 16(1)e0244657:1-79 https://journals.plos.org/plosone/article/file?id subgenus Deltochilum (Deltohyboma) Lane, 1946: Division into species- =10.1371/journal.pone.0244657&type=printa groups ble

Gunter, N.L. & Weir, T.A. 2021 Revision of Australian species of the dung beetle genus Lepanus: review of Zootaxa 4923(1):1-66 the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species

Gusakov, A.A. & Klimenko, A.A. 2021 Novyy vid khrushchey, Tanyproctus (Phalangonyx) legezini, iz Tsentral’nogo Humanity space. International almanac https://cyberleninka.ru/article/n/a-new- Irana 10(4):486-492 species-of-chafer-beetles-tanyproctus- phalangonyx-legezini-coleoptera- scarabaeidae-melolonthinae-pachydemini- from-central/viewer Gusakov, A.A. & Ustinov, V.E. 2021 Pervaya nakhodka Adoretus afghanus Machatschke v Tadzhikistane Humanity space. International almanac https://cyberleninka.ru/article/n/the-first- 10(4):493-496 record-of-adoretus-afghanus-machatschke- coleoptera-scarabaeidae--from- tajikistan/viewer Guzmán-Vásquez, H.M., Sánchez-García, 2021 Los escarabajos Dynastinae de Oaxaca, México: lista actualizada de Revista Mexicana de Biodiversidad http://revista.ib.unam.mx/index.php/bio/article J.A., Hernández-Cruz, J., Rös, M. & Deloya, especies y análisis de su distribución 92(e923419):1-29 /download/3419/2174 C.

Han, G.Y., Choi, J.B. & Park, J.K. 2021 A checklist of subfamily Scarabaeinae from Cambodia Journal of Asia-Pacific Biodiversity 14(1):132-139

Hauser, G., Wermelingen, B., Roth, N. & 2021 Redécouverte d'Aesalus scarabaeoides (Panzer, 1793) en Suisse Entomo Helvetica 14:135-139 Chittaro, Y.

Hensen, M.C., Hernández, M.I.M. & Lobo, 2021 Predicted distributions could suggest regional sympatry in spite of local Entomological Science 24(2):143-151 J.M. allopatry: The case of the dung beetle Canthon rutilans Castelnau, 1840

Hoermann, C., Lackner, T., Sommer, D., 2021 Carcasses at Fixed Locations Host a Higher Diversity of Necrophilous 12(5):1-18 https://www.mdpi.com/2075- Heurich, M., Benbow, M.E. & Müller, J. Beetles 4450/12/5/412/pdf

Holguin, C.M. & Mira, R.H. 2021 Report of Astaena pygidialis Kirsch, the main chafer beetle causing damage Florida Entomologist 104(1):36-41 https://journals.flvc.org/flaent/article/view/117 to avocado fruit and young leaves in Antioquia Department, Colombia 687/130204

International Commission on Zoological 2021 Case 3812 – Choeridium latum Boucomont, 1928 (currently Ateuchus latus): Bulletin of Zoological Nomenclature 78(1):6- Nomenclature proposed conservation of the specific name 16

International Commission on Zoological 2021 Opinion 2477 (Case 3768) – Hybosorus illigeri Reiche, 1853 (Insecta, Bulletin of Zoological Nomenclature 78:144- Nomenclature Coleoptera): name conserved 146

Jákl, S. 2021 A new species of Glycosia Schoch, 1896 from South Vietnam Studies and Reports. Taxonomical Series https://sar.fld.czu.cz/cache/article- 17(1):37-41 data/SaR/Published_volumes/2021-1/37- 42_jakl_1_kor_2.pdf

Jákl, S. 2021 Taxonomical notes about Liocola C. G. Thomson, 1859 and Macroliocola Studies and Reports. Taxonomical Series https://sar.fld.czu.cz/cache/article- Alexis & Delpont, 1998, subgenera of Protaetia Burmeister, 1842, with 17(1):43-55 data/SaR/Published_volumes/2021-1/43- description of new species 56_jakl_2_kor_2.pdf

