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Home , Acalles, Acanthinus, Acanthocerus, Acanthocinus, Acanthoderes, Acanthoderini

of Florida and Neighboring Land Areas

Volume 18

A Checklist of the of with Data on Distributions and Bionomics (Insecta: Coleoptera)

Stewart B. Peck

Florida Department of Agriculture and Consumer Services Charles H. Bronson, Commissioner

Division of Plant Industry Richard Gaskalla, Director P.O. Box 147100 Gainesville, FL 32614-7100

Arthropods of Florida and Neighboring Land Areas

Volume 18

A Checklist of the Beetles of Cuba with Data on Distributions and Bionomics (Insecta: Coleoptera)

Stewart B. Peck Department of Biology 1125 Colonel By Drive Carleton University, Ottawa, K1S 5B6, Canada

Florida Department of Agriculture and Consumer Services Charles H. Bronson, Commissioner

Division of Plant Industry Richard Gaskalla, Director P.O. Box 147100 Gainesville, FL 32614-7100 Fernando de Zayas (1912-1983), Entomologist, Cuban Academy of Sciences Table of Contents

Abstract ...... Introduction ...... Summary of the history of Cuban studies ...... Construction of a checklist of Cuban beetles ...... Summary and origins of the beetle fauna of Cuba ...... Annotated list of the beetles (Coleoptera) of Cuba* ...... Suborder 4. Micromalthidae ...... Suborder 9. ...... 10. Carabidae (+Cicindelinae) ...... 11. Gyrinidae...... 12. ...... 14. ...... 17. ...... Suborder Series Superfamily 18. ...... 21. ...... Superfamily 22. ...... 23. ...... 25. ...... 26. Scydmaenidae ...... 28. Staphylinidae (+ and Pselaphinae) ...... Series Scarabaeiformia Superfamily 31. ...... 33. ...... 35. ...... 37. ...... 38. ...... 39. Ceratocanthidae ...... 41. ...... Series Superfamily 47. ...... Superfamily 51. ...... Superfamily 53. ...... 54. ...... 55. Lutrochidae ...... 56. Limnichidae ...... 57. ...... 58. Psephenidae...... 60. ...... 61. ......

* Families known to be present in Cuba, numbered following the world list of beetle families in Arnett and Thomas (2000). Superfamily 67. ...... 69. Elateridae ...... 73. ...... 76. Lampyridae ...... 78. Cantharidae...... Series Superfamily 81. ...... 82. ...... 83. (+Lyctinae) ...... 84. Anobiidae (+Ptininae)...... Series Superfamily Lymexyloidea 85. ...... Superfamily 87. ...... 89. ...... 93. (+Malachiinae) ...... Superfamily 95. ...... 97. Nitidulidae ...... 98. ...... 99. (+Rhizophaginae) ...... 103. ...... 104. ...... 106. ...... 108. ...... 111. ...... 113. Languriidae ...... 114. ...... 116. ...... 117. ...... 118. ...... 120. ...... 121. ...... 122. ...... 123. Corylophidae ...... 124. ...... Superfamily 125. ...... 128. ...... 130. ...... 131. ...... 132. Rhipiphoridae ...... 133. Colydiidae ...... 134. Monommatidae ...... 135. ...... 140. Tenebrionidae (+Alleculinae and Lagriinae) ...... 143. ...... 145. Meloidae ...... 146. ...... 151. ...... 152. ...... 153. ...... 154. ...... Superfamily 155. Cerambycidae ...... 156. Bruchidae ...... 159. Chrysomelidae ...... Superfamily Curculionoidea 161. ...... 162. ...... 163. ...... 164. ...... 167. ( + Scolytinae and Platypodinae) ......

ORDER STREPSIPTERA ...... Suborder Stylopidia 2. Corioxenidae ...... 4. Myrmecolacidae ...... 5. Halictophagidae Abstract

This summary of the beetle fauna of Cuba lists 87 families, 954 genera, and 2,673 species plus three families, three genera, and three species of strepsipterans. Many families in Cuba are poorly or not at all studied and additional species will be found. The fauna contains 1,495 endemic species - species which are naturally limited to Cuba. There are 1,092 native species - these have natural distributions in adjacent or surrounding land masses (the Bahama , Florida and the southeastern , , the , México and , and and other islands to the east). There are 86 species which are recognized to have been introduced by human activities and many of these are pests of agriculture, forestry, or stored products. The fauna is disharmonic (unbalanced), as is expected for an oceanic fauna which has long been isolated by water. Several families that occur on the continental margins of the Caribbean are apparently not present in Cuba. Geological history suggests that Cuba has existed as an island and has been available for colonization for the past 35 million years. Species movement to and from Cuba may have been more frequent at times of past low sea levels during Pleistocene glacials, when the neighboring continental margins and Caribbean islands were more exposed and the water gaps between them and Cuba were smaller. Twenty-eight genera contain 10 or more species limited to Cuba and may represent island “species swarms.” A Checklist of the Beetles of Cuba 1

Figure 1. Outline map of Cuba, showing present provinces.

Introduction

A fundamental part of the patrimony or natu- Many groups, such as vascular plants and birds ral heritage of every nation is the diversity of its are now well known in Cuba. Less well known are biological species. Cuba is a tropical island nation the invertebrate , especially the . Of which is exceptionally well endowed with a rich course there are exceptions to this. Some insects array of plant and species in both marine have received much study and their species and and terrestrial environments. However, this island distributions are well known. Two such examples with the richest known biota in the are the butterflies (Alayo and Hernandez 1987) and remains little known by most entomologists, biolo- the plant-feeding aphids (Holman 1974) of Cuba. gists, and conservationists outside of Cuba. Reviews of various groups of terrestrial inverte- The Republic of Cuba includes more than 1,600 brates are in the newsletter ”Cocuyo”, published islands and covers an area of 110,922 km2. Cuba since 1994 by the National Museum of Natural harbours the greatest species diversity of the West History, , Cuba. Indies (Woods 1989), with more than 6,000 species It is estimated that Cuba was about 60% cov- of plants, 552 species of terrestrial vertebrates, 778 ered by various kinds of forest formations at the species of fish, probably more than 15,000 species time of its discovery by Columbus in 1492. Four of insects, and an additional large number of myri- hundred years of Spanish colonial occupation did apods, terrestrial crustaceans, land snails, and not much change this. In the late 1800’s forests arachnids. still covered an estimated 56% of the island (Smith Genaro and Tejuca (1999) provide a count of 1954). With the independence of Cuba from Spain 8,312 species of insects in the 29 insect orders now in 1898, and the creation of a preferential tariff for known in Cuba, and they provide an introduction the importation of sugar into the USA, the rate of to a review of Cuban insects in general. Cuba also clearing of native forest cover was greatly acceler- has the highest amount of West Indian endemism ated. By 1959 more than 4.4 million ha (hectares) (species unique to the country); more than 32% of of forests had been destroyed or greatly altered and its vertebrates (approximately 90% of the amphib- forested areas had been reduced to only 14% of Cuba’s ians and 85% of the reptiles), and 50% of its flower- land area. The deforestation process has now been ing plants naturally occur only on this island. And reversed, and natural and plantation forest cover many of the endemics are limited to only parts of has increased to approximately 18% of the land area. the island. Of the some 6,000 species of flowering The Food and Agriculture Organization of the plants, 1,575 occur only in eastern Cuba (López United Nations identifies Cuba as now having the Almirall 1998). Vales et al. (1998) estimate that lowest annual deforestation rate (0.1%) in Latin some 30% of Cuba’s insects are endemic. America. 2 Arthropods of Florida and Neighboring Land Areas, Vol. 18

The first Cuban national park, Parque Nacional ing of the insect fauna is more advanced for Cuba Pico Cristal, was established in 1930, and only five than for any other island in the Caribbean. more had been established by 1959. Then, nine Cuba is the natural biological and cultural additional national parks were established under crossroads of the . An understand- Public Law 239 in 1959. The first attempt to clas- ing of the dynamics of insect movement into and sify and preserve the country’s remaining natural out of Cuba over evolutionary and historical time forests was that of Samek (1968). Later, Miller is important for applied biology and management (1974) provided the basis for establishing the present purposes. Knowledge of insect pests is important system of protected areas. To insure adequate rep- for their control. Knowledge of insects in general is resentation of all natural landscapes, geographers an important part of the cultural heritage of all and biologists evaluated their protected status. societies, because all these species are fellow occu- These efforts resulted in a law establishing a Na- pants of the biosphere. tional System of Protected Areas, which now has more than 200 protected areas that cover 12% of References the country. However, only 1-2% of the country is strictly protected and some reserves appear to be Alayo D., P. and Hernandez, L.R. 1987. Atlas too small to effectively preserve the biota they con- de las mariposas diurnas de Cuba (: tain. It is estimated that Cuba now has about two Rhopalocera). Editorial Cientifico-Téchnica, La million ha of forests, of which 1.7 million ha (85%) Havana, Cuba. 148 pp. consist of natural forests and the remainder is for- Cuevas, J.R. and García, F. 1982. Los recursos estry plantations. Of these forested lands, national naturales y su conservación. Editorial Pueblo parks cover 5.1%, wildlife conservation areas are y Educación, La Havana, Cuba. 44 pp. 24.3%, watershed protection areas are 17.0%, Genaro, J. A. and Tejuca, A. E. 1999. Datos coastal protection areas are 18.2%, production for- cuantitativos, endemismo y estado actual del ests are 32.7%, and other categories are 2.7% conocimiento de los insectos cubaños. Cucuyo (Santana 1991, Cuevas and García 1982). 8: 24-28. The purposes of this checklist of Cuban beetles Holman, J. 1974. Los aphidos de Cuba. Instituto are several. It may be used to help evaluate pos- Cubano del Libro, La Havana, Cuba. sible pest status of beetles moving into and out of López Almirall, A. 1998. Algunas characterísticas Cuba through human activities. The advantages del endemismo en la flora de Cuba Oriental. of this are evident. Some beetles are pests of agri- Pp. 47-82. In Halffter, G. (ed.). La diversidad cultural, horticultural, and forest plants; some are biológica de Iberoamerica. Vol. 2. Acta Zooogica pests of stored agricultural products, or are destruc- Mexicana, nueva series, volumen especial 1998. tive to man-made structures; and some are actual Miller, K. 1974. Manejo y desarrollo integral de or potential vectors of diseases of livestock, crops, las áreas naturales y culturales. Centro de and humans. To unify information on these makes Investigaciones y Capacitación Forestal, La the data more accessible and useful to the benefit Havana, Cuba. 102 pp. of human society. However, most beetles (as is also Samek, V. 1968. La protección de la naturaleza true for almost all insects) have little or no direct en Cuba. Serie Transformación de la impact on human activities. These species comprise Naturaleza. Academia de Ciencias de Cuba, La most of the richness of the vibrant natural com- Havana, Cuba. No. 7. 23 pp. munities of the island of Cuba. Santana, C.E. 1991. Native conservation and sus- This list may help to stimulate and encourage tainable development in Cuba. Conservation additional research activity in summarizing Biology 5: 13-16. information on the beetles of Cuba. All Smith, E. E. 1954. The forests of Cuba. Maria the species of Cuban insects have been shaped and Moors Cabot Foundation Publ. 2, Cambridge formed by the biological and physical processes of Forest, Petersham, Massachusetts, and the evolution by natural selection. The geographic dis- Atkins Garden and Research Laboratory, tributions of the species within and beyond Cuba Cienfuegos, Cuba. 98 pp. have been shaped by their genetic adaptations to Vales, M. A., Montes, L. and Alayo, R. 1998. the ecological conditions of the local environments Estado del cononcimiento de la biodiversidad and physiography, and human changes to the land- en Cuba. pp. 239-249. In Halffter, G. (ed.). La scape. All of this is a part of the patrimony of Cuba. Biodiversidad biológica de Iberoamerica. Acta. The history of study of insects (and especially Zool. Mexicana, of beetles) in Cuba is long and rich. The science of Woods, C.A. 1989. Biogeography of the West insect in Cuba is the most highly devel- Indies: past, present and future. Sandhill Crane oped of any nation in the Caribbean. An understand- Press, Gainesville, Florida. A Checklist of the Beetles of Cuba 3

Figure 2. Percentages of the total of 1,092 native Cuban beetle species shared with adjacent lands in and around the Caribbean. These are an indicator of the relative frequency of successful natural “over-water” colonizations to and from Cuba and other land masses. These numbers are subject to change as additional research on the identity and geographic distribution of Caribbean beetles continues. Number of native species shared only with one other geographic area are as follows: Bahamas, 17; Jamaica, 27; Cayman Islands, 1; Hispaniola, 175; Mexico, 46; USA, 120; , 15. Summary of the history of Cuban beetle studies

Numerous naturalists and entomlogists con- the collecting activities of this most famous and tributed to the historical development of knowledge influential Cuban naturalist. He published during of the beetles of Cuba. Many of these were Euro- the period from 1831 to 1868. His best known work pean taxonomists who first published the names is “Memorias sobre la Historia Natural de la Isla and scientific descriptions of the species themselves. de Cuba” in two volumes (1851-1861) (Norman I here briefly introduce the most important people 1938). In this he treated some entomological taxa, who actually did the field work and collecting of especially butterflies and beetles. Chapter 25 in his the specimens. This was the foundation activity in “Memorias” (1851) is about beetles, listing what was exploring and documenting the beetle fauna of Cuba. known at that time but containing no new infor- Data on the early European or American taxono- mation. mists who described the species can be found in Poey provided many of the specimens of beetle sources such as Horn and Kahle (1935). species described by Jacquelin du Val (1856-1857) Felipe Poey y Aloy (1799-1891) is considered in Sagra’s (1856-1857) “History of the Island of to be the father of Cuban zoology. The general study Cuba.” He is still honored in the name of the bio- of insects and other animals in Cuba started with logical journal “Poeyana,” published by the Instituto 4 Arthropods of Florida and Neighboring Land Areas, Vol. 18 de Ecologia y Systematica of the Academia de tion in Havana does not contain any designated Ciencias de Cuba. He exchanged an historically type specimens of beetles, but is mostly composed important collection of identified beetles and moths of specimens compared to “types”, or specimens with the Academy of Natural Sciences of Philadel- which represent part of the series from which the phia. The beetles are now incorporated into the gen- “types” came. The specimens in the Gundlach col- eral beetle collections in Philadelphia. The moths lection bear no labels except for numbers which were exchanged with the Carnegie Museum of Pitts- are coded to their identity as published in volume 3 burgh. More data on Sagra and Poey and other of his “Contribucion.” Ragués (1914) presented a naturalists mentioned below can be found in Alvarez catalogue of the collection, consisting of species (1958), Chardon (1949), and Papavero (1971: 179- names and their Gundlach code numbers. 185). The Gundlach insect collection is now housed Juan (Johannes) Gundlach (1810-1896) con- in glass-topped drawers in large wooden cases in a tributed greatly to the foundations of a tradition of dehumidified room in the Institute of Ecology and Cuban natural history studies (Reyes et al. 2002). Systematics (IES) of the Cuban Academy of Sci- He published from 1866 to 1894. He can be consid- ences, located on Carretera de Varona km 3½, ered as the father of Cuban entomology. He was Capdevila, , La Havana, Cuba. This is about born in Marburg, Germany and attained a Ph.D. 10 km SW of the old center of Havana. The degree in Marburg in 1838. He was then invited to Gundlach collection is kept separate from the Cuba by Mr. Charles Booth of Fundador de Canimar Institute’s general (non-historical) beetle collection. on the Canimar River in Matanzas Province where The entire insect collection of the Institute has a he worked as a professional collector. In 1841 Booth total of about 67,550 insect specimens (Reyes and and Gundlach left Canimar and moved to the es- Hernandez 1995). The many “syntypes” or “cotypes” tate “San Juan” near the town of Cárdenas, near of Cuban species described by Chevrolat and many the city of Matanzas. In 1846 he established a mu- other European taxonomists are scattered in many seum to hold his collections on the farm “El Refugio” European museums. Their possible location can near Cárdenas. From this he radiated throughout often be deduced by consulting Horn and Kahle Cuba on many collecting trips. In 1864 he moved (l935-1937). I have not tried to verify the existence his museum to the Fermina sugar estate of the or location of this early type material. Cárdenas family. He continued to collect through- E.A. Schwarz (1844-1928) was perhaps the out the island until the “Ten Years Revolution” next most important beetle collector to work in (starting in 1868), when it became too dangerous Cuba. He was employed by the U.S. Department of to do field work. In 1876 his collection was pur- Agriculture to investigate the status of the cotton chased by the Academy of Natural Sciences in Ha- boll (Anthonomus) in Cuba in 1902, and from vana. In 1873, 1875 and 1881 he made prolonged February 1903 to June 1904. He was stationed field trips to . He then formed a second mostly at the plantation of La Magdalena, better Cuban collection, which was deeded to the Museum known as “Cayamas”, about 10 km WNW of of Havana in 1892. More details of his life and bib- Yaguaramas (N 22º 14’; W 80º 48’), about 40 km liography are given by Ramsden (1915). NW of Cienfuegos (Schwarz 1905). Schwarz did His major publication about beetles is volume much general beetle collecting in the vicinity and 3 (404 pages) of his “Contribucion a la Entomologia Cayamas is the type locality for more Cuban beetle Cubana” (Gundlach 1891). This summarizes the species than any other site. Type specimens of all beetle fauna then known, and his 46 years of obser- or most of these should be in the U.S. National vations upon it. Apparently pages 259-344 were Museum of Natural History in Washington, D.C. actually published in 1894 and are cited in some A series of letters by Schwarz (1905, 1929) describes publications with this date. Unfortunately, this his Cuban localities and the often extremely un- volume has many typographical errors, especially pleasant and taxing field conditions he experienced in the Latin names: these are attributable to or endured. Howard et al. (1928) give a biography Gundlach’s failing eyesight and inability to correct and bibliography of Schwarz. Holland and Schwarz proofs of the volume. (1917) give a list of beetles known from the collec- Gundlach was an exceedingly generous person. tions of G.A. Link on the Isle of Pines (now Isla de Many of the species first collected by him were de- la Juventud). The specimens are in the Carnegie scribed by others, especially in an important series Museum (Pittsburgh). of papers by Chevrolat (1838-1870) and Chevrolat Carlos (Charles) Ramsden de la Torre (1876- and Fauvel (1863). The present Gundlach collec- 1950) was a rancher and versatile amateur natu- A Checklist of the Beetles of Cuba 5

ralist. He became the colleague and then son-in- beetles. He also collected many other beetles and law of the influential naturalist Professor Carlos other insects in the mountains of central Cuba de la Torre of Santiago de Cuba (who knew Juan around Cienfuegos and , and in eastern Gundlach well). Ramsden made an extensive natu- Cuba (then called Oriente Province). He may have ral history collection in eastern Cuba, especially been the first foreign insect collector to work the around Guantánamo Bay, and this is now housed summit areas of Pico Turquino, the highest point in the Museo Carlos Ramsden of the Universidad in Cuba, in the Sierra Maestra. The collections are de Oriente in Santiago de Cuba, which was estab- in the Museum of Comparative Zoology and his field lished in 1953. The museum houses a large insect notes are preserved in the files of the Department collection, including 20 drawers (about 3000 speci- of Entomology of the Museum of Comparative Zool- mens) of beetles. It is maintained as an historically ogy. He wrote several papers describing Cuban spe- important collection, and other more recent mate- cies, and the most important are Darlington (1934 rial is kept separate. Ramsden was a prolific corre- and 1937). Ball (1985) is a source of information spondent with and host to visiting American natu- about the work and contributions of Darlington. ralists. He provided beetles to many specialists in The center of activity of Darlington’s Cuban the United States, mostly collected around trips was the Atkins Botanical Garden of Harvard Guantánamo, especially at the family farm University, at Soledad in the Arimao River Valley, “Estancia San Carlos.” Many of his specimens are 18 km E of Cienfuegos, at about N 22º 5', W 80º 18'. in the American Museum of Natural History, New It was estsablished as a biological laboratory in 1924 York City. (Darlington 1938). Soledad is the type locality for Alexander Bierig (1884?-1963) was born in many beetle species. Barbour (1945) gives descrip- Karlsruhe, Germany. As a student and young man tions of conditions when the Garden was operated he studied graphic arts in Germany and Paris. In as a field station. Many young American biologists the difficult economic times following WW I, Bierig first experienced tropical biology in the Soledad area moved to Havana in 1919, where he found good from the 1930’s to the 1950’s. Of its 101 ha area, employment as a graphic artist. He taught draw- about 7 ha of the Garden are still preserved as a ing and natural science at the German School in natural example of tropical lowland semi-decidu- Havana, as well as his own private art school. His ous forest. The site was still operated as a botani- previous insect interests became well developed at cal garden and tourist attraction until it was ex- this time, and he published 27 papers on Neotropi- tensively damaged by a hurricane in the mid- cal staphylinid beetles by 1940. He collected mostly 1990’s. in western Cuba around Havana and in Pinar del Stephen Cole Bruner (1891-1953) was an Rio Province. He described more than 150 new beetle American entomologist who became director of re- species, many of them from Cuba, and almost all search at the Agronomic Experimental Station were very attractively and accurately illustrated. (EEA) at , some 12 km SW of In 1939 he moved to San José, Costa Rica, and be- the center of . This was once the main came a professor of entomology at the University of center for Cuban applied insect research. After the Costa Rica. There he mostly specialized in the biol- closure of the Experimental Station the collections ogy and control of insects injurious to tropical crops were transferred to the Institute of Ecology and until 1954, when he stopped entomological work. Systematics of the Cuban Academy of Sciences. After his death his collection, library, and collec- Bruner, Scaramuzza and Otero (1975) provided a tion of paintings suffered some damage and loss. summary catalog of the insects which attack plants His large and important collection of Staphylinidae of economic importance in Cuba. This has been and Pselaphidae from Cuba and other countries was updated with a list by Vàzquez et. al (1992). moved in 1966 to the Field Museum of Natural Fernando de Zayas (1912-1983) is the last of History in Chicago. Co-types of at least some of his the important founders of the study of beetles in Cuban species were exchanged with R. E. Cuba. He worked as an entomologist for the Cuban Blackwelder at the U.S. National Museum, Wash- Academy of Sciences, and traveled and collected ington Additional data are in Genaro and Gutiérrez widely throughout Cuba. He also published exten- (1999). sively on many orders of insects, and his major Philip J. Darlington, Jr. (1904-1983) of the work is the 8 volume series “Entomofauna Cubana” Museum of Comparative Zoology (MCZ), Harvard (seemingly only volumes 3, 6, 7, 8 are published). University, made four collecting trips to Cuba (in Among his many works are a major revision of the 1926, 1929, 1934, 1936), especially for carabid Cerambycidae of Cuba (Zayas 1975) and descrip- 6 Arthropods of Florida and Neighboring Land Areas, Vol. 18

tions of about 200 new beetle species (Zayas 1988). in the , and a bibliography of his pub- These last descriptions were extracted from the very lications, Pp. 17-30. In: G. E. Ball, (ed). Tax- large manuscript for volume 4, on Cuban beetles, onomy, phylogeny and zoogeography of beetles completed in l965, for the Entomofauna Cubana and : A volume dedicated to the memory of series. Regrettably, this still remains unpublished. Philip Jackson Darlington, Jr. (1904-1983). His very large collection is still at his old family Series Entomologica, vol. 33. Dr W. Junk Publ., home in Havana (Ivie 1991). Dordrecht, The . Barbour, T. 1945. A Naturalist in Cuba. Little, Other Workers and Collections Brown and Co., Boston. In addition to the collectors and collections men- Bruner, S. C., Scaramuzza, L. C., and Otero, tioned above there are others of lesser importance A. R. 1975. Catalogo de los insectos que atacan in the study of Cuban beeetles. Additional informa- a las plantas economicas de Cuba. Second edi- tion on past and present locations and conditions of tion, revised and augmented. Institiuto de insect and other zoological collections is in Aguayo Zoologia, Academia de Ciencias de Cuba, La (1950), Alvarez (1958), Chardon (1949), Hernández Havana. (1972) and Silva (1994). Working reference collec- Chardon, C.E. 1949. Los naturalistas en la tions are in various applied research institutes such América Latina. Ciudad Trujillo (Santo as The Forestry Institute in Havana. Domingo), Républica Dominicana. 386 pp. The Manuel Barro collection contains about Chevrolat, L.A.A. 1838. Insectes coléoptères 2,500 insects of which 85% are chrysomelid beetles. inédits, découverts par M. Lanier dans It is held as an historical collection in the National l’intérieur de l’île de Cuba. Rev. Zool., 1838: Museum of Natural History, La Havana. 279-286. The Romanian and Cuban Academies of Sci- Chevrolat, L.A.A. 1858. Descriptions de ences sponsored a joint series of expeditions to in- coléoptères de la partie mèridionale de l’île de vestigate the cave, litter, soil, groundwater and Cuba. Rev. Mag. Zool., ser. 2, 10: 209-212. other invertebrate faunas of Cuba. The expeditions Chevrolat, L.A.A. 1862. Coléoptères de l’île de were from 9 March to 18 June, l969; 3 November Cuba. Notes, synonymies et descriptions to 12 December, 1970; and 3 February to 4 June, d’espéces nouvelles. Familles des cérambycides 1973. The results were published in a series of four et des parandrides. Ann. Soc. Ent. , ser. volumes by the Romanian Academy of Sciences 4, 2: 245-280. (Orghidan et al. 1973, 1977, 1981, 1983), and vol- Chevrolat, L.A.A. 1863. Coléoptères de l’île de umes 1 and 2 provide lists and descriptions of the Cuba. (Suite) Notes, synonymies et descriptions collecting sites. Several papers are on cave and non- d’espéces nouvelles. Familles des cicindélètes, cave collections of beetles. The cave beetles have carabiques, dytiscides, gyrinides et palpicornes. been summarized by Peck et al. (1998). While many Ann. Soc. Ent. France, ser. 4, 3: 183-210. species of cave specialized arthropods occur in Cuba, Chevrolat, L.A.A. 1864a. [? 1865]. Description no known cave-specialized species of beetles occur d’un nouveau genre et diverses espèces there. The beetle specimens from these expeditions d’insectes coléoptères de l’île de Cuba. Rev. Mag. are scattered in the collections of the authors of the Zool., ser. 2, 16: 179-182 papers describing the species or may be in Roma- Chevrolat, L.A.A. 1864b. Coléopteres de l’île de nia, at the “Emile Racovitza” Institute of Speleol- Cuba. (Suite) Notes, synonymies et descriptions ogy in Bucharest. Not all collections from these ex- d’espèces nouvelles. Quatrième mémoire. peditions have been studied and remain in Famille des histériens, phalacrides, Bucharest. nitidulaires, trogositaires, colydiens, rhyzodides, cucujipes, mycètophagides, dermestins, References byrrhiens et chélonarrides. Ann. Soc. Ent. France, ser. 4, 3: 589-620. [March 23, 1864] Aguayo, C.G. 1950. Bosquejo historico de la Chevrolat, L.A.A. 1864c. Coléoptères de l’île de zoología de Cuba. Boletín de Historia de la Cuba. (Suite) Notes, synonymies et descriptions Sociedad “Felipe Poey” 1: 3-31. d’espèces nouvelles. Cinquième mémoire. Alvarez C., J. 1958. Historia de la Zoologia en Famille des parnides, hétérocérides, passalides Cuba. Havana. 369 pp. et lamellicornes (Tribus des coprides, Ball, G. E. 1985. The contributions of Philip J. aphodiides, hybosorides, géotrupides et Darlington, Jr. to the study of carabid beeles A Checklist of the Beetles of Cuba 7

Table 1. Abbreviations used here for the presently rec- Darlington, P.J., Jr. 1937. West Indian Carabidae ognized provinces of Cuba. III: new species and records from Cuba, with a brief discussion of the mountain fauna. Mem. Ca Provincia de Camagüey Soc. Cubana Hist. Natur. ‘Felipe Poey’ 11: 115- CA Provincia de Ciego de Ávila 136. Ci Provincia de Cienfuegos Darlington, P.J., Jr. 1938. Experiments on mim- Gr Provincia de Granma icry in Cuba, with suggestions for future study. Gu Provincia de Guantánamo Trans. R. Ent. Soc. London 87: 681-695. Ha Provincia de La Habana (including Provincia Genaro, J. A. and Gutiérrez, E. 1999. Alexander de Ciudad de la Habana) Bierig: biografía, su colleción entomológica y Ho Provincia de Holguín publicaciones. Cocuyo 9: 27-29. IJ Provincia de Isla de la Juventud (= Isla de Gundlach, J. 1881-1891. Contribucíon a la Pinos, or Isle of Pines) Entomología Cubana. Anales Acad. Cien. Med. LT Provincia de Las Tunas Fis. Nat. (La Havana).[This runs through sev- M Provincia de Matanzas eral years as supplements to the Anales]. Also Or Unknown locality in old “Oriente” province published in book form in three vols., as follows: PR Provincia de Pinar del Rio Vol. I, 1881, Lepidoptera; Vol. II, 1886, Hy- SC Provincia de Santiago de Cuba menoptera, Neuroptera and ; Vol. III, SS Provincia de Sancti Spíritus (in 5 parts) 1891, 404 pp., Coleoptera. (microfilm VC Provincia de Villa Clara copy in MCZ Library, Harvard, University box 2, no. 78-3504) trogides). Ann. Soc. Ent. France, ser. 4, 4: 405- Hernández, O.J. 1972. La zoologia en Cuba (desde 418. 1868 a 1968). Academia de Ciencias de Cuba, Serie Chevrolat, L.A.A. 1865. Coléoptères de l’île de Biologica 44: 1-77. Cuba. (Suite) Notes, synonymies et descriptions Holland, W.J. and Schwarz, E.A. 1917. List of d’espèces nouvelles. Sixième mémoire. Famille the Coleoptera collected on the Isle of Pines by des lamellicornes (tribus des mélolonthides, Gustav A. Link, Sr., 1912-1913. Ann. Carnegie rutélides, dynastides et cétonides, avec les in- Mus. (Pittsburgh) 11: 333-345. dications des tribus, soustribus et groupes du Horn, W. and I, Kahle. 1935-1937. Über genera de Lacordaire). Ann. Soc. Ent. France, entomologische Sammlungen, Entomologen und ser. 4, 5: 21-36. Entomo-Museologie (ein Beitrag zur Geschichte Chevrolat, L.A.A. 1867. Coléoptères de l’île de der Entomologie). Entomologische Beihefte, Band Cuba. (Suite) Notes, synonymies et descriptions 2 (1935), 3 (1936), 4 (l937). Berlon-Dahlem, Ber- d’espèces nouvelles. Septième mémoire. Famille lin. des buprestides, throscides, eucnémides et Howard, L.O., Barber, H S., and Busck, A. 1928. élaterides. Ann. Soc. Ent. France, ser. 4, 7: 571- Dr. E. A. Schwarz. Proc. Ent. Soc. Washington 616. 30: 152-183. Chevrolat, L.A.A. 1870. Coléoptères de l’île de Ivie, M.A. 1991. Taxonomic notes on a little known Cuba. (Suite) Notes, synonymies et descriptions publication: Zayas, 1988, Entomofauna Cubana. d’espèces nouvelles. Huitième mémoire. Famille Orden Coleoptera. Coleop. Bull. 45: 399-401. des dascyllides et malacodermes. Ann. Soc. Ent. Jacquelin du Val, P.N.C. 1856-1857. Insectes. France, ser. 4, 10: 67-78. Ordre des coléoptères, Lin. In: Sagra, Ramon de Chevrolat, L.A.A. and Fauvel, A. 1863. la (ed.). 1856-1857. Histoire physique, politique Coléoptères de l’île de Cuba. (Suite) Notes, syn- et naturelle de l’île de Cuba. vol. 7. Animaux onymies et descriptions d’espèces nouvelles. articules, insectes, Coleoptera. pp. 1-136 published Trosième mémoire. Famille des staphyliniens. in 1856; pp. 137-328 published in 1857. Paris Ann. Soc. Ent. France, ser. 4, 3: 427-446. (French edition, in Special Collections, MCZ Li- Darlington, P.J., Jr. 1934. New West Indian brary, Harvard University; Spanish edition pub- Carabidae, with a list of the Cuban species. lished in Madrid, 20 September, 1857, pp. 1-136, Psyche 41: 66-131. in a larger page size and with a different pagina- tion from the French edition). Norman, J.R. 1938. Notes on the dates, etc. of the ‘Memorias’, ‘Repertorio’, ‘Synopsis’, and 8 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Table 2. A summary list of important classical collecting sites which are not easily found on most presently available maps. These are placed into present provinces with the use of the Atlas de Cuba (1978).

Aguadores. SC; a Darlington coastal locality east of the mouth of the bay at Santiago de Cuba. Almendares. Ha; a riverside locality of Bierig in Havana. Baragua. CV: 20 km southeast of city of Ciego de Avila. Boniato. SC; town north of Santiago de Cuba; contains Rio Frio and the Boniato Mountians to the north. Buenos Aires. Probably SS; an unlocated Darlington and Zayas collecting site in the Trinidad Mts., about 800 m elev., maybe near Topes de Collantes. Caimito. Ha; a village in Sierra de Anafe, also called Caimito del Guayabal. A Bierig and Zayas locality. Canimar. M; a Gundlach locality on the Canimar river. Cárdenas. M; the most important Gundlach locality. Cauto el Cristo. Gr; on the Cauto River; a Darlington locality. Cayamas. Ci; 40 km NW of Cienfuegos; the most important Schwarz locality; N 22º 15’; W 80º 48’. Central Baraguá. see Baraguá. Central Fermina. see Fermina. Central Jaronú. CA; now the town of Jaronú. Cienaga de Zapata. M; a large partly flooded swampy , with saw-grass, pinelands, and hardwood- hammock forest, similar to the region of the Everglades of southern Florida. Daiquiri. SC; a playa (beach) below Gran Piedra; upstream is Vinent, another Darlington locality. Fermina. M; a sugar mill (central) 10 km E of Jovellanos, south of Cardenas, a Gundlach locality. Gran Piedra. SC; a mountain National Park, east of Santiago de Cuba, approached through the town of Caney. Hanabanilla Falls. VC; up Hanabanilla. River, N slope Trinidad Mts.; a dam and lake are now at the site. Loma del Gato (= Cobre Peak). SC; in Sierra del Cobre (3690' = 1097 m), the first mountain range west of Santiago de Cuba; Loma de San Juan is at the other end of the range. Mina Carlota. Probably SS. an unlocated Zayas collecting site in the Trinidad Mountains. Peninsula de Zapata; see Cienaga de Zapata. Pico Turquino. SC; the highest peak in Cuba, in the Sierra Maestra. Ascents are from the south so collections were made in Santiago de Cuba Province, rather that Granma Province on the north slope. Rangel (Sierra de Rangel). PR; a village in the Sierra del Rosario, NW of San Cristobal; an important Bierig locality and a Darlington locality; with a nearby peak at 655 m. San Blas. Ci; on the west slope of the Trinidad Mts, northwest of Trinidad City; also a Zayas locality and village in Matanzas Province at the SE edge of the Cienaga Zapata. Santa Clara. VC; an important Zayas locality. Sierra del Cobre. SC; See Loma del Gato above. sn important Zayas locality. Sierra Cristal. Ho; a national park, and an important Zayas locality. Soledad. Ci; the Atkins (or Harvard) Botanical Garden which is now a tourist locality; in Arimao River valley; Darlington locality; east of Cienfuegos. At about N 22º 5', W 80º 18'. Topes de Collantes. SS; a Zayas locality at about 800 m elevation in the Trinidad Mountains. Trinidad. usually referring to the mountains north and west of the City of Trinidad. The mountains are also called the Sierra Escambry, and are in Cienfuegos, Santa Clara and Sancti Spíritus Provinces. Most collecting sites were probably in Sancti Spritus Province. Zapata Peninsula; see Cienaga de Zapata. ‘Enumeratio’ of Filipe Poey. J. Soc. Bibliog. Nat. Papavero, N. 1971. Essays on the history of Neo- Hist. 1: 135-137. tropical Dipterology. Vol. 1. Museu de Zoologia, Orghidan, T., Núñez Jiménez, A, Botosaneanu, Universidade de São Paulo, Brazil. L., Decou ,V., , St., and Viña Bayes, Peck, S.B., Ruiz-Baliú, A.E., and Garcés N. 1973-1981. Résultats des expéditions Gonzaléz, G.F. 1998. The cave-inhabiting biospéologiquess Cubano-Roumaines à Cuba. beetles of Cuba (Insecta: Coleoptera): diversity, Editura Academiei Republicii Socialiste România, distribution, and ecology. J. Cave Karst Studies Bucuresti. Vol. 1, 1973; vol. 2, 1977; vol. 3, 1981. 60: 156-166. A Checklist of the Beetles of Cuba 9

Poey, F. 1851-l855. Ch. 25. Conspectus familiarum Schwarz, E.A. 1905. A letter from Cuba. Proc. Ent. Coleopterorum, aspecto de los familias de los Soc. Washington 5: 286-289. insectos Coleopteros. Memorias sobre la historia Schwarz, E.A. 1929. “Letters from Cuba”. J. New natural de la Isla de Cuba. La Havana, vol. 1; pp. York Ent. Soc. 37: 326-359. 302-337 (seen). Reprinted l975, Antiquariaat Silva Taboada, G. 1994. Zoological collections and Junk, Lochem, Netherlands. collecting in Cuba during the Twentieth Century. Ragués, P. V. 1914. Catalogo general, Museo Cubana Curata 37: 90-98. “Gundlach,” Zoología. Instituto de Segunda Vàzquez, L.L., Valdés, E., and Amor, J.C. 1992. Enseñanza de la Havana. Inprenta “Cuba Intel- New manifestations of pests on economically im- lectual,” La Havana. portant plants during the period from 1970 to Ramsden, C.T. 1915. Juan Gundlach. Ent. News 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo 26: 241-260. Silvestri 49: 41-52. Reyes, E. and Hernández, L.M. 1995. Tipos de Zayas, F. de. 1975. Revision de la familia insecta depositados en la coleción del Instituto de Cerambycidae (Coleoptera, Phytophagoidea). Ecología y Sistemática. Cocuyo 3: 2-3. Instituto de Zoologia, Academia de Ciencias, La Reyes, E.E., M. Hidalgo-Gato, A.D. Alvarez. 2002. Havana, Cuba. 443 pp. Juan Gundlach y su colección entomológica. Zayas, F. de. 1988. Entomofauna Cubana. Orden Cocuyo 12: 20-21. Coleoptera. Separata. Descripcion de nuevas Sagra, Ramon de la. 1856-1857. Histoire physique, especies. Editorial Cientifica-Technica. La Ha- politique et naturelle de l’île de Cuba. Paris. vana. 212 pp.

Construction of a checklist of Cuban beetles

This checklist is a summary of present knowl- (1914, 1917). Bruner (1951) contributed Cuban edge of the diversity, habits, and distributions of records missed by Blackwelder. the Cuban beetle fauna. It is designed to be a com- The beetle species described by the Cuban en- panion volume to the distributional checklist of the tomologist Fernando de Zayas (1988) are included beetles of Florida (Peck and Thomas 1998). Much here, as well as the suggested synonomies of Ivie work remains to be done to completely establish (1991). Other general works on beetles of Cuba or the identity, validity, and accuracy of presently used the West Indies are few. An exception are the chap- names of Cuban beetles. Additionally, much more ters on aquatic beetle families by Spangler (1982). field and laboratory study is required to document Although these are a valuable entry to the litera- the unreported or undescribed species, and their ture, they contain little direct or new information distributions and bionomics in Cuba. about Cuban beetles. The Zoological Record has been searched from 1950-2003. I have also searched Bio- Methods and Materials logical Abstracts on CD-ROM (from 1989-1995), The body of this book is a species checklist, Cambridge Scientific Abstracts (1994 to 2000), with summarized information on the distribution AGRICOLA (from 1969 to 2000), and Entomology and bionomics of the beetles of Cuba. The list was Abstracts (vol. 1 to 31, 2000). The primary key constructed from two kinds of primary sources. The words in the searches were “Cuba” and “Coleoptera.” first was modern taxonomic revisions or summa- After I prepared preliminary lists, I then sought ries of the beetle fauna of Cuba or the West Indies, specialists in Cuba or elsewhere who could check or adjacent land masses. These are mentioned their speciality families for accuracy and complete- where appropriate for each family unit, and are ness. The contribution of each specialist is acknowl- cited in full at the end of each family unit. If no edged for each family or section they reviewed. modern literature is known to me, a preliminary When possible, data have been added from col- list was constructed from Blackwelder’s (1944, 1957) lections in Cuba, and some in such monumental checklist of the Beetles of Latin as the Field Museum (Chicago), and the Museum America and the update of this by Arnett (1983). of Comparative Zoology (Harvard University). The Blackwelder’s checklist was partly formed by build- list is intended to be as complete as I have been ing upon the earlier lists by Leng and Mutchler able to make it with the time and resources avail- able. It is constructed so that it can serve the needs 10 Arthropods of Florida and Neighboring Land Areas, Vol. 18 of a multiplicity of users; from agronomists, to ecolo- Catalogs. I do not cite catalogs or lists of names gists, land-use managers, taxonomists, quarantine if no new taxonomic, distributional, or bionomic entomologists, ecologists and biogeographers. Us- information is given in them. I do not give cita- ers can insert their own new data as they come tions to the older primary literature which is in- upon it. cluded in Blackwelder (1944, 1957; reprinted in 1982). The literature cited in Blackwelder is from Organization 1944 and earlier. To include all these older refer- The core of the work is a list of valid species ences here would greatly increase the length of this names, synonyms, and the more recent references listing. about these species in the scientific literature. The References. It is my intention to give refer- following explains the selection and organization of ences for “younger” literature only; i.e., that pub- the included data. lished since Blackwelder (1944, 1957) which has Higher classification. The checklist gener- contributed data on Cuban species. I have also tried ally follows the sequence and phylogenetic arrange- to include references to recent faunal summaries ment of the world list of beetle families, subfami- or revisions which may not contain direct informa- lies, and tribes of Lawrence and Newton (1995 and tion on Cuban species but which may contain keys, at http:www. coleopsoc.org/). Arnett and Thomas descriptions, or illustrations or other information (2000), and Arnett et al. (2002). Data on distribu- on species in Cuba or which may be expected in tion and habits of beetles outside of Cuba are gen- Cuba. Primary older literature not cited in erally from Lawrence (1982). The families are num- Blackwelder can be found in appropriate issues of bered so that finding them in sequence is easier. “Zoological Record” or in the several catalogues Numbering from the world list is used because sev- cited. The relevant family chapters in Arnett and eral families not now known from Cuba may be Thomas (2000) and Arnett et al. (2002) are valu- expected to be found in the future. able references for all beetle families in Cuba, and Included species. The list contains all spe- are usually not cited for each family. cies of beetles which are known or reported to oc- cur in Cuba. If the known distribtion of a species Types. Type status and depository are given (if suggests that it very probably occurs in Cuba, but known). Older species usually do not have desig- there are seemingly no records for it, its name and nated types. Co-types or syntypes from which lec- references are enclosed in parentheses ( ). Some totypes may be designated may be in collections in lists or catalogues only vaguely give distributions Havana (the Gundlach collection), or in Paris as “West Indies” or “Antilles.” If the total distribu- (Chevrolat and Jaquelin du Val), or elsewhere (see tion suggests that this includes Cuba I have con- table 3). Jacquelin du Val published new species servatively included the species. The function of names from specimens in both the Chevrolat and such additions is to alert future workers to species Guérin-Méneville collections, so those will probably which are named but not yet explicitly recorded have appropriate type material. Horn and Kahle from Cuba. Such citations could undoubtedly be (1935-1937) may be of help in locating historically far more numerous than those I have included. If important collections and specimens. the species has been recorded in the past from Cuba, but the application of the name is now known to be Type localities. I list published type localities (t.l.) an error, the name and references are enclosed in only for endemic species. In many older works, no brackets [ ]. I ask that users of the catalogue bring type localities were specified. In these cases “Cuba” errors of omission or inclusion to my attention. can be seen to serve as a type area. If this is the Citations. The author, year of publication, and case, I usually do not cite “Cuba.” page of publication are given (when known) for each valid species. These data will help to locate the origi- Distributions. Distributions of the species are nal references. When a species was originally placed presented in two ways. in a different than now, I give this original (A). They are first summarized as Cuba and genus name when it is known to me. then as a list of the known major islands or coun- Synonomies. I list synonym names below the tries outside of Cuba from where the species is valid name of the species. I do not give complete known. Distribution within the USA is by abbre- lists of synonyms for the species in this checklist. viations of states at the edges of the range in the My intention is to give synonomies only for names USA. “Endemic” is used as a term to indicate that which have been applied to Cuban records. the species is naturally limited to Cuba. “Intro- A Checklist of the Beetles of Cuba 11

Table 3. Locations, available codens, addresses, and annotations for major collections known to have identified Cuban beetles. Codens for collections follow usage of Arnett et al. (1997).

Amsterdam. ZMAN. Instituut voor Taxonomische Zoologie, Zoologisch Museum, Universiteit van Amsterdam, Plantage Middenlaan 64, 1018 DH, Amsterdam, The Netherlands. Aylmer. CMNC. Entomology Collections, Canadian Museum of Nature, Aylmer, Quebec, (P.O. Box 3443, Station D, Ottawa ON K1P 6P4), Canada. Basel. NHMB. Entomology Department, Naturhistorisches Museum, Augustinergasse 2, 4001, Basel, Switzer- land. Especially containing the large collection of G. Frey and W. Wittmer. Berkeley. EMEC. Essig Museum of Entomology, Department of Integrative Biology, University of California, Berkeley, California, 94720. USA. Berlin. ZMHB. Museum für Naturkunde der Humbolt Universitat zu Berlin, Bereich Zoologisches Museum, Invalidenstrasse 43, 1040, Berlin, Germany. Bozeman. MTEC. Montana State University Entomology Collection, Entomology Research Laboratory, Depart- ment of Biology, Montana State University, Bozeman, Montana, 59715, USA. Containing the important collection of West Indian insects of M. A. Ivie. Bruner. S.C. Bruner collection, from the Santiago de las Vegas Agriculture experiment station (EEA), supposedly now in Havana at the Institute of Ecology and Systematics (IES). Brno. MMBC. Department of Entomology, Moravské Museum, Preslova ul. 659 37, Brno, Czech Republic. Brussels. ISNB. Institut Royal des Sciences Naturelles de Belgique, 29 rue Vautier, B1040, Brussels, Belgium. Bucarest. Institute of Speologie “Emile Racovitza”, Str. Mikail Moxa 9, Bucarest 12, Romania. This may contain material from the Cuban-Roumanian cave biology expeditions. Cambridge. MCZC. Entomology Department, Museum of Comparative Zoology, Harvard University, 22 Oxford St., Cambridge, Massachusetts, 02138, USA. A collection strong in Cuban material from field work of P. J. Darlington, Jr, and others. Champaign. INHS. Illinois Natural History Survey Insect Collection, 607 E. Peabody Dr., Champaign, Illinois 61820, USA. With Cuba material collected by M. W. Sanderson. Chicago. FMNH. Insect Collection, Division of Insects, Field Museum of Natural History, Roosevelt Road and Lake Shore Drive, Chicago, Illinois, 60605, USA. Containing the important Bierig collection of Staphylinoidea. Columbus. OSUC. Ohio State University Collection of Insects and Spiders, 1735 Neil Avenue, Columbus, Ohio, 43210, USA. Eberswalde. DEIC. Deutsche Entomologische Institut, Eberswalde Finow 1, 1300, Germany. Franz. H. Franz collection, Mödling, Austria. Now placed in the Vienna Museum, Austria. Geneva. MHNG. Department des Insectes, Museum d’Histoire Naturelle, CP 434, CH-1211, Geneva 6, Switzer- land. Gundlach. J. Gundlach collection. Now a part of the collections of the Institute of Ecology and Systematics (IES) in Havana. The Gundlach code number relates the specimen to the name and locality as published in volume 3 of this “Contribucion a la Entomologia Cubana.” Garrido. Orlando Garrido, Havana, Cuba. A private collection specializing in Tenebrionidae. Gainesville. FSCA. Florida State Collection of Arthropods, Division of Plant Industry, 1911 34th St. SW, PO Box 147100, Gainesville, Florida, 32614, USA. The Strohecker collection of Endomychidae and other large collec- tions from throughout the Caribbean are here. This institution also houses the important and large private collection of scarabs and other families of beetles of R. E. Woodruff Havana. IZAC. Collections of the Institute of Ecology and Systematics (IES) of the Academia de Ciencias de Cuba, at Carretera de Varona km 3 ½, Capdevila, Boyeros, La Habana, Cuba; containing the historically important Gundlach collection, and the former collection of the Instituto de Zoologia de la Academia de Ciencias de Cuba (IZAC), Havana. It may also contain some or all of (1) the former collections of the Estacion Experimental de Agronomia (EEA), once located at Santiago de las Vegas, near Havana, (2) the reference collections of the Cuban Sugar Club (CSC), (3) of the Department of Speleology, Academy of Sciences, with some of the specimens from the Cuban-Roumanian cave-biology expeditions, and (4) the tenebrionid collec- tion of O. Garrido. Part or all of these collections may move to the National Museum of Natural History, located in downtown Old Havana (Habana Vieja). Halle. MLUH. Wissenschaftsbereich Zoologie, Sektion Biowissenschaften Martin-Luther Universitat Halle, WB Zoologie, Domplatz 4, 4020 Halle (Salle), Germany. Possibly contains Cuban species in the large and impor- tant collections of Burmeister, Germar, and especially that of Suffrian. Helsinki. MZHF. Zoological Museum, Finnish Museum of Natural History, Universitets Helsinki, P. Rautienkatu 13, SF-00100, Helsinki 10, Finland . Howden. HAHC. Henry and Anne Howden Collection, Ottawa, Ontario, Canada. Placed in the Entomology Divi- sion, Canadian Museum of Nature, Aylmer, Quebec, (PO Box 3443, Station D, Ottawa, ON K1P 6P4), Canada. 12 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Table 3. Continued.

Lafayette. PURC. Entomology Research Collection, Department of Entomology, Perdue University, West Lafayette, Indiana, 47907, USA. Of special importance because of the W.A. Blatchley collection. Leiden. RMNH. Nationaal Natuurhistorische Museum, Raamsteeg 2, Leiden, 2311 P, The Netherlands. London. BMNH. Department of Entomology, The Natural History Museum (formerly the British Museum (Natu- ral History)), London, SW7 5BD,. Medvedev. E.L.Medvedev collection (Chrysomelidae), Moscow, Russia. Miami. University of Miami, Miami, Florida, USA. Possibly contains parts of the Stroehecker collection not at Gainesville. Munich. ZSMC. Zoologische Staatssamslungen, Munchhausenstrasse 21, D-8000 München 60, Bayern, Ger- many. New York. AMNH. Department of Entomology Collection, American Museum of Natural History, Central Park West at 79th St., New York City, NY, 10024, USA. O’Brien. CWOB. Charles W. O’Brien collection, of Tallahassee, FL; to be deposited in the collections of the California Academy of Sciences, San Francisco. See San Francisco below. Ottawa. CNCI. Canadian National Collection of Insects, ECORC, Biological Research Division, Central Experi- mental Farm, Agriculture Canada, Ottawa, Ontario, K1A OC6, Canada. Paris. MNHN. National Collection of Insects, Museum National d’Histoire Naturelle, 45 rue Buffon, Paris 75005, France. Probably contains Cuban specimens in the important collections of Chevrolat and Jacquelin Du Val, and possibly others. Philadelphia. ASNP. Department of Entomology, Academy of Natural Sciences, 19th and the Parkway, Philadel- phia, PA, 19103, USA. Poey exchange collection containing possible Chevrolat co-types Pittsburgh. CMHN. Section of Insects and Spiders, Carnegie Museum of Natural History, 900 Forbes Ave., Pittsburgh, Pennsylvania, 15213, USA. Prague. NMPC. Department of Entomology, National Museum (Natural History), 148 00 Praha 4, Kunratice 1, Czech Republic. Ramsden. Charles Ramsden collection, Museo de Carlos Ramsden de la Torre, Universidad de Oriente, Santiago de Cuba, Cuba. Ratcliffe. BCRC. Brett C. Ratcliffe collection, Systematic Research Collections, W436 Nebraska Hall,University of Nebraska, Lincoln, Nebraska, 68588-0514, USA Sacramento. CDAE. California State Collection of Arthropods, Analysis and Identification Unit, California Department of Food and Agriculture, 1220 N St., Rm. 340, Sacramento, California, 95814, USA. San Francisco. CASC. Department of Entomology, California Academy of Sciences, Golden Gate Park, San Fran- cisco, California, 94118, USA. Sao Paulo. MZSP. Museu de Zoologia da Universidade de São Paulo, Biblioteca 7172, 01.051 São Paulo, SP, Brazil. Thérond. J. Thérond collection, Nimes, France. Histeridae. Verona. MSNV. Museo Civico di Storia Naturale, Lungadige Porta Vittoria 9, Verona, I-37129, Italy. Washington. USNM. United States National Entomological Collection, Department of Entomology, U.S. Na- tional Museum of Natural History, (formerly the United States National Museum), Washington, DC 20560, USA. Holding the important collections of E. A. Schwarz made around Cayamas, Cuba, and the many types resulting from this. Warsaw. ZMPA. Museum of the Institute of Zoology, Polish Academy of Science, Wilcza 64, 00-679, Warsaw, Poland. Zayas. Fernando de Zayas collection, La Habana, Cuba. A private collection, not presently available for study. duced” is used as a term if the species occurs in Sometimes, only the imprecise words “Antilles, Cuba because it was probably brought there Caribbean or West Indies” have been given in ear- through either accidental or intentional human lier literature. activity. Many more species are probably introduced (B). Secondly, data are given on species distri- than are indicated here. I have indicated a species butions within Cuba. If a species is common and as introduced only when a publication has explic- known from about 10 or more published localities, itly stated this. I have checked that the taxon and I usually indicate this with the term “widespread” distribution data used here agree with that in the and give no additional data. Province abbreviations summary list of the beetles of Florida (Peck and only are given in some cases. Some catalogues (e. Thomas 1998). Where distributional details are g., Blackwelder 1957, Arnett 1983, O’Brien and lacking, I may indicate this with a question mark. Wibmer 1982) or taxonomic papers indicate a spe- A Checklist of the Beetles of Cuba 13 cies presence in Cuba without listing specific lo- references on these topics is given at the end of calities or collections with voucher specimens. In each family. Data on insects of agricultural impor- such cases I cannot give these details. tance or status as a pest in other ways in Cuba are Table 1 is a list of abbreviations for the pres- often drawn from Bruner et al. (1975), which is ently recognized Provinces of Cuba. The First Con- usually not specifically cited. Additions to this list gress of the Communist Party of Cuba in Decem- are in Vàzquez et al. (1992). I have cited host plant ber 1975 increased the number of provinces from names as reported, either as common name or ge- the former number of six to the present fourteen nus and/or species. Family names for genera and (figure 1). The six pre-1975 provinces were Pinar genera for English common names may be found del Rio, Havana, Matanzas, Las Villas, Camagüey, in a botanical reference such as Heywood (1978) or and Oriente. For example, many older published Willis (1980). records are from the former Province of Oriente, Collections. Locations of voucher specimens an area which is now composed of the provinces of documenting the distribution records are usually Santiago de Cuba, Granma, Guantánamo, and indicated by the name of the city or collection which Holguín. Thus, older literature or insect labels may contains the material (not by the now common prac- use old province names which may not correspond tice of an abbreviation or coden). Table 3 is a list of to the present provinces. I have attempted to place these cities or collections as published or known to the old locality records into the present provinces. us or reviewing specialists which hold identified When I have not been able to place a locality from Cuban material. While there is a growing trend to the former Oriente Province into its present prov- use codens for identifying insect collections, I find ince I have left it as “Or.” these to be often confusing, not easily memorized, and not “user-friendly,” especially to non-taxono- Localities. Within a province, Cuban usage also mists and insect workers outside of Canada and can place localities within a municipality, which the USA. I think my use of cities or collections to roughly corresponds to a county as used in the be more readily and immediately recognizable. Even United States. I list localities within a province from if no repositories of specimens are known to me I larger (municipalities) down to smaller (town) geo- insert the heading and leave space in which notes graphic units, and separate them with a comma. can be added by users. Separate localities within a province are separated by a semicolon. Generally more than 10 localities Acknowledgements are simply summarized as “widespread.” A frequently used Cuban word that can con- I am indebted to the many people who have fuse non- is “Central.” Labels may abbrevi- given of their time and knowledge to help complete ate this as “Cent.” The term does not mean “in the this checklist. The specialists who reviewed family middle of.” Rather, in Cuba, a “Central” is a sugar or other sections of early drafts are thanked in those mill or sugar refinery which was used as a place sections. Joyce Cook proofread the entire manu- name for a collecting locality. Many of these are script. Curators of the many collections holding indicated on the most recent Atlas of Cuba (1978), Cuban beetles have allowed access to the collections but some names have been changed and the sites under their care. Colleagues in Cuba, especially are no longer identifiable. Table 2 is a list of rela- Julio Antonio Genaro at the Cuban National Mu- tively obscure but important localities which may seum of Natural History, Havana, were generous not be easily located by users outside of Cuba. Even in sharing their knowledge of their beetle fauna. if no precise localities are known to me, I insert the My field, museum and library research were sup- heading “Cuba localities” and leave a blank space ported by operating grants from the Natural Sci- so that notes or records can be added by users. ences and Engineering Research Council of Canada. Published localities which I have been unable to locate are indicated with “?” References Bionomics. After information on the distri- bution of a species within Cuba, I present notes on Arnett, R.H., Jr. 1983. Checklist of the beetles of habitat, hosts, field biology, pest status, and refer- North and Central America and the West ences to the natural history or ecology and other Indies. Florida and Fauna Publs., Gainesville, important data when known. I use the term bion- Fla. Vols. 1-10, 2173 pp., actually published in omics to be roughly equivalent to a summary of parts between 1977 and 1984. the natural history of the species. Full citation for 14 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Arnett, R. H., Jr., Samuelson, G.A., and Lawrence, J.F. 1982. Coleoptera. pp. 482-533. In: Nishida, G.M. 1997. The insect and spider S. Parker (ed.). Synopsis and Classification of collections of the world. 2nd ed. Flora and Fauna Living Organisms, vol. 2. McGraw Hill, N.Y. Handbook no. 11. CRC Press, Boca Raton, FL. Lawrence, J.F. and Newton, A.F. 1995. Fami- 310 pp. lies and subfamilies of Coleoptera (with selected Arnett, R. H., Jr. and Thomas, M. C. 2000. genera, notes, references and data on family American Beetles: Archostemata, , group names), pp. 779-1006. In J. Pakaluk and Adephaga, Polyphaga: Staphyliniformia. Vol. S.A. Slipinski (eds.). Biology, phylogeny, and 1. CRC Press, Boca Raton, FL. 443 pp. classification of Coleoptera: papers celebrating Arnett, R. H., Jr., Thomas, M. C., Skelley, P. the 80th birthday of Roy A. Crowson. E., and Frank, J. H. 2002. American beetles, Leng, C.W., and Mutchler, A.J. 1914. A prelimi- Polyphaga: Scarabaeoidea through Curcul- nary list of the Coleoptera of the West Indies as ionidea. Vol. 2. CRC Press, Boca Raton, FL. recorded to January l, 1914. Bull. American 861 pp. Mus. Nat. Hist. 33: 391-493. Atlas de Cuba. 1978. Instituto de Geodesia y Leng, C.W. and Mutchler, A.J. 1917. Supple- Cartagrafia, La Havana, 143 pp. ment to preliminary list of the Coleoptera of Blackwelder, R.E. 1944. Checklist of the co- the West Indies. Bull. American Mus. Nat. leopterous insects of México, Central America, Hist. 37: 191-220. the West Indies, and South America. U.S. Natl. O’Brien, C.W., and Wibmer, G.J. 1982. Anno- Mus. Bull. 185(1-5) I-XII, 1-925. Reprinted in tated checklist of the (Curculionidae 1982. sensu lato) of Nortlh America, Central America, Blackwelder, R.E. 1957. Checklist of the co- and the West Indies (Coleoptera: leopterous insects of México, Central America, Curculionidae). Mem. Amer. Ent. Inst. 34. 382 the West Indies, and South America. U.S. Natl. pp. Mus. Bull. 185(6): I-IV + 927-1492. Reprinted Peck, S.B. and Thomas, M.C. 1998. A distribu- 1982. tional checklist of the beetles (Coleoptera) of Bruner, S.C. 1951. Algunas adiciones a la lista de Florida. Arthropods of Florida and neighbor- Coleopteros de Cuba. Mem. Soc. Cubana Hist. ing land areas, vol. 16. 180pp. Natur. 20: 81-88. Spangler, P.J. 1982. Coleoptera, pp. 328-397. In: Bruner, S.C., Scaramuzza, L.C., and Otero, Hurlbert, S.H., and Villalobos-Figueroa, A. A.R. 1975. Catalogo de los insectos que atacan (eds.). Aquatic biota of Mexico, Central America a las plantas economicas de Cuba. Second edi- and the West Indies. San Diego State Univer- tion, revised and augmented. Institiuto de sity, San Diego, CA. Zoologia, Academia de Ciencias de Cuba, La Vàzquez, L.L., Valdés, E. and Amor, J.C. 1992. Havana. New manifestations of pests on economically Heywood, V.H. (ed.). 1978. Flowering plants of important plants during the period from 1970 the world. Mayflower Books, New York. to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo Horn, W. and Kahle, I. 1935-1937. Über Silvestri 49: 41-52. entomologische Sammlungen, Entomologen Willis, J.C. 1980. A dictionary of the flowering und Entomo-Museologie (ein Beitrag zur plants and ferns. Eighth ed. (1973), revised by Geschichte der Entomologie). Entomologische H.K. Airy Shaw. Cambridge University Press, Beihefte, Band 2 (1935), 3 (1936), 4 (l937). New York. Berlon-Dahlem, Berlin. Zayas, F. de. 1988. Entomofauna Cubana. Orden Ivie, M.A. 1991. Taxonomic notes on a little known Coleoptera. Separata. Descripcion de nuevas publication: Zayas, 1988, Entomofauna especies. Editorial Cientifica-Technica. La Ha- Cubana. Orden Coleoptera. Coleop. Bull. 45: 399- vana. 212 pp. 401. A Checklist of the Beetles of Cuba 15

Table 4. Families, genera, and species numbers of beetles known to occur or probably occurring in Cuba. The total number of genera and species known for the family in Cuba is given: followed by the number which are endemic (limited to Cuba), native (naturally occurring in Cuba and other countries), and introduced (whose presence in Cuba is because of accidental or intentional transport by human activities). Predicted species not yet documented for Cuba but are expected because of their distribution. Erroneous species are species names recorded in the past for Cuba but now known to be in error. Predicted and erroneous species are not included in the species totals. It is of interest to note the number of families known in Cuba from only one or a few genera and one or a few species. This is an indicator of the disharmonic nature of the island’s fauna.

Total Total End. Native Intro Predict. Erroneous ORDER COLEOPTERA Genera Species Species Species Species Species Species ______

Suborder Archostemata 4. Micromalthidae 1101000 Suborder Adephaga 9. Rhysodidae 2550000 10. Carabidae 69 186 73 111 2 0 4 11. Gyrinidae 3624000 12. Haliplidae 2752000 14. Noteridae 6 10 19000 17. Dytiscidae 18 47 9 38 0 1 2 Suborder Polyphaga Staphyliniformia Hydrophiloidea 18. Hydrophilidae 16 46 7 39 0 1 0 21. Histeridae 16 38 10 28 0 1 1 Staphylinoidea 22. Hydraenidae 3422001 23. Ptiliidae 1110000 25. Leiodidae 5660000 26. Scydmaenidae 3 12 12 0000 28. Staphylinidae 114 282 109 162 11 14 2 Scarabaeiformia Scarabaeoidea 31. Passalidae 1211006 33. Trogidae 2312000 35. Geotrupidae 1211000 37. Ochodaeidae 1101000 38. Hybosoridae 2201100 39. Ceratocanthidae 1440000 41. Scarabaeidae 26 156 117 32 7 0 0 Elateriformia Scirtoidea 47. Scirtidae 3990000 Buprestoidea 51. Buprestidae 25 57 36 17 4 1 2 Byrrhoidea 53. Elmidae 4523000 54. Dryopidae 1101000 55. Lutrochidae 1101000 56. Limnichidae 4413000 57. Heteroceridae 4523000 58. Psephenidae 1110000 60. Ptilodactylidae 1770000 61. Chelonariidae 1651000 Elateroidea 67. Eucnemidae 8 16 10 6000 69. Elateridae 25 68 34 34 0 2 1 16 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Table 4. Continued.

Total Total End. Native Intro Predict. Erroneous ORDER COLEOPTERA Genera Species Species Species Species Species Species ______

73. Lycidae 6 11 92000 76. Lampyridae 8 37 29 8000 78. Cantharidae 12 19 19 0000 Bostrichiformia Bostrichoidea 81. Nosodendridae 1110000 82. Dermestidae 5817070 83. Bostrichidae 13 18 0 14 4 3 0 84. Anobiidae 17 23 12 10 1 3 0 Cucujiformia Lymexyloidea 85. Lymexylidae 1101000 Cleroidea 87. Trogossitidae 5 10 36160 89. Cleridae 11 20 17 0321 93. Melyridae 5871000 Cucujoidea 95. Sphindidae 2220000 97. Nitidulidae 15 32 1 28 3 3 0 98. Smicripidae 1101000 99. Monotomidae 2431000 103. Silvanidae 6 12 37260 104. Passandridae 2202010 106. Laemophloeidae 6725030 108. Phalacridae 7 10 73000 111. Cryptophagidae 1101000 113. Languriidae 3413000 114. Erotylidae 2651001 116. Biphyllidae 1110000 117. Bothrideridae 3404000 118. Cerylonidae 5624000 120. Discolomatidae 1220000 121. Endomychidae 1101000 122. Coccinellidae 31 43 19 22 2 0 1 123. Corylophidae 2431000 124. Latridiidae 5633010 Tenebrionoidea 125. Mycetophagidae 3303010 128. Ciidae 4 20 7 13 0 1 0 130. Melandryidae 1110000 131. Mordellidae 3 10 82000 132. Rhipiphoridae 2504110 133. Colydiidae 13 17 3 14 0 1 0 134. Monommatidae 2202000 135. Zopheridae 2312000 140. Tenebrionidae 48 174 104 63 7 4 0 143. Oedemeridae 5835020 145. Meloidae 4734000 146. Mycteridae 3532000 151. Salpingidae 1101000 152. Anthicidae 8 20 4 13 3 2 1 153. Aderidae 2202000 154. Scraptiidae 2220000 A Checklist of the Beetles of Cuba 17

Table 4. Continued.

Total Total End. Native Intro Predict. Erroneous ORDER COLEOPTERA Genera Species Species Species Species Species Species ______

Chrysomeloidea 155. Cerambycidae 42 230 144 80 6 9 9 156. Bruchidae 13 36 8 24 4 0 0 159. Chrysomelidae 85 374 306 68 0 9 6 Curculionoidea 161. Anthribidae 12 29 23 5120 162. Belidae 1110000 163. Attelabidae 4 13 67000 164. Brentidae 9 20 14 5110 167. Curculionidae 134 385 232 131 22 12 0 ______Totals 954 2673 1495 1092 86 98 38

ORDER STREPSIPTERA

Corioxenidae 1101010 Myrmecolacidae 1101000 Halictophagidae 1101000

______Totals 3103010

Summary and origins of the beetle fauna of Cuba

Summary of diversity harmonic or unbalanced fauna. It is a result of the The Cuban beetle fauna is now known to con- long oceanic isolation of Cuba as an island. If there tain 87 families, 954 genera, and 2,673 species had been free and obstructed movement of faunas (Table 4). This diversity can be compared with the between all these and Cuba it would be ex- known beetle fauna of other areas, such as the pected that the higher-level taxonomic makeup of United States with 131 families; Canada with 129 the beetles of Cuba would be similar to that of the families, 3526 genera, and 25,160 species; Mexico neighboring continental lands. with 114 families; Brazil with 104 families, 4,351 The actual dissimilarity is strong evidence that genera, and 26,755 species; and with 113 Cuba has not had a direct land connection to these families (Costa 2000, Marske and Ivie 2003, neighboring areas and, as a corollary, that Cuba Navarrete-Heredia and Fierros-López 2001). It is has been long isolated from these lands by a sig- clear that additional families, genera, and species nificant oceanic barrier to beetle dispersal and colo- remain to be discovered in Cuba. There are some nization. Islands which have always been so iso- evident patterns in the Cuban beetle fauna, as dis- lated are called oceanic islands. Table 5 is a list of cussed below. beetle families which are known in continental warm-temperate Nearctic and tropical Neotropical An unbalanced fauna lands adjacent to Cuba and which might either be The family, subfamily, tribal and generic com- eventually discovered in Cuba, or whose absence position of the Cuban beetle fauna is dissimilar in helps to show the unbalanced nature of the Cuban many ways from that found in the neighboring con- fauna. tinental lands of the , Mexico, and Central America. The most conspicu- Generic and specific endemism and species ous feature is that many taxonomic groups are ab- swarms sent. This comparative difference is called a dis- 18 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Table 5. Beetle families with warm-temperate and tropical distibutions in the Neotropical and Nearc- tic biogeographic realms and which are seemingly “absent” from Cuba, and the likelyhood that they will be found to occur in Cuba. The families predicted to be present show the incomplete family level knowl- edge of the Cuban beetle fauna, and the Neotropical families which are absent help to show the history of isolation of the island.

Archostemata 1. in USA, Brazil and Argentina; a relict group, really absent 2. Ommatidae in Brazil and Argentina; a relict group, really absent Myxophaga 5. Lepiceridae México to S. America; probably absent 6. Microsporidae North and Central America; probably absent 7. Hydroscaphidae México to Brazil; probably absent 8. Torridinicolidae se Brazil; probably absent Polyphaga Staphylinoidea 24. North America; a relict group, probably absent 27. N. to S. America, in Hispaniola; really absent Hydrophiloidea 20. Syntellidae México; probably absent Scarabaeoidea 30 Lucanidae widespread Neotropical; probably absent Scirtoidea 45. Neotropical; probably present 46. Neotropical; probably present 48. Dascillidae arid Neotropical; really absent 49. Rhipiceridae Neotropical; really absent Byrrhoidea 52. Byrridae mostly temperate; really absent 55. Lutrochidae USA to Brazil; really absent 63. Callirhipidae Neotropical; really absent Elateroidea 64. Artematopidae Neotropical; maybe present 65. Known from amber in Hispaniola; possibly present 66. Neotropical; really absent 74. Telegeusidae sw USA to nw South America; probably absent 75. Neotropical, Acladocera in Hispaniola; probably present Bostrichoidea 79. Neotropical; probably present 81. Nosodendridae Neotropical; poorly studied, really absent Tenebrionoidea 126. Archeocrypticidae Neotropica; possibly present 149. Ischyomius is Neotropical; possibly present 150. Pyrochroidae questionable Neotropical, mostly temperate; probably absent Curculionoidea 153. temperate Neotropical; relict group, probably absent If oceanic islands have been in isolation long past, and if the colonization was sufficiently long enough, some ancient colonists have had opportu- ago, one or more colonizing species of each genus nity to evolve into genera which are endemic (lim- developed species level differences of its own, pro- ited) to the island. Table 6 is a list of the 18 genera ducing one or more species endemic to Cuba. presently thought to be Cuban endemics. However, Additionally, some genera, possibly ones which most beetle genera in Cuba also occur elsewhere. colonized the island relatively early in its history, Most of these genera arrived at some time in the have developed a great many species, often as an A Checklist of the Beetles of Cuba 19

Table 6. A. Beetle genera thought to be endemic to Cuba. There are some 190 beetle endemic genera in the entire West Indies.

Cerambycidae Calliclytus Fisher 1932 Cubaecola Lameere 1912 Cupeyalia Zayas 1975 Enodes Fisher 1926 Fisherostylus Gilmour 1963. Lethes Zayas 1975 Maisi Zayas 1988 Pygmaleptostylus Gilmour 1963 Trinoplon Zayas 1975

Coccinellidae Geodimmockius Chapin 1930 Neotina Gordon 1977

Curculionidae Doleropus Buchanan 1947 Nesolordops Marshall 1934 Pseudocaecocossonus Osella 1977

Heteroceridae Filiolus Pacheco 1964

Tenebrionidae Garridoa Marcuzzi 1985: 180 Orgidania Ardoin 1977 Trimytantron Ardoin 1977 (= Bielawskia Marcuzzi 1985)

B. Beetle genera previously thought to be endemic to Cuba, and which are now known to occur elsewhere.

Curculionidae Decouanellus Osella 1977: 399, now known in Bahamas, Puerto Rico, . Scarabaeidae Planophileurus Chapin 1932, now known in Bahamas. Staphylinidae Cubanotyphlus Coiffait and Decou 1972, now known in Florida. adaptive and evolutionary response to ecological op- probably between the southeastern USA and Cuba, portunity and the presence of “unfilled” niches. and between Mexico and Cuba. But even though These assemblages can be called “species swarms,” the affinities of the Cuban beetle fauna are a mix- and they are excellent groups for further study, to ture, it is still mostly Neotropical in its overall com- either verify their taxonomic status or to gain in- position. sights into the origin and action of speciation pro- cesses and the morphology and ecology of adaptive Native species in adjacent land masses radiation. Table 7 lists the 28 genera which are Many species which originated on Cuba may presently recognized as having 10 or more Cuban have naturally dispersed to other lands, or other species apiece. species from other lands have naturally moved into Cuba. These species naturally shared between Cuba A mixing of Neotropical and Nearctic faunas and other lands are called native species. Figure 2 Cuba shares with Mexico the property of being shows relative proportions of the 1092 native spe- an important mixing zone of Nearctic and Neotro- cies which are shared with other neighboring land pical beetle faunas at the level of genera and higher masses. All these species must have moved to or taxa. Some Nearctic genera and tribes are at their from Cuba at some time in the past (in the Pleis- southern distributional limits in the Caribbean in tocene or Recent) by crossing the barriers of the Cuba. Likewise, some Neotropical genera and tribes oceanic water gaps which isolated Cuba. are at their northern distribtional limits in the It is sometimes possible to deduce which direc- Caribbean in Cuba. Both these patterns suggest tion the movement has been by studying the other older (Tertiary) north and south faunal movements, members of the genus. The movements may have 20 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Table 7. Genera with 10 or more endemic species, showing significant speciation within Cuba.

Family Genus number of Species Carabidae Platynus 14

Staphylinidae Osorius 15

Scarabaeidae Phyllophaga (Cnemarachis) 68 Anoplosiagum 15

Lampyridae Callopisma 12 Photinus 11

Tenebrionidae Diastolinus 20 18 Cyrtosoma 13 Strongylium 11

Cerambycidae Elaphidion 29 Leptostylus 10

Chrysomelidae Lema 18 Cryptocephalus 62 Pachybrachis 16 Chlamisius 12 Metachroma 28 Leucocera 11 Oedionychus 12 Stoiba 11

Anthribidae 15

Curculionidae Lachnopus 30 Exophthalmus 20 Conotrachelus 10 Anthonomus 20 Acalles 12 Pseudomus 11 Cryptorhynchus 37 been through the air by wind and flight, on the sea accidental actions of humans. Table 8 lists the spe- surface by rafting on floating materials, and by cies presently thought to be introduced to Cuba. being transported by other animals such as birds. Many of these originate in the , and many Such movements were likely most frequent in the are important pests of agricultural crops, trees, and past during times of Pleistocene continental gla- stored products. The number of introductions of ciation, when sea levels were lower than at present, such pests will continue to grow without efforts to and the margins of surrounding islands and conti- prevent this. nents were closer to Cuba than now. Caribbean Insect Biogeography Introduced species. The West Indies are among the world’s biologi- A significant number of species of beetles (86 cally most important island groups. Cuba is the are presently recognized) have undoubtedly been largest island in the West Indies. It lies in the midst brought to Cuba through either the intentional or of other islands or island groups (Bahamas, A Checklist of the Beetles of Cuba 21

Table 8. Beetles recognized in the scientific literature as introduced by human activities (adventive) to Cuba, and approved English common name (from the list of approved common names of insects, Entomological Society of America).

Carabidae Leichenum canaliculatum variegatum Klug nigriceps (Dejean) Palorus ratzeburgii (Wissmann). The smalleyed flour pallens (Fabricius) beetle Staphylinidae Palorus foveicollis Blair tetracarinatus (Block) Taurocerus cornuta Fabricius Anotylus vinsoni (Cameron) Tribolium castaneum (Herbst). The red flour beetle Carpelimus corticinus (Gravenhorst) Tribolium confusum Jacquelin du Val. The confused Carpelimus politus Kiesenwetter flour beetle Gabronthus maritimus (Motschulsky) Anthicidae Oxytelus incisus Motschulsky floralis (Linnaeus). The narrow necked Phacophyllus parumpunctatus (Gyllenhal) grain beetle Philonthus discoideus (Gravenhorst) Sticticomus tobias (Marseul) Philonthus varians (Paykull) vicinus (Laferté-Sénectère) Philonthus ventralis (Gravenhorst) Cerambycidae Remus pruinosus Erichson (Say) Hybosoridae Monochamus titillator (Fabricius). The southern pine Hyposorus illigeri Reiche sawyer beetle Scarabaeidae Neoclytus acuminatus (Fabricius). The redheaded ash Aphodius lividus Olivier borer Ataenius stercorator Fabricius Neoptychodes trilineatus (Linnaeus) Cyclocephala notata (Illiger) Philematum femorale (Olivier) Onthophagus gazella Fabricius. The brown dung Stromatium fulvum (Villers) beetle Bruchidae Phyllophaga (Cnemarachis) analis (Burmeister) Acanthoscelides obtectus Say. The Phyllophaga (Cnemarachis) bruneri Chapin Bruchus pisorum (Linnaeus). The pea weevil. Pseudataenius walterhorni (Balthasar) Bruchus rufimanus (Boheman). The broadbean wee- Buprestidae vil Buprestis (Sterosa) apricans Herbst. The turpentine Zabrotes subfasciatus (Boheman). The Mexican bean borer weevil Chalcophora virginiensis (Drury) Anthribidae Dicerca tuberculata Laporte and Gory coffeae Fabricius. The coffee bean weevil Melanophila acuminata (Degeer) Brentidae Bostrichidae Cylas formicarius (Fabricius). The sweetpotato beetle Apate monacha Fabricius Curculionidae Dinoderus minutus (Fabricius). The bamboo Caulophilus oryzae (Gyllenhal). The broadnosed grain powderpost beetle weevil Heterobostrycus aequalis (Waterhouse) Euscepes postfasciatus (Fairmaire). The West Indian Minthea obstita (Wollaston) sweetpotato weevil Anobiidae Sitophilus granarius (Linnaeus). The granary weevil Gibbium aequinoctiale Boieldieu Sitophilus oryzae (Linnaeus). The rice weevil Trogossitidae Sitophilus zeamais Motschulsky. The maize weevil. Tenebriodes mauritanicus (Linnaeus). The cadelle Stenommatus musae Marshall beetle Scolytinae. Bark and ambrosia beetles. Cleridae Cocotrypes advena Blandford Necrobia ruficollis Fabricius. The redshouldered ham Cocotrypes carpophagus (Hornung) beetle Cocotrypes dactyliperda (Fabricius) Necropbia rufipes Degeer. The redlegged ham beetle Cocotrypes robustus Eichhoff Thanoclerus buquet (Lefebvre) Hypothenemus birmanus (Eichhoff). The kiawe Nitidulidae scolytid Carpophilus fumatus Boheman Hypothenemus columbi Hopkins Omosita colon (Linnaeus) Hypothenemus crudiae (Panzer) Urophorus humeralis (Fabricius) Hypothenemus javanus (Eggers) Silvanidae Hypothenenus mangiferae (Stebbing). The mango Ahasverus advena (Waltl). The bark beetle Oryzaephilus surinamensis (Linnaeus). The saw- Hypothenemus setosus (Eichhoff) toothed grain beetle Pagiocerus frontalis (Fabricius) Coccinellidae Premnobius cavipennis Eichhoff Catana clauseni Chapin Scolytogenes knabi (Hopkins) Rhodalia cardinalis (Mulsant). The vedalia beetle Trischidius atoma (Hopkins) Rhipiphoridae Xyloborinus andrewesi (Blandford) Rhipidius pectinicornis Thunberg Xylosandrus compactus (Eichhoff). The black twig Tenebrionidae borer Epitragus arulentus Kirsch 22 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Hispaniola, Jamaica, Caymans) and near continen- Geological tal land of North America (Florida) and Central The (and Cuba) were formerly America (Yucatan peninsula ). Most study of the thought to be composed of blocks of continental rock patterns and processes in Caribbean biogeography and to be relatively fixed in their location through have been based on vertebrates and plants (Woods geological time. The islands are composed of a com- 1989). Only recently have insects been used to help plex of volcanic rocks and marine limestones. It is understand the , and the now known that the islands have had a complex impact of the Caribbean on the . history of location, submergence, and emergence. Examples of biogeographic analyses of Carib- Donnelly (1988, 1989) has summarized the mod- bean insect groups are butterflies (Millere and Miller ern understanding of the geological history of the 1989; Scott 1972), lygaeid bugs (Slater 1988), Caribbean, but the Caribbean Basin remains one auchenorhynchous bugs (Ramos 1988), scaritine of the most controversial areas in the world for a and platynine carabid beetles (Nichols 1988, reconstruction of its past history. The best modern Liebherr 1988), caddis (Hamilton 1988), overview, from the point of biogeography, is that of drosophilid flies (Grimaldi 1988), ants (Wilson 1988) Iturralde-Vinent and MacPhee (1999). These refer- and sweat (Eichwort 1988). Genaro and Tejuca ences have been used to construct the following (2001) provide a discussion of patterns in the Cu- summary. ban insect fauna in general. Cuba seems to be composed mainly of a ter- A variety of biogeographic patterns exists be- rane of slivers of oceanic material stacked by com- yond those noted by Darlington (1957) for verte- pressive deformation of linear masses of sufficient brates. There are no fossil beetles to help in a re- thickness to achieve emergence. This seems to be construction of their history in Cuba. Only a re- the result of lateral compression at the edge of the cent dytiscid and a scarab have been recovered from Caribbean tectonic plate as it moved from an east- asphalt tar seeps in Matanzas Province (Iturralde- ern Pacific location into a location between the North Vinent et al. 2000; Valdés 1999). and South American plates in late and early Tertiary (Eocene) times. Cuba also seems to Biogeographic History of Cuba include parts of the Florida-Bahamas continental The biogeographic history of Cuba has long margin, and possible fragments of continental mar- stimulated interest, speculation, and investigation. gin derived from the south (the Chortis block). The basics of Cuban zoogeography, with data drawn All these fragments, island arc sediments, and from the vertebrate animals, were summarized by volcanics seem to have been consolidated in the late Darlington (1957), who categorized the West Indies Cretaceous with renewed compressive deformation as a “fringing archipelago.” The vertebrate fauna in the Eocene. Thus, Cuba possibly provides a small is unbalanced in its diversity. The absence of strictly exception to the strictly non-continental origin of fresh-water fishes, the limited diversity of amphib- the Greater Antilles. But there is no evidence for ians and reptiles, and the virtual absence of land any direct, emergent, continental land connection mammals is evidence that Cuba has always been at any time. However, there is evidence that there isolated by oceanic water gaps, just as is the case were very early but limited land connections to for the rest of the Greater Antilles (Hedges 2001). what is now Hispaniola and Puerto Rico, but not There is also a distributional structure or pat- Jamaica. tern in the vertebrate fauna. There is a primary After its origin by consolidation, the history of pattern in relict-endemic vertebrate groups (with Cuba was probably one of isolation from the rest of small distributional ranges) which shows an east- the Greater Antilles. From 40 Ma (million years ward diminution in overall species diversity, sug- ago) to 10 Ma it lay north of the east-west strike- gesting colonization primarily from the Yucatan slip faults that continued to juxtapose different sliv- Peninsula of Mexico directly into Cuba. There is a ers of the other islands (especially Hispaniola). secondary pattern suggesting dispersal to Cuba Cuba is presently composed of three connected through Jamaica. Some of these groups then moved blocks: Western Cuba, West Central Cuba, and from Cuba into the islands further to the east Eastern Cuba. Available evidence suggests that (Hispaniola and Puerto Rico). A full understanding these have existed separately as subaerial land since of the historical development of distributional pat- the Eocene-Oligocene boundary, at about 35 Ma, terns requires some knowledge of the geological and that the three blocks have been joined into the history of Cuba. present island since the mid-Miocene (after 8 Ma). A Checklist of the Beetles of Cuba 23

In summary, Cuba has had a complex geologi- Biogeography of the West Indies; patterns and cal origin and history. It is partly derived from two perspectives. CRC Press, Boca Raton, FL. former continental margins. It has mostly been Grimaldi, D.A. 1988. Relicts in the Drosophilidae submerged. It may have had potential terrestrial (Diptera). Pp. 183-213. In: Liebherr, J.K. (ed.). (or smaller water gap) connections to the Yucatan. Zoogeography of Caribbean Insects. Cornell These geological potentials, in combination with the Univ. Press, Ithaca, New York. zoogeographic data, are consistent with a Tertiary Hamilton, S.W. 1988. Historical biogeography of emergent history, with continued isolation by wa- two groups of Caribbean Polycentropus ter gaps, with the greatest source of over-water colo- (Trichoptera: Polycentropidae). Pp. 153-182. In: nists being Central America (directly with Yucatan Liebherr, J.K. (ed.). Zoogeography of Caribbean and via Jamaica from ). The closing of Insects. Cornell Univ. Press, Ithaca, New York. the Central American (Panamanian) isthmus in Hedges, S.B. 2001. Biogeography of the West the Pliocene modified oceanic circulation and formed Indies: an overview. Pp. 15-33. . In: Woods, C.A. the Gulf Stream, which must have altered over- and F.E. Sergile (eds.), Biogeography of the West water colonization processes. Lower glacial sea-levels Indies; patterns and perspectives. CRC Press, exposed vastly greater areas of land, and dimin- Boca Raton, FL. ished the size of the isolating water gaps. The Pleis- Iturralde-Vinent, M. A., and MacPhee, R. D. tocene changes of climate in the Caribbean are E. 1999. Paleogeography of the Caribbean Re- poorly understood, but seemingly lower sea levels gion: implications for Cenozoic biogeography. were coincident with increased aridity, and this Bull. American Museum of Natural History no. decreased the dispersal and colonization potential 238, 95 pp. of mesic terrestrial biotas. Iturralde-vinent, M. A., MacPhee, R. D. E., Díaz-Franco ,S., Rojas-Consuegra, R., References Súarez, W. and Lomba, A.. 2000. Las Breas de San Felipe, a Quaternary fossiliferous as- Costa, C. 2000. Estado de conocimineto de los Co- phalt seep near Martí (Matanzas Province, leoptera Neotropicales. Pp. 99-114. In: Martín- Cuba). Caribbean Journal of Science 36: 300- Piera, F., Morrone, J. J., and Melic, A. (eds). 313. Hacia un proyecto CYTED para el inventario y Liebherr, J.K. 1988. Biogeographic patterns of estimación de la diversidad entomologica en West Indian Platynus carabid beetles. Pp. 121- Iberoamérica: PrIBES-2000. M3m: Monografías 152. In: Liebherr, J.K. (ed.). Zoogeography of Tercer Milenio, vol. 1, SEA, Zaragoza, Spain. Caribbean Insects. Cornell Univ. Press, Ithaca, Darlington, P.J., Jr. 1957. Zoogeography. John New York. Wiley and Sons, New York. Marske, K.A. and Ivie, M.A. 2003. Beetle fauna Donnelly, T.W. 1988. Geologic constraints on of the United States and Canada. Coleopterists Caribbean biogeography. Pp. 15-37. In: Bulletin 57: 495-503. Liebherr, J.K. (ed.). Zoogeography of Caribbean Miller, L.D. and Miller, J.Y. 1989. The biogeog- Insects. Cornell Univ. Press, Ithaca, New York. raphy of West Indian butterflies (Lepidoptera: Donnelly, T.W. 1989. History of marine barriers Papilionoidea: Hesperioidea): a vicariance and terrestrial connections: Caribbean paleo- model. Pp. 229-262. In: Woods, C.A. (ed.), Bio- geographic inference from pelagic sediment geography of the West Indies: Past, Present analysis. Pp 103-118. In: Woods, C.A. (ed.). Bio- and Future. Sandhill Crane Press, Gainesville, geography of the West Indies; Past, Present FL. and Future, Sandhill Crane Press, Gainesville, Navarrete-Heredia, J. L. and Fierros-López, FL. H. E. 2001. Coleoptera de México: situación Eichwort, G.C. 1988. Distribution patterns and actual y perspectivas de estudio. Pp. 1-21. In: biology of West Indian sweat bees (Hy- Navarete-Heredia, J. L., Fierros-López, J. L., menoptera: Halictidae). Pp. 231-253. In: and Burgos-Soloria, A. (eds.). Tópicos sobre Co- Liebherr, J.K. (ed.). Zoogeography of Caribbean leoptera de Mexico. Universidad de Insects. Cornell Univ. Press, Ithaca, New York. Guadalajara-Universidad Autónoma del Estado Genaro, J.A. Tejuca, A.E. 2001. Patterns of en- de Morelos, Guadelajara, Mexico. demism and biogeography of Cuban insects. Pp. Nichols, S.W. 1988. Kaleidoscopic biogeography 77-83. In: Woods, C.A. and F.E. Sergile (eds.), of West Indian (Coleoptera: Carabidae). Pp. 71-120. In: Liebherr, J.K. (ed.). 24 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Zoogeography of Caribbean Insects. Cornell Liebherr, J.K. (ed.). Zoogeography of Caribbean Univ. Press, Ithaca, New York. Insects. Cornell Univ. Press, Ithaca, New York. Ramos, J.A. 1988. Zoogeography of the Valdés, P. 1999. Primero insectos fósiles cubanos auchenorrynchous Homoptera of the Greater (Coleoptera: Scarabaeidae: Dytiscidae). Cocuyo Antilles (). Pp. 61-70. In: Liebherr, 9: 17-18. J.K. (ed.). Zoogeography of Caribbean Insects. Wilson, E.O. 1988. The biogeography of the West Cornell Univ. Press, Ithaca, New York. Indian ants (). Pp. 214-230. In: Scott, J.A. 1972. Biogeography of Antillean but- Liebherr, J.K. (ed.). Zoogeography of Caribbean terflies. Biotropica 4: 32-45. Insects. Cornell Univ. Press, Ithaca, New York. Slater, J.A. 1988. Zoogeography of the West In- Woods, C.A. (ed.). 1989. The biogeography of the dian Lygaeidae (Hemiptera). Pp. 38-60. In: West Indies: Past, Present and Future. San- dhill Press, Gainesville, FL. A Checklist of the Beetles of Cuba 25 An Annotated Distributional Checklist of the Beetles of Cuba

ORDER COLEOPTERA SUBORDER MYXOPHAGA This suborder does not occur in Cuba or else- SUBORDER ARCHOSTEMATA where in the West Indies. Arce Pérez (1997, Only one species in this small suborder occurs Dugesiana 4: 41-50) gives a summary of the fauna in Cuba. of México with notes and references on the Neotro- pical fauna. 1. FAMILY CUPEDIDAE This family occurs in North and South America, 5. FAMILY LEPICERIDAE , , and Australia. None are known from Two species are known from flood debris along the West Indies. The larvae live in rotten wood. streams, from México to northern South America.