Jiang, R.X., Ballerio, A., Liu, H.Y. & Wang, S. 2021 Description of the male of Pterorthochaetes yunnanensis Ballerio, 2014 Zootaxa 4950(1):196-200 https://www.mapress.com/j/zt/article/view/zoo taxa.4950.1.12/43312

Josso, J.F. 2021 Description de deux nouveaux Onthophagini de Zambie Catharsius. La Revue 22:10-14

Juřena, D. & Trojan, I. 2021 Amphimallon burmeisteri in the Czech Republic, Austria and Slovakia Klapalekiana 57:115-124

Kakizoe, S., Jiang, R.X., Cai, H.H. & Wang, 2021 Sphaeraphodius gaohani, a new genus and species of Aphodiini from Zootaxa 4949(2):333-340 S. Guangxi, China

Kalawate, A.S. & Hillert, O. 2021 Two new species of the genus Bolboceras Kirby, 1819 from India with an Zootaxa 4964(3):559-570 updated identification key of the Bolboceras nigricans Westwood, 1848 species group Kalawate, A.S., Mukhopadhyay, B., Pawar, 2021 Some new records of scarab beetles of the genus Onthophagus Latreille, Journal of threatened taxa 13(1):17580- https://www.threatenedtaxa.org/index.php/Jo S.V. & Shinde, V.D. 1802 from northern Western Ghats, Maharashtra, with a checklist 17586 TT/article/view/5695/7308

Kharel, B.P. & Sarkar, S.K. 2021 World checklist and key to Indian species with a new record of Tiniocellus Journal of Asia-Pacific Biodiversity 14:321- Péringuey, 1901 from India 327

Kon, M., Johki, Y., Ueno, H., Araya, K., 2021 Leptaulax species collected from Lambir Hills National Park, Sarawak, Giornale Italiano di Entomologia 16(66):169- Meleng, P., Kaliang, C.H. & Itioka, T. Malaysia, with descriptions of two new species 178

Krajčik, M. 2021 A list of some cetonid beetles preserved in the Natural History Museum Animma.X 93:1-8 London, with remarks on the taxonomy for interesting taxa Part I

Krajčik, M. 2021 A list of some cetonid beetles preserved in the Natural History Museum Animma.X 95:1-4 London, with remarks on the taxonomy for interesting taxa. Part III

Krajčik, M. 2021 A list of some cetonid beetles preserved in the Natural History Museum Animma.X 94:1-4 London, with remarks on the taxonomy for interesting taxa. Part II. Illustrated books for determination

Krajčik, M. 2021 A list of some cetonid beetles preserved in the Natural History Museum Animma.X 96:1-5 London, with remarks on the taxonomy for interesting taxa. Part IV

Král, D. & Sommer, D. 2021 Faunistic Records from the Czech Republic - 508 Klapalekiana 57:163-165

Král, D., Kopr, D. & Bezděk, A. 2021 First country records of Erytus aequalis and Euorodalus tersus from Albania Ecologica Montenegrina 41:35-40 https://www.biotaxa.org/em/article/download/ and Montenegro em.2021.41.5/66052

Král, D., Lu, Y. & Bai, M. 2021 Airapus rakovici, a new species from Fujian, China Zootaxa 4920(1):140-144

Král, D., Mencl, L. & Rakovič, M. 2021 A new Carinaulus species from Nepal Studies and Reports Taxonomical Series https://sar.fld.czu.cz/cache/article- 17(1):69-75 data/SaR/Published_volumes/2021-1/69- 76_kral_kor_2.pdf

Krátký, J. 2021 New distributional data on beetle species from the Canary Islands Weevil News 90:1-6 https://d-nb.info/122833126X/34

Krell, F. 2021 Taxonomy and distribution of Orubesa in the Arabian Peninsula and Zoology in the Middle East :1-12 online first surrounding areas

Lago, P.K. 2021 A review of Central American Astaena, with descriptions of new species Transactions of the American Entomological Society 147:209-471

Leo, P., Garagnani, P. & Sabatinelli, G. 2021 Le Anoxia italiane del sottogenere Mesanoxia Medvedev, 1951: Giornale Italiano di Entomologia considerazioni tassonomiche e descrizione di una nuova specie di Sardegna 16(66):211-230 e Corsica

Lin, J.Z. 2021 Description of Lucanus yulaoensis sp. nov., a new species stag beetle from Faunitaxys 9(2):1-5 https://www.faunitaxys.fr/app/download/1188 northern Taiwan 8001697/Faunitaxys_f85.pdf?t=1610405743 Lin, J.Z. & Chou, W.I. 2021 Description of a new species of the genus Neolucanus Thomson, 1862 from Faunitaxys 9(7):1-9 https://www.faunitaxys.fr/app/download/1190 Taiwan, with new localities record of N. Taiwanus 5517697/Faunitaxys_f90.pdf?t=1616942939

Martínez, I.M., Rivera-Gasperín, S.L., 2021 Comparative aspects of the internal reproductive system of males in species Acta Zoológica Mexicana (nueva serie) https://azm.ojs.inecol.mx/index.php/azm/articl Carrillo-Ruiz, H. & Morón, M.A. of Melolonthinae, Dynastinae, and Rutelinae from Mexico 37:1-17 e/view/2419/2855

McLeod, L. & Colville, J.F. 2021 Observations on unusual feeding and mating behaviour of a monkey beetle African Entomology 29(1):301-306 genus Amblymelanoplia Dombrow

Mello, C.A.A., Amorim, I.C., Silva, A.F., 2021 Mitogenome of Coprophanaeus ensifer and phylogenetic analysis of the Genetics and Molecular Biology, 44(3) https://www.scielo.br/j/gmb/a/xFBDZM7jJQcc Medeiros, G.R., Wallau, G.L. & Moura, R.C. Scarabaeidae family e20200417:1-8 7VCXr3bLFpy/?format=pdf&lang=en

Menis, F.T., Fuhrmann, J. & Rodrigues, S.R. 2021 Identification and description of the antennal sensilla of Liogenys suturalis Revista Brasileira de Entomologia https://www.scielo.br/j/rbent/a/HWqd3jq9mNX 65(3)(e20210047):1-5 tvbRbgSs5XRd/?format=pdf&lang=en

Mertlik, J. 2021 Druhy čeledí Geotrupidae, Scarabaeidae a Trogidae východních Čech – Elateridarium 15:42-93 http://www.elateridae.com/clanky/mertlik_kop výsledky faunistického mapování v roce 2020 rofagove-vc_-1d_16_2_2021.pdf

Milani, L. 2021 Descrizione di Pycnoschema kisoa Dechambre maschio, rara specie Giornale Italiano di Entomologia 16(66):1-12 endemica del Madagascar

Milani, L. 2021 Nota sul genere Golofa Hope con descrizione di una nuova specie dal Perù Giornale Italiano di Entomologia 16(66):13- 24

Minkina, L., Miessen, G. & Král, D. 2021 Mecynodes zagrosensis n.sp. from Iran Lambillionea 121(1):5-14

Moctezuma, V. 2021 El género Onthophagus Latreille, 1802 de México Dugesiana 28(2):175-220 http://dugesiana.cucba.udg.mx/index.php/DU G/article/view/7166/6499

Moctezuma, V. & Halffter, G. 2021 Taxonomic revision of the Phanaeus endymion species group, with the European Journal of Taxonomy 747:1-71 https://europeanjournaloftaxonomy.eu/index. descriptions of five new species php/ejt/article/view/1333/4099

Moctezuma, V. & Halffter, G. 2021 Species redescriptions and new species of the Onthophagus mexicanus Zoological Studies 60(30):1-30 https://zoolstud.sinica.edu.tw/Journals/60/60- species group, with notes on distribution and rodent-dung beetle 30.pdf associations