2. FAMILY OMMATIDAE 6. FAMILY MICROSPORIDAE This family occurs in southern Brazil and This family contains the single genus northern Argentina, and eastern Australia. Microsporus Kolenati, known from North and Cen- tral America, , Asia Minor, southeast Asia, 3. FAMILY CROWSONIELLIDAE Africa, Madagascar, and Australia. Adults and lar- This family is known only from a minute, wing- vae occur in mud, gravel and leaf litter on stream less, small eyed, soil dwelling species from central edges, or in moss in bogs. Italy. 7. FAMILY HYDROSCAPHIDAE 4. FAMILY MICROMALTHIDAE This family occurs in western North America, The telephone-pole beetles México, Panamá, Brazil, , North Africa, This family contains only one species which is southeast Asia, and Madagascar. Larvae live in native to and widespread in the southeastern USA; streams, where they feed on algae. larvae bore in partly rotted damp wood, such as pine logs, or in rotten structural timber. It is now 8. FAMILY widely introduced to many parts of the world. Since About 6 genera are known from Brazil, south- larvae are known as amber fossils from the Do- ern Africa, and Madagascar. Larvae live on wet minican Republic (Philips 2001) the family may be rock surfaces. a natural component of the Cuban fauna. SUBORDER ADEPHAGA Micromalthus LeConte 1878: 613 M. debilis LeConte 1878: 613; Silvestri 1941: 2; 9. FAMILY RHYSODIDAE Lawrence 1982; 486; Philips 2001: 159 The wrinkled bark beetles Distribution. Cuba, México to Brazil, Ha- Adults and larvae occur deep within the damp waii, Hong Kong, Africa. Native to se USA interior of logs and rotten wood. Adults force their Cuba localities. M: Grotta (cueva) Bellamar, way between layers of heartwood and seemingly larvae in wooden table feed on slime molds. A key to the Cuban species is Collections. in Bell and Bell (1995). Bell (2001) discusses the biogeography of the West Indian fauna. REFERENCES Philips, T. K. 2001. A record of Micromalthus TRIBE OMOGLYMMINI debilis (Coleoptera: Micromalthidae) from Cen- tral America and a discussion of its distribu- Plesioglymmius Bell & Bell 1978: 70 tion. Florida Ent. 84: 159-160. P. compactus Bell & Bell 1979: 437; 1995: 187 Silvestri, F. 1941. Distribuzione geografica del Rhysodes cubanus Zayas 1988: 9 (synonomy by Micromalthus debilis LeConte (Coleoptera Ivie 1991: 400); Ho: Altiplano de Mayari (t.l.), Micromalthidae). Boll. Soc. Ent. Italiana 73:- Zayas (type) 2. Distribution. Cuba. Endemic 26 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Gu. Mte. Iberia, (t.l. not Bell, R. T. and J. R. Bell. 1979. Rhysodini of the specified). Ho: altiplano de Mayari world. Part II. Revisions of the smaller genera Collections. Pittsburgh. Cambridge (type) (Coleoptera: Carabidae or Rhysodidae). Quaest Ent. 15: 377-446. TRIBE CLINIDIINI Bell, R.T. and J.R. Bell. 1995. The Rhysodini (Insecta; Coleoptera; Carabidae) of Cuba. Ann. Clinidium Kirby 1830: 6 Carnegie Mus. 64: 185-195. C. (Protainoa) extrarium Bell and Bell 1978: 63; Ivie, M. A. 1991. Taxonomic notes on a little known 1995: 189 publication: Zayas, 1988, Entomofauna Distribution. Cuba. Endemic Cubana. Order Coleoptera. Coleop. Bull. 45: 399- Cuba localities. PR: Lomas del Rio. (t.l.) 401. Collections. Pittsburgh C. (Tainao) curvicosta Chevrolat 1873: 215; Bell 10. FAMILY CARABIDAE and Bell 1995: 190 The ground beetles C. guildingi Kirby, Chevrolat 1864: 611; Reviewed by G. E. Ball, March, 2001 Gundlach 1891: 116 These are mostly active predators on plant sur- Distribution. Cuba. Endemic faces, and in litter. Some are phytopha- Cuba localities. Gu: N of Imias, 3-4000'. SC: gous. The larvae are usually liquid-feeding preda- Cobre Range; Gran Piedra, 6 km NE Siboney, tors. Larvae of and are ecto- Rio Juragua; Loma del Gato 8000' (t.l.) parasitic. The main reference and introduction to Collections. Gatineau. Pittsburgh. Gundlach the West Indian carabid fauna is the list by Erwin no. 1181. Cambridge (type) and Sims (1984). This is a most important paper C. (T.) chevrolati Reitter 1880: 30; Bell and Bell because it gives keys to the genera known in Cuba. 1995: 191 Keys to all Neotropical genera are in Reichardt C. (T.) turquinense Bell 1970: 318; SC: Pico (1977). Darlington (1937) is an itemization of Cu- Turquino, 3-5000' (t.l.), Cambridge (type) ban carabids known to him, and includes several Distribution. Cuba. Endemic keys for identification. Mateu (1977) has provided Cuba localities. “New Granada” (t.l.). SC: a list of carabids found in Cuban caves, but none of Pico Turquino; Broche; Palma Mocha. Gu: the species are cave limited (Peck et al. 1998). Valdés Nibujon, Gu: Baracoa (1999) provides a list of Cuban Cicindelinae. The Collections. Cambridge. Pittsburgh. San higher classification used here follows that in Francisco Bousquet and Larochelle (1993). Ball (1992a) pro- C. (C.) humeridens Chevrolat 1873: 215; Bell and vides an excellent framework by which to analyze Bell 1995: 193 the entire Cuban carabid beetle fauna. Distribution. Cuba. Endemic Cuba localities. SUBFAMILY PAUSSINAE Collections. TRIBE OZAENINI

REFERENCES Pachyteles Perty 1830: 3 Bell, R.T. 1970. The Rhysodini of North America, P. gyllenhali (Dejean) 1825: 436 (); Chevrolat Central America, and the West Indies (Co- 1863: 190 (Goniotropis); Gundlach 1891: 18; leoptera, Carabidae or Rhysodidae). Misc. Publ. Ball and McCleve 1990: 94 Ent. Soc. Amer. 6: 289-324. Goniotropis pallida Chevrolat 1863: 190 Bell, R.T. 2001. Rhysodine beetles in the West Distribution. Cuba, México, , Indies. Pp. 117-125. In: Woods, C.A. and F.E. Surinam, Brazil, sw USA (AZ) Sergile (eds.), Biogeography of the West Indies; Cuba localities. Ci: Cayamas patterns and perspectives. CRC Press, Boca Collections. Cambridge Raton, FL. Bell, R. T. and J. R. Bell. 1978. Rhysodini of the SUBFAMILY CARABINAE world. Part I. A new classification of the tribe, TRIBE CARABINI and a synopsis of Omoglymmius, subgenus Nitiglymmius, new subgenus (Coleoptera: Calosoma Weber 1801: 20 Carabidae or Rhysodidae). Quaest. Ent. 14: 43- C. (Castrida) sayi Dejean 1826: 198; Gidaspow 1963: 88. 300 A Checklist of the Beetles of Cuba 27

C. alternans Fabricius; Jacquelin du Val 1856: C. boops Dejean 1831: 258; Jacquelin du Val 1856: 19; Chevrolat 1863: 186: Gundlach l891: 12 5; Chevrolat 1863: 185; Gundlach 1891: 11 Distribution. Cuba, Bahamas?, Jamaica, Distribution. Cuba, Hispaniola, Puerto Rico, Hispaniola, Puerto Rico, México, Guatemala, Caicos Islands, Jamaica, Virgin Islands USA (CA-ND-FL) Cuba localities. M: Cárdenas Cuba localities. Widespread. Ci: Cienfuegos. Collections. Gu: Guantánamo. Ho: Holquin. LT: Jobabo. C. cardini Leng and Mutchler 1916: 689 PR: Guanahacabibes Distribution. Cuba. Endemic Collections. Cuba localities. C. (Calodrepa) splendidum Dejean 1831: 558; Collections. Jacquelin du Val 1856: 19; Chevrolat 1863: 186; C. cubana Leng and Mutchler 1916: 698 Gundlach 1891: 12.; Gidaspow 1963: 283 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola, USA (GA, Cuba localities. FL). Collections. Cuba localities. Gu: Guantánamo. LT: [C. dorsalis Say 1817: 20; see Valdés 1999: 13] Jobabo. At lights Distribution. Cuba in error; México, eastern Collections. USA C. marginata Fabricius 1775: 226; Gundlach 1891: SUBFAMILY CICINDELINAE 11 (see Freitag 1992) Distribution. Cuba, Bahamas, widespread TRIBE MEGACEPHALINI eastern USA Cuba localities. M: between Matanzas and Megacephala Latreille 1802: 79 Punta de Jeacos, on beaches M. acutipennis Dejean 1825: 13; Jacquelin du Val Collections. 1856: 4; Gundlach 1891:9 C. olivacea Chaudoir 1854: 118; Chevrolat 1863: M .a. adonis Laporte 1834: 83; Jacquelin du 185; Gundlach 1891: 10 Val 1856: 4; Chevrolat 1863: 184 (Tetracha) Distribution. Cuba, USA (FL) Distribution. Cuba, Hispaniola, Puerto Rico Cuba localities. Ha: Havana, on coast Cuba localities. Widespread Collections. Collections. C. trifasciata Fabricius 1781: 286 M. carolina Linné 1766: 657; Jacquelin du Val C. tortuosa Dejean 1825: 87; Jacquelin du Val 1856: 3 1856: 5; Chevrolat 1863: 185; Gundlach 1891: M. occidentalis Klug 1829: 11; Chevrolat 1863: 10 184 (Tetracha); Gundlach 1891: 9 Distribution. Cuba, Bahamas, Grand Cay- Distribution. Cuba, Jamaica, Bahamas, man, Jamaica, Hispaniola, Puerto Rico, St. Grand Cayman, México, Guatemala, Nicara- Thomas, St. John, St. Martin, , gua, USA (CA-TX-VA-FL) , , St. Barthélemy, , Cuba localities. Widespread St. Croix, Virgin Islands, USA (TX-NJ-FL) Collections. Cuba localities. Widespread. On beaches [M. rutilans Thomson 1857a: 35; see Valdés 1999: Collections. Carnegie (as tortuosa) 13] C. viridicollis Dejean 1831: 265; Jacquelin du Val M. r. infuscata Mannerheim 1837: 6; Jacquelin 1856: 6; Chevrolat 1863: 185; Gundlach 1891: du Val 1856: 3; Chevrolat 1863: 184 (Tetracha) 11 Distribution. Cuba record an error?, C. v. carbonaria Zayas 1988: 11 (Zayas, type) Hispaniola, Puerto Rico, St. Thomas, St. John, Distribution. Cuba. Endemic St. Croix, St. Martin, St. Barthélemy, USA(?) Cuba localities. Ci: Soledad. Ha: Havana. SC: Gran Piedra. SS: Buenos Aires. .; Mina Carlota; TRIBE CICINDELINI San Blas; Trinidad Mts. On sandy roads and at lights Cicindela Linné 1758:407 Collections. Zayas C. acuniae Mutchler 1924: 1; Barreto 1924: 167 Distribution. Cuba. Endemic SUBFAMILY SCARITINAE Cuba localities. Ca: Sierra Cubitas (t.l.) (see Nichols 1988a, 1988b) Collections. New York (type) TRIBE 28 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Ardistomis Putzeys 1846: 637 C. (P.) bipustulata Fabricius 1801: 125 (); A. elongatulus Putzeys 1866: 208 Jacquelin du Val 1856: 17; Gundlach 1891:24 Distribution. Cuba. Endemic (Aspidoglossa) Cuba localities. Ci: Soledad. Marsh and Distribution. Cuba, Jamaica, Cayman Is- swamps in lowlands; winged lands, Hispaniola, México, eastern USA, Collections. Cambridge Canada (ON) A. nitidipennis Darlington 1934: 70. Cuba localities. Widespread. Ca: Central Distribution. Cuba. Endemic Jaronú. Ci: Soledad. Ha: Almendares. PR: San Cuba localities. Ci: Soledad (t.l.). SC: Cauto Vicente. SC: Aguadores El Cristo; Cauto River; Cruces. PR: Sierra del Collections. Rangel. VC: Hanabanilla Falls. Mesophile C. (P.) fasciata Putzeys 1846: 624 Collections. Cambridge (type) Dyschirius insularis Jacquelin du Val 1857: 13; A. ramsdeni Darlington 1937: 120 Chevrolat 1863: 191; Gundlach 1891: 21 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Jamaica, Cay- Cuba localities. SC: Pico Turquino (t.l.), 1800 man Islands, Hispaniola, Puerto Rico, Lesser m. Vestigial wings; wet forest humicole Antilles, Yucatan, México to South America, Collections. Cambridge (type) USA (FL) Cuba localities. CA: Baraguá. Ci: Cayamas. Aspidoglossa Putzeys 1846: 626 Gr: Cauto El Cristo. Ha: Havana. M: Cárdenas. A. mexicana Chaudoir 1837:18 Common at lights A. vulnerata Putzeys 1846: 633; Jacquelin du Val Collections. Cambridge 1856: 17 (Not Cuba, but Puerto Rico, South C. (Semiclivina) cubae Darlington 1934: 68 America) Distribution. Cuba. Endemic A. comma Putzeys 1846: 634; Jacquelin du Val Cuba localities. Ca: Camaguëy. CA: Baraguá. 1856: 18; Chevrolat 1863: 193; Gundlach 1891: Ci: Cayamas; Soledad (t.l.). SC: Aguadores. In 24 river gravels Distribution. Cuba, Bahamas, México Collections. Cambridge (type) Cuba localities. Ca: Central Jaronú. CA: C. (S.) dentipes Dejean 1825: 415; Jacquelin du Val Baraguá. Ci: Cayamas; Soledad. Gu: Maisi. Ha: 1856: 15; Chevrolat 1863: 192 Almendares; Sierra de Anafe, Caimito. .PR: Distribution. Cuba, Jamaica, México, se Sierra de Rangel. SC: Calabazas; coast below USA Pico Turquino Cuba localities. CA: Baraguá. Ci: SS: Buenos Collections. Cambridge Aires; Soledad. Gr: Cauto El Cristo. Ha: Camoa; ; Sierra de Anafe, Caimito. PR: San Clivina Latreille 1802: 96 Diego. Winged hygrophile C. (Isoclivina) limbipennis Jacquelin du Val 1857: Collections. 16 C. simplex Chevrolat 1863: 192 Dyschiriodes Jeannel 1941: 300 Distribution. Cuba, Hispaniola, Puerto Rico D. larochellei (Bousquet) 1988: 374 (Dyschirius) Cuba localities. Ci: Cienfuegos; Soledad. Gr: D. erythrocerus Leconte, Mateu 1977: 377 Guane. Ha: Coama. PR: San Vicente. SC: Distribution. Cuba, Bahamas, Hispaniola, Aquadores Yucatan, México, USA (FL) Collections. Cuba localities. Ha: mud of Lago . C. (Paraclivina) biguttata Putzeys 1866: 155. PR: Hoyo de Fania. Halobiontic, flightless C. bisignata Putzeys 1846: 624; Chevrolat 1863: Collections. Bucharest? 193 D. sublaevis (Putzeys) 1846: 562 (Dyschirius) Distribution. Cuba, Jamaica Distribution. Cuba, México (Yucatan), USA Cuba localities. CA: Baraguá. Ci: Cayamas; (FL) Soledad. Ha: Almendares. IJ. SC: Aguadores. Cuba localities. Ha: Havana, on saline mud Eurytopic hygrophile; agricultural associate; flats. Halobiontic. A probable predator on winged Bledius staphylinids (Thayer and Purrington Collections. 2004: 441) Collections. A Checklist of the Beetles of Cuba 29

Halocoryza Allaud 1919: 100 TRIBE SCARITINI H. arenaria (Darlington) 1939: 84 Distribution. Cuba, Bahamas, Jamaica, Cay- Scarites Fabricius 1801: 123 man Islands, Virgin Islands, , S. (S.) alternans Chaudoir 1843: 729; Jacquelin du México, Central America, South America, Af- Val 1856: 13; Chevrolat 1863: 191; Gundlach rica, USA (s FL, Keys). Intertidal, winged 1891:20 Cuba localities. Distribution. Cuba. Endemic in highlands Collections. (disjunct distribution?) Cuba localities. Ci: Soledad. Gu: San Ger- Putzeys 1866: 103 man. Ho: Aguayo. SC: Aguadores; Santiago. O. cristalensis Zayas1988: 13 Or: Central Palma. Winged Distribution. Cuba. Endemic Collections. Cuba localities. Ho: Sierra Cristal (t.l.) S. (S.) illustris Chaudoir 1880: 66 Collections. Zayas (type) Distribution. Cuba, México, Swan Island O. cubanus Zayas 1988: 12 (Honduras), Panamá and south Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Pico Cuba (t.l.) Collections. Collections. Zayas (type) S. (S.) marinus Nichols 1986: 258 O. rufus (Putzeys) 1846: 564 (Dyschirius); Distribution. Cuba, Yucatan, Bahamas, USA Jacquelin du Val 1856: 14; Chevrolat 1863: 191; (FL) Mateu 1977:377 Cuba localities. Ha: Havana (t.l.). Brac- Dyschirius brevicarinatus Putzeys 1846: 571; hypterous, intertidal halobiont of seaweed wash- Jacquelin du Val 1856: 15; Chevrolat 1863: 192 up on sandy beaches Distribution. Cuba, Bahamas, Cayman Is- Collections. Cambridge (type) lands, Jamaica, Puerto Rico, Yucatan, México, S. (S.) subterraneus Fabricius 1775: 249; Jacquelin USA (FL) du Val 1856: 12; Chevrolat 1863: 191; Cuba localities. Ci: Soledad. Ha: Almendares; Gundlach 1891:21 Marianao; Sierra de Anafe, Caimito. M: S. (S.) patruelis LeConte 1845: 207 Matanazas. PR: Lago Canoa; Cabañas. SC: edge Distribution. Cuba, Yucatan, México, USA ( of Laguna de Juragua; Aquadores. Winged, at GA, FL) lake and swamp edges Cuba localities. Widespread. Winged, Collections. Bucharest? halophobic, hygrobiontic; subtropical moist and dry lowlands Putzeys 1846: 649 Collections. Cambridge S. arimao Darlington 1934: 71 S. (Taeniolobus) cubanus Bänninger 1937: 321 Distribution. Cuba, Jamaica, Hispaniola Distribution. Cuba. Endemic Cuba localities. Ci: Santa Clara, Cuba localities. Ci: SS: Buenos Aires, Cumanayagua; Soledad (t.l.). Riparian in gravel Trinidad Mts., c. 1000 m, (t.l.). Flightless on shorelines halophobic hygrobiont; lower montane wet for- Collections. Cambridge (type) est Collections. Cambridge (type) Kult 1850: 301 S. cyaneolimbatus (Chevrolat) 1863: 194 SUBFAMILY TRECHINAE Ardistomis gundlachi Gundlach 1891: 25; Leng TRIBE TRECHINI and Mutchler 1914: 395 Distribution. Cuba, Cayman Islands, Schaum 1860: 663 Hispaniola, México. P. columbus Darlington 1934: 86 Cuba localities. CA: Baraguá. Ci: Cayamas; Distribution. Cuba. Endemic Soledad. IJ. Gu: Maisi. Gr: Cauto El Cristo. Cuba localities. Ci: Soledad (t.l.). PR: Sierra SC: Aguadores; Santiago de Cuba. Riparian de Rangel. In gravel bars Collections. Collections. Cambridge (type) 30 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE BEMBIDIINI Cuba localities. Ci: Cayamas; Santa Clara; Soledad (t.l.). Mesophile Bembidion Latreille 1802: 82 Collections. Cambridge (type) B. affine Say 1823: 86; Jacquelin du Val 1856: 23; Chevrolat l863: 198; Gundlach 1891: 32 Micratopus Casey 1914a: 42 Distribution. Cuba, USA (AL-TX-OH-DE) M. parviceps Darlington 1934: 85 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 413 Cuba localities. SS: Jatibonico (t.l.). In field B. cubanum Darlington 1937a: 121 soils Distribution. Cuba. Endemic Collections. Cambridge (type) Cuba localities. PR: Sierra Rangel (t.l.).; SC: Gran Piedra; Sierra de Cobre, 1000 m. Winged, Bates 1882: 144 at streamsides M. autumnalis (Bates) 1882: 137 Collections. Cambridge (type) Distribution. Cuba, , Guadeloupe, B. (Emphanes) darlingtoni Mutchler 1934: 3; Mateu México, Guatemala, , Panamá 1977: 377 Cuba localities. Gu: Sierra de Purial. SC: Distribution. Cuba, Puerto Rico Sierra de Cobre. SS: Sierra de Trinidad. Winged Cuba localities. SS: Sierra de Trinidad. Ci: hygrophile Soledad (t.l.). Ha: Laguna de Ariguanabo. Collections. Winged hygrophile Collections. Cambridge (type). Bucharest? Paratachys Casey 1918a: 174 B. (Notaphus) sparsum Bates 1882: 151; Mateu P. abruptus (Darlington) 1934: 80 (Tachys); Mateu 1977: 377 1977: 378 Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba, Hispaniola, Guadeloupe, México to Nicaragua, South America Cuba localities. Ha: Laguna de Ariguanabo Cuba localities. SC: Cueva Jibara; Pico (lake edge) Turquino (t.l.); Sierra de Cobre. Winged; Collections. Bucharest? hygrophile B. turquineum Darlington 1937: 122. Collections. Cambridge (type). Bucharest? Distribution. Cuba. Endemic P. cubax (Darlington) 1934: 78 (Tachys); Mateu Cuba localities. SC: Pico Turquino, c. 1600 1977: 377 m (t.l.). Along mountain brooks, wings vesti- Distribution. Cuba. Endemic gial Cuba localities. Ci: Cayamas; Santa Clara; Collections. Cambridge (type) Soledad (t.l.). Gu: San Carlos Estancia. SC: B. viridicolle (Laferté-Sénectère) 1841: 48 Cueva Atabex; Cueva Jibara. SS: Jatibonico; B. apicale Jacquelin du Val 1856: 23; Chevrolat Sierra de Trinidad 1863: 198; Gundlach 1891: 32 Collections. Cambridge (type). Bucharest? Distribution. Cuba, Puerto Rico, México, P. filax (Darlington) 1934: 83. widespread eastern North America Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. Ci: Cayamas (t.l.); Santa Collections. Gundlach no. 413 (B. apicale) Clara. SS: Sierra de Trinidad Collections. Cambridge (type) Elaphropus Motschulsky 1839: 73 P. paulax (Darlington) 1934: 80 E. yunax (Darlington) 1939: 87 (Tachys); Mateu Distribution. Cuba. Endemic 1977: 377 Cuba localities. Ci: Cayamas (t.l.); Santa Distribution. Cuba, Hispaniola Clara Cuba localities. SC: Baconao, in rotting al- Collections. Cambridge (type) gae on beach P. striax (Darlington) 1934: 82 (Tachys); Mateu Collections. Cambridge (type from 1977: 378 Hispaniola). Bucharest? Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas; Santa Clara; Lymnastis Motschulsky 1862: 27 Soledad (t.l.). Ho; Cueva Bariay (many in L. americana Darlington 1934: 83 guano) Distribution. Cuba. Endemic Collections. Cambridge (type) A Checklist of the Beetles of Cuba 31

P. vorax (LeConte) 1852: 194 (Tachys); Mateu 1977: D. rutilus (Chevrolat) 1863: 197 (); 378 Gundlach 1891: 25, 3l; Bousquet and Laplante Distribution. Cuba, Puerto Rico, México, 1997: 708 USA (CA-AZ-NM-TX) Distribution. Cuba, Cayman Islands, Cuba localities. Ha: Laguana de Ariguanabo. Hispaniola, México, Colombia, , USA PR: Hoyo de Fania. SC: Laguna Juragua. SS: (NJ-FL-AZ) Sierra de Trinidad Cuba localities. Gu: Baitiquiri; Guantánamo Collections. Bay; Savana 1a Mar. On saline mudflats or saline soil LeConte 1851: 191 Collections. Cambridge. Gainesville P. elegantulus Laferté-Sénectère 1841: 46; Mateu 1977: 377 SUBFAMILY P. blandulus Schaum 1860: 202; Chevrolat 1863: TRIBE MORIONINI 198; Gundlach 1891:33 (Bembidion); Mateu 1977: 377 Morion Latreille 1810: 159 Distribution. Cuba, Puerto Rico M. monilicornis (Latreille) 1806: 206 (Harpalus); Cuba localities. Gu: Rio Sabana-la-mar at Chevrolat 1863: 190; Gundlach 1891: 20 San Antonio. SC: Laguana Juragua Distribution. Cuba, South America, wide- Collections. spread in e and se USA P. immaculatus Bates 1871b: 246 Cuba localities. Distribution. Cuba, México, Honduras, Costa Collections. Rica, Panamá, South America Cuba localities. Ci: Cayamas; Soledad TRIBE LOXANDRINI Collections. P. jamcubanus Erwin 1974b: 57 Loxandrus LeConte 1852: 250. See Allen and Distribution. Cuba, Jamaica Ball 1980 Cuba localities. Ci: Soledad; San Blas L. celeris Dejean 1828: 246; Mateu 1977: 378 (Trinidad Mts.). Gr: Cauto El Cristo. Gu: Maisi L. cruentatus Chevrolat 1858: 209 Collections. Distribution. Cuba, Bahamas, Puerto Rico, México, se USA (TX-SC-FL) Polyderis Motschulsky 1862: 27 Cuba localities. Ha: Laguna de Ariguanabo P. ridiculus Schaufuss 1879: 552 Collections. Bucharest? Limnastis capito Bates 1884: 287; Mateu 1977: L. crenatus LeConte 1852: 252 378 Distribution. Cuba, se USA (LA-SC-FL) Distribution. Cuba, St. Thomas, Virgin Is- Cuba localities. lands, Guatemala Collections. Cuba localities. PR: Quemado de Pineda L. cubanus Tschitschérine 1903: 60 Collections. Distribution. Cuba, Bahamas, Hispaniola, Puerto Rico, México, Costa Rica Tachys Stephens 1828b: 4 Cuba localities. Ci: Soledad T. bradycellinus Hayward 1900: 224 Collections. Distribution. Cuba, , Jamaica, USA L. nocticolor Darlington 1934: 91 (TX-LA-FL) Distribution. Cuba. EndemicSister species of Cuba localities. L. rectangulus of México, Grand Cayman, and Collections. USA (FL) T. translucens Darlington 1937: 123 Cuba localities. Ci: Soledad (t.l.). In flood Distribution. Cuba. Endemic debris Cuba localities. Gu: Guantánamo, Boquerón Collections. Cambridge (type) (t.l.) (L. rectangulus LeConte 1878: 377) Collections. Cambridge (type) Distribution. Hispaniola, Grand Cayman, México, USA (TX, FL) TRIBE POGONINI Cuba localities. Collections. Diplochaetus Chaudoir 1871: 43 32 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE PTEROSTICHINI Distribution. Cuba, Bahamas (Mayaguana), Cayman Islands, Jamaica, Hispaniola, Puerto Poecilus Bonelli 1810: synoptic table Rico, Central America P. chalcites Say 1823: 56 Cuba localities. Gu: Guantánamo Bay. SC: Distribution. Cuba, widespread USA Santiago, Roig Botanical Gardens Cuba localities. Collections. Gatineau. Gainesville Collections. D. cinctus (Putzeys) 1878: 45 Distribution. Cuba. Endemic Pterostichus Bonelli 1810: synoptic table Cuba localities. P. cubensis Darlington 1937: 123. Collections. Distribution. Cuba. Endemic Cuba localities. Ci: Santa Clara, near Selenophorus Dejean 1829: 80 (see Ball 1992a) Buenos, 3000'. (t.l.); Trinidad Mts. Winged S. alternans Dejean 1829: 86 mesophile Distribution. Cuba, Bahamas, Hispaniola, Collections. Cambridge (type) Puerto Rico, Guadeloupe, South America, México TRIBE HARPALINI Cuba localities. Collections. Acupalpus Latreille 1829: 291 S. chalybaeus Dejean 1829: 110; Jacquelin du Val A. convexulus Darlington 1934: 112 1856: 22; Chevrolat 1863: 196; Gundlach 189l: Distribution. Cuba. Endemic 29; Mateu 1977: 378 Cuba localities. Ca: Camagüey (t.l.). CA: Distribution. Cuba, Bahamas, Jamaica, Baraguá Hispaniola, Puerto Rico, Antigua, Guadeloupe Collections. Cambridge (type) Cuba localities. Widespread. Ci: Soledad. Ho: A. iridens Motschulsky 1864: 201 Cueva de Jagüey IJ: Nueva Gerona. Or: Distribution. Cuba. Endemic Guardalavaca. PR: Cueva del Indio. SC: Matias. Cuba localities. At lights Collections. Collections. Bucharest? Pittsburgh S. flavilabris Dejean 1829: 97; Ball l992a: 96 Dejean 1829: 62 S. flavilabris cubanus Darlington 1935: 203 (=S. A. puncticollis Putzeys 1878: 34 cubanus cubanus Darlington) Distribution. Cuba, Hispaniola, Cayman Is- Distribution. Cuba, Bahamas (Mayaguana), lands, Jamaica, Puerto Rico Jamaica Cuba localities. CA: Baraguá Cuba localities. SS: Sierra de Trinidad (t.l. Collections. of subspecies). Winged mesophile Collections. Cambridge (type) Erichson 1837: 64 S. discopunctatus Dejean 1829: 92; Jacquelin du B. cubanus Darlington 1934: 110 Val 1856: 22; Chevrolat 1863: 196; Gundlach Distribution. Cuba. Endemic 189l: 30 Cuba localities. Ci: Cayamas (t.l.) S. cuprinus Dejean 1829: 96; Chevrolat 1863: 196; Collections. Cambridge (type) Gundlach 1891: 29; Blackwelder l944: 49 B. festinans Casey 1914: 257 Distribution. Cuba, Bahamas, Cayman Is- Distribution. Cuba, USA (TX-AR-SD-WY) lands, Jamaica, Hispaniola, Puerto Rico, Cuba localities. Ci: Cayamas (t.l.); Antigua, Central and South America, USA (FL) Cumanayagua; Santa Clara; Soledad Cuba localities. Widespread. IJ: Nueva Collections. Washington (type) Gerona B. velatus Darlington 1934: 111 Collections. Gatineau. Pittsburgh Distribution. Cuba, Puerto Rico S. integer (Fabricius) 1801: 196 (Harpalus) Cuba localities. Ci: Soledad (t.l.); in flood Distribution. Cuba, Bahamas, Jamaica, debris Hispaniola, Puerto Rico Collections. Cambridge (type) Cuba localities. Collections. LeConte 1835: 381 (see Ball 1992a) S. nonseriatus Darlington 1934: 109 D. beauvoisi (Dejean) 1829: 98 Distribution. Cuba, Jamaica, Hispaniola A Checklist of the Beetles of Cuba 33

Cuba localities. Gu: Guantánamo Bay. PR: Stenomorphus Dejean 1831: 696. See Ball et al. Sierra de Rangel. Winged hygrophile. 1991: 932 Collections. Gainesville S. cubanus Darlington 1937: 135 S. pyritosus Dejean 1829: 84; Jacquelin du Val Distribution. Cuba. Endemic 1856: 21; Chevrolat 1863: 195; Gundlach 1891: Cuba localities. Gr: Cauto El Cristo (t.l.).; in 28; Mateu 1977: 378 wet ditch Distribution. Cuba, Bahamas, Jamaica, Collections. Cambridge (type) Hispaniola, Puerto Rico, México, Panamá, South America TRIBE LICININI Cuba localities. Widespread. Ci: Soledad; Gu: Guantánamo Bay. Ha: Cueva Cinco Cuevas; Badister Clairville 1806: 90 (see Ball 1992b; 377) IJ: Nueva Gerona; Sierra de las Casas. SC: B. (Baudia) reflexus LeConte 1880: 165 Siboney, caves Distribution. Cuba, Jamaica, Cayman Is- Collections. Bucharest? Gainesville. Pitts- lands, Bahamas, Hispaniola, Mexico, eastern burgh USA S. sinuatus Gyllenhal 1806: 203 Cuba localities. CA: Baraguá. Ci: Soledad. S. parumpunctatus Gundlach 1891: 29; Collections. Blackwelder 1944: 50; Mateu 1977: 378 Distribution. Cuba, Bahamas, Cayman Is- Brullé 1834: 407 lands, Jamaica, Hispaniola, Puerto Rico, D. major LeConte 1848: 418 Antigua, Guadeloupe D. m. melissisa Ball 1959: 78 Cuba, USA (FL- Cuba localities. Ci: Soledad. Gu: AL-LA-TX) Guantánamo Bay Distribution. Cuba, widespread southeastern Collections. Bucharest? Gainesville USA S. solitarius Darlington 1934: 106 Cuba localities. Ha: Camoa; Somorrostro Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Cayamas; Santa Clara. SS: Zaza del Medio (t.l.) TRIBE PANAGAEINI Collections. Cambridge (type) S. striatopunctatus Putzeys 1878: 33 Brullé 1835: 433 Distribution. Cuba, Bahamas, Cayman Is- C. effeminata Darlington 1934: 89 lands, Jamaica, Hispaniola, Puerto Rico, Distribution. Cuba. Endemic México, Leeward and , South Cuba localities. Ca: Camaguëy; Central America, se USA (FL) Jaronú. Ci: Soledad. (t.l. CA: Baraguá. At lights Cuba localities. Collections. Cambridge (type) Collections. C. quadrisignatus Chevrolat 1835: 187 Distribution. Cuba, Puerto Rico, St. Thomas, Stenolophus Stephens 1827: 67 México S. cruentatus Chevrolat 1858: 209; 1863: 197; Cuba localities. Gundlach 1891: 30 Collections. Distribution. Cuba. Endemic C. sauricollis Darlington 1934: 88 Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Soledad (t.l.). In flood S. infuscatus Dejean 1829: 54 debris Distribution. Cuba, e USA (FL) Collections. Cambridge (type) Cuba localities. Collections. TRIBE CHLAENINI S. ochropezus Say 1825: 54 S. cruentatus Chevrolat 1858: 209 Chlaenius Bonelli 1810: synoptic table Distribution. Cuba, Puerto Rico, México C. cubanus Chaudoir 1876a: 238 (Baja), USA (CA) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. 34 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. gundlachi Chaudoir 1876a: 148; Gundlach 1891: USA (NY to TX) 26 Cuba localities. Ci: Soledad. Ha: Camoa. IJ: Distribution. Cuba. Endemic Punta Pedernales Cuba localities. M: Cárdenas Collections. Bucharest? Collections. [S. gilvipes Chaudoir 1883: 504]. Replacement name C. niger Randall 1838: 34; Gundlach 1891: 26 for S. pallipes Reiche 1846 (not Brullé 1838). Distribution. Cuba, widespread USA Not cited by Bousquet (1996) as a northern Cuba localities. M: Cienaga de Zapata Neotropical-southern Nearctic species. Cuba Collections. record in Blackwelder 1944: 45 C. perplexus Dejean 1831: 655 Distribution. Not Cuba. Northern South Chalaenius poeyi Chevrolat 1863: 194; Gundlach America 1891: 25 S. insulana (Jacquelin du Val) 1856: 20 (Oodes); C. circumcinctus Say, Gundlach 1891: 26; Leng Chevrolat 1863: 195; Gundlach 1891: 28; and Mutchler 1914: 395 Bousquet 1996: 481 Distribution. Cuba, Puerto Rico, Hispaniola, Distribution. Cuba, Grand Cayman, México, México, USA (TX-FL) Trinidad, , USA (TX) Cuba localities. Ci: Soledad. M: Cárdenas; Cuba localities. Ci: Santa Clara; Soledad. Ha: Cienaga de Zapata Camoa Collections. Collections. Cambridge S. (Dejean) 1826: 379; Mateu 1977: 378 TRIBE OODINI (see Bousquet 1996) S. agilis Laferté-Sénectère 1851: 273 Distribution. Cuba, Puerto Rico, México to Anatrichius LeConte 1853:391 and Brazil A. oblonga Horn 1891: 37; Bousquet 1996: 457 Cuba localities. Gu: Rio Sabana la Mar, San Distribution. Cuba, México, USA (TX) Antonio Cuba localities. Ci: Soledad. Collections. Bucharest? Collections. Cambridge S. olivacea (Bates) 1878: 589; Bousquet 1996: 489 Distribution. Cuba, Hispaniola, México, Oodinus Motschulsky 1865: 353 Puerto Rico O. darlingtoni Bousquet 1996: 465. Cuba localities. Ci: Soledad. Gr: Cauto El Distribution. Cuba. Endemic Cristo. Gu: Maisi; Sabana la Mar. Ho: Holguín Cuba localities. Ci: Soledad. Gr: Cauto El Collections. Cambridge. New York Cristo. (t.l.). S. mexicana (Chevrolat) 1835: 102; Bousquet 1996: Collections. Cambridge (type). 486 O. pseudopiceus Bousquet 1996: 462 S. sulcatus Chevrolat 1863: 195; Leng and [O. mexicanus Chaudoir 1882: 323 Mutchler 1914: 395; Gundlach 1891: 28 (misidentification for Cuba)] Distribution. Cuba, México, USA (NJ-TX) Distribution. Cuba, Hispaniola, Bahamas, Cuba localities. Widespread USA (FL). Collections. Cuba localities. Gr: Cauto El Cristo. S. tibialis (Chevrolat) 1834: 46 (Amara); Chevrolat Collections. Cambridge. Washington 1863: 194 (Stenous); Bousquet 1996: 493 Stenous pallipes Brullé 1838: 32; Gundlach 1891: Oodes Bonelli 1810: synoptic table 27 O. amaroides Dejean 1831: 674; Gundlach 1891: Distribution. Cuba, Hispaniola, Jamaica, 27; Bousquet 1996: 471 Puerto Rico, México, Guatemala, South Distribution. Cuba, eastern USA America Cuba localities. Gr: Cauto El Cristo Cuba localities. Widespread Collections. Collections. Cambridge. Carnegie

Stenocrepis Chaudoir 1857: 39 TRIBE PENTAGONICINI S. duodecimstriata (Chevrolat) 1835: 173; Mateu 1977: 378; Bousquet 1996: 485 Schmidt-Goebel 1846: 47 (see Bell Distribution. Cuba, Hispaniola, Jamaica, 1985) Bahamas. México, Nicaragua, Guatemala, se A Checklist of the Beetles of Cuba 35

P. flavipes (LeConte) 1853: 377; Gundlach 1891: 19 P. bucheri (Darlington) 1937: 130 (Rhombodera); Mateu 1977: 378 Distribution. Cuba. Endemic Rhombodera bicolor Gundlach 1891: 20 Cuba localities Gu: Mts. north of Imias (t.l.); Distribution. Cuba, Bahamas, Hispaniola, Sierra de Purial. SC: Gran Piedra. Hygrophile; Jamaica, Puerto Rico, Guadeloupe, , Costa winged Rica, Panamá, South America, USA (SC-FL- Collections. Gatineau. Cambridge (type) LA) P. carabiai (Darlington) 1937: 129 (); Cuba localities. Widespread Mateu 1977: 378 Collections. Gatineau. Bucharest? Gundlach Distribution. Cuba. Endemic no. 42. Cuba localities. Gu: Mts. north of Imias; Si- P. nigricornis Darlington 1934: 121. erra de Purial (t.l.). Ho: Cueva Bariay. SC: Distribution. Cuba, Cayman Islands, Cen- Cueva Jibara. Gu: Maisi; above Los Llanos. tral America, Colombia, USA (FL); in flood de- Hygrophile bris Collections. Bucharest? Cambridge (type) Cuba localities. Ci: Soledad (t.l.). Gu: N of P. fratrorum (Darlington) 1937: 129 Imias; Sierra de Purial. PR: Sierra Rangel. SC: Distribution. Cuba. Endemic Gran Piedra. Winged Cuba localities. SC: Cobre Range, Loma del Collections. Cambridge (type) Gato (t.l.). Hygrophile; winged Collections. Cambridge (type) TRIBE P. mediopterus (Darlington) 1937: 130 Distribution. Cuba. Endemic Bonelli 1810: synoptic table Cuba localities. SC: Pico Turquino, 1000-2000 A. decorum (Say) 1823: 53 m (t.l.). Hygrophile, wings vestigial A. (Anchomenus) extensicolle Say 1825: 54; Collections. Cambridge (type) Chevrolat 1863: 197; Gundlach 1891: 31 P. medius (Darlington) 1937: 130 (Colpodes); Mateu A. extensicolle cubanum Darlington 1934: 97; 1977: 378 Mateu 1977: 378 Distribution. Cuba. Endemic A. n.sp. Zayas 1988: 14 Cuba localities. Ha: Laguna Ariguanabo. SC: Distribution. Cuba, Grand Cayman, - Gran Piedra (t.l.) maica, Hispaniola, México, USA (widespread, Collections. Bucharest? Cambridge (type) FL) P. pinarensis (Darlington) 1937: 128 Cuba localities. Ca: Camagüey. CA: Baraguá. Distribution. Cuba. Endemic Ci: Cayamas; Cumanayagua; Marcata, Rio Cuba localities. PR: Rio Taco Taco; Sierra Bayamo; Santa Clara; Soledad. Gr: Cauto El de Rangel, 500 m (t.l.). Hygrophile; winged Cristo. Ha: Laguna Ariguanabo; Camoa. At Collections. Cambridge (type) lights, in river gravel P. subangustus (Darlington) 1937: 131 Collections. Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino, 4500- Platynus Bonelli (see Liebherr 1987, 1992) 5500' (t.l.). Hygrophile; wings vestigial baragua group Collections. Cambridge (type) P. baragua (Darlington) 1935: 197 (Colpodes); P. turquinensis (Darlington) 1937: 131 Mateu 1977: 378 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino, 1000-2000 Cuba localities. CA: Baraguá (t.l.). Ci: m (t.l.). Hygrophile; wings vestigial Cienfuegos; Soledad; Sumidero de Vega Alta at Collections. Cambridge (type) Topes de Collantes. Ha: Havana. SS: Buenos Aires. , Trinidad Mts. Hygrophile; winged. degallieri group (not ovatulus group) Collections. Bucharest? Cambridge (type) P. coptoderoides (Darlington) 1937: 134 P. bruneri (Darlington) 1937: 132 Distribution. Cuba; Bahamas (Andros, Distribution. Cuba. Endemic Greater Exuma, Nassau, S. Bimini) Cuba localities. SC: Cobre Range, Loma del Cuba localities. Gu: Cabo Maisi, Maya River Gato, 3000-3600' (t.l.). Mesophile; vestigial (t.l.). PR: Pinar del Rio SC: Rio Turquino; Si- wings erra del Cobre. Mesophile; winged Collections. Cambridge (type) Collections. Cambridge (type) 36 Arthropods of Florida and Neighboring Land Areas, Vol. 18

jaegeri group Distribution. Cuba, Jamaica, México, Gua- P.acuniai (Darlington) 1937: 133 temala Distribution. Cuba. Endemic Cuba localities. Ci: Soledad; Trinidad Mts., Cuba localities. SC: Pico Turquino (t.l.), 1600 San Blas, 300-1000 m. Gr: Cauto El Cristo. Gu: m. Wings vestigial Baracoa; Imias; Maisi; upper Ovando River. PR: Collections. Cambridge (type) Sierra Rangel. SC: Aguadores; Daiquiri; coast P. cubensis (Darlington) 1937: 132 below Turquino Distribution. Cuba. Endemic Collections. Cambridge Cuba localities. Ci: Trinidad Mts., SS: Buenos Aires (t.l.), 650-1200 m. Wings vesti- Eucaerus LeConte 1853: 386 gial E. insularis Darlington 1934: 120; Liebherr 1988: Collections. Cambridge (type) 22 Distribution. Cuba. Endemic memnonius group Cuba localities. Ci: Cayamas; Santa Clara; P. aequinoctialis (Chaudoir) 1850: 382 Soledad. (t.l.); in flood debris Colpodes silviae Zayas 1988: 15; Ivie 1991: 400 Collections. Cambridge (type) Distribution. Cuba, Jamaica, Hispaniola, México, Central and South America Euphorticus Horn 1881: 144 Cuba localities. None given E. pubescens Dejean 1831: 30; Darlington 1941: Collections. Zayas (type) 13; Liebherr 1988: 28 E. p. aeneolus Bates 1883: 156. Cuba, México, Sericoda Kirby 1837:14 Guatemala, South America S. montana Liebherr 1991 Distribution. Cuba, Jamaica, México, Gua- Distribution. Cuba. Endemic temala, Honduras, Colombia, Trinidad, se USA Cuba localities. SC: Sierra Maestra (t.l.), (LA-OH-SC-FL) 1200-1300 m Cuba localities. CA: Baraguá. Ha: Santiago Collections. Cambridge (type) de las Vegas. IJ: Nueva Gerona Collections. Cambridge. Warsaw TRIBE PERIGONINI Dejean 1831: 28; Darlington 1934: Perigona Laporte 1835: 151 98; 1941: 13; Liebherr 1988: 34 P. laevigata Bates 1872: 200 L. leucopterus Chevrolat 1863: 198: Gundlach 1891: Distribution. Cuba, México, Central America 37 (); Liebherr 1988: 34 Cuba localities. Distribution. Cuba, Puerto Rico, Jamaica, Collections. Hispaniola P. nigriceps (Dejean) 1831: 44; Mateu 1977: 378 Cuba localities. Ci: Soledad, gravel bar; SS: Distribution. Cuba (introduced), Puerto Rico, Trinidad Mts. Gr: Cauto El Cristo; Bayamo. Guadeloupe, , widespread USA; cos- Gu: Baracoa; Maisi. PR: San Diego; Sierra mopolitan, introduced from Old World Rangel. SC: Aguadores; Daiquiri; Imias. VC: Cuba localities. IJ: Santa Fe. At lights Hanabanillo Falls Collections. Bucharest? Cambridge Collections. Cambridge P. picea Darlington 1934: 98 Distribution. Cuba, Hispaniola, Guadeloupe TRIBE ODACANTHINI Cuba localities. Ci: Cayamas; Santa Clara. Gu: San Carlos Estancia; Sierra de Purial; Colliuris DeGeer 1874: 79 Yateras. SC: Pico Turquino; Sierra de Cobre. C. gundlachi Darlington 1934: 122 Winged, mesophile Distribution. Cuba. Endemic Collections. Cambridge Cuba localities. Ci: Soledad (t.l.); in damp places TRIBE LACHNOPHORINI (see Liebherr l988) Collections. Cambridge (type) C. noah Darlington 1934: 123 Anchonoderus Reiche 1843: 38 Distribution. Cuba. Endemic A. subtilis Bates 1871: 33; Darlington 1934: 98; Cuba localities. Ci: Soledad. (t.l.), in flood 1941: 13; Liebherr 1988: 27 debris A Checklist of the Beetles of Cuba 37

Collections. Cambridge (type) Distribution. Cuba, Bahamas, Cayman Is- C. picta Chaudoir 1843: 697; Gundlach 1891: 18 lands, Jamaica (Casnonia) Cuba localities. M: Cárdenas. SS: Trinidad C. p. concluda Liebke 1930: 689 Collections. C. p. extrema Liebke 1930: 689 A. d. delicata Darlington 1934: 118 Distribution. Cuba, México, USA (AZ, TX) A. strandi Liebke 1939: 119 (Ball in litt.) Cuba localities. Gu: between Manati and Los Distribution. Cuba, Hispaniola; A. d. cay- Canos man in Cayman Islands Collections. Cuba localities. Ci: Cayamas; Santa Clara; Soledad (t.l.). SC: Santiago, Roig Botanical Latreille 1829: 371 Garden; 6 km NE Siboney. Sifting and in fields L. compressus Zayas 1988: 20 Collections. Gatineau. Cambridge (type) Distribution. Cuba. Endemic A. lata Darlington 1934: 119 Cuba localities. M: Cienaga de Zapata (t.l.) Distribution. Cuba, Bahamas, Hispaniola Collections. Zayas (type) Cuba localities. Ci: Soledad (t.l.) L. cubanus Zayas 1988: 19 Collections. Cambridge (type) Distribution. Cuba. Endemic A. parallela Dejean 1825: 218 (Cymindis); Cuba localities. Ca: Santa Lucia (t.l.) Jacquelin du Val 1856: 7; Chevrolat 1863: 187; Collections. Zayas (type) Gundlach 1891: 15 L. dorsalis Fabricius 1801: 220; Gundlach 1891: Distribution. Cuba, Bahamas (not Puerto 19 Rico, México) Distribution. Cuba, e USA Cuba localities. M: Cárdenas. SS: Trinidad Cuba localities. Collections. Collections. A. scobifera Darlington 1953: 12; Mateu 1977: 378 Distribution. Cuba, Jamaica TRIBE CYCLOSOMINI Cuba localities. SC: Siboney; at lights Collections. Bucharest? Aephnidius MacLeay 1825: 23. Data from Ball, A. sulcicollis Jacquelin du Val 1857: 8; Chevrolat in litt. 1863: 188; Gundlach 1891: 15 A. (Aephnidius) ciliatus Mutchler 1934: 130 Distribution. Cuba, Bahamas Distribution. Cuba, Cayman Islands, Baha- Cuba localities. M: Cárdenas mas (Mayaguana Island) Hispaniola, Puerto Collections. Rico, to Argentina Cuba localities. Ci: Cayamas; Santa Clara; Chaudoir 1846: 12 Soledad. Gu: Maisi. SC: 10 km NE Caney A. sericeus Darlington 1934: 116 Collections. Gatineau Distribution. Cuba. Endemic A. (Macracanthus) sericatus (Chaudoir) 1846: 541 Cuba localities. Ci: Soledad (t.l.). On hot river (Aephenidius) sand banks A. brevicillus (Chevrolat) 1863: 189 (Masoreus); Collections. Cambridge (type) Gundlach 1891: 17; Mateu 1977: 378 Distribution. Cuba, Hispaniola, Puerto Rico Dejean 1825: 220 Cuba localities. Ci: Soledad. Ha: Cueva C. rubricollis Dejean 1825: 225; Jacquelin du Val Mariana Quivican. Ho: Cueva de los Santos. 1856: 9 M: Cárdenas. PR: Viñales, San Vicente. SC: 10 C. elegans Chaudoir 1844: 469; Chevrolat 1863: km NE Caney, Arroyo Grovert, 300 m; Vista 187; Gundlach 1891: 14 Alegre. Often in caves, winged Distribution. Cuba. Endemicnot USA Collections. Gatineau. Bucharest? Cambridge Cuba localities. Ci: Soledad. SC: Sierra de Cobre; Santiago, Roig Botanical Gardens. SS: TRIBE LEBIINI Sierra de Trinidad. Mesophile; winged, at lights Collections. Gatineau Apenes LeConte 1851: 174 (see Ball 1992a) C. tinctula Darlington 1934: 117 A. coriacea Chevrolat 1863: 188; Gundlach 1891: Distribution. Cuba. Endemic 15 Cuba localities. Ci: Cayamas (t.l.); Santa Clara. SC: Sierra de Cobre. Winged mesophile 38 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Cambridge (type) Collections. Zayas (type) L. dorsovittata Zayas 1988: 16 Dejean 1825: 173 (see Shpeley and Ball Distribution. Cuba. Endemic 1993) Cuba localities. SC: costera de Juraguá (t.l.) C. festiva Dejean 1825: 174; Jacquelin du Val 1856: Collections. Zayas (type) 11; Chevrolat 1863: 189; Gundlach 1891: 18; L. esurialis Casey 1920: 257; Mateu 1977: 378 Shpeley and Ball 2000: 146 Distribution. Cuba; USA (TX, SD) Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. SC: Siboney. At lights Puerto Rico, México to Panamá, Venezuela, Collections. Ecuador, Peru, Brazil, USA (FL) L. marginicollis Dejean 1825: 271 Cuba localities. Ci: Soledad.; Trinidad Mts. L. cyanea Dejean 1831: 386; Jacquelin du Val M: Cárdenas. SC: Boniato Mts., Rio Frio; Gran 1856: 11; Chevrolat 1863: 188; Gundlach 1891: Piedra 16 Collections. Cambridge Distribution. Cuba, México, Guatemala, C. picea Dejean 1826: 458 Nicaragua, widespread USA C. unicolor Chevrolat 1834: 40; Gundlach 1891: Cuba localities. M: Cárdenas 18 Collections. Distribution. Cuba, México to Panamá, wide- spread South America, USA (FL) Microlestes Schmidt-Goebel 1846: 41 Cuba localities. Ca: Camaguëy. SC: Santiago M. poeyi (Jacquelin du Val) 1857: 10 (Dromius); de Cuba Chevrolat 1863: 188 (Blechrus); Gundlach 1891: Collections. 16 Distribution. Cuba. Endemic Euproctinus Leng and Mutchler 1927: 14 (see Cuba localities. M: Cárdenas. SC: Aguadores Shpeley 1986) Collections. Cambridge E. trivittatus LeConte 1878: 373; Shpeley 1986: 295- 300 Chaudoir 1869: 145 Distribution. Cuba, USA (FL) (dispersed P. costata Darlington 1937: 135 from Cuba to FL) Distribution. Cuba. Endemic Cuba localities. Ha: Santa Maria del Mar; Cuba localities. SC: Sierra del Cobre (t.l.); Santaiago de las Vegas Loma del Gato. Mesophile; winged Collections. Collections. Cambridge (type) P. dealata Darlington 1937: 136 Galerucidia Chaudoir 1872: 416 Distribution. Cuba. Endemic G. dimidiata Chaudoir 1872a: 420 Cuba localities. SC: Pico Turquino (t.l.), Distribution. Cuba. Endemic 1000-2000 m. Wings vestigial Cuba localities. Collections. Cambridge (type) Collections. P. imitatrix Darlington 1934: 114. Distribution. Cuba. Endemic Lebia Latreille 1802: 85 Cuba localities. Ci: Cayamas (t.l.); Santa L. abdominalis Chaudoir1843: 704; Gundlach 1891: Clara. SC: Sierra de Cobre. SS: Sierra de 16 Trinidad. Mesophile; winged Distribution. Cuba, Nicaragua to e USA Collections. Cambridge (type) Cuba localities. P. plagiata Darlington 1934: 114 Collections. Distribution. Cuba. Endemic L. bitaeniata Chevrolat 1834: fascicle 2; Gundlach Cuba localities. Gu: Guantamano (t.l.) 1891: 16 Collections. Cambridge (type) Distribution. Cuba, Costa Rica to s USA (s P. schwarzi Darlington 1934: 115 TX) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Ci: Cayamas. (t.l.); Santa Collections. Clara L. darlingtoni Zayas 1988: 17 Collections. Cambridge (type) Distribution. Cuba. Endemic Cuba localities. SC: Ciudamar (t.l.) A Checklist of the Beetles of Cuba 39

Plochionus Latreille & Dejean 1824: 150 T. bierigi Liebke 1934: 387 P. bicolor Notman 1919: 234 Distribution. Cuba. Endemic Distribution. Cuba, USA (FL) Cuba localities. Ha: Havana (t.l.) Cuba localities. Collections. Liebke? (type) Collections. T. dorsalis Brullé 1834: 181 P. pallens (Fabricius) 1775: 244; Chevrolat 1863: Distribution. Cuba, Jamaica, USA (wide- 189; Gundlach 1891: 17 spread) Plochionus bonfilsii Dejean, Jacquelin du Val Cuba localities. 1856: 10 Collections. Distribution. Cuba (introduced), Bahamas, T. marginicollis (Darlington) 1934: 126 México, Central America, South America, USA (Pseudaptinus): Mateu 1977: 378 (FL, MA, PA); cosmopolitan, native to Old Distribution. Cuba, Hispaniola World Cuba localities. CA: Baraguá. Ci: Soledad Cuba localities. M: Cárdenas (t.l.). PR: Sierra de Rangel; Arroyo de los Collections. Guineos. SC: 6km ne Siboney. In flood debris, P. rubrofasciatus Zayas 1988: 18 at lights; winged Distribution. Cuba. Endemic. Probably con- Collections. Gatineau. Bucharest? Cambridge specific with P. bicolor, but not established (G. (type) Ball in litt.) T. pygmaeus (Dejean) 1826: 460 Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba, USA (LA-AR-SC-FL) Collections. Zayas (type) Cuba localities. Collections. TRIBE ZUPHIINI Zuphium Latreille 1806: 198 Pseudaptinus Laporte 1835: 56 Z. bierigi Liebke 1933: 467 P. cubanus Chaudoir 1877: 252 Distribution. Cuba. Endemic Distribution. Cuba, USA (FL) Cuba localities. LT?: Lago de Canoa (t.l.) Cuba localities. Ci: Soledad Collections. Liebke? (type) Collections. Z. cubanum Liebke 1933: 470 P. deceptor Darlington 1934: 128 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Santa Clara; Cruces (t.l.) Cuba localities. Ci: Soledad (t.l.). Flood de- Collections. Liebke? (type) bris Z. mexicanum Chaudoir 1862: 314 Collections. Cambridge (type) Distribution. Cuba, México, Costa Rica P. insularis Mutchler 1934a: 4 Cuba localities. Distribution. Cuba, Puerto Rico, Trinidad, Collections. Brazil Cuba localities. Ci: Soledad TRIBE GALERITINI Collections. P. salebrosus Liebke 1934: 375 Fabricius 1801: 214 Distribution. Cuba. Endemic G. lecontei tenebricosa (Klug) 1834: 65; Reichardt Cuba localities. Ci: Santa Clara, Cruces (t.l.) 1967: 36; Ball and Nimmo 1983: 318 Collections. Liebke? (type) G. vetula Chevrolat 1863: 186; Gundlach 1891: 13 Thalpius LeConte 1851: 174 Distribution. Cuba, Cayman Islands, T. apicalis (Darlington) 1934: 125 Hispaniola, not Honduras Distribution. Cuba. Endemic Cuba localities. Cuba localities. Ci: Soledad (t.l.) Collections. Collections. Cambridge (type) G. ruficollis Dejean 1825: 191; Reichardt 1967: 61 T. arrogans Liebke 1934: 385 G. erthrodera Brullé 1834b: 103; Jacquelin du Distribution. Cuba. Endemic Val 1856: 7; Chevrolat 1863: 186; Gundlach Cuba localities. Ha: Havana (t.l.) 1891: 13 Collections. Liebke? (type) Distribution. Cuba, México to Panamá, Ja- maica 40 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Ci: Soledad. SS: Trinidad Mts Ball, G.E. and A.P. Nimmo. 1983. Synopsis of Collections. Cambridge the species of subgenus Progaleritina Jeannel, [G. unicolor Latreille & Dejean 1823: 117; including reconstructed phylogeny and geo- Reichardt 1967: 106] graphical history (Coleoptera: Carabidae: Distribution. Not Cuba (mislabeled). South Galerita Fabricius). Trans. Amer. Ent. Soc. America 109: 295-356. Ball, G. E., D. Shpeley, and D.C. Currie. 1991. SUBFAMILY BRACHININAE The New World genus Stenomorphus Dejean (Coleoptera: Carabidae: Harpalini); classifica- Brachinus Weber 1801: 22 tion, allometry, and evolutionary consider- B. adustipennis Erwin 1970: 81 ations. Canadian Ent. 123: 933-988. B. lateralis Dejean, Jacquelin du Val 1856: 127; Bell, R.T. 1985. Pentagonica of the West Indies Chevrolat 1863: 187; Gundlach 1891: 13 (Coleoptera: Carabidae). Coleop. Bull. 39: 321- Distribution. Cuba, México, Panamá, east- 327. ern USA (including FL) Bousquet, Y. 1996. Taxonomic revision of Nearc- Cuba localities. Ci: Soledad. Ha: Camoa. M: tic, Mexican, and West Indian Oodini (Co- Cárdenas. SC: Santiago de Cuba leoptera: Carabidae). Canadian Ent. 128: 433- Collections. Cambridge 537. B. brunneus Laporte 1835: 59 Bousquet, Y. and S. LaPlante. 1997. Taxonomic B. gilvipes Mannerheim, Gundlach 1891: 14 review of the New World Pogonini (Coleoptera: Distribution. Cuba, Hispaniola, Puerto Rico, Carabidae). Canadian Ent. 129: 699-731. Virgin Islands, Costa Rica, French Guiana Bosquet, Y. and A. Larochelle. 1993. Catalogue Cuba localities. CA: Baraguá. M: Cárdenas. of the Geadephaga (Coleoptera: Trachy- Colon pachidae, Rhysodidae, Carabidae including Collections. Cambridge Cicindelini) of America north of México. Mem. Ent. Soc. Canada 167, 397 pp. REFERENCES Darlington, P.J., Jr. 1934. New West Indian Allen, R.T. and G.E. Ball. 1980 (1979). Synopsis Carabidae, with a list of the Cuban species. of Mexican taxa of the Loxandrus series (Co- Psyche 41: 66-131. leoptera: Carabidae: Pterostichini). Trans. Darlington, P.J., Jr. 1937. West Indian Carabidae Amer. Ent. Soc. 105: 481-576. III: new species and records from Cuba, with a Ball, G.E. 1959. A taxonomic study of the North brief discussion of the mountain fauna. American Licinini with notes on the Old World Memorias de la Sociedad Cubana de Historia species of the genus Diplocheila Brullé (Co- Natural ‘Felipe Poey’ 11: 115-136. leoptera). Mem. Amer. Ent. Soc. No. 16, 258 Darlington, P.J., Jr. 1939. West Indian pp. Carabidae. V. New forms from the Dominican Ball, G.E. 1992a. Geographical distribution and Republic and Puerto Rico. Memorias de la evolution of the Selenophorini (Harpalini) and Sociedad Cubana de Historia Natural ‘Felipe Apenes LeConte (Lebiini) in the Antilles (Co- Poey’ 13: 79-101. leoptera: Carabidae). pp 79-121:. In: G.R. Darlington, P.J., Jr. 1941. West Indian Noonan, G.E. Ball, and N.E. Stork (eds). The Carabidae. VI. The Jamaican species, and their biogeography of ground beetles (Coleoptera: wings. Psyche 48: 10-15. Carabidae) of mountains and islands. Intercept Darlington, P. J., Jr. 1953. West Indian Ltd., Andover, Hants, UK. Carabidae (IX). More about the Jamaican spe- Ball, G.E. 1992b. The tribe Licinini (Coleoptera: cies. Occ. Pap. Mus. Institute Jamaica Carabidae); a review of the genus-groups and (Kingston) no. 8, 1-14. of the species of selected genera. J. New York Erwin, T.L. and L.L. Sims. 1984. Carabid beetles Enstomol. Soc. 100: 325-380. of the West Indies (Insecta: Coleoptera): a syn- Ball, G.E. and S. McCleve. 1990. The Middle opsis of the genera and checklists of tribes of American genera of the tribe Ozaenini, with Caraboidea, and of the West Indian species. notes about the species in southwestern United Quaest. Ent. 20: 341-466. States and selected species in México. Quaest. Freitag, R. 1992. Biogeography of West Indies ti- Ent. 26: 30-116. ger beetles (Coleoptera, Cicindelidae), pp. 123- 157. In: G.R. Noonan, G.E. Ball, and N.E. Stork A Checklist of the Beetles of Cuba 41

(eds.). The biogeography of ground beetles of Shpeley, D. 1986. Genera of the subtribe mountains and islands. Intercept Ltd., U.K. Metallicina and classification, reconstructed Gidaspow, T. 1963. The genus Calosoma in Cen- phylogeny and geographical history of the spe- tral America, the Antilles, and South America cies of Euproctinus Leng and Mutchler (Coleoptera, Carabidae). Bull. Amer. Mus. Nat. (Coleoptea: Carabidae: Lebiini). Quaest. Ent. Hist. 124 (7): 275-314. 22: 261-349. Ivie, M. A. 1991. Taxonomic notes on a little known Shpeley, D. and G.E. Ball. 1993. Classification, publication: Zayas, 1988, Entomofauna reconstructed phylogeny, and geographical his- Cubana. Order Coleoptera. Coleop. Bull. 45: 399- tory of the New World species of Coptodera 401. Dejean (Coleoptera: Carabidae; Lebiini). Proc. Liebherr, J.K. 1987. A taxonomic revision of the Ont. Ent. Soc. 124: 1-182. West Indian Platynus beetles (Coleoptera: Shpeley, D. and G.E. Ball. 2000. A taxonomic Carabidae). Trans. Amer. Ent. Soc. 112: 289- review of the subtribe Pericalina (Carabidae: 368. Lebiini) in the , with de- Liebherr, J. K. 1988. Redefinition of the supertribe scriptions of new species and notes about clas- Odacanthitae, and revision of the West Indian sification and zoogeography. Insecta Mundi 14: Lachnophorini (Coleoptera: Carabidae). Quaest. 1-185 Ent. 24: 1-24. Thayer, M.K., F.F. Purrington. 2004. Bledius Liebherr, J.K. 1991. Phylogeny and revision of beattyi Blackwelder (Coleoptera: Staphylinidae: the Anchomenus clade: the genera , ), new to the United States, and co- Anchomenus, Sericoda, and Elliptoleus (Co- occurrences of some Bledius Leach and leoptera; Carabidae; Platynini). Bull. Amer. Dyschiriodes Jeannel (Coleoptera: Carabidae: Mus. Nat. Hist. 202: 1-163. Scaratinae) species in south Florida. Coleop. Liebherr, J.K. 1992. Phylogeny and revision of Bull. 589: 439-442. the Platynus degallieri species group (Co- Valdés R., P. 1999. Lista anotada de los leoptera: Platynini). Bull. Amer. Mus. Nat. escarabajos tigres (Coleoptera: Carabidae) de Hist. 214, 114 pp. Cuba. Cocuyo 8: 13-14. Mateu, J. 1977. Carabiques récoltés par les expe- ditions biospéologiques cubano-roummaines à 11. FAMILY GYRINIDAE Cuba (1969 et 1973). Resultats des expéditions The whirligig beetles biospéologiques Cubano-Roumaines à Cuba. 2: Adults are commonly seen swimming rapidly 376-379. in circles on surfaces of ponds. They prey on organ- Nichols, S. W. 1988a. Systematics and biogeogra- isms falling on the water surface. The genus phy of West Indian Scaritini (Coleoptera: is nocturnal and species are hidden by day. Carabidae). Doctoral thesis, Cornell University. UMI Disserttion Services, Order no. 8804612, SUBFAMILY GYRININAE Ann Arbor, Michigan xiii + 193 pp. TRIBE GYRININI Nichols, S.W. 1988b. Kaleidoscopic biogeography of West Indian Scaritinae (Coleoptera: Dineutus Macleay 1825: 30 Carabidae), pp. 71-120. In: Liebherr, J.K. (ed.). D. americanus (Linnaeus) 1767: 568; Spangler Zoogeography of Caribbean Insects. Cornell 1981: 147 Univ. Press, Ithaca, NY. D. metallicus Aubé, Chevrolat 1863: 203; Peck, S.B., A.E. Ruiz-Baliú, and G.F. Garcés Gundlach 1891:41 Gonzaléz. 1998. The cave-inhabiting beetles Distribution. Cuba, Bahamas, Jamaica, of Cuba (Insecta: Coleoptera): diversity, distri- Hispaniola to Virgin Islands, USA (FL) bution, and ecology. J. Cave Karst Studies 60: Cuba localities. Widespread 156-166. Collections. Cambridge. Washington Reichardt, H. 1967. A monographic revision of D. longimanus Olivier 1792: 11; Jacquelin du Val the American Galeritini (Coleoptera, 1856: 30: Chevrolat 1863: 203; Gundlach 1891: Carabidae). Arquivos de Zoologia (Sao Paulo) 40; Spangler 1973: 353; 1981: 147 15: 1-176. Distribution. Cuba, Jamaica, Hispaniola, Reichardt, H. 1977. A synopsis of the genera of Puerto Rico, Costa Rica. Cuban specimens are Neotropical Carabidae (Insecta: Coleoptera). the subspecies D. l. cubensis Ochs 1926:87 Quaest. Ent. 13: 346-493. Cuba localities. Widespread 42 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Cambridge. Washington Haliplus Latreille 1802: 77 H. bierigi Guignot 1936: 115; Spangler 1981: 167 TRIBE OREOTOCHILINI Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. Gr: Cauto El Gyretes Brullé 1834: 241 Cristo. Ha: Playa Marianao (t.l.) G. cubensis Régimbart 1883: 392; Spangler 1981: Collections. Guignot (type) 168 H. carinatus Guignot 1936: 116; Spangler 1981: G. vulneratus Aubé 1838: 752; Chevrolat 1863: 146 203; Gundlach 1891: 41 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: SoledadHa: Playa Cuba localities. IJ: Arroyo la Talega. M: Marianao (t.l.). SC: Siboney Cárdenas; Cienaga Zapata Collections. Guignot (type). Washington Collections. Cambridge H. cubensis Chapin 1930: 10; Spangler 1981: 167 G. darlingtoni Ochs 1938: 90; Spangler 1981: 168 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. Gu: Sierra Imias (1-1300 m) Collections. Washington (type) (t.l.); upper Ovando River; SC: Cobre H. havanensis Wehncke 1880: 74; Spangler 1981: Range 167. There is a note on specimens in Gainesville Collections. Cambridge (type) and Cambridge by F. Young; to the effect that H. havaniensis is a senior synonym of H. Gyrinus Geoffroy 1862: 193 confluentus Roberts [USA:SC to FL & Keys] G. elevatus LeConte 1868: 368; Spangler 1981: 147 Distibution. Cuba, USA (FL) G. cubensis Régimbart 1883: 151; synonomy by Cuba localities. Spangler 1981: 147 Collections. Distribution. Cuba, USA (TX-FL) H. nanus Guignot 1936: 115; Spangler 1981: 167 Cuba localities. IJ: Laguna Base; Represa Distribution. Cuba. Endemic La Fé Cuba localities. Collections. Washington Collections. Guignot (type) G. rugifer Régimbart 1883: 179; Spangler 1973: 353 H. tumidus LeConte 1880: 166; Spangler 1981: 146 ? G. lateralis Aubé, Gundlach 1891: 42 Distribution. Cuba, Bahamas (Cat. Island); Distribution. Cuba, Jamaica, Hispaniola, USA (TX) Puerto Rico, Guadeloupe Cuba localities. Ho: Arroyo Jarahueca; Cuba localities. Ci: Soledad.; San Blas. LV: Mayari Arriba Arroyo Vegas Grandes. SS: Buenos Aires. . VC: Collections. Washington Mataguá Collections. Cambridge. Washington Peltodytes Régimbart 1878: 450 P. darlingtoni Young 1961: 221; Spangler 1981: REFERENCES 167 Spangler, P.J. 1973. Aquatic Coleoptera collected Distribution. Cuba. Endemic by the Biospeological expeditions. Resultats des Cuba localities. PR: Candelaria (t.l.) expeditions biospéologique Cubano-Roumaines Collections. Cambridge (type) à Cuba 1: 353-358. Spangler, P.J. 1981. Supplement to the aquatic REFERENCE and semiaquatic Coleoptera of Cuba. Resultats Spangler, P.J. 1981. Supplement to the aquatic des expeditions biospéologiques Cubano- and semiaquatic Coleoptera of Cuba. Resultats Roumaines à Cuba 3: 145-171. des expeditions biospéologiques Cubano- Roumaines à Cuba 3: 145-171. 12. FAMILY HALIPLIDAE The crawling water beetles 13. FAMILY Haliplid beetles live among aquatic vegetation These beetles live in the Holarctic region and at the edges of ponds, lakes, and streams. The southern South America. adults are slow moving, and the larvae feed by suck- ing on algal cells. 14. FAMILY NOTERIDAE The burrowing water beetles A Checklist of the Beetles of Cuba 43

This family inhabits shallow margins of stag- Distribution. Cuba, Hispaniola, Bahamas nant or slow-running pools or streams. The larvae (Cat Island), USA (SC-FL-LA) live in mud and feed on algae. Cuba localities. M: Playa Larga; Palpite Collections. Washington SUBFAMILY NOTERINAE Suphisellus Crotch 1873: 187 Say 1825: 105 S. binotatus (Fleutiaux & Sallé) 1889: 398; H. advena Sharp 1882: 281; Young, 1985: 96 Spangler 1981: 149 H. iricolor Say 1825: 105; Spangler 1981: 148 Distribution. Cuba, Hispaniola, Guadeloupe Distribution. Cuba, Hispaniola, Puerto Rico, Cuba localities. Ci: Soledad. IJ: Laguna Guadaloupe, widespread eastern USA. Collections. Washington Cuba localities. Gu: Arroyo Landivar at S. insularis Sharp 1882: 270; Spangler 1981: 167 Finca Pozon. IJ: Laguna, Represa La Fé. PR: Distribution. Cuba, Hispaniola, Puerto Rico, Viñales México, USA (FL) Collections. Washington Cuba localities. Ci: Soledad H. oblongus Sharp 1882: 280; Spangler 1981: 167; Collections. Young 1985: 97 S. nigrinus (Aubé) 1838: 411; Spangler 1981: 149 Distribution. Cuba, Bahamas, USA (FL) Distribution. Cuba, Guadeloupe, Colombia, Cuba localities. Ci: Soledad. PR: Candelaria Brazil, Argentina, Paraguay Collections. Cambridge Cuba localities. Gu: Arroyo Landivar at Finca Pozon. IJ: Laguna Mesonoterus Sharp 1882: 4 Collections. Washington M. laevicollis Sharp 1882a: 4; Spangler 1981: 148 S. tenuicornis (Chevrolat) 1863: 199 Distribution. Cuba, Hispaniola, Puerto Rico, (Hydrocanthus); Spangler 1981: 149 Virgin Islands, México to Panamá, Ecuador, Distribution. Cuba. Endemic Brazil Cuba localities. Gu: Arroyo Landivar at Cuba localities. Ci: Soledad. Gu: Arroyo Finca Pozon Landivar at Finca Pozon. IJ: Laguna; Represa Collections. Washington La Fé Collections. Washington REFERENCES Spangler, P.J. 1981. Supplement to the aquatic Sharp 1882: 260 and semiaquatic Coleoptera of Cuba. Resultats N. sharpi Balfour-Browne 1939: 98; Spangler 1981: des expeditions biospéologiques Cubano- 148 (replacement name for N. politus Sharp.) Roumaines à Cuba 3: 145-171. Distribution. Cuba, Bahamas, Jamaica, Young, F. 1985. A key to the American species of Puerto Rico, , Virgin Is- Hydrocanthus Say, with descriptions of new lands, México to Costa Rica, Trinidad, Ecua- taxa (Coleoptera: Noteridae). Proc. Acad. Nat. dor, USA (FL) Sci. Philadelphia 137: 90-98. Cuba localities. Ca: Estacion Experimental Club Azucarero.Gu: Arroyo Jarahueca at 15. FAMILY AMPHIZOIDAE Mayari Arribab; Rio Ceiba. IJ: Laguna Base. These beetles occur in northwestern North PR: Quemado de Pineda America, and Tibet. They crawl on submerged Collections. Washington rocks, logs, and vegetation, and are presumed predators. Sharp 1882: 263 P. addendus (Blatchley) 1920: 261; Spangler 1981: 16. FAMILY HYGROBIIDAE 167 These occur in Europe, China, and Australia. Distribution. Cuba, USA (FL) They are predacious bottom dwellers in muddy stag- Cuba localities. Ci: Soledad nant water. Collections. Cambridge (labeled P. punctipennis Sharp, F. Young) 17. FAMILY DYTISCIDAE The predacious diving beetles Aubé 1836: 209 This family of predaceous water beetles lives S. inflatus (LeConte) 1863: 22; Young 1954: 126 in a variety of aquatic habitats, especially lakes 44 Arthropods of Florida and Neighboring Land Areas, Vol. 18

and ponds. Both adults and larvae are good swim- C. posticatus (Fabricius) 1801: 268; Chevrolat 1863: mers and active predators. 200; Gundlach 1891: 36; Spangler 1981: 153 Distribution. Cuba, Jamaica, Hispaniola, SUBFAMILY Puerto Rico, Lesser Antilles, Central and South America Copelatus Erichson 1832: 18 Cuba localities. Ci: Soledad. Gu: Rio Baracoa [C. anastomosans Guignot 1952: 30. (Rev. Fr. Ent. at Tinta; upper Ovando River (300-600 m). PR: 19: 17-31)] Entronque de Manuel Sanguili; Hoyo de Fania; Distribution. Brazil (t.l.). Cuba record needs Quemado de Pineda. SS: Buenos Aires (1000 verification m) Cuba localities. No locality data with descrip- Collections. Washington tion Collections. SUBFAMILY C. barbouri Young 1942: 88; Spangler 1981: 167 Distribution. Cuba. Endemic Laccodytes Régimbart 1895: 345 Cuba localities. Gu: Mts. n of Imias (3-4000') L. pumilio (LeConte) 1878: 596; Spangler 1981: 150 (t.l.) Distribution. Cuba, Hispaniola, USA (FL) Collections. Cambridge (type) Cuba localities. IJ: Laguna. CA: Baraguá C. caelatipennis angustatus Chevrolat 1863: 201; Collections. Washington Young 1963: 62, 73; Spangler 1981: 153 C. angustatus Chevrolat; Gundlach 1891: 37 Laccophilus Leach 1818: 69 (placed as a subspecies by Young, 1963) L. bifasciatus Chevrolat 1863: 200; Gundlach 1891: Distribution. Cuba; other subspecies in Ba- 35; Spangler 1973: 354; 1981: 150 hamas, Central and South America, USA Distribution. Cuba, Hispaniola Cuba localities. Ci: Soledad. Gu: Matias Cuba localities. Gu: Cabezadas de Rio Indio; Collections. Washington Arroyo de la Poa at Sabonilla; Arroyo de la C. cubaensis Schaeffer 1908: 18; Young 1963: 75 Idalia, Gran Piedra.; Yunque de Baracoa (300- Distribution. Cuba, USA (FL) 600 m); upper Ovando River. SC: Pico Cuba localities. Ci: Cayamas (t.l.). Soledad Turquino. (3000 m); Loma del Gato (1000 m) Collections. Washington (type) Collections. Washington C. darlingtoni Young 1942: 87; Young 1942: 87; L. gentilis suavis Sharp 1882: 13; Zimmerman Spangler 1981: 167 1970: 229; Spangler 1981: 150 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola, México to Cuba localities. PR: Rangel Mts. (t.l.), 1500' Panamá, Bahamas. L. g. gentilis LeConte is Collections. Cambridge (type). Washington in USA (FL) C. glyphicus Say 1825a: 99; Young 1963: 62; Cuba localities. CA: Baraguá. Ci: Cayamas. Spangler 1981: 167 Gu: Arroyo de los Berros; Laguna la Chanco C. decemstriatus Aubé 1838: 374 Redondo Distribution. Cuba, Guadeloupe, Central Collections. Washington America, USA (CA-NY-FL) L. ovatus zapotecus Zimmermann 1970: 231; Cuba localities. Spangler 1981: 150 Collections. Distribution. Cuba, México to Panamá C. insolitus Chevrolat 1863: 201; Gundlach 1891: (nominate subspecies in SA) 37; Young 1942: 83; Spangler 1981: 153 Cuba localities. IJ: Laguna; Represa de la Distribution. Cuba, Jamaica, Hispaniola Fé. PR: Hoyo de Fania Cuba localities. Gr: Bayamo. SC: Cabezada Collections. Washington de Rio Baconao; Loma del Gato; Gran Piedra, L. proximus Say 1823: 101; Chevrolat 1863: 149; 2000-3000' Gundlach 1891: 35; Zimmerman 1970: 91; Collections. Washington Spangler 1973: 353; 1981: 149 C. montivagus Young 1942: 89; Spangler 1981: 167 L. americanus Aubé; Jacquelin du Val 1856: 27 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Jamaica, Cuba localities. Gu: upper Ovando River Hispaniola, Puerto Rico, Virgin Islands, Bar- (300-600 m). SC: Pico Turquino (500 m) bados, México, USA (FL and widespread to Collections. Cambridge. Washington Canada) A Checklist of the Beetles of Cuba 45