Moctezuma, V., Sánchez-Huerta, J.L., Vaz- 2021 Revalidation and redescription of Deltochilum (Calhyboma) burmeisteri, with Revista Mexicana de Biodiversidad http://www.revista.ib.unam.mx/index.php/bio/ de-Mello, F. & Halffter, G. a key to related species 92(e923712):1-9 article/download/3712/2223

Montoya-Molina, S. & Vaz-de-Mello, F.Z. 2021 Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species European Journal of Taxonomy 734:1-64 https://europeanjournaloftaxonomy.eu/index. group php/ejt/article/view/1233/3577

Montoya-Molina, S., González-Alvarado, A., 2021 Ateuchus fedescobari – a new dung beetle species from Colombia, and Journal of Natural History 55(1-2):115-124 https://www.tandfonline.com/doi/pdf/10.1080/ Giraldo-Echeverri, C. & Vaz-de-Mello, F.Z. redescription of the rare A. punctatissimus (Génier, 2010) from Brazil 00222933.2021.1895348

Moretto, P. 2021 Un nouveau Garreta Janssens, 1940, d’Afrique centrale forestière Catharsius. La Revue 22:1-9 Moretto, P. 2021 Description d’un nouvel Heliocopris Hope, 1837, d’Afrique tropicale Catharsius. La Revue 22:24-33

Moretto, P. 2021 Description d’un nouvel Onitis Fabricius, 1798, du Tchad Catharsius. La Revue 22:37-42

Moretto, P. & Josso, J.F. 2021 Synonymie d’Onthophagus fradei Gomes Alves, 1956, et Onthophagus Catharsius. La Revue 22:34-36 obenbergeri Balthasar, 1965

Moretto, P., Cosson, B., Takano, H., 2021 Un refuge forestier menacé : La forêt d’altitude à Parinari du Mont Tonkoui Catharsius. La Revue Hors-série 1:1-123 Basquin, P., Bordat, P., Boucher, S., Bouyer, en Côte d’Ivoire. Évaluation de la biodiversité entomologique, botanique et T., Danflous, S., Dérozier, V., Eitschberger, ornithologique. Valeurs de conservation du site U., Juhel, P., Leroy E., Limbourg, P., Massa, B., Meunier, J., Miles, W., Orbach, E., Robiche, G., Rojkoff, S. & Silvestre, G. Mückstein, P. 2021 A new species and a new country record in the genus Protaetia Burmeister, Zootaxa 4952(1):192-200 1842 from Vietnam

Nazaré-Silva, E.E. & Silva, F.A.B. 2021 A taxonomic revision of the South American species of Pseudocanthon Zootaxa 5027(1):61-86 Bates, 1887

Nazaré-Silva, E.E. & Silva, F.A.B. 2021 A revision of the subgenera Euhyboma Kolbe, 1893, Parahyboma Paulian, Revista Brasileira de Entomologia https://www.scielo.br/j/rbent/a/xYJsrkRYpQhJ 1938, and Rubrohyboma Paulian, 1939 of Deltochilum Eschscholtz, 1822 65(3)(e20200100):1-19 4QxJNqwxSGy/?lang=en&format=pdf

Neita-Moreno, J.C. 2021 A review of the black species of Cyclocephala Dejean from Colombia Zootaxa 5026(1):1-58

Neita-Moreno, J.C., Castro-Vargas, M.I., 2021 Descripción de los estados inmaduros de Platycoelia puncticollis, código de Revista Mexicana de Biodiversidad http://www.revista.ib.unam.mx/index.php/bio/ Ibarra-Polesel, M.G. & Pulido-Santacruz, P. barras de ADN y clave para las larvas de tercer estadio del género 92(e923752):1-13 article/download/3752/2245

Ocampo, F.C. 2021 Two new species of the Neotropical genus Anaides from Peru Revista de la Sociedad Entomológica https://www.biotaxa.org/RSEA/article/view/69 Argentina 80(2):12-16 883/67481

Ochi, T. & Kon, M. 2021 Notes on the Coprophagous Scarab Beetles from Southeast Asia. XXXIV Giornale Italiano di Entomologia 16(66):149- Four new species of Copris Müller from Myanmar, Laos and Cambodia 158

Ochi, T. & Sakai, K. 2021 A revisional study of the genus Phaedimus Waterhouse, 1841 from the Giornale Italiano di Entomologia 16(66):25- Philippines 48

Ochi, T., Kon, M. & Bai, M. 2021 Four new species of the genera Odontotrypes and Phelotrupes from China Giornale Italiano di Entomologia 16(66):133- 142

Ochi, T., Kon, M. & Kakizoe, S. 2021 Three new species of Nothochodaeus Nikolajev from Myanmar and Laos Giornale Italiano di Entomologia 16(66):179- 188

Özgen, I. & Çam, A. 2021 An endemic species very important for eco-tourism: Pygopleurus koniae and International Journal of Fauna and https://www.faunajournal.com/archives/2021/ new nomenclature “Furry wedding beetle” and the Turkish name: “Kürklü Biological Studies 8(2):113-116 vol8issue2/PartB/8-3-4-868.pdf Düğün Böceği” Pacheco, T.L., Monné, M.L., Vaz-de-Mello, 2021 Revision of type specimens of Astaena described by L.W. Saylor European Journal of Taxonomy 750:94-123 https://europeanjournaloftaxonomy.eu/index. F.Z. & Ahrens, D. php/ejt/article/view/1365/4265

Perissinotto, R. & Clennell, L. 2021 Census of the fruit and flower chafers of the Macau SAR, China ZooKeys 1026:17-43 https://zookeys.pensoft.net/article/60036/dow nload/pdf/524439

Pham, P.V. & Pham, N.T. 2021 A remarkable new species of the genus Macronotops Krikken, 1977 from Zootaxa 5039(4):593-598 Vietnam

Poloni, R. & Morelli, A. 2021 New or interesting beetles from the Abruzzi region (Central Italy) Bollettino della Società entomologica italiana 153(1):37-41

Popa, A.F., Motoc, R.M., Iorgu, E.I., 2021 Morphological variability and teratologies in the stag beetle Lucanus cervus Biologia 76(8):2181-2193 Costache, M. & Popa, L.O. from Romania

Prandi, M. & Grossi, P.C. 2021 A new species of Eupatorus Burmeister, 1847 related to Eupatorus Zootaxa 4966(1):29-40 birmanicus Arrow, 1908 from southwestern China

Prušáková, D., Peska, V., Pekár, S., 2021 Telomeric DNA sequences in beetle taxa vary with species richness Scientific Reports 11(13319):1-15 https://www.nature.com/articles/s41598-021- Bubenik, M., Čížek, L., Bezděk, A. & 92705-y.pdf Frydrychová, R.C.

Qi, Z.H. 2021 Lucanus moae sp. nov., a new species from Sichuan, China Faunitaxys 9(23):1-7 https://www.faunitaxys.fr/app/download/1200 7563797/Faunitaxys_f106.pdf?t=1632404617

Qiu, J.Y., Zhao, M.Z. & Xu, H. 2021 Description of a new species of the genus Didrepanephorus Wood-Mason, Zootaxa 4933(4):493-513 1878 from China with notes on allied species

Rakovič, M. & Anichtchenko, A. 2021 Two new species of the genus Odochilus Harold, 1877 Zootaxa 4942(3):447-450

Ramírez-Ponce, A. & Galindo, O. 2021 A new species of Heterosternus Dupont from Mexico Studies on Neotropical Fauna and Environment :1-7 online first

Ramírez-Ponce, A., Delgado, L. & Soto, Y. 2021 A new species of Epectinaspis Blanchard from Guatemala and rediscovery Studies on Neotropical Fauna and of Balanogonia constricta Paucar-Cabrera Environment :1-8 online first

Ratcliffe, B.C. & Seidel, M. 2021 A new species of Chalepides Casey, 1915 from the Pantanal of Brazil Evolutionary Systematics 5:189-192