Cuba localities. Widespread Distribution. Cuba, USA (AL, NC, FL) Collections. Washington Cuba localities. Ci: Soledad (the record is L. quadrivittatus Aubé 1838: 428 based on 1 female) Distribution. Cuba, Jamaica, Hispaniola Collections. Cambridge Cuba localities. Ci: Soledad. Gu: Baracoa C. slossoni Mutchler 1918: 85; Young 1979: 823 Collections. Gainesville Distribution. Cuba, Mexico, USA (FL-AR- L. vacaensis Young 1953: 31; Zimmerman 1970: MD) 158 Cuba localities. Distribution. Cuba, Bahamas, Hispaniola, Collections. Jamaica, México, Guatemala, USA (AZ-TX-FL) Cuba localities. SC: Vista Alegre TRIBE HYDROVATINI Collections. Washington L. venustus Chevrolat 1863: 200; Gundlach 1891: Motschulsky 1855: 82 35; Spangler 1973: 354; 1981: 150 H. horni Crotch 1873: 378; Spangler and Vega 1982: Distribution. Cuba. Endemic 37 Cuba localities. Ci: Soledad; San Blas. LV: Distribution. Cuba, México, Guatemala, Arroyo Vegas Grandes; Rio Caburny. PR: USA (TX) Quemado de Pineda; Sierra de Rangel. Gu: Cuba localities. PR: Viñales Cueva Jibera; Pozo Caliente; Sierra de Imias; Collections. Havana. Washington upper Ovando River. SC: Daiquiri. SS: Buenos H. pustulatus compressus Sharp 1882: 324; Aires. VC: Hanabanillo Falls Spangler 1981: 151 Collections. Washington Distribution. Cuba, Dominica, USA (FL) Cuba localities. SC: Laguna de Juragua SUBFAMILY HYDROPORINAE Collections. Washington TRIBE VATELLINI TRIBE HYPHYDRINI Sharp 1882: 286 D. lentus (Wehncke) 1876: 357; Spangler 1981: 151 Babington 1841: 18. See Young Distribution. Cuba, Hispaniola, Jamaica, 1980 for subgenera. Trinidad, South America, USA (FL) D. aspera Young 1981: 4 Cuba localities. CA: Cueva de los Lagos Distribution. Cuba, Bahamas, USA (AL?, Collections. Washington FL) Cuba localities. CA: Baraguá. Ci: Soledad TRIBE METHLINI Collections. Cambridge. Gainesville. New York. San Francisco. Washington Celina Aubé 1836: 219 [“D. cuspidatus Kunze”: Gundlach 1891: 34; in la- C. angustata Aubé 1838: 447; Young 1979: 823 goons at Cárdenas and Trinidad; Gundlach no. Distribution. Cuba, USA (FL) 567; species identity not known] Cuba localities. D. darlingtoni Young 1989: 307 Collections. Distribution. Cuba, Jamaica, Hispaniola, C. cubensis Guignot 1947: 222; Spangler 1981: 167 Colombia Distribution. Cuba. Endemic Cuba localities. Gu: Baracoa Cuba localities. Ha: Rio Almendares. (t.l.) Collections. Cambridge Collections. Guignot (type). Washington D. glabella Young 1981: 7 C. grossula (LeConte) 1863: 22; Young 1979: 821; Distribution. Cuba. Endemic Spangler 1981: 167 Cuba localities. IJ: Laguna Base (t.l.) Distribution. Cuba, USA (FL) Collections. Washington (type) Cuba localities. Ci: Soledad (D. striola Sharp 1887: 752; Young 1990b: 226.) Collections. Distribution. Widespread South and Central C. imitatrix Young 1979: 824 America, and USA (FL including Keys) Distribution. Cuba, USA (TX-IN-VA-FL) Cuba localities. Cuba localities. Ci: Soledad. CA: Baraguá Collections Collections. Cambridge. Washington D. tarda Spangler 1973: 354; Spangler 1981: 167 C. palustris Young 1979: 830 Distribution. Cuba. Endemic 46 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. SC: Cabezadas de Rio Indio; Guignot 1939: 53 Gran Piedra (t.l.) L. affinis (Say) 1823: 104; Spangler 1981: 152 Collections. Cambridge. Gainesville. Wash- Distribution. Cuba, Bahamas, North, Cen- ington (type) tral and South America Cuba localities. Ho: Pinares de Mayari Sharp 1882: 338 Collections. Washington P. obniger (Chevrolat) 1863: 199 (Hyphydrus); Gundlach 1891: 33; Spangler 1981: 168 Young 1967: 79 Distribution. Cuba, Bahamas, Puerto Rico, N. pullus floridanus (Fall) 1917: 168; Spangler USA (FL) 1981: 152 Cuba localities. Ci: Soledad (as P. princeps Distribution. Cuba, Hispaniola?, Jamaica?, Blatchley) Bahamas, USA ( FL). Notes: Nominate sub- Collections. Cambridge species from western FL to México Cuba localities. IJ: Laguna TRIBE BIDESSINI Collections. Washington

Anodocheilus Babington l838: 132 SUBFAMILY COLYMBETINAE A. exiguus (Aubé) 1838: 490; Young 1974: 10; TRIBE COLYMBETINI Spangler 1981: 152 Distribution. Cuba, USA (TX-FL) Rhantus Boisduval and Lacordaire 1835: 309 Cuba localities. Ho: Arroyo Landivar at R. calidus (Fabricius) 1792: 193 (Dytiscus); Finca Pozon. PR: Quemada de Pineda. SC: Chevrolat 1863: 200 (Colymbetes); Gundlach Laguna de Juragua 1891: 36; Spangler 1981: 168 Collections. Washington Distribution. Cuba, Bahamas, Jamaica, Hispaniola, México, Central and South Bidessonotus Régimbart 1895: 331 America, Lesser Antilles, USA (NY-TX-FL) B. browneanus Balfour-Browne 1947: 441; Spangler Cuba localities. Gu: upper Ovando River. M: 1981: 152; Young 1990a: 376 Cárdenas; Cienaga de Zapata. PR: Candelaria. Distribution. Cuba, Jamaica, Hispaniola, SS: Buenos Aires Puerto Rico Collections. Cambridge Cuba localities. Widespread Collections. Washington SUBFAMILY B. caraibus (Chevrolat) 1863: 199 (Hydroporus); TRIBE HYDATICINI Gundlach 1891: 34; Spangler 1981: 167; Young 1990: 360 (indicating possible conspecificity Hydaticus Leach 1817: 69 with B. pulicarius (Aubé)) H. bimarginatus (Say) 1834: 442 Distribution. Cuba, Belize, USA (FL?) Distribution. Cuba, Bahamas, USA (TX-AR- Cuba localities. Ci: Cayamas. M: Zapata NY-FL) Swamp Cuba localities. Ci: Soledad. CA: Baraguá. Collections. Washington Gr: Cauto El Cristo B. fallax Balfour-Browne 1947: 429; Young 1990: Collections. Cambridge 373 H. riehli Wehncke 1876: 195; Spangler 1981: 167 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Zapata Swamp. PR: Cuba localities. Pinar del Rio. Ha: Havana. Ci: Cayamas (t.l.) Collections. Collections. Washington (type) H. rimosus Aubé 1838: 182; Chevrolat 1863: 202; Gundlach 1891: 40; Spangler 1981: 167 Zimmermann 1919: 134 Distribution. Cuba, México, Hispaniola, Cen- B. apicatus (Clark) 1862: 184; Spangler 1981: 152 tral America, Guadeloupe Distribution. Cuba, Hispaniola, México, Cen- Cuba localities. CA: Baraguá. Gr: Cauto El tral America, USA (FL) Cristo. Gu: upper Ovando River. M: Cárdenas. Cuba localities. PR: Rangel Mts. SS: Buenos Aires Collections. Collections. Cambridge A Checklist of the Beetles of Cuba 47

TRIBE Cybister lherminieri Guérin, Jacquelin du Val 1856: 25; Chevrolat 1863: 201; Gundlach 1891: Dejean 1833: 53 37 T. basillaris (Harris) 1829: 1; Spangler 1981: 154 Distribution. Cuba, Bahamas, Hispaniola, incisus Aubé, Gundlach 1891: 39 Puerto Rico, México to Argentina, Lesser Distribution. Cuba, Bahamas, Jamaica, Antilles Hispaniola, Puerto Rico, México, Guatemala, Cuba localities. Ci: Soledad Lesser Antilles, South America, USA (CA- OK- Collections. Washington NY-FL) Cuba localities. Ci: Soledad. Ha: Havana. IJ: REFERENCES Laguna. M: Cárdenas; Cienaga de Zapata. PR: Note: The extensive bibliography of Spangler (1981) Candelaria is not repeated here and should be consulted Collections. Washington for additional sources. T. circumscriptus (Latreille) 1809: 223 (Acilius); Spangler, P.J. 1973. Aquatic Coleoptera collected Jacquelin du Val 1856: 27; Chevrolat 1863: 202; by the biospeleological expeditions to Cuba by Gundlach 1891: 38; Spangler 1981: 154 the Academies of Science of Cuba and Roma- Hydaticus insularis Laporte 1834: 96 nia. Résultats des expéditions biospéologiques Hydaticus havaniensis Laporte 1834: 96 Cubano-Roumaines à Cuba 1: 353-358. Distribution. Cuba, Bahamas, Jamaica, Spangler, P.J. 1981. Supplement to the aquatic Puerto Rico, Lesser Antilles, México to Brazil and semiaquatic Coleoptera of Cuba. Résultats Cuba localities. Widespread des expéditions biospéologiques Cubano- Collections. Washington Roumaines à Cuba 3: 145-171. T. margineguttatus Aubé 1838: 149; Gundlach Spangler, P.J. and A. Vega. 1982. First record 1891: 39; Spangler 1981: 154 of Hydrovatus horni Crotch from the Antilles Distribution. Cuba, Hispaniola, Puerto Rico, with notes on its known distribution and sta- México to Brazil, Lesser Antilles, South tus (Coleoptera: Dytiscidae). Ent. News 93: 37- America 41. Cuba localities. CA: Baraguá. Ci: Soledad. Young, F.N. 1974. Review of the predaceous water Gr: Cauto El Cristo. .M: Cárdenas; Cienaga de beetles of genus (Coleoptera: Zapata Dytiscidae: Hydroporinae). Occ. Pap. Mus. Zool. Collections. Washington Univ. Michigan 670, 28 pp. Young, F. 1979. A key to the Nearctic species of TRIBE CYBISTRINI Celina with descriptions of new species (Co- leoptera, Dytiscidae). J. Kansas Ent. Soc. 52: Cybister Curtis 1827: 151 820-830. C. occidentalis Aubé 1838: 67; Jacquelin du Val Young, F. 1980. Predaceous water beetles of the 1856: 26; Chevrolat 1863: 202; Gundlach 1891: genus Desmopachria Babington: the subgen- 38; Spangler 1981: 154 era with descriptions of new taxa (Coleoptera: C. laevigatus Fabricius, Gundlach 1891: 38. Dytiscidae). Rev. Biol Trop. 28: 305-321. Distribution. Cuba, Bahamas, USA (FL) Young, F.N. 1981. Predaceous water beetles of the Cuba localities. Ci: Soledad Ha: Havana. SS: genus Desmopachria: the convexa-grana group Buenos Aires; Trinidad Mts (Coleoptera: Dytiscidae). Occ. Pap. Florida St. Collections. Washington Colln. Arthrop. 2: 1-9. Young, F. N. 1989. A new species of Desmopachria Megadytes Sharp 1882: 704 (s.str.) from the Antillean-Caribbean region M. fraternus Sharp 1882b: 708; Spangler 1981: 154 (Coleoptera: Dytiscidae). Psyche 96: 307-309. Distribution. Cuba, Bahamas, Jamaica, Young, F. N. 1990a. A review of classification of Hispaniola, Puerto Rico, Guatemala, Panamá, the water beetles of the New World genus Lesser Antilles, USA (FL) Bidessonotus Régimbart (Coleoptera: Cuba localities. IJ: La Fé Dytiscidae: Hydroporinae: Bidessini). Quaest. Collections. Washington Ent. 26: 355-381. M. gigantea Laporte 1835: 99; Spangler 1981: 154 Young, F. N. 1990b. Predaceous water beetles of the genus Desmopachria Babington: the sub- 48 Arthropods of Florida and Neighboring Land Areas, Vol. 18

genus Pachriostrix Guignot (Coleoptera: Collections. Washington Dytiscidae). Coleop. Bull. 44: 224-228. B. exigus (Say) 1825: 189; Van Tassell 1966: 149 Zimmerman, J. R. 1970. A taxonomic revision of Distribution. Cuba, Bahamas, widespread the beetle genus Laccophilus (Dytiscidae) of USA North America. Mem. Amer. Ent. Soc. 26: 1- Cuba localities. Gu: Guantanamo. PR: Pinar 275. del Rio Collections. New York? SUBORDER POLYPHAGA B. peregrinus (Herbst) 1797: 314; Van Tassell 1966: SERIES STAPHYLINIFORMIA 163. SUPERFAMILY HYDROPHILOIDEA Distribution. Cuba, widespread USA Cuba localities. PR: Pinar del Rio 18. FAMILY HYDROPHILIDAE Collections. New York? The water scavenger beetles B. quadridens Chevrolat 1863: 206; Gundlach 1891: These occur either in water or in moist habi- 47; Spangler 1981: 156 tats. The larvae are usually predators while the Distribution. Cuba, México, Central America adults feed on decomposing plant matter. Cuba localities. Ci: Soledad. Ho: Arroyo Landivar at Finca Pozon. IJ: Laguna. M: SUBFAMILY HYDROCHINAE Cárdenas Collections. New York, Washington Leach 1817: 190 B. stribalus d’Orchymont 1946: 13; Spangler 1981: H. naraini Makhan 1993: 67 156 Distribution. Cuba. Endemic Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. “Cuba.” A specimen previ- Puerto Rico ously labeled as H. pallipes Chevrolat Cuba localities. Ho: Arroyo Landivar at Collections. Budapest (type) Finca Pozon; Matias. IJ: Laguna. PR: Manuel H. pallipes Chevrolat 1863: 207; Gundlach 1891: Sanguili 49; Spangler 1981: 155 Collections. Washington Distribution. Cuba, Hispaniola, Puerto Rico B. trilobus Chevrolat 1863: 207; Gundlach 1891: Cuba localities. IJ: Represa La Fé 47; Spangler 1973: 354; 1981: 156 Collections. Washington Distribution. Cuba. Endemic H. Mulsant 1844: 373; Spangler 1981: 168 Cuba localities. Widespread Distribution. Cuba, USA (MA-FL) Collections. Washington Cuba localities. B. undatus (Fabricius) 1792: 185; Spangler 1981: Collections. Washington 156 H. tarsalis Chevrolat 1863: 207; Gundlach 1891: Distribution. Cuba, México 48; Spangler 1981: 155 Cuba localities. Ho: La Aguada. SC: Laguna Distribution. Cuba, Colombia? de Juroqua; Siboney Cuba localities. Ci: Soledad. IJ: La Fé; La- Collections. Washington guna. M: Cárdenas. Gr: Cauto El Cristo. Ho: Arroyo Landivar at Finca Pozon; El Cayo at Derallus Sharp 1882: 77 Boire. PR: Candelaria; Quemado de Pineda. SC: D. altus (LeConte) 1855: 366; Van Tassell 1966: Arroyo Colorado 282 Collections. Washington Distribution. Cuba, Dominican Republic, Puerto Rico, USA (TX-NJ-FL) SUBFAMILY Cuba localities. PR: Pinar del Rio TRIBE BEROSINI Collections. New York? D. rudis Sharp 1887: 765; Spangler 1981: 168 Berosus Leach 1817: 92 Distribution. Cuba, México, Brazil, Argen- B. chevrolati Zaitzev 1908: 358 (replacement name tina, Guadeloupe for B. aculeatus Chevrolat); Spangler 1981: 168 Cuba localities. B. aculeatus Chevrolat 1863: 207 (not LeConte) Collections. Distribution. Cuba. Endemic Cuba localities. Widespread. See Spangler 1981: 155 for localities A Checklist of the Beetles of Cuba 49

TRIBE ANACAENINI TRIBE LACCOBIINI

Anacaena Thomson 1859: 18 Laccobius Erichson 1837: 202 A. debilis (Sharp) 1882: 66; Spangler 1973: 356; H. (Laccophilus) antillensis Spangler 1968: 752; 1981: 160 1981: 160 Distribution. Cuba, México to Argentina Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Gu: Arroyo de la Poa at Puerto Rico Sabonilla; Baracoa; El Cayo at Baire; Rio Cuba localities. Ci: Soledad. SC: Rio Contramaestra at Pozo Caliente. Ho: Pinares Contramaestra at Pozo Caliente; Rio Mogote at de Mayari. IJ: Arroyo Talega. LV: Arroyo Ve- Matias gas Grandes Collections. Washington Collections. Washington TRIBE HYDROPHILINI Paracymus Thomson 1867: 119 (see key to SUBTRIBE ACIDOCERINA West Indian species in Wooldridge 1971) P. acutipenis Wooldridge 1971: 400; Spangler 1973: Enochrus Hope 1859: 18 356; 1981: 160 E. aequalis (Sharp) 1882: 68; Spangler 1973: 355; Distribution. Cuba. Endemic 1981: 157 Cuba localities. Type locality not specified, Distribution. Cuba, Lesser Antilles, México possibly Lago Aquianaba. IJ: La Fé, Laguna to Nicaragua Collections. Cambridge (type). Washington Cuba localities. Widespread P. confusus Wooldridge 1966: 719; Spangler 1981: Collections. Washington 158 E. ochraceus (Melsheimer) 1844: 101; Spangler Distribution. Cuba, Bahamas, México, USA 1981: 157 (SC-FL) E. coriaceus (Chevrolat) 1863: 206 (Phylhydrus); Cuba localities. IJ: Laguna. Ho: Arroyo Gundlach 1891: 47 Landivar at Finca Pozon. SC: Arroyo Colorado; Distribution. Cuba, Bahamas, Jamaica, Arroyo de los Berros, Laguna Charco Redondo; Hispaniola, México, Central America, wide- El Cayo at Baire; Loma da Bandeca, Mayari spread USA. Abajo Cuba localities. IJ: La Fé; Laguna Collections. Washington Collections. “Cuba” in Cambridge. Washing- P. delatus Wooldridge 1971: 401; Spangler 1981: ton 159 E. pseudochraceus Gunderson 1977: 256; Spangler Distribution. Cuba, Puerto Rico, Virgin Is- 1981: 157 lands, Dominica Distribution. Cuba, widespread West Indies, Cuba localities. SC: Mayari Abajo; Matias; México, Central America to west coast of South Pinares de Mayari America Collections. Washington Cuba localities. IJ: Laguna. Ho: Arroyo P. indigens Wooldridge 1969: 419; Spangler 1981: Jarahueca at Mayari Arriba; Arroyo Landicar 159 at Finca Pozon. PR: Quemado de Pineda Distribution. Cuba, México to Costa Rica Collections. Washington Cuba localities. Ha: Laguna Ariguanabo. Ho: E. pygmaeus (Fabricius) 1792: 186; Spangler 1981: Arroyo Landivar. Gu: Baracoa at Tinta, Rio 157 San Juan. IJ: La Fé, Laguna. PR: Quemado de Distribution. Cuba, Bahamas, México, USA Pineda (CA-FL) Collections. Washngton Cuba localities. Ho: Arroyo Jarahueca at P. lodingi (Fall) 1910: 98; Spangler 1981: 159 Mayari Arriba; Rio Contramaestra; Rio Mogote Distribution. Cuba, Bahamas, México, USA at Matias (AL-FL) Collections. Washington Cuba localities. Ha: Laguna de Ariguanabo. E. sagrae Knirsh 1924: 214 (replacement name for IJ: La Fé, Laguna. PR: Quemado de Pineda Philhydrus melanocephalus Olivier); Spangler Collections. Washington 1981: 168 Philhydrus melanocephalus Olivier 1792: 121, Jacquelin du Val 1856: 53; Gundlach 1891: 46 50 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba. Endemic Tropisternus Solier 1834: 302 Cuba localities. T. apicipalpis Chevrolat 1834: 54; Spangler 1981: Collections. 168 Distribution. Cuba, México to Brazil, Lesser Helobata Bergroth 1882: 221 Antilles. H. striata Brullé 1841: 58; Spangler 1981: 168 Cuba localities. Distribution. Cuba, Bahamas, México to Bra- Collections. zil, se USA T. chalybeus Laporte 1840: 53; Spangler 1981: 168 Cuba localities. T. blandus Chevrolat 1863: 205; Gundlach 1891: Collections. 45 Distribution. Cuba, Puerto Rico, México to Helochares Mulsant 1844: 197 Brazil, Lesser Antilles H. abbreviata (Fabricius) 1801: 251; Spangler 1981: Cuba localities. M: Cárdenas 158 Collections. Distribution. Cuba, Virgin Islands, Lesser T. collaris (Fabricius) 1775: 229; Jacquelin du Val Antilles, México to Argentina 1856: 53; Chevrolat 1863: 205; Gundlach 1891: Cuba localities. IJ: Laguna 45; Spangler 1981: 161 Collections. Washington T. collaris proximus Sharp 1883: 97; Spangler 1981: 161. Cuban endemic subspecies SUBTRIBE HYDROPHILINA Distribution. Cuba, Hispaniola, Puerto Rico, México to Brazil Hydrobiomorpha Blackburn 1888: 814 Cuba localities. SC: Arroyo Colorado; Laguna H. media (Brullé) 1837: 54 (Hydrous); Spangler de Juragua 1981: 168 Collections. Hydrous tenebroides Jacquelin du Val 1856: T. lateralis (Fabricius) 1775: 228; Jacquelin du Val 50; Gundlach 1891: 46 1856: 52: Chevrolat 1863: 205; Gundlach 1891: Distribution. Cuba. Endemic 45; Spangler 1981: 160 Cuba localities. Distribution. Cuba, Bahamas, Hispaniola, Collections. Jamaica, Virgin Islands, Monserrat, México to Argentina, widespread e USA. Two subspecies Hydrophilus Geoffroy 1862: 180 are named: T. l. lateralis and T. l. nimbatis H. (Dibolocelus) smaragdinus Brullé 1837: 54; (Say) Spangler 1981: 168 (as Dibolocelus) Cuba localities. Widespread. See Spangler Distribution. Cuba, México, South America 1981: 160 for localities Cuba localities. Collections. Washington Collections. T. mergus (Say) 1835: 171; Spangler 1981: 161 H. (H.) ater Olivier 1792: 125 Distribution. Cuba, México to Argentina H. a. intermedius Jacquelin du Val 1856: 48; Cuba localities. Gu: Baracoa at Tinta. PR: Chevrolat 1863: 204; Gundlach 1891: 43; Hoyo de Fania; Quemado de Pineda Spangler1981: 168 Collections. Washington H. picicornis Chevrolat, Gundlach 1891: 44 (T. quadristriatus Horn 1871: 33l) H. violaceo-nitans Jacquelin du Val 1856: 49; Distribution. Cuba, Grand Cayman, Baha- Chevrolat 1863: 205; Gundlach 1891: 44 mas, México, USA (MA-FL) Distribution. Cuba, Hispaniola, Puerto Rico, Cuba localities. A brackish water species St. Lucia Collections. Washington Cuba localities. Widespread Collections. Carnegie SUBFAMILY SPHAERIDIINAE H. (H.) insularis Laporte 1840: 50; Jacquelin du Val 1856: 47; Chevrolat 1863: 204; Gundlach These beetles live in terestrial habitats, usu- 1891: 43; Spangler 1981: 168 ally in very wet decaying organic matter. Smetana Distribution. Cuba, Bahamas, Puerto Rico, (1978, 1984) revised the North American fauna. Central America, USA (FL) The genera Pelosoma (Spheridiini), Omicrus and Cuba localities. Aculomicrus (Omicrini) occur in Florida and/or the Collections. A Checklist of the Beetles of Cuba 51

West Indies and are to be expected in Cuba Distribution. Cuba, Jamaica, Bahamas, (Smetana 1975, 1984). Hispaniola, West Indies, México, widespread USA TRIBE COELOSTOMATINI Cuba localities. Ha: Laguna de Ariguanabo Collections. Washington Dactylosternum Wollaston 1854: 99 (C. variegatus Sharp 1882: 107) D. abdominale (Fabricius) 1792: 99; Spangler 1981: Distribution. Jamaica, Puerto Rico, México, 168 Central America, South America, USA (TX, Distribution. Cuba, Bahamas, Hispaniola, NC-FL) Jamaica, México, Central America, s USA (CA- Cuba localities. FL) Collections. Cuba localities. IJ: Nueva Gerona. Probably widespread in moist decaying plant matter. Oosternum Sharp 1882: 112 Collections. Carnegie O. costatum Sharp 1882: 113; Spangler 1981: 168 D. flavicorne (Mulsant) 1844: 174 (Cyclonotum); Distribution. Cuba, Puerto Rico, México to Jacquelin du Val 1856: 53; Chevrolat 1863: 208; Panamá, USA (widespread SE) Gundlach 1891: 49; Spangler 1981: 168 Cuba localities. Ci: Soledad. SC: Cueva de Distribution. Cuba, Jamaica, Hispaniola, los Majaes, in moist guano Puerto Rico Collections. Gatineau. Cuba localities. Collections. REFERENCES D. picicorne Mulsant 1844: 175; Spangler 1981: 168 The extensive bibliography of Spangler (l981) is not Distribution. Cuba, Jamaica, Puerto Rico repeated here. Cuba localities. Makhan, D. 1993. Nine new Hydrochus species Collections. from South and Central America (Coleoptera: D. subdepressum Laporte 1840: 58; Spangler 1981: Hydrophilidae). Ann. Hist.-Natur. Mus. Nat. 168 Hung. 85: 65-70. Distribution. Cuba, widespread tropical Smetana, A. 1975. Revision of the New World gen- America era of the tribe Omicrini trib. no. of the Cuba localities. hydrophilid subfamily Sphaeridiinae (Co- Collections. leoptera). Stud. Neotrop. Fauna 10: 153-182. Smetana, A. 1978. Revision of the subfamily Phaenonotum Sharp 1882: 97 Sphaeridiinae of America north of México (Co- P. exstriatum (Say) 1835: 171; Spangler 1981: 161 leoptera: Hydrophilidae)). Mem. Ent. Soc. Distribution. Cuba, Jamaica, Puerto Rico, Canada 105. Lesser Antilles, México to Brazil, USA (wide- Smetana, A. 1984. Revision of the subfamily spread) Sphaeridiinae of America north of México (Co- Cuba localities. SC: Pedrera de la Waren leoptera: Hydrophilidae)). Supplementum 2. Collections. Canadian Ent. 116: 555-566. (P. minor Smetana 1978: 20) Spangler, P.J. 1973. Aquatic Coleoptera collected Distribution. USA (AL-FL) by the biospeleological expeditions. Resultats Cuba localities. des expeditions biospéologique Cubano- Collections. Roumaines à Cuba 1: 353-358. Spangler, P.J. 1981. Supplement to the aquatic TRIBE MEGASTERNINI and semiaquatic Coleoptera of Cuba. Resultats des expeditions biospéologique Cubano- Cercyon Leach 1817: 95 Roumaines à Cuba 3: 145-171. C. insulare Chevrolat 1863: 208; Gundlach 1891: Van Tassell, E. 1966. Taxonomy and biology of 50; Spangler 1981: 168 the subfamily Berosinae of North and Central Distribution. Cuba. Endemic? American and the West Indies. PhD thesis, Cuba localities. Catholic University of America. University Collections. Microfilms 66-15,059. C. praetextus Say 1825: 190; Spangler 1981: 161 C. centrimaculatum Sturm, Gundlach 189l: 50 52 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Wooldridge, D.P. 1971. Two new Paracymus from Horn 1873: 356 the West Indies. J. Kansas Ent. Soc. 44: 400- A. (Aeletes) assimilis Wenzel 1944: 74 403. Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) 19. FAMILY SPHAERITIDAE Collections. Washington (type) This family occurs in , north- A. (A.) poeyi (Marseul) 1862: 695 (); western North America, and China. Adults may Chevrolat 1864: 598; Gundlach 1891: 83 come to birch sap, and larvae may be in soil soaked Distribution. Cuba. Endemic in birch sap. Cuba localities. M: Cárdenas Collections. 20. FAMILY SYNTELIDAE A. (A.) rugiceps Wenzel 1944: 68 This family occurs in México, Japan, , Distribution. Cuba. Endemic and . Adults and larvae are predators under Cuba localities. Ci: Cayamas (t.l.) rotted bark, or in rotted cactus. Collections. Washington (type) A. (A.) schwarzi Wenzel 1944: 67 21. FAMILY HISTERIDAE Distribution. Cuba. Endemic The clown beetles Cuba localities. Ci: Cayamas (t.l.) Reviewed by S. Mazur, March, 2002 Collections. Washington (type) These live in a variety of habitats and are preda- A. (Acritinus) floridae (Marseul) 1862: 693 (Acritus) tors as adults or larvae. They may be predators on Distribution. Cuba, USA (FL, LA, NC, SC) wood-boring beetles, on maggots in carrion and Cuba localities. dung, and some are myrmecophiles. Decou and Collections. Thérond (1977) give Cuban records of species col- lected in caves. Primary references: Mazur 1984 TRIBE BACANIINI and Johnson et al. 1991. The higher classification follows Mazur (2001), which contains a key for iden- Bacanius LeConte 1853: 291 tification to the genera of Mexico, and all Cuban B. (Bacanius) scalptus Lewis 1888: 237; Wenzel genera are included. 1944: 96 Distribution. Cuba, México to Argentina, SUBFAMILY ABREINAE USA (FL?) TRIBE Cuba localities. Ci: Cayamas (t.l) Collections. Washington (type) Acritus LeConte 1853: 288 B. (Gomyister) debilitans Casey 1893: 560 A. analis LeConte 1853: 290; Chevrolat 1864: 599; Distribution. Cuba, USA (FL) Gundlach 1891: 83; Decou and Thérond 1977: Cuba localities. 403 Collections. Distribution. Cuba, Puerto Rico, Guatemala, México SUBFAMILY SAPRININAE Cuba localities. Ha: Cueva Galera. PR: Cueva Incognita Euspilotus Lewis 1907: 320 Collections. Bucharest. Havana. Thérond E. (Euspilotus) cubaecola (Marseul) 1855: 467 A. atomus LeConte 1853: 291 (Saprinus); Jacquelin du Val 1856: 111; A. atomulus Marseul 1862: 694; Chevrolat 1864: Chevrolat 1864: 596; Gundlach 1891: 79 598; Gundlach 1891: 82; Wenzel 1944: 58 Distribution. Cuba, USA (FL) Distribution. Cuba, USA (LA, FL) Cuba localities. M: Cienaga de Zapata Cuba localities. Ha: Caisuita; Havana; Collections. Marianao. M: Cárdenas E. (Hesperosaprinus) arrogans (Marseul) 1855: 487 Collections. Cambridge (Saprinus) A. ignobilis (Lewis) 1888: 238; Wenzel 1944: 58 S. tarnieri Marseul 1862: 480; Chevrolat 1864: Distribution. Cuba, Puerto Rico, Panamá, 597; Gundlach 1891:80 Colombia, USA (FL) Distribution. Cuba, México, Guatemala, Ven- Cuba localities. Ha: Marianao ezuela, Galapagos Islands, USA (TX) Collections. Chicago Cuba localities. Collections. A Checklist of the Beetles of Cuba 53

E. sterquilinus (LeConte) 1860: 315 (Saprinus); C. (C.) troglodytes (Paykull) 1811: 46 (Hister); Chevrolat 1864: 597; Gundlach 1891: 81; Decou Jacquelin du Val 1856: 109; Chevrolat 1864: and Thérond 1977: 403; all as Saprinus 594; Gundlach 1891: 76; Decou and Thérond Distribution. Cuba, se USA? 1977: 403 Cuba localities. Ha: Havana. SC: Cueva de Distribution. Cuba, West Indies (where?); Cativar; Cueva del Humo; Cueva de la Patana; tropicopolitan Cueva el Mudo; Cueva de Guanaraya; Cueva Cuba localities. Gr: Cueva de Fustete. Ha: de la Virgen; Cueva de los Majes. PR: near Cueva el Mudo. Ho: Cueva Cuatrocientas Rozas. Cueva Clara. Abundant in caves in bat guano IJ: Nueva Gerona. M: Cueva La Eloisa. PR: Collections Gatineau. Bucharest. Havana. Hoyo de Fania. SC: Cueva de los Majaes. SS: Thérond Cueva de Guanayara. Abundant in bat caves Collections. Bucharest. Carnegie (as C. Saprinus Erichson 1834: 172 tenellus?). Havana. Thérond S. (Saprinus) cavalieri Marseul 1855: 462; Jacquelin du Val 1856: 110; Chevrolat 1864: 596; Paromalus Erichson 1834: 167 Gundlach 1891: 78 P. (Isolomalus) hispaniolae Marseul 1870: 101 Distribution. Cuba, Argentina P. nanulus (Casey) 1916: 246 (Isolomalus); Ha: Cuba localities. Widespread Havana (t.l.); Washington (type) Collections. Distribution. Cuba, Hispaniola S. (Saprinus) guayanensis Marseul 1855: 453; Cuba localities. Ha: Havana Chevrolat; 1864b: 596; Gundlach 1891: 79. Collections. Cambridge Distribution. Cuba, México, Brazil, Argen- P. (I.) parallelus LeConte 1860: 314 tina P. laterirectus Marseul 1862: 712; Chevrolat Cuba localities. 1864: 595; Gundlach 1891: 77 Collections. Isolomalus parallelus Bickhardt 1917: 120 Distribution. Cuba (t.l.), Antilles (where?) Xerosaprinus Wenzel 1962: 374. Cuba localities. X. fulgidus (LeConte) 1860: 316 (Saprinus); Collections. Cambridge (type) Chevrolat 1864: 598; Gundlach 1891: 82. P. (I.) productus Marseul 1855: 113; Jacquelin du Distribution. Cuba. Endemic Val 1856: 109; Chevrolat 1864: 595; Gundlach Cuba localities. 1891: 77 Collections. Distribution. Cuba, Colombia X. viator (Marseul) 1855: 499 (Saprinus); Jacquelin Cuba localities. du Val 1856: 112; Chevrolat 1864: 597; Collections. Gundlach 1891: 80 Distribution. Cuba. Endemic SUBFAMILY TRYPANAEINAE Cuba localities. Collections. Trypanaeus Eschscholtz 1829: 10 [T. breviculus Marseul 1857: 401] SUBFAMILY DENDROPHILINAE Distribution. Cuba(?), French Guyana TRIBE PAROMALINI Cuba localities. Doubtful record from Mazur 1984 Carcinops Marseul 1855: 83 T. flavipennis Marseul 1860: 841; Chevrolat 1858: C. (Carcinops) parvulus LeConte 1860: 314; 209 (Tripanidius) Chevrolat 1864: 594; Gundlach 1891: 76. T. pallidipennis Marseul 1860: 841; Chevrolat Distribution. Cuba, Antilles 1864: 591; Gundlach 1891: 71 Cuba localities. Distribution. Cuba, Panamá, Guatemala, Collections. Surinam (C. (C.) pumilio (Erichson) 1834: 169.) Cuba localities. Widespread Distribution. North America. Nearly cosmo- Collections. politan T. luteivestis Marseul 1860: 842 Cuba localities. Probably present but no Distribution. Cuba, México, Guatemala, records are known Guadeloupe Collections. Cuba localities. 54 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Cuba localities. Ci: Soledad. Gr: Bayamo Collections. Cambridge SUBFAMILY HISTERINAE TRIBE HOLOLEPTINI TRIBE HISTERINI

Hololepta Paykull 1811: 101 Atholus Thomson 1859: 76 H. (Hololepta) cubensis Erichson 1834: 93; A. confinis (Erichson) 1834: 154 (Hister); Jacquelin Jacquelin du Val 1856: 102; Chevrolat 1864: du Val 1856: 105; Chevrolat 1864: 592; 590; Gundlach 1891: 69 Gundlach 1891: 73 Distribution. Cuba, Haiti, Brazil? Distribution. Cuba, Puerto Rico, St. Vincent, Cuba localities. Ci: Soledad. M: Cárdenas Guadeloupe, USA (FL) Collections. Cuba localities. M: Cárdenas H. interrupta Marseul 1853: 214; Jacquelin du Val Collections. 1856: 103 Leionota ruptistria Marseul, Chevrolat 1864: 590; Hister Linnaeus 1758: 358 Gundlach 1891: 70 (Leioderma) H. coenosus Erichson 1834: 140; Jacquelin du Val Distribution. Cuba, Puerto Rico, USA (FL) 1856: 105; Chevrolat 1864: 593; Gundlach 1891: Cuba localities. Widespread. Ci: Soledad 74; Caterino 1999: 59 Collections. Cambridge Distribution. Cuba. Jamaica, Hispaniola, H. (Leionota) minuta Erichson 1834: 96 México to Panamá, USA (FL-VA-AR-CA) Leionota rimosa Marseul 1853: 218; Jacquelin Cuba localities. IJ: Nueva Gerona. M: du Val 1856: 103; Chevrolat 1864: 590; Cienaga de Zapata. PR Gundlach 1891: 70 (Leioderma) Collections. Carnegie Distribution. Cuba, Brazil, México, Panamá, H. servus Erichson 1834: 147; Jacquelin du Val French Guyana, USA (TX, FL) 1856: 106; Chevrolat 1864: 593; Gundlach 1891: Cuba localities. M: Cárdenas 74 Collections. Distribution. Cuba, México, Central America, USA (AZ-IN-MD-FL) TRIBE OMALODINI Cuba localities. Collections. Omalodes Erichson 1834: 114 O. laevigatus Quensel in Schönherr 1806: 90; SUBFAMILY ONTHOPHILINAE Jacquelin du Val 1856: 104; Chevrolat 1864: 491 Epierus Erichson 1834: 158 Distribution. Cuba, Central America, west- E. (Epierus) antillarum Marseul 1854: 700; ern North America Jacquelin du Val 1856: 107; Chevrolat 1864: Cuba localities. 594; Gundlach 1891: 75; Decou and Thérond Collections. 1977: 404 O. ruficlavis Marseul 1853: 531; Jacquelin du Val Epierus cubensis Casey 1916: 238 (Ci: Cayamas 1856: 104; Chevrolat 1864: 591; Gundlach 1891: (t.l.); type in Washington) 72 Distribution. Cuba, Hispaniola, Puerto Rico, O. klugii Marseul, Chevrolat 1864: 592; Gundlach México, USA (SE states) 1891:73 Cuba localities. Gu: Cueva de la Patana. Ho: Distribution. Cuba, Central America, Ven- Cueva Los Cuatrocientas Rozas; Rio ezuela Cuyaguateje; M: Cárdenas. Cuba localities. Collections. Bucharest. Havana. Thérond Collections. E. (E.) cylindricus Wenzel 1944: 118 Distribution. Cuba. Endemic TRIBE EXOSTERNINI Cuba localities. PR: Pinar del Rio (t.l.) Collections. Chicago (type) Phelister Marseul 1853: 462 E. pulicarius Erichson 1834: 162; Decou and P. riehli Marseul 1862: 697; Chevrolat 1864: 591; Therond 1977: 404 Gundlach 1891: 71 Distribution. Cuba, widespread eastern USA Distribution. Cuba. Endemic Cuba localities. SC: Cueva Atabex A Checklist of the Beetles of Cuba 55

Collections. Bucharest. Havana. Thérond Hydraena Kugelann 1794: 579 E. (E.) smaragdinus Marseul 1862: 698; Chevrolat H. decui Spanger 1980: 329; Perkins 1980: 144; 1864: 593; Gundlach 1891: 75 Spangler 1980: 329; 1981: 162 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Guantánamo Cuba localities. Gu: Sabanilla, Arroyo de la Collections. Poa (t.l.) Collections. Washington (type) Idolia Lewis 1885: 214 [H. marginicollis Kiesenwetter 1849: 177] I. laevissima (LeConte) 1852: 40 (Coerosternus); Distribution. se USA, erroneously reported Chevrolat 1864: 595 (Tribalus); Gundlach 1891: from Cuba and Virgin Islands according to Jach 78. 1994 Tribalus laevigatus (Paykul) (Hister); Jacquelin H. perkinsi Spangler 1980: 331; Perkins 1980: 172; du Val 1856: 108 Spangler 1980:331; 1981: 162 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Mina Carlota . Gu: up- Cuba localities. Gr: Laguna la Charco per Ovando River. M: Cárdenas Redondo. Gu: Arroyo de los Berros. Ha: Laguna Collections. Cambridge Ariguanabo. Ho: Mayari Arriba; Mayari Abajo. IJ: Laguna; Represa la Fé. SC: El Cayo at Baire. REFERENCES PR: Laguna Esperanza; Laguna La Cochinata; Caterino, M. S. 1999. The taxonomy and Quemado de Pineda (t.l.) phylogenetics of the Coenosus group of Hister Collections. Brussels. Bucharest (Decu). Lon- Linnaeus (Coleoptera: Histeridae). Univ. Cali- don. San Francisco. Ottawa. Paris. Washing- fornia Publ. Ent. 110. 75 pp. ton (type) Decou, V. and Therond, J. 1977. Histérides hypogés de Cuba. Resultats des Expéditions SUBFAMILY OCHTHEBIINAE Biospeologique cubano-roumaines à Cuba 2: 403- TRIBE OCHTHEBIINI 405. Mazur, S. 1984. A world catalogue of Histeridae. Gymnochthebius d’Orchymont 1943: 38 Polskie Pismo. Ent. 54 (3-4): 1-379. G. fossatus (LeConte) 1855: 362 (Ochthebius); Mazur, S. 2001. Review of the Histeridae (Co- Perkins 1980: 277; Spangler 1981: 163 leoptera) of Mexico. Dugesiana 8: 17-66. Ochthebius nitiduloides d’Orchymont 1943: 43; Johnson, S.A., R.W. Lundgren, A.F.Newton, Spangler 1981: 162 (Hydraena) Jr., M.K. Thayer, R.L. Wenzel, and M.R. Distribution. Cuba, Jamaica, Dominican Wenzel. 1991. Mazur’s world catalogue of Republic, Puerto Rico, México to Brazil and Histeridae: emendations, replacement names Argentina, USA (CA-FL) for homonyms, and an index. Polskie Pismo. Cuba localities. Ci: Soledad. Gr: Laguna La Ent. 61 (2): 1-100. Charco Redondo. Ha: Laguna de Ariguanabo. Wenzel, R. 1944. On the classification of the Ho: Arroyo Landivar; Mayari Arriba; Rio Histerid Beetles. Field Museum Zool. Ser. 28 Seboruco. IJ: Represa Fé. OR: Arroyo de los (no. 545): 51-151. Berros. SC: El Cayo at Baire; Laguna Juragua Collections. Washington SUPERFAMILY STAPHYLINOIDEA 22. FAMILY HYDRAENIDAE Ochthebius Leach 1815: 95 The minute moss beetles O. attritus LeConte 1878: 380; Perkins 1980: 346; This family occurs in aquatic and semi-aquatic Spangler 1981: 164 habitats, in sand and gravel along stream edges, Distribution. Cuba, Dominican Republic, in aquatic vegetation, and brackish pools. The Puerto Rico, Yucatan, Colombia, Brazil, USA fauna of the Western Hemisphere was revised by (TX-FL) Perkins (1980). Cuba localities. Ho: Loma la Bandera; Mayari Abajo SUBFAMILY HYDRAENINAE Collections. Washington TRIBE HYDRAENINI 56 Arthropods of Florida and Neighboring Land Areas, Vol. 18

REFERENCES poorly studied and unreported genera are present. Jach, M. 1994. Descriptions of new species of The fauna of the Neotropical Region is summarized Hydraenidae (Coleoptera) from Texas, Indiana, by Peck et al. (1998). and Oklahoma, with faunistic and taxonomic notes on the family in Texas. Coleop. Bull. 48: SUBFAMILY 301-308. TRIBE ANEMADINI Perkins, P.D. 1980. Aquatic beetles of the family Hydraenidae in the Western Hemisphere: clas- Dissochaetus Reitter 1884: 39 sification, biogeography and inferred phylogeny D. cubensis Peck 1999: 606 (Insecta: Coleoptera). Quaest. Ent. 16: 3-554. Distribution. Cuba. Endemic Spangler, P. J. 1980. Two new species of aquatic Cuba localities. SC: Gran Piedra, 1000 m, beetles of the genus Hydraena from Cuba (Co- forests (t.l.) leoptera: Hydraenidae). Proc. Ent. Soc. Wash- Collections. Gatineau (type). Cambridge. ington 82: 329-333. Chicago. Gainesville Spangler, P.J. 1981. Supplement to the aquatic and semiaquatic Coleoptera of Cuba. Resultats TRIBE PTOMAPHAGINI des expeditions biospéologiques Cubano- Roumaines à Cuba 3: 145-171. Proptomaphaginus Szymczakowski 1969: 87. The genus is a relictual cave, forest, and soil 23. FAMILY PTILIIDAE inhabitant in México, Cuba, Hispaniola, and The feather-winged beetles Puerto Rico This family of minute beetles lives in leaf litter P. apodemus Szymczakowski 1969: 88; Decou 1973: and in association with decaying wood, and fungi, 367 (); Peck 1983: 257; 1999: 609; Peck et and even in spore tubes of bracket fungi. They are al. 1998: 159 certainly present and diverse in Cuba but no iden- Distribution. Cuba. Endemic tified species records have been published. They Cuba localities. Widespread; common in have been recovered from guano in caves. caves on bat guano; also in lowland forests Collections. Bucharest. Gatineau. Cam- SUBFAMILY ACROTRICHINAE bridge. Chicago. Gainesville. Krakow (type). Pittsburgh Acrotrichis Motschulsky 1848: 569 P. darlingtoni (Jeannel) 1936: 92; Peck 1970: 241.; A. species; Genaro and Gutiérrez 2001: 10 1999: 608 Distribution. Cuba. Endemic? Distribution. Cuba. Endemic Cuba localities. Gr: Alegria de Pio Cuba localities. Ci: Soledad (t.l.). SC: Gran Collections. Havana Piedra (600-1100m, forests only) Collections. Gatineau. Cambridge (type). REFERENCE Chicago. Gainesville Genaro, J. A. and E. Gutiérrez. 2001. La fa- milia Ptiliidae en Cuba (Insecta: Coleoptera). SUBFAMILY Cocuyo 11: 10-11. TRIBE LEIODINI

24. FAMILY AGYRTIDAE Zeadalopus Broun 1903: 614. This scavenging family is mostly Holarctic in Z. sp. distribution, and ranges from the Mexican Plateau Distribution. Cuba. Endemic northwards in the Nearctic. It does not occur in Cuba localities. SC: Gran Piedra; Santiago, the West Indies or from Guatemala to southern Jardin Botanico; 10 km NE Caney, Arroyo South America. Grovert Collections. Peck 25. FAMILY LEIODIDAE The small carrion and round beetles TRIBE PSEUDOLIODINI This family has diverse habits, and members feed on fungi, carrion and animal dung. They may Aglyptinus Cockerell 1906: 240 live in leaf litter and caves and may be in nests of A. sp. vertebrates and social insects. The Cuban fauna is Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 57

Cuba localities. PR: Cueva la Barca, in fruit Euconnus Thomson 1862: 88 bat guano. SC: Gran Piedra; Santiago, Jardin E. breviceps (Schaufuss) 1867: 58 (Scydmaenus); Botanico Gundlach 1891: 88; Franz 1991:41 Collections. Havana. Peck Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. Gr: Bayamo. M: TRIBE SCOTOCRYPRINI Cárdenas. SS: Mina Carlota Collections. Eberswalde (type). Franz. Paris. Creagrophorus Matthews 1887: 82 Urbana C. sp. E. cienfuegosi Franz 1991: 45 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Gran Piedra; Santiago, Cuba localities. Ci: Soledad (t.l.) Jardin Botanico Collections. Franz. Urbana (type) Collections. Peck E. cubaicola Franz 1991: 44 Distribution. Cuba. Endemic REFERENCES Cuba localities. SS; Mina Carlota (t.l.) Decou, V. Gh. 1973. Recherches sur les Coléoptères Collections. Urbana (type). Franz hypogés de Cuba. III. Catopidae-Catopinae: E. cubanus Franz 1991: 44 Protomaphaginus apodemus Szymczakowski. Distribution. Cuba. Endemic Résultats des expéditions biospéologiquess Cuba localities. SS: Mina Carlota (t.l.) Cubano-Roumaines à Cuba. 1. 367-372. Collections. Urbana (type) Peck, S.B. 1970. The Catopinae (Coleoptera: E. globulicollis (Schaufuss) 1867: 64 (Scydmaenus); Leiodidae) of Puerto Rico. Psyche 77: 237-242. Gundlach 1891: 88 Peck, S.B. 1983. A new cavernicolous Distribution. Cuba. Endemic Proptomaphaginus from Hispaniola and Cuba localities. M: Cárdenas México (Coleoptera: Leiodidae: Cholevinae). Collections. Paris Florida Ent. 66: 254-260. E. gundlachianus Franz 1991: 41 Peck, S.B. 1999. New species and records of “small Distribution. Cuba. Endemic carrion beetles” from Cuba and Hispaniola (Co- Cuba localities. Ci: Soledad (t.l.) leoptera; Leiodidae; Cholevinae). Canadian Ent. Collections. Urbana (type) 131: 605-611. E. gundlachii (Schaufuss) 1867: 59 (Scydmaenus); Peck, S.B., P. Gnaspini, and A.F. Newton. Gundlach 1891: 88 1998. Catalogue and generic keys for the Distribution. Cuba. Endemic Leiodidae of México, West Indies, and Central Cuba localities. M: Cárdenas and South America. Giornale Italiana di Collections. Riehl collection in Cassel? Entomologia 9: 37-72. E. patens (Schaufuss) 1867: 66 (Scydmaenus) Peck, S.B., A.E. Ruiz-Baliú, and G.F. Garcés Distribution. Cuba. Endemic González. 1998. The cave-inhabiting beetles Cuba localities. of Cuba (Insecta: Coleoptera): diversity, distri- Collections. Riehl collection in Cassel? bution and ecology. J. Cave Karst Studies 60: E. schaufussi Csiki 1919: 62 (replacement name) 156-166. Eumicrus brevicornis Schaufuss 1867: 93 Szymczakowski, W. 1969. Découverte d’un Distribution. Cuba. Endemic représentant des Ptomaphagini à Cuba (avec Cuba localities. une esquisse de la systématique et la géonémie Collections. de cette tribu) (Coleoptera, Catopidae). Acta Zool. E. soledadensis Franz 1991: 42 Cracoviensia 14: 87-97. Distribution. Cuba. Endemic Cuba localities. Ci: Soledad (t.l.). SS: Mina 26. FAMILY SCYDMAENIDAE Carlota The antlike stone beetles Collections. Urbana (type). Franz Reviewed by Sean T. O’Keefe, February, 2001 This is a poorly known family of leaf-litter in- Homoconnus Sharp 1887: 64 habiting predators. H. dentipes (Schaufuss) 1867: 71 (Scydmaenus); Gundlach 1891: 88 SUBFAMILY Distribution. Cuba. Endemic TRIBE CYRTOSCYDMINI Cuba localities. 58 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Eberswalde seum, Chicago and it contains many additional Cuban specimens that are unidentified or with TRIBE SCYDMAENINI manuscript names. Cuban records of species in this museum are listed here, from data provided by A. Scydmaenus Latreille 1802: 116 F. Newton, but identifications and/or distributions S. pubescens (Schaufuss) 1867: 76 (Eumicrus); need to be confirmed. Full synonym lists and dis- Gundlach 1891: 88 tributional details are in the monumental catalogue Distribution. Cuba. Endemic of Herman (2001), but this does not cover the sub- Cuba localities. Widespread families , , Scaphidiinae Collections. Esberswalde and Pselaphinae. Most genera in Cuba are covered in the keys to the Mexican and Central American REFERENCE fauna in Navarrete et al. (2002). Franz, M. 1991. Weiterer Beitrag zur Kenntnis des Scydmaeniden der Karibischen Inseln (Co- SUBFAMILY leoptera: Scydmaenidae). Koleopterologische TRIBE OMALIINI Rundschau 61: 25-48. Omalium Gravenhorst 1802: 111 27. FAMILY SILPHIDAE O. cubanum Blackwelder 1943: 55 This family does not occur in Cuba. One spe- Distribution. Cuba. Endemic cies of Nicrophorus occurs in the highlands of Cuba localities. Hispaniola (Sikes and Peck 2000), and this genus Collections. and others ring the Caribbean from the USA to México, Central America, and throughout South Phloeonomus Heer 1839: 184 America (Peck and Anderson 1985). P. pedicularius (Erichson) 1840: 887 Distribution. Cuba, Jamaica, Hispaniola, REFERENCES Puerto Rico, Guadeloupe, , Trinidad Peck, S.B. and R.S. Anderson. 1985. Taxonomy, Cuba localities. phylogeny, and biogeography of the carrion Collections. beetles of (Coleoptera, Silphidae). Quaest. Ent. 21: 247-317. SUBFAMILY PSELAPHINAE Sikes, D.S. and S.B. Peck. 2000. Description of The -like beetles Nicrophorus hispaniola, new species, from Reviewed by D. Chandler, March, 2001 Hispaniola (Coleoptera: Silphidae) and a key to These small beetles live in forest leaf litter and the species of Nicrophorus of the New World. feed on mites, Collembola, and other small inverte- Ann. Ent. Soc. Amer. 93: 391-397. brates. The Cuban fauna is very poorly known. Park et al. (1976) is the last treatment of the Cuban fauna. 28. FAMILY STAPHYLINIDAE This group is considered to be a subfamily of The rove beetles and allies Staphylinidae by Newton and Thayer (1995) and Checked and Reviewed by J. H. Frank, April, Lawrence and Newton (1995). We use here their 2001; A.F. Newton, March and July, 2002. classification within the subfamily Pselaphinae. This is a very large family of mostly predatory beetles. The following list has been mostly drawn SUPERTRIBE EUPLECTITAE from Blackwelder (1943), and the accuracy of iden- TRIBE EUPLECTINI tification and distribution for many species that have not been recently revised may be highly sus- Lemelba Park 1953: 1 pect. The Blackwelder collection is in Washington, L. millsi Park 1976: 40 and contains many misidentifications. While the Distribution. Cuba. Endemic North American fauna is now becoming well stud- Cuba localities. Ci: Soledad. (t.l.) ied, the West Indian fauna is still very poorly known. Collections. Chicago. Urbana (type) Frank (1986) provided a checklist for Florida. The higher classification follows that of Lawrence and SUPERTRIBE GONIACERITAE Newton (1995). Many species described by Bierig TRIBE BRACHYGLUTINI may need lectotypes designated. The large and im- portant Bierig collection is now at the Field Mu- A Checklist of the Beetles of Cuba 59

Berdura Reitter 1882: 206 Cuba localities. Gu: near Imias (t.l.), and B. leavitti Park 1976: 49 Upper Yara Valley Distribution. Cuba. Endemic Collections. Cambridge (type). Chicago Cuba localities. Ci: Soledad (t.l.) R. truncata (Schaufuss) 1887: 128 Collections. Cambridge (type). Chicago Distribution. Cuba. Endemic Cuba localities. “Cuba” Briaraxis Brendel 1894:158 Collections. B. depressa Brendel 1894: 159 Distribution. Cuba, Jamaica, Virgin Islands, SUPERTRIBE PSELAPHITAE , Panama, Venezuela, USA (FL) TRIBE TYRINI Cuba localities. “Cuba.” Collections. Chicago Ephimia Reitter 1883: 34 E. cubensis Park 1976: 67 Bythinogaster Schaufuss 1887: 111 Distribution. Cuba. Endemic B. bisphaeroides Schaufuss 1887: 297; Park et al. Cuba localities. Ci: Soledad (t.l.) 1976: 54 Collections. Cambridge (type). Chicago Distribution. Cuba. Endemic E. sp. Zayas 1988: 22 Cuba localities. CA: Baraguá (t.l.) Distribution. Cuba. Endemic Collections. Berlin (type). Cambridge. Chi- Cuba localities. cago Collections. Zayas B. parsonsi Park 1976: 54 Distribution. Cuba. Endemic TRIBE CTENISTINI Cuba localities. Ci: Soledad (t.l.) Collections. Cambridge (type). Chicago Ctenistodes Raffray 1896: 247 C. weberi Park 1976: 64 Decarthron Brendel 1865: 30 Distribution. Cuba. Endemic D. arcuatum Park 1976: 60 Cuba localities. Ci: Soledad (t.l.) Distribution. Cuba. Endemic Collections. Cambridge (type). Chicago Cuba localities. Ci: Soledad (t.l.) Collections. Cambridge (type). Chicago TRIBE ARHYTODINI D. pectinale Park 1976: 61 Distribution. Cuba. Endemic Caccoplectus Sharp 1887: 22 Cuba localities. CA: Baraguá. Ci: Soledad C. schwarzi Chandler and Wolda 1986: 485 (t.l.) Distribution. Cuba. Endemic Collections. Chicago. Cambridge (type) Cuba localities. Ci: Cayamas (t.l.) D. unifovealatum (Schauffuss) 1887: 125 (Bryaxis) Collections. Chicago. Washington (type) Distribution. Cuba. Endemic Cuba localities. “Cuba” SUPERTRIBE CLAVIGERITAE Collections. TRIBE CLAVIGERINI

Pselaptus LeConte 1880: 184 Fustiger LeConte 1866: 109 P. longiclavus (Schaufuss) 1887: 126 (Bryaxis); F. barroi (Bierig) 1945: 15 (Pseudofustiger); Park Park et al. 1976: 52 et al. 1976: 72 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. “Cuba” (t.l.) Collections. Chicago Collections. Chicago (type) F. schwarzi Mann 1918: 105; Park et al. 1976: 73 Reichenbachia Leach 1826: 451 Distribution. Cuba. Endemic R. bisinuata (Schaufuss) 1882: 130 Cuba localities. Ci: Cayamas. (t.l.); in ant Distribution. Cuba. Endemic nest Cuba localities. “Cuba.” Collections. Chicago. Washington (type) Collections. Chicago F. stricticornis (Reitter) 1883: 33 (Articerus); Bierig R. darlingtoni Park 1976: 57 1945: 13; Park et al. 1976: 72 Distribution. Cuba. Endemic 60 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, St. Thomas (Virgin Is- Collections. Cambridge. Chicago. Gundlach lands) no. 85 (C. infimus). Washington Cuba localities. PR: Sierra del Rosario C. rutilus (Erichson) 1839: 253 (Tacninus); Collections. Chicago Blackwelder 1943: 522; Campbell 1975: 195 Distribution. Cuba, Jamaica, Hispaniola, SUBFAMILY TACHYPORINAE Puerto Rico, St. Thomas to Trinidad, South TRIBE TACHYPORINI America, Mexico, Central America, USA (TX- LA) Coproporus Kraatz 1857: 399 Cuba localities. Ci: Cayamas; Soledad. M: C. apicalis (Erichson) 1839: 250 (Tachinus); Cárdenas Blackwelder 1943: 514 Collections. Cambridge. Chicago. Gundlach Distribution. Cuba, Jamaica, Hispaniola, no. 412 (C. terminalis). Washington Puerto Rico C. segnis (Sharp) 1883: 301 (Erchomus) Cuba localities. Ci: Soledad. SC: Loma del Distribution. Cuba, Mexico, Guatemala, Gato; Pico Turquino Costa Rica, se USA (FL) Collections. Cambridge. Chicago. Washing- Cuba localities. ton Collections. Chicago C. cinctiventris (Sharp) 1883: 304; Navarrete et al. C. terminalis (Erichson) 1839: 250 (Tachinus); 2002: 110 Jacquelin du Val 1856: 32; Chevrolat and Distribution. Cuba, Argentina, Colombia, Fauvel 1863: 429 (Coproporus); Gundlach Costa Rica, Guatemala, Mexico 1891: 55; Campbell 1975: 197 Cuba localities. Ha: Havana Distribution. Cuba, Puerto Rico Collections. Chicago Cuba localities. Ha: Havana C. hepaticus (Erichson) 1839: 249 (Tachinus); Collections. Chicago Blackwelder 1943: 514; Campbell 1975: 189 C. vicinus Sharp 1883: 305; Navarrete et al. 2002: C. convexus (Erichson) 1839:249; Chevrolat and 111 Fauvel 1863: 429; Gundlach 1891: 55 Distribution. Cuba, Mexico, Guatemala, (Coproborus) Costa Rica Distribution. Cuba, Puerto Rico, Trinidad, Cuba localities. Ha: Havana South America, Mexico, Central America, USA Collections. Chicago (CA-TX, FL) Cuba localities. Ci: Cayamas; Soledad. San Sepedophilus Gistel 1856: 267 Antonio de los Baños S. interruptus (Erichson) 1839: 225 (Conurus); Collections. Cambridge. Chicago. Washing- Blackwelder 1943: 525 (Conosomus) ton S. caribeanum (Bierig) 1937: 276 (Conosoma) C. nitidulus (Erichson) 1839: 247 (Tachinus) S. cubanum (Bierig) 1937: 277 (Conosoma) C. flavicollis (Scriba) 1855: 297 (Tachyporus); S. conforme (Bierig) 1937: 278 (Conosoma) Blackwelder 1943: 518 Distribution. Cuba, Jamaica, Hispaniola, Distribution. Cuba, Jamaica, Puerto Rico, Puerto Rico, Monserrat to Trinidad, Columbia South America, Mexico, Central America. Cuba localities. Ha: Sierra de Bonilla. PR: Cuba localities. Ci: Cayamas; Soledad El Rangel; Cueva del Indio; Pico Somorrostro; Collections. Cambridge. Chicago. Washing- Aspiro. Ci: Cayamas; Soledad. SC: Pico ton Turquino C. pulchellus (Erichson) 1839: 247 (Tachinus); Collections. Cambridge. Chicago. Washing- Blackwelder 1943: 520; Campbell 1975: 185. ton Tachinus infimus Jacquelin du Val 1857: 33; Chevrolat and Fauvel 1863: 430 (Coproporus); TRIBE MYCETOPORINI Gundlach 1891: 55 Distribution. Cuba, Jamaica, Hispaniola, Bryoporus Kraatz 1858: 452 Puerto Rico, Antigua to Trinidad, South B. aciculatus Blackwelder 1943: 529 America, Central America, México, USA (AL- Distribution. Cuba. Endemic FL) Cuba localities. Gu: Mts. N of Imias (t.l.) Cuba localities. Widespread Collections. Chicago. Cambridge (type) A Checklist of the Beetles of Cuba 61

B. cubanus Blackwelder 1943: 530 Collections. Gundlach no. 1734 Distribution. Cuba. Endemic A. (Xenochara) puberula Klug 1833: 139; Cuba localities. Ci: Soledad (t.l.) Blackwelder 1943: 561 Collections. Chicago. Cambridge (type). A. dubia Fauvel in Chevrolat and Fauvel 1863: Washington 428; Gundlach 1891:54 B. seriatus (Erichson) 1839: 272 (Boletobius) Distribution. Cuba, México, USA. Cosmopoli- Distribution. Cuba, Mexico, Costa Rica, Co- tan-pantropical-subtropical (records in lombia, Venezuela, Brazil, Argentina Klimaszewski et al. 2000: 233) Cuba localities. Cuba localities. Collections. Chicago Collections. Chicago. Gundlach no. 890 (A. dubia) SUBFAMILY ALEOCHARINAE A. (Xenochara) verberans Erichson 1839: 164; Reviewed by J. Klimaszewski, April, 2001 Chevrolat and Fauvel 1863: 428; Gundlach 1891: 54; Blackwelder 1943: 561; Klimaszewski TRIBE DEINOPSINI and Génier 1986: 116 (lectotype designation and redescription) Adinopsis Cameron 1919: 242 Distribution. Cuba, South America, Central A. myllaenoides (Kraatz) 1857:38 (); America Klimaszewski 1979: 72; 1980:120 Cuba localities. Distribution. Cuba, Jamaica, St. Lucia, Collections. Chicago México, Central America, Trinidad, Venezuela, Brazil, USA (MS, LA, FL) TRIBE HOPLANDRINI Cuba localities. Ci: Cayamas Collections. Chicago.Washington Hoplandria Kraatz 1857: 4 H. (Genosema) aemula (Bierig) 1939: 27 (Genosema) TRIBE MESOPORINI Distribution. Cuba. Endemic Cuba localities. Ha: Jamaica. PR: Aspiro; Anacyptus Horn 1877: 87 Caimito del Guayabal A. testaceus (LeConte) 1863: 30 (Hypocyptus): Collections. Chicago (type) Blackwelder 1943: 532 H. pacei Hanley 1998: 386 (replacement name for Distribution. Cuba, Montserrat, Mexico, H. puncticollis Pace 1990: 166) USA (AR-MI-NC-FL) H. puncticollis Pace 1990: 166 Cuba localities. Ci: Cayamas Distribution. Cuba. Endemic Collections. Chicago. Washington Cuba localities. Ha: Playa Marianao (t.l.) Collections. Chicago (type) TRIBE Sharp 1883: 170 Gravenhorst 1802: 67 T. cavicollis Sharp 1883: 170 A. (Aleochara) lateralis Erichson 1837: 161; Distribution. Cuba, Guatemala, Nicaragua, Chevrolat and Fauvel 1863: 428; Gundlach Paraguay, Argentina, USA (FL) 1891: 53; Blackwelder 1943: 560; Klimaszewski Cuba localities. et al. 1987: 255, Klimaszewski 1990: 173 Collections. Chicago Distribution. Cuba, México, South America Cuba localities. M: Cárdenas; Cienaga de TRIBE Zapata Collections. Chicago. Gundlach no. 894 Sharp 1887: 779 A. (Coprochara) notula Erichson 1839: 167; M. albicincta (Erichson) 1839: 56; Blackwelder 1943: Jacquelin du Val 1856: 32; Chevrolat and 548 Fauvel 1863: 429; Gundlach 1891: 54; Distribution. Cuba, Jamaica, Grenada, Blackwelder 1943: 560 México, South America Distribution. Cuba to Mustique, México, Cuba localities. South America (Klimaszewski et al. 1987, Collections. Chicago Klimaszewski 1990), USA (CA-TX-NE-PA-FL) Cuba localities. M: Cárdenas 62 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE Collections. Chicago (type) H. zonata (Bierig) 1934: 115 (); Blackwelder Neolara Sharp 1883: 231 1943: 538; Frank 1972: 132 N. cubana Casey 1906: 189; Blackwelder 1943: 548 Distribution. Cuba, Jamaica, USA (FL) Distribution. Cuba. Endemic Cuba localities. PR: Aspiro-Rangel; Cuba localities. PR: Bahia Honda (t.l.) Portugues. Ha: Sierra Bonilla (t.l.) Collections. Washington (type). Chicago Collections. Chicago (type)

Phloeopora Erichson 1837: 311 Oligota Mannerheim 1830: 72 (see revision of P. occidentalis Cameron 1923: 388; Blackwelder Frank 1972) 1943: 557 [O. centralis Sharp 1883: 293; Bierig 1934: 114; Distribution. Cuba, Hispaniola Blackwelder 1943: 536; Frank 1972: 137] Cuba localities. Distribution. Guatemala, Colombia. Re- Collections. Chicago. London corded in error by Bierig (1934) from Ha: , see O. minuta below TRIBE HYPOCYPHTINI (O. chrysopyga Kraatz 1859:45; Frank 1972: 131) Distribution. West Indies, USA (FL); Solier 1849: 335 pantropical H. cadaverina (Bierig) 1934: 113 (Oligota); Cuba localities. Blackwelder 1943: 536; Frank 1972: 132 Collections. Distribution. Cuba, Virgin Islands O. maculicornis Cameron 1922: 124; Bierig 1934: Cuba localities. Ha: Playa Marianao; Mariel; 117; Blackwelder 1943: 536; Frank 1972: 142 Sierra Bonilla, Cueva Indio. PR: Rangel (t.l.) Distribution. Cuba, Hispaniola Collections. Chicago (type) Cuba localities. PR: Aspiro; Pico H. caribae (Frank) 1972: 134 (Oligota) Somorrostro; Sierra Anafe; Sierra del Rosario, Distribution. Cuba. Endemic El Sabicu Cuba localities. Ha: Sierra Rangel (t.l.). PR: Collections. Chicago Aspiro O. parva Kraatz 1862: 300; Bierig 1934: Frank Collections. Chicago (type) 1972: 140 H. humboldti (Frank) 1972: 135 (Oligota) Distribution. Cuba, Jamaica, Tortola, St. Distribution. Cuba. Endemic Vincent, Grenada, Central America, South Cuba localities. Ci: Trinidad, El Cubanito America, USA (TX-MO-MA-FL), Europe, Ori- (t.l.) ent Collections. Chicago (type) Cuba localities. Ha: Havana, under dog H. luteicornis (Bierig) 1934: 119 (Oligota); corpse (Bierig 1934) Blackwelder 1943: 536; Frank 1972: 138 Collections. Chicago Distribution. Cuba. Endemic O. rhopalocera Bernhauer 1923: 145; Bierig 1934: Cuba localities. Ha: Cruz de Piedra; La Lisa; 116; Blackwelder 1943: 537; Frank 1972: 139 Mariel; Sierra de Anafe, Caimito del Guayabal. Distribution. Cuba, St. Thomas PR: Aspiro; Sierra de Candela, Güines; Sierra Cuba localities. Ha: Sierra Bonilla, Cueva del Rosario del Indio. PR: Cruz de Piedra; Sierra de Can- Collections. Chicago dela, Güines; Sierra Rangel; Tepaste H. minuta (Cameron) 1931: 82 (Oligota); Frank Collections. Chicago 1972: 136 (O. testaceorufa Bernhauer 1923: 144) Distribution. Cuba, Jamaica, Trinidad Distribution. St. Thomas, USA (FL) Cuba localities. Ha: Havana; Santiago de las Cuba localities. Vegas; reported by Bierig (1934) under leaves Collections. of beans and on Cedrela infested with mites, O. tricolor Bierig 1934: 116; Blackwelder 1943: 538; under the name of O. centralis Frank 1972: 139 Collections. Chicago Distribution. Cuba. Endemic H. multicarinata (Frank) 1972: 134 (Oligota) Cuba localities. Ha: (t.l.) Distribution. Cuba. Endemic Collections. Chicago (type) Cuba localities. PR Aspiro; El Rangel; Si- erra del Rosario (t.l.) A Checklist of the Beetles of Cuba 63

TRIBE MYLLAENINI D. obscuricollis (Bernhauer) 1908: 344 (Apheloglossa); as var. luederwaldti) Erichson 1837: 382 Distribution. Cuba, Brazil, Argentina, Para- (M. potowatomi Klimaszewski 1982: 192) guay Distribution. Haiti, Jamaica, México, USA Cuba localities. (CA-TX-WI-FL) Collections. Chicago D. (Apheloglossa) melanura (Erichson) 1839: 117 TRIBE (); Blackwelder 1943: 553; Pace 1986: 421 Brachychara Sharp 1883: 267 Distribution. Cuba, Puerto Rico, St. John B. palliditarsis Bierig 1939: 17 Cuba localities. Distribution. Cuba. Endemic Collections. Chicago Cuba localities. PR: Sierra del Rosario, Aspiro D. sperata Sharp 1876: 47; Blackwelder 1943: 544 (t.l.) Distribution. Cuba, Jamaica, Hispaniola, Collections. Chicago (type) Guadeloupe to Grenada, Guatemala, Costa Rica, South America Mannerheim 1830: 75 Cuba localities. B. cinctigastra Bierig 1937: 280; Blackwelder 1943: Collections. Chicago 546 Distribution. Cuba. Endemic Eumicrota Casey 1906: 280 Cuba localities. Ha: Sierra Bonilla. PR: El E. cornuta Casey 1906: 285 Rangel Distribution. Cuba. Endemic Collections. Chicago (type) Cuba localities. Ci: Cayamas. (t.l.) Collections. Chicago. Washington (type) Cephaloxynum Bernhauer 1907: 282 C. rambouseki Bierig 1931: 424; Blackwelder 1943: Mannerheim 1830: 74 459; Newton 1988: 256; Frank et al. 1989: 717 G. atomaria Cameron 1922: 647 Distribution. Cuba, USA (FL) Distribution. Cuba, Jamaica, Grenada Cuba localities. Ha: Camoa; Sierra Bonilla, Cuba localities. Tepaste (t.l.). PR: Sierra del Rosario, Rangel; Collections. Chicago SS: Buenos Aires Collections. Cambridge. Chicago (type). Heterota Mulsant and Rey 1874: 194 Washington (H. plumbea (Waterhouse) 1858: 6074) Distribution. Jamaica, México, coastal Eu- Coenonica Kraatz 1857: 45 rope to , USA (FL); marine lit- (C. puncticollis Kraatz 1857:46) toral; possibly introduced to New World (see Distribution. Grenada, St. Vincent, Guyana, Frank and Thomas, 1984, Fla. Ent. 67: 409- Brazil, USA (FL); pantropical, from Old world; 417) subcortical (see Frank and Thomas, 1984, Fla. Cuba localities. Ent. 67: 409-417) Collections. Cuba localities. Collections. Phanerota Casey 1906: 285 P. cubensis Casey 1906: 288; Blackwelder 1943: Diestota Mulsant and Rey 1870: 194 541 (Gyrophaena) D. ambiguum (Notman) 1920: 715 (Elachistarthron Distribution. Cuba, USA (FL) Distribution. Cuba, USA (FL) Cuba localities. Ci: Cayamas. (t.l.) Cuba localities. Collections. Chicago. London, Cameron col- Collections. Chicago lection. Washington (type) D. braziliana Bernhauer 1907: 288 Distribution. Cuba, Brazil, Argentina Cuba localities. Stictalia Casey 1906: 264 Collections. Chicago S. unicolor (Fenyes) 1909: 198 (Bolitochara) D. luederwaldti (Bernhauer) 1908: 343 Distribution. Cuba, USA (CA), Canada (BC) (Apheloglossa) Cuba localities. 64 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Chicago TRIBE LOMECHUSINI

Xenobiota Bierig 1938: 144 Stephens 1832: 430 X. bernhaueri Bierig 1938: 145; Blackwelder 1943: Z. distinctus Bierig 1937: 281; Blackwelder 1943: 54 556 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Viñales; Sumidero Rio Cuba localities. PR: El Rangel (t.l.). Pre- Taco Taco; El Sabicu; Rangel sumed guest in nests of ants Tetramorium Collections. Chicago (type) simillimum Smith Collections. Chicago TRIBE PLACUSINI SUBFAMILY SCAPHIDIINAE Euvira Sharp 1883: 278 Many more species of this family of fungus- E. cubana Bierig 1934: 121; Blackwelder 1943: 546; feeding beetles should occur in Cuba than are re- Bierig 1939: 25 corded. See Leschen and Löbl (1995) for classifica- E. flavipes Bierig 1934: 123 ( t.l., Sierra de tion within the subfamily. Rosario) Distribution. Cuba. Endemic TRIBE CYPARINI Cuba localities. Ha: Sierra Bonilla, Cueva del Indio. PR: near Rangel Cyparium Erichson 1845: 3 Collections. Chicago (type) C. flavosignata Zayas 1988: 23 E. debilis Sharp 1883: 281 Distribution. Cuba. Endemic Distribution. Cuba, Costa Rica, Guatemala, Cuba localities. None given USA (FL) Collections. Zayas (type) Cuba localities. C. f. bicolor Zayas 1988: 25 Collections. Chicago Distribution. Cuba. Endemic Cuba localities. None given TRIBE Collections. Zayas (type) C. f. funebris Zayas 1988: 24 Thomas 1859: 39 Distribution. Cuba. Endemic A. (Datomicra) sordidula (Erichson) 1839: 335 Cuba localities. None given (Homalota); Blackwelder 1943: 554 Collections. Zayas (type) Distribution. Cuba, widespread North C. f. splendidum Zayas 1988: 25 America, Europe Distribution. Cuba. Endemic Cuba localities. Cuba localities. None given Collections. Chicago. London Collections. Zayas (type) A. (Xenota) longifrons Sharp 1883: 178 Distribution. Cuba, Guatemala, ?Mexico TRIBE SCAPHISOMATINI Cuba localities. Collections. Chicago Scaphisoma Leach 1815: 89 S. bicinctum Zayas 1988: 27 Trichiusa Casey 1893: 339 Distribution. Cuba. Endemic T. ursina Notman 1920: 725 Cuba localities. None given Distribution. Cuba, USA (FL) Collections. Zayas (type) Cuba localities. S. cubense Reitter 1879 (1880): 48 Collections. Chicago Distribution. Cuba. Endemic Cuba localities. None given TRIBE Collections. Zayas (type) S. nitidulum Zayas 1988: 26 Guajira Bierig 1938: 146 Distribution. Cuba. Endemic G. cubana Bierig 1938: 146; Blackwelder 1943: 555 Cuba localities. None given Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. PR: Sierra del Rosario (t.l.) Collections. Chicago (type) A Checklist of the Beetles of Cuba 65

SUBFAMILY TRIBE THORACOPHORINI

Hypotelus Erichson 1840: 840 Clavilispinus Bernhauer 1926: 255 H. insulanus Bierig 1934: 343; Blackwelder 1943: C. exiguus (Erichson) 1840: 830; Irmler 1991: 86 42 Paralispinus exiguus (Erichson), Blackwelder Distribution. Cuba, Jamaica, Hispaniola, St. 1943: 158 Vincent Lispinus piceus Fauvel, of Leng and Mutchler; Cuba localities. Ci: Cayamas. PR: Aspiro Blackwelder 1943: 159 (Paralispinus) (t.l.), Sierra del Rosario Distribution. Cuba, Jamaica, Hispaniola, Collections. Chicago (type). Washington Puerto Rico, St. Croix, Guadeloupe, St. Lucia, Grenada, Columbia, Bolivia, Brazil, Guatemala, Piestus Gravenhorst 1806: 223 (see USA (LA-FL); Old World Scheerpeltz 1952) Cuba localities. Ci: Cayamas. Ha: Sierra de (P. mexicanus Castelnau 1834: 130) Anafe, Caimito Distribution. West Indies (islands not Collections. Chicago. London. Washington named), México to Venezuela and NW Brazil C. megacephalus Fauvel 1864: 57 (Ancaeus); P. minutus Erichson 1840: 834; Herman 2001: 1792 Blackwelder 1943: 160 (Paralispinus) Distribution. Cuba, Mexico to Argentina C. crepusculus (Blackwelder) 1943: 160 Cuba localities. (Paralispinus) Collections. Distribution. Cuba, Jamaica, Puerto Rico, P. penicillatus (Dalman) 1821: 375 (Zirophorus); St. Lucia, Trinidad, Mexico, Guatemala, Ven- Blackwelder 1943: 46 ezuela to Paraguay P. erythropus Erichson 1840: 834; Jacquelin du Cuba localities. Ci: Cayamas Val 1856: 46; Chevrolat and Fauvel 1863: 444; Collections. Chicago. Washington Gundlach 1891: 69 C. politus (Sharp) 1887: 718; Irmler 1991: 86 Distribution. Cuba, Jamaica, Hispaniola to Clavilispinus rulomus (Blackwelder) 1945: 160 Trinidad, México to French Guyana and ne Distribution. Cuba, Jamaica, St. Vincent, Brazil Guadeloupe, St. Lucia, México, Guatemala, Cuba localities. Ci: Soledad. Gu; Costa Rica, Panama, Brazil, Peru Guantánamo; Upper Ovando River. SS: Buenos Cuba localities. Gr: Yara Aires Collections. Chicago Collections. Cambridge. Gundlach no. 305. New York. Washington Espeson Schaufuss 1882: 168 (P. pygmaeus Castelnau 1834: 130) E. moratus Schaufuss 1882: 168; Blackwelder 1943: Distribution. Hispaniola, Lesser Antilles, 146 México to Argentina, Uruguay Distribution. Cuba, Jamaica, Puerto Rico, Cuba localities. widespread Lesser Antilles, Trinidad Collections. Cuba localities. Ci: Soledad P. sulcatus (Gravenhorst) 1806: 224; Blackwelder Collections. Chicago. Cambridge 1943: 44 Distribution. Cuba, Dominica to Trinidad, Lispinus Erichson 1839: 31 Brazil L. cordobensis Bernhauer 1929: 187; Irmler 1991: Cuba localities. 59; 2000: 83 Collections. Distribution. Cuba, Hispaniola, México, Guadeloupe, Costa Rica, Guatemala, St. SUBFAMILY Vincent, Grenada TRIBE ELEUSININI Cuba localities. Collections. Chicago Eleusis Laporte 1834: 131 L. cubensis Irmler 1994: 63 E. darlingtoni Blackwelder 1943: 156 Distribution. Cuba, Jamaica Distribution. Cuba. Endemic Cuba localities. PR: Sierra “Rampe” Cuba localities. Ci: Soledad (t.l.) (=Rangel) (t.l.) Collections. Cambridge (type). Washington Collections. Chicago (type) 66 Arthropods of Florida and Neighboring Land Areas, Vol. 18

L. insularis Fauvel in Chevrolat and Fauvel 1863: Distribution. Cuba, Brazil, Bolivia 442; Gundlach 1891: 67; Blackwelder 1943: 136; Cuba localities. Irmler 1994: 63; 2000: 85 Collections. Chicago Distribution. Cuba, Hispaniola, Jamaica, N. nigrifrons( Fauvel in Chevrolat and Fauvel 1863: widespread Lesser Antilles; not Panamá, 443 (Lispinus); Gundlach 189l: 68; Blackwelder México, Costa Rica 1943: 128 (Pseudolispinodes Cuba localities. Ha: Caimito. M: Cárdenas; Distribution. Cuba, Hispaniola, Montserrat, Yumuri-Tal [Valley] Guadeloupe, St. Lucia, Mexico, Costa Rica, Collections. Chicago. Gundlach no. 607 Panama, Colombia to Brazil L. laticollis Erichson 1840: 828; Jacquelin du Val Cuba localities. Ci: Cayamas 1856: 45; Chevrolat and Fauvel 1863: 442; Collections. Chicago. Gundlach no. 678. Gundlach 1891: 67; Blackwelder 1943: 131; Washington Irmler 1994: 58 N. opacus (Fauvel) 1864: 51 (Lispinus) Distribution. Cuba (type locality), Puerto Distribution. Cuba, Jamaica, Mexico, Gua- Rico, Panamá, Costa Rica, Brazil temala, Costa Rica, Ecuador, Peru, Bolivia, Cuba localities. Widespread Brazil, sw USA (AZ) Collections. Cambridge. Chicago. Gundlach Cuba localities. no. 616. London. New York. Washington Collections. Chicago L. striola Erichson 1840: 829; Jacquelin du Val N. spegazzinii (Bernhauer) 1933: 326 (Lispinus) 1856: 46; Gundlach 1891: 67; Blackwelder 1943: Distribution. Cuba, Costa Rica, Panama, 135; Irmler 1994: 57 Colombia to Surinam and Argentina Distribution. Cuba, Puerto Rico, México to Cuba localities. Nicaragua, Trinidad, Colombia, Peru, Brazil Collections. Chicago Cuba localities. Ci: Cayamas. M: Cárdenas N. tenellus (Erichson) 1840: 830 (Lispinus); Collections. Chicago. Gundlach no. 94. Wash- Chevrolat and Fauvel 1863: 443Gundlach 1891: ington 68; Blackwelder 1942: 82 (Nacaeus),1943: 126 (Pseudolispinodes Nacaeus Blackwelder 1943: 82 Distribution. Cuba, Jamaica, Hispaniola, N. dejectus (Sharp) 1887: 721 (Lispinus) Puerto Rico, Lesser Antilles, México to Costa Distribution. Cuba, Mexico, Guatemala, Rica, Colombia to Brazil, widespread se USA Costa Rica, Panama, Colombia (AZ-PA-FL) Cuba localities. Cuba localities. Ci: Cayamas. Ha: Havana Collections. Chicago Collections. Washington N. fauvelli Sharp 1887: 720 (Lispinus) N. tenuis (LeConte) 1863: 60 (Lispinus) Distribution. Cuba, Mexico, Belize, Guate- Distribution. Cuba, Mexico, Costa Rica, Co- mala, Colombia lombia, USA (CA-FL-SC) Cuba localities. Cuba localities. Collections. Chicago Collections. Chicago N. impar (Cameron) 1913: 322; Blackwelder 1943: 122 (Pseudolispinodes) Thoracophorus Motschulsky 1837: 98 Distribution. Cuba, Jamaica, Hispaniola, T. brevicristatus (Horn) 1871: 332 (Glyptoma); Montserrat, St. Lucia, St. Vincent, Trinidad Blackwelder 1943: 150; Irmler 1985: 48 Cuba localities. Distribution. Cuba, Martinique, St. Thomas, Collections. Washington Guadeloupe, México, Costa Rica, USA (AZ, FL) N. irregularis (Blackwelder) 1943: 124 Cuba localities. Ci: Cayamas (Pseudolispinodes) Collections. Chicago. Washington Distribution. Cuba, Jamaica, Hispaniola, T. guadelupensis Cameron 1913: 323; Blackwelder Puerto Rico, Guadeloupe, Mexico, Panama, 1943: 151; Irmler 1985: 54 Brazil Distribution. Cuba, St. Thomas, Guadeloupe Cuba localities. Ci: Cayamas; Soledad. Ha: to Grenada, México to Brazil Havana Cuba localities. Ci: Cayamas. SS: Mina Collections. Cambridge. Chicago. Washing- Carlota. PR ton Collections. Chicago. Washington N. laetus (Sharp) 1876: 417 (Lispinus) A Checklist of the Beetles of Cuba 67

TRIBE OSORIINI O. haitiellus Darlington 1937: 299; Blackwelder 1943: 191 Holotrochus Erichson 1839: 30 Distribution. Cuba, Hispaniola H. minor Fauvel in Chevrolat and Fauvel 1863: Cuba localities. Gu: Mts. N of Imias 437; Gundlach 189l: 63; Blackwelder 1943: 165 Collections. Cambridge. Washington (Neotrochus) O. latipes (Gravenhorst) 1906: 198 (Oxytelus); Distribution. Cuba, Jamaica, Grenada, Blackwelder 1943: 188 Trinidad, Venezuela, Brazil Distribution. Cuba, USA (TX-SD-PA-FL) Cuba localities. Ci: Cayamas; Soledad Cuba localities. Ci: Soledad Collections. Cambridge. Washington Collections. Cambridge O. manni Notman 1925: 9: Blackwelder 1943: 183 Mimogonus Fauvel 1903: 261. Distribution. Cuba. Endemic M. fumator (Fauvel) 1889: 246 (Osorius); Cuba localities. Ho: Pinares (t.l.). Gu: Blackwelder 1943: 172; Irmler 1981: 143 Yunque de Baracoa Distribution. Cuba, Jamaica, Hispaniola, Collections. Cambridge. Washington (type) Puerto Rico, Guadeloupe, Mexico, Africa, Ori- O. micros Sharp 1887: 682 ental Region Distribution. Cuba, Guatemala, Honduras, Cuba localities. Panama, Brazil Collections. Cambridge Cuba localities. Collections. Chicago Osorius Latreille 1829: 438 O. oriente Blackwelder 1943: 187 (O. buscki Notman 1925: 19) Distribution. Cuba. Endemic Distribution. Cuba? Hispaniola Cuba localities. Gu: Mts. N of Imias (t.l.) Cuba localities. Collections. Cambridge (type). Washington Collections. Chicago O. schwarzi Notman 1925: 7; Blackwelder 1943: O. crenulifrons Notman 1925: 9; Blackwelder 1943: 191 178 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. PR: San Diego de los Baños Collections. Cambridge. Chicago. Washing- (t.l.) ton (type) Collections. Washington (type) O. socors Darlington 1937: 294; Blackwelder 1943: O. cubensis Blackwelder 1943: 181 183 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Soledad.. Gu: Imias. PR: Cuba localities. Ci: Soledad (t.l.) San Antonio de los Baños (t.l.); Rangel Mts. Collections. Chicago. Cambridge (type) SS: Buenos Aires O. strictus Blackwelder 1943: 182 Collections. Cambridge. Washington (type) Distribution. Cuba. Endemic O. darlingtoni Blackwelder 1943: 177 Cuba localities. Gu: Mts. N of Imias (t.l.) Distribution. Cuba. Endemic Collections. Chicago. Cambridge (type) Cuba localities. Gu: Upper Ovando River. SC: O. turquinus Blackwelder 1943: 188 Pico Turquino (t.l.) Distribution. Cuba. Endemic Collections. Cambridge (type). Washington Cuba localities. SC: Pico Turquino (t.l.) O. eggersi Bernhauer 1904; 19; Blackwelder 1943: Collections. Cambridge (type). Chicago. 196 Washington Distribution. Cuba, Jamaica, Hispaniola, St. Thomas, Panama SUBFAMILY OXYTELINAE Cuba localities. SC: Santiago (see Herman 1970) Collections. New York O. exiguus Notman 1925: 7; Darlington 1937: 300; TRIBE THINOBIINI Blackwelder 1943: 199 Distribution. Cuba. Endemic Bledius Leach 1819: 174 Cuba localities. Ci: Cayamas (t.l.) B. beattyi Blackwelder 1943: 117; Herman 1972: Collections. Washington (type) 160; Thayer and Purrington 2004:441 68 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Puerto Rico, Virgin Is- Collections. Chicago. Washington (type) lands, Mexico, Colombia, USA (FL Keys) C. corticinus (Gravenhorst) 1806: 192 (Oxytelus) Cuba localities. Ha: Almendares; Playa C. fulvipennis (Fauvel) in Chevrolat and Fauvel Marianao 1863: 440 (Troglophloeus); Gundlach 1891: 66; Collections. Chicago. Cambridge Blackwelder 1943: 86 B. ceratus Blackwelder 1943: 118; Herman 1972: Distribution. Cuba (introduced), Jamaica, 164 cosmotropical, native to Old World Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. M: Cárdenas Bahamas, USA (FL) Collections. Chicago Cuba localities. Gu: Boqueron C. cubensis (Bierig) 1935: 15 (Troglophloeus); Collections. Cambridge. Chicago Blackwelder 1943: 71 B. cubensis Blackwelder 1943: 115 Distribution. Cuba, Jamaica Distribution. Cuba. Endemic Cuba localities. Ha: Rio Almendares; Playa Cuba localities. Ci: Cayamas (t.l.) Jaimanitas; Valle Yumurí. PR: near Rangel Collections. Chicago. Washington (type) Collections. Chicago (type) [B. jamaicensis Blackwelder 1943: 117 (Cuba C. demmeli (Bierig) 1935: 16 (Troglophloeus); record); Herman 1986: 156 (Cuba not men- Blackwelder 1943: 85 tioned, but not excluded)] Distribution. Cuba. Endemic Distribution. Jamaica, not Cuba Cuba localities. Ha: Sierra de Anafe, Caimito (B. mandibularis Erichson 1840: 765; Herman del Guayabal; Vedado 1972: 191) Collections. Chicago (type) Distribution. Bahamas, Cayman Islands, C. discipennis (Bierig) 1935: 16 (Troglophloeus); USA (widespread FL-TX-UT-MA), Canada Blackwelder 1943: 80 Cuba localities. Distribution. Cuba, Jamaica Collections. Cuba localities. Rio Almendares; Playa B. semiferrugineus LeConte 1863: 52; Herman 1972: Marianao. PR: near Cayajabos 205, 1986: 176 Collections. Chicago (type) Distribution. Cuba, e USA, Canada C. dissonus (Bierig) 1935: 11 (Troglophloeus); Cuba localities. Based on old records, needs Blackwelder 1943: 83 confirmation Distribution. Cuba. Endemic Collections. Berlin Cuba localities. Ha: Rio Almendares; Laguna La Canoa; Playa Marianao. PR: Aspiro Carpelimus Samouelle 1819: 174 Collections. Chicago (type) C. aequalis (Jacquelin du Val) 1857: 44 C. flavipes (Erichson) 1840: 808 (Troglophloeus); (Troglophloeus); Chevrolat and Fauvel 1863: Jacquelin du Val 1856: 44; Chevrolat and 441; Blackwelder 1943: 82 Fauvel 1863: 441; Gundlach 1891: 66; Troglophloeus amplicollis (Bierig) 1935: 17 (Ha: Blackwelder 1943: 77 Marianao (t.l.), Chicago, type) Troglophloeus hiemalis (Bierig) 1935: 13 (Ha: Distribution. Cuba, Bermuda, Bahamas, Playa Marianao (t.l.), Chicago (type)) Jamaica Troglophloeus flavibasis (Bierig) 1935: 13; Cuba localities. Ha: Playa Marianao Cayajobos, t.l. Collections. Chicago. Gundlach no. 1736 Distribution. Cuba, Bahamas to Grenada, C. aridus (Jacquelin du Val) 1857: 43 Mexico, Central America (Trogophloeus); Chevrolat and Fauvel 1863: Cuba localities. Ci: Cayamas. Ha: Playa 439; Gundlach 189l: 65; Blackwelder 1943: 73 Marianao. PR: Sierra Rosario, near Cayajabos; Distribution. Cuba, , Grenada, Aspiro Trinidad Collections. Chicago. Gundlach no. 1332. Cuba localities. Gu: Guantánamo; La Washington Coronella. Ci: Soledad C. fulvipes (Erichson) 1840: 804 (Troglophloeus); Collections. Cambridge. Chicago. Gundlach Jacquelin du Val 1856: 43; Chevrolat and no. 1552. Washington Fauvel l863: 439; Gundlach 1891: 65; C. conformis Blackwelder 1943: 82 Blackwelder 1943: 62 Distribution. Cuba. Endemic Troglophloeus aequalis, Gundlach 189l: 66 Cuba localities. Ha: El Cano (t.l.) (misidentification of aequalis Jacquelin du Val) A Checklist of the Beetles of Cuba 69

Troglophloeus rubripennis Fauvel in Chevrolat Thinobius Kiesenwetter 1884: 355 and Fauvel 1863: 440 T. nitidulus Bernhauer 1905: 11; Blackwelder 1943: Distribution. Cuba to Grenada, Brazil, Ar- 109 gentina, Mexico, Central America, USA (FL, Distribution. Cuba, Jamaica, Puerto Rico, HI, TX) Grenada Cuba localities. CA: Baraguá. Ci: Cayamas; Cuba localities. Ha: Mariel. PR: Aspiro; Soledad. Ha: Rio Almendares; Cayajabos. M: Cayajabos; Rangel; Sierra Rosario Cárdenas. SC: Cauto El Cristo. SS: Trinidad Collections. Chicago. Cambridge. Washing- Mts ton Collections. Cambridge. Chicago. Gundlach T. opaculus Cameron 1923: 396; Blackwelder 1943: no. 563. Washington 106 C. haplomus Blackwelder 1943: 79 T. morio Bierig 1934: 217 (t.l., Ha: Rio Distribution. Cuba, St. Croix Almendares; Chicago, type) Cuba localities. T. nebulosus Bierig 1934: 218 (t.l., Playa Collections. Chicago Marianao; Chicago, type) C. imitator (Bierig) 1935: 14 (Troglophloeus); T. cubensis Bierig 1934: 219 (t.l. not specified; Blackwelder 1943: 72 Chicago, type) Distribution. Cuba to Guadeloupe Distribution. Cuba, Jamaica, Hispaniola Cuba localities. Ci: Cayamas.PR: Aspiro, Cuba localities. Ha: Laguna Somorrostro; Rio Sierra Rosario. Ha: Rio Almendares; Cayajabos Almendares; Playa Marianao; Laguna Canoa; Collections. Chicago. Washington Sierra de Anafe del Guayabal. PR: Aspiro C. phaios Blackwelder 1943: 75 Collections. Chicago, Washington Distribution. Cuba, Jamaica T. ornatus Cameron 1923: 397; Blackwelder 1943: Cuba localities. Ci: Cayamas. PR: Viñales 105 Collections. Chicago. Washington T. tenebrophilus Bierig 1934: 215 (t.l. not speci- C. politus Kiesenwetter 1850: 221 fied; Chicago, type) Distribution. Cuba (introduced?), Japan, T. flavifer Bierig 1934: 216 (t.l. not specified; Chi- China, Poland cago, type) Cuba localities. Distribution. Cuba, Jamaica, Hispaniola Collections. Chicago Cuba localities. Ha: Rio Almendares; Sierra C. prolixus (Bierig) 1935: 12 (Troglophloeus); de Anafe, Caimito del Guayabal; La Lisa; Blackwelder 1943: 79 Vedado. PR: Sierra Organos, San Vicente Distribution. Cuba. Endemic Collections. Chicago Cuba localities. Ha: Rio Almendares; Laguna T. torrei (Bierig) 1934: 214 (Torrentomus); La Canoa; Marianao. PR: Aspiro Blackwelder 1943: 87. Collections. Chicago (type). Washington Distribution. Cuba. Endemic C. sericeus (Cameron) 1923: 392; Bierig 1935: 10; Cuba localities. Ha: Sierra del Grillo (t.l.) Blackwelder 1943: 63 Collections. Chicago (type) Troglophloeus sextagonalis (Bierig) 1935: 10 (t.l. not specified, Chicago (type)) Thinodromus Kraatz 1858: 866 Distribution. Cuba, Jamaica, St. Croix T. croceipes (Fauvel) 1863: 438; Blackwelder 1943: Cuba localities. Ha: Rio Almendares; Laguna 61 (Carpelimus) La Canoa; Sierra de Anafe, Caimito del Trogophlaeus arcuatus Fauvel in Chevrolat and Guayabal. PR: Aspiro, Sierra Rosario Fauvel 1863: 438; Gundlach 1891: 64 Collections. Chicago. Washington Distribution. Cuba, Jamaica, Hispaniola, C. varicornis (Bernhauer) 1904: 24; Blackwelder Montserrat, Guadeloupe 1943: 73 Cuba localities. PR: Sierra del Rosario; Distribution. Cuba, Hispaniola to Grenada, Hongolosongo. SC: Daiquiri. SS: San Blas, Trinidad Trinidad Mts Cuba localities. Ci: Soledad. Gu: Maisi; Collections. Cambridge. Chicago. London Sabana la Mar. SC: Cauto El Cristo (type). Washington Collections. Cambridge. Chicago. Washing- T. smithi (Bernhauer) 1908: 229 (Troglophloeus); ton Blackwelder 1943: 67; Herman 2001: 1774 70 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Hispaniola to Trinidad, Distribution. Haiti, temperate South Bolivia, Argentina America, Canada (BC), USA (CA-IN-FL); world- Cuba localities. Ci: Soledad. Ha: Guanabo; wide in temperate regions; probably native to Hongolosongo. PR: Aspiro. Africa Collections. Cambridge. Chicago. Washing- Cuba localities. ton Collections.