Reddy, P.V.R. & Kurian, R.M. 2021 Occurence of fruit chafer beetle Protaetia alboguttata (Vigors) on fig Indian Journal of Entomology 83(e20174):1- 3

Rivera-Gasperin, S.L., Escobar-Hernández, 2021 An update on the feeding preferences of the dung beetle Onthophagus The Coleopterists Bulletin 75(1):247-255 F. & Favila, M.E. moroni Zunino and Halffter from the Cuetzalan caves, Puebla, Mexico

Rizzotto, M., Negro, M. & Barbero, E. 2021 Some traits of the biology and epigean behavior of Sisyphus schaefferi The Coleopterists Bulletin 75(1):75-85 Rossini, M. 2021 Additional mislabeling in African Onthophagus Latreille, 1802: the case of Zootaxa 5032(2):262-274 Onthophagus viviensis and Onthophagus laevatus

Rossini, M., Montreuil, O., Grebennikov, V. & 2021 Genome sequencing reveals extraordinary cephalic horns in the ZooKeys 1033:63-79 https://zookeys.pensoft.net/article/63527/dow Tarasov, S. Madagascan dung beetle genus Helictopleurus: insight from a revision of nload/pdf/534907 fungicola species group

Ruchin, A.B., Egorov, L.V. & Polumordvinov, 2021 Coleoptera of the Penza region, Russia based on fermental crown trap Biodiversitas 22(4):1946-1960 https://smujo.id/biodiv/article/view/7847/4743 O.A.

Ruchin, A.B., Egorov, L.V., Alekseev, S.K., 2021 Notes on the fauna of beetles adjacent to the territory of the Mordovia State Amurian Zoological Journal 16(1):12-35 https://azjournal.ru/index.php/azjournal/article Semishin, G.B. & Esin, M.N. Nature Reserve /view/601/544

Sabatinelli, G. & Pham, P. 2021 Taxonomic notes on the genus Cyphochilus (part 4) with description of eight Revue suisse de Zoologie 128(1):157-172 new species from Indochina and China

Seidel, M. 2021 New records of Anomala tibialis Lansberge, 1886 and a new replacement Zootaxa 4966(4):494-496 name for Anomala tibialis Schaeffer, 1906

Seidel, M. & Reid, C.A.M. 2021 Taxonomic changes resulting from a review of the types of Australian Zootaxa 4908(2):225-238 https://www.mapress.com/j/zt/article/downloa Anoplognathini housed in Swedish natural history collections d/zootaxa.4908.2.4/42501

Sergeeva, E.V. & Stolbov, V.A. 2021 The fauna of lamellicorn beetles of the Tyumen Region Amurian Zoological Journal 16(1):124-145 https://azjournal.ru/index.php/azjournal/article /view/630/551

Shabalin, S.A. 2021 Distribution of two species of dung-beetles in Primorsky Krai A.I. Kurentsov's Annual Memorial Meetings 32:88-92

Silva, F.A.B. & Valois, M. 2021 Description of larva and adult female of Golofa claviger (Linnaeus, 1771), Austral Entomology 60(2):351-363 with notes on taxonomy and biology

Skrylnik, Y.E. 2021 A new species of Ceratolontha Arrow, 1948 from Peru Humanity space International almanac 10(4):513-517

Skrylnik, Y.E. & Pak, O.V. 2021 A new species of Glaphyrus Latreille, 1802 (subgenus Eoglaphyrus Humanity space International almanac Semenov, 1926) from Afghanistan 10(4):518-528

Smith, A.B.T. 2021 Two New Species of Phalangogonia Burmeister, 1844 The Coleopterists Bulletin 75(1):259-265

Sommer, D., Hillert, O., Hrůzová, L. & Král, 2021 Bolbelasmus (Bolbelasmus) zagrosensis, a new species from Iran, along Zootaxa 4920(3):380-394 D. with an updated key to the western Palaearctic species of the subgenus

Sommer, D., Král, D. & Týr, V. 2021 Faunistické zprávy ze západních Čech – 19. Coleoptera: Scarabaeidae: Západočeské entomologické listy 12:16-18 http://www.entolisty.cz/2021/12-02- Aphodiinae. (Faunistic records from western Bohemia - 19) Sommer.pdf

Sousa, R. & Seidel, M. 2021 Review of the Inca irroratus species group with description of two new European Journal of Taxonomy 748:15-35 https://europeanjournaloftaxonomy.eu/index. species of Inca LePeletier & Serville, 1828 php/ejt/article/view/1335/4119 Stanbrook, R.A., Harris, W.E., Wheater, C.P. 2021 Evidence of phenotypic plasticity along an altitudinal gradient in the dung PeerJ 9:e10798:1-15 https://peerj.com/articles/10798.pdf & Jones, M. beetle Onthophagus proteus

Strümpher, W.P. & Daniel, G.M. 2021 A new dung beetle species of the genus Stiptopodius Harold, 1871 recorded Zootaxa 4999(4):335-342 from xeric savanna in Namibia

Strümpher, W.P. & Stals, R. 2021 Proper cleaning of keratin beetles reveals a cryptic species: Phoberus Zootaxa 4942(4):543-557 https://www.mapress.com/j/zt/article/downloa fumarius (Haaf, 1953) from southern Africa is reinstated as valid d/zootaxa.4942.4.3/43111

Tauzin, P. 2021 Commentaires sur Fornasinius insignis ab. funereus Burgeon 1937 Le Coléopteriste 24(1):14-15

Tello, F., Verdú, J.R., Rossini, M. & Zunino, 2021 Onthophagus pilauco sp. nov.: evidence of beetle extinction in the ZooKeys 1043:133-145 https://zookeys.pensoft.net/article/61706/dow M. Pleistocene–Holocene transition in Chilean Northern Patagonia nload/pdf/555634

Tonelli, M., Giménez-Gómez, V.C., Verdú, 2021 Dung Beetle Assemblages Attracted to Cow and Horse Dung: The Life 11(873):1-13 https://www.mdpi.com/2075- J.R., Casanoves, F. & Zunino, M. Importance of Mouthpart Traits, Body Size, and Nesting Behavior in the 1729/11/9/873/pdf Community Assembly Process

Trócoli, S. 2021 Nuevas citas de coleópteros para el Parque Natural de Sant Llorenç del Revista gaditana de Entomología 12:35-40 https://archive.org/download/rg-e-vol-xii-35- Munt i Serra de l’ Obac (Barcelona, Península Ibérica) 2ª Nota 40.-nuevas-citas-pn-s.-llorenc-del- munt/RgE%20vol%20XII%2C%2035- 40.%20Nuevas%20citas%20PN%20S.%20Ll oren%C3%A7%20del%20Munt%20.pdf Valan, M., Vondráček, D. & Ronquist, F. 2021 Awakening a taxonomist’s third eye: exploring the utility of computer vision Systematic Entomology 46(4):757-766 https://onlinelibrary.wiley.com/doi/epdf/10.111 and deep learning in insect systematics 1/syen.12492

Vallejos-Garrido, P. & Fuentes-Olivares, P. 2021 Nuevo caso teratológico mandibular en un ejemplar de Chiasognathus Revista Chilena de Entomología 47(2):385- http://www.insectachile.cl/rchen/pdfs/2021v47 grantii Stephens, 1831 de Chiloé, Chile 389 -2/Vallejos-Garrido_y_Fuentes- Olivares_2021.pdf

Wang, F.L. 2021 A new species of the genus Anomala Samouelle, 1819 from China Far Eastern Entomologist 424:14-20 https://www.biosoil.ru/Files/FEE/00002035.pd f

Wang, F.L. 2021 Description of four new species of genus Anomala Samouelle from South Studies and Reports. Taxonomical Series https://sar.fld.czu.cz/cache/article- China 17(1):189-200 data/SaR/Published_volumes/2021-1/189- 200_wang_kor.pdf