TRIBE Platystethus Mannerheim 1830: 46 P. spiculus Erichson 1840: 784; Blackwelder 1943: Anotylus Thomson 1859: 44 110 A. glareosus Wollaston 1854: 610 P. exiguus Jacquelin du Val 1856: 41; Chevrolat Distribution. Cuba, Jamaica, Grenada, St. and Fauvel 1863: 438; Gundlach 1891: 63 Vincent Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Ci: Cayamas Lesser Antilles, Mexico, Guatemala, Panama, Collections. Chicago Colombia, Venezuela, USA (TX, FL) A. insignitus (Gravenhorst) 1806: 188; Jacquelin Cuba localities. du Val l856: 20; Chevrolat and Fauvel 1863: Collections. Chicago 438; Gundlach 1891: 64; Blackwelder 1943: 92 (all Oxytelus); Herman 1970: 418; Hammond SUBFAMILY MEGALOPSIDIINAE 1976: 174 Distribution. Cuba, Jamaica, Hispaniola, Megalopinus Eichelbaum 1915: 104 México to Argentina, e Canada, USA M. rambouseki (Bierig) 1934: 220 (Megalopsida); Cuba localities. M: Cárdenas; Matanzas. Ci: Blackwelder 1943: 208 (Stylopodus) Cayamas. Soledad Distribution. Cuba. Endemic Collections. Chicago. Gundlach no. 409. Cam- Cuba localities. Ha: Sierra Bonilla. PR: Rio bridge. Washington Hondo, Carretera de Batanaó; Aspiro, Sierra A. scorpio Fauvel 1904: 107 Rosario Distribution. Cuba. Endemic Collections. Chicago Cuba localities. Collections. Chicago SUBFAMILY A. tetracarinatus (Block) 1799: 116 (Staphylinus) Distribution. Cuba (introduced), Canada, Stenus Latreille 1797: 77 USA; native to the Palearctic S. bakeri Bernhauer 1910: 364; Blackwelder 1943: Cuba localities. 213; Puthz 1973: 50, 1974: 48 Collections. Chicago S. mutchleri Blackwelder 1943: 218. PR: 7 km n A. vinsoni (Cameron) 1936: 201; Hammond Viñales (t.l.). Candelaria. Ci: Soledad. Cam- 1976:177 bridge. New York. Washington (type) Distribution. Cuba (introduced), Jamaica, Distribution. Cuba. Endemic Old World tropics, Pacific Islands Cuba localities. Ci: Soledad. Ha: Havana Cuba localities. (t.l.), Almendares; Laguna la Canoa. PR: Collections. Aspiro; Viñales Collections. Cambridge. Chicago (type) Oxytelus Gravenhorst 1802: 101 S. callosus Erichson 1840: 737 O. incisus Motschulsky 1857: 504; Blackwelder S. callosus juanae Puthz 1984: 138. Cuba; en- l943: 96 demic subspecies Distribution. Cuba (introduced), Bermuda, Distribution. Cuba, the nominate subspecies Jamaica, Hispaniola, Puerto Rico, widespread occurs in the eastern USA Lesser Antilles, Mexico, Panama, Canada, USA Cuba localities. Ci: Soledad. Ha: Havana; (TX, FL); cosmotropical-subtropical Laguna La Canoa. PR: San Vicente (t.l.). SC: Cuba localities. Ci: Cayamas; Soledad. CA: St. Clara. SS: Sierra Escambray Baraguá Collections. Amsterdam. Cambridge. Chi- Collections. Cambridge. Chicago. New York. cago. London. Washington (type) Washington S. columbus Puthz 1984: 122 (O. sculptus Gravenhorst 1806: 191 Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 71

Cuba localities. PR: El Sabicú (t.l.); Aspiro- Collections. Chicago (type) Rangel; El Rangel E. infidus Puthz 1997: 142 Collections. Chicago (type) Distribution. Cuba. Endemic S. cubensis Bernhauer 1910: 364; Blackwelder Cuba localities. Ha: Playa Marianao (t.l.) 1943: 225 Collections. Chicago (type) Distribution. Cuba, USA (AL-NY-FL) E. juanae Puthz 1997: 143 Cuba localities. Ci: Soledad. Ha: Havana (t.l.) Distribution. Cuba. Endemic Collections. Cambridge. Chicago (type?). Cuba localities. PR: Aspiro (t.l.). CA?: Sabicú Washington Collections. Chicago (type) S. gentilis Sharp 1886: 52; Puthz 1973: 52 S. cubanus Blackwelder 1943: 221; Puthz 1974: Euaesthetus Gravenhorst 1806:201; key to spe- 56. Ci: Cayamas (t.l.). Ha: Playa Guanabo; Rio cies in Puthz 2001: 32 Almendares; Rio Cristal. Laguna la Canoa. E. americanus Erichson 1840: 747 Washington (type) Distribution. Cuba, México?, Panama?, S. hispaniolus Blackwelder 1943: 224. Ci: Canada (ON, PQ), USA (widespread eastern) Soledad. SS: Buenos Aires. .PR: Hongolosongo. Cuba localities. Cambridge. Washington Collections. Chicago Distribution. Cuba, Hispaniola, Mexico, Gua- E. chantali Puthz 1998: 236 temala, Belize, Costa Rica, Panama, Ecuador Distribution. Cuba, Canada (ON), USA (MA, Cuba localities. As above MI, NH, TX) Collections. Chicago. Chicago. Washington Cuba localities. “Cuba” S. paludivagans Puthz 2000: 172 Collections. Chicago Distribution. Cuba. Endemic E. columbus Puthz 1998: 226 Cuba localities. M: Cienaga Zupata [Zapata], Distribution. Cuba. Endemic Playa Larga; 2 km NE Palpite. PR: San Vicente Cuba localities. Ha: Sierra de Anafe, Caimito (t.l.) del Guayabal (t.l.) Collections. Geneva. Schlitz (Puthz). Wash- Collections. Chicago (type) ington (type) E. cubanus Puthz 1998: 224 S. podagricus Sharp 1886: 667; Puthz 1984: 69, 90 Distribution. Cuba. Endemic S. dyeri Blackwelder 1943: 220 ( Washington Cuba localities. Ha: Rio Ariguanabo; Rio (type) Cristal. PR: Sierra del Rosario, Aspiro (t.l.). Distribution. Cuba, Mexico, Honduras, Nica- Laguna La Canoa ragua Collections. Chicago (type); London Cuba localities. E. juanaensis Puthz 1998: 240 Collections. Chicago Distribution. Cuba. Endemic S. sectilifer Casey 1884: 110 Cuba localities. PR: Sierra del Rosario, Aspiro S. odius Blackwelder 1943: 213. Ci: Soledad (t.l.). (t.l.). Sister species of E. atomus Casey 1884: Cambridge (type) 28 of Florida, wrongly reported from Cuba Distribution. Cuba. Jamaica, Mexico, USA Collections. Chicago (type) (DC-FL) E. pugetensis Hatch 1957: 241; Puthz 2001: 30 Cuba localities. Distribution. Cuba, Canada (widespread), Collections. Chicago USA (widespread) Cuba localities. Ha: Caimito del Guayabal; SUBFAMILY Rio Almendares; Punta Brava. PR: Aspiro Collections. Chicago Edaphus Motschulsky 1857: 7 E. cubanus Puthz 1997: 138 Octavius Fauvel 1873: 62 Distribution. Cuba. Endemic O. bermudezi Bierig 1934: 222 (Exoctavius); Cuba localities. Ha: Playa Marianao (t.l.) Blackwelder 1943: 227 Collections. Chicago (type) Distribution. Cuba. Endemic E. ferdinandae Puthz 1997: 145 Cuba localities. PR: Sierra del Rosario, Distribution. Cuba. Endemic Sumidero Rio Taco Taco (t.l.); Aspiro Cuba localities. PR: Sierra del Rosario; Collections. Chicago (type). Washington Rangel (t.l.) 72 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Tamotus Schaufuss 1877: 289 Collections. Cambridge. Chicago. Washing- T. femoratus Schaufuss 1874: 289; Blackwelder ton 1943: 227 H. solus Blackwelder 1943: 327 Distribution. Cuba. Endemic Distribution. Cuba, Jamaica, Hispaniola Cuba localities. Cuba localities. Ci: Cayamas (t.l.). Gu: Mt. Collections. Berlin? Chicago N of Imias. SC: Loma del Gato; Pico Turquino; Gran Piedra SUBFAMILY Collections. Cambridge. Chicago. Washing- TRIBE NEOTYPHLINI ton (type) H. testaceipes (Bierig) 1935: 39 (Gastrolobium); Cubanotyphlus Coiffait and Decou 1972: 131; Blackwelder 1943: 330 Frank and Thomas 1984: 1412 Gastrolobium congener Bierig 1935: 40 (PR: Si- C. jimenezi Coiffait and Decou 1972: 132 erra del Rosario, Rio Taco-Taco, t.l.; Chicago, Distribution. Cuba. EndemicAn eyeless soil type) species Distribution. Cuba, Hispaniola Cuba localities. SC: Gran Piedra, valle Rio Cuba localities. Ci: Soledad. PR: Sierra del Indio (t.l) Rosario, Rio Taco Taco. Guanabo. Ha: Sierra Collections. Bucharest (type) de Anafe, Caimito del Guayabal. Aspiro.. Gu: C. poeyi Decou 1973: 359 Maisi. PR: Rangel Mts. SS: Buenos Aires Distribution. Cuba. EndemicAn eyeless soil Collections. Cambridge. Chicago. Washing- species ton Cuba localities. Gu: Sierra Baracoa, Arroyo La Jaiba at Palenque (t.l.) Ochthephilum Stephens 1829: 287 Collections. Bucarest (type) O. atlanticum (Bierig) (Cryptobiella) 1935: 41; Blackwelder 1943: 332 SUBFAMILY PAEDERINAE Distribution. Cuba. Endemic TRIBE PAEDERINI Cuba localities. Ha: Playa Marianao; mouth SUBTRIBE ASTENINA of Rio Cojímar, in algae Collections. Chicago Astenus Stephens 1832: 275 O. darlingtoni Blackwelder 1943: 334 A. cubensis Blackwelder 1943: 367 Distribution. Cuba. Endemic Distribution. Cuba, Jamaica Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. Ci: Cayamas (t.l.); Limones; Collections. Cambridge (type). Chicago Soledad. SS: Jatibanico Collections. Cambridge. Chicago. Washing- SUBTRIBE ECHIASTERINA ton (type) Echiaster Erichson 1840: 636 SUBTRIBE CRYPTOBIINA E. microps Blackwelder 1943: 370 Distribution. Cuba, Montserrat Biocrypta Casey 1905: 26 Cuba localities. Ci: Cayamas B. cubensis Blackwelder 1943: 340 Collections. Chicago. Washington Distribution. Cuba. Endemic E. waterhousei Cameron 1913: 336; Blackwelder Cuba localities. SC: Pico Turquino (t.l.) 1943: 371 Collections. Cambridge (type). Washington Distribution. Cuba, Jamaica, Hispaniola, St. Vincent, Grenada Homaeotarsus Hochhuth 1851: 34 Cuba localities. Ha: Marianao H. albipes (Erichson) 1840: 566 (Cryptobium); Collections. Chicago. London? Blackwelder 1943: 328 Distribution. Cuba, Jamaica, Hispaniola, Sphaeronum Sharp 1876: 225 Puerto Rico, St. Vincent, Grenada, South S. berberum Blackwelder 1943: 375 America, Mexico, Guatemala Distribution. Cuba. Endemic Cuba localities. CA: Baraguá. Ci: Cayamas; Cuba localities. Ci: Soledad (t.l.) Soledad. Gr: Cauto El Cristo; Marcata, Rio Collections. Cambridge (type). Chicago Bayamo.. Gu: Imias. SC: Aguadores. A Checklist of the Beetles of Cuba 73

SUBTRIBE LATHROBIINA Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.); Soledad Lathrobium Gravenhorst 1802: 51 Collections. Cambridge. Chicago. Washing- L. rubidum (Fauvel) in Chevrolat and Fauvel 1863: ton (type) 436 (Lithocharis) 1863: 436; Gundlach 1891: A. suturalis (Blackwelder) 1943: 254 (Aderocharis) 62; Blackwelder 1943: 314 Distribution. Cuba, Jamaica, Hispaniola, Distribution. Cuba, Jamaica, Hispaniola, Puerto Rico Mexico? Cuba localities. Ci: Soledad Cuba localities. Ci: Cayamas; Soledad. M: Collections. Cambridge. Chicago Cárdenas Collections. Cambridge. Chicago. Washing- Lithocharis Boisduval and Lacordaire 1835: ton 431 L. tetricum (Casey) 1905: 110 (Tetartopeus); L. dorsalis Erichson 1840: 616; Blackwelder 1943: Watrous 1980: 320 247 L. canoaensis Bierig 1934: 325 (Tetartopaeus); Distribution. Cuba, Jamaica, Hispaniola, Blackwelder 1943: 309; Laguna La Canoa, t.l.; Puerto Rico to Trinidad Chicago, type Cuba localities. Ci: Cayamas Distribution. Cuba, USA (FL) Collections. Chicago. Washington Cuba localities. PR: Laguna La Canoa. L. heres Blackwelder 1943: 249 Collections. Chicago. Washington (type) Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.); Soledad Lobrathium Mulsant and Rey 1878: 78 Collections. Cambridge. Chicago. Washing- L. discolor (Erichson) 1840: 598; Blackwelder 1943: ton (type) 313 Distribution. Cuba, Hispaniola, St. Thomas Medon Stephens 1832: 273 Cuba localities. Ci: Soledad. SC: Aguadores M. cordatum Blackwelder 1943: 276 Collections. Cambridge. Chicago. Washing- Distribution. Cuba. Endemic ton Cuba localities. SS: Buenos Aires. (t.l.) L. odium Blackwelder 1943: 318 Collections. Chicago. Cambridge (type) Distribution. Cuba. Endemic M. cubanum Blackwelder 1943: 274 Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Chicago. Washington (type) Cuba localities. SC: Loma del Gato (t.l.); Pico Turquino Pseudolathra Casey 1905: 129 Collections. Cambridge (type). Chicago. P. margipallens (Jacquelin du Val) 1857: 41 Washington (Lathrobium); Blackwelder 1943: 316 M. schwarzi Blackwelder 1943: 275 Distribution. Cuba, Jamaica Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas; Soledad. Gu: Cuba localities. Ci: Cayamas (t.l.); Soledad. Maisi PR: Rangel Collections. Cambridge. Chicago. Washing- Collections. Chicago. Cambridge. Washing- ton ton (type) P. nitidum (Erichson) 1840: 599 (Lathrobium); Blackwelder 1943: 315 Stilomedon Sharp 1886: 565 Distribution. Cuba, Hispaniola, St. Croix, St. S. audanti Blackwelder 1943: 258 Lucia, Colombia, Uruguay, Argentina, USA Distribution. Cuba, Hispaniola, Puerto Rico, (FL, MI) St. Croix Cuba localities. SC: Aguadores Cuba localities. CA: Baraguá. Ci: Cayamas Collections. Cambridge. Chicago. Washing- (t.l.); Soledad ton Collections. Cambridge. Chicago. Washing- ton (type) SUBTRIBE MEDONINA S. connexus (Sharp) 1876: 254 (also spelled convexa) (Lithocharis); Blackwelder 1943: 257; 262 Achenomorphus Motschulsky 1858: 647 A. cubensis (Blackwelder) 1943: 251 (Aderocharis) 74 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Jamaica, Hispaniola, Distribution. Cuba. Endemic Guadeloupe to Trinidad, South America, Mexico, Cuba localities. SC: Pico Turquino (t.l.) Panama Collections. Champaign (type). Zayas Cuba localities. Collections. Chicago. Washington SUBTRIBE SCOPAEINA S. insularum (Cameron) 1923: 399 (Medon); Blackwelder 1943: 256 Orus Casey 1884: 136 Distribution. Cuba, Jamaica, Hispaniola O. cameroni Blackwelder 1943: 278 Cuba localities. Ha: Santiago de las Vegas. Distribution. Cuba, Jamaica, Hispaniola, Ci: Cayamas Grenada Collections. Chicago. Washington Cuba localities. CA: Baraguá. Ci: Cayamas Collections. Chicago. Washington Sunius Stephens 1829: 287 S. debilicornis (Wollaston) 1857: 194 (Lithocharis); Scopaeus Erichson 1840: 604 Blackwelder 1943: 267 S. angusticollis Cameron 1913: 346; Blackwelder Distribution. Cuba, Jamaica, St. Croix to 1943: 289 Trinidad, South America, Mexico, Central S. illustris Chevrolat and Fauvel 1863: 436; America, USA (SC, TX, FL) Gundlach 1891: 62 (misidentified, not from Cuba localities. Ci: Cayamas; Soledad Cuba? Blackwelder 1943: 280) Collections. Cambridge. Chicago. Washing- Distribution. Cuba, Grenada ton Cuba localities. SC: Aguadores S. ferrugineus (Bierig) 1934: 326 (Medonella); Collections. Cambridge. Chicago. Gundlach Blackwelder 1943: 262 no. 940 Distribution. Cuba. Endemic S. antennalis Cameron 1913: 349; Blackwelder Cuba localities. Ha: Playa Marianao, in al- 1943: 284 gae on beach (t.l.) Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Chicago St. Croix, Guadeloupe, St. Lucia S. occipitalis (Bierig) 1934: 329 (Xenocharis); Cuba localities. CA: Baraguá. Ci: Cayamas; Blackwelder 1943: 263 Soledad. SC: Aguadores Distribution. Cuba. Endemic Collections. Cambridge. Chicago. Washing- Cuba localities. Ci: Cayamas. Ha: Cruz de ton Piedra. PR: San Vicente, Sierra Organos; near S. darlingtoni Blackwelder 1943: 291 Rangel; Aspiro; Bejucal Distribution. Cuba. Endemic Collections. Chicago. Washington Cuba localities. SS: Trinidad Mts., Buenos Aires (t.l.) Thinocharis Kraatz 1859: 142 Collections. Cambridge (type). Chicago T. bakeri Casey 1910: 188; Blackwelder 1943: 233 S. filum Sharp 1886: 542; Blackwelder 1943: 288 Distribution. Cuba. Endemic Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Ci: Cayamas (t.l.) Guatemala Collections. Chicago. Washington (type) Cuba localities. CA: Baraguá. Ci: Cayamas; T. exilis Erichson 1840: 627; Blackwelder 1943: 237 Soledad T. pertenuis (Casey) 1910: 188 (Ci: Cayamas. t.l.; Collections. Cambridge. Chicago. Washing- Washington, type) ton Distribution. Cuba, Antigua, Dominica, St. S. illustris Chevrolat and Fauvel 1863: 436; Lucia, Trinidad, Guatemala, Panama, Colom- Gundlach 1891: 62; Blackwelder 1943: 280 (as bia, Brazil, Argentina., USA (AL, FL) suspected misidentification needing confirma- Cuba localities. Ci: Cayamas tion) Collections. Chicago. Washington Distribution. Cuba Cuba localities. SUBTRIBE PAEDERINA Collections. Gundlach no. 940 S. marginatus Cameron 1913: 349; Blackwelder Paederus Fabricius 1775: 268 1943: 286 P. zayasi Sanderson 1967: 61 (with key to species Distribution. Cuba, Jamaica of Greater Antilles) Cuba localities. Ci: Soledad A Checklist of the Beetles of Cuba 75

Collections. Cambridge. Chicago Distribution. Cuba. Hispaniola S. rambouseki (Bierig) 1934: 24 (Scopaeopsis); Cuba localities. Blackwelder 1943: 292 Collections. Chicago Distribution. Cuba. Endemic Cuba localities. CA: Baraguá. Ha: Morro; Rio Suniosaurus Bierig 1938: 139 Ariguanabo; Sierra de Anafe, Caimito del S. cuadriceps Bierig 1938: 140; Blackwelder 1943: Guayabal 361 Collections. Chicago. Washington Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. Ha: La Lisa; SUBTRIBE STILICINA Mariel; Sierra Anafe. PR: Aspiro Collections. Cambridge. Chicago Rugilus Samouelle 1819: 173 R. coprophilus (Bierig) 1936: 138 (Stilicus); TRIBE PINOPHILINI Blackwelder 1943: 305 SUBTRIBE PINOPHILINA Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas; Soledad. IJ Araeocerus Nördlinger 1836: 7 (t.l.); PR: Rangel, Sierra del Rosario A. parvipennis Bierig 1935: 30 Collections. Cambridge. Chicago. Washing- Distribution. Cuba. Endemic ton Cuba localities. Ha: La Lisa; Playa Mariano; Sierra Bonilla.. PR: Aspiro; El Rangel; Rangel SUBTRIBE STILICOPSINA Mountains. SC: Gran Piedra; Loma del Gato. SS: Buenos Aires Stamnoderus Sharp 1886: 607 Collections. Cambridge. Chicago S. labeo (Erichson) 1840: 648 (Sunius); Blackwelder 1943: 352 Pinophilus Gravenhorst 1802: 201 S. incarenatus Bierig 1936: 140 P. aguayoi Bierig 1935: 29; Blackwelder 1943: 384 S. caribaeus Bierig 1936: 141 Distribution. Cuba. Endemic S. truncatus Bierig 1936: 142 Cuba localities. Ci: Cumanayagua; Soledad. Distribution. Cuba, Jamaica, Hispaniola, Ha: Vedado. PR: Aspiro; El Rangel. SC: Puerto Rico, St. John, Guadeloupe, Dominica Aguadores Cuba localities. Ci: Cayamas; Soledad. Gu: Collections. Cambridge. Chicago. Washing- Baracoa. Ha: Sierra Bonilla. IJ: Sierra de las ton Casas. PR: Aspiro; El Rangel; Sitio Perdido; P. cubanus Blackwelder 1943: 379 Sierra del Rosario; Sierra del Grillo; El Sabicú. Distribution. Cuba. Endemic SS: Buenos Aires Cuba localities. Ci: Cayamas (t.l.); Soledad Collections. Cambridge. Chicago. Washing- Collections. Cambridge. Chicago. Washing- ton ton (type) P. flavipes Erichson 1940: 674; Blackwelder 1943: Stiliphacis Bierig 1938: 141 383 S. occipitalis Bierig 1938: 142 Distribution. Cuba, Bahamas, Cayman Is- Distribution. Cuba. Endemic lands, Hispaniola, Puerto Rico Cuba localities. PR: Sierra del Rosario (t.l.) Cuba localities. Ca: Central Jaronú. CA: Collections. Chicago Baraguá. Ci: Cayamas; Soledad. Ha: Laguna La Canoa Stilosaurus Blackwelder 1943: 348 Collections. Cambridge. Chicago. Washing- S. coronalis Blackwelder 1943: 350 ton Distribution. Cuba. Endemic P. schwarzi Blackwelder 1943: 386 Cuba localities. Ci: Cayamas (t.l.); Soledad Distribution. Cuba, Cayman Islands Collections. Cambridge. Chicago. Washing- Cuba localities. Ci: Cayamas; Soledad ton (type) Collections. Cambridge. Chicago. Washing- ton Suniophacis Blackwelder 1943: 345 P. subterraneus Blackwelder 1943: 380 S. thoracica(Cameron) 1923: 398; Blackwelder Distribution. Cuba. Endemic 1943: 346 Cuba localities. SS: Buenos Aires (t.l.) 76 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Cambridge (type). Chicago Distribution. Cuba, Jamaica, Hispaniola, Puerto Rico to Trinidad, Central America, SUBTRIBE PROCIRRINA South America, North America Cuba localities. Ci: Cayamas; Soledad. PR: Palaminus Erichson 1840: 681 Aspiro P. anacoreta Bierig 1935: 34; Blackwelder 1943: Collections. Cambridge. Chicago. Washing- 388 ton Distribution. Cuba. Endemic Cuba localities. Ha: Portugalete (t.l.) TRIBE XANTHOLININI Collections. Chicago (type) P. bermudezi Bierig 1935: 35; Blackwelder 1943: Lithocharodes Sharp 1876: 204 389 L. cameroni Blackwelder 1943: 497 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: , Sierra del Rosario (t.l.); Cuba localities. Ci: Cayamas (t.l.) El Rangel Collections. Chicago. Washington (type) Collections. Chicago (type) L. floridanus (Leconte) 1880: 170 (Metaponcus) P. bruchianus Bierig 1935: 36; Blackwelder 1943: Distribution. Cuba, Mexico, se USA 389 Cuba localities. Distribution. Cuba. Endemic Collections. Chicago Cuba localities. Ha: Playa Marianao (t.l.) L. rambouseki Blackwelder 1943: 500 Collections. Chicago (type) Distribution. Cuba. Endemic P. coriaceus Cameron 1913: 331; Blackwelder 1943: Cuba localities. Ha: Rio Almendares (t.l.) 389 Collections. Chicago. Washington (type) Distribution. Cuba, Jamaica, Hispaniola Cuba localities. Ci: Cayamas Neohypnus Coiffait and Sáiz 1964: 522 Collections. Chicago. Washington N. attenuatus (Erichson) 1839: 330 (Xantholinus); P. hylaeus Bierig 1935: 32; Blackwelder 1943: 390 Chevrolat and Fauvel 1863: 430; Gundlach Distribution. Cuba. Endemic 1891: 56; Blackwelder 1943: 478 Cuba localities. Ha: Playa Marianao; Sierra Distribution. Cuba, Jamaica, Puerto Rico to Bonilla. PR: Aspiro,;Cayajabos; Coralillo Trinidad, Mexico, Venezuela, Brazil, Paraguay, Collections. Chicago (type) Argentina P. peralutaceus Bierig 1935: 37; Blackwelder 1943: Cuba localities. M: Cárdenas; Soledad. Ha: 391 Almendares Distribution. Cuba. Endemic Collections. Cambridge. Chicago. London? Cuba localities. PR: El Rangel (t.l.) Gundlach no. 495 Collections. Chicago (type) N. humeralis (Erichson) 1840: 327 (Xantholinus); P. pilum Bierig 1935: 33; Blackwelder 1943: 391 Blackwelder 1943: 479 Distribution. Cuba. Endemic Xantholinus puncticeps Jacquelin du Val 1857: Cuba localities. Ha: Sierra Anafe; 35; Chevrolat and Fauvel 1863: 430; Gundlach Portugalete (t.l.). PR: Sierra del Rosario 1891: 57 Collections. Chicago (type, female) Xantholinus punctiger Gemminger and Harold P. rosariensis Bierig 1935: 38; Blackwelder 1943: 1868: 603 392 Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba. Endemic St. John, St. Croix, Lesser Antilles Cuba localities. PR: Cayajabos; Sumidero Rio Cuba localities. Ci: Cayamas Taco Taco Collections. Chicago. Washington Collections. Chicago (type) Neoxantholinus Cameron 1944: 783 SUBFAMILY STAPHYLININAE N. testaceipennis (Fauvel) in Chevrolat and Fauvel TRIBE DIOCHINI 1863: 431 (Leptacinus); Blackwelder 1943: 507 (Oligolinus) Diochus Erichson 1839: 307 Leptacinus pallidipennis Fauvel 1863: 431; D. nanus Erichson 1839: 301; Chevrolat and Fauvel Gundlach 1891: 57 1863: 430: Blackwelder 1943: 454 Distribution. Cuba, Hispaniola A Checklist of the Beetles of Cuba 77

Cuba localities. Widespread A. insulanus Bierig 1938: 125; Blackwelder 1943: Collections. Cambridge. Chicago. Gundlach 468 no. 64. Washington Distribution. Cuba. Endemic N. variabilis (Blackwelder) 1943: 508 (Oligolinus) Cuba localities. Ha: Mariel; Laguna de Distribution. Cuba, Hispaniola Canoa. IJ. Cuba localities. Gu: Mts. N of Imias Collections. Cambridge. Chicago?Washington Collections. Chicago. Cambridge Heterothops Stephens 1829: 284 Phacophallus Coiffait 1956: 50 H. rambouseki Blackwelder 1943: 465 P. parumpunctatus (Gyllenhal) 1827: 481 Distribution. Cuba, Jamaica (Staphylinus); Blackwelder 1943: 494 Cuba localities. Ci: Cayamas. Ha: (Leptacinus) Havana(t.l.) Distribution. Cuba, Jamaica, Puerto Rico, Collections. Chicago. London. Washington St. Kitts to Grenada, North America, etc.; prob- ably introduced from Palearctic SUBTRIBE TANYGNATHININA Cuba localities. Collections. Chicago. Washington Atanygnathus Jacobson 1809: 521 A. laticollis (Erichson) 1839: 289 (Tanygnathus); Stenolinus Bierig 1937: 272 Blackwelder 1943: 472 S. macrothrichus Bierig 1937: 275; Blackwelder Distribution. Cuba, Hispaniola, Puerto Rico, 1943: 503 Antigua, St. Vincent, Trinidad Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. PR: Rangel Mts. Cuba localities. PR: El Rangel (t.l.) VC: Hanabanillo Falls Collections. Chicago (type) Collections. Cambridge. Chicago. Washing- ton Xantholinus Dejean 1821: 23. See Smetana 1982. The following species of Xantholinus may be- SUBTRIBE STAPHYLININA long to other genera. X. beattyi Blackwelder 1943: 485 Creophilus Samouelle 1819: 172 Distribution. Cuba, Hispaniola, St. Croix C. maxillosus (Linnaeus) 1758: 421; Blackwelder Cuba localities. Ci: Soledad. SC: Gran Pie- 1943: 447 dra; Loma del Gato. SS: Buenos Aires Staphylinus villosus Gravenhorst 1802: 160; Collections. Cambridge. Chicago. Washing- Jacquelin du Val 1856: 36 ton C. villosus (Gravenhorst); Chevrolat and Fauvel X. cubensis Blackwelder 1943: 480 1863: 432; Gundlach 1891: 57 Distribution. Cuba. Endemic Distribution. Cuba, Jamaica, Central Cuba localities. Gu: Imias. Ha: Sierra Bonilla America, widespread North America, etc. (t.l.). SC: Gran Piedra. SS: Trinidad Mts Cuba localities. IJ: Los Indios Collections. Cambridge. Chicago. Washing- Collections. Carnegie (C. villosus). Chicago. ton Gundlach no. 491 X. impunctus Blackwelder 1943: 487 Distribution. Cuba. Endemic Platydracus Thomson 1858: 29 Cuba localities. Ci: Soledad (t.l.) P. tomentosus (Gravenhorst) 1802: 161 Collections. Cambridge (type). Chicago (Staphylinus) P. antiquus Nordmann 1837: 35 (Staphylinus) TRIBE P. antiquus of Blackwelder 1943: 446 SUBTRIBE QUEDIINA (misidentified, not P. antiquus Nordmann 1837: 35) Acylophorus Nordmann 1837: 127 Ocypus cubae Jacquelin du Val 1857: 36; A. hatuey Bierig 1938: 120; Blackwelder 1943: 469 Chevrolat and Fauvel 1863: 432; Gundlach 1891: Distribution. Cuba. Endemic 58 Cuba localities. PR: Aspiro; El Rangel; La- Distribution. Cuba, Trinidad, South guna Somorrostro America, Central America, eastern Nearctic Collections. Chicago (type) Cuba localities. 78 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Chicago Distribution. Cuba, Jamaica, Mona, Puerto Rico, St. Croix, St. Kitts, Antigua, Montserrat, SUBTRIBE PHILONTHINA Guadeloupe, Dominica, USA (FL) Cuba localities. Ha: Playa Marianao Belonuchus Nordmann 1837: 129 Collections. Chicago. Washington B. agilis Erichson 1840: 423; Jacquelin du Val 1856: 37; Chevrolat and Fauvel 1863: 433; Gundlach Gabronthus Tottenham 1955: 178 1891: 58; Blackwelder 1943: 327 G. maritimus (Motschulsky) 1858: 661 (Gabrius) B. chevrolati Fauvel in Chevrolat and Fauvel Distribution. Cuba (introduced?), Europe, 1863: 432 Congo, South Africa, Mauritius, Reunion, Phil- Distribution. Cuba, Jamaica ippines Cuba localities. Widespread Cuba localities. Collections. Chicago. Gundlach no. 302. Collections. Chicago Washington (G. mgogoricus Tottenham 1955: 183; Smetana B. gagates Erichson 1840: 424; Jacquelin du Val 1995: 484) 1856: 38: Chevrolat and Fauvel 1863: 433; Distribution. Jamaica, Hispaniola, Puerto Gundlach 1891: 59; Blackwelder 1943: 424; Rico, Virgin Islands, USA (FL, MS, NC), in- Smetana 1995: 737 troduced from Africa; in many kinds of decay- Distribution. Cuba, Bahamas, Jamaica, ing organic materials Hispaniiola, Puerto Rico to Montserrat, USA Cuba localities. (s FL) Collections. Cuba localities. Ci: Cayamas; Soledad. Ha: Marianao; Havana; Santiago de la Vega Neobisnius Ganglbauer 1895: 484 Collections. Cambridge. Chicago. Gundlach N. demmeli Bierig 1933: 54; Frank 1981: 32 no. 305. London Washington Distribution. Cuba. Endemic (B. rufipennis (Fabricius) 1801: 597; Smetana 1995: Cuba localities. PR: Cayajabos; Sierra del 726) Rosario (t.l.). Ci: Soledad Distribution. Jamaica, Central America, Collections. Cambridge. Chicago, (type). South America, USA (AZ, IN-VA-FL) Washington Cuba localities. N. humilis (Erichson) 1840: 512 (Philonthus); Collections. Chevrolat and Fauvel 1863: 435; Gundlach 1891: 61; Blackwelder 1943: 441 (Erichsonius); Cafius Stephens 1829: 283 Frank 1981: 18 C. bistriatus (Erichson) 1840: 502; Blackwelder Lathrobium margipallens Jacquelin du Val 1856: 1943: 438; Frank 1985: 61; Frank et al. 1986: 41 149 Distribution. Cuba, Jamaica, Hispaniola, Distribution. Cuba. Bahamas, Jamaica, Puerto Rico to Grenada Puerto Rico to Trinidad, South America, USA Cuba localities. Widespread (ME-FL), Canada Collections. Cambridge. Chicago. Gundlach Cuba localities. Ha: Playa de Marianao no. 564. Washington Collections. Chicago N. nigrocoeruleus (Cameron) 1922: 118; Frank C. caribeanus Bierig 1934: 68; Blackwelder 1943: l981: 26 437; Frank 1985: 61 Neobisnius carbonarius (Bierig) 1933: 49 (Sierra Distribution. Cuba, Jamaica, Puerto Rico to del Rosario, t.l.; Chicago, type) Grenada, Trinidad, South America, Mexico, Distribution. Cuba, Jamaica, Hispaniola Panama, USA (FL) Cuba localities. PR: Cayajabos; El Rangel; Cuba localities. Ha: Playa de Marianao Aspiro; Rio Taco Taco Collections. Chicago. Washington Collections. Chicago. Washington C. rufifrons Bierig 1934: 68; Frank et al. 1986: 153 Distribution. Cuba, USA (FL) Philonthus Stephens 1929: 279 Cuba localities. Ha: Playa de Marianao (t.l.) P. alumnus Erichson 1840: 490; Smetana 1995: Collections.. Chicago (type) 434 C. subtilis Cameron 1922: 121; Blackwelder 1943: Distribution. Cuba, Cayman Islands, 436; Frank 1985: 62 Hispaniola, Puerto Rico A Checklist of the Beetles of Cuba 79

Cuba localities. Collections. Chicago. Washington Collections. Washington (P. neptunus Smetana 1995: 436) P. caribaeus Bierig 1940: 141; Blackwelder 1943: Distribution. Jamaica, Bahamas Islands, 415 USA (s FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. PR. Ha. T.l. not specified Collections. Collections. Chicago (type) P. rufulus Horn 1884: 197; Smetana 1995: 430 P. discoideus (Gravenhorst) 1802: 38 (Staphylinus); Distribution. Cuba, Bahamas, México, wide- Jacquelin du Val 1856: 40; Chevrolat and spread USA Fauvel 1863: 435; Gundlach 1891: 61; Cuba localities. Ci: Cayamas Blackwelder 1943: 407; Smetana 1995: 184 Collections. Chicago. Washington Distribution. Cuba (introduced), Jamaica, P. varians (Paykull) 1789: 45; Jacquelin du Val Hispaniola, Puerto Rico to Grenada, widespread 1856: 39; Chevrolat and Fauvel 1863: 435; North America; a Palaearctic species which is Gundlach 1891: 60; Blackwelder 1943: 412 now cosmopolitan (Smetana 1995: 184) Distribution. Cuba (introduced) St. John, St. Cuba localities. Vincent, North America, etc., native to the Old Collections. Chicago. Gundlach no. 1828. World. The Neotropical records need confirma- London tion (Smetana 1995: 237) P. figulus Erichson 1840: 464; Chevrolat and Cuba localities. Fauvel 1863: 434; Gundlach 1891: 61; Collections. Chicago. Gundlach no. 787 Blackwelder 1943: 409 P. ventralis (Gravenhorst) 1802: 174 (Staphylinus); P. amazonicus Sharp 1876: 165, in Leng and Blackwelder 1943: 404; Smetana 1995: 190 Mutchler 1914: 406 Distribution. Cuba (introduced), Jamaica, Distribution. Cuba, Puerto Rico, St. Vincent, Hispaniola, Puerto Rico to Trinidad, Central Trinidad, South America, Central America America, South America, widespread North Cuba localities. Ha: Havana America, etc.; native to the Old World Collections. Chicago Cuba localities. Cayamas; Soledad (P. flavolimbatus Erichson 1840: 471; Smetana Collections. Cambridge. Chicago. Washing- 1995: 200) ton Distribution. West Indies, México, Central America, South America, widespread in s USA Remus Holme 1837: 64 Cuba localities. R. pruinosus (Erichson) 1840: 510 (Cafius); Bierig Collections. 1934: 67 P. havaniensis (Laporte) 1835: 116; Blackwelder Distribution. Cuba (introduced), native to 1943: 418; Smetana 1995: 434 Europe P. obscurus Fauvel in Chevrolat and Fauvel Cuba localities. 1863: 434; Fauvel 1891: 115 (synonomy); Collections. Chicago Gundlach 1891: 60 Distribution. Cuba, Jamaica, Hispaniola, SUBTRIBE HYPTIOMINA Puerto Rico to Antigua Cuba localities. Ci: Cayamas; Soledad Holisus Erichson 1839: 298 (see Newton 1988) Collections. Cambridge. Chicago. Washing- H. debilis Erichson 1839: 299; Blackwelder 1943: ton 461 P. hepaticus Erichson 1840: 451; Blackwelder 1943: Holisus cubensis (Casey) 1906: 362 (Havana, t.l.); 402; Smetana 1995: 175 Leng and Mutchler 1914: 406 P. vilis Erichson 1840: 451; Jacquelin du Val Distribution. Cuba, Bahamas, Hispaniola, 1856: 39; Chevrolat and Fauvel 1863: 433; Puerto Rico to St. Vincent Gundlach 1891: 59 Cuba localities. Ci: Soledad Distribution. Cuba, Bahamas, Jamaica, Collections. Cambridge. Chicago. Washing- Hispaniola, Puerto Rico to Trinidad, South ton America, Central America, North America, etc. H. laevis Blackwelder 1943: 463 Some Neotropical records may be in error (see Distribution. Cuba, Jamaica Smetana 1995: 177) Cuba localities. Gu: Mts. N of Imias Cuba localities. Ci: Cayamas Collections. Cambridge. Chicago 80 Arthropods of Florida and Neighboring Land Areas, Vol. 18

REFERENCES Hanley, R.S. 1998. Corrections to the nomencla- Blackwelder, R.E. 1943. Monograph of the West ture of a species of Neotropical Hoplandria Indian beetles of the family Staphylinidae. U.S. Kraatz (Coleoptera: Staphylinidae: National Museum Bull. 182. 658 pp. Aleocharinae: ). Coleop. Bull. 52: Campbell, J. M. 1975. A revision of the genera 386-387. Coproporus and Cilea (Coleoptera: Herman, L.H., Jr. 1970. Phylogeny and reclassi- Staphylinidae) of America north of Mexico. Can. fication of the rove-beetle subfamily Oxytelinae Ent. 107: 175-216. of the world (Coleoptera, Staphylinidae). Bull. Chandler, D.S. and Wolda, H. 1986. Seasonal- Amer. Mus. Nat. Hist. 142: 343-454. ity and diversity of Caccoplectus, with a re- Herman, L.H., Jr. 1972. Revison of Bledius and view of the genus and description of a new ge- related genera. Part I. The aequatorialis, nus, Caccoplectinus (Coleoptera: Pselaphidae). mandibularis, and semiferrugineus groups and Zoologische Jahrbücher, Abteilung für two new genera (Coleoptera, Staphylinidae, Systematik, Ökologie und Geographie der Tiere Oxytelinae). Bull. Am. Mus. Nat. Hist. 149(2). 113: 469-524. Herman, L.H., Jr. 1986. Revison of Bledius and Coiffait, H. and Decou, V. 1972. Recherches sur related genera. Part IV. Classification of spe- les Coléopterès hypogés de Cuba. I. cies groups, phylogeny, natural history, and Staphylinidae-Leptotyphlinae: Cubanotyphlus catalogue (Coleoptera, Staphylinidae, jimenezi n. g., n. sp. Rev. Ecol. Biol. Sol 9: 131- Oxytelinae). Bull. Am. Mus. Nat. Hist. 184: 138. 13-67. Decou, V. G. 1973. Recherches sur les Coléoptères Herman, L.H., Jr. 2001. Cataogue of the hypogés de Cuba. II. Staphylinidae- Staphylinidae (Insecta: Coleoptera) 1758 to the Leptotyphlinae: Cubanotyphlus poeyi n.sp. end of the second millenium. Bull. Amer. Mus. Résultats des expeditions biospéologiquess Cu- Natur. Hist. 265, 4218 pp. ban-Roumaines á Cuba. 1: 359-363. Irmler, U. 1981. Neue Arten der Gattung Frank, J.H. 1972. The genus Oligota in the Car- Mimogonia Coiffait (1978) aus der Neotropis ibbean region (Coleoptera: Staphylinidae). (Coleoptera, Staphylinidae). Ent. Blätter 77: Coleop. Bull. 26: 125-141. 143-152. Frank, J.H. 1981. A revision of the New World Irmler, U. 1985. Neue Arten der Gattungen species of the genus Neobisnius Ganglbauer. Aneucamptus und Thoracophorus (Col., Occ. Pap. Fla. St. Colln. Arthrop. 1: 1-160. Staphylinidae) aus der Neotropis. Ent. Blätter Frank, J.H. 1985. New records of Cafius 81: 41-58. caribeanus and C. subtilis in Florida and Ven- Irmler, U. 1991. Neue Arten des Gattung ezuela (Coleoptera: Staphylinidae). Ent. News Clavilispinus Blackwelder (Col., Staphylinidae) 96: 61-62. aus der Neotropis. Ent. Blätter 87: 85-91. Frank, J.H. 1986. A preliminary checklist of the Irmler, U. 1994: Taxonomie und Verbreitung Staphylinidae (Coleoptera) of Florida. Florida neotropischer Lispinus Er. (Coleoptera, Ent. 69: 363-382. Staphylinidae). Beitr. Ent. 44: 53-82. Frank, J.H., Carlysle, T.C., and Rey, J.R. 1986. Irmler, U. 2000. Lectotype designation of Lispinus Biogeography of the seashore Staphylinidae granadensis Fauvel, 1865 and Lispinus Cafius bistriatus and C. rufifrons (Insecta: insularis Chevrolat & Fauvel, 1863 with de- Coleoptera). Florida Scientist 49: 148-161. scription and distribution of the related species Frank, J.H., J. Klimaszewski, and S.B. Peck. (Coleoptera: Staphylinidae: Osoriinae). Bull. 1989. Leptagria perexilis and Myllaena audax Inst. Royal Sci. Natur. Belgique, Entomologie (Coleoptera: Staphylinidae) in Florida. Florida 70: 81-88. Ent. 72: 717-718. Klimaszewski, J. 1979. A revision of the Frank, J.H. and M.C. Thomas. 1984. and Deinopsini of the world (Co- Cubanotyphlus largo, a new species of leoptera, Staphylinidae, Aleocharinae). Agric. Leptotyphlinae (Coleoptera: Staphylinidae) Canada Monog. 25: 1-169. from Florida. Canadian Ent. 116: 1411-1417. Klimaszewski, J. 1980. Two new species of Hammond, P.M. 1976. A review of the genus Deinopsini from the Afrotropical and Nearctic Anotylus C. G. Thomson (Coleoptera: regions, with notes on two other species of this Staphylinidae). Bull. Brit. Mus. (Nat. Hist.), tribe (Coleoptera, Staphylinidae). Polskie Pismo Ent. 33: 139-187. Ent. 50: 109-120. A Checklist of the Beetles of Cuba 81

Klimaszewski, J. 1982. Stydies of Myllaenini (Co- Pace, R. 1990. Nuovi Falagriini, Hoplandriini et leoptera: Staphylinidae, Aleocharinae). 1. Sys- Aleocharini della regine Neotropica (Coleoptera, tematics, phylogeny, and zoogeography of Ne- Staphylinidae) (LXXX contributo alla Myllaena Erichson. Canadian Ent. 114: conoscenza delle Aleocharinae). Giornale 181-242. Italiano di Entomologia 5: 157-180. Klimaszewski, J. 1990. Two new species and new Park, O., J.A. Wagner, and M.W. Sanderson. records of Aleochara from Latin America (Co- 1976. Review of the pselaphid beetles of the leoptera: Staphylinidae). Ann. Transvaal Mus. West Indies (Coleoptera: Pselaphidae). Fieldiana 35: 171-176. Zoology 68: 1-90. Klimaszewski, J., F. Génier. 1986. Lectotype Puthz, V. 1973. Zur Synonymie und Stellung designation and redescription of Aleochara einiger Stenus-Arten IV Coleoptera, verberans (Coleoptera: Staphylinidae). Ent. Staphylinidae). 136. Beitrag zur Kenntnis der News 97: 115-118. Steninen. Ent. Tidskr. 94: 48-55. Klimaszewski, J., Génier, F., and Uhlig, M. Puthz, V. 1974. Studies on the Neotropical spe- 1987. Review of Erichson’s types of Aleochara cies of Stenus described by Bernhauer (Co- from México, West Indies, and South America. leoptera, Staphylinidae); Seventy-eighth contri- Florida Ent. 70: 249-259. bution to the knowledge of Steninae. Fieldiana Klimaszewski, J., Uhlig, M. and Maus, C. Zoology 65: 45-72. 2000. Diversity of Aleochara species in Mada- Puthz, V. 1984. Über neue und alte neotropische gascar (Coleoptera Staphylinidae Aleocharinae). Stenus (Hypostenus)-Arten (Coleoptera, Belgian J. Ent. 2: 227-256. Staphylinidae). 198. Beitrag zur Kenntnis der Lawrence, J.F. and A.F. Newton. 1995. Fami- Steninen. Ent. Blättter 79: 65-140. lies and subfamilies of Coleoptera with selected Puthz, V. 1997. Beiträge zur Kenntnis der genera, notes, references and data on family Euaesthetinen LXXVIII. Neotropische group names. 779-1106. In: J. Pakaluk and Edaphus-Arten mit Atriumtaschen an den S.A. Slipinski (eds.). Biology, phylogeny, and Abdominaltergiten 2-4 (Staphylinidae, Co- classification of Coleoptera: papers celebrating leoptera). Philippia 8: 137-145. the 80th birthday of Roy A. Crowson. Muzeum Puthz, V. 1998. Beiträge zur Kenntnis der i Instytut Zoologii PAN, Warsaw. Euaesthetinen LXXIX. Neuweltliche Leschen, R.A.B., and I. Löbl. 1995. Phylogeny Euaesthetus-Arten (Staphylinidae, Coleoptera). of Scaphidiinae with redefinition of tribal and Philippia 8: 223-244. generic limits (Coleoptera: Staphylinidae). Rev. Puthz, V. 2000. Beiträge zur Kenntnis der Suisse Zool. 102: 425-474. Steninen. CLXII. Neue und alte neotropische Navarrete-Heredia, J.L., A. F. Newton, M.K. Arten der Gattung Stenus Latreille Thayer, J.S.Ashe, D.S. Chandler. 2002. (Staphylinidae, Coleoptera). Philippia 9: 165- Guía ilustrada para los géneros de Staphylinidae 213. (Coleoptera) de México. Illustrated guide to the Puthz, V. 2001. Neotropical Euaesthetus genera of Staphylinidae (Coleoptera) of Mexico. Gravenhorst (Coleoptera: Staphylinidae) (82nd Universidad de Guadalajara, CONABIO, contribution to the knowledge of Mexico, D.F. Euaesthetinae). Dugesiana 8: 29-36. Newton, A.F. 1988. Fooled by flatness: subfamily Sanderson, M.W. 1967. Revision of the Greater shifts in subcortical Staphylinidae (Coleoptera). Antilean species of Pederus (Coleoptera: Coleop. Bull. 42: 255-262. Staphylnidae). Coleop. Bull. 21: 59-62. Newton, A.F. and M.K. Thayer. 1995. Sheerpeltz, O. 1952. Revision der Gattung Piestus Protopselaphinae new subfamily for Gravh. (Coleoptera Staphylinidae). Rev. Chilena Protopselaphus new genus from Malaysia, with Ent. 2: 281-305. a phylogenetic analysis and review of the Smetana, A. 1982. Revision of the subfamily Omaliine group of Staphylinidae including Xantholininae of America north of Mexico (Co- Pselaphidae (Coleoptera). 219-320. In: J. leoptera: Staphylinidae). Mem. Ent. Soc. Pakaluk and S.A. Slipinski (eds.): Biology, phy- Canada 120. 389 pp. logeny, and classification of Coleoptera: papers Smetana, A. 1995. Rove beetles of the subtribe celebrating the 80th birthday of Roy A. Philonthina of America north of México (Co- Crowson. Muzeum i Instytut Zoologii PAN, leoptera: Staphylinidae); Classification, phylog- Warsaw. 82 Arthropods of Florida and Neighboring Land Areas, Vol. 18

eny and taxonomic revision. Mem. Ent. [P. coniferus Escholtz 1829: 16] Internat. 3: 1-946. Distribution. Ecuador, Brazil, Peru, Bolivia; Thayer, M.K., F.F. Purrington. 2004. Bledius not Cuba (rejected by Reyes-Castillo et al. 1995: beattyi Blackwelder (Coleoptera: Staphylinidae: 32) Oxytelinae), new to the United States, and co- P. interstitialis Escholtz 1829: 18; Jacquelin du Val occurrences of some Bledius Leach and 1857: 137; Chevrolat 1864c: 408; Gundlach Dyschiriodes Jeannel (Coleoptera: Carabidae: 1891: 153 Scaratinae) species in south Florida. Coleop. Passalus convexus Schönherr 1817: 335; Bull. 589: 439-442. Jacquelin du Val 1857: 138; Chevrolat 1864c: Watrous, L. 1980. Lathrobium (Tetartopeus): 408; Gundlach 1891: 153 (no. 45) natural history, phylogeny, and revision of the Distribution. Cuba, Jamaica, México to Ar- Nearctic species (Coleoptera, Staphylinidae). gentina; ecology (Rodriguez 1985, Rodriguez and Syst. Ent. 5: 303-338. Zorilla 1986 Cuba localities. Widespread. SERIES SCARABAEIFORMIA Collections. Gundlach no. 46. Pittsburgh. SUPERFAMILY SCARABAEOIDEA [P. morio Percheron 1835: 83] Reviewed by H. F. Howden, September, 2002. Distribution. México to Argentina.; not Cuba (rejected by Reyes-Castillo et al. 1995: 29) 29. FAMILY LUCANIDAE [P. pentaphyllus Palisot de Beauvois 1805: 2; This family occurs in the USA through México Jacquelin du Val 1857: 137; Chevrolat 1864c: to South America but does not occur in the West 410; Gundlach 189l:155; Ivie and Gillogly 1998: Indies. Larvae feed on rotted wood of old logs and 2.] stumps. Distribution. Hispaniola; not Cuba P. pertyi Kaup 1869: 22 30. FAMILY DIPHYLLOSTOMATIDAE Passalus pelliculatus Percheron 1835: 77; This family is limited to three species of Jacquelin du Val 1857: 139; Chevrolat 1864c: Diphyllostomus in California. Females are flight- 409; Gundlach 1891: 154 less, and larvae probably live in soil. Distribution. Cuba. Endemic: records from Hispaniola doubtful ( Reyes-Castillo et al. 1995: 31. FAMILY PASSALIDAE 32) The bess beetles Cuba localities. Reviewed by J. Schuster, June 1994 Collections. These beetles are found in sub-social groups in rotting logs in forests. The genus Spasalus has more Spasalus Kaup 1869: 27 than 3 lamellae in their antennae, and old records [S. crenatus (MacLeay) 1819: 106 (Paxillus)] of these for Cuba need to be confirmed before being Distribution. Puerto Rico, Virgin Islands, accepted. S. Peck has a manuscript key to the Lesser Antilles; not Cuba, Hispaniola (Ivie and known Cuban species by J. Schuster. Ivie and Gillogly 1998: 1) Gillogly (1998) is the principal reference. Cuba localities. Collections. SUBFAMILY PASSALINAE TRIBE PASSALINI REFERENCES Ivie, M. A. and Gillogly, A.R. 1998. Passalidae Passalus Fabricius 1792: 240 of the West Indies. http://virgin.msu. [P. affinis Percheron 1835: 72; Chevrolat 1864c: montana.edu/WestIndies/Polyphaga/ 409; Gundlach 1891: 154] Scarabaeoidea/WIPassalidae.html Distribution. Hispaniola; not Cuba (rejected Reyes-Castillo, P., Maes, J.M. and Guerrero, by Reyes-Castillo et al. 1995: 30) K.A. 1995. Los Passalidae (Coleoptera: [P. binominatus Percheron 1844: 23; Jacquelin du Scarabaeoidea) de la Espanola, Grandes Val 1857: 138; Chevrolat 1864c: 409; Gundlach Antillas. Rev. Soc. Mex. Hist. Nat. 46: 29-34. 189l: 154) Rodriguez, M.E. 1985. The role of Passalus Distribution. French Guiana, Brazil, Argen- interstitialis (Coleoptera: Passalidae) in the tina; not Cuba (rejected by Reyes-Castillo et al. initiation of wood decomposition at the Sierra 1995: 29) del Rosario Ecological Station. I. Activity un- A Checklist of the Beetles of Cuba 83

der natural conditions. Cien. Biol. no. 13: 29- 34. FAMILY PLEOCOMIDAE 38. This family contains only the genus Pleocoma, Rodriguez, M.E. and M.A. Zorilla. 1986. The which is limited to western North America. The role of Passalus interstitialis (Coleoptera: larvae feed on roots. Passalidae) in the initiation of wood decompo- sition at the Sierra del Rosario Ecological Sta- 35. FAMILY GEOTRUPIDAE tion. II. Activity under laboratory conditions. The earth-boring scarab beetles Cien. Biol. 16: 69-76. Adults of this family dig deep burrows, and store food (such as fungi, leaves, litter, dung, or carrion) 32. FAMILY GLARESIDAE for their larvae. The family is composed of the genus , with some 50 species worldwide except Australia. SUBFAMILY GEOTRUPINAE These beetles usually live in sandy areas in arid TRIBE ATHYREINI regions. They are probably absent from the Carib- bean Islands. Neoathyreus Howden and Martinez 1963: 347 N. androsensis Howden 1996: 1505 33. FAMILY TROGIDAE Distribution. Cuba, Bahamas (Andros) The hide beetles Cuba localities. CA: Cayo Coco Adults and larvae feed on old dry carcasses of Collections. Howden birds or mammals, and on acculumations of fur N. castaneus (Guérin Ménèville) 1830: 83 and feathers. (Athyreus); Jacquelin du Val 1856: 122; Chevrolat 1864c: 415; Gundlach 1891: 132; Omorgus Erichson 1847: 111. Howden 1995: 1507 O. suberosus (Fabricius) 1775: 31; Vaurie 1955: 60 (Trox) Athyreus angulatus Klug 1843: 28 Trox crenatus Olivier, Gundlach 1891: 132 Athyreus tridentatus castaneus Guérin-Ménèville Trox punctatus Germar; misidentified in 1829: 83 Jacquelin du Val 1856: 121; Chevrolat 1864c: Distribution. Cuba. Endemic 416 Cuba localities. Widespread Distribution. Cuba, widespread West Indies, Collections. Gatineau. Gundlach no. 263. México to Brazil, widespread USA Paris (type?) Cuba localities. Widespread Collections. Gundlach no. 1 (T. crenatus) REFERENCES O. tytus (Robinson) 1941: 228; Vaurie 1955: 57 Howden, H.F. 1996. West Indian Neoathyreus and (Trox) their over-water dispersal (Coleoptera: Distribution. Cuba, USA (AR, OK, GA); in Scarabaeidae: Geotrupinae). J. Nat. Hist. 30: barn owl nests 1503-1515. Cuba localities. CA: Baraguá Collections. New York 36 FAMILY BELOHINIDAE This family is based on a single species in the Trox Fabricius 1775: 31 genus Belohinus from Madagascar. T. insularis Chevrolat 1864c: 416; Vaurie 1955: 41 Distribution. Cuba. Endemic 37. FAMILY OCHODAEIDAE Cuba localities. CA: Baraguá. Ci: Soledad The sand-loving scarab beetles Collections. Gundlach no. 133. New York This family occurs in North, Central and South America but in the West Indies it is known only in REFERENCES Cuba. Adults to lights and larvae live in soil. Scholtz, C.H. 1982. Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Republic South SUBFAMILY OCHODAEINAE Africa Dept. Agric. Fisheries Ent. Mem. 54: 27 pp. Ochodaeus Serville 1825: 360 Vaurie, P. 1955. A revision of the genus Trox in O. grandiceps Fairmaire 1897: 241 North America (Coleoptera, Scarabaeidae). Bull. Distribution. Cuba, “West Indies” Amer. Mus. Nat. Hist. 106: 1-89. Cuba localities. Ci: Soledad 84 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Howden Cuba localities. Widespread Collections. Gatineau. Gundlach no. 872 38. FAMILY HYBOSORIDAE C. turquinensis (Zayas) 1988: 28 () The scavenger scarab or hybosorid beetles Distribution. Cuba. Endemic. Possibly a syn- The family is widely distributed in the world onym of C. chalceus tropics. Cuba localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) Apalonychus Westwood 1846: 168 A. waterhousei Westwood 1846: 169; Chevrolat REFERENCES 1864c: 415 (Hapalonychus); Gundlach 1891: Howden, H.F. and Gill, B.D. 2000. Tribes of New 136 World Ceratocanthinae, with keys to genera Distribution. Cuba, Hispaniola and descriptions of new species (Coleoptera: Cuba localities. M: Cárdenas Scarabaeidae). Sociobiology 35: 281-329. Collections. Gundlach no. 137 40. FAMILY Hybosorus MacLeay 1819: 120 This family contains 8 genera, three of which H. illigeri Reiche 1853: 88; Chalumeau 1981: 173; occur in Peru and Chile. It is not in the West Indies. Ocampo 2002: 318 Distribution. Cuba (introduced, oldest record 41. FAMILY SCARABAEIDAE from 1954), Jamaica, USA (TX-KY-FL); native The scarab beetles to ; at lights Reviewed by Bruce Gill, May, 1998 Cuba localities. SC: Matias; Uvira; Playa This is a large family with diverse habits. They Ingles; Playa Larga feed on roots, leaves, rotted wood, litter, dung, Collections. fruits, etc. Some may be pests of forestry or agri- culture. Chalumeau (1981) reports on the collec- REFERENCES tions from the Cuban-Roumanian cave biology ex- Ocampo, F.C. 2002. Hybosorids of the United peditions. States and expanding distribution of the intro- duced species Hybosorus illigeri (Coleoptera: SUBFAMILY APHODIINAE Scarabaeoidea: Hybosoridae). Ann. Ent. Soc. TRIBE Amer. 95: 316-322. Aphodius Illiger 1798: 115. Keys in Chapin 1940, 39. FAMILY CERATOCANTHIDAE and Woodruff 1973 The pill scarabs or ceratocanthid beetles A. cuniculus Chervolat 1864c: 411: Gundlach 1891: These can be found under bark and by beating 148; Chapin 1940: 7 dead branches. Some live with termites and can be Distribution. Cuba, Jamaica, Hispaniola, taken using broken termite nests as baits. Howden Puerto Rico, Lesser Antilles, México, Guate- and Gill (2000) present keys to New World genera. mala, USA (FL-TX) Cuba localities. M: Cienaga de Zapata Ceratocanthus White 1842: 84 Collections. Gundlach no. 1880. Washington C. chalceus (Germar)1843: 114 (Sphaeromorphus); A. lividus Olivier 1789: 86; Jacquelin du Val 1856: Gundlach 1891: 133 (Cloeotus) 116; Chevrolat 1864c: 412; Gundlach 1891: 149; Distribution. Cuba. Endemic Chapin 1940: 6; Chalumeau 1981: 174 Cuba localities. Widespread Distribution. Cuba (introduced), Jamaica, Collections. Gundlach no. 178 Hispaniola, Puerto Rico, Lesser Antilles, wide- C. gundlachi Harold 1874: 34 (Cloeotus) spread USA, México to Argentina; native to the Distribution. Cuba. Endemic Old World Cuba localities. Cuba localities. Widespread Collections. Collections. Gundlach no. 196. Washington C. semistriatus (Germar) 1843: 120 A. quadridentatus Harold 1861: 107; Chevrolat (Sphaeromorphus); Jacquelin du Val 1856: 120; 1864c: 412; Gundlach 1891: 149; Chapin 1940: Chevrolat 1864c: 417; Gundlach 1891: 134 4 (Cloeotus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. A Checklist of the Beetles of Cuba 85

Collections. Cuba localities. Ci: Soledad (t.l.). Ha: Ha- A. n. sp. Zayas 1988: 28 vana Distribution. Cuba. Endemic Collections. Brussels. Paris (type) Cuba localities. A. haroldi Steinheil 1872: 556; Chapin 1940: 15 Collections. Zayas Distribution. Cuba, Hispaniola, Puerto Rico, México, Argentina TRIBE Cuba localities. Collections. Cambridge? Ataenius Harold 1867: 100. Keys in Cartwright A. havanensis Balthasar 1938: 56; Chapin 1940: (1973), Chapin (1940) and Woodruff (1973) 41 A. bicolour Petrovitz 21963: 642; Stebnicka 2002: Distribution. Cuba, Barbados, St. Kitts, St. 268 Croix, USA (FL (Lower Keys only)) Distribution. Cuba, Jamaica Cuba localities. Ha: Havana (t.l.) Cuba localities. Ha: Havana. PR: Joabel Collections. Prague (type?) Rubio; Viñales. SC: Boniato A. imbricatus Melsheimer 1844: 136; Chapin 1940: Collections.Berlin 21; Chalumeau 1981: 175 A. brevinotus Chapin 1940: 39 Distribution. Cuba, Hispaniola, Bahamas, Distribution. Cuba. Endemic St. Croix, widespread USA, México to Argen- Cuba localities. CA: Baraguá (t.l.) tina Collections. Cambridge (type) Cuba localities. CA: Baraguá. Ci: Cayamas. A. corrosus Chapin 1940: 19 IJ: Represa La Fé Distribution. Cuba. Endemic Collections. Cambridge. Washington Cuba localities. SC: Aguadores (t.l.) A. jardinensis Stebnicka 2002: 278 Collections. Cambridge (type) Distribution. Cuba. Endemic A. cribrithorax Bates 1887: 95; Chapin 1940: 28 Cuba localities. SC: Santiago, Jardin Distribution. Cuba, Jamaica, St. Thomas, Botanico (t.l.) México to Panamá Collections. Gatineau (type) Cuba localities. A. liogaster Bates 1887: 94; Hinton 1937: 193l; Collections. Cambridge? Chapin 1940: 29 A. darlingtoni Hinton 1937: 179 Distribution. Cuba, Jamaica, Guadeloupe, Distribution. Cuba, Jamaica, Hispaniola, México to Panamá, Ecuador (Galapagos Islands) Puerto Rico, Lesser Antilles Cuba localities. Ha: Havana Cuba localities. Collections. Cambridge Collections. Cambridge? A. miami Cartwright 1934: 200; Chapin 1940: 22 A. edwardsi Chapin 1940: 26; Chalumeau 1981: Distribution. Cuba, Puerto Rico, Barbados, 174 se USA Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Puerto Rico, Lesser Antilles Collections. Cuba localities. PR: Quemado de Pineda A. picinus Harold 1867: 281; Woodruff 1973: 124; Collections. Chalumeau 1981: 175 A. gracilis Melsheimer 1844: 137; Jacquelin du Val Distribution. Cuba, Jamaica, Hispaniola, 1856: 119 (); Chevrolat 1864c: 414; Puerto Rico, Lesser Antilles, Venezuela, se USA Gundlach 1891: 152; Chapin 1940: 25; Cuba localities. IJ: Base Mella Chalumeau 1981: 174 Collections. Distribution. Cuba, Jamaica, Hispaniola, A. rhyticephalus (Chevrolat) 1864c: 413 (Auperia); Puerto Rico, Lesser Antilles, USA, Colombia Gundlach 1891: 150; Chalumeau 1981: 175 to Argentina A. strigicauda Bates 1887: 96; Chapin 1940: 31 Cuba localities. Widespread Distribution. Cuba, Jamaica, Hispaniola, Collections. Gatineau. Cambridge? Gundlach Puerto Rico, Lesser Antilles, México, Central no. 84 and South America, USA (FL-SC-TX) A. gruneri Chalumeau 1979: 229, 1981: 174; Cuba localities. IJ: Matias. SC: 6 km NE Stebnicka 2002: 274 Siboney Distribution. Cuba. Endemic Collections. Gatineau. Cambridge 86 Arthropods of Florida and Neighboring Land Areas, Vol. 18

A. scutellaris Harold 1867: 82; Chalumeau 1981: Pseudataenius Brown 1927: 288 175 P. walterhorni (Balthasar) 1938: 55 (Ataenius); Distribution. Cuba, Antilles, Venezuela, Cen- Woodruff 1973: 133; Chapin 1940: 41; tral America Cartwright 1974: 12; Chalumeau 1981: 175 Cuba localities. IJ: Matias Distribution. Cuba. Endemic (USA: FL?, Collections. believed introduced to Key West) A. stercorator Fabricius 1775: 20; Jacquelin du Val Cuba localities. Ci: Soledad. Ha: Havana 1856: 118; Chevrolat 1864c: 413; misidentified (t.l.). IJ: Base Mella. PR: Quemado de Pineda; by Horn 1887: 83?; Gundlach 1891: 150 El Embarcadero (Auperia) Collections. Washington Distribution. Cuba (introduced ), Puerto Rico, South America, USA; native to the Old World TRIBE Cuba localities. Widespread Collections. Gundlach no. 193 Odontopsammodius Gordon and Pittino 1992: A. sulcatulus (Chevrolat) 1864c: 413 (Auperia); 265 Gundlach 1891: 151; Chapin 1940: 37 O. decuiella (Chalumeau) 1979: 229, 1981: 176 Distribution. Cuba. Endemic (Psammodius) Gordon and Pittino 1992: 266 Cuba localities. CA: Baraguá. Ci: Cayamas. Distribution. Cuba. Endemic Ha: Santiago de las Vegas. PR: Bahia Honda. Cuba localities. Gu: Rio Brazo Seco (t.l.) SC: Santiago, Roig Botanical Garden Collections. Paris (type, supposedly de- Collections. Gatineau. Gundlach no. 1831. stroyed) Washington A. terminalis (Chevrolat) 1864c: 414 (Auperia); Platytomus Mulsant 1842: 310 Gundlach 1891: 151; Chapin 1940: 38; P. parvulus (Chevrolat) 1864c: 415 (Psammodius); Chalumeau 1981: 175; Stebnicka 2002: 267 Gundlach 1891: 152; Chapin 1940: 8 Distribution. Cuba, Jamaica (Pleurophorus); Gordon and Pittino 1992: 268 Cuba localities. Widespread Distribution. Cuba, Jamaica, Hispaniola to Collections.Berlin. Cambridge. Gundlach no. Grenada 195. Ottawa Cuba localities. M: Cienaga de Zapata A. walterhorni Balthasar 1938: 55; Stebnicka 2002: Collections. Gundlach no. 946. Washington 265 Distribution. Cuba, USA (PA-FL-LA) TRIBE RHYPARINI Cuba localities. Ci:Cayamas; Cienfuegos; Rio Caburni. Ha: Havana. SC. Ca: Baragua Rhyparus Westwood 1845: 93 Collections. Eberswalde (type). Washington R. zayasi Cartwright and Woodruff 1969: 7 Distribution. Cuba, Jamaica Euparixia Brown 1927: 288 Cuba localities. SC: Turquino, Cardero; at E. bruneri Chapin 1940: 40; Woodruff and light Cartwright 1967: 12 Collections. Gatineau. Gainesville. Havana. Distribution. Cuba. Endemic Ottawa. Washington Cuba localities. Ca: Camagüey (t.l.). CA: Baraguá. Ha: Havana; Santiago de las Vegas. SUBFAMILY SCARABAEINAE Probably in nests of Atta insularis leafcutting TRIBE CANTHONINI ants (bibijaguas) Collections. Washington (type) Canthochilum Chapin 1934: 99 C. anacaona Zayas and Matthews 1966: 8; Myrmecaphodius Martinez 1952: 85 Matthews 1966: 71; Matthews 1969: 121 M. excavaticollis (Blanchard) 1846: 184; Chalumeau Distribution. Cuba. EndemicInterior high- 1981: 174 land forests of eastern Cuba Distribution. Cuba, se USA to Argentina; Cuba localities. .Gu: Mt. N of Imias; Yunque lives in nests of Solenopsis fire ants de Baracoa (t.l.). Ho: Las Mulas; Sierra Cristal Cuba localities. Ho: Arroyo Guayabo. SC: Collections. Cambridge. Zayas (type). Wash- Matias ington Collections. A Checklist of the Beetles of Cuba 87

C. baracutey Zayas and Matthews 1966: 11; Cuba localities. SC: Loma del Gato, 1100 m Matthews 1966: 69 (t.l.); Gran Piedra (elfin forest, 1000m). SC: Distribution. Cuba. EndemicInterior high- Pico Turquino land forests of eastern Cuba Collections. Gatineau. Cambridge. Zayas Cuba localities. SC: Florida Blanca, Songo (type) (t.l.) Collections. Zayas (type) Canthonella Chapin 1930: 2 C. cemi Zayas and Matthews 1966: 7; Matthews C. pygmaea (Harold) 1869: 96; Chapin 1930: 2; 1966: 69 Zayas and Matthews 1966: 16; Matthews 1966: Distribution. Cuba . EndemicInterior high- 77; 1969: 123 land forests of eastern Cuba Distribution. Cuba. EndemicLowland and Cuba localities. . Gu: Resucia. SC: Florida coastal scrub forest, eastern and western Cuba Blanca, Songo; Palma Mocha, Sierra Maestra (not Puerto Rico or Hispaniola) (t.l.); Santa Maria de Loreto Cuba localities. Ha: Cueva del Indio. Ho: Las Collections. Ramsden. Washington. Zayas Mulas; Sierra Cristal. PR: Monte El Pinar (type) Collections. Paris (type). Washington. Zayas C. guayaca Zayas and Matthews 1966: 8; Matthews 1966: 70 TRIBE ONITICELLINI Distribution. Cuba. EndemicInterior high- land forests of eastern Cuba Oniticellus Serville 1825: 356 Cuba localities. SC: Pico Cuba, 1810 m (t.l.) O. cubiensis Laporte 1840: 92; Jacquelin du Val Collections. Washington. Zayas (type) 1856: 116; Chevrolat 1864c: 410; Gundlach C. gundlachi (Harold) 1968: 80; Chapin 1934: 101; 1891: 147; Matthews 1966: 27; 1969: 114 Zayas and Matthews 1966: 5; Matthews 1966: Distribution. Cuba, Jamaica, Bahamas (not 71 Florida); pastures and roadsides (Matthews Distribution. Cuba. EndemicLowland and 1969) coastal scrub forest, western Cuba Cuba localities. Widespread Cuba localities. M: Dolina del Compamento. Collections. Gundlach no. 324. London. Paris PR: Guanabacabibes. Ca: Playa El Ingles, Santa (type not found). Pittsburgh. Washington. Lucia Zayas Collections. Paris (type). Zayas C. histeroides (Harold) 1868: 80; Chapin 1934: 101; TRIBE ONTHOPHAGINI Zayas and Matthews 1966: 4; Matthews 1966: 70; Chalumeau 1981: 174 Onthophagus Latreille 1802: 141 Distribution. Cuba. EndemicLowland coastal O. gazella Fabricius 1787: 377. The brown dung scrub and forest, north central coast and west- beetle ern Cuba Distribution. Cuba (introduced); widespread Cuba localities. M: Dolina del Campamento. in New World, across USA and México; native Ha: Rio Santa Ana. PR: Guanahacabibes. to Old World (Hoebeke and Beucke, 1997, Ent. Collections. Paris (type). Zayas News 108: 345-362) C. pijirigua Zayas and Matthews 1966: 13; Cuba localities. SC: Santiago, Roig Botani- Matthews 1966: 71 cal Garden (first Cuba record) Distribution. Cuba. EndemicLowland and Collections. Gatineau. Gainesville. Havana coastal scrub forest, north central coast and O. marginatus Laporte 1840: 87; Jacquelin du Val western Cuba 1856: 115; Chevrolat 1864c: 410; Gundlach Cuba localities. PR: El Pinar, Los Palacios 1891: 148; Matthews 1966: 9; Chalumeau 1981: (t.l.). Ha: Rio Santa Ana. PR: Guanahacabibes. 174 Ca: Playa El Ingles; M: Pen. Zapata Distribution. Cuba. Endemic Collections. Zayas (type). Washington Cuba localities. Widespread; in pastures and C. tureya Zayas and Matthews 1966: 12; Matthews roadsides 1966: 69; 1969: 121 Collections. Gatineau. Gundlach no. 270. Distribution. Cuba. EndemicInterior high- London. Paris? (type not found by Chalumeau). land forests of eastern Cuba Washington. Zayas 88 Arthropods of Florida and Neighboring Land Areas, Vol. 18

SUBFAMILY MELOLONTHINAE A. subsericans (Jacquelin du Val) 1856: 132; Chevrolat 1865: 27; Gundlach 1891: 141 TRIBE MELOLONTHINI Distribution. Cuba. Endemic; introduced to Panamá?; intercepted in USA (Miami), a pest Phyllophaga Harris 1826: 6; subgenus of sugarcane Cnemarachis Saylor 1942: 159 (=Ancylonycha Cuba localities. CA: Baraguá. Ci: Cayamas.; Blanchard 1845). Constancia. Ha: Alquizar; Hoyo Colorado; Santiago de las Vegas. IJ: Punta Pedernales; This genus probably has the greatest impact of Santa Fe; Nueva Gerona any beetle on Cuban agriculture. The adults feed Collections. Gundlach no. 1059 (A. on the leaves of many plants, and the larvae feed in bifoveolata); no. 1118 (A. subsericans). Halle? the soil on plant roots (Fernandez Garcia 1998). (type?). Washington Chapin (1932) gives a key to species known at that P. (C.) angusta (Blanchard) 1850: 138 time (see Garcia Vidal, 1975), and Muslera Ramos (Ancylonycha); Chevrolat 1865: 25; Gundlach and Fernandez Garcia (1998) provide a summary 1891: 139; Chapin 1932: 205 of the Cuban fauna; see also Woodruff and Beck Distribution. Cuba. Endemic (1989) on Florida species. Cuba localities. Collections. Paris (type?) P. (C.) acuñai Chapin 1937: 3 P. (C.) baracoana Chapin 1932: 188 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: near Pico Turquino (t.l.) Cuba localities. Gu: Baracoa (t.l.) Collections. Washington (type). Havana Collections. Washington (type) P. (C.) aeneotincta Chapin 1932: 201 P. (C.) baraguensis Chapin 1932: 185 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. IJ: Santa Fe (t.l.) Cuba localities. CA: Baraguá (t.l.) Collections. Washington (type) Collections. Washington (type) Notes: Possibly the species upon which Chevrolat P. (C.) bicavifrons Chapin 1932: 187 (1865: 28) based his Isle of Pines record for P. Distribution. Cuba. Endemic suturalis (Chapin 1932: 201) Cuba localities. SC: Sierra Maestra, 1070- P. (C.) aeruginosa (Burmeister) 1855: 338 1350 m (t.l.) (Ancylonycha); Chevrolat 1865: 26; Gundlach Collections. Washington (type) 1891: 140 (acruginosa): Chapin 1932: 204 P. (C.) bihammifrons Saylor 1940: 305; Chalumeau Distribution. Cuba. Endemic 1989: 339 Cuba localities. Ci: Cayamas. Ha: Santiago Distribution. Cuba. Endemic de las Vegas. M: Cárdenas Cuba localities. IJ. Nueva Gerona Collections. Gundlach no. 1807. Halle (type?). Collections. Prague Washington P. (C.) bilyi Chalumeau 1989: 338 P. (C.) ahlbrandti Garcia Vidal 1988: 104 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR, Vinaloa (t.l.); Sierra Cuba localities. Gr: Rio Yara (t.l.) Rosario; Taco Taco) Collections. San Francisco (type) Collections. Prague (type) P. (C.) alayoi Garcia 1978: 3 P. (C.) bimaculata Garcia 1984: 313 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gr: Cabo Cruz (t.l.) Cuba localities. SC: Puerto Boniato (t.l.) Collections. Havana. Zayas (type) Collections. San Francisco (type) P. (C.) alquizara Chapin 1932: 204 P. (C.) brevicornis Garcia Vidal 1988: 101 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ha: Alquizar (t.l.) Cuba localities. PR: Viñales (t.l.) Collections. Washington (type) Collections. Zayas (type) P. (C.) analis (Burmeister) 1855: 340 (Ancylonycha); P. (C.) bruneri Chapin 1932: 203 Chevrolat 1865: 28; Gundlach 1891: 143; Distribution. Cuba Endemic; (USA, intro- Chapin 1932: 197; Chalumeau 1981: 177 duced into .s FL and established) A. bifoveolata (Jacquelin du Val) 1856: 133; Cuba localities. Ha: Santiago de las Vegas Chevrolat 1865: 27; Gundlach 1891: 141 (t.l.) A Checklist of the Beetles of Cuba 89

Collections. Washington (type) Cuba localities. Ci: Soledad. PR: Loma de P. (C.) caneyensis Garcia 1984: 315 Soroa; Rancho Mundito, Sierra Organos (t.l.); Distribution. Cuba. Endemic Viñales Cuba localities. SC: Gran Piedra (t.l.) Collections. Zayas (type); Havana Collections. San Francisco (type). Washing- P. (C.) expansa Chapin 1932: 182 ton Distribution. Cuba. Endemic P. (C.) cardini Chapin 1932: 197; Garcia Vidal 1988: Cuba localities. SC: Constancia (t.l.) 101 Collections. Washington (type) Distribution. Cuba. Endemic P. (C.) explanicollis Chapin 1935: 69 Cuba localities. Gu: Baracoa (t.l.). SC: Pico Distribution. Cuba. Endemic Turquino; Loma de Cala to Palma Mocha, Si- Cuba localities. PR: Santo Tomas (t.l.). A erra Maestra; Loma del Gato. (Not Ca: Jaronú) former pest of young pine trees (Vàzquez et. al. Collections. Washington (type). Zayas 1992: 44) P. (C.) castineirasi Garcia Vidal 1978: 5 Collections. Havana? Washington (type) Distribution. Cuba. Endemic P. (C.) fimbriata Chapin 1932: 182 Cuba localities. IJ: Nuevo Gerona (t.l.); Distribution. Cuba. Endemic Reparto Nazareno Cuba localities. Ca: Jaronú (t.l.) Collections. Zayas (type); Havana Collections. Washington (type) P. (C.) chalumeaui Garcia Vidal 1984: 317 P. (C.) franciscana Garcia Vidal 1984: 319 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Sierra Maestra, Loma Cuba localities. Ha: Sierra del Grillo (t.l.) de Cala to P. Mocha (t.l.) Collections. San Francisco (type) Collections. San Francisco (type) P. (C.) hardyi Garcia Vidal 1984: 319 P. (C.) complexipennis Garcia Vidal 1984: 317 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Tortuguilla ( t.l.) Cuba localities. PR: Viñales (t.l.) Collections. San Francisco (type). Washing- Collections. San Francisco (type) ton P. (C.) crenaticollis (Blanchard) 1850: 137 P. (C.) hernandezi Garcia Vidal 1984: 321 (Ancylonycha); Chevrolat 1865: 23; Gundlach Distribution. Cuba. Endemic 1891: 136; Chapin 1932: 183 Cuba localities. SC: Sierra Maestra (t.l.) Distribution. Cuba. Endemic Collections. San Francisco (type). Washing- Cuba localities. Ha: Rio Almendares; Ha- ton vana; Vedado. M: Bolondron P. (C.) imitatrix Chapin 1932: 199; Chalumeau Collections. Gundlach no. 443. Paris (type). 1981: 177 Washington Distribution. Cuba. Endemic P. (C.) cubana Chapin 1932: 194 Cuba localities. IJ: Santa Barbara (t.l.); Distribution. Cuba. Endemic Represa de La Fé Cuba localities. CA: Baraguá (t.l.) Collections. Washington (type) Collections. Washington (type) P. (C.) independentista Garcia Vidal 1984: 322 P. (C.) dissimilis (Chevrolat) 1865: 26 Distribution. Cuba. Endemic (Ancylonycha); Gundlach 1891: 140; Chapin Cuba localities. Ha; Santiago de las Vegas 1932: 201; Chalumeau 1981: 177 (t.l.). Los Jazmines Distribution. Cuba. EndemicIntercepted in Collections. San Francisco (type). Washing- USA (Miami); a pest of sugarcane ton Cuba localities. Ca: Jaronú. CA: Baraguá. P. (C.) insulaepinorum Garcia Vidal 1978: 6 Ci: Cayamas; Constancia; Santa Clara; Distribution. Cuba. Endemic Soledad. IJ: Matias. M: Cárdenas; Perico. LT: Cuba localities. IJ: El Abra (t.l.) Jobabo Collections. Havana (type?) Collections. Cotypes in Berlin and Philadel- P. (C.) jaronua Chapin 1932: 194 phia (Poey?). Gundlach no. 1115. Washington Distribution. Cuba. Endemic P. (C.) dubitatus Garcia Vidal 1978: 8 Cuba localities. Ca: Jaronú (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) P. (C.) laboriosa Garcia Vidal 1988: 102 Distribution. Cuba. Endemic 90 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Ca: Jaronú (t.l.) Distribution. Cuba, Hispaniola, St. Kitts, Collections. San Francisco (type, female). Guadeloupe Washington Cuba localities. Ca: Galbis. Ci: Cayamas; P. (C.) labrata Chapin 1932: 199 Constancia. SS: Trinidad. Distribution. Cuba. Endemic Collections. Gundlach no. 28. J. Moser, Ber- Cuba localities. PR: Herradura (t.l.) lin (type?). Washington Collections. Washington (type) P. (C.) persimilis Chapin 1935: 70 P. (C.) maestrensis Garcia Vidal 1984: 323 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Baracoa (t.l.) Cuba localities. Gu: “Capitolio,” Rio Yara Collections. Washington (type) (t.l.). SC: Loma de Cala, Sierra Maestra P. (C.) pilotoensis Garcia Vidal 1987: 304 Collections. San Francisco (type). Washing- Distribution. Cuba. Endemic ton Cuba localities. Ho: Batey de Moa (t.l.); P. (C.) marinus Garcia Vidal 1987: 303 Piloto, Moa Distribution. Cuba. Endemic Collections. Urbana (type). San Francisco Cuba localities. SC: Cuidamar (t.l.) P. (C.) potrerillo Garcia Vidal 1987: 304 Collections. Urbana (type) Distribution. Cuba. Endemic P. (C.) microsoma Chapin 1932: 202 Cuba localities. LV: Pico Potrerillo (t.l.) Distribution. Cuba. Endemic Collections. Urbana (type) Cuba localities. Ca: Camagüey (t.l.) P. (C.) pseudomicans Chapin 1935: 73 Collections. Washington (type) Distribution. Cuba. Endemic P. (C.) migratoria Garcia Vidal 1984: 324 Cuba localities. PR: Sierra Rangel (t.l.) Distribution. Cuba. Endemic Collections. Havana ?. Washington (type) Cuba localities. PR: San Vincente (t.l.).; P. (C.) puberula (Jacquelin du Val) 1856: 130 Viñales (Ancylonycha); Chevrolat 1865: 24; Gundlach Collections. San Francisco (type) 1891: 137; Chapin 1932: 189 P. (C.) nasalis Chapin 1935: 72 Distribution. Cuba. Endemic. Introduced Distribution. Cuba. Endemic into USA (collected once at Miami, FL) Cuba localities. PR: Sierra Rangel, Las Cuba localities. CA: Baraguá; Jaronú. Ci: Animas (t.l.) Cayamas. Ha: Alquizar; Santiago de las Ve- Collections. Washington (type) gas. M: Hato Nuevo. PR: Cabanas. SC: P. (C.) onchophora Chapin 1932: 184; Chalumeau Constancia. SS: Trinidad. The main pest of 1981: 177 pineapple crops (Vàzquez et al. 1992: 44) Distribution. Cuba. EndemicIntroduced to Collections. Berlin? (type?). Gundlach no. 20. USA (GA) Washington Cuba localities. Gr: Paso Estancia (t.l.). Ha: P. (C.) rangelana Chapin 1935: 71 Santiago de las Vegas. SC: Trinidad. M:Perico Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. PR: Sierra Rangel, 500 m P. (C.) parallela (Blanchard) 1850: 137 (t.l.) (Ancylonycha); Chevrolat 1865: 24; Gundlach Collections. Washington (type) 1891: 138; Chapin 1932: 205 P. (C.) sandersoni Garcia Vidal 1988: 104 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Loma del Gato.(t.l.) Collections. Gundlach no. 1858 Collections. Champaign. San Francisco P. (C.) parcesetifera Chapin 1935: 71 (type). Washington Distribution. Cuba. Endemic P. (C.) santaclarae Chapin 1932: 189 Cuba localities. PR: Sierra Rangel (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. SC: Manacas (t.l.) P. (C.) patruelis (Chevrolat) 1865: 25 Collections. Washington (type) (Ancylonycha); Gundlach 1891: 139; Chapin P. (C.) santiaguensis Garcia Vidal 1987: 303 1932: 191 Distribution. Cuba. Endemic P. tenuicalcaris Chapin 1932: 192 (Ca: Galbis, Cuba localities. SC: Cuidamar (t.l.) t.l.; type in Washington) Collections. Champaign. San Francisco (type) P. (C.) scaramuzzai Garcia Vidal 1988: 106 A Checklist of the Beetles of Cuba 91

Distribution. Cuba. Endemic P. (C.) tuberculifrons (Chevrolat) 1865: 23 Cuba localities. PR: Guanahacabibes (t.l.) (Ancylonycha); Gundlach 1891: 137; Chapin Collections. San Francisco (type) 1932: 186; Chalumeau 1989: 339 P. (C.) schwarzi Chapin 1932: 184 Distribution. Cuba. Endemic Distribution. Cuba Endemic Cuba localities. M: Santo Tomas, Zapata Cuba localities. Ci: Cayamas (t.l.) Collections. Gundlach no. 1120. Paris. Phila- Collections. Washington (type) delphia (Poey, cotypes?) P. (C.) siboneyensis Garcia Vidal 1988: 103 P. (C.) vermiculata Chapin 1932: 195 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Siboney (t.l.) Cuba localities. SC: Palma Mocha (t.l.) Collections. Washington (type) Collections. Washington (type) P. (C.) signaticollis (Burmeister) 1855: 337 P. (C.) villaclarensis Garcia Vidal 1987: 302 (Ancylonycha); Jacquelin du Val 1856: 56; Distribution. Cuba. Endemic Chevrolat 1865: 25; Chapin 1932: 190; Cuba localities. M: Cienega de Zapata (t.l.) Chalumeau 1981: 177 Collections. Urbana (type) Ancylonycha confusa Jacquelin du Val 1856: 131 P. (C.) zayasi Garcia Vidal 1978: 10 (type in Berlin?); Chevrolat 1865: 24; Gundlach Distribution. Cuba. Endemic 1891: 138 Cuba localities. IJ: Represa La Demajagua Distribution. Cuba. Endemic (t.l.). Attacking citrus and casuarina trees Cuba localities. Ci: Cayamas.Ha: Alquizar; Collections. Havana. Zayas (type) Havana; Hoyo Colorado; Santiago de las Vegas Collections. Gundlach no. 261. Halle? (type?); TRIBE MACRODACTYLINI Washington P. (C.) speculifera (Chevrolat) 1865: 27; Gundlach Anoplosiagum Blanchard 1850: 119 1891: 142; Chapin 1935: 69 A. cubensis Garcia Vidal 1982: 330 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Sierra Rangel; Las Cuba localities. M: Central Cuba (t.l.) Animas, (500 m) Collections. Zayas (type) Collections. Gundlach no. 1138. Moser, Ber- A. flavicollis Chapin 1935: 68 lin (cotype?). Philadelphia, (Poey cotype?). Distribution. Cuba. Endemic Washington Cuba localities. Ci: Pico Potrerillos, Trinidad P. (C.) spinicola Garcia Vidal 1984: 324 Mts. (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. SC: Gran Piedra (t.l.) A. oteroi Garcia Vidal 1982: 332 Collections. Champaign. San Francisco Distribution. Cuba. Endemic (type). Washington Cuba localities. Ha: Estacion Experimental P. (C.) suturalis (Chevrolat) 1865: 29 Agronomica Finca Coca (t.l.); Santiago de las (Ancylonycha); Gundlach 1891: 142; Chapin Vegas 1932: 200 Collections. Zayas (type) Distribution. Cuba. Endemic A. pallidulum Blanchard 1850: 120; Chevrolat 1865: Cuba localities. Ci: Cayamas. SS: Jatibonico. 21; Gundlach 1891; 134; Chapin 1932: 177 IJ: (?) Distribution. Cuba. Endemic Collections. Gundlach no. 1140. J. Moser, Cuba localities. Ci: Cayamas; Soledad. M: Berlin (type?). Washington Cárdenas P. (C.) tetracera Chapin 1932: 193 Collections. Gundlach no. 1139. Paris (type); Distribution. Cuba. Endemic Washington Cuba localities. SC: Seboruco (t.l.) A. pauliani Chalumeau 1979: 229; 1981: 176 Collections. Washington (type) Distribution. Cuba. Endemic P. (C.) trinitariensis Garcia Vidal 1987: 306 Cuba localities. IJ: Represa de La Fé (t.l.) Distribution. Cuba. Endemic Collections. Paris (type). Washington Cuba localities. SS: Lomas de Trinidad A. peltatum Chapin 1932: 176 ,Topes, (t.l.) Distribution. Cuba. Endemic Collections. Urbana (type) Cuba localities. Ha: San Antonio de las Baños (t.l.); Santiago de las Vegas 92 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Washington (type) TRIBE RUTELINI A. rufum Chapin 1932: 177 Distribution. Cuba. Endemic Rutela Latreille 1802: 151 Cuba localities. SS: Central Jatibonico (t.l.) R. formosa Burmeister 1844: 383; Jacquelin du Val Collections. Washington (type) 1856: 130; Chevrolat 1865: 29; Gundlach 1891: A. rutilum (Chevrolat) 1865: 22 (Clavipalpus); 144; Chapin 1932: 283; Chalumeau 1981: 177 Gundlach 1891: 135; Chapin 1932: 178 Distribution. Cuba, Bahamas, USA (FL) Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. SC: Pico Turquino, 4-5000' Collections. Halle (type?). Pittsburgh. Wash- Collections. Gundlach no. 553 (type?). Wash- ington ington A. scabrosum Chapin 1932: 175 SUBFAMILY DYNASTINAE Distribution. Cuba. Endemic TRIBE CYCLOCEPHALINI Cuba localities. PR: Cabanas (t.l.) Collections. Washington (type) Cyclocephala Latreille 1829: 552; Key in Chapin A. scaramuzzai Chapin 1935: 67 1932: 287 Distribution. Cuba. Endemic C. atricolor Chapin 1932: 289 Cuba localities. Ha: Santiago de la Vegas, Distribution. Cuba. Endemic (t.l.) Cuba localities. CA: Baraguá (t.l.). Gu: Collections. Havana? Washington (type) Baracoa A. simplicipes Chapin 1932: 175 Collections. Washington (type) Distribution. Cuba. Endemic C. cardini Chapin 1935: 74 (replacement name) Cuba localities. CA: Baraguá (t.l.) C. signatoides Chapin 1932: 287 (not Höhne 1923) Collections. Washington (type) Distribution. Cuba. Endemic A. swezeyi Garcia Vidal 1982: 333 Cuba localities. PR: San Juan y Martinez Distribution. Cuba. Endemic (t.l.). Gu: Baracoa Cuba localities. PR: Las Martinas (t.l.); San Collections. Washington (type) Julian C. cubana Chapin 1932: 291; Chalumeau 1981: 178 Collections. San Francisco. Zayas (type) C. signata of authors (in part) A. turquinensis Garcia Vidal 1982: 334 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. SC: Pico Turquino (t.l.); Collections. Washington (type) Palma Mocha to Pico Joaquin C. frontalis Chevrolat 1844: 90; Jacquelin du Val Collections. San Francisco. Zayas (type) 1856: 129; Chevrolat 1865: 30. Gundlach 1891: A. variabile Chevrolat 1865: 22; Gundlach 1891: 145 135 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. IJ: Santa Barbara Cuba localities. Widespread Collections. Collections. Gundlach no. 1137 C. fusiformis Chapin 1932: 287 A. zayasi Garcia Vidal 1982: 335 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. IJ: Santa Barbara (t.l.) Cuba localities. Gu: La Glorieta (t.l.) Collections. Washington (type) Collections. San Francisco. Zayas (type) C. notata (Illiger) 1806: 235 (Melolontha); Chapin 1932: 288 SUBFAMILY Distribution. Cuba (introduced), Jamaica, TRIBE ANOMALINI Hispaniola. Note: Probably introduced to Cuba from Hispaniola (Chapin 1932: 288) Samouelle 1819: 191 Cuba localities. Gu: Guantánamo A. calceata Chevrolat 1865: 28; Gundlach 1891: 143; Collections. Havana. Washington Chapin 1932: 284 C. rangelana Chapin 1935: 75; Joly 1995: 58 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. PR: Las Animas, Sierra Collections. Washington Rangel (t.l.) Collections. Washington (type). Havana? A Checklist of the Beetles of Cuba 93

C. signata Fabricius 1781: 39; Jacquelin du Val TRIBE PENTODONTINI 1856: 128; Chevrolat 1865: 30; Gundlach 189l: 144 Ligyrus Burmeister 1847: 542; key in Chapin Distribution. Cuba, Jamaica, Grenada, 1932: 297. The genus Tomarus Erichson 1847 Belize to Trinidad seems to have priority over Ligyrus but has Cuba localities. Widespread not been used in a Cuban context. Collections. Gundlach no. 262 L. cuniculus Fabricius 1801: 20 C. verticalis Burmeister 1847: 51; Jacquelin du Val Distribution. Cuba, Jamaica to Puerto Rico, 1856: 129; Chevrolat 1865: 31; Gundlach 189l: widespread Lesser Antilles, México to Trinidad, 145; Chapin 1932: 290 USA Distribution. Cuba. Endemic Cuba localities. Cuba localities. Widespread Collections. Collections. Gundlach no. 36. Halle (type?). L. fossor (Latreille) 1833: 11; Chapin 1932: 297 Havana. Washington L. juvencus of authors Distribution. Cuba, Puerto Rico, Panamá to Dyscinetus Harold 1869: 123. Key in Chapin 1932: Argentina and Venezuela 293 Cuba localities. Ci: Cayamas. Ha: Santiago. D. frater Bates 1888: 312; Chapin 1932: 295 M: Colón. PR: San Blas Martinéz; Pinar del D. picipes Holland 1917: 340 (misidentification) Rio; San Diego Distribution. Cuba, México to Panamá Collections. Havana. Washington Cuba localities. IJ: Neuva Jerona. Ci: L. gibbosus DeGeer 1874: 322. The carrot beetle Trinidad Distribution. Cuba, Jamaica, México, USA ( Collections. Pittsburgh. Washington FL) D. laevipunctatus Bates 1888: 311; Chapin 1932: Cuba localities. 295 Collections. Chalepus trachypygus Chevrolat 1865: 31 L. tumulosus (Burmeister) 1847: 101 Distribution. Cuba, México to Panamá (Heteronychus); Chevrolat 1865: 31; Gundlach Cuba localities. Ca: Jaronú. Ci: Baños Ciego 1891: 131; Chapin 1932: 298 Montera; Cayamas; Chaparra. Ha: Santiago de Distribution. Cuba, México, sw USA las Vegas. IJ: Neuva Gerona Cuba localities. Ca: Jaronú. Gu: Collections. Washington Guantánamo. Ho: Port Tanamo. IJ: Nueva D. minor Chapin 1935: 74 Gerona. M: Cárdenas. SC: San Nicolas. SS: Distribution. Cuba. Endemic Trinidad Cuba localities. IJ: Neuva Gerona. PR: San Collections. Gundlach no. 1085. Havana. Juan y Martinéz (t.l.); El Rangel Washington Collections. Havana? Washington (type) D. morator Fabricius 1798: 24 TRIBE ORYCTINI Chalepus trachypygus Burmeister 1847: 79; Chevrolat 1865: 31; Gundlach 1981: 146 Strategus Hope 1837: 87; key in Chapin 1932: Distribution. Cuba, Puerto Rico, e USA 299, 300 Cuba localities. PR: Rangel S. ajax (Olivier) 1889: 27 (Scarabaeus); Chapin Collections. Gundlach no. 1314 1932: 300; Ratcliffe 1976: 106 D. picipes (Burmeister) 1847: 79 (Chalepus); S. titanus, Jacquelin du Val 1856: 125; Chevrolat Chevrolat 1865: 31; Gundlach 1891: 145; 1865: 32; Gundlach 1891: 129; Ratcliffe l976: Chapin 1932: 293 106; 1988: 54 Chalepus geminatus (Fabricius) Jacquelin du Val Distribution. Cuba. Bahamas 1856: 127 Cuba localities. Widespread. Breeds in some Distribution. Cuba. Hispaniola, Puerto Rico, agaves, and may attack sugarcane Guadeloupe, México Collections. Cambridge. Chicago. Gundlach Cuba localities Widespread and common no. 436 (S. titanus). Howden. London. New Collections. Gundlach no. 35. Washington York. Pittsburgh. San Francisco. Washington S. anachoreta Burmeister 1847: 134; Jacquelin du Val 1856: 124; Chevrolat 1865: 33; Gundlach 94 Arthropods of Florida and Neighboring Land Areas, Vol. 18

1891: 130; Chapin 1932: 301; Ratcliffe 1976: Phileurus quadrituberculatus (Palisot de 114 Beauvois) 1806: 42 (Scarabaeus); Jacquelin du Distribution. Cuba. Endemic Val 1856: 124; Chevrolat 1865: 34; Gundlach Cuba localities. Widespread. A pest of coco- 1891: 129; distributed from México to Brazil, but nut palms, and may attack sugarcane also in Puerto Rico. Above early records from Collections. Cambridge. Halle (type). Cuba are probably misidentification of H. Gundlach no. 447. London. New York. Ottawa. cubanus, Chapin 1932: 304 Pittsburgh. San Francisco. Washington Distribution. Cuba. Endemic S. sarpedon (Klug) 1829: 11; Ratcliffe 1976: 140 Cuba localities. Ca: Camaguëy. Gu: Podalgus sarpedon Burmeister, Jacquelin du Val Guantánamo (t.l.). IJ: Nueva Gerona. M: 1856: 126 Cárdenas. PR: Rangel Scatophilus sarpedon Chevrolat 1865: 32; Collections. Gundlach no. 474 (P. Gundlach 1891: 131 quadrituberculatus). Pittsburgh. Washington Distribution. Cuba. Endemic Cuba localities. Widespread: May attack Phileurus Latreille 1807: 103 sugarcane P. valgus (Linnaeus) 1758: 347; Jacquelin du Val Collections. Type unknown. Cambridge. 1856: 123; Chevrolat 1865: 33; Gundlach 1891: Gundlach no. 100. Howden. London. New York. 128; Ratcliffe 1988: 52 Ottawa. Washington Distribution. Cuba, Jamaica, Puerto Rico, S. symphenax Ratcliffe 1976: 148 St. Croix, Guadeloupe, St. Vincent, Barbados, Distribution. Cuba. Endemic Grenada, México to Argentina, USA (southern) Cuba localities. PR: Las Martinas (t.l.) Cuba localities. Ca: Jaronú. CA: Baraguá. Collections. Ratcliffe. Washington (type) Ci: Cayamas. Ha: Havana; Santiago. M: Cárdenas TRIBE PHILEURINI Collections. Gundlach no. 130. Washington

Hemiphileurus Kolbe 1910: 336; see key in Planophileurus Chapin 1932: 307 Ratcliffe and Ivie 1998 P. planicollis (Chevrolat) 1865: 34 (Phileurus); H. cribratus (Chevrolat) 1844: 90 (Phileurus); Gundlach 1891: 129; Endrödi 1985: 704; Jacquelin du Val 1856: 123; Chevrolat 1865: 33; Ratcliffe 1988: 54 Gundlach 1891: 128; Chapin 1932: 306 Epiphileurus planicollis Kolbe 1910: 342 (Epiphileurus); Endrödi 1985: 713 Distribution. Cuba, Bahamas Distribution. Cuba. Endemic Cuba localities. Ha: Havana; Santiago. PR: Cuba localities. Widespread Heradura Collections. Gundlach no. 475. Paris (type?). Collections. Gundlach no. 150. Philadelphia, Washington Poey. Washington H. cubaensis (Chalumeau) 1979: 230 (Epiphileurus)., 1981: 178 (E. cubensis) SUBFAMILY CETONIINAE Distribution. Cuba. Endemic TRIBE GYMNETINI Cuba localities. “Cuba” (t.l.) Collections. Paris (type) Gymnetis MacLeay 1819: 152 H. laminicornis Dupuis and Dechambre 2000: 29 G. lanius (Linnaeus) 1766: 557 Distribution. Cuba. Endemic Gymnetis sternalis Chevrolat 1865: 35; Gundlach Cuba localities. SC: Gran Piedra (t.l.) 1891: 14; Chapin 1932: 310 Collections. Paris (type) Distribution. Cuba, eastward to St. Lucia H. parvus Dupuis and Dechambre 2000: 31 Cuba localities. M: Cienaga de Zapata. Ha: Distribution. Cuba. Endemic Havana. SC: Manacas. VC: in caves with dry Cuba localities. “Cuba” guano as larval food Collections. Paris (type) Collections. Gundlach no. 1203. Washington

Homophileurus Kolbe 1910: 336; key in Chapin Tiarocera Burmeister 1842: 247 1932: 304 T. cornuta (Gory and Percheron) 1833: 69 H. cubanus Prell 1911: 106; Chapin 1932: 304; (Gymnetis); Jacquelin du Val 1857: 136; Chapin Endrödi 1985: 711 1932: 309 A Checklist of the Beetles of Cuba 95

Allorhina (Tiarocera) cornuta, Chevrolat 1865: trations of female genitalia of twelve other 34; Gundlach 1891: 146 Phyllophaga. Pan-Pac. Ent. 60: 313-325. Distribution. Cuba. Endemic Garcia Vidal, M. 1987. The genus Phyllophaga Cuba localities. Widespread. CA: Baraguá. Harris 1826 (Coleoptera: Scarabaeidae) in Cuba. Gu: Guantánamo. Ha: in cave with dry guano IV. Description of six new species. J. New York as larval food. PR: Cerro Cabras. SC: Santiago Ent. Soc. 95: 302-306. Collections. Gatineau. Gundlach no. 437. Garcia Vidal, M. 1988. The genus Phyllophaga Washington Harris 1826 (Coleoptera: Scarabaeidae) in Cuba. III. Descriptions of six new species and the male REFERENCES of P. chapini. Pan-Pac. Ent. 64: 101-106. Cartwright, O. L. 1973. Ataenius, Aphotaenius, Gordon, R.D. and R. Pittino. 1992. Current sta- and Pseudataenius of the United States and tus of the American genera and species of Canada (Coleoptera; Scarabaeidae; Aphodiinae). Psammodiini (Coleoptera: Scarabaeidae: Smithsonian Cont. Zool. 154: 105 pp. Aphodiinae). Coleop. Bull. 46: 260-273. Cartwright, O.L. and R.E. Woodruff. 1969. Ten Howden, H.F. and B.D. Gill. 2000. Tribes of New Rhyparus from the Western Hemisphere (Co- World Ceratocanthinae, with keys to genera leoptera: Scarabaeidae: Aphodiinae). and descriptions of new species (Coleoptera: Smithsonian Cont. Zool. 21: 1-20. Scarabaeidae). Sociobiology 35: 281-329. Chalumeau, F. 1979. Diagnose preliminaire de Joly, L.J. 1995. Cyclocephala rangelana Chapin y quatre nouveaux Scarabaeidae de Cuba. Nouv. Cyclocephala vidua Endrödi, dos especies Rev. Ent. 9: 229-230. Antillanas estrechamente relacionadas (Co- Chalumeau, F. 1981. Scarabaeoidea (Coleoptera) leoptera, Melolonthidae, Dynastinae, recoltés par V. Decou. Résultats des expéditions Cyclocephalini). Bol. Ent. Venez. N. s. 10: 57- biospéologiques Cubano-Roumaines à Cuba 3: 67. 173-180. Matthews, E.G. 1966. A taxonomic and zoogeo- Chalumeau, F. 1989. Phyllophaga Harris 1826 graphic survey of the Scarabaeinae of the des Antilles (Melolonthinae): espece nouvelle Antilles (Coleoptera: Scarabaeidae). Mem. designation de lectotypes et synonomies. Nouv. Amer. Ent. Soc. 21. 134 pp. Rev. Ent. 6: 330-340. Matthews, E.G. 1969. New data on Antillean Dupuis, F. and R.-P. Dechambre. 2000. Dix Scarabaeine beetles, and two new species from nouvelles espèces d’Hemiphileurus Kolbe, 1910 Hispaniola. Psyche 76: 114-125. (Coleoptera, Dynastidae). Cóleoptères 6: 21-32. Muslera Ramos, L. and Fernandez Garcia, I. Endrödi, S. 1985. The Dynastinae of the World. 1998. Actual stsate of knowledge of genus Series Entomol., vol. 28. Dr. W. Junk Publ., Phyllophaga (Coleoptera: Scarabaeidae) in Cuba Dordrecht. 800 pp. – Estado actual de conocimiento del genero Fernandez Garcia, I. 1998. Algunos aspectos Phyllophaga (Coleoptera: Scarabaeidae) en ecologicos de la communidad de coleopteros de Cuba. Poeyana 461: 1-22. un agroecosistema canero en la provinicia de Ratcliffe, B.C. 1976. A revision of the genus La Habana. Poeyana 460: 1-14. Strategus (Coleoptera: Scarabaeidae). Bull. Garcia Vidal, M. 1975. Compilacion sobre los Univ. Nebraska St. Mus. 10: 93-204. principales aspectos relativos al genero Ratcliffe, B.C. 1988. New species and distribu- Phyllophaga. Revista Agrotec. Cuba 7: 17-22. tions of Neotropical Phileurini and a new Garcia Vidal, M. 1978. El Genero Phyllophaga Phileurine from Burma (Coleoptera: Harris 1826 (Coleoptera: Scarabaeidae) en Cuba Scarabaeidae: Dynastinae). Coleop. Bull. 42: 43- I. Descripcion de cinco neuvas especias. 55. Poeyana 182: 1-14. Ratcliffe, B.C., and M.A. Ivie. 1998. New spe- Garcia Vidal, M. 1982. The genus Anoplosiagum cies of Hemiphileurus Kolbe (Coleoptera: Blanchard (Coleoptera, Scarabaeidae) in Cuba. Scarabaeidae: Dynastinae) from the Domini- Part I. Descriptions of five new species. Pan- can Republic with a key to the West Indian Pac. Ent. 58: 330-335. species of Hemiphileurus. Coleop. Bull. 52: 201- Garcia Vidal, M. 1984. The genus Phyllophaga 208. Harris 1826 (Coleoptera: Scarabaeidae) in Cuba. Stebnicka, Z. 2002. The New World species of II. Descriptions of eleven new species and illus- Ataenius Harold, 1867. II. Revision of the West Indian A. terminalis-group (Coleoptera: 96 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Scarabaeidae: Eupariini). Acta Zool. Cracov. 45: REFERENCES 259-281. Endrody-Younga, S. 1981. The American species Vàzquez, L.L., E. Valdés, and J.C. Amor. 1992. of the Familia Clambidae (Coleoptera, New manifestations of pests on economically Eucinetoidea). Ent. Generalis 7: 33-67 important plants during the period from 1970 Endrody-Younga, S. 1998. The genus Clambus to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo in Central and South America (Coleoptera: Silvestri 49: 41-52. Eucinetoidea: Clambidae). Ann. Transvaal Woodruff, R.E. 1973. The scarab beetles of Florida. Museum 36: 373-403 Arthropods of Florida Vol. 8. 220 pp. Woodruff, R.E. and B.M. Beck. 1989. The scarab 47. FAMILY SCIRTIDAE beetles of Florida. Part II. The May beetles (ge- The marsh beetles nus Phyllophaga). Arthropods of Florida Vol. This family usually occurs on vegetation near 13. 225 pp. water or in damp places. Ora and Scirtes are mostly Woodruff, R.E. and O.L. Cartwright. 1967. A tropical, and have enlarged hind legs for jumping. review of the Genus Euparixia with descrip- The larvae usually live in water in streams, ponds, tion of a new species from nests of leaf-cutting swamps, and even bromeliad bases, and feed on a ants in Louisiana. Proc. U.S. Nat. Mus. 123: diversity of organic matter. 1-21, no. 3616. Zayas, F. de and E.G. Matthews. 1966. Revi- SUBFAMILY HELODINAE sion de los Cantoninos de Cuba (Coleoptera; Scarabaeidae) con la descripción de seis nuevas Elodes Latrielle 1796: 44 especias. Poeyana 14: 1-24. E. angustata Chevrolat 1870: 68; Gundlach 1891: 245 SERIES ELATERIFORMIA Distribution. Cuba. Endemic 42. FAMILY PODABROCEPHALIDAE Cuba localities. SC: Santiago de Cuba This family is based on Podabrocephalus of Collections. Gundlach no. 1777 southern India. E. xanthura Chevrolat 1870: 68; Gundlach 1891: 244 43. FAMILY RHINORHIPIDAE Distribution. Cuba. Endemic This family is based on Rhinorhipus of east- Cuba localities. M: Cárdenas; Cienaga de ern Australia. Zapata Collections. Gundlach no. 1041 SUPERFAMILY SCIRTOIDEA 44. FAMILY DECLINIIDAE Ora Clark 1865: 385 This family is known only from the Soviet Far O. bicolor Zayas 1988: 30 East. Distribution. Cuba. Endemic Cuba localities. SC: Siboney (t.l.) 45. FAMILY EUCINETIDAE Collections. Zayas (type) The family is almost world-wide in distribu- O. guttata Zayas 1988: 30 tion and probably occurs in Cuba. Adults and lar- Distribution. Cuba. Endemic vae are found in leaf litter, and under bark. They Cuba localities. Ha: Havana (t.l.) feed on fungus or slime molds. Collections. Zayas (type) O. interrupta (Chevrolat) 1870: 69 (Scirtes); 44. FAMILY CLAMBIDAE Gundlach 1891: 246 The minute beetles Distribution. Cuba. Endemic This family probably occurs in Cuba. It is Cuba localities. M: Cárdenas known from North, Central and South America, Collections. Gundlach no. 556 but in the West Indies only on Hispaniola (Endrody- O. sexlineata (Chevrolat) 1870: 69 (Scirtes); Younga 1981, 1998). Species are usually found in Gundlach 1891: 245 debris piles or leaf litter, where they probably feed Distribution. Cuba. Endemic on fungal spores; some may be predators on scale Cuba localities. M: Cienaga de Zapata insects. Collections. Gundlach no. 962 A Checklist of the Beetles of Cuba 97

Scirtes Illiger 1807: 301 P. (P.) angulosa Jacquelin du Val 1856: 62; S. apicalis Chevrolat 1870: 70; Gundlach 1891: 246 Chevrolat 1867: 582; Gundlach 1891: 166; Distribution. Cuba. Endemic Fisher 1925: 20 Cuba localities. M: Cárdenas P. excavata Kerremans 1904: 518 Collections. Gundlach no. 692. Distribution. Cuba, Bahamas, USA (AL-GA- S. cinctipennis Chevrolat 1870: 69; Gundlach 1891: FL, Cazier 1952) 246 Cuba localities. Gu: Caimanera Distribution. Cuba. Endemic Collections. Gundlach no. 807. New York. Cuba localities. M: Cárdenas Philadelphia. Washington Collections. Gundlach no. 900 P. (P.) chevrolati Thomson 1878: 43; Kerremann S. fusca Chevrolat 1870: 70; Gundlach 1891: 246 1906: 488 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas; Cienaga de Cuba localities. Zapata Collections. London (type) Collections. Gundlach no. 652 P. (P.) goryi Saunders 1871: 58; Fisher 1925: 12; Cobos 1981: 78 SUPERFAMILY DASCILLOIDEA Distribution. Cuba, Bahamas, Mexico 48. FAMILY DASCILLIDAE (Yucatan), Colombia, Venezuela, French This family occurs in North, Central, and South Guyana America but is seemingly not reported from the Cuba localities. Caribbean. Collections. P. (Tularensis) cubae Chevrolat 1838: 55; 1867: 582; 49. FAMILY RHIPICERIDAE Gory 1840: 141; Jacquelin du Val in Sagra 1856: This family occurs in North, Central, and South 62; Gundlach 1891: 165; Kerremans 1906: 500; America but is seemingly not the Caribbean re- Fisher 1925: 10. gion. Larvae of species of Sandalus are Distribution. Cuba. Endemic ectoparasitoids on cicadas, which do occur in the Cuba localities. Gu: Guantánamo Caribbean region. Collections. Gundlach no. 861. New York. Washington SUPERFAMILY BUPRESTOIDEA 50. FAMILY TRIBE This family only occurs in the southwestern USA, and larvae may be root feeders. Eschscholtz 1829: 9 A. cubaecola Jacquelin du Val 1856: 57; Chevrolat 51. FAMILY BUPRESTIDAE 1867: 583; Gundlach 1891: 167; Kerremanns The metallic wood-boring beetles 1906: 61; Fisher 1925: 39 Reviewed by Richard Westcott, March, 1997; Distribution. Cuba, USA (FL) Charles L. Bellamy, November, 2001; T. C. Cuba localities. Gu: Caimanera MacRae, December, 2001. Collections. Gundlach no. 1414. London. Adults are active daytime flyers. They can be Philadelphia. Washington found feeding on leaves, pollen, petals and on nec- A. marginenotata Chevrolat 1867: 583; Gundlach tar of flowers. The larvae bore in both living and 1891: 167; Kerremanns 1906: 56; Fisher 1925: dead plant tissue under bark, in twigs, in roots of 34 trees and shrubs, and stems of herbaceous plants. A. cubaecola Fall 99: 35 (not A. cubaecola See Cazier (1952) for key to genera in Bahamas Jacquelin du Val) and Fisher (1925) for a review of the West Indies Distribution. Cuba, Bahamas, USA (FL) fauna, keys to species, and Cuba localities and col- Cuba localities. Ci: Cienfuegos. Gu: lections. Caimanera Collections. Gundlach no. 809 SUBFAMILY A. pulcherrima Jacquelin du Val 1856: 56; TRIBE POLYCESTINI Chevrolat 1867: 583; Gundlach 1891: 166; Fall 1899: 16; Kerremanns 1906: 53; Fisher 1925: Polycesta Solier 1833: 281 42 Distribution. Cuba. Endemic 98 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Gu: Caimanito; Guantánamo SUBTRIBE PRISTIPTERINA Collections. Gundlach no. 808. New York. Philadelphia, Washington Chrysesthes Solier 1833: 290 A. subcylindrica Fisher 1925: 30 C. lanieri (Chevrolat) 1838: 280 (Buprestis); Distribution. Cuba. Endemic Jacquelin du Val 1856: 61; Chevrolat 1867: 574 Cuba localities. Ci: Cayamas (t.l.) (Pelecopselaphus); Gundlach 1891: 157; Fisher Collections. Washington (type) 1925: 67 Chrysesthes persimeri Obenberger 1926: 154 TRIBE TYNDARINI Distribution. Cuba. Endemic Cuba localities. Ca: Camaguëy. M: Cárdenas. Tyndaris Thomson 1857: 168 Larvae live in royal palms T. (Knulliella) mojito Bily 1987: 45 Collections. Gundlach no. 244. Philadelphia. Distribution. Cuba. Endemic Washington Cuba localities. SC: Daiquiri. On Coccoloba Collections. Prague? Halecia Laporte and Gory 1837: 108 H. verecunda Chevrolat 1867: 578; Gundlach 1891: TRIBE MASTOGENIINI 156; Fisher 1925: 79 Distribution. Cuba. Endemic Mastogenius Solier 1852: 507 Cuba localities. Gr: Bayamo (M. castlei Champlain and Knull 1922: 145; Collections. Gundlach no. 792 Westcott 1990: 256) M. antennatus Cazier 1952: 6 SUBTRIBE EUPLECTALECINA Distribution. Cuba?; Bahamas, USA (FL). Reared from pods of Tamarindus indica Euplectalecia Obenberger 1924: 20 Cuba localities. E. quadricolor (Chevrolat) 1867: 574 (Halecia); Collections. Gundlach 1891:157 Distribution. Cuba. Endemic Micrasta Kerremans 1893: 115 Cuba localities. Gr: Bayamo M. cubensis Fisher 1930: 10 Collections. Gundlach no. 1417 Distribution. Cuba. Endemic Cuba localities. SUBTRIBE EUCHROMATINA Collections. M. subcylindrica Fisher 1939: 165 Euchroma Solier 1833: 284 Distribution. Cuba. Endemic E. gigantea Linnaeus 1764: 85; Fisher 1925: 62 Cuba localities. E. g. harperi Sharp 1881: 298. Cuba, Guadeloupe, Collections. Guyana Distribution. Cuba, Jamaica, Guadeloupe, SUBFAMILY México to Argentina TRIBE CHALCOPHORINI Cuba localities. SUBTRIBE CHALCOPHORINA Collections. London

Chalcophora Solier 1833: 278 SUBTRIBE AGAEOCERINA C. virginiensis (Drury) 1872: 66 (Buprestis); Chevrolat 1867: 573; Gundlach 1891: 155; Peronaemis Waterhouse 1887: 178 Fisher 1925: 76 Paracinyra Fisher 1925: 133; Bellamy 2000:163 Distribution. Cuba (introduced?), México, P. elegans (Fisher) 1930: 127 (Paracinyra) Guatemala, Honduras, USA (widespread), Distribution. Cuba. Endemic Canada Cuba localities. Cuba localities. Gundlach (1891) states that Collections. the species was introduced by ships to Cuba P. monticola (Fisher) 1936: 343 (Paracinyra) from the USA. It feeds on pine trees Distribution. Cuba. Endemic Collections. Cuba localities. SC: Pico Turquino (t.l.) Collections. Havana (Bruner?). Washington (type) A Checklist of the Beetles of Cuba 99

P. viridimaculata (Fisher) 1925: 134 (Paracinyra); TRIBE DICERCINI Bellamy 2000: 163 SUBTRIBE DICERINA Distribution. Cuba. Endemic Cuba localities. Gu: Guantánamo (t.l.) Dicerca Eschscholtz 1829: 9 Collections. New York (type) D. tuberculata Laporte and Gory 1837: 99; P. viridithorax Zayas 1988: 39; possibly a synonym Chevrolat 1867: 577; Gundlach 1891: 159. The of P. monticola (Fisher) (Bellamy 2000: 163) turpentine beetle Distribution. Cuba. Endemic Distribution. Cuba (introduced), widespread Cuba localities. SC: Paso de Calas, below Pico USA (not FL), Canada Turquino (t.l.) Cuba localities. Ha: Havana. Gundlach Collections. Zayas (type) (1891) states that the species was imported into Cuba in lumber TRIBE PSILOPTERINI Collections. Gundlach no. 1542

Hilarotes Thomas 1778: 39 SUBTRIBE PHRIXIINA H. chalcoptera (Jacquelin du Val) 1856: 59 (Buprestis); Chevrolat 1867: 578; Gundlach Spectralia Casey 1909: 175 1891: 161; Fisher 1925: 73 S. costulifera (Chevrolat) 1867: 579 (Cinyra); Distribution. Cuba. Endemic Gundlach 1891: 162; Fisher 1925: 138 Cuba localities. Distribution. Cuba. Endemic Collections. Gundlach no. 308. London. Cuba localities. Widespread Philadelphia. Washington Collections. Gundlach no.105. New York. Philadelphia. Washington Lampetis Spinola 1837: 120. S. multipunctata (Olivier) 1790: 86 (Buprestis); [L. aurifer (Olivier) 1790: 13 (Buprestis); Herbst Jacquelin du Val 1856: 61; Chevrolat 1867: 579; 1801: 185; Fabricius 1801: 191; Schonherr Gundlach 1891: 162; Fisher 1925: 140 1817: 219; Jacquelin du Val 1856: 58; Chevrolat Distribution. Cuba. Endemic 1867: 575; Gundlach 1891: 158; Kerremanns Cuba localities. Ha: Havana; Bahia Honda. 1910: 57; Fisher 1925: 54 (all as Psiloptera)] M: Fermina Distribution. Hispaniola, South America (?) Collections. Gundlach no.1261. London. Cuba localities. Doubtful old records (see Washington Fisher 1925: 56) S. sulcicollis (Chevrolat) 1867: 580 (Cinyra); Collections. Gundlach 1891: 163; Fisher 1925: 137 L. straba (Chevrolat) 1867: 575 (Psiloptera); Distribution. Cuba. Endemic Gundlach 1891: 158; Kerremans 1910: 58 (all Cuba localities. Gu: Caimanera as Psiloptera) Collections. Gundlach no.1418 Distribution. Cuba. Endemic Cuba localities. IJ: Nueva Gerona. Gu: San TRIBE BUPRESTINI Francisco; Vuelta Abaja. PR: Viñales. Feeds on pine needles Buprestis Linnaeus 1758: 408 Collections.. Gundlach no. 1257. New York.. B. (Buprestis) cubensis Fisher 1925: 157 Philadelphia. Pittsburgh. Washington Distribution. Cuba. Endemic L. torquata (Dalman) 1823: 54 (Buprestis); Cuba localities. PR: Viñales (t.l.) Jacquelin du Val 1856: 59; Castelnau and Gory Collections. New York 1837: 36; Chevrolat 1867: 576; Gundlach 1891: B. (Buprestis) decipiens Fisher 1925: 158 159; Kerremanns 1910: 55; Fisher 1925: 51 (all Distribution. Cuba. Endemic as Psiloptera) Cuba localities. IJ: Nueva Gerona. PR: Distribution. Cuba, Jamaica Viñales (t.l.) Cuba localities. Ci: Cayamas. IJ: Nueva Collections. Carnegie. New York. Washing- Gerona. M: Zapata ton Collections. London. New York. Pittsburgh. B.(Buprestis) lineata Fabricius 1775: 217; Chevrolat Washington 1867: 577; Gundlach 1891: 160; Fisher 1925: 152 100 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Ancylochira lineata Castelnau and Gory 1838: Distribution. Cuba. Endemic 143; Gundlach 1891: 623 Cuba localities. SC: Loma del Gato (t.l.) Distribution. Cuba, Puerto Rico, Guadeloupe, Collections. Washington (type) Argentina, USA (TX-IN-FL), Canada Cuba localities. Ca: Camaguëy. M: Cienaga Xenorhipis LeConte 1866: 384 de Zapata. SC: Santiago. Larvae live in pine X. vauriei Cazier 1952: 3 trees Isophaenus israeli Zayas 1988: 40; Ivie 1991: 400 Collections. Gundlach no. 1416 Distribution. Cuba; Bahamas B. (Buprestis) zayasi Fisher 1952: 329 Cuba localities. Gu: Guantánamo, Distribution. Cuba. Endemic Tortuguilla (t.l.) Cuba localities. Ho: Pinares de Moa (t.l.) Collections. Zayas (type) Collections. Washington B. (Sterosa) apricans Herbst 1801: 125; Fisher SUBTRIBE ANTHANIINA 1925: 147 Distribution. Cuba (introduced), USA (TX- Anthaxia Eschscholtz 1829: 9 NY-FL) [A. fulgurans (Schrank) 1789: 85; Bily 1991: 126] Cuba localities. Ha: Havana. Probably in- A. bifoveata Fisher 1925: 168. New York (type) troduced to Cuba in pine lumber Distribution. A European species; the Cuba Collections. Washington record is an error in labeling B. (Stereosa) decora Fabricius 1775: 217; Chevrolat A. viridifrons Gory 1841: 248 1867: 578; Gundlach 1891: 161; Fisher 1925: A. subsinuata Gory 1841: 289; Jacquelin du Val 148 1856: 65; Chevrolat 1867: 581; Gundlach 1891: Ancylochira decora Castelnau and Gory 1838: 164; Fisher 1925: 169; synonomy in Bily 1991: 143; Gundlach 1891: 623 136 Distribution. Cuba, Puerto Rico, Guadeloupe, Distribution. Cuba, widespread eastern USA USA (TX-NJ-FL) Cuba localities. “Cuba” Cuba localities. Collections. Paris (type) Collections. Gundlach no.1630. Philadelphia TRIBE CHRYSOBOTHRINI TRIBE MELANOPHILINI SUBTRIBE CHRYSOBOTHRINA

Melanophila Eschscholtz 1829: 9 Chrysobothris Eschscholtz 1829: 9 M. acuminata (DeGeer) 1874: 133 (Buprestis) C. cubensis Théry 1927: 31 M. longipes Say 23-164; Chevrolat 1867: 581; Distribution. Cuba. Endemic Gundlach 1891: 164 Cuba localities. Distribution. Cuba, Bahamas, Hispaniola, Collections. México to Honduras, USA (CA-FL-NJ-MO), C. fisheri Théry 1927: 31 introduced to Old World? Distribution. Cuba. Endemic Cuba localities. Ca: Camaguëy. SC: Brazo el Cuba localities. Cauto Collections. Collections. Gundlach no. 1853 C. lepida Laporte and Gory 1837: 16; Jacquelin du M. notata (Laporte and Gory) 1837: 4; Chevrolat Val 1856: 64; Chevrolat 1867: 585; Gundlach 1867: 580; Gundlach 1891: 163; Fisher 1925: 1891: 168; Fisher 1925: 110 171 Distribution. Cuba, Bahamas, Puerto Rico Distribution. Cuba, Bahamas, México, USA Cuba localities. Widespread (e, FL) Collections. Gundlach no. 505. New York. Cuba localities. IJ Philadelphia. Washington Collections. Gundlach 1705 C. sexfasciata Schaeffer 1918: 212; Fisher 1925: 127 TRIBE ANTHAXIINI Distribution. Cuba, Bahamas, Jamaica, USA SUBTRIBE XENORHIPIDINA (FL) Cuba localities. Ca: Camagüey. Ci: Cayamas Isophaenus Waterhouse 1889: 175 Collections. Washington I. acunai Fisher 1936: 272 A Checklist of the Beetles of Cuba 101

C. tranquebarica (Gmelin) 1788 (1790): 1932 Leiopleura Deyrolle 1864: 219 (Buprestis); Fisher 1925: 96. The Australian L. compactilis Chevrolat 1867: 588; Gundlach 1891: Pine borer 171; Fisher 1925: 194 C. impressa Chevrolat 1867: 586; Gundlach 1891: Distribution. Cuba. Endemic 169 (not Fabricius) Cuba localities. M: Guamarco, between Distribution. Cuba, Bahamas, Hispaniola, Cárdenas and Mantanzas Puerto Rico, Dominica, Grenada, Guadeloupe, Collections. Gundlach no. 862 Surinam, USA (FL). Larvae bore in Casuarina L. cupeyali Zayas 1988: 44. (possibly Neotrachys, and Rhizophora; also in Cassia, Conocarpus, Ivie 1991: 400) and Pinus pallustris (Benoit, 1966, Can. Ent. Distribution. Cuba. Endemic 98: 324-330) Cuba localities. SC: Cupeyal (t.l.) Cuba localities. Collections. Zayas (type) Collections. L. cyanea Zayas 1988: 44. (possibly Neotrachys, C. tumida Chevrolat 1867: 585; Gundlach 1891: Ivie 1991: 400) 168; Fisher 1925: 99 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Cupeyal (t.l.) Cuba localities. Collections. Zayas (type) Collections. Gundlach no. 839. Philadelphia. L. otero (Fisher) 1935: 52 (Enbrachys); Nelson et Washington al. 1981: 144 Distribution. Cuba. EndemicUSA (FL, a SUBTRIBE ACTENODINA probable introduction which has died out) Cuba localities. Ha: Santiago de las Vegas Actenodes Lacordaire 1857: 72 (t.l.) A. auronotatus (Laporte and Gory) 1837: 20 Collections. Washington (type) (Chrysobothris); Jacquelin du Val 1856: 64; Chevrolat 1867: 584; Gundlach 1891: 168; SUBTRIBE BRACHYINA Fisher 1925: 88 Distribution. Cuba, Jamaica, Hispaniola, Taphrocerus Solier 1833: 314 USA (FL-GA) T. aeneocupreus Fisher 1925: 190 Cuba localities. Widespread. Distribution. Cuba. Endemic Collections. Gundlach no. 201. London. New Cuba localities. Ci: Cayamas (t.l.) York. Philadelphia. Pittsburgh. Washington Collections. Washington (type) A. caray Zayas 1988: 41 T. bruneri Fisher 1930: 8. Distribution. Cuba. Endemic T. brunneri Obenberger 1934: 50 Cuba localities. SC: Ciudad Mar (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. CA: Central Baraguá (t.l.) Collections. Havana. Washington (type) SUBFAMILY T. laesicollis Chevrolat 1867: 587; Gundlach 1891: TRIBE AGRILINI 170; Fisher 1925: 183 Distribution. Cuba. Endemic Curtis 1825: 67 Cuba localities. A. denticornis Chevrolat 1867: 586; Gundlach 1891: Collections. 170; Fisher 1925: 200 T. tao Zayas 1988: 41 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cienaga de Zapata Cuba localities. PR: Viñales (t.l.) Collections. Gundlach no. 959 Collections. Zayas (type) A. dominicanus Thomas 1878: 83; Fisher 1925: 198 T. timidus Chevrolat 1867: 587; Gundlach 1891: Distribution. Cuba, Hispaniola 171; Fisher 1925: 191; Fernandez Garcia 1998: Cuba localities. Gu: Guantánamo, Rio Seco 3 Collections. London. New York Distribution. Cuba. Endemic Cuba localities. M: Cienaga de Zapata. PR: TRIBE TRACHINI Viñales. On Juncu SUBTRIBE LEIOPLEURINA Collections. Gundlach no. 1053. New York. Washington 102 Arthropods of Florida and Neighboring Land Areas, Vol. 18

REFERENCES C. quadrata (Darlington) 1927: 95 (Helmis); Bellamy, C. L. 2000. New synonymy and combi- Darlington 1936: 81 (Helmis); Spangler 1973: nations in Antillean Buprestidae (Coleoptera). 356; 1981: 169 Ann. Transvaal Mus. 37: 163-164. Distribution. Cuba. Endemic Bily, S. 1991. Taxonomical notes on Anthaxia with Cuba localities. Ci: Soledad (t.l.). PR: Rio Los descriptions of new taxa (Coleoptera, Palacios Buprestidae). Acta Ent. Bohemoslovaca 88: 121- Collections. Cambridge (type) 137. Cazier, M. 1952. Additions to the Buprestid fauna Neoelmis Musgrave 1935: 34 of the Bahama Islands, British West Indies N. minima (Darlington) 1927: 93 (Helmis); (Coleoptera: Buprestidae). Amer. Mus. Darlington 1936: 79 (Helmis caesa minima); Novitates 1562. 10 pp. Spangler 1973: 357; 1981: 165 Fernandez Garcia, I. 1998. Algunos aspectos Distribution. Cuba, Jamaica, Hispaniola ecologicos de la comunidad de coleopteros de un Cuba localities. Ci: Soledad (t.l.). Gu: Arroyo agroecosistema cañero en la provincia de La de la Poa; Baracoa; Rio Contramaestra; Rio Habana. Poeyana no. 460, 114. Yumuri.. Ho: Mayari; Rio Brazo Seco; Rio Fisher, W.S. 1925. A revision of the West Indian Seboruco. IJ: Arroyo La Talega. PR: Rio Los Coleoptera of the family Buprestidae. Proc. U.S. Palacios Nat. Mus. 65(8): 1-207. Collections. Cambridge (type). Washington Ivie, M. A. 1991. Taxonomic notes on a little known publication: Zayas, 1988, Entomofauna Cubana. Xenelmoides Hinton 1935: 5 Order Coleoptera. Coleop. Bull. 45: 399-401. X. simplex (Darlington) 1927: 96; 1936: 78 (Helmis); Spangler 1973: 357; 1981: 164 SUPERFAMILY BYRRHOIDEA Distribution. Cuba. Endemic 52. FAMILY Cuba localities. Ci: Soledad (t.l.). Gu: Rio The family is mostly temperate in distribution. Baracoa at Tinta. IJ: Arroyo de Talega. PR: It is not known from the West Indies. Martinius Rio Los Palacios, Las Villas; Rio Cuburny; Rio ripisaltator Spilman of Cuba (which Spilman called Cuyaguateje a byrrhid (1959, Coleop. Bull. 13: 118)) is now placed Collections. Cambridge (type). Washington in the family Limnichidae. SUBFAMILY LARAINAE 53. FAMILY ELMIDAE The riffle beetles Phanocerus Sharp 1882: 128 These beetles occur where there is running P. clavicornis Sharp 1882: 129; Spangler 1973: 356; water. Adults and larvae of most species occur on 1981: 164; Spangler and Santiago-Fragoso 1992: submerged rocks and in debris and gravel. 13 Darlington (1927) gives an outdated key to the Cu- Distribution. Cuba, Jamaica, Hispaniola, ban species. Puerto Rico, México to Panama, USA (TX) Cuba localities. Widespread SUBFAMILY ELMINAE Collections. Cambridge. Washington TRIBE ELMINI REFERENCES Cylloepus Erichson 1847: 521 Darlington, P.J., Jr. 1927. Four new Helmidae C. filiformis (Darlington) 1927: 94 (Helmis); from Cuba, with notes on other West Indian Darlington 1936: 79 (Helmis); Spangler 1973: species. Psyche 34: 91-97. 357; 1981: 165 Spangler, P.J. 1973. Aquatic Coleoptera collected Distribution. Cuba; Jamaica. C. filiformis in the biospeleological expeditions. Resultats des jamaicensis (Darlington, 1927) occurs in Ja- expeditions biospéologiques Cubano-Roumaines maica à Cuba 1: 353-358. Cuba localities. Ci: Soledad (t.l.). Gu: Rio Jojo, Spangler, P.J. 1981. Supplement to the aquatic Veguita del Sur; Rio Yumuri. PR: Rio Los and semiaquatic Coleoptera of Cuba. Resultats Palacios des expeditions biospéologiques Cubano- Collections. Cambridge (type). Washington Roumaines à Cuba 3: 145-171. A Checklist of the Beetles of Cuba 103

Spangler, P. J., and S. Santiago-Fragoso. 1992. Throscinus LeConte 1874: 51 The aquatic beetle subfamily Larainae (Co- T. aethiops Darlington 1936: 75; Wooldridge 1981: leoptera: Elmidae) in México, Central America, 220 and the West Indies. Smith. Cont. Zool. 528: 1- Distribution. Cuba, Hispaniola, Jamaica, 74. Puerto Rico, USA (FL) Cuba localities. None given 54. FAMILY DRYOPIDAE Collections. Cambridge (type) The long-toed water beetles This family has aquatic larvae and some adults SUBFAMILY LIMNICHINAE are also aquatic. They occur in streams, in rotten TRIBE LIMNICHINI logs, leaf litter, and flood debris, and adults come to lights. Eulimnichus Casey 1889: 146. Key in Wooldridge 1979 Pelonomus Erichson 1847 E. ater (LeConte) 1854: 117 (Limnichus); P. obscurus gracilipes Chevrolat 1864c: 406; Wooldridge 1978: 166 Gundlach 1891: 51; Darlington 1936: 77 Distribution. Cuba, Jamaica, México, Cen- Distribution. Cuba, Bahamas, USA (OK-DE- tral America, Colombia and Venezuela to Ar- FL). P. o. obscurus is in México, Guatemala, gentina, USA (TX-IN-FL) and Texas Cuba localities. Cuba localities. Ci: Soledad. M: Cárdenas; Collections. Cambridge Cienaga de Zapata Collections. Cambridge Limnichites Casey 1889: 146 L. rudis Wooldridge 1977: 183 55. FAMILY LUTROCHIDAE Distribution. Cuba, Hispaniola, Jamaica The travertine beetles Cuba localities. Ci: Soledad (t.l.) Both adults and larvae of this family are Collections. Cambridge (type) aquatic, and occur on rocks in rapids and flowing areas of streams with high calcium content. They SUBFAMILY THAUMASTODINAE occur from the southern United States to Brazil. Martinius Spilman 1959: 118 Erichson 1847: 509 M. ripisaltator Spilman 1966: 124 L. geniculatus Chevrolat 1864c: 406; Gundlach Distribution. Cuba. Endemic 1891: 51; Darlington 1936: 75; Spangler 1981: Cuba localities. Ha: Havana (t.l.), seashore. 166 Adults and larvae may be in moist sand or de- Distribution. Cuba; Hispaniola bris at the high tide line Cuba localities. Gu: Guantánamo, Rio Collections. Washington (type) Guasco; Rio Mogote at Matias Collections. Cambridge. Washington REFERENCES Spilman, T. J. 1966. A new species of Martinius REFERENCES from Cuba (Coleoptera: Limnichidae). Coleop. Spangler, P.J. 1981. Supplement to the aquatic Bull. 20: 123-125. and semiaquatic Coleoptera of Cuba. Resultats Wooldridge, D. P. 1977. New World Limnichinae des expeditions biospéologique Cubano- III. A revision of Limnichites Casey (Coleoptera: Roumaines à Cuba 3: 145-171. Limnichidae). Great Lakes Ent. 10: 179-189. Wooldridge, D. P. 1978. New World Limnichinae 56. FAMILY LIMNICHIDAE IV. Eulimnichus Casey. A. Synonomies, lecto- The minute marsh-loving beetles type designations, and redescription (Co- Adults often come to lights at night. Adults and leoptera: Liminichidae). Great Lates Ent. 11: larvae may live in soil under moss and algae, at 163-173. stream edges, and in flood debris. Wooldridge, D. P. 1979. New World Limnichidae IV. Eulimnichus Casey. B. Descriptions of new SUBFAMILY CEPHALOBYRRHINAE species (Coleoptera: Limnichidae). Great Lakes Ent. 12: 1-11. 104 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Wooldridge, D.P. 1981. Three new species of INCERTAE SEDIS Throscinus LeConte, with notes on other spe- cies (Coleoptera: Limnichidae, Cephalo- Heterocerus Fabricius 1792: 262 byrrhinae). Coleop. Bull. 35: 217-221. H. decimmaculatus Chevrolat 1864c: 407; Gundlach 1891: 53; Pacheco 1964: 122 (the iden- 57. FAMILY HETEROCERIDAE tity of this species is unknown) The variegated mud-loving beetles Distribution. Cuba. Endemic? Both adults and larvae construct tunnels in Cuba localities. M: Cárdenas sand or mud at the edges of streams and ponds, Collections. Gundlach no. 710 where they feed on algae or diatoms (see Pacheco 1964). REFERENCES Bameul, F. 1995. Un nouveau Tropicus Pacheco SUBFAMILY HETEROCERINAE de la Guadeloupe (Coleoptera, Heteroceridae). TRIBE HETEROCERINI Bull. Soc. Ent. France 100: 475-480. Ivie, M. A. and J. B. Stribling. 1984. Taxonomic Filiolus Pacheco 1964: 84 (endemic genus) and nomenclatorial notes on Caribbean F. guttatus (Kiesenwetter) 1851: 294 (Heterocerus); Tropicus Pacheco (Coleoptera: Heteroceridae). Chevrolat 1864c: 407; Gundlach 1891: 52; Proc. Ent. Soc. Washington 86: 946-950. Pacheco 1964: 84 Mascagni, A. 1989. Tropicus alcicornis sp. n. Distribution. Cuba. Endemic dell’America meridionale. Boll. Soc. Ent. Ital. Cuba localities. Ci: Cayamas (t.l.) 120: 187-190. Collections. Washington (type) Pacheco, F. 1964. Sistematica, filogenia y distribucion de los Heteroceridos de America Neoheterocerus Pacheco 1964: 54 (key to spe- (Coleoptera: Heteroceridae). Monografias del cies) Colegio de Post-Graduados: no. l, Chapingo, N. angustatus (Chevrolat) 1864c: 407 México. Pp. 155. (Heterocerus); Gundlach 1891: 52; Pacheco 1964: 62; Spangler 1981 58. FAMILY PSEPHENIDAE Distribution. Cuba, Bahamas, Haiti, USA The water penny beetles (IA-NY-FL) Larvae feed on algae on submerged rocks in Cuba localities. Ca: California. CA: Baraguá. streams, and adults occur on vegetation at stream M: Cárdenas edges, or at lights. Collections. Gundlach no. 662 SUBFAMILY PSEPHENINAE TRIBE TROPICINI Pheneps Darlington 1936: 67 Tropicus Pacheco 1964: 102; see Bameul 1995 P. cubanus Darlington 1936: 68; Spangler 1973: T. bilineatus (Chevrolat) 1864c: 407 (Heterocerus); 357; 1981: 167 Gundlach 1891: 53; Pacheco 1964: 122; Ivie and Distribution. Cuba. Endemic Stribling 1984: 948; Mascagni 1989: 189 Cuba localities. Ca: Rio El Manantiales. Ci: Distribution. Cuba, Costa Rica Soledad (t.l.). Gu: Rio Baracoa, at Tinta; Cuba localities. PR: Soroa Cabezades de Rio Judio. Other records in Collections. Gundlach no.688. Havana. Spangler 1981: 167 Washington Collections. Cambridge (type) T. pusillus (Say) 1823: 200 (Heterocerus); Pacheco 1964: 106; Spangler 1981 REFERENCES Distribution. Cuba, Jamaica, Antigua, Spangler, P.J. 1973. Aquatic Coleoptera collected México, Honduras, Costa Rica, Panamá, USA on the biospeleological expeditions. Resultats (CA-PA-FL), Canada (ON) des expeditions biospéologique Cubano- Cuba localities. Roumaines à Cuba 1: 353-358. Collections. Spangler, P.J. 1981. Supplement to the aquatic and semiaquatic Coleoptera of Cuba. Resultats des expeditions biospéologique Cubano- Roumaines à Cuba 3: 145-171. A Checklist of the Beetles of Cuba 105

59. FAMILY CNEOGLOSSIDAE Chelonarium Fabricius 1801: 101 This family is known only from Central and C. convexum Méquignon 1832: 246 South America. Larvae and habits are unknown. Distribution. Cuba. Endemic Cuba localities. 60. FAMILY PTILODACTYLIDAE Collections. The ptilodactylid beetles C. costatipenne Méquignon 1832: 244 Adults may occur on vegetation and are at- Distribution. Cuba. Endemic tracted to lights. Larvae occur in leaf litter, rotted Cuba localities. logs, and debris at stream edges. Collections. C. maculatum Méquignon 1834: 54 SUBFAMILY PTILODACTYLINAE Distribution. Cuba. Endemic Cuba localities. Ptilodactyla Illiger 1807: 342 Collections. P. annulicornis Chevrolat 1870: 71; Gundlach 1891: C. problematicum Méquignon 1832: 244 249 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas. Collections. Collections. Gundlach no. 619 C. punctatum Fabricius 1801: 101; Jacquelin du P. carbonaria Chevrolat 1870: 72; Gundlach 1891: Val 1856: 68; Chevrolat 1864: 618 250 Distribution. Cuba, Puerto Rico Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Santiago de Cuba Collections. Collections. Gundlach no. 1877 C. sublaeve Méquignon 1833: 46 P. emarginata Chevrolat 1870: 71; Gundlach 1891: Distribution. Cuba. Endemic 248 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. M: Cienaga de Zapata Collections. Gundlach no. 1886 62. FAMILY EULICHADIDAE P. militaris Chevrolat 1870: 72; Gundlach 1891: This family occurs in northern California, 249 China, India, and southeast Asia. The larvae live Distribution. Cuba. Endemic in streams. Cuba localities. Ha: Cojímar Collections. Gundlach no. 1308 63. FAMILY CALLIRHIPIDAE P. ramicornis Chevrolat 1870: 70; Gundlach 1891: The family occurs in North, Central, and South 247 America but seemingly not in the West Indies. Distribution. Cuba. Endemic Cuba localities. M: Cienaga de Zapata SUPERFAMILY ELATEROIDEA Collections. Gundlach no. 1884 64. FAMILY ARTEMATOPODIDAE P. simplex Chevrolat 1870: 70; Gundlach 1891: 248 In the New World, this family occurs from Distribution. Cuba. Endemic North America, through México to South America. Cuba localities. M: Cienaga de Zapata Larvae may feed on moss, lichens, or leaf litter. Collections. Gundlach no. 1885 They are unknown in the Caribbean. P. turquinensis Zayas 1988: 32 Distribution. Cuba. Endemic 65. FAMILY BRACHYPSECTRIDAE Cuba localities. SC: Pico Turquino (t.l.) These beetles live under bark in arid western Collections. Zayas (type) North America, India, and Malaysia. A larva has been found in Miocene amber in the Dominican 61. FAMILY CHELONARIIDAE Republic, so they might yet be found in the West The turtle or chelonariid beetles Indies. Little is known of the biology of adults or lar- vae of this family. The genus Chelonarium is abun- 66. FAMILY CEROPHYTIDAE dant in the Neotropics, Asia, and Australia. They This family is known from Europe, eastern live in moist to dry litter or under bark, and some North America, and Central and South America. may live in the refuse heaps of ants. 106 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Adults come to lights, and can jump. Larvae may Cuba localities. SC: Brazo de Cauto, west of be in rotten wood. This family may occur in Cuba. Santiago Collections. Gundlach no. 1852 67. FAMILY EUCNEMIDAE The false click beetles SUBFAMILY MACRAULACINAE The adults resemble Elateridae, the true click TRIBE MACRAULACINI beetles. Larvae bore in dead wood and may feed on slime molds or fungal decay products (Muona 1993). Dromaeolus Kiessenwetter 1858: 197 D. ischiodontoides Chevrolat 1867: 592; Gundlach SUBFAMILY MELASINAE 1891: 177 (Dromocolus) TRIBE DIRHAGINI Distribution. Cuba. Endemic Cuba localities. Adelothyreus Chevrolat 1867: 593 Collections. Gundlach no. 1838 A. quadrimaculatus Chevrolat 1867: 593; Gundlach 1891: 177 Fornax Laporte 1835: 167 Distribution. Cuba. Endemic F. bicolor Melsheimer 1846: 149 Cuba localities. M: Fermina Fabricius lurida Chevrolat 1867: 591, not Dejean; Collections. Gundlach no. 353 Gundlach 1891: 175 Distribution. Cuba, eastern and central USA Arrhipis Bonvouloir 1871: 71 Cuba localities. Gu: Yateras A. cubanus Fisher 1945: 88 Collections. Gundlach no. 101 Distribution. Cuba. Endemic F. ebenina Fleutieux 1897: 253 Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. A. insularis Fisher 1945: 89 Collections. Distribution. Cuba. Endemic F. poeyi Fisher 1945: 84 Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. Ci: Cayamas (t.l.) A. lanieri (Guérin-Méneville) in Chevrolat 1838: 279 Collections. Washington (type) (Hylochares); Gundlach 1891: 178 F. repulsa Chevrolat 1867: 571; Gundlach 1891: Hypocaelus lasnieri Jacquelin du Val 1856: 65 175 Hylochares lanieri jacquelini Chevrolat 1867: 593 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico, USA (FL) Cuba localities. Ci: Trinidad Mts Cuba localities. ? Cabo de Hacha. ? Guaban Collections. Gundlach no. 1432 Collections. Gundlach no. 545 TRIBE NEMATODINI Microrhagus Dejean 1833 M. albofasciatus (Fisher) 1945: 90 (Dirhagis) Nematodes Berthold 1827: 335 Distribution. Cuba, Hispaniola. N. exiguus Fisher 1945: 92 Cuba localities. Gu: Baracoa. Ho: Felton (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. Ci: Cayamas (t.l.) M. phyrrhopus Chevrolat 1867: 592; Gundlach Collections. Washington (type) 1891: 176 N. rugicollis Chevrolat 1867: 594; Gundlach 1891: Distribution. Cuba. Endemic 179 Cuba localities. M: Cienaga de Zapata Distribution. Cuba, Brazil Collections. Gundlach no. 960 Cuba localities. Gr: Bayamo; Braza del Cauto Collections. Gundlach no. 1420 SUBFAMILY EUCNEMINAE N. simulans Chevrolat 1867: 595; Gundlach 1891: TRIBE DENDROCHARINI 180 Distribution. Cuba. Endemic Dendrocharis Guérin-Méneville 1843: 193 Cuba localities. D. bombycina Guérin-Méneville 1843: 193; Collections. Gundlach no. 1878 Chevrolat 1867: 591; Gundlach 1891: 174 Distribution. Cuba, Colombia A Checklist of the Beetles of Cuba 107

Neomathion Fleutiaux 1830: 45 M. sturmi (Germar) 1844: 188 (Aphanobius); N. leprieuri (Laporte) 1835: 174 (Galba); Gundlach Chevrolat 1867: 609; Gundlach 1891: 196; Leng 1891: 179 (Nematodes) and Mutchler 1914: 428 (as Crigmus) Distribution. Cuba, México to Peru and Elater cribricollis Sturm 1826: 136 French Guiana Distribution. Cuba, Central America Cuba localities. Cuba localities. M: Cárdenas Collections. Gundlach no. 1867 Collections. Gundlach no. 62 M. taeniata Candèze 1865: 30; Chevrolat 1867: 609; REFERENCE Gundlach 1891: 197 Muona, J. 1993. Review of the phylogeny, classifi- Distribution. Cuba. Endemic cation and biology of the family Eucnemidae Cuba localities. M: Cárdenas. PR: Rangel (Coleoptera). Ent. Scand. Suppl. 44: 1-113. Collections. Gundlach no. 501

68. FAMILY THROSCIDAE TRIBE AGRIOTINI Larvae of this family may feed on rotted wood, or fungi or roots. Drapetes has been placed in this Drasterius Eschscholtz 1829: 33 family but is listed here in Elateridae: Lissominae. D. elegans (Fabricius) 1792: 230 (Elater); Chevrolat 1867: 606 (); Gundlach 1891: 193. 69. FAMILY ELATERIDAE Cryptohypnus circumscriptus Germar 1824: 46; The click beetles Jacquelin du Val 1856: 75 These are called “cocuyos” in Cuba. Adults are Distribution. Cuba, Puerto Rico, USA often collected on vegetation or at lights. Larvae Cuba localities. M: Cárdenas occur in many habitats, including soil, litter, and Collections. Gundlach no. 243 rotten wood. They may be predators, phytophagous, or saprophagous. Some may be pests of agriculture Glyphonyx Candèze 1863: 448 (Vàzquez et al. 1992). G. fuscula (Erichson) 1842: 119; Chevrolat 1867: 613; Gundlach 1891: 203 SUBFAMILY THYLACOSTERNINAE Adrastus fusculus Germar; Jacquelin du Val 1856: 75 Thylacosternus Bonvouloir 1871: 76 Distribution. Cuba, Hispaniola T. maestralis Zayas 1988: 38 (as Tylacosternus) Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 380 Cuba localities. SC: Pico Turquino (t.l.) G. gundlachi Candèze 1863: 453; Chevrolat 1867: Collections. Zayas (type) 613; Gundlach 1891: 203 Distribution. Cuba. Endemic SUBFAMILY Cuba localities. Widespread TRIBE ELATERINI Collections. Gundlach no. 300 G. quadraticollis Champion 1896: 536 Neotrichophorus Jacobson 1813: 741 Distribution. Cuba, Antigua, México to Costa N. havaniensis (Candèze) 1863: 309 (Ludius); Rico, USA Chevrolat 1867: 613; Gundlach 1891: 202 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Widespread G. recticollis (Say) 1823: 168; Chevrolat 1867: 614; Collections. Gundlach no. 500 Gundlach 1891: 203 Distribution. Cuba, México, USA (CT-FL) TRIBE MEGAPENTHINI Cuba localities. Widespread Collections. Gundlach no. 785 Megapenthes Kiesenwetter 1858: 229 G. scabriuscula (Chevrolat) 1867: 614 (Silesus); M. opacula Candèze 1865: 29; Chevrolat 1867: 609; Gundlach 1891: 204 Gundlach 1891: 197 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas Collections. Gundlach no. 1847 Collections. Gundlach no. 293 G. superba Zayas 1988: 33 Distribution. Cuba. Endemic 108 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. SC: Gran Piedra (t.l.) 1891: 185 Collections. Zayas (type) A. seminiger Candèze 1859: 71 Distribution. Cuba. Endemic TRIBE Cuba localities. Collections. Gundlach no. 482 (sic) LeConte 1853: 459 Notes. This species seems to be part of a Batesian A. rufescens Candèze 1859: 404; Chevrolat 1867: mimicry complex centered around species of 608; Gundlach 1891: 196 Thonalmus (Lycidae) (Darlington 1938: 689) Distribution. Cuba. Endemic Cuba localities. Widespread Blauta LeConte 1853: 472 Collections. Gundlach no. 185 B. cribraria (Germar) 1844: 307 A. rufiventris Candèze 1859: 401; Chevrolat 1867: B. cauta LeConte 1853: 473 608; Gundlach 1891: 195 Distribution. Cuba, USA (GA-FL) Distribution. Cuba. Endemic Cuba localities. IJ: Nueva Gerona Cuba localities. Collections. Carnegie Collections. Gundlach no. 1846 Dicrepidius Eschscholtz 1829: 31 Anoplischius Candèze 1859: 9 D. ramicornis (Palisot de Beauvois) 1805: 10 A. cubitas Zayas 1988: 36 (Elater); Jacquelin du Val 1856: 69; Chevrolat Distribution. Cuba. Endemic 1867: 600; Gundlach 1891: 186 Cuba localities. Ca: Sierra de Cubitas (t.l.) Distribution. Cuba, Guadeloupe, México, Collections. Zayas (type) Nicaragua, Brazil, Argentina, USA (se, FL) A. depressipennis Candèze 1859: 69; Chevrolat 1867: Cuba localities. Widespread 599; Gundlach 1891: 184 Collections. Gundlach no. 493. Pittsburgh Distribution. Cuba. Endemic Cuba localities. Dipropus Germar 1839 Collections. Gundlach no. 915 D. antennatus (Candèze) 1859: 108 (Ischiodontus); A. dimidiatus Zayas 1988: 36 Chevrolat 1867: 601; Gundlach 1891: 187 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola Cuba localities. Ho: Sierra Cristal (t.l.) Cuba localities. Collections. Zayas (type) Collections. Gundlach no. 1851 A. pyronotus Candèze 1859: 71; Chevrolat 1867: D. simplex (LeConte) 1853: 462 (Ischiodontus) 599; Gundlach 1891: 184 I. oblitus (Candèze) 1859: 101; Chevrolat 1867: Distribution. Cuba. Endemic 601; Gundlach 1891: 186 Cuba localities. Distribution. Cuba, USA (TX) Collections. Gundlach no. 482 Cuba localities. A. ruficeps Candèze 1865: 23; Chevrolat 1867: 600; Collections. Gundlach no. 1850 Gundlach 1891: 185 D. striatus (Candèze) 1859: 117 (Ischiodontus); Distribution. Cuba. Endemic Gundlach 1891: 187 Cuba localities. Distribution. Cuba. Endemic Collections. Gundlach no. 1840 Cuba localities. A. sagranianus (Jacquelin du Val) 1856: 69 Collections. Gundlach no. 193 (Dicrepidius); Gundlach 1891: 185 A. lineipennis Candèze 1859: 72 SUBFAMILY CARDIOPHORINAE A. sangraninus Schenkling 1925: 80 Distribution. Cuba. Endemic Esthesopus Eschscholtz 1829: 32 Cuba localities. Widespread E. hepaticus Erichson 1840: 336; Chevrolat 1867: Collections. Gundlach no. 63 610; Gundlach 1891: 199 A. suturalis Zayas 1988: 35 Distribution. Cuba, México to Panamá, Bra- Distribution. Cuba. Endemic zil, Argentina Cuba localities. SC: Pico Turquino ( t.l.) Cuba localities. M: Zarabanda Collections. Zayas (type) Collections. Gundlach no. 1061 A. venustus (Jacquelin du Val) 1857: 70 E. humilis Candèze 1860: 284 (Dicrepidius); Chevrolat 1867: 599; Gundlach E. murinus Candèze 1860: 284 A Checklist of the Beetles of Cuba 109

Distribution. Cuba, México to Panamá, Co- SUBFAMILY lombia, Brazil TRIBE Cuba localities. Collections. Latreille 1829: 451 E. poedicus Candèze 1860: 270; Chevrolat 1867: (A. modesta Boisduval 1832: 108) 610; Gundlach 1891: 199 Distribution. West Indies; cosmopolitan Distribution. Cuba, Guadeloupe Cuba localities. Cuba localities. Gu: Baracoa Collections. Collections. Gundlach no. 1422 A. subcostata Candolle 1857: 69; Chevrolat 1867: 596; Gundlach 189l: 181 Horistonotus Candèze 1860: 104 Distribution. Cuba, Antigua, Guadeloupe H. asthenicus Candèze 1860: 266; Chevrolat 1867: Cuba localities. M: Cárdenas 610; Gundlach 1891: 198 Collections. Gundlach no. 499 H. a. sallei Fleutiaux 1911: 260 Distribution. Cuba. Endemic Meristhus Candèze 1857: 19 Cuba localities. M. scobinula Candèze 1857: 164 Collections. Gundlach no. 372 M. setarius Chevrolat 1867: 596; Gundlach H. badius Candèze 1881: 84; Chevrolat 1867: 609; 1891: 181 Gundlach 1891: 198 Distribution. Cuba, México, Guatemala, Co- Distribution. Cuba. Endemic lombia, USA (?) Cuba localities. Cuba localities. Gr: Bayamo Collections. Collections. Gundlach no. 1555 H. cruxnigra Chevrolat 1867: 610; Gundlach 1891: 198 TRIBE Distribution. Cuba. Endemic Cuba localities. M: Fermina Aeolus Eschscholtz 1829: 33 Collections. Gundlach no. 1845 A. binotatus Candèze 1859: 287; Chevrolat 1867: 606; Gundlach 1891: 194 SUBFAMILY LISSOMINAE Distribution. Cuba, Puerto Rico TRIBE LISSOMINI Cuba localities. Collections. Gundlach no. 329 Drapetes Redtenbacher 1849: 290 A. discicollis Candèze 1859: 303; Chevrolat 187: D. azurea Jacquelin du Val 1856: 67; Chevrolat 607; Gundlach 1891: 194 1867: 590; Gundlach 1891: 174 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Widespread Collections. Collections. Gundlach no. 198 A. maculatus DeGeer 1774: 163 D. bicolor Laporte 1835: 180; Chevrolat 1867: 589; Elater angulatus Fabricius 1801: 243; Chevrolat Gundlach 1891: 173 1867: 607 (Aeolus); Gundlach 1891: 194 D. cyanipennis Jacquelin du Val 1856: 67 Distribution. Cuba, Hispaniola Distribution. Cuba. Endemic Cuba localities. Cuba localities. Widespread. Collections. Gundlach no. 1421 Collections. Gundlach no. 197 D. nigripennis Jacquelin du Val 1856: 66; Chevrolat Conoderus Eschscholtz 1829: 31 1867: 588; Gundlach 1891: 172 C. bifoveatus (Palisot de Beauvois) 1805: 78 (Elater); Distribution. Cuba, México Chevrolat 1867: 603; Gundlach 1891: 190 Cuba localities. M: Cárdenas lobatus (Say), Jacquelin du Val Collections. Gundlach no. 331 1856: 71 D. tunicata Bonvouloir 1859: 43; Chevrolat 1867: M. lividus Chevrolat 1867: 603 588; Gundlach 1891: 172 Distribution. Cuba, Hispaniola, Bahamas, Distribution. Cuba, Hispaniola Puerto Rico, Guadeloupe Cuba localities. Widespread Cuba localities. Widespread. The larvae are Collections. Gundlach no. 384 a pest of sugarcane (Vàzquez et. al. 1992: 44) Collections. Gundlach no. 275. Pittsburgh 110 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. dorsalis (Say) 1823: 167 (Elater); Chevrolat 1867: Latreille 1834: 155 605 (Aeolus); Gundlach 1891: 192 H. amplicollis (Gyllenhal) 1817: 41; Chevrolat 1867: Distribution. Cuba. Endemic 607; Gundlach 1891: 195 Cuba localities. M: Cárdenas Distribution. Cuba, Puerto Rico, Guadeloupe, Collections. Gundlach no. 1677 Martinique, Dominica, South America, USA C. lividus (DeGeer) 1774: 162 (Elater); Chevrolat (CA-TX-AL-FL) 1867: 603 (Monocrepidius); Gundlach 1891: 189 Cuba localities. Distribution. Cuba, Bahamas, Hispaniola, Collections. Puerto Rico, Guadeloupe, México, USA (TX-IL- NC-FL) TRIBE Cuba localities. Widespread Collections. Gundlach no. 1902. Pittsburgh Ignelater Costa 1975: 105. C. memorabilis (Candèze) 1859: 249 I. brunneus Costa 1980: 158 (Monocrepidius); Chevrolat 1867: 605; Distribution. Cuba. Endemic Gundlach 1891: 19l Cuba localities. SC: N. of Imias (t.l.); Loma Distribution. Cuba, Puerto Rico, Martinique del Gato Cuba localities. Collections. Cambridge Collections. Gundlach no. 798 I. havaniensis (Laporte) 1840: 236 (); C. parallelus Candèze 1859: 246 Jacquelin du Val 1856: 73; Chevrolat 1867: 612; Distribution. Cuba, México Gundlach 1891: 201; Costa 1975: 106; 1980: Cuba localities. 160 Collections. Distribution. Cuba, Bahamas, Hispaniola, C. pinguis (Candèze) 1859: 207 (Monocrepidius); Mexico, Colombia, , USA Chevrolat 1867: 602; Gundlach 1891: 189 (FL) Distribution. Cuba, Puerto Rico Cuba localities. Widespread Cuba localities. Collections. Cambridge. Gundlach no. 38. Collections. Gundlach no. 1839 London. New York. Pittsburgh C. posticus Eschscholtz 1822: 76 I. novoae.Fernandez and Lozada 1998: 58 C. sticturus (Candèze) 1859: 260 (Monocrepidius); Distribution. Cuba. Endemic Gundlach 1891; 192; Peck et al. 1998: 160 Cuba localities.. SC: Loma del Gato (t.l.); Distribution. Cuba, Guadeloupe, México to Playa Juraguá; Sierra Maestra Brazil Collections. Havana (type and paratypes) Cuba localities. Widespread. Ha: in Cueva I. paveli Fernandez and Lozada 1998: 59 de los Murcielagos, larvae feeding in guano Distribution. Cuba. Endemic Collections. Gundlach no. 465 Cuba localities.. Gu: Guantánamo (t.l.) (C. rufidens (Fabricius) 1801: 234) Collections. Ramsden Distribution. Caribbean, USA (FL) I. phosphoreus (Linnaeus) 1758: 404 (Elater); Costa Cuba localities. 1975: 106; 1980: 163 Collections. Pyrophorus longipennis Schwarz 1906: 154 C. sericans Candèze 1891: 71 (Cayamas, t.l.); Leng and Mutchler 1914: 428 Distribution. Cuba. Endemic P. lynchiferus Germar 1841: 32; Chevrolat 1867: Cuba localities. 612; Gundlach 1891: 201 Collections. P. lynchnus Candèze 1863: 26; Chevrolat 1867: C. sericatus (Chevrolat): 1857; 604 (Monocrepidius); 612; Gundlach 1891: 200 not Candèze 1859: 249; Gundlach 1891: Distribution. Cuba; Hispaniola, Puerto Rico, 190 Virgin Islands Distribution. Cuba, Hispaniola. Cuba localities. Gu: Baracoa. Ha: Los Indios. Cuba localities. IJ: Nueva Gerona. M: Cárdenas Collections. Gundlach no. 1899 Collections. Gundlach no. 1430 (P. lychnus). C. sticticus Erichson 1847: 78 Pittsburgh Distribution. Cuba, Hispaniola, Colombia to Peru Pyrophorus Illiger 1809: 141 Cuba localities. P. mellifluus Costa 1972: 211; 1976: 147 Collections. A Checklist of the Beetles of Cuba 111