Wang, F.L. & Zorn, C. 2021 Description of a new species and new faunistic records of the genus Contributions to Entomology 71(1):147-159 Anomala Samouelle from China and neighboring regions

Warner, W.B. 2021 Two new scarab beetles from the southwestern USA Insecta Mundi 855:1-15 https://centerforsystematicentomology.org/ins ectamundi/0855_Warner_2021.pdf

Wojas, T. & Kopacz, A. 2021 Nowe dane o rozmieszczeniu Chaetopteroplia segetum (Herbst, 1783) w Wiadomości Entomologiczne 40(1):5-6 http://pte.au.poznan.pl/we/2021/3N-Wojas-&- południowej Polsce Kopacz.pdf

Yang, H., You, C.J., Tsui, C.K.M., Tembrock, 2021 Phylogeny and biogeography of the Japanese rhinoceros beetle, Trypoxylus Ecology and Evolution 11(1):153-173 https://www.onlinelibrary.wiley.com/doi/epdf/1 L.R., Wu, Z.Q. & Yang, D.P. dichotomus based on SNP markers 0.1002/ece3.6982

Ying, Y., Zhan, Z.H. & Wan, X. 2021 New color patterns and new synonym of Odontolabis sinensis (Westwood, Zootaxa 4926(2):263-275 1848): insights from a multilocus phylogeny and species delimitation Yuan, J.J., Chen, D. & Wan, X. 2021 A multilocus assessment reveals two new synonymies for East Asian ZooKeys 1021:65-79 https://zookeys.pensoft.net/article/58832/dow Cyclommatus stag beetles nload/pdf/515661

Zhao, M.Z. 2021 Contribution to the genus Anomala Samouelle, 1819 of China and adjacent Zootaxa 4903(4):578-590 regions. Part I: descriptions of two new species and remarks on four species

Zhao, M.Z. 2021 On the genus Cyphochilus Waterhouse, 1867 from Hainan Island, China Faunitaxys 9(18):1-8 https://www.faunitaxys.fr/app/download/1198 7584297/Faunitaxys_f101.pdf?t=1623361803

Zhao, M.Z. 2021 On the genus Mimela Kirby, 1823 from China and adjacent countries, with Zootaxa 4995(2):201-230 description of five new species

Ziani, S. 2021 Subsequent spelling ritchiei for Mnematium ritchii Macleay, 1821 to be Insecta Mundi 856:1-5 https://centerforsystematicentomology.org/ins maintained for prevailing usage ectamundi/0856_Ziani_2021.pdf

Zídek, J. 2021 Revision of Philacelota Heller, 1900 and Engertia Dalla Torre, 1912, with Telnov, D. et al. (eds): Biodiversity, description of a new genus Biogeography and Nature Conservation in Wallacea and New Guinea, IV :397-411

Zilli, A. 2021 On the date of publication of Linnaeus’ second edition of “Fauna Svecica” Nota lepidopterologica 44:69-71

Zinchenko, V.K. 2021 A new record of rare dung beetle from Kazakhstan Euroasian Entomological Journal 20(1):52- https://kmkjournals.com/upload/PDF/EEJ/20/ 54 20_1_052_054_Zinchenko.pdf

Zothansanga, C. 2021 A review of scarab beetles diversity in India Science Vision 21(2):43-49 https://www.sciencevision.org/storage/journal - articles/June2021/KhI85WDqPS7TXQC1m2Ij .pdf Zurawlew, P. & Bunalski, M. 2021 Stwierdzenia kwietnicy okazałej Protaetia speciosissima (Scopoli, 1786) z Wiadomości Entomologiczne 40(2):3-4 http://pte.au.poznan.pl/we/2021/5N- dala od miejsca potencjalnego rozwoju oraz informacje o jej występowaniu w Zurawlew-i-Bunalski.pdf powiecie pleszewskim (Nizina Wielkopolsko-Kujawska)