Distribution. Cuba, Hispaniola, Martinique, Distribution. Cuba, Hispaniola, México to Ar- St. Lucia, Trinidad, México, Guatemala, gentina, USA (southeastern states) Panamá, Colombia Cuba localities. Ci: Cienfuegos. IJ: Nueva Cuba localities. Widespread Gerona Collections. Cambridge. Paris. Pittsburgh. Collections. Gundlach no. 1631. Pittsburgh Washington [P. noctilucus (Linnaeus) 1758: 404 (Elater); SUBFAMILY PROSTERNINAE Jacquelin du Val 1856: 72; Chevrolat 1867: 611; TRIBE ATHOINI Gundlach 1891: 200; Costa 1975: 113. Costa 1976: 148 shows the species does not occur in Hemicrepidius Germar 1839: 212 Cuba] (H. pictipes Chevrolat 1843: 230) Cuba localities. Material under this name Distribution. West Indies, México was reported from: IJ: Nueva Gerona. In caves Cuba localities. in SC, SS, VC, Ha, PR where larvae occur in Collections. guano Collections. Pittsburgh. Gundlach no. 22 Platycrepidius Candèze 1859: 165 P. tuberculifer Escholtz 1829: 32; Costa 1972: 203; P. cyanipennis (Candèze) 1859: 167 (Eudactylus); 1975: 114; 1976: 146 Chevrolat 1867: 602; Gundlach 1891: 188 P. hesperus Candèze 1863: 18; Chevrolat 1867: Distribution. Cuba. Endemic 611; Gundlach 1891: 200 Cuba localities. Gr: Bayamo. M: Cárdenas Distribution. Cuba, Guadeloupe, México, Collections. Gundlach no. 289 Honduras, Brazil, Paraguay, Argentina P. schaumi (Candèze) 1859: 167 (Eudactylus); Cuba localities. IJ: Nueva Gerona. Cuba Chevrolat 1867: 602; Gundlach 1891: 188 records need confirmation. Distribution. Cuba. Endemic Collections. London. Pittsburgh Cuba localities. M: Cárdenas; Guamutas Collections. Gundlach no. 106 TRIBE HEMIRHIPINI Notes. This species seems to be part of a Batesian mimicry complex centered around species of Eschscholtz 1829: 33 Thonalmus (Lycidae) (Darlington 1938: 689) A. patricius (Jacquelin du Val) 1856: 73; Candèze P. trinitatis Zayas 1988: 34 1857: 24 () Distribution. Cuba. Endemic Calais patricius Jacquelin du Val 1857: 33; Cuba localities. SS: Lomas de Trinidad, Al- Chevrolat 1867: 597; Gundlach 1891: 182 tiplano de Buenos Aires (t.l.) Distribution. Cuba, USA (FL) Collections. Zayas (type) Cuba localities. Widespread Collections. Gundlach no. 473. Pittsburgh REFERENCES A. primarius (Candèze) 1865: 18 (Calais); Gundlach Casari, S.A. 1986. Revision of Pherimius 1891: 182 Fleutiaux, 1942, with establishment of Distribution. Cuba. Endemic Saltamartinus, new genus (Coleoptera, Cuba localities. Gu: Yateras Elateridae, Pyrophorinae, Hemirhipini). Papéis Collections. Gundlach no. 1429 Avulsos de Zoología (São Paulo), 39: 379-403. Costa, C. 1972. Genero Pyrophorus. 6. Redescricão Hemirrhipus Latreille 1829: 402 de P. tuberculifer Eschscholtz, P. H. viduus Chevrolat 1867: 598; Gundlach 1891: phosphorescens Castelnau e descricão de 183 quatorze espécies novas (Col. Elateridae). Papéis Distribution. Cuba. Endemic Avulsos de Zoologia (São Paulo) 25: 199-227. Cuba localities. M: Cienaga de Zapata Costa, C. 1975. Systematics and evolution of the Collections. Gundlach no. 1844 tribes Pyrophorini and Heligmini, with descrip- tion of Campyloxeninae, new subfamily (Co- Pherhimius Fleutiaux 1942: 112 leoptera, Elateridae). Arquivos de Zoologia (São P. fascicularis (Fabricius) 1787: 171; Chevrolat Paulo) 26: 49-190. 1867: 598; Gundlach 1891: 183 (Hemirrhipus); Costa, C. 1980. Novas espécies dos géneros Casari 1986: 402 Ignelater e Opselater (Coleoptera, Elateridae, 112 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Pyrophorinae). Papéis Avulsos de Zoologia (São Distribution. Cuba. Endemic Paulo) 33: 157-176. Cuba localities. Widespread Darlington, P.J., Jr. 1938. Experiments on mim- Collections. Gundlach no. 859. New York. icry in Cuba, with suggestions for future study. Pittsburgh. Prague. Washington Trans. R. Ent. Soc. London 87: 681-695. T. aulicus (Jacquelin du Val) 1856: 77 (Calopteron); Fernandez Garcia, I and Lozada Piña, A. 1998. Chevrolat 1870: 76; Gundlach 1891: 254; Leng Descripción de dos nuevas especiaes de Ignelater and Mutchler 1922: 424; Mutchler 1923: 2; (Coleoptera: Elateridae; Pyrophorinae) de Cuba. Wojciechowska and Slipinski 1985: 268 Avicennia 8/9: 57-60. Calopteron distinguendenum Jacquelin du Val Vàzquez, L.L., E. Valdés, and J.C. Amor. 1992. 1856: 80; Chevrolat 1870: 77 New manifestations of pests on economically Calopteron elegantulus Jacquelin du Val 1856: important plants during the period from 1970 78; Chevrolat 1870: 76; Gundlach 1891: 255 to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo Calopteron nigritarse Chevrolat 1870: 77; Silvestri 49: 41-52. Gundlach 1891: 255 Distribution. Cuba. Endemic 70. FAMILY PLASTOCERIDAE Cuba localities. Widespsread This family occurs in Asia minor and south- Collections. Gundlach no. 513 (C. aulicus); east Asia. no. 926 (C. elegantulum); no. 277 (C. nigritarse); no. 922 (C. distinguendum). New 71. FAMILY DRILIDAE York. Philadelphia. Pittsburgh. Prague. Wash- This family occurs in southern Eurasia, Asia ington Minor, India, and Africa. The larvae are predators Notes. Darlington (1938) conducted experiments on snails. on the mimicry complex centered on T. aulicus and T. suavis at Soledad, the Atkins Biological 72. FAMILY OMALISIDAE Station, near Cienfuegos This family occurs in southern Europe. T. bicolor (Linnaeus) 1763: 395 (Cantharis); Leng and Mutchler 1922: 419 73. FAMILY LYCIDAE Calopteron bicolor (Fabricius); Jacquelin du Val The net-winged beetles 1856: 77; Chevrolat 1870: 73 Adults are short lived and feed on pollen and Distribution. Cuba, Jamaica, Hispaniola, nectar. They have bright warning coloration and Guadeloupe are the base of mimicry complexes. Larvae occur Cuba localities. under bark and in rotted wood and probably feed Collections. on slime molds or fungal products. Leng and T. mariani Wojciechowska and Slipinski 1985: 266 Mutchler (1922) reviewed the fauna of the West T. orientalis Zayas 1988: 46 ( type locality not Indies. given; Zayas, type); Ivie 1991: 400 Distribution. Cuba. Endemic SUBFAMILY PLATERODINAE Cuba localities. Ho: Piloto, Moa, Breña; Si- erra Cristal. SC: Gran Piedra (t.l.) Plateros Bourgeois 1879: XIX Collections. Prague (type). Zayas P. bilineatithorax Pic 1925: 18 T. suavis (Jacquelin du Val) 1856: 80 (Calopteron); Distribution. Cuba. Endemic Chevrolat 1870: 75; Gundlach 1891: 254; Leng Cuba localities. and Mutchler 1922: 423; Mutchler 1923: 2; Collections. Wojciechowska and Slipinski 1985: 265 Distribution. Cuba. Endemic SUBFAMILY LYCINAE Cuba localities. Widespread, most abundant TRIBE THONALMINI in spring Collections. Gundlach no. 858. New York. Thonalmus Bourgeois 1883: 376 Philadelphia. Washington T. amabilis (Jacquelin du Val) 1856: 82 (Calopteron); Chevrolat 1870: 75; Gundlach TRIBE CALOPTERINI 1891: 254; Leng and Mutchler 1922: 423; Mutchler 1923: 2; Wojciechowska and Slipinski Calopteron Guérin-Méneville 1830: 70 1985: 269 A Checklist of the Beetles of Cuba 113

C. albicolle Chevrolat 1858: 209; 1870: 77; 75. FAMILY PHENGODIDAE Gundlach 1891: 256 The family occurs in North, Central, and South Distribution. Cuba. Endemic America and in the West Indies only in Hispaniola Cuba localities. SS: Trinidad Mts (Acladocera Wittmer). It should be expected in Collections. Gundlach no. 1478 Cuba. The genus Pseudophengodes of South C. semiflavum Chevrolat 1870: 78; Gundlach 1891: America has light organs on the male . 256; Leng and Mutchler 1922: 329 Males often come to lights, and females and larvae Distribution. Cuba. Endemic are predators on millipeds and other litter Cuba localities. PR: Rangel arthropods. Collections. Gundlach no. 1260 76 FAMILY LAMPYRIDAE Mesopteron Bourgeois 1905: 123 The beetles M. pecticorne (Chevrolat) 1870: 74 (Calopteron) Firefly adults often use light organs to produce Distribution. Cuba, Guadeloupe mating signals. Larvae live in litter and soil and Cuba localities. are predators on various invertebrates. Collections. SUBFAMILY LAMPYRINAE Metapteron Bourgeois 1905: 126 TRIBE LAMPROCERINI M. testaceipenne Pic 1922: 26 Distribution. Cuba. Endemic Alecton Laporte 1833: 135 Cuba localities. A. discoidale Laporte 1833: 135; Jacquelin du Val Collections. 1856: 83; Gundlach 1891: 253; Leng and Mutchler 1922: 433; Mutchler 1923: 3 SUBFAMILY LEPTOLYCINAE A. dorsalis Lacodaire 1857: 320 TRIBE LEPTOLYCINI Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. Ha: Santiago Leptolycus Leng and Mutchler 1922: 430 de las Vegas. M: Cárdenas; Delicias L. parda Zayas 1988: 47 Collections. Gundlach no. 6. New York Distribution. Cuba. Endemic A. flavum Leng and Mutchler 1922: 434; Mutchler Cuba localities. SC: Loma del Gato (t.l.) 1923: 3 Collections. Zayas (type) Distribution. Cuba. Endemic Cuba localities. “Cuba” (t.l.). PR: TacoTaco REFERENCES Collections. Philadelphia (type). New York Darlington, P.J., Jr. 1938. Experiments on mim- A. improvisus Olivier 1909: 221; Leng and Mutchler icry in Cuba, with suggestions for future study. 1922: 433 Trans. R. Ent. Soc. London 87: 681-696. Distribution. Cuba. Endemic Ivie, M.A. 1991. Taxonomic notes on a little known Cuba localities. Ha: Havana (t.l.). PR: publication: Zayas, 1988, Entomofauna Viñales Cubana. Order Coleoptera. Coleop. Bull. 45: 399- Collections. New York (type) 401. A. x-nigra Zayas 1988: 48 Leng, C.W., and A.J. Mutchler. 1922. The Distribution. Cuba. Endemic Lycidae, Lampyridae, and Cantharidae of the Cuba localities. PR: Guanahacabibes (t.l.) West Indies. Bull. Amer. Mus. Nat. Hist. 46: Collections. Zayas (type) 413-499. Wojciechowska, A. and S.A. Slipinski. 1985. TRIBE CRATOMORPHINI Notes on the Thonalmus Bourgeois (Coleoptera, Lycidae) of Cuba with description of a new spe- Aspisoma Laporte 1833: 127 cies. Polskie Pismo Ent. 55: 265-270. A. ignita Linnaeus 1767: 645 Distribution. Cuba, Hispaniola, widespread 74. FAMILY TELEGEUSIDAE Lesser Antilles, México to Venezuela, Trinidad The family is known from the southwestern Cuba localities. US to Panamá, and northwestern South America. Collections. It might possibly occur in Cuba. 114 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE PHOTINI Distribution. Cuba, Puerto Rico Cuba localities. Gr: Naguas. Gu: Callopisma Motschulsky 1853: 43 Guantánamo, Rio Seco. M: Cienaga de Zapata. Notes. Some species in this genus seem to be part PR: Taco Taco of a Batesian mimicry complex centered around Collections. New York species of Thonalmus (Lycidae) (Darlington C. monticola Mutchler 1923: 8 1938: 689) Distribution. Cuba. Endemic C. adjuncta E. Olivier 1899: 90 (Lychnuris); Leng Cuba localities. SC: Pico Turquino (t.l.) and Mutchler 1922: 443; Mutchler 1923: 9 Collections. New York (type) Distribution. Cuba. Endemic C. ramsdeni Leng and Mutchler 1922: 442; Cuba localities. Ci: Cayamas. SC: Sierra Mutchler 1923: 10 Maestra, Palma Mocha Distribution. Cuba. Endemic Collections. Philadelphia. Washington Cuba localities. Gu: Belona (t.l.); C. bellicosa (E. Olivier) 1899: 91 (Lychnuris); Leng Guantánamo. Gr: Naguas and Mtchler 1922: 444; Mutchler 1923: 10 Collections. New York (type). Philadelphia Distribution. Cuba. Endemic C. rufa Olivier 1790: 28; Leng and Mutchler 1922: Cuba localities. PR: Viñales; San Vincente 441 Collections. New York. Philadelphia Lychnuris dimidiatipennis Jacquelin du Val C. demissa (E. Olivier) 1899: 90 (Lychnuris); Leng 1856: 84; Gundlach 1891: 251 and Mutchler 1922: 444 Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba. Endemic St. Thomas Cuba localities. IJ: Los Indios Cuba localities. M: Cárdenas; Matanzas. Gu: Collections. Philadelphia. Pittsburgh. Wash- Baracoa; Guantánamo. Ha: Havana ington C. flavilabris (E. Olivier) 1899: 20 (Lychnuris); Diphotus Barber 1941: 12 Leng and Mutchler 1922: 444 D. glaucus Olivier 1890: 13; Mutchler 1923; 1 Distribution. Cuba. Endemic (Photinus) Cuba localities. Distribution. Cuba, Jamaica?, Hispaniola Collections. Philadelphia Cuba localities. SC: Sierra Maestra C. fuscotermina Mutchler 1923: 9 Collections. Distribution. Cuba. Endemic Cuba localities. PR: Taco Taco (t.l.) Heterophotinus E. Olivier 1894: 24 Collections. New York (type) H. limbipennis (Jacquelin du Val) 1856: 86 C. janthinipennis (Jacquelin du Val) 1857: 83 (Photinus); Gundlach 1891: 252; Leng and (Lychnuris); Gundlach 1891: 251; Leng and Mutchler 1922: 476; Mutchler 1923: 6 Mutchler 1922: 445; Mutchler 1923: 10 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Ci: Cayamas. Gr: Bayamo. Collections. Gundlach no. 10. New York. Ha: Havana; Santiago de las Vegas Washington Collections. Gundlach no. 1518. New York. Washington Lecontea E. Olivier 1899: 37 C. maestra Mutchler 1923: 8 L. gamma (Jacquelin du Val) 1856: 85 (Photinus); Distribution. Cuba. Endemic Gundlach 1891: 251; Leng and Mutchler 1922: Cuba localities. SC: Pico Turquino (t.l.); Si- 454; Mutchler 1923: 13 (Pyractomena) erra Maestra Distribution. Cuba, Jamaica Collections. New York (type) Cuba localities. Widespread C. militaris (E. Olivier) 1899: 91 (Lychnuris); Leng Collections. Gundlach no. 540. New York. and Mutchler 1922: 445 Philadelphia. PittsburghLucidota Laporte 1833: Distribution. Cuba. Endemic 136 Cuba localities. Gu: Guantánamo L. bruneri Mutchler 1923: 6 Collections. New York Distribution. Cuba. Endemic C. miniatocollis (Chevrolat) 1858: 209 (Lychnuris) Cuba localities. SC: Pico Turquino, summit Lucidota miniatocollis, Leng and Mutchler 19222: (t.l.) 437; Mutchler 1923: 5 Collections. New York (type) A Checklist of the Beetles of Cuba 115

L. chevrolati Mutchler 1923: 7 Collections. Philadelphia. Washington Distribution. Cuba. Endemic P. nefarius E. Olivier 1912: 25; Leng and Mutchler Cuba localities. SC: Sierra Maestra, 2000- 1922: 471; Mutchler 1923: 5 3000 feet (t.l.) P. blandus Jacquelin du Val 1857: 87; Gundlach Collections. New York 1891: 252 L. discoidalis Laporte 1833: 144 Distribution. Cuba. Endemic Distribution. Cuba, Brazil Cuba localities. Widespread Cuba localities. Collections. Gundlach no. 223. New York. Collections. Philadelphia. Washington L. fulvotinctus Mutchler 1923: 4 P. pygmaeus E. Olivier 1899: 92; 1912: 29 L. fulvotinctus flavicollis Mutchler 1923: 5 (Gu: (Pygolampis); Mutchler 1923: 2 Baracoa (t.l.), type in Washington) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ca: Camaguëy. Ci: Cayamas; Cuba localities. Ci: Cayamas (t.l.) Soledad. Gr: Naguas. SC: Pico Turquino; Si- Collections. Washington (type) erra Maestra L. subdubitata Mutchler 1923: 6 Collections. New York. Washington Distribution. Cuba. Endemic P. singularis Leng and Mutchler 1922: 477 Cuba localities. SC: Pico Turquino, summit Distribution. Cuba. Endemic (t.l.) Cuba localities. M: Maria de Loreto (t.l.) Collections. New York (type) Collections. New York (type) P. sublateralis Mutchler 1923: 6 Photinus Laporte 1833: 140 Distribution. Cuba. Endemic P. albicollis (Chevrolat) 1858: 209 (Calopteron); Cuba localities. SC: Pico Turquino (t.l.) Gundlach 1891: 2565: Leng and Mutchler 1922: Collections. New York (type) 480 P. vitiosus Gemminger 1870: 120; Leng and Distribution. Cuba. Endemic Mutchler 1922: 477 Cuba localities. SS: Trinidad Mts Distribution. Cuba, Jamaica, Hispaniola Collections. Philadelphia Cuba localities. P. apoplecticus E. Olivier 1899: 92; E. Olivier 1912: Collections. 28 (Pygolampis) Distribution. Cuba. Endemic SUBFAMILY PHOTURINAE Cuba localities. Ci: Cayamas. IJ: Los Indios; Nueva Gerona Photuris LeConte 1851: 337 Collections. Philadelphia. Pittsburgh. Wash- P. brunnipennis Jacquelin du Val 1856: 89; ington Gundlach 1891: 253; Leng and Mutchler 1922: P. blandus Motschoulsky 1854: 25; Leng and 482; Mutchler 1923: 7 Mutchler 1922: 469 (uncertainty about the Distribution. Cuba. Endemic name) Cuba localities. Widespread. Ci: Cayamas. Distribution. Cuba, Hispaniola Gu: Baracoa; Guantánamo. Ha: Havana; PR: Cuba localities. Cabañas; Taco Taco. SC: Santiago Collections. Collections. Gundlach no. 82. New York. P. cubanus Leng and Mutchler 1922: 463 Washington Distribution. Cuba. Endemic Cuba localities. IJ: Nueva Gerona (t.l.) REFERENCES Collections. Philadelphia (type). Washington Darlington, P.J., Jr. 1938. Experiments on mim- P. magnus Mutchler 1923: 3 icry in Cuba, with suggestions for future study. P. m. turquino Mutchler 1923: 4 Trans. R. Ent. Soc. London 87: 681-695. Distribution. Cuba. Endemic Mutchler, A.J. 1923. Notes on West Indian Cuba localities. SC: Sierra Maestra (t.l.) Lycidae and Lampyridae (Coleoptera), with Collections. New York (type) descriptions of new forms. Amer. Mus. P. nanus (E. Olivier) 1912: 29 (Pygolampris); Leng Novitates 60: 1-13. and Mutchler 1922: 462 Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas 116 Arthropods of Florida and Neighboring Land Areas, Vol. 18

77. FAMILY OMETHIDAE T. camagueyanus Zayas 1988: 53 The family occurs in southeast Asia, Japan, Distribution. Cuba. Endemic and North America. Their habits are unknown. Cuba localities. Ca: Camagüey (t.l.) Collections. Zayas (type) 78. FAMILY CANTHARIDAE T. cubanus Leng and Mutchler 1922: 489 The soldier beetles Distribution. Cuba. Endemic Adults are usually are found on flowers, where Cuba localities. Ci: Cayamas (t.l.) they feed on nectar or pollen, but some may be Collections. New York. Washington (type) predators. Larvae occur in soil or litter and are T. darlington Mutchler 1937: 2 usually predators. Distribution. Cuba. Endemic Cuba localities. Ci: SS: Buenos Aires (t.l.) SUBFAMILY CANTHERINAE Collections. Cambridge (type). New York T. (Thinalmus) gracilis Wittmer 1991: 117 Cantharis Linnaeus 1758: 400 Distribution. Cuba. Endemic C. cubanus Zayas 1988: 51 Cuba localities. Ca: Camagüey. SS: Distribution. Cuba. Endemic Mayajigua (t.l.) Cuba localities. SC: Guamá, Cobre (t.l.) Collections. Basel (type). Washington Collections. Zayas (type) T. insularis Wittmer 1991: 123 Distribution. Cuba. Endemic SUBFAMILY SILINAE Cuba localities. Ho: Felton. (t.l.) TRIBE SILINI Collections. Washington (type) T. maestralis Zayas 1988: 53 Silis Charpentier 1825: 194 Distribution. Cuba. Endemic S. apicalis Zayas 1988: 52 Cuba localities. SC: Sierra Maestra (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Ca: Sierra de Cubitas (t.l.) T. rubidus Mutchler 1937: 4 Collections. Zayas (type) Distribution. Cuba. Endemic S. impressa Pic 1916: 8 (Hapalocrosilis); Leng and Cuba localities. Ci: SS: Buenos Aires (t.l.) Mutchler 1922: 492: Mutchler 1923: 9 Collections. Cambridge (type). New York Distribution. Cuba. Endemic T. rutilis Mutchler 1937: 2 Cuba localities. Gu: Guantánamo. PR: Distribution. Cuba. Endemic Viñales. SC: Pico Turquino Cuba localities. Ci: Soledad (t.l.) Collections. New York. Washington Collections. Cambridge (type) S. marginella Jacquelin du Val 1856: 90; Gundlach 1891: 257; Leng and Mutchler 1922: 492; SUBFAMILY CHAULIOGNATHINAE Mutchler 1923: 9 TRIBE ICHTHYURINI Distribution. Cuba. Endemic Cuba localities. Ca: Camaguëy. Ci: Cayamas; Belotus Graham 1881: 99 Soledad. PR: Rangel; Viñales B. antillarum Leng and Mutchler 1922: 486 Collections. Gundlach no. 1262. New York. Distribution. Cuba. Endemic Philadelphia. Washington Cuba localities. Ci: Cayamas (t.l.) Collections. New York. Philadelphia. Wash- TRIBE TYTTHONYXINI ington (type) B. balloui Mutchler 1923: 8 Tytthonyx LeConte 1851: 347 Distribution. Cuba. Endemic Notes. Some species in this genus seem to be part Cuba localities. SC: Sierra Maestra, Palma of a Batesian mimicry complex centered around Mocha (t.l.) species of Thonalmus (Lycidae) (Darlington Collections. 1938: 689) B. cacumenum Mutchler 1923: 7 T. bicolor Zayas 1988: 54 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. Ha: La Havana. LV: Las Collections. New York (type) Villas. Type locality not given B. flavithorax Zayas 1988: 49 Collections. Zayas (type) Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 117

Cuba localities. SC: Pico Turquino (t.l.) Adults and larvae feed on a variety of plant Collections. Zayas (type) and animal materials, and often on stored prod- B. minutus Leng and Mutchler 1922: 486 ucts. Foods range from pollen and nectar, to dry Distribution. Cuba. Endemic animal carcasses, to household fabrics, and stored Cuba localities. Ci: Cayamas (t.l.) grains and cereals. An alternative higher classifi- Collections. Washington (type) cation is that of Mroczkowski (1968). B. turquinensis Zayas 1988: 50 Distribution. Cuba. Endemic TRIBE DERMESTINI Cuba localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) Dermestes Linnaeus 1758: 354 D. ater DeGeer 1774: 223. The black larder beetle REFERENCES D. cadaverinus Fabricius, Jacquelin du Val Darlington, P.J., Jr. 1938. Experiments on mim- 1856: 101; Chevrolat 1864: 616; Gundlach 1891: icry in Cuba, with suggestions for future study. 85 Trans. R. Ent. Soc. London 87: 681-695. Distribution. Cuba, West Indies, México to Wittmer, W. 1991. Zur Kenntnis der Gattung Argentina, USA, cosmopolitan; widespread Tytthonyx Le Conte, 1851 Subgenus stored products pest in dried fish, cheese, etc. Thinalmus Gorham 1881. Coleop. Cantharidae, Cuba localities. SC: Santiago subfamily Silinae, tribus Tytthonyxini. Mitt. Collections. Schweiz Ent. Ges. 64: 115-126. D. carnivorus Fabricius 1775: 55; Jacquelin du Val 1856: 100; Chevrolat 1864: 615; Gundlach 1891: SERIES BOSTRICHIFORMIA 84 79. FAMILY JACOBSONIIDAE Distribution. Cuba, West Indies, México to This family is represented in the West Indies South America, USA; cosmopolitan by Derolathrus sharpi Grouvelle from Guadeloupe, Cuba localities. Widespread; stored products and this or another (probably introduced) species pest occurs in south Florida as well as one in northern Collections. Florida. The family is probably present in Cuba. (D. frischi Kugelann 1792: 478) The adults are so small they are often overlooked. Distribution. Cuba?; a pest in dried fish Adults and larvae occur in litter or under bark. Cuba localities. Collections. SUPERFAMILY DERODONTOIDEA (D. lardarius Linneaus 1758: 354. The larder beetle) 80. FAMILY Distribution. Cuba?; cosmopolitan; a pest of This family is restricted to the temperate parts dried meat, fish, and cheese of the northern and southern hemispheres. They Cuba localities. feed on fermenting areas under bark or on fungi. Collections. D. maculatus DeGeer 1774: 223. The hide beetle SUPERFAMILY BOSTRICHOIDEA D. vulpinus Fabricius, Chevrolat 1864: 615; 81. FAMILY NOSODENDRIDAE Gundlach 1891: 84 The wounded-tree or nosodendrid beetles Distribution. Cuba, West Indies, México to These beetles occur in forested areas, and in- Argentina, USA; cosmopolitan. Pest of stored habit slime fluxes and tree wounds, and probably skins and hides feed on fermentation products. Cuba localities. SS: Trinidad Collections. Nosodendron Latreille 1804: 146 D. marmoratus Say 1823: 197; Jacquelin du Val N. punctatostriatum Chevrolat 1864: 618; 1856: 101; Chevrolat 1864: 616; Gundlach Gundlach 1891: 87 1891: 85 Distribution. Cuba. Endemic Distribution. Cuba, México, USA Cuba localities. Widespread Cuba localities. Widespread Collections. Collections. (D. unicolor Lepesme 1850: 44) 82. FAMILY DERMESTIDAE Distribution. Antilles, Baja California The skin and larder beetles Cuba localities. Collections. 118 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE THORICTINI Collections. (T. bicinctum Reitter 1880: 38.) Thylodrias Motschulsky 1839: 76 Distribution. West Indies (T. contractus Motschulsky 1839: 76). The odd Cuba localities. beetle Collections. Distribution. Cosmopolitan; in homes and T. insulare Chevrolat 1864: 617; Gundlach 1891: museums, on animal skins; the females are 86 wingless Distribution. Cuba, Puerto Rico Cuba localities. Cuba localities. Ha: Havana. M: Cárdenas Collections. Collections. T. subfasciata Chevrolat 1864: 617; Gundlach 1891: TRIBE ATTAGENINI 87 Distribution. Cuba. Endemic Attagenus Latreille 1802: 121 Cuba localities. Widespread A. fasciatus (Thunberg) 1795: 105. The wardrobe Collections. beetle Distribution. Cuba; nearly cosmopolitan; a REFERENCES stored products pest? Beal, R. S., Jr. 1998. Taxonomy and biology of Cuba localities. Nearctic species of (Coleoptera: Collections. Dermestidae). Trans. Amer. Ent. Soc. 124: 271- A. unicolor (Brahm) 1791: 144. The black carpet 332. beetle Mroczkowski, Maciej. 1968. Distribution of the Atagenus Fabricius, Chevrolat 1864: Dermestidae (Coleoptera) of the world with a 616; Gundlach 1891: 86 catalogue of all known species. Annales Distribution. Cuba, México, Puerto Rico, Zoologicae (Polska Akademia Nauk) 26(3): 15- USA, cosmopolitan; a stored products pest? 191. Cuba localities. Collections. 83. FAMILY BOSTRICHIDAE The horned powder-post or bostrichid beetles TRIBE ANTHRENINI Adults and larvae of this family bore into wood or wood products and a few are pests of stored Anthrenus Fabricius 1775: 61 grains. (A. verbasci Linnaeus 1767: 568). The varied car- pet beetle SUBFAMILY POLYCAONINAE Distribution. Cosmopolitan; in various dried stored products; see Beal 1998: 310 Heterarthron Guérin 1844: 186. The genus name Cuba localities. Melalgus Dejean may apply to these species Collections. H. femoralis Fabricius 1792: 361; Fisher 1950: 6 Distribution. Cuba, St. Croix, USA (TX) Motschulsky 1858: 48 Cuba localities. O. fulvipes (Guérin-Méneville) 1838: 138 Collections. (Globulicornis); Chevrolat 1864: 617 H. gonagrum (Fabricius) 1798: 156; Fisher 1950: (); Gundlach 1891: 87 6 Distribution. Cuba, Puerto Rico, Guadeloupe, Distribution. Cuba, widespread in Greater Brazil, USA (FL), cosmotropical; introduced to and Lesser Antilles Old World Cuba localities. Cuba localities. Collections. Collections. H. plicatum LeConte 1874: 65; Fisher 1950: 7 Distribution. Cuba, Jamaica, México, USA Trogoderma Latreille 1829: 511 (TX) (T. anthrenoides (Sharp) 1902: 649.) Cuba localities. Distribution. West Indies, México to Collections. Panamá, Colombia, USA (TX, Hawaii) Cuba localities. A Checklist of the Beetles of Cuba 119

SUBFAMILY Collections. TRIBE APATINI Tetrapriocera Horn 1878: 544 Apate Fabricius 1775: 54 T. longicornis (Olivier) 1795: 15; Fisher 1950: 102 A. monacha Fabricius 1775: 54 Distribution. Cuba, Isla de Pinos, A. carmelita Fabricius 1801: 379; Jacquelin du Hispaniola, West Indies, México to Brazil, USA Val 1857: 239; Gundlach 1891: 99 (FL); boring in wood of many trees, roots and Distribution. Cuba (introduced), Puerto Rico, wooden boxes native to Africa Cuba localities. Cuba localities. Collections. Collections. Xylomeira Lesne 1801: 502 TRIBE X. tridens (Fabricius) 1792: 362 (Apate); Fisher 1950: 120 Amphicerus LeConte 1861: 208 Tetrapriocera tridens (Fabricius) A. bicaudatus (Say) 1824: 320; Fisher 1950: 67 Distribution. Cuba, widespread West Indies, Distribution. Cuba, USA (IN-KS-FL) México, USA (TX, FL); boring in several tree Cuba localities. species Collections. Cuba localities. IJ: Los Indios A. cornutus Pallas 1772: 8; Fisher 1950: 69 Collections. Carnegie Distribution. Cuba, México to South America; boring in many tree species SUBFAMILY DINODERINAE Cuba localities. IJ: unspecified (Bruner, 1951: 83) Dinoderus Stephens 1830: 352 Collections. (D. bifoveolatus (Wollaston) 1858: 409; Fisher 1950: (A. hamatus Fabricius 1787: 33; Fisher 1950: 68) 25) Distribution. USA Distribution. Tropicopolitan; boring into Cuba localities. many wood products Collections. Cuba localities. Collections. Heterobostrychus Lesne 1899: 443 D. minutus (Fabricius) 1775: 54. (Apate); Fisher H. aequalis (Waterhouse) 1884: 215; Fisher 1950: 1950: 30. The bamboo powerpost beetle 85 Distribution. Cuba (introduced), widespread Distribution. Cuba (introduced), probably West Indies, Guatemala to Argentina, USA tropicopolitan; USA (FL); a pest of wood struc- (FL, CA), native to Old World tropics; boring tures and products in bamboo and other woods, plus sugarcane, Cuba localities. Ha: Marianao (only West rattan, packing cases and stored products Indian record, Bruner 1951: 83), not estab- Cuba localities. Widespread (Bruner 1951: lished? 83) Collections. Collections.

Micrapate Casey 1898: 66 Rhizopertha Stephens 1830: 354 M. fusca Lesne 1801: 502 R. dominica Fabricius 1792: 359; Fisher 1950: 34 Distribution. Cuba, Panamá Distribution. Cuba, Puerto Rico, México to Cuba localities. Brazil, USA?; cosmopolitan in stored products Collections. Cuba localities. Collections. TRIBE XYLOPERTHINI SUBFAMILY LYCTINAE Dendrobiella Casey 1898: 67 TRIBE LYCTINI D. sericea Mulsant and Wachanru 1852: 14 Distribution. Cuba, Jamaica, Hispaniola, Lyctus Fabricius 1792: 507 Guatemala L. brunneus Stephens 1830: 117. The brown Cuba localities. powderpost beetle 120 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, West Indies, USA, cos- 84. FAMILY ANOBIIDAE mopolitan; in dry seeds and spices The death-watch and spider beetles Cuba localities. Collections. SUBFAMILY PTININAE (L. caribeanus Lesne 1831: 96) Reviewed by T. Keith Philips, July 2001 Distribution. West Indies, Central America These beetles live in both moist and dry areas Cuba localities. and feed on a variety of materials of plant and ani- Collections. mal origin. Some are stored products pests. L. villosus Lesne 1911: 537; Gerberg 1957: 29 Distribution. Cuba, Puerto Rico, Mexico, TRIBE GIBBIINI Central America, Surinam, USA (AZ, FL) Cuba localities. Ha: Havana (in cedar furni- Gibbium Scopoli 1775: 505 ture) G. aequinoctiale Boieldieu 1854: LXXXIV Collections. Washington G. chevrolati Boieldieu 1854: LXXXV; Gundlach 1891: 116 Minthea Pascoe 1866: 97 Distribution. Cuba (introduced), USA (wide- M. obstita (Wollaston) 1867: 112 (Lyctus); Gerberg spread), New and Old World; cosmopolitan 1957: 31 Cuba localities. Old Cuban records of cos- Distribution. Cuba (introduced), Central mopolitan G. psylloides Czempinski 1778 are America, USA; native to Africa probably of this species Cuba localities. Ha: Santiago de las Vegas Collections. Gundlach no. 815 Collections. Washington G. cubense Zayas 1988: 68 M. rugicollis Walker 1856: 206 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola, USA (FL?, Cuba localities. SC: Pico Cuba (t.l.) intercept from Guantánamo, Cuba), cosmopoli- Collections. Zayas (type) tan Cuba localities. Gu: Guantánamo TRIBE PTININI Collections. Washington Fabrasia Martinez and Viana 1964: 8 TRIBE TROGOXYLONINI Cubaptinus Zayas 1988: 69 F. cubana (Zayas) 1988: 69 (Cubaptinus); Philips Trogoxylon LeConte 1862: 209 1997: 52 T. aequale (Wollaston) 1867: 111 (Lyctus); Gerberg Distribution. Cuba. Endemic 1957: 37 Cuba localities. None cited T. curtulum Casey 1891: 13 Collections. Zayas (type) Distribution. Puerto Rico, Central America Cuba localities. Ha: Havana (in Bucida Pitnus Gorham 1883: 197 buceras wood) P. huesanus Fisher 1919: 295; Belles 1992: 178 Collections. Distribution. Cuba, Bahamas, USA (FL). Feeds on sea grape and sea lavender REFERENCES (Icthyomethia piscipula and Tournefortia Bruner, S.C. 1951. Algunas adiciones a la lista de gnaphalodes); probably a leaf miner in the host coleópteros de Cuba. Mem. Soc. Cubana Hist. plants (Philips et al. 1998) Nat. Felipe Poey 20: 81-86. Cuba localities. Gr: Cayo Manzanillo Fisher, W.S. 1950. A revision of the North Ameri- Collections. Washington can species of beetles belonging to the family Bostrichidae. US Dept. Agric. Misc. Publ. 698. Ptinus Linnaeus 1766: 537 157 pp. P. niveicollis Boieldieu 1856: 661 Gerberg, E. J. 1957. A revision of the New World Distribution. Cuba, Hispaniola species of Powder-post beetles belonging to the Cuba localities. family Lyctidae. US Dept. Agric. Tech. Bull Collections. 1157. 65 pp., 14 plates. (P. tectus Boieldieu 1856: 652) P. ocellus Brown 1929: 109 A Checklist of the Beetles of Cuba 121

Distribution. West Indies (introduced) New Xyletinus Latreille 1809: 376 and Old World, cosmopolitan (X. castaneus Laporte 1840: 295) Cuba localities. Distribution. West Indies Collections. Cuba localities. Collections. SUBFAMILY ANOBIINAE The majority of these beetles feed in dead wood, TRIBE LASIODERMINI but some attack seed pods and stored products. The classification used here is after White (1982). Lasioderma Stephens 1835: 417 L. serricorne (Fabricius) 1792: 241. The cigarette TRIBE GASTRALLINI beetle Distribution. Cuba, Puerto Rico, Grenada, Falsogastrallus Pic 1914: 10 México to Paraguay, USA; a pest of tobacco and F. librinocens (Fisher) 1938: 44 (Neogastrallus) a wide variety of dry organic materials Distribution. Cuba, USA (FL). Damages Cuba localities. books Collections. Cuba localities. Ha: San Antonio de Las Ve- gas (Bruner 1951: 84) Megorama Fall 1905: 206 Collections. M. cristalensis Zayas 1988: 75 Distribution. Cuba. Endemic TRIBE NICOBIINI Cuba localities. Ho: Sierra Cristal (t.l.) Collections. Zayas (type) LeConte 1861: 204 T. grisacea Zayas 1988: 71 SUBFAMILY Distribution. Cuba. Endemic TRIBE CALYMMADERINI Cuba localities. Ca: Playa Santa Lucía (t.l.) Collections. Zayas (type) Calymmaderus Solier 1849: 472 T. juragua Zayas 1988: 72 C. bibliothecarum (Poey) 1851: 228 (); Distribution. Cuba. Endemic Gundlach 1891: 115 (Dorcatoma) Cuba localities. SC: Juraguá, near Santiago Distribution. Cuba, Puerto Rico, Guadeloupe (t.l.) Cuba localities. Ha: Havana; described from Collections. Zayas (type) specimens infesting books in Poey’s library T. orientalis Zayas 1988: 70 Collections. Gundlach no. 79 Distribution. Cuba. Endemic C. testaceipes Pic 1905: 92 Cuba localities. Ca: Florida Blanca (t.l?) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. T. teresae Zayas 1988: 73 Collections. Distribution. Cuba. Endemic C. whitei Zayas 1988: 76 Cuba localities. Gr: Cabo Cruz (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. SS: Sierra Escambray, San Blas (t.l.) TRIBE STEGOBIINI Collections. Zayas (type)

Stegobium Motschulsky 1860: 154 TRIBE DORCATOMINI (S. paniceum (Linnaeus) 1758: 357). The Caenocara Thomson 1859: 90 Distribution. Puerto Rico, Guadeloupe, Cen- C. sp.; Genaro 1996: 9 tral America, se USA; cosmopolitan; a pest in Distribution. a wide variety of stored organic materials Cuba localities. M: Macunayagua; phoretic Cuba localities. on a female of the ground nesting Collections. Anthophora atrata Cresson Collections. Havana SUBFAMILY TRIBE XYLETININI 122 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE PROTHECINI Genaro, J.A. 1996. Foresis de Caenocara sp. (Co- leoptera: Anobiidae) en Anthophora atrata (Hy- Protheca LeConte 1865: 241 menoptera: Anthophoridae). Melissa 9: 9. P. pilosa Zayas 1988: 77 Philips, T.K. 1997. Cubaptinus Zayas, a new syn- Distribution. Cuba. Endemic onym of Fabrasia Martinez and Viana (Co- Cuba localities. Type locality not given leoptera; Anobiidae, Ptininae). Coleop. Bull. 51: Collections. Zayas (type) 52. Philips, T.K., M.A. Ivie, and L.L. Ivie. 1998. TRIBE MESOCOELOPINI Leaf mining and grazing in spider beetles (Co- leoptera: Anobiidae: Ptininae): an unreported Mesocoelopus Jacquelin du Val 1860: 143 mode of larval and adult feeding in the M. rudepubens Pic 1948: 13 Bostrichoidea. Proc. Ent. Soc. Wash. 100: 147- Distribution. Cuba. Endemic 153. Cuba localities. White, R.E. 1982. A catalog of the Coleoptera of Collections. America north of Mexico. Family: Anobiidae. USDA Agric. Handbook 529-70. TRIBE TRICORYNINI White, R.E. 1984. A revision of the American ge- nus Cryptorama (Coleoptera: Anobiidae). Trans. Cryptorama Fall 1965: 256 Amer. Ent. Soc. 110: 77-127. C. antillensis White 1984: 85 Distribution. Cuba, Virgin Islands?, SERIES CUCUJIFORMIA Guadeloupe? SUPERFAMILY LYMEXYLOIDEA Cuba localities. Ci: Cayamas. SS: Trinidad 85. FAMILY LYMEXYLIDAE Collections. Washington The ship-timber beetles C. rufescens White 1984: 117 These beetles are unique in having reduced Distribution. Cuba, USA (FL) elytra and no transverse fold in the hind wing. The Cuba localities. Ci: Cayamas larvae bore in dead wood and are associated with Collections. Washington ambrosia fungi.

Tricorynus Waterhouse 1849: 68 SUBFAMILY LYMEXYLINAE T. herbarius (Gorham) 1883: 207 (Catorama) Distribution. Cuba, México, S USA (CA-FL- Atractocerus Palisot de Beauvois 1801: 3 MA) A. braziliensis Lepeltier and Audinet-Serville 1825: Cuba localities. Ha: Almendares; Marianao. 309; Gundlach 1891: 115 A stored products or household pest Distribution. Cuba, Jamaica, Puerto Rico, Collections. Lesser Antilles, México, Central and South T. tabaci (Guérin-Méneville) 1850: 437 America Distribution. Cuba, Guatemala, México, USA Cuba localities. M: Cárdenas (FL); a stored products pest Collections. Gundlach no. 840 Cuba localities. Collections. SUPERFAMILY CLEROIDEA 86. FAMILY PHLOIOPHILIDAE TRIBE INCERTAE SEDIS This family occurs in Europe, and the larvae feed on the wood-rotting fungus Phlebia. Neosothes White 1967: 43 N. granulatus White 1967: 48 87. FAMILY TROGOSSITIDAE Distribution. Cuba. Endemic The bark-gnawing beetles and cadelles Cuba localities. SC: Caney; in mango (t.l.) The species are often predators, and one is a Collections. Washington (type) stored products pest. The Peltinae often occur in bracket fungi. Keys are in Barron (1971). REFERENCES Belles, X. 1992. Sistematica, historia natural y SUBFAMILY LOPHOCATERINAE biogeografia del genero Pitnus Gorham, 1990 (Coleoptera, ). Eos 68: 167-192. Lophocateres Olliff 1883: 180 A Checklist of the Beetles of Cuba 123

(L. pusillus (Klug) 1832: 159; Barron 1971: 43) Collections. Distribution. Panamá, Curaçao, USA (FL), T. floridanus Schaeffer 1918: 199; Barron 1971: cosmopolitan. A stored products pest 111 Cuba localities. Distribution. Cuba, Jamaica, Hispaniola, Collections. Puerto Rico, Guadeloupe, México, Brazil, Surinam, Bahamas USA (FL, LA) SUBFAMILY TROGOSSITINAE Cuba localities. Ci: Soledad. PR: Guanahacabibes Airora Reitter 1876: 18 Collections. New York (A. striatopunctata Reitter 1876: 20) T. mauritanicus (Linnaeus) 1758: 417; Jacquelin Distribution. West Indies, Brazil du Val 1857: 254; Chevrolat 1864: 606; Cuba localities. Gundlach 1891: 97; Barron 1971: 93. The cadelle Collections. beetle Distribution. Cuba (introduced), USA (FL), Calanthasoma Reitter 1876: 10 cosmopolitan; in flour, meal, and grains. An (C. flavomaculata Reitter 1876: 11) important stored products pest Distribution. West Indies Cuba localities. Ha: Havana Cuba localities. Collections. Collections. T. punctulata Reitter 1875: 74 Distribution. Cuba, Puerto Rico, Guadeloupe Temnoschila Westwood 1830: 231; (=Temnochila Cuba localities. Erichson 1844: 449) Collections. (T. borrei Reitter 1875: 37) T. soror (Jacquelin du Val) 1857: 252 (Trogosita); Distribution. West Indies, Colombia Chevrolat 1864: 604; Gundlach 1891: 95; Cuba localities. Barron 1971: 101 Collections. Distribution. Cuba, Bahamas, USA (FL T. elongata Zayas 1988: 67 Keys) Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. M: Cárdenas Cuba localities. SC: Pico Turquino (t.l.) Collections. New York Collections. Zayas (type) T. sulcifrons (Jacquelin du Val) 1857: 253 T. hubbardi Léveillé 1889: 417; Barron 1971: 75 (Trogosita); Chevrolat 1864: 606; Gundlach Distribution. Cuba, Bahamas, USA (FL) 1891: 96 Cuba localities. Distribution. Cuba, México to Brazil Collections. Cuba localities. (T. insignis Reitter 1875: 10) Collections. Distribution. West Indies, Colombia T. transversicollis (Jacquelin du Val) 1857: 255 Cuba localities. (Trogosita); Chevrolat 1864: 606: Gundlach Collections. 1891: 97 Distribution. Cuba, Puerto Rico, Guadeloupe Tenebroides Piller and Mitterpacher 1883: 87 Cuba localities. Widespread (T. bipustulata Fabricius 1801: 152) Collections. Distribution. West Indies, México to Argen- tina REFERENCES Cuba localities. Barron, J. R. 1971. A revision of the Trogositidae Collections. of America north of México (Coleoptera: T. elongatula (Jacquelin du Val) 1857: 251 Cleroidea). Ent. Soc. Canada Mem. 75. (Trogosita); Chevrolat 1864: 605; Gundlach 1891: 96 88. FAMILY CHAETOSOMATIDAE Distribution. Cuba. Endemic This family is limited to New Zealand, and the Cuba localities. M: Cárdenas larvae live in the burrows of wood-inhabiting beetles. Collections. T. flaviclavis Reitter 1875: 78 89. FAMILY CLERIDAE Distribution. Cuba. Endemic The checkered beetles Cuba localities. Reviewed by Weston Opitz, August, 2002. 124 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Adults of some species are predators on larvae Cuba localities. of wood-boring beetles and others feed on pollen. Collections. Some feed on dead animal or plant matter. SUBFAMILY HYDNOCERINAE SUBFAMILY THANOCLERINAE Phyllobaenus Dejean 1837: 127 Thaneroclerus Lefebvre 1838: 13 P. monticola Zayas 1988: 62 T. buquet (Lefebvre) 1835: 577 (Clerus) Distribution. Cuba. Endemic Distribution. Cuba (introduced), Guadaloupe, Cuba localities. PR: Lomas de Cajálbana USA (intercept at ports), native to the Old (t.l.); found in leaf litter World. These may occur in bracket fungi Collections. Zayas (type) Cuba localities. Collections. SUBFAMILY EPIPHLOEINAE

SUBFAMILY TILLINAE Epiphloeus Spinola 1841: 75 E. nebulosus Chevrolat 1876: 28 Monophylla Spinola 1841: 75 Distribution. Cuba. Endemic M. cinctipennis (Chevrolat) 1874: 281 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. E. quadristigma Chevrolat 1874: 320 Collections. Distribution. Cuba. Endemic Cuba localities. Onychotillus Chapin 1945: 596 Collections. O. cubana Zayas 1988: 57 Distribution. Cuba. Endemic Madoniella Pic 1935: 10 Cuba localities. Gr: Cabo Cruz. SC: Pico M. flavicollis ( Zayas) 1988: 61 (Phlogistosternus); Turquino (t.l.) Opitz 1997: 62 Collections. Zayas (type) Distribution. Cuba. Endemic O. dimidiata Zayas 1988: 58 Cuba localities. Gr: Cabo Cruz. SS: Trinidad. Distribution. Cuba. Endemic Type locality not given Cuba localities. M: Canasí (t.l.) Collections. Zayas (type) Collections. Zayas (type) M . orientalis (Zayas) 1988: 60 (Phlogistosternus) O. minuta Zayas 1988: 57 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Gran Piedra; Loma del Cuba localities. Gu: Yateras (t.l.) Gato; Pico Turquino. Type locality not given Collections. Zayas (type) Collections. Zayas (type) O. trinitatis Zayas 1988: 56 Distribution. Cuba. Endemic SUBFAMILY ENOPLIINAE Cuba localities. SS: Lomas de Trinidad (t.l.) Collections. Zayas (type) Corinthiscus Fairmaire and Germain 1861: 4 C. axinoides Walcott 1923: 3 SUBFAMILY CLERINAE Distribution. Cuba. Endemic Cuba localities. Gu: Rio Seco; San Carlos (t.l.) Aulicus Spinola 1841: 74 Collections. New York (type) A. alboguttulatus Chevrolat 1874: 299 Distribution. Cuba. Endemic Neorthopleura Barr 1976: 2 Cuba localities. [N. thoracica (Say) 1823: 188] Collections. Distribution. Not in Cuba, or México; wide- A. basicollis Chevrolat 1874: 299 spread in USA) Distribution. Cuba. Endemic N. lepida Klug 1842: 359 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Cayamas. PR A. bilineatus Chevrolat 1874: 300 Collections. Distribution. Cuba. Endemic (N. nesiotes Barr 1941: 359) A Checklist of the Beetles of Cuba 125

Distribution. Cayman Brac, Bahamas 92. FAMILY Cuba localities. This family occurs in the southern Palearctic, Collections. Asia, and Africa. N. zayasi Barr 1976: 10 Distribution. Cuba. Endemic 93. FAMILY MELYRIDAE Cuba localities. Ci: Cumanayagua The soft-winged flower beetles Collections. Reviewed by Douglas G. Le Doux,and Adriean Mayor, March, 1997 Pelonium Spinola 1844: 341 Adults occur on flowers and feed on pollen, P. piciventre Chevrolat 1874: 325 spores or other insects. Both adults and larvae may Distribution. Cuba. Endemic be very substrate specific, occurring on coastal sand Cuba localities. dunes, tidal flats, or on specific types of rock sub- Collections. strates such as sandstones, igneous, and calcare- ous rock outcrops. Larvae are predators and may SUBFAMILY KORYNETINAE occur in litter, soil, or under bark or rocks.

Necrobia Olivier 1795: 76 SUBFAMILY MALACHIINAE N. ruficollis Fabricius 1775: 57; Jacquelin du Val TRIBE MALACHIINI 1856: 92 (Corynetes); Gundlach 1891: 244. The redshouldered ham beetle Ablechrus Waterhouse 1877: 79 Distribution. Cuba (introduced), West Indies, A. cubanus Wittmer 1984: 232 México to Argentina, cosmopolitan; in stored Distribution. Cuba. Endemic products Cuba localities. PR: Guane (t.l.); Pinar del Cuba localities. Widespread Rio; Viñales Collections. Gundlach no. 544 Collections. New York (type); Basel N. rufipes DeGeer 1775: 165. The redlegged ham beetle Anthocomus Erichson 1840: 97 Distribution. Cuba (introduced), West Indies, A. dimidiatus Erichson 1840: 111; Jacquelin du México to Chile, USA (FL), cosmopolitan; found Val 1856: 91; Gundlach 1891: 250 on dry animal carcasses (Acanthocomus) Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 339 Cuba localities. M: Cárdenas; Cienaga de (N. violacea (Linnaeus) 1758: 356) Zapata Distribution. Cosmopolitan Collections. Gundlach no. 458 Cuba localities. Collections. Attalomimus Wittmer 1976: 290 A. purpureipennis Wittmer 1976: 290 REFERENCES Distribution. Cuba. Endemic Barr, W.F. 1976. Taxonomy of the new clerid ge- Cuba localities. Ci: Soledad (t.l.) nus Neorthopleura (Coleoptera). Melanderia 24: Collections. Basel. Cambridge 1-14. Opitz, W. 1997. Classification, natural history, and Attalus Erichson 1840: 89 evolution of the Epiphloeinae (Coleoptera: A. dimidiatus Erichson 1840: 11; Fall 1917: 81 Cleridae). Part I. The genera of Epiphloeinae. Distribution. Cuba, USA Insecta Mundi 11: 51-96. Cuba localities. Collections. Berlin? (type) 90. FAMILY ACANTHOCNEMIDAE A. nigra Zayas 1988: 64 This family naturally occurs in Australia, and Distribution. Cuba. Endemic has been introduced widely to other countries. Cuba localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) 91. FAMILY PHYCOSECIDAE A. sp. Zayas 1988: 63 This family lives in coastal sands of Australia Distribution. Cuba. Endemic and New Zealand, and they scavenge on detritus Cuba localities. SC: Pico Turquino (t.l.) or dead fish. Collections. Zayas 126 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Malachius Fabricius 1775: 207 97. FAMILY NITIDULIDAE M. metalicus Zayas 1988: 65 The sap-feeding beetles Distribution. Cuba. Endemic Reviewed and corrected by J. Jelinek, May 1995 Cuba localities. SC: Gran Piedra (t.l.) Members of this family feed on a variety of Collections. Zayas (type) materials, including decaying fruits, dried fruits, M. turquinensis Zayas 1988: 66 pollen, and leaf litter. Material mentioned below Distribution. Cuba. Endemic which was seen by Nicholas Evans(1977) is in Ha- Cuba localities. SC: Pico Turquino (t.l.) vana or Prague (J. Jelinek, pers. comm.). Collections. Zayas (type) SUBFAMILY CILLAEINAE REFERENCES Colopterus Erichson 1842: 149 (the genus was Wittmer, W. 1976. Zur Kenntnis der Malachiidae revised in an unpublished thesis by L. Watrous, der Antillen (Col.). (33 Beitrag zur Kenntnis which may contain new information) der Neotropischen Malachiidae). Mitteilungen C. amputatus Erichson 1843: 243; Jacquelin du Val der Schweizerischen Entomologischen 1856: 94 (Colastes); Chevrolat 1864: 600; Gesellschaft 49: 281-291. Gundlach 1891: 89 Wittmer, W. 1984. Über die Gattungen Ablechrus Distribution. Cuba, Bahamas, Puerto Rico, Waterhouse and Tucumanius Pic (Coleoptera: Guadeloupe, México to Argentina Malachiidae). Ent. Basilensia 9: 226-243. Cuba localities. Widespread Collections. SUPERFAMILY CUCUJOIDEA C. posticus Erichson 1843: 237 94. FAMILY Distribution. Cuba (Nicholas Evans 1977), This family occurs only in Chile, Argentina, Central America and Australia. Cuba localities. Collections. 95. FAMILY SPHINDIDAE C. truncatus Randall 1838: 18 The cryptic slime mold beetles Distribution. Cuba, Puerto Rico to Grenada, Adults and larvae feed on spores of slime molds. México to Argentina, Canada (BC-PQ), USA (widespread) SUBFAMILY SPHINDINAE Cuba localities. Collections. Carinisphindus McHugh 1990: 309 C. bicolor (Fisher) (Eurysphindus) 1936: 201 Conotelus Erichson 1843: 249 (see key in Jelinek Distribution. Cuba. Endemic and Nicholas Evans 1982) Cuba localities. CA: Baraguá (t.l.) (C. conicus Fabricius 1801: 603; Jelinek and Nicho- Collections. Havana (Bruner). Washington las Evans 1982: 234) (type) Distribution. Not in Cuba, but Puerto Rico, Hispaniola? Vigin Islands, Lesser Antilles, Sphindus Chevrolat 1833: 8 Mexico? S. cubensis Reitter 1878: 202 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. C. fuscipennis Erichson 1843: 249; Jacquelin du Collections. Val 1856: 94; Chevrolat 1864: 600; Gundlach 1891: 90; Jelinek and Nicholas Evans 1982: 230 REFERENCES Distribution. Cuba, Bahamas, USA (FL and McHugh, J.V. 1990. Carinisphindus, a new ge- Keys) nus and three new species of Neotropical Cuba localities. Widespread Sphindidae (Coleoptera: Clavicornia). Coleop. Collections. Berlin. Cambridge. Havana. Bull. 44: 307-322. Prague C. zayasi Jelinek and Nicholas Evans 1982: 231 96. FAMILY BRACHYPTERIDAE Distribution. Cuba, Jamaica, Hispaniola This family occurs in most regions, including Cuba localities. Gu: Yunque de Baracoa. SC: the United States, but not the Neotropics. Sierra Maestra (t.l.). SS: Topes de Collantes Collections. Havana (type). Prague A Checklist of the Beetles of Cuba 127

SUBFAMILY CARPOPHILINAE C. hemipterus (Linnaeus) 1758: 358; Jacquelin du Val 1856: 95; Chevrolat 1864: 601; Gundlach Amphicrossus Erichson 1843: 346; key to US 1891: 90. The dried fruit beetle species in Parsons 1972 Distribution. Cuba, widespread West Indies, A. ciliatus (Olivier) 1811: 210 México to Argentina, USA, Old World Distribution. Cuba, Panamá, Canada (ON), Cuba localities. Widespread USA (widespread) Collections. Cuba localities. C. maculatus Murray 1864: 372; Nicholas Evans Collections. 1977 Distribution. Cuba; Oriental Region, Pacific Carpophilus Stephens 1829: 81 islands including Hawaii, Central America, C. craigheadi Dobson 1972: 145 West Indies (Connell 1977) Distribution. Cuba (Nicholas Evans 1977); Cuba localities. USA (FL). On Harrisia cactus flowers Collections. Cuba localities. C. marginellus Motschulsky 1858: 40; Nicholas Collections. Evans 1977 C. dimidiatus (Fabricius) 1792: 261; Jacquelin du Distribution. Cuba; cosmopolitan Val 1856: 96; Chevrolat 1864: 601; Gundlach Cuba localities. 1891: 91. The corn Collections. Distribution. Cuba, Bahamas, Puerto Rico C. mutilatus Erichson 1843: 258; Jacquelin du Val to Grenada, México to Argentina, Canada (PQ), 1856: 96; Chevrolat 1864: 601; Gundlach 1891: USA (CA, TX-NY-FL); cosmopolitan 91 Cuba localities. IJ: Los Indios = hemipterus (Fabricius 1792): 261, not (Linnaeus Collections. Carnegie 1758): 359; Nicholas Evans 1977. Note. C. dimidiatus as listed in Coleopterorum Distribution. Cuba, USA (widespread); cos- Catalogus represents a complex of sibling spe- mopolitan cies (e.g. multilatus, fumatus, maculatus, Cuba localities. vittiger, etc.) and the true identity of many Collections. listed synonyms is uncertain without the revi- C. obsoletus Erichson 1843: 259; Nicholas Evans sion of type species. We recommend accepting 1977 only the reliable synonymy given by Audisio Distribution. Cuba, USA (widespread); cos- (1993). See Williams et al. 1983 mopolitan C. floralis Erichson 1843: 261 Cuba localities. Distribution. Cuba (Nicholas Evans 1977), Collections. México, eastern USA (Parsons 1943) C. pilosellus Motschulsky 1858: 41; Nicholas Evans Cuba localities. 1977 Collections. Distribution. Cuba; cosmopolitan C. freemani Dobson 1956: 41 Cuba localities. Distribution. Cuba (Nicholas Evans 1977); Collections. cosmopolitan C. tempestivus Erichson 1843: 260; Jacquelin du Cuba localities. Val 1856: 96; Chevrolat 1864: 601; Gundlach Collections. 1891: 9 C. fumatus Boheman 1851: 564 Distribution. Cuba, Puerto Rico, USA (LA- C. ochropterus Klug 1862: 209; Jacquelin du Val AR-GA-FL) 1856: 96 (as C. tempestivus); Chevrolat 1864: Cuba localities. M: Cárdenas 602; Gundlach 189l: 92; Nicholas Evans 1977 Collections. Distribution. Cuba (introduced); Africa, Por- (C. vitraci Grouvelle 1902: 757) tugal, Azores, Central America, Caribbean Is- Distribution. Cuba? (Nicholas Evans 1977), lands (which?), USA (FL). Apparently of Guadeloupe (type not seen) Afrotropical origin, and introduced into other Cuba localities. countries Collections. Cuba localities. Collections. 128 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Epuraea Erichson 1843: 267 Distribution. Cuba, México, Guatemala, Co- E. (Haptoncus) luteola Erichson 1843: 272; lombia Jacquelin du Val 1856: 97; Chevrolat 1864: 602; Cuba localities. Gundlach 1891: 92. The yellowbrown sap beetle Collections. Distribution. Cuba, widespread West Indies, México, Central America, USA (TX-FL); Prometopia Erichson 1843: 279 cosmotropical P. cf. rauca Sharp 1890: 316 Cuba localities. M: Cárdenas Distribution. Cuba (unpublished), México, Collections. Central America (type not seen but the genus is in Cuba) Urophorus Murray 1864: 342 Cuba localities. U. humeralis (Fabricius) 1798: 74 Collections. Distribution. Cuba (introduced) (Nicholas Evans 1977), Puerto Rico, Uruguay, Ceylon, Pocadius Erichson 1843: 318 Malaysia, China P. brevis Reitter 1873: 94 Cuba localities. Distribution. Cuba. Endemic (maybe conspe- Collections. cific with P. helvolus?) Cuba localities. SUBFAMILY NITIDULINAE Collections. Notes. Taxonomy of Cuban Pocadius is not clear. Lobiopa Erichson 1843: 291 Jelinek did not see any material from Cuba. L. insularis Laporte 1840: 10 The data on the occurrence of P. helvolus in L. decumana Erichson 1843: 295; Jacquelin du South America apparently refer to other spe- Val 1856: 97; Chevrolat 1864: 602; Gundlach cies (Jelinek 1979). It occurs in México, east- 1891: 93 ern USA and Canada (Parsons 1943). Accord- Distribution. Cuba to Grenada, México to Ar- ing to that author the status of Cuban species gentina, USA (Gulf States) is doubtful Cuba localities. Widespread P. helvolus Erichson 1843: 320 Collections. P. ferrugineus (L.) Chevrolat 1863b: 604; Gundlach 1891: 95 Mystrops Erichson 1843: 234 (see Gillogly 1955. Distribution. Cuba, México to Argentina, Rev. Bras. Ent. 3: 191-204) USA M. insularis Grouvelle 1898: 352 Cuba localities. Distribution. Cuba (Nicholas Evans 1977), Collections. Martinique; in palm flowers Cuba localities. Stelidota Erichson 1843: 300 Collections. Taxonomy of Neotropical Stelidota is very confused and their revision is badly needed. Therefore it Omosita Erichson 1843: 298 is difficult to give the exact distribution of par- O. colon (Linnaeus) 1758: 362 ticular species. Nevertheless, the list corre- Distribution. Cuba (Nicholas Evans 1977) (in- sponds to the present knowledge. Bibliography: troduced), North America; native to the Pale- Weiss and Williams (1980) arctic Region S. coenosa Erichson 1843: 303; Jacquelin du Val Cuba localities. 1856: 98; Chevrolat 1864: 604; Gundlach 1891: Collections. 95 Distribution. Cuba, South America Pallodes Erichson 1843: 348 Cuba localities. P. ruficollis Reitter 1873: 134 Collections. Distribution. Cuba, Hispaniola (unpublished) (S. ferruginea Reitter 1873: 14) Cuba localities. IJ: Nueva Gerona Distribution. Cuba? (Nicholas Evans 1977), Collections. Carnegie West Indies, Central America, South America, USA (AZ-MI-NJ-FL) Platychora Erichson 1843: 284 Cuba localities. P. lebasi Erichson 1843: 285 Collections. A Checklist of the Beetles of Cuba 129

S. geminata (Say) 1825: 181; Jacquelin du Val Erichson (Coleoptera: Nitidulidae, Nitidulinae). 1856: 98; Chevrolat 1864: 603; Gundlach 1891: Ohio Agric. Res. and Development Center Res. 94. The strawberry sap beetle Circular 255, Wooster, 37 pp. Distribution. Cuba, Puerto Rico to Grenada, Williams, R.N., D.S. Fickle, M. Kehat, D. México to Brazil Blumberg and M.G. Klein. 1983. Bibliogra- Cuba localities. Widespread phy of the genus Carpophilus Stephens (Co- Collections. leoptera: Nitidulidae). Ohio Agric. Res. and Dev. S. ruderata Erichson 1843: 303; Jacquelin du Val Center Res. Circular 278, Wooster, 95 pp. 1856: 99; Chevrolat 1864b; Gundlach 1891: 94. Distribution. Cuba, Hispaniola, Puerto Rico 98. FAMILY SMICRIPIDAE to Guadeloupe The palmetto beetles Cuba localities. Widespread Adults and larvae occur in decaying flowers, Collections. leaf litter, and under bark.

Teichostethus Sharp 1891: 354 Smicrips LeConte 1878: 399 T. sp. indet. Cuba (Nicholas Evans 1977; only a S. palmicola LeConte 1878: 399 single unidentifiable specimen available) S. hypocoproides Reitter 1876: 301 Distribution. Distribution. Cuba, Puerto Rico, USA (FL). Cuba localities. Found in association with flowers of Sabal pal- Collections. metto. Cuba localities. REFERENCES Collections. Audisio, P. 1993. Coleoptera: Nitidulidae- . Fauna d’Italia, vol. 32, xvi + 971 99. FAMILY MONOTOMIDAE pp. Ed. Calderini, Bologna. The root eating beetles Connell, W.A. 1977. A key to Carpophilus sap These beetles feed on the spores of fungi and in beetles associated with stored foods in the United various kinds of decaying vegetation. States (Coleoptera: Nitidulidae). U.S. Dept. Agric. Coop. Plant Pest Rep. 2(23): 398-404. SUBFAMILY MONOTOMINAE Jelinek, J. 1975. New genus of Oriental TRIBE MONOTOMINI Meligethinae with note on supergeneric classi- fication of Nitidulidae (Coleoptera, Nitidulidae). Monotoma Herbst 1793: 22 Annot. Zool. Bot., Bratislava, no. 102: 1-9. M. sp. Jelinek, J. 1979. Revision of South American spe- Distribution. Cuba cies of the genus Procadius Er. with descrip- Cuba localities. SC: Gran Piedra tion of new genus (Coleoptera, Nitidulidae). Collections. Gatineau Acta Ent. Mus. Nat. Pragae 39 (1977): 29-44. Jelinek, J. and A.N. Evans. 1982. Species of the TRIBE EUROPINI genus Conotelus from the West Indies (Co- leoptera, Nitidulidae). Acta Ent. Bohemoslovaca Bactridium LeConte 1861: 86 79: 226-237. B. angulicolle Reitter 1872: 33 Nicholas Evans, A. 1977. Nitidulidae of Cuba (Co- Distribution. Cuba, Colombia? leoptera). Entomological Institute, Cuba localities. Chechoslovakia Academy of Sciences, Prague. Collections. Unpublished CSc thesis, J. Jelinek Supervisor. B. cubensis (Chevrolat) 1864b: 604 (Rhizophagus); Specimens in Prague and Havana. Not seen. Gundlach 1891: 98 Parsons, C.T. 1943. A revision of Nearctic Distribution. Cuba. Endemic Nitidulidae (Coleoptera). Bull. Mus. Comp. Zool. Cuba localities. Widespread 92, no. 3: 121-278, 13 pls. Collections. Parsons, C.T. 1972. On the mesosternum in some B. heydeni Reitter 1872: 33 Nitidulidae (Coleoptera) with a key to the New Distribution. Cuba. Endemic World Amphicrossus. Coleop. Bull. 26: 103-115. Cuba localities. Weiss, M. J. and Williams, R.N. 1980. An an- Collections. notated bibliography of the genus Stelidota 130 Arthropods of Florida and Neighboring Land Areas, Vol. 18

100. FAMILY Collections. This family occurs in Australia, southern South T. quadripunctatus Schaufuss 1876: 387 America, and South Africa. Distribution. Cuba. Endemic Cuba localities. 101. FAMILY Collections. This family occurs in the warmer parts of the Old World from Africa to Japan. They feed on sap SUBFAMILY from tree wounds. Ahasverus Gozis 1881: CXXVII 102. FAMILY PHLOEOSTICHIDAE A. advena (Waltl) 1832: 169 (Cryptophagus); Tho- This family occurs in Europe, Australia, New mas 1993: 32. The foreign grain beetle Zealand and Chile. They occur with moss, or un- Distribution. Cuba (introduced), widespread der bark.. Neotropics, USA (FL); cosmopolitan; on moldy stored products 103. FAMILY SILVANIDAE Cuba localities. The silvanid flat bark beetles Collections. Reviewed by M. C. Thomas, October, l998 A. rectus (LeConte) 1854: 78 (Silvanus): Thomas This family commonly occurs under bark and 1993: 34 in litter, where members feed on fungi. Some are Distribution. Cuba, Puerto Rico, USA (AZ- pests of stored products. MO-FL) Cuba localities. SUBFAMILY BRONTINAE Collections. TRIBE PSAMMOECINI Cathartosilvanus Grouvelle 1912: 340 Cryptamorpha Wollaston 1854: 156 (C. opaculus (LeConte) 1854: 78; Thomas 1993: 28) (C. desjardinsii (Guérin-Ménèville) 1829: 196 Distribution. Jamaica, Hispaniola, Virgin (Psammoecus); Thomas 1993: 13) Islands, Grenada, Guadeloupe, St. Vincent, Distribution. St. Vincent, Grenada, USA Trinidad, Central America south to Brazil and (FL); cosmopolitan Bolivia, USA (FL, CA, AZ) Cuba localities. Cuba localities. Collections. Collections.

Telephanus Erichson 1846: 329 Cathartus Reiche 1854: 77 T. apicalis Grouvelle 1877: 207 C. quadricollis (Guérin-Méneville) 1829: 198 Distribution. Cuba. Endemic (Silvanus); Thomas 1993: 35. The squarenecked Cuba localities. grain beetle Collections. C. cassiae Reiche 1854: 78; Chevrolat 1864: 614; T. bipunctatus Schaufuss 1876: 385 Gundlach 1891: 101 Distribution. Cuba. Endemic Sylvanus gemellatus Jacquelin du Val 1857: 250; Cuba localities. Chevrolat 1864: 614 (Cathartus) Collections. Distribution. Cuba, widespread Neotropics, T. cubanus Nevermann 1932: 28 USA (FL); cosmopolitan Distribution. Cuba, Puerto Rico Cuba localities. Widespread Cuba localities. Collections. Collections. T. megacephalus Nevermann 1932: 12 Monanus Sharp 1879: 207 Distribution. Cuba, Puerto Rico M. concinnulus (Walker) 1858: 207 (Monotoma); Cuba localities. Thomas 1993: 31 Collections. Distribution. Cuba, widespread Neotropics, T. pallidus Reitter 1874: 523 USA (FL); cosmopolitan; stored products and Distribution. Cuba, Puerto Rico (not T. moldy plant wastes pallidulus Chevrolat 1863: 612, which occurs Cuba localities. only in Puerto Rico) Collections. Cuba localities. A Checklist of the Beetles of Cuba 131

Nausibius Lentz 1857: 75 104. FAMILY PASSANDRIDAE N. clavicornis (Kugelann) 1794: 511 (); The parasitic flat bark beetles Thomas 1993: 26 Members of this family are ectoparasitic on lar- Corticaria dentatus Marsham 1802: 108; vae of wood boring insects Jacquelin du Val 1857: 250 (Silvanus); Chevrolat 1864: 613 (Nausibius); Gundlach Catogenus Westwood 1830: 221 1891: 101 (C. rufus Fabricius 1798: 123) Distribution. Cuba, widespread Neotropics; Distribution. Cuba? The species occurs in cosmopolitan south Florida and others are in the Caymans Cuba localities. Widespread and Puerto Rico, so the genus should occur in Collections. Cuba (Slipinski 1989; Thomas 1993) Cuba localities. Oryzaephilus Ganglbauer 1899: 45 Collections. (O. acuminatus Halstead 1980: 309) Distribution. USA (FL); cosmopolitan; in Passandra Dalman 1818: 146 stored products P. fasciata Gray 1832: 93; Chevrolat 1864: 611; Cuba localities. Gundlach 1891: 112; Slipinski 1986: 576 Collections. Distribution. Cuba, México to Argentina (O. mercator Fauvel 1889: 132). The merchant Cuba localities. Widespread grain beetle Collections. Gundlach no. 5 Distribution. Cuba?; cosmopolitan; in stored products Taphroscelidia Crotch 1873: 44 Cuba localities. T. linearis LeConte 1863: 70 Collections. Distribution. Cuba, Bahamas, México, Bra- O. surinamensis (Linnaeus) 1758: 357; Chevrolat zil, USA (CA-FL) 1864: 613 (Silvanus); Gundlach 1891: 100. The Cuba localities. saw-toothed grain beetle Collections. O. s. affinis Chevrolat (Leng and Mutchler 1914: 416) REFERENCES Distribution. Cuba (introduced), widespread Slipinski, S.A. 1986. A review of the Passandridae Neotropics; cosmopolitan of the world (Coleoptera: Cucujoidea) I. Genus Cuba localities. Passandra Dalman. Ann. Mus. Civ. Storia Nat. Collections. Genova 86: 553-603. Slipinski, S.A. 1989. A review of the Passandridae Silvanopropus Reitter 1911: 45 of the world (Coleoptera: Cucujoidea) II. Genus (S. scuticollis (Walker) 1859: 53) Catogenus Westwood. Polskie Pismo Ent. 59: Distribution. USA (OK-WV-FL); cosmopoli- 85-129. tan Thomas, M.C. 1993. The flat bark beetles of Cuba localities. Florida (Coleoptera: Silvanidae, Passandridae, Collections. Laemophloeidae). Arthropods of Florida, Vol 15, 93 pp. Silvanus Latreille 1804: 158 (S. sp.; Thomas 1993: 23.) 105. Distribution. Cuba?; should be under bark These commonly occur under bark and are fac- of trees, also in stored products ultative predators. They do not occur in the Carib- Cuba localities. bean. Collections. 106. FAMILY LAEMOPHLOEIDAE REFERENCES The lined flat bark beetles Thomas, M.C. 1993. The flat bark beetles of Members of this family occur under bark where Florida (Coleoptera: Silvanidae, Passandridae, they feed upon fungi, and some are pests of stored Laemophloeidae). Arthropods of Florida, Vol 15, food products (Thomas 1993, 1991). 93 pp. 132 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Charaphloeus Casey 1916: 127 P. modestus (Say) 1827: 268 C. bituberculatus (Reitter) 1876: 316 Distribution. Cuba, Jamaica, Trinidad, Distribution. Cuba, Bahamas, Jamaica, México to Panamá, USA (AZ-NY-FL) Hispaniola, Puerto Rico, Virgin Islands, USA Cuba localities. (FL) Collections. Cuba localities. (P. politissimus (Wollaston) 1867: 268) Collections. Distribution. Greater and Lesser Antilles, México to Brazil, USA (FL), Africa Cryptolestes Ganglbauer 1899: 608 Cuba localities. C. bicolor (Chevrolat) 1864b: 612 (Laemophloeus); Collections. Gundlach 1891: 112 Laemophloeus concavus Reitter 1778: 192 REFERENCES Distribution. Cuba. Endemic Thomas, M.C. 1993. The flat bark beetles of Cuba localities. M: Cárdenas Florida (Coleoptera: Silvanidae, Passandridae, Collections. Gundlach no. 396 Laemophloeidae). Arthropods of Florida, Vol 15, C. pusillus (Schönherr) 1817-55; Chevrolat 1864: 93 pp. 612; Gundlach 1891: 113 Thomas, M.C., and S.B. Peck.1991. Survey of Distribution. Cuba, widespread West Indies; insects of south Florida and the Florida Keys; cosmopolitan Flat Bark Beetles (Coleoptera: Cucujidae (sens. Cuba localities. M: Cárdenas lat.) {Laemophloeidae: Passandirdae: Silvan- Collections. Gundlach no. 397 idae}. Florida Ent. 74: 536-543. (C. unicicornis (Reitter) 1876: 45) Distribution. Greater and Lesser Antilles, 107. FAMILY PROPALTICIDAE México to Argentina, USA (se and FL) This family is restricted to the Old World, from Cuba localities. Africa to Australia, Japan, and Hawaii. They oc- Collections. cur under bark of dead trees.

Laemophloeus Laporte 1840: 384 108. FAMILY PHALACRIDAE L. chevrolati Grouvelle 1878: 264. (a synonym of The shining flower or shining mold beetles L. lecontei Grouvelle?) Adults often occur with flowers or vegetation Distribution. Cuba. Endemic and larvae of some feed in flower heads of compos- Cuba localities. ites, while others feed on spores or ergots of various Collections. rusts, smuts and pyrenomycete fungi.

Lathropus Erichson 1845: 307 Acylomus Sharp 1888: 256 (L. pictus Schwarz 1878: 358) A. cubensis Casey 1916: 76 Distribution. Guadeloupe, México, Virgin A. cubiensis Leng and Mutchler 1914: 199 Islands, USA (FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Ha: Havana (t.l.) Collections. Collections. Washington (type) A. detractus Casey 1916: 75 Parandrita LeConte and Horn 1883: 133 Distribution. Cuba. Endemic P. permixtus (Grouvelle) 1912: 303 Cuba localities. PR: Bahia Honda (t.l.) Distribution. Cuba, Virgin Islands, Collections. Washington (type) Guadeloupe, Panamá, USA (FL) A. quadrispinosus Casey 1916: 75 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. PR: Bahia Honda (t.l.) Collections. Washington (type) Placonotus MacLeay 1871: 186 P. macrognathus Thomas 1984: 12 Heterolitus Guillebeau 1893: 278 Distribution. Cuba, USA (FL Keys) H. strigellus Guillebeau 1894: 297 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Collections. A Checklist of the Beetles of Cuba 133

Litolibrus Sharp 1889: 258 Distribution. Cuba, USA, Old World. A L. princeps Schwarz 1878: 361 (Olibrus) stored products pest Distribution. Cuba, USA (FL) Cuba localities. Cuba localities. Collections. Collections. 112. FAMILY LAMINGTONIIDAE Phalacrus Paykull 1800: 438 This family is restricted to central eastern Austra- P. flavangulus Chevrolat 1863: 599 lia. Distribution. Cuba. Endemic Cuba localities. 113. FAMILY LANGURIIDAE Collections. The lizard beetles P. politus Melsheimer 1846: 102 The Cuban species probably occur in rotten veg- Distribution. Cuba, USA etation. Leschen and Wegrzynowicz (1998) have Cuba localities. IJ: Nueva Gerona reviewed the higher classification of the genera of Collections. Pittsburgh the world.

Stilboides Guillebeau 1894: 282 SUBFAMILY TORAMINAE S. grouvellei Guillebeau 1894: 307 Toramus Grouvelle 1916: 26 Distribution. Cuba, Brazil T. acutus Reitter 1875: 86 Cuba localities. Distribution. Cuba, USA Collections. Cuba localities. Collections. Stilbus Seidleitz 1872: 35 S. univestris Guillebeau 1894: 309 SUBFAMILY XENOSCELINAE Distribution. Cuba. Endemic TRIBE XENOSCELINI Cuba localities. Collections. Hapalips Reitter 1877: 122 H. filum Reitter 1877: 124 Xanthocomus Guillebeau 1893: 291 H. tenuis Reitter 1877: 124 X. floralis Guillebeau 1894: 306 Distribution. Cuba, Grenada, México to Co- Distribution. Cuba. Endemic lombia Cuba localities. Cuba localities. Collections. Collections.

109. FAMILY HOBARTIIDAE TRIBE LOBERINI This family occurs in Australia and South America. Loberus LeConte 1861: 98 L. mutatus Grouvelle 1916: 26 110. FAMILY CAVOGNATHIDAE L. insularis Casey 1900: 83 This family occurs in Australia, New Zealand, Distribution. Cuba, Bahamas, Puerto Rico and Chile. Cuba localities. Collections. 111. FAMILY CRYPTOPHAGIDAE L. parvus Bruce 1954: 173 The silken fungus beetles Distribution. Cuba. Endemic This family generally feeds on molds and de- Cuba localities. PR: Pinar del Rio. LT: caying vegetation. Some occur in animal nests and Hormigero (t.l.) with stored products. Collections. Amsterdam

SUBFAMILY CRYPTOPHAGINAE REFERENCES TRIBE CRYPTOPHAGINI Bruce, N. 1954. Einige Coleoptera (Cryptophagidae) aus Ost- und Westindien und Südafrika im Cryptophagus Herbst 1892: 172 Zoologischen Museum, Amsterdam. Ent. C. acutangulus Gyllenhal 1828: 285 Berichten 15: 172-175. 134 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Leschen, R. A.B. and P. Wegrzynowicz. 1998. REFERENCES Generic catalogue and taxonomic states of Skelley, P. E. 1998. Revision of the genus Ischyrus Languriidae (Cucujoidea). Annales Zoologici Lacordaire (1842) of North and Central America (Warszawa) 48: 221-243. (Coleoptera; Erotylidae; Tritominae). Occ. Pap. Florida State Colln. Arthropods 8: 1-135. 114. FAMILY EROTYLIDAE The pleasing fungus beetles 115. FAMILY Reviewed by Paul Skelley, February 1997 This family lives in North America, Eurasia, This family occurs almost exclusively on the and the Greater Sunda Islands. Some eat fruits of fruiting bodies of higher fungi growing on rottted Rubus and catkins of Quercus. wood. 116. FAMILY BIPHYLLIDAE SUBFAMILY TRITOMINAE The false skin beetles Some members of this family live under bark, Eptyus Dejean 1837: 428 and eat spores or fermentation products. E. brunnipes (Kuhnt) 1910: 231 Distribution. Cuba. Endemic Anobocoelus Sharp 1902: 625 Cuba localities. A. costulatus (Chevrolat) 1863: 615 (Diplocoelus); Collections. Gundlach 1891: 102 E. cyaneus (Duponchel) 1824: 31 Distribution. Cuba. Endemic Oocyanus violaceus (Sturm) 1826: 82; Jacquelin Cuba localities. du Val 1857: 322; Gundlach 1891: 127 Collections. Distribution. Cuba. Endemic Cuba localities. Widespread 117. FAMILY BOTHRIDERIDAE Collections. Gundlach no. 284 The bothriderid beetles E. flavitarsis (Lacordaire) 1842: 130; Jacquelin du The larvae of these beetles are fungal feeders Val 1857: 321 (Ischyrus); Gundlach 1891: 127 or are ectoparasitic on the larvae of wood boring Distribution. Cuba. Endemic insects (Stephan 1989). Cuba localities. Widespread Collections. Gundlach no. 285 SUBFAMILY BOTHRIDERINAE E. gundlachi (Zayas) 1988: 81 (Oocyanus) TRIBE BOTHRIDERINI Distribution. Cuba. Endemic Cuba localities. Type locality not given Bothrideres Erichson 1845: 287 Collections. Gundlach no. 1514 (type) B. dentata Chevrolat 1864: 609; Gundlach 1891: E. tarsalis (Lacordaire) 1842: 106 (Oocyanus) 106; Slipinski et al. 1989: 176 O. tarsatus Lacordaire 1842: 196; Gundlach 1891: Distribution. Cuba, St. Vincent 127 Cuba localities. Distribution. Cuba, Hispaniola Collections. Gundlach no. 311. Paris Cuba localities. Widespread (Grouvelle; Chevrolat) Collections. Gundlach no. 179 B. planus Chevrolat 1864: 610; Gundlach 1891: 106; Slipinski et al. 1989: 178 Ischyrus Lacordaire 1842: 89 Distribution. Cuba, Guadeloupe I. sapphirus Skelley 1998: 58 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 1105. Paris Cuba localities. Ci: Soledad (t.l.). Gu: Baracoa (Grouvelle) Collections. Chicago. Cambridge (type). Washington Lithophorus Sharp 1894: 487 L. gemmatus (Reitter) 1877: 347 (Prolyctus); Tritoma Fabricius 1775: 570 Slipinski et al. 1989: 143 [T. pulchra Say 1826: 301] Distribution. Cuba, Hispaniola (T. sellata Kuhnt 1910: 247) Cuba localities. Distribution. Not Cuba, but widespread east- Collections. Berlin (Humboldt) ern USA A Checklist of the Beetles of Cuba 135

TRIBE DERETAPHRINI Philothermus Aubé 1843: 93 P. parmulus (Chevrolat) 1864b: 600 (Discoloma); Sosylus Erichson 1845: 287 Gundlach 1891: 108 S. costipennis (Jacquelin du Val) 1856: 103 Distribution. Cuba. Endemic (Nematidium); Chevrolat 1864: 609; Gundlach Cuba localities. Widespread 1891: 105 Collections. Gundlach no. 628 Distribution. Cuba, Grenada P. puberulus Schwarz 1878: 361; Lawrence and Cuba localities. Widespread Stephan 1975: 153 Collections. Gundlach no. 537 Distribution. Cuba, Puerto Rico, Hispaniola, se USA; probably not México, Central America REFERENCES Cuba localities. Slipinski, S.A., R.D. Pope, and R.J.W. Collections. Cambridge Aldridge. 1989. A review of the world Bothriderini (Coleoptera: Bothrideridae). Tyrtaeus Champion 1913: 76 Polskie Pismo Ent. 59: 131-202. T. rufus Champion 1913: 17 Stephan, K.S. 1989. The Bothrideridae and Distribution. Cuba, Guatemala Colydiidae of America north of México (Co- Cuba localities. leoptera, Clavicornia and Heteromera). Occ. Collections. Pap. Fla. St. Colln. Arthrop. 6: 1-65. REFERENCES 118. FAMILY CERYLONIDAE Lawrence, J.F. and K. Stephan. 1975. The The minute bark beetles North American Cerylonidae (Coleoptera: These beetles are usually found in leaf litter Clavicornia). Psyche 82: 131-166. and rotten wood, where they probably feed on fungi. Some occur in bat guano in West Indian caves. 119. FAMILY This family occurs only in Eurasia. SUBFAMILY EUXESTINAE 120. FAMILY DISCOLOMATIDAE Euxestus Wollaston 1858: 411 The discolomatid beetles E. erithacus (Chevrolat) 1864b: 599 (Olibrus); Peck These beetles usually occur under bark, and et al. 1998: 160 some may feed on fungi. Distribution. Cuba, Jamaica, Hispaniola, Puerto Rico, Guadeloupe, Grenada, México, SUBFAMILY DISCOLOMATINAE Guatemala, USA (FL), Surinam, Old World Cuba localities. SC: Cueva Atabex, in bat Discoloma Erichson 1845: 242 guano D. cubanum John 1947: 97 Collections. Gatineau Distribution. Cuba. Endemic Cuba localities. SUBFAMILY CERYLONINAE Collections. Eberswalde (type). Hamburg D. parvulum Pascoe 1863: 99; John 1947: 90 Cerylon Latreille 1802: 228 D. erichsoni Reitter 1877: 176 C. amaroides Chevrolat 1864: 610; Gundlach 1891: D. circulare (Chevrolat) 1863: 603 (Thyreosoma); 107 (Cerilon) Gundlach 1891: 93; John 1944: 100; not Cuba; Distribution. Cuba, Grenada but Costa Rica and Panamá Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 660 Cuba localities. M: Cárdenas Collections. Berlin. Dresden (type). Gundlach Mychocerus Erichson 1845: 2192. Casey 1890: no. 315. Hamburg 317 M. cubanus (Hinton) 1936: 18. (Lapethus) REFERENCES Distribution. Cuba. Endemic John, H. 1944. Revision der Gattung Discoloma. Cuba localities. Ci: Cayamas (t.l.) Arb. Morphol. Taxon. Ent. Berlin-Dahlem 11: Collections. Washington (type) 81-108. 136 Arthropods of Florida and Neighboring Land Areas, Vol. 18

121. FAMILY ENDOMYCHIDAE Cuba localities. On citrus at an unspecified The handsome fungus beetles location; Bruner 1951: 84 This family occurs in North, Central, and South Collections. Havana? America. The only previously published West In- D. pusillus (LeConte) 1852: 135 dian records seem to be from the Lesser Antilles. Distribution. Cuba, USA (MA-FL-west to CA) Most species seem to feed on the spores and soft Cuba localities. Ha: Santiago de Las Vegas, tissues of fungi. They may be collected on and un- attacking Aleyrodidae (Bruner 1951: 84) der bark or in leaf litter, and may graze on bark Collections. Havana? surfaces. Some are stored product pests. TRIBE STICHOLOTIDINI SUBFAMILY EUPSILOBIINAE Neaptera Gordon 1991: 309 Eidoreus Sharp 1885: 146 N. cubensis Gordon 1994b: 237 (E. politus Casey 1895: 454 (Eupsilobius)) Distribution. Cuba. Endemic Distribution. Guadeloupe, USA (FL; known Cuba localities. Locality not specified from Keys) Collections. Washington (type) Cuba localities. Collections. Nelasa Gordon 1991: 299 N. cubensis Gordon 1991: 303 REFERENCES Distribution. Cuba. Endemic Pakaluk, J. and S.A. Slipinski. 1990. Review of Cuba localities. PR: Bahia Honda (t.l.) Eupsilobiinae (Coleoptera, Endomychidae) with Collections. Washington (type) descriptions of new genera and species from N. schwarzi Gordon 1991: 303 South America. Rev. Suisse Zool. 97: 705-728. Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) 122. FAMILY COCCINELLIDAE Collections. Washington (type) The lady beetles Reviewed by R. Gordon and N. Vandenberg, Neotina Gordon 1977: 213. Endemic genus November 1994 N. cariba Gordon 1977: 213; 1994b: 236 Most species are predators as adults and lar- Distribution. Cuba. Endemic vae. The most common prey are aphids, mealybugs, Cuba localities. Gu: Baracoa (t.l.) and scale insects. Some Epilachninae feed on leaves Collections. Washington (type) of solanaceous or cucurbitaceous plants, and N. schwarzi Gordon 1994b: 236 Psylloborini are mildew feeders. An important early Distribution. Cuba. Endemic reference on the Cuban fauna is Dimmock (1906). Cuba localities. Ci: Cayamas (t.l.) Fernandez Garcia (1998) lists species in sugar cane Collections. Washington (type) fields. SUBFAMILY SCYMNINAE TRIBE SCYMNILLINI SUBFAMILY STICHOLOTIDINAE TRIBE SERANGIINI Zilus Mulsant 1850: 958 Z. bruneri (Chapin) 1930: 490 (Scymnillodes) Catana Chapin 1940: 266 Distribution. Cuba. Endemic C. clauseni Chapin 1940: 267 Cuba localities. Ha: Santiago de las Vegas Distribution. Cuba (introduced in 1930), Ba- (t.l.) hamas?, Sumatra, Malaya Collections. Washington (type) Cuba localities. Ha: Santiago de las Vegas; Z. caseyi (Chapin) 1930: 492 (Scymnillodes). In Caimito Dimmock (1906) as Bura sp. Collections. Havana. Washington Delphastus violaceus Casey 1924: 170, not Sicard 1922 Delphastus Casey 1899: 111 Distribution. Cuba, Puerto Rico D. pallidus (LeConte) 1878: 400; Chapin 1940: 266 Cuba localities. Ci: Cayamas (t.l.); Simpatia. Distribution. Cuba, USA (FL) ? Cabada Collections. Washington (type) A Checklist of the Beetles of Cuba 137

Z. eleutherae (Casey) 1899: 115 (Scymnillus) Diomus bruneri Chapin 1930: 494. Ci: Cayamas Distribution. Cuba, Bahamas, USA (FL) (t.l.). Washington (type). Havana. London Cuba localities. Ha: Havana Distribution. Cuba. Endemic Collections. Cuba localities. Widespread Z. iris (Chapin) 1930: 491 (Scymnillodes) Collections. Distribution. Cuba. Endemic Cuba localities. Ha: Havana (t.l.) TRIBE HYPERASPIDINI Collections. Washington (type) Z. splendidus (Chapin) 1930: 491 (Scymnillodes) Hyperaspis Redtenbacher 1844: 8 Distribution. Cuba. Endemic H. apicalis Weise 1885: 167 Cuba localities. Widespread. CA: Baraguá H. festivus apicalis Weise (t.l.) Distribution. Cuba, Hispaniola, Puerto Rico Collections. Havana. London. Washington Cuba localities. Or: Jobabo (Bruner 1951: 85) (type) Collections.

TRIBE STETHORINI Thalassa Mulsant 1850: 506 T. flaviceps Mulsant 1850: 513; Jacquelin du Val Stethorus Weise 1885: 65 1857: 326 S. utilis Horn 1895: 107 T. prasina Mulsant 1850: 516: Gundlach 189l: Distribution. Cuba; widely distributed in 403 Caribbean, southern US Distribution. Cuba. Endemic Cuba localities. Cuba localities. Widespread Collections. Collections. Gundlach no. 290. Havana T. pentaspilota Chevrolat 1835: 124 TRIBE SCYMNINI T. pentastigma Mulsant 1850: 512 T. prasina Mulsant, Jacquelin du Val 1857: 327; Decadiomus Chapin 1933: 96 Gundlach 1891: 403 D. bahamicus (Casey) 1899: 159; Chapin 1933: 97; Distribution. Cuba, México, Guatemala Bruner 1951: 84 Cuba localities. Distribution. Cuba, Bahamas, Bermuda, Collections. Gundlach no. 290 USA (FL) Cuba localities. Ci: Cayamas. Ha: San An- TRIBE BRACHIACANTHINI tonio de los Baños; Santiago de Las Vegas Collections. Havana. Washington Brachiacantha Dejean 1837: 458 D. peltatus (Chapin) 1933: 99 B. decora Casey 1899: 119 Distribution. Cuba. Endemic B. bistripustulata Fabricius(in Bruner 1951: 84) Cuba localities. Ha: Santiago de las Vegas Distribution. Cuba, Jamaica, widespread to (t.l.) Brazil, USA (AZ-FL) Collections. Havana. Washington (type) Cuba localities. PR: Sierra Rangel. SC: Santiago de Cuba Diomus Mulsant 1850: 95 Collections. D. ochroderus (Mulsant) 1850: 951 Distribution. Cuba, Puerto Rico, Lesser SUBFAMILY Antilles TRIBE CHILOCORINI Cuba localities. Collections. Arawana Leng 1908: 34. D. roseicollis (Mulsant) 1853: 270 A. cubensis (Dimmock) 1906: 322 (Exochromus); Distribution. Cuba, Puerto Rico, Lesser Gordon 1994c: 133 Antilles, USA (FL) Distribution. Cuba. Endemic Cuba localities. Ha. IJ: Nueva Gerona. M Cuba localities. Collections. Pittsburgh Collections.

Scymnus Kugelann 1894: 545 Chilocorus Leach 1815: 116 S. distinctus Casey 1924: 176 C. cacti Linnaeus 1767: 584 138 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, México to South America Collections. Cuba localities. Ha: Havana Collections. Geodimmockius Chapin 1930: 489; Gordon 1994a: 223. Endemic genus Cladis Mulsant 1850: 1033 G. explanatus Chapin 1930: 490 C. nitidula Fabricius 1792: 286 Distribution. Cuba. Endemic Exochomus uva Mulsant 1850: 479; Jacquelin Cuba localities. SC: San Nicholas (t.l.), on du Val 1857: 326 Coffea arabica Distribution. Cuba, Lesser Antilles Collections. London. Washington (type) Cuba localities. Ha: Havana Collections. TRIBE NOVIINI

Curinus Mulsant 1850: 472 Rodolia Mulsant 1850: 902 C. peleus Mulsant 1853: 191 R. cardinalis (Mulsant) 1850: 906. The vedalia beetle Distribution. Cuba. Endemic Distribution. Cuba (introduced), s USA (CA- Cuba localities. SC); introduced from Australia for biocontrol Collections. purposes Cuba localities. Widespread. Predator on Egius Mulsant 1850: 452 Icerya purchasi (the cottony cushion scale) E. platycephalus Mulsant 1850: 464; Jacquelin du Collections. Val 1857: 325; Gundlach 1891: 402 Distribution. Cuba. Endemic SUBFAMILY COCCINELLINAE Cuba localities. Widespread TRIBE COCCINELLINI Collections. Gundlach no. 964 Coccinella Linnaeus 1758: 36 Exochomus Redtenbacher 1843: 11 C. maculata (DeGeer) 1775: 392; Jacquelin du Val E. marginipennis (LeConte) 1824: 173 1857: 325 (Megilla); Bruner 1951: 84 Distribution. Cuba, USA (NY-FL-TX- KS) Megilla novemnotata cubensis Casey 1908: 394. Cuba localities. Ha: Santiago de Las Vegas; Cuba Víbora (Bruner 1951: 85) C. floridana Leng 1903: 38 (Bruner 1951: 84) Collections. Distribution. Cuba, Lesser Antilles, México to Chile, se USA SUBFAMILY COCCIDULINAE Cuba localities. Widespread TRIBE COCCIDULINI Collections. Gundlach no. 374. Havana. Pitts- (Key to genera in Gordon 1994a) burgh

Botynella Weise 1891: 286 Coelophora Mulsant 1850: 390 B. quadripunctata Weise 1891: 287; Gordon 1994a: C. inaequalis (Fabricius) 1775: 80 223 Distribution. Cuba, Caribbean, USA (FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. B. quinquepunctata Weise 1891: 287; Gordon Cycloneda Crotch 1871: 6 1994a: 223 C. sanguinea Linnaeus 1763: 10 Distribution. Cuba. Endemic C. s. limbifer Casey 1899: 92 (Cuba and Florida Cuba localities. subspecies) Collections. Distribution. Cuba, Bahamas, widespread West Indies, México to Chile, USA Bura Mulsant 1850: 374 Cuba localities. Widespread in Cuba [B. cuprea Mulsant 1850: 374; Dimmock 1906: 287- Collections. Pittsburgh 392] C. n.sp. Zayas 1988: 80 Distribution. Hispaniola Distribution. Cuba. Endemic Cuba localities. Doubtful record ( see Gor- Cuba localities. Ho: Pinares de Mayarí don 1994a: 223) Collections. Zayas A Checklist of the Beetles of Cuba 139

Hipppodamia Mulsant 1846: 30 Cuba localities. Widespread. Pest of cucurb- H. convergens Guérin-Méneville 1842: 321. The con- its vergent lady beetle Collections. Gundlach no. 276. Havana Distribution. Cuba, West Indies, México to Honduras, USA REFERENCES Cuba localities. Common in western Cuba Chapin, E.A. 1957. Records of coccinellid beetles (Bruner 1951: 85) from the Cayman Islands, with descriptions of Collections. new species from the West Indies. Ent. Mon. Mag. 93: 89-91. Olla Casey 1899: 84 Dimmock, G.W. 1906. Algunas Coccinellidae de O. v-nigrum (Mulsant) 1866:64 (Harmonia) Cuba. Inf. An. Est. Centr. Agron., 1904-1905 Procula orientalis Zayas 1988: 80; SC: Santiago (1906): 287-392. de Cuba (t.l.), Zayas (type); Ivie 1991: 400 Fernandez Garcia, I. 1998. Algunos aspectos Distribution. Cuba (widespread), México to ecologicos de la comunidad de coleopteros de un South America, USA (widespread) (dark form agroecosistema cañero en la provincia de La has been called O. oculata Fab., and Coccinella Habana. Poeyana no. 460, 114. oculata Thunberg) Gordon, R.D. 1991. West Indian Coccinellidae IV Cuba localities. (Coleoptera): new genera and species of Collections. Sticholotidini. Proc. Ent. Soc. Washington 93: 298-316. Procula Mulsant 1850: 374 Gordon, R.D. 1994a. West Indian Coccinellidae. P. cubensis Chapin 1941: 167 V. (Coleoptera): a review of Coccidulini and ad- Distribution. Cuba. Endemic ditions to Psorolyma Sicard. J. New York Ent. Cuba localities. Ha: Sierra de Anafe, Caimito Soc. 102: 222-231. (t.l.); Santiago de Las Vegas. Ci: Cayamas Gordon, R.D. 1994b. West Indian Coccinellidae. Collections. Washington (type) VI. (Coleoptera): new genera and species of Sticholotidini and a cladistic analysis of included TRIBE HALYZIINI genera. J. New York Ent. Soc. 102: 232-241. Gordon, R.D. 1994c. Arawana scapularis (Co- Psyllobora Chevrolat 1836: 458 leoptera: Coccinellidae) in the United States and P. nana Mulsant 1850: 181; Jacquelin du Val 1857: new locality records for species of Arawana. Ent. 325; Gundlach 1891: 402; Chapin 1957: 89 News 105: 133-136. Distribution. Cuba, Bahamas, Cayman Is- Ivie, M.A. 1991. Taxonomic notes on a little known lands, Jamaica, Hispaniola to St. Thomas, USA publication: Zayas, 1988, Entomofauna (FL) Cubana. Order Coleoptera. Coleop. Bull. 45: 399- Cuba localities. H. M: Cárdenas. PR 401. Collections. Havana, Gundlach no. 431 P. schwarzi Chapin 1957: 89 123. FAMILY CORYLOPHIDAE Distribution. Cuba, Bahamas, Hispaniola, The minute hooded beetles Cayman Islands These beetles appear to feed on various fungal Cuba localities. Ca: Jaronú. Ci: Cayamas; spores, and some are common on moldy grass cut- Limones; Soledad. Ha: Santiago de la Vega. PR: tings. Chirigota (t.l.) Collections. Cambridge. Washington (type) SUBFAMILY RYPOBIINAE TRIBE RYPOBIINI SUBFAMILY EPILACHNINAE Hoplicnema Matthews 1899: 161 Epilachna Redtenbacher 1843: 7 H. cubensis Pakaluk 1987: 89 E. tredecimnotata (Latreille) 1833: 67. The squash Distribution. Cuba. Endemic beetle Cuba localities. Ci: Cayamas (t.l.); Soledad E. borealis Fabricius, in Jacquelin du Val 1857: Collections. Cambridge. Washington (type) 327; Gundlach 1891: 404 H. hesperia Pakaluk 1987: 93 Distribution. Cuba, México to Argentina, Distribution. Cuba. Endemic USA (widespread) 140 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Ci: Cayamas (t.l.). PR: Ba- Melanophthalma Motschoulsky 1866: 269 hia Honda M. americana (Mannerheim) 1844: 50 Collections. Washington (type) Distribution. Cuba, Argentina, USA (FL) H. spiniventer Pakaluk 1987: 108 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Cayamas (t.l.). PR: Ba- M. flavicula Motschoulsky 1866: 290 hia Honda Distribution. Cuba. Endemic Collections. Cambridge. Washington (type) Cuba localities. Collections. SUBFAMILY CORYLOPHINAE M. signata Belon 1887: 289 TRIBE PARMULINI Distribution. Cuba. Endemic Cuba localities. Clypastrea Haldeman 1842: 191. Collections. C. instabile (Matthews) 1894: 336 (Sacium) Distribution. Cuba, Grenada Metophthalmus Wollaston 1854: 192 Cuba localities. M. cuba Andrews 1998: 60 Collections. Distribution. Cuba. Endemic Cuba localities. Ho: Bahia Barclay (t.l.) REFERENCES Collections. Bozeman. Sacramento. Washing- Pakaluk, J. 1987. Revision and phylogeny of the ton Neotropical genus Hoplicnema Matthews (Co- leoptera, Corylophidae). Trans. Amer. Ent. Soc. REFERENCES 113: 73-116. Andrews, F. 1998. A new species of Metophthalmus Wollaston (Coleoptera: 124. FAMILY LATRIDIIDAE Latriidae) from Cuba. Coleop. Bull. 52: 60-62. The minute brown scavenger beetles All members of this family appear to feed on SUPERFAMILY TENEBRIONOIDEA spores and are often associated with molds. Some 125. FAMILY MYCETOPHAGIDAE are stored products pests. The hairy fungus beetles These beetles mostly feed on fungi on and un- SUBFAMILY LATHRIDIINAE der bark and in moldy plant materials.

Cartodere Thomson 1859: 93 SUBFAMILY MYCETOPHAGINAE (C. constricta (Gyllenhal) 1827: 138). The plaster beetle Litargus Erichson 1846: 405 Distribution. USA (FL), cosmopolitan L. balteatus LeConte 1856: 14 Cuba localities. Distribution. Cuba, Puerto Rico, Montserrat, Collections. México to Panamá, USA (FL); cosmopolitan; occasionally in stored corn and grain Eufallia Muttkowski 1810: 162 Cuba localities. Enicmus Thomson 1859: 93 Collections. E. seminiveus Motschoulsky 1866: 265 Distribution. Cuba, Puerto Rico, México, Typhaea Stephens 1830: 58 USA (FL) T. semirufa Chevrolat 1864: 614; Gundlach 1891: Cuba localities. 102 Collections. Distribution. Cuba, Puerto Rico Cuba localities. M: Cárdenas SUBFAMILY CORTICARIINAE Collections. Gundlach no. 326 (T. stercorea Linnaeus 1758: 357). The hairy fun- Corticaria Marsham 1802: 106 gus beetle C. ferruginea Marshall 1902: 111 Distribution. Puerto Rico, Lesser Antilles, Distribution. Cuba, USA (FL), cosmopolitan México to Guatemala, USA (FL); cosmopolitan; Cuba localities. feeds on fungi in moldy grains, seeds, and pea- Collections. nuts A Checklist of the Beetles of Cuba 141

Cuba localities. Cuba localities. Ci: Cayamas. Gu: Collections. Guantánamo. SS: Buenos Aires; Trinidad Collections. Cambridge. Gundlach no. 725 SUBFAMILY BERGININAE C. pullulus (Casey) 1898: 90; Lawrence 1967: 121; 1971: 501 Berginus Erichson 1846: 405 Distribution. Cuba, Jamaica, Puerto Rico, B. bahamicus Casey 1900: 129 Virgin Islands, se USA (NC-LA) Distribution. Cuba, Bahamas, USA (FL) Cuba localities. Ci: Soledad. SS: Buenos Cuba localities. Aires; Mina Carlotta; Trinidad Collections. Collections. Cambridge C. punctulatus rubriculus Lawrence 1967: 124; 126. FAMILY ARCHEOCRYPTICIDAE Lawrence 1967: 124; 1971: 501 This family is not known to occur in the West Distribution. Cuba, Jamaica, e USA Indies, but it is in the southern USA, and Central Cuba localities. PR: Baños de San Vicente and South America. They are found in leaf litter (the only Cuba record) and may feed on fungi. Collections.

127. FAMILY PTEROGENIIDAE Cis Latreille 1796: 50 This family occurs in southeast Asia and the C. bipartita Jacquelin du Val 1857: 241; Gundlach . They feed on hard polypore fungi. 1891: 109 Distribution. Cuba. Endemic 128. FAMILY CIIDAE Cuba localities. M: Cárdenas The minute tree-fungus beetles Collections. Gundlach no. 394 Both larvae and adults of this family bore in C. cayensis Lawrence 1971: 448 hard fungal fruiting bodies. Lawrence (1971, 1982) Distribution. Cuba, USA (s FL) revised and catalogued the North American gen- Cuba localities. SS: Buenos Aires; Trinidad era and species. Collections. Cambridge C. creberrimus Mellié 1848: 357: Lawrence 1971: SUBFAMILY CIINAE 452 Distribution. Cuba, Jamaica, Bahamas, Ceracis Mellié 1848: 375 (see Lawrence 1967) Grenada, México, Central and South America, C. curtus (Mellié) 1848: 267 (Ennearthron); USA (FL) Gundlach 1891: 110; Lawrence 1967: 110; 1971: Cuba localities. PR: Rinar del Rio 497 Collections. Cambridge Distribution. Cuba, Jamaica, Hispaniola, C. crinitus Lawrence 1971: 453 Puerto Rico, USA (TX-FL) Distribution. Cuba, Jamaica, Bahamas, Cuba localities. Ci: Cayamas; Soledad. IJ Puerto Rico Collections. Cambridge. Gundlach no. 1335. Cuba localities. Ci: Cayamas Washington Collections. Washington C. minutus Dury 1917: 25: Lawrence 1967: 114; C. floridae Drury 1917: 9: Lawrence 1971: 460 1971: 499 Distribution. Cuba, USA (GA-FL) Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Ci: Cayamas Bahamas, USA (TX-FL) Collections. Washington Cuba localities. Ci: Cayamas; Soledad C. hirsutus Casey 1898: 83; Lawrence 1971: 461 Collections. Cambridge. Washington Distribution. Cuba, Jamaica, Hispaniola, C. monocerus Lawrence 1967: 115, replacement Puerto Rico, USA (FL) name for C. unicorne Casey 1898: 90 Cuba localities. PR: Baños de San Vicente Distribution. Cuba, USA (LA-FL) Collections. Cuba localities. C. hirtella Jacquelin du Val 1856: 241; Gundlach Collections. 1891: 109 C. multipunctatus (Mellié) 1848: 368 Distribution. Cuba. Endemic (Ennearthron); Gundlach 1891: 110; Lawrence Cuba localities. 1967: 116; 1971: 499 Collections. Distribution. Cuba, Jamaica, USA (AL-FL) C. murina Mellié 1848: 252; Gundlach 1891: 109 142 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba. Endemic USDA Agriculture Handbook 529-105. Cuba localities. Collections. Gundlach no. 720 129. FAMILY C. superba Kraus 1908: 75 This family occurs in North America, Europe, Distribution. Cuba. Endemic India, the East Indies, and Japan, but not the West Cuba localities. Ci: Cayamas (t.l.) Indies. They occur in the softer fruiting bodies of Collections. Washington (type) some Basidiomycete fungi.

Ennearthron Mellié 1847: 110 130. FAMILY MELANDRYIDAE E. annulatum Kraus 1908: 80 The false darkling beetles Distribution. Cuba. Endemic Most larvae of this family bore in rotten wood, Cuba localities. Ci: Cayamas (t.l.) and some feed in fungal fruiting bodies. Collections. Washington (type) E. castaneipennis (Mellié) 1848: 376 (Ceracis); SUBFAMILY MELANDRYINAE Jacquelin du Val 1857: 246; Gundlach 1891: TRIBE ORCHESIINI 111 Ceracis nitidula Mellié 1848: 334 Orchesia Latreille 1807: 194 Distribution. Cuba, Guadeloupe O. sp. (Leng and Mutchler 1917: 215) Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 579 Cuba localities. E. delicatulum Jacquelin du Val 1857: 244; Collections. New York Gundlach 1891: 110 Distribution. Cuba, Puerto Rico 131. FAMILY MORDELLIDAE Cuba localities. M: Cienaga de Zapata The tumbling flower beetles Collections. Gundlach no. 1027 Reviewed by J.A. Jackman, April, 1995 E. taurulus Jacquelin du Val 1857: 245; Gundlach Larvae usually feed in rotten wood or stems of 1891: 111 herbaceous plants, and a few bore into fungi. Adults Ceracis bison Reitter 1878: 37 are common on flowers. Distribution. Cuba. Endemic Cuba localities. M: Cardenas; M: Cienaga de SUBFAMILY MORDELLINAE Zapata TRIBE MORDELLINI Collections. Gundlach no. 581 E. variabilis (Mellié) 1848: 380 (Ceracis); Jacquelin Mordella Linnaeus 1758: 420 du Val 1857: 246; Gundlach 1891: 111 M. ornata Zayas 1988: 83 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. PR: Soroa (t.l.) Collections. Gundlach no. 580 Collections. Zayas (type) M. scutellaris Fabricius 1801: 123 Orthocis Casey 1898: 84 Distribution. Cuba, Puerto Rico, México to (O. huesanus Kraus 1908: 77; Lawrence 1971: 485) Brazil Distribution. USA (FL, Keys only) Cuba localities. Cuba localities. Collections. Collections. Tomoxia Costa 1854: 8 REFERENCES T. bella Zayas 1988: 84 Lawrence, J.F. 1967. Delimitation of the genus Distribution. Cuba. Endemic Ceracis (Coleoptera: Ciidae) with a revision of Cuba localities. Ha: Santiago de Las Vegas North American species. Bull. Mus. Comp. Zool. (t.l.) 136: 91-144. Collections. Zayas (type) Lawrence, J.F. 1971. Revision of the North Ameri- T. trimaculata Zayas 1988: 85 can Ciidae (Coleoptera). Bull. Mus. Comp. Zool. Distribution. Cuba. Endemic 142: 419-522. Cuba localities. Ho: Sierra Cristal Lawrence, J.F. 1982. A catalog of the Coleoptera Collections. of America north of México. Family Ciidae. A Checklist of the Beetles of Cuba 143

TRIBE MORDELLISTENINI M. sordidum Gerstaecker 1855: 28 Distribution. Cuba, Puerto Rico, Guatemala Mordellistena Costa 1854: 16 to Brazil M. corralillo Zayas 1988: 87 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities.: Corralillo (t.l.) (M. spinosum Fabricius 1798: 126) Collections. Zayas (type) Distribution. West Indies, Brazil, Argentina M. crucina Zayas 1988: 88 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Gr: Cabo Cruz (t.l.) Collections. Zayas (type) SUBFAMILY RHIPIDIINAE M. graciliformis Ray 1941: 302 Distribution. Cuba. Endemic Ripidius Thunberg 1806: 5 Cuba localities. Ca: Central Jaronu. CA: R. pectinicornis Thunberg 1806:5; Falin: 2001: 196 Central Jagueyal. M: Matanzas; Coliseo (t.l.) Distribution. Cuba (introduced), USA (FL, Collections. Washington (type) HI; intercepts); tropicopolitan, native to India? M. nigricans Melsheimer 1840: 313 Cuba localities. Ha: Havana (intercept on Distribution. Cuba, widespread USA ship from Havana to Miami, FL); of Cuba localities. IJ: Nueva Gerona Blatella germanica L. cockroaches Collections. Pittsburgh Collections. Washington? M. siboney Zayas 1988: 86 Distribution. Cuba. Endemic REFERENCES Cuba localities. SC: Cuidamar (t.l.) Falin, Z. H. 2001. Notes on the occurrence of Collections. Zayas (type) Ripidius pectinicornis Thunberg (Coleoptera: M. tetramaculata Zayas 1988: 86 Rhipiphoridae) in the continental United States Distribution. Cuba. Endemic and Hawaii. Coleop. Bull. 55: 194-197. Cuba localities. SC: Gran Piedra; Loma del Vaurie, P. 1955. A review of the genus Gato (t.l.). SS: Buenos Aires Macrosiagon in México, with notes on Collections. Zayas (type) Rhipiphorus (Coleoptera: Rhipiphoridae). Amer. Mus. Novitates no. 1717. 19 pp. 132. FAMILY RHIPIPHORIDAE The wedge-shaped beetles 133. FAMILY COLYDIIDAE The larvae are endoparasites on larvae of bees The cylindrical bark beetles or colydiid beetles and wasps. These beetles occur in dead wood, under bark, in leaf litter, and sometimes on fungi. Some are SUBFAMILY RHIPIPHORINAE predators on wood-boring beetles, and some feed on TRIBE MACROSIAGONINI fungus spores and hyphae (Slipinski and Lawrence 1997). The higher classification is used differently Macrosiagon Hentz 1830: 462 (see key in Vaurie in different sources (Ivie and Slipinski 1990). The 1955) following are placed in Zopheridae in Slipinski and M. basale Gerstaecker 1855: 21 Lawrence (1999). Distribution. Cuba, Puerto Rico Cuba localities. TRIBE ACROPINI Collections. M. cruentum Germar 1824: 168 Ethelema Pascoe 1860: 107 Distribution. Cuba, México, USA (CA-IN-FL) E. cubana Zayas 1988: 78 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. M: Peninsula Zapata. PR. M. discicolle (Gerstaecker) 1855: 32 SC: Pico Turquino (t.l?) Distribution. Cuba, Puerto Rico, Guadeloupe, Collections. Zayas (type) Hispaniola, Jamaica, México to Brazil Cuba localities. TRIBE ADIMERINI Collections. 144 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Monoedus Horn 1882 Bitoma Herbst 1793: 25 (M. guttatus LeConte 1882: 116; Stephan 1989: 61) B. carinata LeConte 1863: 68 Distribution. West Indies, USA (FL; in Distribution. Cuba, Hispaniola, USA (MS- Metastelma (= Cynanchum) milkweed) VA-FL) Cuba localities. Cuba localities. Collections. Collections. B. discolor Schaeffer 1907: 138 TRIBE COLYDIINI Distribution. Cuba, USA (FL) Cuba localities. Ci: Cayamas Aulonium Erichson 1845: 275 Collections. Washington A. bidentatum (Fabricius) 1801: 556; Jacquelin du B. quinquecarinata (Chevrolat) 1864: 609 Val 1857: 248; Chevrolat 1864: 607; Gundlach (Eulachus); Gundlach 1891: 105 1891: 105 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico, Guadeloupe, Cuba localities. Gr: Brazo de Cauto. SC: México to Brazil Cobre Cuba localities. Widespread Collections. Gundlach no. 1889 Collections. Gundlach no. 664 B. semifuliginosa (Chevrolat) 1864: 608 (Eulachus); Gundlach 1891: 104 Colydium Fabricius 1792: 459 Distribution. Cuba, Puerto Rico C. thomasi Stephan 1989: 57 Cuba localities. Widespread Distribution. Cuba, USA (FL) Collections. Gundlach no. 312 Cuba localities. Ci: Cayamas B. undata Guérin-Méneville 1844: 194 Collections. Washington Distribution. Cuba, Puerto Rico Cuba localities. TRIBE NEMATIDIINI Collections.

Nematidium Erichson 1845: 275 Lobogestoria Reitter 1878: 31 N. cylindricum Fabricius 1801: 557 L. gibbicollis Reitter 1878: 32; Schaeffer 1907: 141 Distribution. Cuba, Colombia, Brazil Aditoma bifida Casey 1897: 630 Cuba localities. Distribution. Cuba, Puerto Rico, se USA (LA- Collections. SC-FL) Cuba localities. TRIBE SYNCHITINI Collections.

Acolobicus Sharp 1894: 452 Microprius Fairmaire 1869: 779 A. erichsoni (Reitter) 1877: 133 M. rufulus (Motschulsky) 1863: 502 (Bitoma) Distribution. Cuba, México, USA (SC, FL) M. cubanus Slipinski 1985: 811; Ivie et al. 2001: Cuba localities. 186 (synonomy) Collections. Distribution. Cosmotropical?; Cuba, Grand Cayman, Puerto Rico, Sri Lanka, Cape Verde Asynchita Sharp 1894: 448 Islands, Egypt, Tanzania, India to Syria, South A. granulata (Say) 1827: 266 Africa,USA (CA). Probably transported by com- Distribution. Cuba, Hispaniola, Puerto Rico, merce in logs Guadeloupe Cuba localities. Ha: Marianao (t.l.); La Cuba localities. Palma. PR: Sierra del Rosario Collections. Collections. Berlin. Geneva. Prague (type). A. rugulosa (Guérin-Méneville) 1844: 189 Warsaw (Synchita) Colobicus rugulosus (Guérin-Meneville); Neotrichus Sharp 1885: 60 Jacquelin du Val 1854: 247; Chevrolat 1864: 607; N. tuberculatus (Chevrolat) 1864b: 607 (Plagiope); Gundlach 1891: 103 Gundlach 1891: 103 Distribution. Cuba, West Indies (where?) Distribution. Cuba, Puerto Rico Cuba localities. Widespread Cuba localities. Widespread Collections. Gundlach no. 70 Collections. Gundlach no. 1170 A Checklist of the Beetles of Cuba 145

Paha Dajoz 1984: 155 Collections. P. laticollis (LeConte) 1863: 66 Distribution. Cuba, USA (OK-NY-FL) Hyporhagus Thomson 1860: 25 Cuba localities. H. marginatus (Fabricius) 1792: 506 (Tritoma); Collections. Jacquelin du Val 1857: 142; Gundlach 1891: 89, 119 (Monomma) Phloeonemus Erichson 1845: 258 Distribution. Cuba, Bahamas, Hispaniola, P. haroldi Reitter 1878: 114 Puerto Rico, South America Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Collections. Gundlach no. 448. Pittsburgh Collections. REFERENCES REFERENCE Slipinski, S.A. and J.F. Lawrence. 1999. Phy- Ivie, M.A. and Slipinski, S.A. 1990. Catalog of logeny and classification of Zopheridae sensu the genera of world Colydiidae (Coleoptera). novo (Coleoptera: Tenebrionoidea) with a re- Annales Zoologici (Warsaw) 43 (suppl. 1) 1 32 view of the genera of Zopherinae (excluding Ivie, M.A., S.A. Slipinski, and P. Monommatini). Annales Zoologici (Warszawa) Wegrzynowicz. 2001. New records and syn- 49: 1-53. onyms in the Colydiinae and Pycnomerini (Co- leoptera: Zopheridae). Insecta Mundi 15: 185- 135. FAMILY ZOPHERIDAE 188. The ironclad beetles Slipinski, S.A. 1985. New and little known spe- Members of the Zopherinae group were tradi- cies of the genus Microprius Fairmaire (Co- tionally placed within the Tenebrionidae. The lar- leoptera, Colydiidae). Polskie Pismo Ent. 55: vae feed in rotten wood or fungus fruiting bodies. 811-816. The Pycnomerini were traditionally placed in the Slipinski, S.A. and Lawrence, J.F. 1997. Gen- Colydiidae. ). The higher classification is used dif- era of Colydiinae (Coleoptera: Zopheridae) of the ferently in different sources. The following are placed Australo-Pacific region. Annales Zoologici in Zopheridae in Slipinski and Lawrence (1999). (Warszawa) 47: 341-440. Slipinski, S.A. and Lawrence, J.F. 1999. Phy- TRIBE PYCNOMERINI logeny and classification of Zopheridae sensu novo (Coleoptera: Tenebrionoidea) with a re- Pycnomerus Erichson 1842:214 view of the genera of Zopherinae (excluding P. biimpressus Reitter 1877: 355; Ivie and Slipinski Monommatini). Annales Zoologici (Warszawa) 1989: 69 49: 1-53. Cerylon exaratum Chevrolat 1863: 610; Gundlach Stephan, K.S. 1989. The Bothrideridae and 1891: 107 Colydiidae of America north of México (Co- Distribution. Cuba, Jamaica, Hispaniola, leoptera, Clavicornia and Heteromera). Occ. Puerto Rico, Lesser Antilles, Central and South Pap. Fla. St. Colln. Arthrop. 6: 1-65. America Cuba localities. SC: Gran Piedra 134. FAMILY MONOMMATIDAE Collections. Cambridge The monommatid beetles P. valentinei Ivie and Slipinski 1989: 76 These beetles occur in decaying plant material, Distribution. Cuba, Hispaniola and may occur in rotting stems of Euphorbia, Pa- Cuba localities. Ci: Soledad (record based on paya, and Yucca, under fermenting tree bark, and single specimen) in rotted yams. This group is treated as a tribe of Collections. Cambridge Zopheridae in Slipinski and Lawrence (1999). TRIBE ZOPHERINI TRIBE MONOMMATINI Meralius Casey 1907: 470 (known only in Cuba Aspathines Champion 1888: 475 and Venezuela) A. aenea Thomson 1860: 27 M. echinatus (Guerin) in Chevrolat 1838: 280 Distribution. Cuba, Hispaniola, Puerto Rico (Nosoderma); Jacquelin du Val 1857: 141; Cuba localities. Gundlach 1891: 118 146 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba. Endemic Tenebrionidae (s.str.) of the Antilles and a supple- Cuba localities. Widespread ment to the catalogue. In an unpublished check- Collections. Gundlach no. 108 list, Garrido and Gutierrez list 125 species for Cuba (excluding Lagriinae and Alleculinae), of which 66 REFERENCES are endemic species, figures which are now out of Ivie, M.A. and S.A. Slipinski. 1989. The date. They give additional distributional data within Pycnomerini (Coleoptera; Colydiidae) of the Cuba, based upon material in the Garrido collec- West Indies. Fla. Ent. 72: 64-80. tion in Havana. Slipinski, S.A. 1985. New and little known spe- cies of the genus Microprius Fairmaire (Co- SUBFAMILY LAGRIINAE leoptera, Colydiidae). Polskie Pismo Ent. 55: TRIBE BELOPINI 811-816. Slipinski, S.A. and J.F. Lawrence. 1999. Phy- Adelonia Laporte 1840: 221 logeny and classification of Zopheridae sensu A. sulcatulus Champion 1885: 121; Ardoin 1977b: novo (Coleoptera: Tenebrionoidea) with a re- 391 view of the genera of Zopherinae (excluding A. sulcatus Leng and Mutchler 1914: 464 Monommatini). Annales Zoologici (Warszawa) Distribution. Cuba (Oriente), México to Co- 49: 1-53. lombia, Ecuador, USA (TX); at lights Stephan, K.S. 1989. The Bothrideridae and Cuba localities. SC: Siboney Colydiidae of America north of México (Co- Collections. Havana (Garrido) leoptera, Clavicornia and Heteromera). Occ. Pap. Fla. St. Colln. Arthrop. 6: 1-65. Rhypasma Pascoe 1862: 325 Derosimus Fairmaire 1904: 62 136. FAMILY R. haitianum Marcuzzi 1953: 82: Spilman 1961b: This family occurs in Australia, New Zealand, 127 and Chile. Distribution. Cuba, Hispaniola Cuba localities. Ca: SC: Cuabitas. Gu: 137. FAMILY PERIMYLOPIDAE Guantánamo. LT: Jobabo. SC: Santiago This family is restricted to southern South Collections. Washington. Zayas America and the Subantarctic Islands. They live under stones and in grass and may be general scav- SUBFAMILY PHRENAPATINAE engers. TRIBE PENETINI

138. FAMILY CHALCODRYIDAE Dioedus LeConte 1862: 238 This family occurs only in Tasmania and New Arrhabaeus Champion 1886: 144 Zealand. They apparently feed on moss on tree D. sp.; 5 species (undescribed) trunks at night. Distribution. Cuba localities. 139. FAMILY TRACHELOSTENIDAE Collections. This family of one genus is limited to Chile. SUBFAMILY PIMELIINAE 140. FAMILY TENEBRIONIDAE TRIBE BRANCHINI The darkling beetles This is a large and common family. They gen- Branchus LeConte 1862: 222 erally occur in arid areas and feed on a variety of B. cubensis Marcuzzi 1998: 153 (nomen nudum?) materials as scavengers. Larvae occur in soil, lit- Distribution. Cuba. Endemic ter, dead wood, or fungi. Some occur in caves and Cuba localities. some are pests of stored products or agriculture Collections. (La Rosa et al. 1987, Vasquez 1986). The cave fau- B. woodi LeConte 1866: 111; Spilman 1961b: 127 nas have been studied by Ardoin (1977a, 1977b), Distribution. Cuba, Bahamas and Peck et al. (1999) The higher categories and Cuba localities. Ha: Marianao, littoral genera are here arranged according to the concepts Collections. Washington. Zayas of relationships used by Aalbu et al. in Arnett et al. (2002). Marcuzzi (1984, 1998) is a catalogue of the A Checklist of the Beetles of Cuba 147

TRIBE EPITRAGINI T. garridoi Marcuzzi 1998: 153 (replacement name) Epitragus Latreille 1804: 322 Distribution. Cuba. Endemic E. arulentus Kirsch 1866: 189: Freude 1967: 156 Cuba localities. SC; Cueva de Cativar, 50 km Distribution. Cuba (introduced, probably W Santiago (t.l.), other caves in Gu, Gr and SC native to South America), Jamaica, , along coast; in dry guano in caves Curaçao, Margarita, México to Colombia, Ven- Collections. Budapest. Warsaw ezuela, Brazil, Ecuador, USA (TX); at lights; T. escambrayensis Garrido and Gutierrez 1997b: apparently becoming an agricultural pest spe- 33 cies in the West Indies (J. Santiago Blay, pers. Distribution. Cuba. Endemic comm., 2002) Cuba localities. SS: Sierra de Escambray, Cuba localities. SC: Versalles. Ca: Camaguey Cima de Pico Potrerillo Collections. Havana: Garido Collections. Havana E. n.sp.? Zayas 1988: 90 (which notes it is different T. littoralis Garrido and Gutierrez 1997b: 30 from above species, and is possibly introduced Distribution. Cuba. Endemic from Jamaica, which would suggest it could be Cuba localities. Gu: 1 km E Tortuguilla (t.l.) E. jamaicensis Champion 1896: 3; Freude 1967: Collections. Havana 159 T. minor Garrido and Gutierrez 1997b: 36 Distribution. Distribution. Cuba. Endemic Cuba localities. Cuba localities. Ha: Boca de Jaruco (t.l.) Collections. Zayas Collections. Havana T. negreai Ardoin 1977b: 387; Garrido and TRIBE EURYMETOPINI Gutierrez 1997b: 30 Distribution. Cuba. Endemic Garridoa Marcuzzi 1985: 180 Cuba localities. SC: Cueva del Guano (t.l.); G. kassabi Marcuzzi 1985: 180 Cueva de los Murcielagos, Cueva de la Patana, Distribution. Cuba. Endemic Cueva de Golondrinas; Cueva de la Virgen Cuba localities. LT: Puerto Manati (t.l.) Collections. Collections. Budapest (type) T. poeyi Ardoin 1977a: 383; 1977b: 388; Garrido and Gutierrez 1997b: 30 Trimytantron Ardoin 1977a: 381 Distribution. Cuba. Endemic Bielawskia Marcuzzi 1985: 179; 1998: 152 Cuba localities. SC: Cueva de la Patana (t.l.), T. armasi Garrido and Gutierrez 1997b: 32 and other caves in Ha, SS, SC provinces Distribution. Cuba. Endemic Collections. Cuba localities. SS: Sierra de Trinidad; T. punctulaticeps Garrido and Gutierrez 1997b: 34 Maisinicú (t.l.) Distribution. Cuba. Endemic Collections. Havana Cuba localities. Matanzas or Trinidad (no T. cavernicolous Garrido and Gutierrez 1997b: 34 type locality) Distribution. Cuba. Endemic Collections. Gundlach no. 1073 Cuba localities. SS: Cueva el Pirata, (t.l.), T. pumilus Garrido and Gutierrez 1997b: 35 Cueva de Colon; Cueva de los Chivos (all on Distribution. Cuba. Endemic Cayo Caguanes); Punta de Judas, Yaguajay Cuba localities. Gu: Navas (t.l.) Collections. Havana (type) Collections. Havana T. cubanum Ardoin 1977b: 388; Garrido and T. sierrae Garrido and Gutierrez 1997b: 31 Gutierrez 1997b: 30 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Bonega Matías (t.l.) Cuba localities. Ho: Cueva de los Panaderos Collections. Havana (t.l.) T. vinai Ardoin 1977b: 388 Collections. Distribution. Cuba. Endemic T. decui Ardoin 1977a: 382; 1977b: 388; Garrido Cuba localities. M: Cueva de la Pluma (t.l.) and Gutierrez 1997b: 30; Marcuzzi 1998: 155 Collections. (Diastolinus) Bielawskia cubana Marcuzzi 1985: 179 (SC: Ocujal del Turquino (t.l.), Warsaw (type)) 148 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE TRIENTOMINI Phaleria Latreille 1802: 162 P. picipes Say 1824: 280; Triplehorn and Watrous Trientoma Solier 1835: 256 1979: 291; Watrous and Triplehorn 1982: 19; T. cayensis Garrido and Gutierrez 1995b: 47 Triplehorn 1991; 266 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Jamaica, Cay- Cuba localities. Ca: Archipélago de Sabana; man Islands, Hispaniola, México, Belize, Hon- Cayo Romano; Cayo Conuco duras, Venezuela, Les Saintes, Aves, Mustique, Collections. Havana. Grenadines, Marguerita, Las Roques, USA (FL- (T. convexipennis Allard 1883: 14) NJ) Distribution. Cuba?. Locality not stated in Cuba localities. Gu: Baracoa description Collections. Cuba localities. P. punctipes LeConte 1878: 421; Triplehorn and Collections. Watrous 1979: 281; Watrous and Triplehorn T. garridoi Marcuzzi 1988: 69 1982: 13; Triplehorn 1991: 268 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Jamaica, Vir- Cuba localities. Gu: Bahia de Guantánamo; gin Islands, Lesser Antilles, USA (FL) Tortuguilla (t.l.) Cuba localities. Ca: Camaguey Collections. Havana (Garrido) Collections. Washington T. kaszabi Marcuzzi 1985: 181 P. testacea Say 1824: 280; Triplehorn and Watrous Distribution. Cuba. Endemic 1979: 289; Watrous and Triplehorn 1982: 19; Cuba localities. SC: Puerto Boniato (t.l.) Triplehorn 1991: 266 Collections. Warsaw (type). Distribution. Cuba, Bahamas, Puerto Rico, T. maisiensis Marcuzzi 1988: 67 Virgin Islands, Lesser Antilles, ne USA to Ar- T. zayasi Marcuzzi 1988: 68. Synonomy in gentina Garrido and Gutierrez 1995b Cuba localities. SC: Baconao Distribution. Cuba. Endemic Collections. Cuba localities. Gu: Guantánamo; (P. thinophila Watrous and Triplehorn 1982: 15) Tortuguilla; Punta de Maisi (t.l.) Distribution. Hispaniola, Jamaica, Virgin Collections. Havana (Garrido) Islands, Puerto Rico, Barbados, Costa Rica, T. siboneyensis Marcuzzi 1988: 71 Venezuela Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Siboney (t.l.) Collections. Collections. Havana (Garrido, type) T. varvasi Solier 1835: 257 TRIBE Distribution. Cuba, Puerto Rico SUBTRIBE ADELININA Cuba localities. Collections. Adelina LeConte 1851: 149. T. zayasi Marcuzzi 1988: 70 A. bidens Schaeffer 1915: 238; Ardoin 1977c: 10 Distribution. Cuba Endemic (Doliema) Cuba localities. Gu: Guantánamo (t.l.) Distribution. Cuba, Hispaniola, Guatemala, Collections. Zayas (type) USA (FL) Cuba localities. SUBFAMILY BOLITOPHAGINAE Collections. TRIBE BOLITOPHAGINI A. pallida Say 1823: 271 SUBTRIBE RHIPIDANDRINA Distribution. Cuba, México to Nicaragua, Puerto Rico, USA (FL) Rhipidandrus LeConte 1866: 236 Cuba localities. R. sulcatus Gorham 1898: 333 Collections. Distribution. Cuba, Hispaniola, St. Vincent A. pici (Ardoin) 1977c: 7 (Doliema) Cuba localities. Distribution. Cuba, Guadeloupe, Venezuela Collections. Cuba localities. “In a ship originating in Cuba.” SUBFAMILY Collections. Paris TRIBE PHALERIINI A Checklist of the Beetles of Cuba 149

A. plana (Olivier) 1795: 94 (Tenebrio); Jacquelin D. hydni Fabricius 1801: 585; Jacquelin du Val du Val 1857: 151; Gundlach 1891: 113; Ardoin 1857: 147; Gundlach 1891: 119 1977c: 3 (Doliema) Distribution. Cuba, Bahamas, Jamaica, Distribution. Cuba, Hispaniola, Mustique, Hispaniola, Puerto Rico, St. Croix, México to Tres Marias, Guadeloupe, México to Colombia, Panamá, widespread eastern USA Venezuela, Brasil, Chile, USA (CA-AZ-CO-FL) Cuba localities. Ci: Cienfuegos. Gr: Upper Cuba localities. Yara Valley. IJ. PR: Baños de San Vicente Collections. Gundlach no. 1318 Collections. Gundlach no. 732

Gnatocerus Thunberg 1814: 47 Heterophylus Klug 1833: 20 G. cornutus (Fabricius) 1798: 51. The broad-horned H. meszarosi Kaszab 1977: 127 flour beetle Distribution. Cuba. Endemic Distribution. Cuba, México to Brazil, USA; Cuba localities. PR: Soroa (t.l.) cosmopolitan; carried by commerce to Old World Collections. Budapest (type) in stored products Cuba localities. Neomida Latreille 1829: 29 Collections. N. armata (Laporte and Brullé) 1831: 345; G. maxillosus (Fabricius) 1801: 155; Jacquelin du Jacquelin du Val 1857: 146 (Hoplocephala); Val 1857: 150; Gundlach 1891: 114. The slen- Gundlach 1891: 123 der-horned flour beetle Distribution. Cuba, French Guyana Distribution. Cuba, Grenada, México to Co- Cuba localities. M: Cardenas lombia, USA; cosmopolitan; in stored products Collections. Gundlach no. 235 Cuba localities. Ha: Havana. M: Cárdenas N. bicornis Fabricius 1777: 215 Collections. Gundlach no. 325 Hoplocephala cornigera (Fabricius.) 1781: 82; G. guatemalensis (Champion) 1886: 147 (Sicinus) Jacquelin du Val 1857: 146; Gundlach 1891: Distribution. Cuba, Central America, USA 122; Marcuzzi 1962: 38; Triplehorn 1965: 377 (FL) Distribution. Cuba, Jamaica, Bahamas, Ber- Cuba localities. Ca: Nuevitas; Playa Santa muda, USA (TX-MN-MA-FL) Lucía Cuba localities. Ci: Cienfuegos. Ha: Havana. Collections. IJ. M: Cienaga Zapata. PR: Guane-Teneria; Sierra de Los Organos; San Vicente. SC: Buenos Iccius Champion 1886: 147 Aires; Pico Turquino. VC: Manicaragua I. cephalotes Champion 1886: 148; Marcuzzi 1962: Collections. Basel (Frey). Gundlach no. 235 38 (sic) Distribution. Cuba, México, Guatemala N. ferruginea LeConte 1866: 128; Triplehorn 1965: Cuba localities. PR: Pinar del Rio 384 Collections. Leiden? Distribution. Cuba, Hispaniola, México, Gua- temala, Honduras, USA (TX-AL-FL) Sitophagus Mulsant 1854: 264 Cuba localities. Ci: Soledad S. hololeptoides (Laporte) 1840: 220 (Uloma); Collections. Cambridge Jacquelin du Val 1857: 149 (Hypogena); Gundlach 1891: 121 Loxostethus Triplehorn 1962: 504 (as Cyphiscus Distribution. Cuba, Puerto Rico, Guadeloupe, Chevrolat in Gundlach collection) México to Panamá, South America, Madeira; a L. fasciatus Triplehorn 1962: 504; Triplehorn and stored products pest Merkl 1997: 738 Cuba localities. Widespread L. quadrimaculata Zayas 1988: 93 (Sierera de Collections. Gundlach no. 184 Trinidad (t.l.), Zayas (type); synonomy by Ivie 1991: 400; Garrido and Gutierrez 1995a: 7 SUBTRIBE DIAPERINA Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.); Diaperis Geoffroy 1862: 337 Cumanayagua. Gu: Mts. N of Imias. Ha: Ja- D. maculata Olivier 1791: 273; Triplehorn 1965: maica; Jaruco; Santiago de las Vegas. SS: 367 Buenos Aires; Sierra de Trinidad 150 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Cambridge. Chicago. Columbus. Collections. Gundlach no. 1231 Washington (type) P. excavatum (Say) 1823: 267; Triplehorn 1965: L. gibbosus Triplehorn and Merkl 1997: 738 395 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola, Puerto Rico, Cuba localities. SC: Pico Turquino., 3000- Isla Roatan, Jamaica, México to Panamá, South 5000 ft. (t.l.) America, widespread eastern USA Collections. Budapest. Columbus. Harvard Cuba localities. Ci: Cienfuegos. Gu: Mts. N (type) of Imias L. gowdeyi (Pic) 1930: 30 (Pentaphyllus); Collections. Triplehorn and Merkl 1997: 739 P. flexuosum Chevrolat 1877: 178 L. opacifrons Triplehorn 1962: 506 (Ci: Cayamas; Distribution. Cuba. Endemic t.l., Washington, type); synonomy in Triplehorn Cuba localities. and Merkl 1997: 739 Collections. Heterophylus mesarosi Kaszab 1977: 123 (PR: P. micans Zimmerman, in Horn 1870: 383; Soroa, t.l., Budapest, type); synonomy in Triplehorn 1965: 430 Triplehorn and Merkl 1997: 739 Distribution. Cuba, Jamaica, Hispaniola, L. baracoae Garrido and Gutiérrez 1995a: 7 (Gu; México, Guyana, Brazil, se USA Yunque de Baracoa, t.l.), Havana, type), Cuba localities. Ci: Cienfuegos synonomy by Triplehorn and Merkl 1997: 739 Collections. Distribution. Cuba, Hispaniola, Jamaica P. nigratum (Motschoulsky) 1873: 478 (Neomida); Cuba localities. Ci: Soledad. Ha: Jamaica. Triplehorn 1965: 421 Gu: Mts. N of Imias. SC: Pico Turquino. SS: Distribution. Cuba, México, Belize to Costa Trinidad Mts, SS: Buenos Aires Rica, USA (AZ-TX-AL-FL) Collections. Cambridge. Chicago. Columbus Cuba localities. CA: Baraguá. Ci: Cienfuegos Collections. Platydema Laporte and Brullé 1831: 350 P. picicorne (Fabricius) 1792: 498 (Mycetophagus); P. antennatum Laporte and Brullé 1831: 366; Jacquelin du Val 1857: 145 Jacquelin du Val 1857: 144; Gundlach 1891: Distribution. Cuba, Hispaniola, Puerto Rico 121 Cuba localities. Distribution. Cuba, Hispaniola Collections. Cuba localities. M: Cárdenas P. punctacostriatum Chevrolat 1877: 178 Collections. Gundlach no. 183 Distribution. Cuba. Endemic P. apicale (Laporte and Brullé) 1831: 359 (Lelegis); Cuba localities. Gundlach 1891: 122; Triplehorn 1962: 503 Collections. Distribution. Cuba, Puerto Rico P. virens Laporte and Brullé 1831: 391; Jacquelin Cuba localities. Ci: Cayamas. Gu: Imias; du Val 1857: 144; Gundlach 1891: 122 Upper Ovando and Rio Frio. SC: Boniato P. flavicorne Chevrolat 1877: 170 Collections. Cambridge. Gundlach no. 1317. Distribution. Cuba, Puerto Rico Washington Cuba localities. P. basicorne Chevrolat 1877: 178 Collections. Gundlach no. 1128 Distribution. Cuba. Endemic Cuba localities. Serrania Garrido 2003: 50. Collections. S. viridula (Zayas) 1988: 92 (Diaperis); Ivie 1991: P. dichrocerum Triplehorn 1962: 502 400; (misidentification); Garrido 2003: 50. Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Trinidad, Mina Carlota Cuba localities. Ci: Soledad. Ma: Cárdenas. (t.l.) PR: Sierra del Rangel. SC: El Cardero, Pico Collections. Harvard (type) Turquino (t.l.). VC: Corralillo P. diophthalma Laporte and Brullé 1831: 383; Collections. Garrrido. Zayas (type) Jacquelin du Val 1857: 143; Gundlach 1891: 121 Ulomoides Blackburn 1888: 274 Distribution. Cuba, Isla Roatan, México to U. ocularis (Casey) 1891: 65 (Palembus); Triplehorn Panamá 1965: 388 Cuba localities. Gu: Yateras A Checklist of the Beetles of Cuba 151

Distribution. Cuba, Jamaica, Puerto Rico, Blapstinus Latreille 1829: 21 Guadelupe, Monserrat, St. Martin, Bahamas, B. cubanus Marcuzzi 1962: 33; Ardoin 1977b: 390 USA (FL) Distribution. Cuba. Bahamas Cuba localities. Ha: Mariel (intercept on boat Cuba localities. Widespread in Tampa, FL) Collections. Amsterdam. London. Trieste Collections. Washington (type) B. decui Ardoin 1977b: 390 SUBFAMILY HYPOPHLOEINAE Distribution. Cuba. Endemic TRIBE HYPOPHLOEINI Cuba localities. IJ: Punta Pedernales (t.l.), under stones. SC; Siboney, at lights Corticeus Piller and Mitterpacher 1883: 87 (see Collections. Triplehorn 1979; 1990) B. fortis LeConte 1878: 420; Marcuzzi 1962: 32 C. rufipes (Fabricius) 1801: 558; Marcuzzi 1962: Opatrinus punctulatus Dejean, Jacquelin du Val 38 (Hypophloeus) 1857: 141; Gundlach 1891: 118 Distribution. Cuba, México to Brazil, Puerto Distribution. Cuba, Bahamas, México to Rico, Guadeloupe Panamá, USA (TX, FL) Cuba localities. ? Cacarajicara Cuba localities. IJ: Punta Pedernales. PR: Collections. Utrecht Pinar del Rio. SS: Cueva Colón; Cueva Grande C. tensicollis Triplehorn 1979: 46 de Caguanes Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Gundlach no. 368 (O. USA (FL). punctatulus) Cuba localities. Ci: Cayamas; Soledad. Gu: B. punctatus Fabricius 1792: 109; Marcuzzi 1962: Upper Ovando River. SS: San Juan de los 36 Remedios Distribution. Cuba, Puerto Rico, St. Thomas, Collections. Cambridge. New York. Washing- St. John ton Cuba localities. IJ: Los Indios C. tuberculatus Triplehorn 1979: 48 Collections. Munich. Pittsburgh Distribution. Cuba. Endemic B. puncticeps Mulsant and Rey 1859: 118; Marcuzzi Cuba localities. Ci: Cayamas (t.l.) 1985: 184 Collections. Columbus. Washington (type) Distribution. Cuba. Endemic Cuba localities. SUBFAMILY TOXICINAE Collections. TRIBE TOXICINI Diastolinus Mulsant and Rey 1859: 138 Wattius Kaszab 1982: 50 D. alfaroi Garrido and Gutierrez 1996: 228 W. cucullatus (Pascoe) 1871: 349 (Calimnus); Distribution. Cuba. Endemic Spilman 1961b: 127 (Calymmus) Cuba localities. Gu: Sierra de la Conasta; C. asperulus Pascoe 1871: 350 Loma la (t.l.) Distribution. Cuba, México to Argentina Collections. Havana Cuba localities. Ca: Camaguëy. Ci: D. armasi Marcuzzi 1988: 72 Cumanayagua. PR: Guanahacabibes Distribution. Cuba. Endemic Collections. Washington. Zayas Cuba localities. SC: Santiago de Cuba, Ar- royo de la Costa (t.l.) SUBFAMILY OPATRINAE Collections. Havana (Garrido, type) TRIBE OPATRINI D. bielawskii Marcuzi 1985: 182 Distribution. Cuba. Endemic Ammodonus Mulsant and Rey 1859: 143 Cuba localities. M: Cienaga de Zapata; Playa A. tropicus Kirsch 1866: 190 Larga (t.l.) Distribution. Cuba, México to Brazil, Collections: Warsaw (type) Guadeloupe D. caguamensis Marcuzzi 1988: 74 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ca: Cayo Caguamas (t.l.) Collections. Havana (Garrido, type) 152 Arthropods of Florida and Neighboring Land Areas, Vol. 18

D. cubanus Marcuzzi 1962: 30 D. puncticeps Mulsant and Rey 1859: 180; Marcuzzi Distribution. Cuba. Endemic 1962: 30 Cuba localities. “Cuba” (t.l.). SC: Siboney, Distribution. Cuba. Endemic at lights Cuba localities. Collections. London (type) Collections. London D. difficilis Marcuzzi 1976: 126 D. rufoclavatus (Zayas) 1988: 91 (Blapstinus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Sierra de Trinidad? Cuba localities. Ci: Cumanayagua (t.l.). Ha: Collections. Havana, littoral litter D. dozieri Marcuzzi 1965: 128 Collections. Zayas (type) Distribution. Cuba, Bahamas (Grand Turk) D. trinitatis Marcuzzi 1976: 127 Cuba localities. Ho: Gibara Distribution. Cuba. Endemic Collections. Cuba localities. SS: Trinidad D. fuscicornis Chevrolat 1877: VIII; Garrido 2002: Collections. 38 D. waterhousei Mulsant and Rey 1859: 152 D. elongatus Marcuzzi 1976: 126 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico ( Mona Is- Cuba localities. land) Collections. Cuba localities. D. zayasi Marcuzzi 1988: 75 Collections. Distribution. Cuba. Endemic D. garciai Marcuzzi 1988: 99 Cuba localities. Gu: Baracoa, Yumurí; Distribution. Cuba, Endemic Guantánamo; Tortuguilla (t.l.); Meseta de Cuba localities. Gu: 30 km se Guantánamo; Maisí, La Asuncion Tortuguilla (t.l.); Yacabo Abajo Collections. Havana (Garrido, type) Collections. Havana (Garrido, type) D. garridoi Marcuzzi 1988: 78 TRIBE PLATYNOTINI Distribution. Cuba. Endemic Cuba localities. Ha: Marianao (t.l.) Opatrinus Latreille 1829: 19 Collections. Havana (Garrido, type) O. puertoricensis Marcuzzi 1977: 23 D. hernandezi Marcuzzi 1988: 72 O. p. armasi Garrido and Guttierez 1994a: 121; Distribution. Cuba. Endemic 1997a: 26; endemic subspecies in Cuba Cuba localities. IJ: Sierra de Caballos (t.l.) Distribution. Cuba, Jamaica, Puerto Rico. Collections. Havana (Garrido, type) nominate subspecies in Puerto Rico D. juraguensis Marcuzzi 1988: 75 Cuba localities. Ha: San Antonio de los Baños Distribution. Cuba. Endemic Collections. Havana (type) Cuba localities. SC: Santiago de Cuba, Playa O. pullus (Sahlberg) 1823: 16; Marcuzzi 1962: 32; Juraguá (t.l.) Garrido and Gutierrez 1997a: 26 Collections. Havana (Garrido, type) Distribution. Cuba, Jamaica, Puerto Rico, D. kaszabi Marcuzzi 1976: 125 México, Guatemala, Colombia, Venezuela Distribution. Cuba. Endemic Cuba localities. Widespread in Cuba Cuba localities. Ha: Santa Maria del Mar Collections. Havana Collections. D. macamboensis Marcuzzi 1988: 77; Garrido and TRIBE LEICHENINI Gutierrez 1996: 226 D. garciai Marcuzzi 1988: 79. Gu: Tortuguilla Leichenum Blanchard 1845: 14 (t.l.) L. canaliculatum variegatum Klug 1833: 88; Distribution. Cuba. Endemic Spilman 1961b: 127 Cuba localities. Gu: Macambo (t.l.); Yacabo Distribution. Cuba (introduced), Guadeloupe, Abajo; 50 km E of Guantánamo se USA; native to Madagascar Collections. Havana (Garrido, type) Cuba localities. PR: Guanahacabibes; D. orientalis Garrido and Gutierrez 1996: 226 Paredon de los Acosta Distribution. Cuba. Endemic Collections. Washington. Zayas Cuba localities. Gu: Baitiquiri (t.l.); Cajobabo Collections. Havana A Checklist of the Beetles of Cuba 153

SUBFAMILY L. oryzae Waterhouse 1880: 148. The longheaded TRIBE ULOMINI flour beetle Distribution. Cuba, México, Argentina, Old Uloma Laporte 1840: 219 World, USA (FL); in stored products U. extraordinaria Spilman 1961a: 113 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. SC: Gran Piedra; Pico Palma (L. prosopis Chittenden 1904: 167) Mocha to Pico Joachin (t.l.); Piloto; Pico Distribution. USA (FL), cosmopolitan Turquino. ? Calas Cuba localities. Collections. Washington (type) Collections. U. moensis Marcuzzi 2001: 286. Distribution. Cuba. Endemic. Lyphia Mulsant and Rey 1859 Cuba localities. Ho: Monte Ibenia (t.l.) (L. tetraphylla Fairmaire 1856: 534) Collections. Garrido. Genoa (type) Distribution. USA (FL); cosmopolitan Cuba localities. TRIBE HELOPINI Collections.

Nautes Pascoe 1866: 475 Mycotrogus Horn 1870: 367 N. asperipennis Allard 1894: 259; Marcuzzi 1962: M. paripunctatus Spilman 1963: 23 39 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. Gr: Hormiguero Collections. Washington (type) Collections. Leiden? N. azurescens (Jacquelin du Val) 1857: 153 (Helops); Orghidania Ardoin 1977a: 383 Gundlach 1891: 124; Ardoin 1977b: 391 O. torrei Ardoin 1977a: 383 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Siboney, at lights Cuba localities. SS: Cueva Colón Collections. Gundlach no. 1325 Collections. N. rufipes Allard 1876: 5 Distribution. Cuba. Endemic Palorus Mulsant 1854: 250 Cuba localities. P. ratzeburgii (Wissmann) 1848: 77. The smalleyed Collections. flour beetle Distribution. Cuba (introduced); spread by Tarpela Bates 1870: 272 commerce, probably native to Africa; cosmo- T. cactivora Zayas 1988: 105 politan Distribution. Cuba. Endemic Cuba localities. Cuba localities. Gu: Imias (t.l.); feeding on Collections. cactus resin P. foveicollis Blair 1930: 133 Collections. Zayas (type) Distribution. Cuba (introduced); spread by T. cuprosa Zayas 1988: 106 commerce; cosmopolitan Distribution. Cuba. Endemic Cuba localities. Cuba localities. Gu: Cuchillas de Toa. Ho: Collections. Cupeyal; Moa. Type locality not given Collections. Zayas (type) Tribolium McLeay 1825: 47 T. granulipennis (Jacquelin du Val) 1857: 154 T. castaneum (Herbst) 1797: 7. The red flour beetle (Helops); Gundlach 1891: 125 Uloma rubens Laporte 1840: 220; Jacquelin du Distribution. Cuba. Endemic Val 1857: 150; Gundlach l891: 114 Cuba localities. Widespread Distribution. Cuba (introduced), widespread Collections. Gundlach no. 1326 West Indies, USA to Argentina, Old World; in stored products; cosmopolitan TRIBE TRIBOLIINI Cuba localities. Collections. Latheticus Waterhouse 1880: 147 T. confusum Jacquelin du Val 1868 181. The con- fused flour beetle 154 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba (introduced), Jamaica, Zophobas Blanchard 1845: 15 (see Tschinkel Hispaniola, Puerto Rico, México to Chile, USA, 1984, Coleop. Bull. 38: 325-333) Old World; in stored products; cosmopolitan Z. atratus (Fabricius) 1775: 256 Cuba localities. Z. rugipes Kirsch 1866: 197; Marcuzzi 1962: 39; Collections. considered a separate and valid species in Cuba by Garrido and Gutierrez 1994b Ulosonia Laporte 1840: 220. Z. cubanus Marcuzzi 1976: 128. “Cuba.” Budapest U. biimpressa Latreille 1833: 17 (type) Distribution. Cuba, Guadeloupe, Curaçao, Z. morio (Fabricius) 1776: 241; Gundlach 1891: Tres Marias Island, México to Colombia and 117 Guiana Distribution. Cuba, México to Ecuador, Bra- Cuba localities. zil, Puerto Rico, USA (CA-FL) Collections. Cuba localities. Widespread, especially in U. tricornis (Dalman) 1823: 59; Jacquelin du Val caves in Gu and VC 1857: 148; Gundlach 1891: 120 (Hypogena) Collections. Gundlach no. 264 (Z. morio). Distribution. Cuba, Tres Marias, Leiden? Washington Revillagigedo, Curaçao, Venezuela, México to Guatemala, USA (FL) TRIBE CENTRONOPINI Cuba localities. M: Cárdenas Collections. Gundlach no. 61 Tauroceras Hope 1840: 130 T. cornuta Fabricius 1775: 256 TRIBE ALPHITOBIINI Distribution. Cuba (introduced), widespread, cosmopolitan; stored products pest? Alphitobius Stephens 1832: 11 Cuba localities. Havana (Bruner 1951: 85) A. diaperinus (Panzer) 1797: 16; Ardoin 1977: 383. Collections. The lesser mealworn beetle T. mulata Zayas 1988: 94 Distribution. Cuba, Jamaica, Puerto Rico, Distribution. Cuba. Endemic México, Trinidad, USA; spread by commerce; Cuba localities. Ca: Sierra Cubitas. Ha: La pest in many kinds of dried materials; cosmo- Havana. PR: Hato Nuevo (t.l.) politan Collections. Zayas (type) Cuba localities. Caves in Ha, SS, and VC, provinces; in dry guano TRIBE AMARYGMINI Collections. A. laevigatus (Fabricius) 1781: 90; Marcuzzi 1962: Cymatothes Solier 1848: 151 38; Ardoin 1977b: 391. The black fungus beetle C. nebulosa (Fabricius) 1781: 158 Heterophaga fagi Curtis 1831: 303; Jacquelin du Distribution. Cuba, México to Argentina, Val 1857: 147 (Heterocera); Gundlach 1891: 12 Hispaniola, Guadeloupe, Martinique Distribution. Cuba, West Indies, México to Cuba localities. Brazil, USA, Old World; in caves, stored prod- Collections. ucts, chicken coops C. tristis (Laporte) 1840: 236 (Pyanisia); Jacquelin Cuba localities. Widespread. Caves in prov- du Val 1857: 152; Gundlach 1891: 126; Ardoin inces of Gr, Ha, SC, SS, and VC; in bat guano 1977b: 391 Collections. Gundlach no. 73. Leiden? Distribution. Cuba, Puerto Rico, Guadelupe, México to Panamá, USA (AL) TRIBE TENEBRIONINI Cuba localities. Widespread Collections. Gundlach no. 428. Pittsburgh Tenebrio Linnaeus 1758: 417 C. undata (Fabricius) 1792: 122 T. molitor Linnaeus 1758: 417. The yellow meal- Distribution. Cuba, Hispaniola, Lesser worm beetle Antilles, México to Panamá, to Brazil and Ar- Distribution. Cuba, widespread West Indies, gentina USA (FL); spread by commerce; in many kinds Cuba localities. of stored products; cosmopolitan Collections. Cuba localities. Collections. A Checklist of the Beetles of Cuba 155

SUBFAMILY ALLECULINAE Distribution. Cuba. Endemic The comb-clawed bark beetles Cuba localities. PR: Pan de Guajaibon (t.l.) This group has traditionally been treated as a Collections. Urbana (type) separate family but modern study shows that it is L. (F.) cayamasensis Campbell 1966: 33 a unit within the Tenebrionidae. Adults are usu- Distribution. Cuba. Endemic ally found by beating vegetation, at lights, or on Cuba localities. Ci: Cayamas (t.l.) tree trunks at night. Collections. Washington (type) L. (F.) emarginata Campbell 1966: 32 TRIBE ALLECULINI Distribution. Cuba. Endemic SUBTRIBE ALLECULINA Cuba localities. SC: Punta Turquino (t.l.) Collections. Miami (type) Hymenorous Mulsant 1851: 201 L. (F.) flavifemoralis Campbell 1966: 29; 1971: 30 H. bahamensis Campbell 1971: 88 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas Cuba localities. SC: Florida Blanca; Alto Cuba localities. Ci: Soledad. Gu: Songo (t.l.); Pico Turquino Guantánamo. Ha: Havana; Sierra de Anafe. Collections. Cambridge. Urbana (type) M: Peninsula de Zapata. PR: Pinar del Rio: ? L. (F.) flavipes (Jacquelin du Val) 1857: 157 Pedrera de Mendoza (Allecula); Gundlach 1891: 123; Campbell 1966: Collections. Cambridge. Gainesville. Ottawa. 25; 1971: 27 Zayas Distribution. Cuba. Endemic H. convexus Casey 1891: 106; Campbell 1971: 72 Cuba localities. Widespread Distribution. Cuba, Bahamas, USA (s FL) Collections. Cambridge. Champaign. Cuba localities. Gu: Baracoa. Ha: Jibacoa. Gundlach no. 519. New York. Ottawa. Wash- M: Peninsula de Zapata. SC: Ciudamar. ington Collections. Champaign. Ottawa. Washing- L. (F.) quadrinotata Campbell 1971: 28 ton. Zayas Distribution. Cuba. Endemic H. cubensis Campbell 1971: 81 Cuba localities.. CA: Baraguá (t.l.). Ci: Cali- Distribution. Cuba. Endemic fornia; Cumanayagua. Gu: Imias; S. de Cuba localities. Ci: Soledad (t.l.). Gu: Upper Cubitas, Pardones Ovando River Collections. Harvard (type). Ottawa. Zayas Collections. Harvard (type). Ottawa L. (F.) schwarzi Campbell 1971: 29 H. darlingtoni Campbell 1971: 83 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. SS: Buenos Aires (t.l.); Collections. Washington (type) Trinidad L. (F.) tibiodentata Campbell 1966: 30; 1971: 36 Collections. Harvard (type) Distribution. Cuba. Endemic H. farri Campbell 1971: 84 Cuba localities. Widespread Distribution. Cuba, Cayman Islands, Ja- Collections. Helsinki. New York. Ottawa. maica Washington (type). Zayas Cuba localities. Ci: Cayamas. M: Peninsula L. (F.) villasensis Campbell 1971: 35 de Zapata Distribution. Cuba. Endemic Collections. Ottawa. Washington. Zayas Cuba localities. Ci: Soledad (t.l.). SS: H. sparsepunctatus Campbell 1971: 97 Trinidad, Yaguanabo Distribution. Cuba. Endemic Collections. Harvard (type). Zayas Cuba localities. Ci. Cayamas (t.l.) L. (Lobopoda) alutacea Campbell 1971: 58 Collections. Washington (type) Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Lobopoda Solier 1835: 233 Collections. Ottawa. Washington (type) L. (Flavipoda) badius Campbell 1971: 31 L. (L.) cubensis Campbell 1966: 157 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Soledad (t.l.). SS: Cuba localities. (locality unknown) Trinidad, SS: Buenos Aires; San Blas Collections. Munich (type) Collections. Harvard (type). Ottawa. Zayas L. (L.) distans Campbell 1971: 48 L. (F.) bicolor Campbell 1966: 28; 1971: 31 Distribution. Cuba. Endemic 156 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. SC: Loma del Gato (t.l.) Cuba localities. Ho: Cupeyal (t.l.) Collections. Harvard (type) Collections. Zayas (type) L. (L.) fallaciosa Campbell 1971: 49 C. ferruginea Garrido and Gutierrez 1996: 282 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas; Soledad (t.l.). Cuba localities. Gu: Baracoa (t.l.); Punta de Gu: Guantánamo Maisí Collections. Gainesville. Harvard (type). Ot- Collections. Havana tawa. Washington C. gebieni Marcuzzi 1976: 139 L. (L.) paracornis Campbell 1971: 53 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Cordillera de los Cuba localities. SC: Pico Turquino (t.l.) Organos, Sitio del Infierno (t.l.) Collections. Harvard (type). Ottawa. Zayas Collections. Basel (type) L. (L.) paracollis Campbell 1971: 55 C. gundlachi Marcuzzi 1998: 160 (replacement Distribution. Cuba. Endemic name for Cnodalon inflatum Zayas 1988: 101) Cuba localities. Gu: Mts. N. of Imias (t.l.). Distribution. Cuba. Endemic SC: Gran Piedra; Loma del Gato Cuba localities. Ho: Cupeyal (t.l.) Collections. Harvard (type). Ottawa. Zayas Collections. Zayas (type) L. (L.) polita Campbell 1971: 52 C. inflatum Marcuzzi 1976: 138 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. “Cuba” (t.l.) Collections. Harvard (type) Collections. Budapest. London L. (L.) sulcaticollis Pic 1933: 1: Campbell 1966: C. iviei Marcuzzi 1998: 160 (replacement name for 163; 1971: 57 Cnodalon trinitatis Zayas 1988: 102) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ca: Monte Imias. Ci: Soledad. Cuba localities. SS: Lomas de Trinidad (t.l.); SC: Vequitas Pico de Potrerillo Collections. Brussels (type). Cambridge. Ot- Collections. Zayas (type) tawa. Washington C. parallelus (Zayas) 1988: 96 (Cnodalon); Marcuzzi 1998: 160 SUBFAMILY COELOMETOPINAE Distribution. Cuba. Endemic TRIBE COELOMETOPINI Cuba localities. Ho: Cupeyal (t.l.) Collections. Zayas (type) Apsida Lacordaire 1859: 309 C. punctatum (Zayas) 1988: 103 (Cnodalon); A. cubanensis Kulzer 1961: 217 Marcuzzi 1998: 160 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ha: Playa Mariano (t.l.). PR: Cuba localities. PR: Rancho Mundito (t.l) Sierra de Rosario Collections. Zayas (type) Collections. Basel (type) C. tumefactum Marcuzzi 1976: 138 Distribution. Cuba. Endemic Cyrtosoma Perty 1830: 59 Cuba localities. “Cuba” (t.l.) C. cristalensis (Zayas) 1988: 99 (Cnodalon); Collections. Marcuzzi 1998: 160 C. turquinensis (Zayas) 1988: 96 (Cnodalon); Distribution. Cuba. Endemic Marcuzzi 1998: 160 Cuba localities. Ho: Sierra Cristal (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. SC: Loma del Gato; Pico C. cuprosa (Zayas) 1988: 98 (Cnodalon); Marcuzzi Turquino (t.l.) 1998: 160 Collections. Zayas (type) Distribution. Cuba. Endemic C. zayasi Marcuzzi 1998: 160 (replacement name Cuba localities. Gr: La Siberia. SC: Pico for Cnodalon tumefactum Zayas 1988: 95) Turquino.. Type locality not given Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Ho: Cupeyal (t.l.) C. elongatus (Zayas) 1988: 101 (Cnodalon); Collections. Zayas (type) Marcuzzi 1998: 160 Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 157

TRIBE TALANINI S. venusta Zayas 1988: 109 Distribution. Cuba. Endemic Talanus Mäklin 1878: 96 Cuba localities. Ha: Santiago de las Vegas T. cribrarius Jaquelin du Val 1857: 156; Gundlach (t.l.) 1891: 125 Collections. Zayas (type) Distribution. Cuba, Grenada S. virescens Zayas 1988: 110 Cuba localities. M: Colón Distribution. Cuba. Endemic Collections. Gundlach no. 1099 Cuba localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) TRIBE STRONGYLIINI REFERENCES Strongylium Kirby 1818: 417 Aalbu, R.L., C. A. Triplehorn, J.M. Campbell, S. amethystinum Guérin-Méneville in Chevrolat K.W. Brown, R.E.Somerby, and D. B. Tho- 1838: 281; Jacquelin du Val 1857: 155; Gundlach mas. 2002.v 106 Tenebrionidae. Pp. 463-509. 1891: 125 (Stenochia amethistina); Marcuzzi In: Arnett, R. H., Jr., M. C. Thomas, P. E. 1962: 39 Skelley, and J. H. Frank (eds.). American Distribution. Cuba. Endemic beetles, Polyphaga: Scarabaeoidea through Cuba localities. Widespread Curculionidea. Vol. 2. CRC Press, Boca Raton, Collections. Gundlach no. 107. Leiden? FL. 861 pp. S. antennale Mäklin 1864: 162 Ardoin, P. 1977a. Tenebrionidae (Coleoptera) Distribution. Cuba. Endemic récoltés en 1969 dans les grottes de Cuba par Cuba localities. Gu: Guantánamo l’expedition biospéologiques Cubano-Roumaine Collections. a Cuba. Résultats des expéditions S. azureum Germar 1824: 153 biospéeologique Cubano-Roumaines à Cuba 2: Distribution. Cuba, Brazil, Bolivia 382-385. Cuba localities. Ardoin, P. 1977b. Tenebrionidae (Coleoptera) Collections. récoltés par la deuxieme expedition S. basiclavis Zayas 1988: 110 biospéologiques Cubano-Roumaine à Cuba Distribution. Cuba. Endemic (1973). Résultats des expéditions Cuba localities. Gu: Yateras (t.l.). Ho: biospéologiques Cubano-Roumaine a Cuba 2: Cupeyal 387-392. Collections. Zayas (type) Ardoin, P. 1977c. Contribution a l’étude des S. cupeyal Zayas 1988: 110 espèces Américaines du genre Doliema Pascoe Distribution. Cuba Endemic (Col. Tenebrionidae). Annls. Soc. Ent. France Cuba localities. Ho: Cupeyal (t.l.) (n.s.) 13: 1-20. Collections. Zayas (type) Campbell, J.M. 1966. A revision of the genus S. eximium Mäklin 1862: 161 Lobopoda (Coleoptera: Alleculidae) in North Distribution. Cuba. Endemic America and the West Indies. Illinois Biol. Cuba localities. Monog. 37, 203 pp. Collections. Campbell, J.M. 1971. A revision of the Alleculidae S. nigra Zayas 1988: 110 (Coleoptera) of the West Indies. Mem. Ent. Soc. Distribution. Cuba. Endemic Can. 81, 140 pp. Cuba localities. Ho: Sierra Cristal; Moa. Type Garrido, O.H. 2002. El género Diastolinus (Co- locality not given leoptera: Tenebrionidae: Pedinini) en Puerto Collections. Zayas (type) Rico, con la designación de un nombre nuevo S. preciosus Zayas 1988: 108 para D. elongatus. Soleonodon 2: 38-41. Distribution. Cuba. Endemic Garrido, O.H. 2003. Diaperis viridula (Coleoptera: Cuba localities. PR: Guanahacabibes; Tenebrionidae: Diaperini) es un táxon válido Viñales. Type locality not given que representa un género nuevo para Cuba. Collections. Zayas (type) Solenodon 3: 49-52. S. turquinensis Zayas 1988: 107 Garrido, O.H. and E. Gutierrez. 1994a. A new Distribution. Cuba. Endemic species of Opatrinus (Coleoptera: Pedinini: Cuba localities. SC: Pico Turquino (t.l.) Tenebrionidae) from Cuba. Insecta Mundi 8: Collections. Zayas (type) 121-124. 158 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Garrido, O.H. and E. Gutierrez. 1994b. Vari- Heteromera). Annali Mus. Civico Storia Natu- ability of Zophobas rugipes Kirsch (Coleoptera: rale “Giacomo Doria” (Genoa). 93: 269-291. Tenebrionidae) in Cuba. Insecta Mundi 8: 243- Marcuzzi, G. 1984. A catalogue of Tenebrionidae 249. beetles (Coleoptera: Heteromera) of the West Garrido, O.H. and E. Gutierrez. 1995a. Nueva Indies. Folia Ent. Hungarica 45(1): 69-108. especie de Loxostethus (Coleoptera: Marcuzzi, G. 1985. New taxa of Neotropical Tenebrionidae: Diaperini) para Cuba. Insecta Tenebrionidae (Coleoptera). Ann. Hist.-Nat. Mundi 9: 7-10. Mus. Nat. Hung. 77: 171-186. Garrido, O.H. and E. Gutierrez. 1995b. Nueva Marcuzzi, G. 1988. New species of Trientoma and especie de Trientoma (Coleoptera: Diastolinus (Coleoptera: Tenebrionidae) from Tenebrionidae: Trientomini) para Cuba con Cuba. Annali. Mus. Civico Storia Naturale consideraciones sobre otras especies. Insecta “Giacomo Doria” (Genova) 87: 67-83. Mundi 9: 47-49. Marcuzzi, G. 1998. Supplement to the catalogue Garrido, O.H. and E. Gutiérrez. 1996. of Tenebrionidae (Coleoptera) of the West Indies. Consideraciones sobre el genero Diastolinus Ann. Hist.-Natur. Musei Nat. Hung. 90: 151- (Coleoptera: Tenebrionidae: Pedinini) en Cuba, 162. con descripcion de dos nuevas especies. Insecta Peck, S.B., Ruiz-Baliú, A.E., and Garcés Mundi 10: 225-229. Gonzaléz, G.F. 1998. The cave-inhabiting Garrido, O. and E. Gutiérrez. 1997a. beetles of Cuba (Insecta: Coleoptera): diversity, Consideraciones sobre el género Opatrinus (Co- distribution, and ecology. J. Cave Karst Stud- leoptera: Tenebrionidae: Pedinini) en Cuba, ies 60: 156-166. Puerto Rico, y Jamaica. Insecta Mundi 11: 25- Spilman, T. J. 1961a. Uloma extraordinaria, a 27. new species from Cuba (Tenebionidae). Coleop. Garrido, O. and E. Gutiérrez. 1997b. Revision Bull. 15: 113-115. del género endemico cubano Trimytantron Spilman, T. J. 1961b. A few tenebrionids new to (Coleoptera:Tenebrionidae: Trimytini), con la Cuba. Coleop. Bull. 15: 127. descripcion de ocho nuevas especies. Insecta Spilman, T. J. 1963. The American genus Mundi 11: 29-41. Mycotrogus: a synopsis, a new species from Ivie, M. A. 1991. Taxonomic notes on a little known Cuba, and a note on a larva. Proc. Ent. Soc. publication: Zayas, 1988, Entomofauna Washington 65: 21-30. Cubana. Order Coleoptera. Coleop. Bull. 45: 399- Triplehorn, C. 1962. New Diaperini (Coleoptera: 401. Tenebrionidae) from the West Indies. Ann. Ent. Kaszab, A. 1977. Neue Tenebrioniden (Coleoptera) Soc. Am. 55(5): 502-507. von den Galapagos und Antillen, sowie aus Triplehorn, C. 1965. Revision of Diaperini of Südamerika und Ostasien. Annales Historico- America North of México with notes on extral- Naturales Musei Nationalis Hungarici 69: 117- imital species (Coleoptera: Tenebrionidae). Proc. 131. U.S. Nat. Mus. 117(3515): 349-358. Kulzer, H. 1961. Neue Tenebrioniden aus Triplehorn, C.A. 1979. Two new species of Südamerika (Col.). Ent. Arb. Museum Frey 12: Corticeus from Florida and the West Indies 205-235. (Coleoptera: Tenebrionidae). Proc. Ent. Soc. La Rosa, J. and L.L. Vázquez. 1987. Wash. 81: 46-50. Distribucion, danos y lucha contra los Triplehorn, C. A. 1990. Review of the genus principales insectos de los productos vegetales Corticeus (Coleoptera: Tenebrionidae) of almacenados en Cuba. En Resumenes Primer America north of México. Ann. Ent. Soc. Am. seminario Cientifico internacional de Sanidad 83: 287-306. vegetal, Septiembre 1987 (not seen). Triplehorn, C.A. 1991. A review of the genus Marcuzzi, G. 1962: Tenebrionid beetles of the West Phaleria Latreille from the Western Hemi- Indies. Stud. Fauna Curaçao Carib. Islands. sphere (Coleoptera: Tenebrionidae: Phalerinae). No. 57. 12: 21-48. Coleop. Bull. 45: 258-270. Marcuzzi, G. 1976. New species of Neotropical Triplehorn, C. A. and O. Merkl. 1997. Review Tenebrionidae (Coleoptera). Ann. Hist. Nat. of the genus Loxostethus Triplehorn, with de- Mus. Natn. Hung. 68: 117-139. scriptions of three new species (Coleoptera: Marcuzzi, G. 2001. New species of tenebrionid Tenebrionidae; Diaperini.). Ann. Ent. Soc. Am. beetles from Central and South America (Col. 90: 736-741. A Checklist of the Beetles of Cuba 159

Triplehorn, C.A. and L.E. Watrous. 1979. A Distribution. Cuba. Endemic synopsis of the genus Phaleria in the United Cuba localities. States and Baja California (Coleoptera: Collections. Tenebrionidae). Coleop. Bull. 33: 275-295. Vasquez, M.L.L. 1986. Plagas de almacen en Cuba. Hypasclera Kirsch 1866: 210 Coleoptera (Cleridae, Trogossitidae, H. costata (Champion) 1896: 40; Arnett 1984: 3 Dermestidae, Bostrichidae, Anobiidae, Distribution. Cuba, Jamaica, Hispaniola, St. Nitidulidae, Cucujidae, Silvanidae, Vincent, México to Panamá; seemingly not es- Mycetophagidae, Tenebrionidae). Rev. Centro tablished in the USA Agricola 13(2): 44-48. Cuba localities. Watrous, L.E. and C.A. Triplehorn. 1982. Collections. Phaleria of the West Indies and H. dorsalis (Melsheimer) 1846: 55; Arnett 1984: 3 Circumcaribbean region (Coleoptera: Distribution. Cuba, Bahamas, México, USA Tenebrionidae). Coleop. Bull. 36: 12-21. (TX-NY-FL) Cuba localities. 141. FAMILY Collections. The family contains Dryocora of New Zealand and Australia, and widespread Prostomis. The fam- Oxacis LeConte 1866: 165 ily is probably absent from the West Indies. O. laeta (Waterhouse) 1878: 306; Arnett 1984: 3 Distribution. Cuba, Hispaniola, Puerto Rico, 142. FAMILY St. Croix, USA (FL) The family contains Synchroa of North America Cuba localities. and Asia, and Mallodrya of North America. The Collections. family is probably absent from the West Indies. Oxycopis Arnett 1951: 318 143. FAMILY OEDEMERIDAE (O. falli (Blatchley) 1928: 63 (Oxacis); Arnett 1984: The false blister beetles 2) Adults are often brightly colored and on flow- Distribution. Bahamas, Hispaniola, Cayman ers, feeding on pollen and nectar, and most abun- Islands, USA (south FL) dant on oceanic coastlines. Larvae live in moist Cuba localities. rotted wood, including driftwood. Arnett (1984) is a Collections. summary of the oedemerid fauna of Florida and O. mcdonaldi (Arnett) 1951: 328; Arnett 1984: 2 the principal reference for this section. That paper Distribution. Cuba, Grand Cayman, Bimini, may include species yet to be found in Cuba. Arnett USA (FL) (1961) is a key to the genera of the New World. Cuba localities. Collections. SUBFAMILY NACERDINAE O. testaceicollis (Jacquelin du Val) 1857: 158 (Asclera); Gundlach 1891: 126 Nacerdes Dejean 1834: 228 Distribution. Cuba. Endemic (N. melanura (Linnaeus) 1758: 403; Arnett 1984: Cuba localities. M: Cienaga de Zapata 2). The wharf borer Collections. Distribution. Bahamas, México to Argentina, O. thonalmus (Darlington) 1936: 102 (Copidita) widespread USA; introduced from Europe. This Distribution. Cuba. Endemic species damages pilings and ship timbers, and Cuba localities. CA: Janorú (Jaronú) (t.l.). breeds in drift wood in fresh and marine wa- SS: Buenos Aires ters Collections. Cambridge (type) Cuba localities. Notes. This species seems to be part of a Batesian Collections. mimicry complex centered around species of Thonalmus (Lycidae) (Darlington 1938: 689) SUBFAMILY OEDEMERINAE O. vittata Fabricius 1775: 125; Leng and Mutchler TRIBE ASCLERINI 1914: 466; Arnett 1984: 2 Distribution. Cuba, Jamaica, Puerto Rico, Asclera Stephens 1839: 336 Vieques, Guadeloupe, Virgin Islands, Bahamas, A. latior Pic 1923: 32 USA (FL) 160 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Distribution. Cuba. Hispaniola, Puerto Rico, Collections. Lesser Antilles, Trinidad, s USA Cuba localities. Gu: Baracoa. ? Nibujón REFERENCES Collections. Havana Arnett, R. H., Jr. 1961. Contribution toward a monograph of the Oedemeridae. 14. A key to SUBFAMILY NEMOGNATHINAE and notes on the New World genera. Coleop. TRIBE HORIINI Bull. 15: 49-64. Arnett, R. H., Jr. 1984. The false blister beetles Cissites Latreille 1804: 154 of Florida (Coleoptera, Oedemeridae). Florida C. maculata Swederus 1887: 196; Selander and Dept. Agric. Cons. Serv., Entomol Circ. Bouseman 1960: 212; Genaro 1996: 384 259: 1-4. Distribution. Cuba, Hispaniola, Puerto Rico, Darlington, P.J., Jr. 1938. Experiments on mim- Lesser Antilles, México, Central America, icry in Cuba, with suggestions for future study. South America; endoparasitic on Xylocopa car- Trans. R. Ent. Soc. London 87: 681-695. penter bees, X. cubaecola Lucas in Cuba Cuba localities. Widespread 144. FAMILY Collections. Havana. Zayas This family is Holarctic in cool temperate habi- tats, and the larvae live in rotten and often moss TRIBE NEMOGNATHINI covered logs. Nemognatha Illiger 1807: 333 145. FAMILY MELOIDAE N. occupata (Blackwelder) 1945: 481 (Zonitis); The blister beetles Selander and Bouseman 1960: 220; Genaro Reviewed by John Pinto, March, 1999. 1996: 385 Adults are usually found feeding on leaves or N. cubaecola Jacquelin du Val 1857: 161; on flowers and some phytophagous species may Gundlach 1891: 258 cause crop damage. Larvae feed on grasshopper Distribution. Cuba. Endemic eggs, or eggs and food stores of bee nests. Genaro Cuba localities. Ha; Santiado de las Vegas; (1996) gives a key to the Cuban species, as well as San Nicolása de Bari; Ceiba del Agua. Gu: data on habits and distribution. Guantánamo; San German. M: Cárdenas. SS: Trinidad SUBFAMILY TETRAONYCINAE Collections. Gundlach no. 1322. Havana. TRIBE TETRAONYCHINI Washington N. punctulata LeConte 1853: 347; Selander and Tetraonyx Latreille 1805: 204 Bouseman 1960: 222, 1961: 191; Genaro 1996: T. cruciata Castelnau 1840: 277; Selander and 385 Bouseman 1960: 210; Genaro 1996: 383 Distribution. Cuba, Bahamas, Cayman Is- T. cubensis Chevrolat 1858: 210 (sic Tetraomyx) lands, Jamaica, se USA Distribution. Cuba. Endemic Cuba localities. Widespread in Cuba. Lar- Cuba localities. Widespread vae develop in nests of Megachile poeyi and M. Collections. Cambridge. Havana. Washing- singularis leaf-cutting bees. It is the most fre- ton quently collected meloid in Cuba T. maestra Selander and Bouseman 1960: 211; Collections. Champaign. Havana. Washing- Genaro 1996: 384 ton Distribution. Cuba. Endemic Cuba localities. Gu: Yunque de Baracoa. Ho: Pseudozonitis Dillon 1952: 181 Sierra Cristal, Moa; Nicaro. SC: Gran Piedra; P. marginata (Fabricius) 1781: 159; Selander and Loma del Gato; Pico Turquino (t.l.) Bouseman 1960: 214; Genaro 1996: 385 Collections. Cambridge (type). Havana Distribution. Cuba, Bahamas, Jamaica, T. quadrimaculata (Fabricius) 1792: 50 (Apalus); Hispaniola, Puerto Rico, St. Croix, Lesser Selander and Bouseman 1960: 205; Genaro Antilles 1996: 384 Cuba localities. Widespread Mylabris ruficollis Olivier 1795: 14 Collections. Havana. Zayas A Checklist of the Beetles of Cuba 161

REFERENCES Cuba localities. Ci: Cayamas. Ho Genaro, J.A. 1996. Resumen del conocimiento Collections. London. Paris (type, Grouvelle sobre las meloides de Cuba (Insecta: Coleoptera). colln.). Washington Carib. J. Sci. 32: 382-386. Selander, R.B. and J.K. Bouseman. 1960. Meloid REFERENCES beetles (Coleoptera) of the West Indies. Proc. Pollock, D. A. 1999. Review of the New World U.S. National Museum 111: 197-226. Hemipeplinae (Coleoptera: Mycteridae) with de- Selander, R.B. and J.K. Bouseman. 1961. scriptions of ten new species. Ent. Scand. 30: Supplementary records of meloid beetles (Co- 47-73. leoptera) of the West Indies. Ent. News 73: 190- 191. 147. FAMILY This family occurs only in the Holarctic region. 146. MYCTERIDAE Larvae live under the bark of decaying conifers. The palm and flower beetles This family occurs in North America, Eurasia, 148. FAMILY and North Africa, and the tropics. These probably This family occurs only in Asia and the East occur in Cuba in leaf axils of palms. Indies. The larvae probably live in rotten wood.

SUBFAMILY LACCONOTINAE 149. FAMILY PYTHIDAE This family occurs in North, Central, and South Conomorphus Champion 1889: 98 America, but is not known from the West Indies. C. sp. The larvae feed under rotten bark. Distribution. Cuba. Endemic Cuba localities. 150. FAMILY PYROCHROIDAE Collections. The fire-colored beetles This family occurs in North America, and has SUBFAMILY HEMIPEPLINAE a species apiece in French Guiana and Chile. It is apparently absent from México, Central America, Hemipeplus Berthold 1827: 384 and the West Indies. Adults are short-lived and may H. marginipennis (LeConte) 1854: 79 (Nemicellus); be collected at lights. The larvae occur under bark. Pollock 1999: 57 [H. microphthalmus (Schwarz) 1878: 360 151. FAMILY SALPINGIDAE (Nemicellus)]; as synonym (Blackwelder 1945: The narrow-waisted bark beetles 423); not in Cuba The larvae of most species occur under bark, Distribution. Cuba, Bahamas, Mexico, USA in decaying wood, or leaf axils. (FL, GA) Cuba localities. Ci: Cayamas. Gu: SUBFAMILY DACODERINAE Guantánamo Collections. Gainesville. Washington Dacoderus LeConte 1858: 74 H. longiscapus Pollock 1999: 61 D. dominicensis Horn 1874: 219 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola Cuba localities. Ci: Cayamas (t.l.) Cuba localities. Collections. Washington (type) Collections. H. glabratus Pollock 1999: 61 Distribution. Cuba. Endemic 152. FAMILY ANTHICIDAE Cuba localities. Ci: Cayamas (t.l.) The ant-like flower beetles Collections. Gainesville. Pollock. Washing- Reviewed by D. Chandler, March, 2001 ton (type) Adults occur on vegetation and in leaf litter. They are predators or scavengers. Many species in Holopeplus Arrow 1930: 230 this family were described by LaFerté-Sénectère H. gundlachi (Grouvelle) 1878: XLIV (Hemipeplus); and the correct publication dates are discussed in Pollock 1999: 56 Chandler (1999). H. cubensis Arrow 1930: 230 Distribution. Cuba, Bahamas 162 Arthropods of Florida and Neighboring Land Areas, Vol. 18

SUBFAMILY MACRATRIINAE Formicillia gracillipes (sic) Buck 1960: 64 Cu- ban specimens in part Macratia Newman 1838: 377 Distribution. Cuba, Hispaniola, Cayman Is- M. insularis (Laferté-Sénectère) 1848: 17 lands, Venezuela, Honduras, Panama, s USA (Macrarthrus); Jacquelin du Val 1857: 159; Cuba localities. PR: Pinar del Rio Gundlach 1891: 259 (Macrarthrius) Collections. Distribution. Cuba. Endemic A. septemnotatus (Laferté-Sénectère) in Champion Cuba localities. Gu: Baracoa. M: Cárdenas 1890: 227 Collections. Gundlach no. 757 Distribution. Cuba, s Mexico to Brazil Cuba localities. SUBFAMILY Collections. TRIBE ANTHICINI Paykull 1798: 253 Laferté-Sénectère 1849: 5 A. antilleorum Werner 1983b: 218 A. albicinctus (Laferté-Sénectère) 1849: 10; Werner Distribution. Cuba, Bahamas, Hispaniola, 1970c: 860 Jamaica, Virgin Islands Distribution. Cuba, e Mexico to Venezuela Cuba localities. CA: Baraguá. Ci: Cayamas; Cuba localities. Soledad. Gu: Imias Mts., California. Collections. Collections. Cambridge. Champaign. Wash- A. angusticollis (Laferté-Sénectère) 1849: 19; ington Werner 1966: 747; 1983b: 231 A. blackwelderi Werner 1983b: 219 Distribution. Cuba, Jamaica, Colombia to Distribution. Cuba, Jamaica, Hispaniola Argentina Cuba localities. Ci: Soledad. SC: coast below Cuba localities. Ca: Camaguey. Ci: Cayamas; Pico Turquino Santa Clara. Ha: Havana. PR: Bahia Honda Collections. Cambridge Collections. Washington A. cubanus Pic 1944: 7 A. concinnus (Laferté-Sénectère) 1849: 14; Werner Distribution. Cuba. Endemic 1970a: 123; 1983b: 231 Cuba localities. Distribution. Cuba, Hispaniola, e México to Collections. Trinidad, Bolivia and Brazil [A. lesnei Pic 1899: 30] Cuba localities. Intercept on ship from Cuba Distribution. “West Indies” is an error, from (dead at light) “Indies occidental” (India or Sumatra) Collections. Washington A. macgillavryi Buck 1960: 69; Werner 1983b: 226 A. ebeninus (Laferté-Sénectère) 1849:16; Werner Distribution. Cuba. Endemic 1970a: 119; 1983b: 231 Cuba localities. CA: Baraguá. Ci: Cayamas. Pseudoleptaleus cubanensis Pic 1917: 8 ?Limones Leptaleus cubaensis Blackwelder 1945: 433 Collections. Cambridge Distribution. Cuba, Guatemala, Venezuela, A. soledad Werner 1983b: 225 Colombia Distribution. Cuba, Jamaica Cuba localities. Cuba localities. Ci: Soledad (t.l.) Collections. Collections. Cambridge (type) A. quinquemaculatus (Laferté-Sénectère) 1849:14; A. subtilis Laferté-Sénectère 1849: 34; Werner Werner 1970a: 121 1983b. 222 Distribution. Cuba, Puerto Rico, Trinidad, Distribution. Cuba, Hispaniola, Venezuela Ecuador, e Mexico to Brazil Cuba localities. Ci: Soledad. SC: Loma del Cuba localities. Gu: Sabanilla Gato Collections. Collections. Cambridge A. schwarzi Werner 1967c: 1232; 1983b: 232 Distribution. Cuba. Endemic Mulsant and Rey 1866: 77 Cuba localities. Ci: Cayamas (t.l.); Soledad. (C. texanus (Laferté-Sénectère) 1849: 301 PR: Pinar del Rio Distribution. Cayman Islands, Hispaniola, Collections. Washington (type) Puerto Rico, ne México, USA (MA-FL-TX) A. scitulus (LeConte) 1852: 94; Werner 1970b: 724 Cuba localities. Formicilla cubana Pic 1944: 9 Collections. A Checklist of the Beetles of Cuba 163

Omonadus Mulsant and Rey 1866: 104 Guadeloupe, México-Panamá, Venezuela, USA O. floralis (Linnaeus) 1758: 420 (Anthicus); (CA-MO-NC-FL) Jacquelin du Val 1857: 160; Gundlach 1891: Cuba localities. Ca: Camaguey. CA: Baraguá; 259 (Anthicus); Werner 1964: 233 (Anthicus). Manicaragua. Ci: Cayamas; Soledad. Ha: Ha- The narrownecked grain beetle vana. SS: Jatibonica Distribution. Cuba (introduced), Jamaica, Collections. Cambridge Puerto Rico, México to Chile, Guadeloupe to Grenada, Canada (BC), USA (CA-ME-FL); cos- REFERENCES mopolitan; native to the Old World; in stored Buck, F. D. 1960. Anthicid beetles from Venezu- grain and dried fruit, and lawn and leaf clip- ela, Colombia, Cuba, and the Netherlands pings Antilles. Studies on the fauna of Curaçao and Cuba localities. Widespread other Caribbean Islands: no. 47, pp 64-71. Collections. Gundlach no. 109 Martinus Nijhoff, The Hague. (O. formicarius (Goeze) 1777: 706); Werner 1964: Chandler, D. S. 1999. Publication dates of papers 234 (Anthicus) on Anthicidae (Coleoptera) by the Marquis F.T. Distribution. Jamaica, Tobago, Mexico, de Laferté-Sénectère. Transactions of the Panama, Canada (BC), USA (CA-ME-FL); in- American Entomological Society 125: 433-439. troduced; cosmopolitan; native to the Old World Werner, F.G. 1961. A revision of the genus Cuba localities. Vacusus Casey (Coleoptera: Anthicidae). Ann. Collections. Ent. Soc. Amer. 54: 798-809. Werner, F.G. 1964. A revision of the North Ameri- Casey 1895: 732 can species of Anthicus s.str. (Coleoptera, S. pallidus (Say) 1826: 245 (Anthicus); Werner Anthicidae). Misc. Publ. Ent. Soc. Amer. 4: 195- 1964: 230; 1983b: 233 (Anthicus) 242. Distribution. Cuba, Hispaniola, Puerto Rico, Werner, F.G. 1966. A revision of Acanthinus (Co- Grenada, Guadeloupe, Dominica, México to leoptera: Anthicidae). II. The angusticollis- Panamá, Venezuela, USA (TX, MS, AL, FL) group. Annals of the Entomological Society of Cuba localities. Gu: Maisi America 59: 746-751. Collections. Werner, F.G. 1967. A revision of Acanthinus (Co- S. teapensis (Champion) 1890: 249 (Anthicus); leoptera: Anthicidae). VI. Annals of the Ento- Werner 1983a: 424; 1983b: 233 mological Society of America 60: 1217-1234. Distribution. Cuba, Jamaica, Hispaniola, Werner, F.G. 1970a. A revision of Acanthinus (Co- Puerto Rico, Virgin Islands, Guadeloupe, leoptera: Anthicidae). VII. Annals of the Ento- México to Brazil mological Society of America 63: 111-128. Cuba localities. Ci: Cayamas. Gu: Baracoa. Werner, F.G. 1970b. A revision of Acanthinus (Co- PR: Viñales leoptera: Anthicidae). IX. The leporinus, Collections. scitulus, cuyabanus, and mycetops-groups. Annals of the Entomological Society of America Stricticomus Pic 1894: 42 63: 719-731. S. tobias (Marseul) 1879: 125 (Anthicus); Werner Werner, F.G. 1970c A revision of Acanthinus (Co- 1983b 233: (Anthicus) leoptera: Anthicidae). X. Albicintus group and Distribution. Cuba (introduced), Jamaica, conclusion. Annals of the Entomological Soci- Cayman Islands, Puerto Rico, Virgin Islands, ety of America 63: 859-876. Dominica, Guadeloupe, México, Panamá, USA Werner, F.G. 1983a. Neotropical Sapintus, with (CA-OR-NH-FL); cosmopolitan a general key to species (Coleoptera: Cuba localities. Anthicidae). Proceedings Ent. Soc. Washing- Collections. ton 85: 405-425. Werner, F.G. 1983b. Anthicidae of the Greater Vacusus Casey 1895: 674 Antilles and a new species from Venezuela (Co- V. vicinus (Laferté-Sénectère) 1849: 56; Werner leoptera) Psyche 90: 211-235. 1961: 799; 1983b: 234 Distribution. Cuba (introduced?), Hispaniola, 153. FAMILY ADERIDAE Jamaica, Puerto Rico, Virgin Islands, The ant-like leaf beetles 164 Arthropods of Florida and Neighboring Land Areas, Vol. 18

This family occurs in North, Central, and South SUPERFAMILY CHRYSOMELOIDEA America, and in the Greater and Lesser Antilles. 155. FAMILY CERAMBYCIDAE The larvae live with rotten wood, and adults are The long-horned beetles most sommonly found crawling on leaves of plants This is a very large family, and it is one of the at dusk. best studied in Cuba. The larvae bore into living and dead plant tissue; in stems, roots, cambium TRIBE ADERINI and sapwood. Adults are often found on flowers, SUBTRIBE ADERINA feeding on pollen and nectar. They are most often collected on flowers, by beating or sweeping vegeta- Stephens 1829: 57 tion, and at lights at night. The classification and A. brunnipennis (LeConte) 1875: 176 synonyms used here generally follow those in Monné Distribution. Cuba, Bahamas, USA (TX-PA- and Giesbert (1995). See Chemsak et al. (1992) for FL) a diferent classification. Zayas (1957) revised the Cuba localities. prionines of Cuba, and (1975) reviewed the entire Collections. cerambycid fauna of Cuba, and gives some keys for identification. Mészáros (1977) and Fernandez SUBTRIBE OLOTELINA García and Fiol Ferrer (1999) give data on host plants and these data are too extensive to repeat Gymnoganascus Werner 1990: 193 here. Monné and Giesbert (1995) is the source of G. stephani Werner 1990: 193 distributions in and beyond Cuba, except where they Distribution. Cuba, México, USA (TX-OK- do not use the synonyms in Zayas (1975). Browne KY-FL) and Peck (1996) have examined the distribution of Cuba localities. Cuban species into Islands and south Collections. Florida.

REFERENCES SUBFAMILY PARANDRINAE Werner, F.G. 1990. Revision of the Aderidae of TRIBE PARANDRINI eastern North America. J. New York Ent. Soc. 98: 187-232. Parandra Latreille 1804: 252 P. cribrata Thomson 1861: 87; Gundlach 1891: 243; 154. FAMILY SCRAPTIIDAE Zayas 1957: 159; 1975: 18 The false flower beetles Distribution. Cuba, Hispaniola, Puerto Rico This is a nearly worldwide family, and occurs Cuba localities. Widespread. At lights especially in drier areas. Adults often occur on flow- Collections. Gundlach no. 1727. Zayas ers and larvae in decaying wood and leaf litter. P. cubaecola Chevrolat 1862: 275; Gundlach 1891: 243; Zayas 1957: 159; 1975: 19 SUBFAMILY SCRAPTIINAE Distribution. Cuba, and Puerto Rico, accord- TRIBE SCRAPTIINI ing to Monné and Giesbert 1995 Cuba localities. Gr: Guamá. Ho: Sierra Scraptia Latreille 1807: 199 Cristal. In trunks of pine trees S. sp., Leng and Mutchler 1917: 215 Collections. Gundlach no. 432. Zayas Distribution. Cuba. Endemic (P. laevis Latreille 1804: 253) Cuba localities. Distribution. Hispaniola, Jamaica Collections. Cuba localities. Not recognized by Zayas (1975) for Cuba, but listed in Monné and SUBFAMILY ANASPIDINAE Giesbert (1995) without documentation TRIBE PENTARIINI Collections.

Pentaria Mulsant 1856: 391 SUBFAMILY PRIONINAE P. multipilis Ray 1839: 303 TRIBE MACROTOMINI Distribution. Cuba. Endemic Cuba localities. Stenodontes Audinet-Serville 1832: 173 Collections. S. (S.) chevrolati Gahan 1890: 23; Zayas 1957: 160; 1975: 21; Chemsak 1969: 185 A Checklist of the Beetles of Cuba 165

S. damicornis (L.); Jacquelin du Val 1857: 257; Collections. Gundlach. Zayas (neotype) Chevrolat 1862: 273; Gundlach 1891: 241 S. cubensis Casey 1912: 224 TRIBE SOLENOPTERINI Distribution. Cuba, Bahamas, USA (FL) Cuba localities. Widespread. At lights in Derancistrus Audinet-Serville 1832: 181 spring D. anthracinus (Gahan) 1890: 31; Zayas 1975: 32 Collections. Brno. Gundlach no. 446. Pitts- Distribution. Cuba, Hispaniola burgh. Zayas Cuba localities. SC: Gran Piedra S. (Mallodon) bituberculatus Beauvois 1805: 216; Collections. Gundlach no. 668. Zayas Zayas 1957: 161; 1975: 22 D. coeruleus Lameere 1912: 170; Zayas 1975: 321 Mallodon carptor Chevrolat 1862: 273; Gundlach Distribution. Cuba. Endemic 1891: 240 Cuba localities. SC: Loma del Gato;. Pico Distribution. Cuba, Jamaica, Hispaniola, Turquino Puerto Rico Collections. Zayas Cuba localities. M: Cárdenas Collections. Gundlach no. 923. Zayas Elateropsis Chevrolat 1862: 269 E. fulvipes Chevrolat 1838: 282 (Solenoptera); Strongylaspis Thomson 1860: 313 Jacquelin du Val 1857: 260; Gundlach 1891: S. corticarius Erichson in Schomberg 1848: 571; 237; Zayas 1957: 172; 1975: 34 Zayas 1957: 160; 1975: 20 Elateropsis fimbriatus Chevrolat 1862: 270; S. scobinatus Chevrolat 1860: 315; Gundlach Gundlach 1891: 237; Zayas 1957: 172 1891: 239 Elateropsis venustus Chevrolat 1862: 270; Distribution. Cuba, Jamaica, Dominica, Gundlach 1891: 237; Zayas 1957: 171 México to n South America, USA (FL) Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Gr: Cabo Cruz. Gu: Yateras. Collections. Gundlach no. 1100. Pittsburgh. Ha: littoral. Ho: Daiquirí. M: Palenque. SC: Zayas Ciudamar Collections. Gundlach no. 997a (E. venustus); TRIBE CALLIPOGONINI no. 779b (E. fimbriatus).; no. 779c (E. fulvipes). Zayas Callipogon Audinet-Serville 1832: 140 E. lineatus (Linnaeus) 1758: 389; Jacquelin du Val C. (Callomegas) sericeum (Olivier) 1795: 16; 1857: 260 (Solenoptera); Gundlach 1891: 236; Jacquelin du Val 1856: 107; Chevrolat 1862: Zayas 1957: 169; 1975: 38 274; Gundlach 1891: 242 (Orthomegas); Zayas E.subpunctatus Chevrolat 1862: 271; Gundlach 1957: 164 1891: 238 (E. subpunetatus) Distribution. Cuba, Hispaniola, Puerto Rico E. punctatus Gahan 1890: 271; Zayas 1957: 171; Cuba localities. Ha. Ca 1975: 39 Collections. Gundlach no. 442. Zayas Distribution. Cuba, Jamaica, Guadeloupe? C. (Dendroblaptus) barbiflavum Chevrolat 1864a: Cuba localities. Widespread 180; Gundlach 1891: 241; Zayas 1957: 164; Collections. Gundlach no. 551. Zayas 1975: 23 (E. quinquenotatus Chevrolat 1862: 271; Zayas Distribution. Cuba, Bahamas 1957: 157; 1975: 37) Cuba localities. IJ: Nueva Gerona. Ho: Si- Distribution. Cuba?, Jamaica erra Cristal. PR. In pine trees; hosts: see Cuba localities. Mészáros 1977) Collections. Collections. Gundlach no. 1585. New York. E. reticulatus Gahan 1890: 30; Zayas 1957: 173; Philadelphia (Poey no. 956). Pittsburgh. Zayas 1975: 36 Distribution. Cuba. Endemic Cubaecola Lameere 1912: 164. Endemic genus Cuba localities. Ho: Sierra Cristal C. hoploderoides Lameere 1912: 164; Zayas 1957: Collections. Zayas 162; 1975: 25 E. rugosus (Gahan) 1890: 28; Zayas 1957: 170 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, USA (FL) Cuba localities. Ca: Florida. PR: Cuba localities. IJ: Nueva Gerona Guanahacabibes. IJ Collections. Carnegie. Washington 166 Arthropods of Florida and Neighboring Land Areas, Vol. 18

E. scabrosus Gahan 1890: 29; Zayas 1957: 170; [S. thomae (Linnaeus) 1767: 623 (Cerambyx); see Chemsak 1969: 186 (Derancistrus) Galileo and Martins1993: 429] Elateropsis fuliginosus Fabricius, Chevrolat 1862: Distribution. Puerto Rico, Virgin Islands; not 271; Jacquelin du Val 1856: 108; Gundlach Cuba 1891: 238 Elateropsis castaneous Zayas 1975: 40. PR: Maria TRIBE PRIONINI la Gorda, Guanahacabibes (t.l.) Distribution. Cuba, USA (FL) Orthosoma Audinet-Serville 1832: 155 Cuba localities. The most abundant prionine; [O. brunneum (Forster) 1771: 37 (Derobrachus); widespread in Cuba Zayas 1975: 44] Collections. Gundlach no. 265. Zayas Distribution. Not Cuba; Hispaniola, E. sericeiventris Chevrolat 1862: 270; Gundlach Guadeloupe, USA 1891: 238; Zayas 1957: 170; 1975: 35 Distribution. Cuba. Endemic TRIBE MEROSCELISINI Cuba localities. M: Cárdenas. PR: Taco-Taco Collections. Gundlach no. 511. Washington Monodesmus Audinet-Serville 1832: 160 M. atratus Fisher 1932: 3; Zayas 1957: 174; 1975: Prosternodes Thomson 1960: 288 42 P. cinnamipennis (Chevrolat) 1838: 281 Distribution. Cuba. Endemic (Solenoptera); Jacquelin du Val 1856: 258; Cuba localities. PR: Sierra Rangel, Rancho Gundlach 1891: 239; Zayas 1957: 166; Zayas Mundito (t.l.). SS: Trinidad Mts, Buenos Aires 1975: 28 Collections. Washington (type). Zayas Distribution. Cuba, Bahamas M. callidioides Audinet-Serville 1832: 161; Cuba localities. Widespread, coastal Chevrolat 1862: 268; Gundlach 1891: 236 (M. Collections. Gundlach no. 44. Zayas callidivides); Zayas 1957: 174; Zayas 1975: 43; P. cubanus Zayas 1975: 29 Chemsak 1969: 186 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Cayman Is- Cuba localities. Gu: Yateras (t.l.). Ho: lands Cupeyal Cuba localities. Widespread Collections. Zayas (type) Collections. Brno. Gundlach no. 283. Pitts- burgh. Zayas Solenoptera Audinet-Serville 1832: 183 [S. bilineata Fabricius 1775: 162]; Galileo and Mar- SUBFAMILY ASEMINAE tins 1993: 432 TRIBE ASEMINI Distribution. Not Cuba; but Virgin Islands, Puerto Rico Arhopalus Audinet-Serville 1834: 77 S. fraudulenta Galileo and Martins 1993: 445 A. cubensis (Mutchler) 1914: 346 (Criocephalus); Distribution. Cuba. Endemic Zayas 1975: 49 Cuba localities. None given. Cuba material Distribution. Cuba. Endemic misidentified as S. scutellata Gahan by Zayas Cuba localities. Gu: Guantánamo (t.l.). Ho: 1957: 165; 1975: 28] Mayarí. SC: Gran Piedra Collections. Brussels. Leiden. São Paulo Collections. New York (type). Zayas [S. metallescens Thomson 1860: 306; Zayas 1975: 31] SUBFAMILY Distribution. Not Cuba; but Lesser Antilles TRIBE SMODICINI S. parandroides Lameere 1885: XII; Zayas 1957: 168; 1975: 30; Galileo and Martins 1993: 446 Metaphrenon Martins 1975: 330 Distribution. Cuba. Endemic M. impressicolle (Lacordaire) 1869: 146 Cuba localities. Gu: Yateras. Ho: Cupeyal; (Smodicum); Zayas 1975: 46 El Johnson; Moa; Piloto Distribution. Cuba, Hispaniola, Puerto Rico Collections. Zayas Cuba localities. Gu: Imias. M: Cienaga de [S. scutellata Gahan 1890: 25; Zayas 1957: 165; Zapata 1975: 28] Collections. Zayas Distribution. Not Cuba; Hispaniola A Checklist of the Beetles of Cuba 167

TRIBE OEMINI Cuba localities. None known or reported

Malacopterus Serville 1833: 565 Enosmaeus Thomson 1878: 9 M. tenellus (Fabricius) 1801: 335; Zayas 1975: 51 E. cubanus Thomson 1878: 10; Zayas 1975: 48 Distribution. Cuba, sw USA (not FL) to Distribution. Cuba, Hispaniola, Puerto Rico South America Cuba localities. Cuba localities. Widespread Collections. Collections. Gundlach no. 1628. Zayas TRIBE EBURIINI TRIBE METHINI Eburia Lepeltier and Audinet-Serville in Methia Newman 1842: 418 Lacordaire 1830: 77 M. necydalea Fabricius 1798: 148; Zayas 1975: 52; E. cinereopilosa Fisher 1932: 13; Zayas 1975: 60; Chemsak 1969: 196; Philips and Ivie 1998: 72 Chemsak et al. 1992: 3 M. punctata LeConte 1873: 240; Zayas 1975: 53 Distribution. Cuba, Guatemala, USA (FL) M. pusilla Chevrolat 1862: 256, not Newman Cuba localities. Ca: Camaguëy; Santa Lucía. 1840 Gu: Guantánamo (t.l.). Ho: Moa; Sierra Cristal. M. pusilla Newman 1840: 18; Gundlach 1891: M: Peninsula de Zapata. PR: Guanahacabibes. 218 SS: Gavilanes Distribution. Cuba, Bahamas, Cayman Is- Collections. Washington (type). Zayas lands, Hispaniola, Puerto Rico, Virgin Islands, E. confusa Zayas 1975: 61 Lesser Antilles, Mexico, USA (TX-FL-VA) Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. SC: Daiquiri (t.l.); Juraguá Collections. Brno. Gundlach no. 52. Havana. Collections. Zayas (type) London. San Francisco. Zayas E. consobrina Jacquelin du Val 1857: 265; Chevrolat M. taina Zayas 1975: 53; Philips and Ivie 1998: 81 1862: 267; Gundlach 1891: 233 (Coeleburia); Distribution. Cuba. Endemic Zayas 1975: 62 Cuba localities. Ho: Sierra Cristal, Las Mulas Distribution. Cuba, Jamaica (t.l.) Cuba localities. Widespread Collections. Zayas (type) Collections. Gundlach no. 438. Zayas E. cubae Fisher 1932: 16; Zayas 1975: 63 Tessaropa Haldeman 1847: 375 Distribution. Cuba, Guatemala, se México T. luctuosa Zayas 1975: 54; Philips and Ivie 1998: Cuba localities. Ca: Cuabitas. CA: Baraguá 81 (t.l.). Gu: Baracoa. Ho: Cayo Mambi, Central Distribution. Cuba. Endemic Tanamo. M: Cienaga de Zapata; Pan de Cuba localities. SC: Sierra Maestra, El Matanzas Brazón del Rio Yara (t.l.) Collections. Washington (type). Zayas Collections. Zayas (type) E. didyma Olivier 1795: 125; Zayas 1975: 64 E. pulverea Chevrolat 1862: 266 TRIBE Distribution. Cuba, Hispaniola Cuba localities. Widespread Serville 1833: 572 Collections. Gundlach no. 1101. Zayas A. surinamum (Linnaeus) 1767: 632; Chevrolat E. elongata Fisher 1932: 18; Zayas 1975: 63 1862: 264; Gundlach 1891: 230; Zayas 1975: Distribution. Cuba. Endemic 55 Cuba localities. Gu: Florida Blanca; Distribution. Cuba, West Indies, México to Guantánamo (t.l.); Yateras. SC: Alto Songo South America, sw USA (not FL) Collections. New York (type). Zayas Cuba localities. M: Cárdenas E. fisheri Russo 1930: 144 Collections. Gundlach no. 469 Distribution. Cuba, Hispaniola Cuba localities. M: Zapata. PR: Araespor Thomson 1873: 14 Guanahacabibes; Rangel, Rancho Mundito. SC: [A. longicollis Thomson 1878: 15; Zayas 1975: 56] Loma del Gato Distribution. Fiji Islands; not Cuba (an er- Collections. Gundlach (as Chion). Zayas ror or a misidentification) 168 Arthropods of Florida and Neighboring Land Areas, Vol. 18

E. longicornis Fisher 1932: 9; Zayas 1975: 66 Cuba localities. Widespread. A pest of citrus Distribution. Cuba. Endemic Collections. Gundlach no. 275. Zayas Cuba localities. CA: Baraguá; Central Florida (t.l.); El Roble. Gu: Baracoa. SS: Eburodacrys White 1853: 93 Gavilanes E. havanensis Chevrolat 1862: 267; Gundlach 1891: Collections. Gundlach no. 472. Washington 234; Zayas 1975: 75 (type). Zayas Distribution. Cuba, México to South America E. marginalis Fisher 1947: 29; Zayas 1975: 67 Cuba localities. M: Cárdenas. SC: Loma del Distribution. Cuba. Endemic Gato. SS: Lomas de Trinidad Cuba localities. Gu: Yateras. M: Central Collections. Gundlach no. 512. Zayas Mercedes (t.l.) Collections. Washington (type). Zayas TRIBE HESPEROPHANINI [E. octomaculata Chevrolat 1862: 265; Zayas 1975: 68] Stromatium Audinet-Serville 1834: 80 Distribution. Cuba a doubtful record; S. fulvum (Villers) 1889: 25; Chevrolat 1962: 265; Guadeloupe, Dominica, Barbados Zayas 1975: 57 E. pellacia Zayas 1975: 69 S. unicolor Olivier, Gundlach 1891: 231 Distribution. Cuba. Endemic Distribution. Cuba (introduced), North Cuba localities. Ca: Santa Lucía (t.l.). Ci: America; native to Europe; cosmopolitan Cumanayagua Cuba localities. Collections. Zayas (type) Collections. Gundlach no. 819 E. pinarensis Zayas 1975: 69 Distribution. Cuba. Endemic TRIBE Cuba localities. PR: Ancón, San Vincente; Viñales (t.l.) Anelaphus Linsley 1936: 464 Collections. Zayas (type) A. cinereus (Olivier) 1795: 69 (Elaphidion); Zayas E. quadrigeminata (Say) 1827: 275; Chevrolat 1975: 95 (as a synonym of A. nanus (Fabri- 1862: 265; Gundlach 1891: 232: Zayas 1975: cius)); Chemsak 1969: 186 (A. subtropicus 71 Casey) Distribution. Cuba (introduced), e USA Distribution. Cuba, Jamaica, Hispaniola, Cuba localities. Bahamas, USA (FL) Collections. Cuba localities. Ha: Cojimar. M: Cárdenas E. quadrimaculata (Linnaeus) 1767: 626 Collections. Brno. Gundlach no. 1826 E. binodosa Gahan 1895: 95 A. guttiventre (Chevrolat) 1862: 261 (Elaphidion); Cerambyx rhamphygea Linnaeus 1767: 633 Gundlach 1891: 335; Zayas 1975: 89; Chemsak Distribution. Cuba, Puerto Rico, Virgin Is- 1969: 186 (Elaphidionoides) lands Distribution. Cuba. Endemic Cuba localities. Cuba localities. Widespread Collections. Collections. Brno. Gundlach no. 492. Zayas E. ramsdemi Fisher 1932: 14; Zayas 1975: 71 (A. inermis Newman 1840: 29) Distribution. Cuba. Endemic Distribution. Bahamas, Greater Antilles, Cuba localities. Gu: Guantánamo (t.l.) México to Honduras, USA (GA-TX) Collections. New York (type) Cuba localities. E. sericia Sallé 1855: 271; Zayas 1975: 72 Collections. Distribution. Cuba, Jamaica, Hispaniola A. nanus (Fabricius) 1792: 300 (Elaphidion); Zayas Cuba localities. Gu: Finca San Carlos; 1975: 95 Yateras Distribution. Cuba, Puerto Rico, Virgin Is- Collections. Zayas lands E. stigma (Olivier) 1795: 126; Jacquelin du Val Cuba localities. Widespread 1857: 265; Zayas 1975: 73 Collections. Zayas E. duvalii Chevrolat 1862: 266; Gundlach 1891: A. parallelus (Newman) 1840: 29 233 Elaphidion pulverulentum Olivier 1895: 69; Distribution. Cuba, Bahamas, Hispaniola, Chevrolat 1862: 262; Zayas 1975: 99 México to Costa Rica, USA (FL) Distribution. Cuba, e USA A Checklist of the Beetles of Cuba 169

Cuba localities. E. conspersum Newman 1841: 110 Collections. Distribution. Cuba, Bahamas, Hispaniola, [A. villosus (Fabricius) 1792: 302; Zayas 1975: 107] Puerto Rico, Virgin Islands, Lesser Antilles, Distribution. Cuba (accidental record?), USA Curaçao, (eastern states-TX) Cuba localities. Cuba localities. Collections. Collections. E. cristalensis Zayas 1975: 82 Distribution. Cuba. Endemic Curtomerus Stephens 1839: 275 Cuba localities. Ca: Aguada. Ho: Sierra C. fasciatus (Fisher) 1932: 49 (Cylindera); Zayas Cristal (t.l.). M: Cienaga de Zapata, San Blas 1975: 117 Collections. Zayas (type) Distribution. Cuba, USA (FL) E. cubae Fisher 1932: 27; Zayas 1975: 83 Cuba localities. Ca: Nuevitas. Gr: Cabo Cruz. Distribution. Cuba. Endemic Gu: Guantánamo (t.l.) Cuba localities. Ci: Cayamas (t.l.). Gu: Imias. Collections. Washington (type). Zayas SC: Daiquiri C. flavus (Fabricius) 1775: 191; Zayas 1975: 118 Collections. New York. Washington (type). (Cylindera) Zayas Lampromerus pilicorne Fabricius 1792: 327; E. depressum Zayas 1975: 83 Gundlach 1891: 228 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Jamaica, Cuba localities. Gu: Baracoa, Loma de la Grand Cayman, Hispaniola, Mona, Puerto Rico, Farola; Imias; Jaragua (t.l.) Lesser Antilles, México to South America, USA Collections. Zayas (type) (FL, HI), Tahiti E. difflatus Zayas 1975: 84 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 1103 (L. Cuba localities. Ca: Santa Lucía (t.l.) pilicornis); Zayas Collections. Zayas (type) C. glabrus (Fisher) 1932: 51 (Cylindera); Zayas E. elongatum Fisher 1942: 14; Zayas 1975: 85 1975: 119 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: coast below Pico Cuba localities. Ci: Cayamas (t.l.) Turquino (t.l.) Collections. Washington (type) Collections. Harvard (type) E. excelsum Gahan 1895: 101; Zayas 1975: 86 Elaphidion Audinet-Serville 1834: 66 Distribution. Cuba, Guadeloupe E. albosignatum Chevrolat 1862: 259; Gundlach Cuba localities. PR: La Pedrera de Mendoza. 1891: 223; Zayas 1975: 78 SS: Lomas de Trinidad, Buenos Aires Distribution. Cuba. Endemic Collections. Zayas Cuba localities. Ho: Sierra Cristal. SC: Brazo E. fasciatum Fisher 1942: 13; Zayas 1975: 87 del Cauto Distribution. Cuba, Bahamas Collections. Gundlach no. 1423. Zayas Cuba localities. Widespread. In fallen E. angustatum Zayas 1975: 79 branches of citrus Distribution. Cuba. Endemic Collections. Zayas. Cuba localities. Ho: Sierra Cristal (t.l.) E. frisevestitum Fisher 1942: 13; Zayas 1975: 88 Collections. Zayas (type) Distribution. Cuba. Endemic E. cayamae Fisher 1932: 24; Zayas 1975: 80 Cuba localities. CA: Jaronú (t.l.). M: Penin- Distribution. Cuba. Endemic sula de Zapata. PR: Guanahacabibes Cuba localities. Ci: Cayamas (t.l.). M: Colón. Collections. Harvard (type). Zayas SC: Daiquiri; considered to be a common pest E. gracilis Fisher 1932: 43; Zayas 1975: 89 of citrus (Vàzquez et al. 1992: 44) Distribution. Cuba. Endemic Collections. Cambridge. Washington (type) Cuba localities. Ca: Camaguey (t.l.). Ci: E. cinnabarinum Fisher 1942: 11; Zayas 1975: 81 Cayamas; Cienfuegos. Ho: Sierra Cristal Distribution. Cuba. Endemic Collections. New York. Washington (type). Cuba localities. Ci: Soledad (t.l.). Ha: Jibacoa Zayas Collections. Harvard (type). Zayas E. irroratum (Linnaeus) 1767: 633 (Cerambyx); Jacquelin du Val 1857: 266; Gundlach 1891: 170 Arthropods of Florida and Neighboring Land Areas, Vol. 18

221; Zayas 1975: 91; Chemsak 1969: 186 Distribution. Greater Antilles Distribution. Cuba, West Indies (where?), Cuba localities. M: Cárdenas México to Costa Rica, USA (FL) Collections. Gundlach no. 368 Cuba localities. Widespread E. tuberculicolle Fisher 1932: 25; Zayas 1975: 104 Collections. Brno. Gundlach no. 282. Pitts- Distribution. Cuba. Endemic burgh. Zayas Cuba localities. Ca: Santa Lucía. Ci: E. jibacoense Zayas 1975: 92 Cayamas (t.l.). Ha: Marianao. M: Zapata, San Distribution. Cuba. Endemic Blas Cuba localities. Ha: Jibacoa (t.l.) Collections. Zayas Collections. Zayas (type) E. uncinatum Zayas 1975: 105 E. lanatum Chevrolat 1862: 200; Gundlach 1891: Distribution. Cuba. Endemic 224; Zayas 1975: 93; Chemsak 1969: 186 Cuba localities. Ho: Moa, Piloto; Sierra Distribution. Cuba. Endemic Cristal (t.l) Cuba localities. Widespread Collections. Zayas (type) Collections. Brno. Gundlach no. 550. Zayas E. unispinosum Fisher 1942: 9; Zayas 1975: 106 E. monticola Fisher 1935: 191; Zayas 1975: 93 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Baitiquirí (t.l.) Cuba localities. Gu: Florida Blanca. Ho: Si- Collections. Harvard (type) erra Cristal. M: San Blas, Zapata; Pan de E. variolosum Fisher 1947: 32; Zayas 1975: 106 Matanzas (t.l.). PR: Guanahacabibes. SC: Alto Distribution. Cuba. Endemic Songo Cuba localities. Ho: Moa (t.l.) Collections. Zayas Collections. Washington (type) E. niveonotatum Zayas 1975: 96 Distribution. Cuba. Endemic Linsleyonides Skiles 1985: 316 Cuba localities. Ho: Sierra Cristal (t.l.). SC: L. albomaculatus (Champlain and Knull) 1922: 146 Daiquiri (Elaphidion); Zayas 1975: 78 Collections. Zayas (type) Distribution. Cuba, USA (FL) E. pauropilosum Zayas 1979: 98 Cuba localities. Gr: Guamá; Jiguaní. SS: Los Distribution. Cuba. Endemic Gavilanes Cuba localities. CA: Central Baraguá (t.l.) Collections. Zayas Collections. Zayas (type) E. quadrituberculatum Chevrolat 1862: 259; Nesanoplium Chemsak 1966: 214 Gundlach 1891: 222; Zayas 1975: 99 (N. puberulum Fleutiaux and Salle 1889: 464 Distribution. Cuba. Endemic Distribution. West Indies (widely distributed) Cuba localities. Cuba localities. Collections. Collections. E. scaramuzzai Fisher 1951: 77; Zayas 1975: 100 Distribution. Cuba. Endemic Nesostizocera Linsley 1961: 170 Cuba localities. M: Central Mercedes (t.l.) N. insulana (Gahan) 1895: 106 (Periboeum); Zayas Collections. New York. Washington (type) 1975: 112 (Stizocera) E. signaticolle Chevrolat 1862: 261; Gundlach 1891: Distribution. Cuba, Jamaica 225; Zayas 1975: 101 Cuba localities. Widespread Distribution. Cuba, Bahamas, Hispaniola Collections. Zayas Cuba localities. Widespread, coastal N. poeyi (Guérin de Meneville in Chevrolat) 1838: Collections. Gundlach no. 490. Pittsburgh. 284 (Elaphidion); Jacquelin du Val 1857: 267; Zayas Gundlach 1891: 235 (Nephalius); Zayas 1975: E. splendidum Fisher 1932: 19 113 (Stizocera) Distribution. Cuba, Hispaniola Distribution. Cuba, Panamá, north and cen- Cuba localities. Ci: California. M: Zapata, tral South America Santo Tomas. PR: Paredón de los Mendozas, Cuba localities. Ci: Cayamas. M: Cardenas; Guane; Rancho Mundito Playa Girón Collections. Zayas Collections. Gundlach no. 19. Washington. E. tomentosum Chevrolat 1862: 260; Gundlach Zayas 1891: 223; Zayas 1975: 103 A Checklist of the Beetles of Cuba 171

Parelaphidion Skiles 1985: 310 M. elongatus Fisher 1932: 54; Zayas 1975: 121 P. mutatum (Gahan) 1870: 32 (Elaphidion); Zayas Distribution. Cuba. Endemic 1975: 94 Cuba localities. Ha: (t.l.). PR. VC Elaphidion tomentosum Chevrolat 1826: 260 in Collections. Washington (type). Zayas part; Gundlach 1891: 223 M. exiguus Zayas 1974: 122 Distribution. Cuba, Puerto Rico, se USA (FL) Distribution. Cuba. Endemic Cuba localities. Ci: Cumanayagua. Ha: Cuba localities. SC: Pico Turquino; Siboney Jibacoa. Ho: Moa. PR: Viñales. A pest of citrus (t.l.) Collections. Zayas Collections. Zayas (type)

Pyrassa Pascoe 1866: 481 TRIBE CURIINI P. jaumei (Fisher) 1935: 193 (Anepsyra); Zayas 1975: 111 Plectromerus Haldeman 1847: 43 Distribution. Cuba. Endemic P. acunai (Fisher) 1936: 344 (Pentomacrus); Zayas Cuba localities. Ca: Santa Lucía, Nuevitas. 1975: 126 Ci: Santa Clara, Coralillo (t.l.). Ha: Cojímar; Distribution. Cuba. Endemic Puente Almendares Cuba localities. Ho: Sierra Cristal. SC: Gran Collections. Washington (type). Zayas Piedra; Loma del Gato (t.l.); Pico Turquino. SS: Buenos Aires; Topes de Collantes Stenosphenus Haldeman 1847: 39 Collections. Havana. Washington (type). S. insulicola Fisher 1942: 24; Zayas 1975: 159 Zayas Distribution. Cuba. Endemic P. dentipes (Olivier) 1790: 268; Zayas 1975: 123 Cuba localities. SC: coast below Pico Distribution. Cuba, USA (LA-FL) Turquino (t.l.) Cuba localities. Widespread Collections. Harvard (type) Collections. Zayas P. exis Zayas 1975: 123 TRIBE IBIDIONINI Distribution. Cuba. Endemic Cuba localities. Gr: Jiguaní. Ho: Sierra Heterachthes Newman 1940: 9 Cristal. PR: Pan de Guajaibón. SC: Loma del H. ebenus Newman 1840: 9; Zayas 1975: 114 Gato (t.l.) Distribution. Cuba, Bahamas, Puerto Rico, Collections. Zayas (type) México to South America, USA (FL-TX) P. fasciatus (Gahan) 1895: 109 (Pentomacrus); Cuba localities. PR: Viñales Zayas 1975: 127 Collections. Zayas Distribution. Cuba, West Indies to Grenada Cuba localities. M: Cienaga de Zapata. PR: Trinoplon Zayas 1975: 115. Endemic genus Guanahacabibes; Paredón de los Mendoza T. bicolor Zayas 1975: 116 Collections. Zayas Distribution. Cuba. Endemic P. ornatus Fisher 1947: 34; Zayas 1975: 124 Cuba localities., M: Cienaga de Zapata, Playa Distribution. Cuba. Endemic Larga (t.l.) Cuba localities. Ho: Moa (t.l.) Collections. Zayas (type) Collections. Washington (type) P. pinicola Zayas 1975: 125 TRIBE CALLIDIOPINI Distribution. Cuba. Endemic Cuba localities. PR: Malas Aguas (t.l.) Merostenus White 1855: 24 Collections. Zayas (type) M. attenuatus Chevrolat 1862: 263; Gundlach 1891: P. punctatus (Fisher) 1932: 55 (Pentomacrus); 228; Zayas 1975: 120 Zayas 1975: 128 Distribution. Cuba, West Indies (where?) Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Ha: Santiago de las Vegas Collections. Gundlach no. 1021; Zayas (t.l.). Ho: Sierra Cristal M. decoratus Zayas 1975: 121 Collections. Havana (Bruner?). New York. Distribution. Cuba. Endemic Washington (type) Cuba localities. PR: Pan de Guajaibón (t.l.) Collections. Zayas (type) 172 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE MOLORCHINI TRIBE CLYTINI

Molorchus Fabricius 1792: 356 Euryscelis Dejean 1835: 331 M. bimaculatus Say 1823: 428 (E. suturalis Olivier 1795: 62) M. b. corni Haldeman 1847: 45 Distribution. Bahamas, Puerto Rico, Cay- M. ruficollis Gahan 1895: 110; Zayas 1975: 129 man Islands, USA (FL) M. sanguinicollis Chevrolat 1862: 257 Cuba localities. Distribution. Cuba, se USA Collections. Cuba localities. Collections. Neoclytus Thomson 1860: 67 N. acuminatus (Fabricius) 1775: 194; Zayas 1975: TRIBE RHINOTRAGINI 135. The redheaded ash borer Distribution. Cuba (introduced), native to e Bromiades Thomson 1864: 165 North America B. brachyptera (Chevrolat) 1838: 285 (Odontocera); Cuba localities. Probably introduced to Cuba Jacquelin du Val 1857: 269; Gundlach 1891: in lumber 221 (O. brachyptora); Fisher 1930: 13; Zayas Collections. Gundlach no. 1619 1975: 131; Chemsak 1969: 187 N. chevrolati (Laporte and Gory) 1835: 26 (Clytus); Distribution. Cuba, México, South America Jacquelin du Val 1857: 269; Gundlach 1891: Cuba localities. Widespread 229; Zayas 1975: 136 Collections. Brno. Gundlach no. 285. Zayas Distribution. Cuba. Endemic Cuba localities. Widespread Odontocera Serville 1833: 546 Collections. Gundlach no. 1458. Zayas O. josemartii Zayas 1956: 113 N. cordifer (Klug) 1829: 13 (Clytus); Zayas 1975: Distribution. Cuba. Endemic 135; Chemsak 1969: 187 Cuba localities. SC: Pico Joaquín, near Clytus rufescens Laporte and Gory 1835: 16; Turquino (t.l.) Jacquelin du Val 1857: 268 Collections. Zayas (type) Clytus devastator Laporte and Gory 1835: 17; Jacquelin du Val 1857: 268; Gundlach 1891: 229 TRIBE CALLICHROMATINI (Neoclytus) Distribution. Cuba, USA (FL-GA) Philematium Thomson 1864: 171 Cuba localities. Widespread P. femorale (Olivier) 1790: 297 (Cerambyx); Collections. Brno. Gundlach no. 259 (N. dev- Jacquelin du Val 1857: 262; Zayas 1975: 132 astator). Pittsburgh. Zayas Callichroma leucodactylum Chevrolat 1862: 257 N. jibacoense Zayas 1975: 137 Distribution. Cuba (introduced); native to Distribution. Cuba. Endemic Africa Cuba localities. Ha: Boca del Rio Jibacoa (t.l.) Cuba localities. Not known to be presently Collections. Zayas (type) established N. longipes (Drury) 1783: 84; Zayas 1975: 138 Collections. Clytus angulatum Fabricius 1792: 335; Chevrolat 1862: 531 Plinthocoelium Schmidt 1824: 300 Distribution. Cuba, Jamaica, USA (GA-FL) P. virens Drury 1873: 89 Cuba localities. None known Callichroma colombina (Guérin-Méneville) in Collections. Chevrolat 1838: 282; Jacquelin du Val 1857: 262; N. pubicollis Fisher 1932: 59; Zayas 1975: 139 Gundlach 1891: 220; Zayas 1975: 132 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, Hispaniola, Cuba localities. Gu: Baracoa; Guantánamo Jamaica (t.l.). SC: Siboney Cuba localities. Widespread. Important in Collections. Washington (type). Zayas that it attacks live jocuma trees in sub-littoral zone of north coast (listed as P. columbinum, Placosternus Hopping 1937: 447 Vàzquez et al. 1992: 45) P. difficilis (Chevrolat) 1862: 263 (Cyllene); Collections. Gundlach no. 251. Zayas Gundlach 1891: 229; Zayas 1975: 133 (Megacyllene) A Checklist of the Beetles of Cuba 173

Distribution. Cuba, Bahamas, México, USA Collections. Washington (type). Zayas (FL-CA) Cuba localities. Widespread TRIBE RHOPALOPHORINI Collections. Gundlach no. 466. Zayas Rhopalophora Audinet-Serville 1834: 100 TRIBE ANAGLYPTINI R. baracoana Zayas 1975: 162 Distribution. Cuba. Endemic Bates 1885: 303 Cuba localities. Gu: Baracoa (t.l.). IJ T. bruneri Fisher 1932: 60; Zayas 1975: 141 Collections. Gundlach no. 737. Zayas (type) Distribution. Cuba. Endemic Cuba localities. Gr: near Bayamesa, Sibe- TRIBE HETEROPSINI ria. SC: Pico Cuba; Pico Turquino (t.l.) Collections. Washington (type). Zayas Heterops Blanchard 1842: 51 T. cubae Fisher 1932: 63 H. dimidiatus (Chevrolat) 1838: 283 (Eburia); Distribution. Cuba. Endemic Jacquelin du Val 1857: 264; Gundlach 1891: Cuba localities. Ho: Pinares de Mayarí (t.l.) 231; Zayas 1975: 154 Collections. Washington (type) Eburia subangulata Chevrolat 1838: 283 T. elongatus Zayas 1975: 143 H. bipartita LaCordaire 1869: 128 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas, USA (FL) Cuba localities. PR: Sierra Rangel (t.l.) Cuba localities. Widespread Collections. Zayas (type) Collections. Cambridge. Gundlach no. 126. T. nivicinctus (Chevrolat) 1862: 264 (Tillomorpha); Zayas Gundlach 1891: 230; Zayas 1975: 143 Notes. This species seems to be part of a Batesian Distribution. Cuba. Endemic mimicry complex centered around species of Cuba localities. Central western Cuba Thonalmus (Lycidae) (Darlington 1938: 690) Collections. Gundlach no. 945. Zayas H. lanieri (Chevrolat) 1838: 283 (Eburia); Zayas T. pilosus Zayas 1975: 144 1975: 156; Chemsak 1969: 187 Distribution. Cuba. Endemic Eriphus lanieri Jacquelin du Val 1856: 263 (not Cuba localities. Ho: Nicaro (t.l.) Chevrolat) Collections. Zayas (type) Eburia lasnieri Chevrolat 1862: 264; Gundlach T. rufipes Fisher 1942: 20; Zayas 1975: 145 1891: 230 Distribution. Cuba. Endemic H. bicolor Fisher 1936: 346 (SC: Pico Turquino, Cuba localities. Ci: Soledad (t.l.). SC: Loma t.l.; Washington, type) del Gato H. duvali Fisher 1947: 35 (SC: Jarahueca, t.l.; Collections. Harvard (type). Washington Washington, type) (paratype). Zayas H. cubaecola Fisher 1947: 36. (PR: Cayos de San Felipe, t.l.; Washington, type) TRIBE TILLOMORPHINI Distribution. Cuba. Endemic Cuba localities. Ci: Soledad (t.l.). Gr: Calliclytus Fisher 1932: 65. Endemic genus Bueycitos. Ho: Sierra Cristal. M: Cárdenas. SS: C. schwarzi Fisher 1932: 66; Zayas 1975: 146 Lomas de Trinidad Tillomorpha nivicincta Chevrolat, Gundlach Collections. Brno. Gundlach no. 18. Cam- 1891: 230 bridge. Washington Distribution. Cuba. Endemic H. loreyi (Duponchel) 1837: 309; Zayas 1975: 158 Cuba localities. Ci: Cayamas (t.l.). .M: Distribution. Cuba. Endemic Cárdenas Cuba localities. Gu: Guantánamo Collections. Gundlach no. 745 (Washington Collections. Zayas (type) Neopoeciloderma Monné and Martins 1981: Pentanodes Schaeffer 1904: 222 186 P. albofasciatus Fisher 1932: 64; Zayas 1975: 140 N. lepturoides (Jacquelin du Val) 1857: 267 Distribution. Cuba. Endemic (Poeciloderma); Gundlach 1891: 219; Zayas Cuba localities. M: Cardenas (t.l.); Cienaga 1975: 160 de Zapata Distribution. Cuba, Hispaniola 174 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Widespread TRIBE TORNEUTINI Collections. Gundlach no. 429. Zayas. 90Chevrolat 1858: 210 Chlorida Audinet-Serville 1834: 31 C. festiva (Linnaeus) 1758: 389; Jacquelin du Val Trichrous Chevrolat 1858: 210 1857: 261; Chevrolat 1862: 267; Gundlach 1891: T. brevicornis Zayas 1975: 148 235; Zayas 1975: 59 Distribution. Cuba. Endemic Distribution. Cuba, West Indies (where?), Cuba localities. Ho: Moa (t.l.) Central and South America, USA (FL) Collections. Zayas Cuba localities. Widespread T. dimidiatipennis (Chevrolat) 1838: 282 (Eriphus); Collections. Gundlach no. 254. Pittsburgh. Jacquelin du Val 1857: 264; Gundlach 1891: Zayas 227; Zayas 1975: 149; Chemsak 1969: 187 Distribution. Cuba. Endemic Knulliana Linsley 1962: 109 Cuba localities. Widespread (K. cincta (Drury) 1873: 85 (Chion); Zayas 1975: Collections. Brno. Cambridge. Gundlach no. 58) 258. Zayas Distribution. Cuba?; Bahamas, México, se Notes. This species seems to be part of a Batesian USA mimicry complex centered around species of Cuba localities. Thonalmus (Lycidae) (Darlington 1938: 691) Collections. Gundlach no. 726. T. divisus Chevrolat 1858: 210; Gundlach 1891: (misidentification of Eburia fisheri Russo) 227; Zayas 1975: 150 Distribution. Cuba. Endemic SUBFAMILY Cuba localities. Widespread TRIBE PARMENIINI Collections. Cambridge. Gundlach no. 796. Zayas Cupeyalia Zayas 1975: 167. Endemic genus Notes. This species seems to be part of a Batesian C. subterranea Zayas 1975: 168 mimicry complex centered around species of Distribution. Cuba. Endemic Thonalmus (Lycidae) (Darlington 1938: 690) Cuba localities. Gu: Yateras (t.l.). Ho: T. irroratus (Olivier) 1795: 70; Chemsak 1969: 187 Cupeyal Distribution. Cuba, Hispaniola Collections. Zayas (type) Cuba localities. Gr: Belic Collections. Brno Maisi Zayas 1975: 168. Endemic genus T. pilipennis Chevrolat 1862: 262; Zayas 1975: 151; M. fisheri Zayas 1975: 169 Gundlach 1891: 227; Chemsak 1969: 187 Distribution. Cuba. Endemic Distribution. Cuba, Bahamas Cuba localities. Gu: Maisi (t.l.) Cuba localities. Widespread Collections. Zayas (type) Collections. Brno. Cambridge. Gundlach no. 479. Zayas Nanilla Fleutiaux and Salle 1889: 476 Notes. This species seems to be part of a Batesian N. globosa Zayas 1975: 171 mimicry complex centered around species of Distribution. Cuba. Endemic Thonalmus (Lycidae) (Darlington 1938: 690) Cuba localities. SC: Gran Piedra (t.l.) T. vittatus Fisher 1932: 69; Zayas 1975: 152 Collections. Zayas (type) Distribution. Cuba. Endemic N. terrestris Zayas 1975: 172 Cuba localities. PR: Sierra Rangel, Las Distribution. Cuba. Endemic Animas (t.l.) Cuba localities. Ho: Sierra Cristal, alto de Collections. Washington (type) Las Mulas (t.l.). PR: Viñales Collections. Zayas (type) TRIBE AGALLISSINI N. tuberculata Fisher 1935: 196; Zayas 1975: 173 Distribution. Cuba. Endemic Zagymnus LeConte 1873: 203 Cuba localities. SS: Trinidad Mts., SS: Z. clerinus LeConte 1873: 321; Zayas 1975: 164 Buenos Aires (t.l.) Distribution. Cuba, USA (GA-FL) Collections. Washington (type) Cuba localities. PR: Dimas Collections. Zayas A Checklist of the Beetles of Cuba 175

TRIBE LAMIINI TRIBE AGAPANTHINI

Monochamus Megerle 1821: 106 Spalacopsis Newman 1842: 303 M. titillator (Fabricius) 1775: 172 (Lamia); Zayas S. filum (Klug) 1829: 13 (Eutheia); Jacquelin du 1975: 177. The southern pine sawyer beetle Val 1857: 277 (Euthuorus); Chevrolat 1862: 256; Distribution. Cuba (introduced), e USA Gundlach 1891: 218; Zayas 1975: 214; Chemsak Cuba localities. Ha: Santiago de las Vegas; 1969: 189 Vibora. Possibly introduced into Cuba in lum- Distribution. Cuba, Bahamas, Jamaica, ber Puerto Rico; USA (FL); hosts (see Mészáros Collections. Gundlach no. 1206. Zayas 1977) Cuba localities. Widespread Neoptychodes Dillon and Dillon 1941: 44 Collections. Brno. Gundlach no. 476. Zayas N. trilineatus (Linnaeus) 1771: 532 (Cerambyse); S. grandis (Chevrolat) 1862: 255 (Euthuorus); Chevrolat 1862: 255; Gundlach 1891: 217 Gundlach 1891: 217; Zayas 1975: 216; Chemsak (Ptychodes); Zayas 1975: 175 1969: 189 Distribution. Cuba (introduced), West Indies Distribution. Cuba. Endemic to South America, USA (FL, sw states) Cuba localities. Ha: Havana (t.l.); Santa Fé Cuba localities. Widespread Collections. Brno Collections. Gundlach no. 470. Zayas S. spinipennis Fisher 1936: 347: Zayas 1975: 216 Distribution. Cuba. Endemic Taeniotes Serville 1835: 90 Cuba localities. SC: Pico Turquino (t.l.) [T. insularis Thomson 1857: 171; Zayas 1975: 174] Collections. Washington (type) Distribution. Guadeloupe; not Cuba, an er- ror TRIBE ONCIDERINI

TRIBE APOMECYNINI Paraclytemnestra Breuning 1974: 239 (P. gigantea Breuning 1974: 240) Acrepidopterum Fisher 1926: 6 Distribution. West Indies (where?) A. acutum Zayas 1975: 187 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. SC: Gran Piedra (t.l.) Collections. Zayas (type) TRIBE PTEROPLIINI A. minutum Fisher 1926: 6; Zayas 1975: 188 Distribution. Cuba, Jamaica Ataxia Haldeman 1847: 54 Cuba localities. Ci: Cayamas (t.l.); A. alboscutellata Fisher 1926: 3; Zayas 1975: 182 Cumanayagua. Gu: Florida Blanca. Ha: Lomas Distribution. Cuba, Puerto Rico, Virgin Is- de Camoa lands Collections. Washington (type). Zayas Cuba localities. Ci: Cayamas (t.l.) Collections. Washington (type) Bebilis Thomson 1864: 110 A. spinicauda Schaeffer 1904: 224; Zayas 1975: 184 B. schwarzi (Fisher) () 1947: 37; Zayas A. spinipennis, Gundlach 1891: 213 1975: 219 Distribution. Cuba, Bahamas, Jamaica, USA Distribution. Cuba. Endemic (FL) Cuba localities. CA: Baraguá. Ci: Cayamas Cuba localities. Widespread (t.l.); Cumanayaga. PR: Viñales Collections. Zayas. Collections. Washington (type) A. spinipennis (Chevrolat) 1862: 252; Zayas 1975: 181 (Proecha) Thomson 1868: 157 Distribution. Cuba, Puerto Rico P. insularis Fisher 1930: 9; Zayas 1975: 179; Cuba localities. Widespread Chemsak 1969: 187 Collections. Gundlach no. 820. Zayas Distribution. Cuba. Endemic A. stehliki Chemsak 1969: 187 Cuba localities. Ha: Havana. SC: Santiago. Distribution. Cuba. Endemic Attacks cow peas (Vignia sinensis) Cuba localities. Ha: Cojimar (t.l.). PR: Los Collections. Brno Jazmines 176 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Berkeley (paratypes). Brno (type) Collections. Gundlach no. 477 (D. lanata). A. variegata Fisher 1925: 1; Zayas 1975: 185 Zayas Distribution. Cuba. Endemic D. longipilis (Fisher) 1926: 7 (Ischnolea); Zayas Cuba localities. Gu: Guantánamo (t.l.); Alto 1975: 192 Songo. SC: Gran Piedra; Pico Turquino Distribution. Cuba. Endemic Collections. New York (type). Zayas Cuba localities. Ci: Cayamas (t.l.); Cumanayagua. SC: Cuabitas. On Guazuma TRIBE POGONOCHERINI tomentosa Collections. Washington (type). Zayas Ecyrus LeConte 1852: 160 E. hirtipes Gahan 1895: 127 Estola Fairmaire and Germain 1859: 24 E. insularis Fisher 1932: 76 (CA: Baraguá, t.l.; E. dilloni Zayas 1975: 205 Washington, type); Zayas 1975: 207; Chemsak Distribution. Cuba. Endemic 1969: 189 Cuba localities. Gu: Imias. M: Peninsula de Distribution. Cuba, Bahamas, Hispaniola, Zapata (t.l.). SC: Ciudamar; Vista Alegre Puerto Rico, Virgin Islands, Guadeloupe, Collections. Zayas (type) Dominica, Martinique, Grenada Cuba localities. Widespread Eupogonius LeConte 1852: 159 Collections. Brno. Washington. Zayas E. annulicornis Fisher 1926: 9; Zayas 1975: 198 Distribution. Cuba, USA (FL) Lypsimena Haldeman 1947: 54 Cuba localities. Ci: Cayamas (t.l.) L. fuscata Haldeman 1847: 54; Chevrolat 1862: 263; Collections. Washington (type) Gundlach 1891: 214; Zayas 1975: 209; Chemsak E. bierigi Melzer 1933: 380; Zayas 1975: 199 1969: 189 Distribution. Cuba. Endemic Distribution. Cuba, México to Panamá, USA Cuba localities. PR: Sierra Rangel; Viñales (FL, CA) (t.l.) Cuba localities. CA: Baraguá. Gr: Bayamo. Collections. Type in Melzer collection Ha: Cojimar. Ho: Sierra Cristal. PR: Cananas. E. griseus Fisher 1926: 11; Zayas 1975: 200 SC: Central Mercedes Distribution. Cuba. Endemic Collections. Brno. Gundlach no. 1456. Zayas Cuba localities. Ci: Cayamas (t.l.). Gu: Glorieta; Imias. M: Peninsula Zapata. SC: Zaplous LeConte 1878: 415 Siboney Z. annulatus (Chevrolat) 1862: 250 (Ecyron); Collections. Washington (type). Zayas Gundlach 1891: 210 (Ecyrus); Zayas 1975: 211 E. lanuginosus (Chevrolat) 1862: 255 (Phidola); Distribution. Cuba, USA (FL) Gundlach 1891: 216; Zayas 1975: 193 Cuba localities. Ci: San Blas. M: Peninsula Distribution. Cuba. Endemic de Zapata. PR: Viñales Cuba localities. None reported Collections. Gundlach no. 1293. Zayas Collections. Z. baracutey Zayas 1975: 212 E. maculicornis (Chevrolat) 1862: 254 (Phidola); Distribution. Cuba. Endemic Gundlach 1891: 216; Zayas 1975: 194; Chemsak Cuba localities. M: Cienaga de Zapata (t.l.). 1969: 188 (Phidola) PR: Viñales Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Widespread Collections. Brno. Gundlach no. 1292. Zayas TRIBE DESMIPHORINI E. nigritarsus Fisher 1926: 10; Zayas 1975: 201 Distribution. Cuba. Endemic Desmiphora Serville 1835: 62 Cuba localities. Ci: Cayamas (t.l.). SC: Dai- D. hirticollis (Olivier) 1795: 11; Zayas 1975: 190 quiri D. lanata Chevrolat 1862: 263; Gundlach 1891: Collections. Washington (type). Zayas 214 E. piceus (Fisher) (Phidola) 1936: 348; Zayas 1975: Distribution. Cuba, West Indies (where?), 195 South America, sw USA Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. SC: Loma del Gato (t.l.) Collections. Havana. Washington (type) A Checklist of the Beetles of Cuba 177

E. pilosulus (Chevrolat) 1862: 254 (Phidola); 1891: 210; Zayas 1975: 271 Gundlach 1891: 215; Zayas 1975: 202; Chemsak Distribution. Cuba, Puerto Rico, USA (FL) 1969: 189 Cuba localities. Widespread E. rubiginosus (Chevrolat) 1862: 254 Collections. Gundlach no. 7. Zayas E. exiguus (Thomson) 1868: 115 Distribution. Cuba. Endemic Eneodes Fisher 1926: 31. Endemic genus Cuba localities. Widespread E. hirsuta Fisher 1926: 32; Zayas 1975: 276 Collections. Brno. Gundlach no. 480 Distribution. Cuba. Endemic E. superbus (Zayas) 1975: 196 (Phidola) Cuba localities. Ci: Cayamas (t.l.). Gu: Distribution. Cuba. Endemic Yateras Cuba localities. Ho: Cupeyal (t.l.) Collections. Washington (type). Zayas Collections. Zayas (type) E. viridulus Fisher 1942: 37; Zayas 1975: 277 Distribution. Cuba. Endemic TRIBE Cuba localities. Ci: Soledad (t.l.) Collections. Gundlach no. 1887. Harvard Serville 1835: 29 (type) A. circumflexa Jacquelin du Val 1857: 270; Chevrolat 1862: 247; Gundlach 1891: 205 Eugamandus Fisher 1926: 33 (Psopharochus); Zayas 1975: 225; Chemsak E. cayamae Fisher 1926: 35; Zayas 1975: 278 1969: 189 Distribution. Cuba. Endemic Distribution. Cuba, West Indies, México to Cuba localities. Ci: Cayamas (t.l.). Gu: South America; hosts (see Mészáros 1977) Baracoa. PR: Pan de Guajaibón. SC Gran Pie- Cuba localities. Widespread dra; Loma del Gato Collections. Brno. Gundlach no. 364. Zayas Collections. Washington (type). Zayas E. schwarzi Fisher 1926: 34; Zayas 1975: 279 Oreodera Serville 1835: 19 Distribution. Cuba. Endemic O. lateralis Olivier 1895: 179: Zayas 1975: 221; Cuba localities. Ci: Cayamas (t.l.) Chemsak 1969: 189 (O. glauca Linn.) Collections. Washington (type) Distribution. Cuba, Puerto Rico, Panamá, E. tuberculatus Fisher 1942: 40; Zayas 1975: 281 South America Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. SC: Pico Turquino (t.l.) Collections. Brno. Zayas Collections. Harvard (type)

Steirastoma Lacordaire 1830: 180 Fisherostylus Gilmour 1963: 60. Endemic genus S. poeyi Chevrolat 1862: 246; Gundlach 1891: 205; F. bruneri (Fisher) 1926: 23 (Leptostylus); Gilmour Zayas 1975: 224 1963: 61; Zayas 1975: 237 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ha: Santiago de las Vegas. Cuba localities. Gr: Nagua (t.l.). Gu: Imias; M: Fermina. PR: San Diego Tortuguilla. Ho: Sierra Cristal. SC: Gran Pie- Collections. Gundlach no. 1702. Zayas dra Collections. Washington (type). Zayas TRIBE Lagocheirus Dejean 1835: 336 Acanthocinus Mergele in Dejean 1821: 106 L. araneiformis (Linnaeus) 1767: 625; Jacquelin A. obsoletus (Olivier) 1895: 310 (Graphisurus); du Val 1857: 271 (); Chevrolat Zayas 1975: 275 1862: 247; Gundlach 1891: 206; Zayas 1975: Distribution. Cuba, e USA (FL) 227 Cuba localities. Ho: Mayarí; Sierra Cristal; L. a. stroheckeri Dillon 1956: 138, limited to s Sierra de Mícara. PR: Sierra de Cajálban Florida and Cuba Collections. Zayas Distribution. Cuba, Jamaica, Hispaniola, Puerto Rico, Lesser Antilles, México to Alcidion Sturm 1843: 254 Panamá, USA (FL); hosts (see Mészáros 1977) A. umbraticum (Jacquelin du Val) 1857: 272 Cuba localities. Widespread (Probatius); Chevrolat 1862: 249; Gundlach Collections. Gundlach no. 266. Zayas 178 Arthropods of Florida and Neighboring Land Areas, Vol. 18

L. dezayasi Dillon 1957: 144; Chemsak 1969: 189 L. castaneovirescens Zayas 1975: 239 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ha: Havana-Alamár Cuba localities. Ho: Sierra Cristal (t.l.) Collections. Brno Collections. Zayas (type) L. dealbatus (Jacquelin du Val) 1857: 274 Leptostylopsis Dillon 1956: 144 (Amniscus); Chevrolat 1862: 248; Gundlach L. albofasciatus (Fisher) 1926: 16 (Leptostylus); 1891: 208: Zayas 1975; 240 Zayas 1975: 232 Distribution. Cuba. Endemic Distribution. Cuba, USA (FL) Cuba localities. Widespread Cuba localities. Widespread. Ci: Cayamas Collections. Gundlach no. 127. Zayas (t.l.) L. heticus Zayas 1975: 243 Collections. Washington (type). Zayas Distribution. Cuba. Endemic L. argentatus (Jacquelin du Val) 1857: 273 Cuba localities. Ca: Camaguëy (t.l.); (Amniscus); Dillon 1956: 145; Zayas 1975: 233 Nuevitas, California (Leptostylus) Collections. Zayas (type) Distribution. Cuba, Bahamas, USA (FL); L. illitus Zayas 1975: 244 hosts (see Mészáros 1977) Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Ho: Sierra Cristal (t.l.) Collections. Gundlach no. 128. Zayas Collections. Zayas (type) L. atromaculatus (Fisher) 1926: 18 (Leptostylus); L. latifasciatus Zayas 1975: 246 Zayas 1975: 234 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Soroa (t.l.) Cuba localities. Ha: Central Constancia (t.l.) Collections. Zayas (type) Collections. Washington (type). Zayas L. nigritus Zayas 1975: 248 L. duvali (Fisher) 1926: 19 (Leptostylus); Zayas Distribution. Cuba. Endemic 1975: 242 Cuba localities. Ho: Moa; Piloto (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. L. pseudocalcarius Zayas 1975: 249 Collections. Washington (type) Distribution. Cuba. Endemic L. incrassatus (Klug) 1829: 13 (Leptostylus); Zayas Cuba localities. Ho: Sierra Cristal (t.l.) 1975: 245; Chemsak 1969: 189 Collections. Zayas (type) Amniscus transversatus Jacquelin du Val 1857: L. sagittatus (Jacquelin du Val) 1857: 272 273; Chevrolat 1862: 248; Gundlach 1891: 207 (Amniscus); Chevrolat 1862: 248; Gundlach Distribution. Cuba. Endemic 1891: 402; Zayas 1975: 251 Cuba localities. Widespread. For hosts see Distribution. Cuba. Endemic Mészáros (1977) Cuba localities. Widespread Collections. Brno. Gundlach no. 256. Zayas Collections. Gundlach no. 128. Zayas L. monticola (Fisher) 1935: 201 (Leptostylus); Zayas 1975: 247 Lepturgotrichona Gilmour 1957: 27 Distribution. Cuba. Endemic L. cubaecola (Fisher) 1942: 36 (Lepturges); Zayas Cuba localities. M: Pan de Matanzas (t.l.) 1975: 268; Gilmour 1957: 27 Collections. Washington (type) Distribution. Cuba. Endemic Cuba localities. Ci: Cumanayagua; Soledad Leptostylus LeConte 1852: 168 Collections. Zayas L. albagniri Zayas 1975: 231 Distribution. Cuba. Endemic Lethes Zayas 1975: 254. Endemic genus Cuba localities. Ho: Sierra Cristal (t.l.) L. humeralis Zayas 1975: 255 Collections. Zayas (type) Distribution. Cuba. Endemic L. calcarius Chevrolat 1862: 248; Gundlach 1891: Cuba localities. Ho: Sierra Cristal (t.l.) 207; Zayas 1975: 238 Collections. Zayas (type) Distribution. Cuba. Endemic L. indignus Zayas 1975: 256 Cuba localities. Ca: Santa Lucía. Ho: Sierra Distribution. Cuba. Endemic Cristal. M: Cárdenas Cuba localities. Ca: Santa Lucía (t.l.) Collections. Gundlach no. 129. Zayas Collections. Zayas (type) A Checklist of the Beetles of Cuba 179

Liopinus Linsley and Chemsak 1995: 88 Cuba localities. Widespread. Ci: Cayamas. L. pantherinus (Zayas) 1975: 264 (Sternidius) (t.l) Distribution. Cuba. Endemic Collections. Washington (type). Zayas Cuba localities. Ho: Sierra Cristal, Las Mulas S. nigronotatus (Zayas) 1975: 261 (Urgleptes) (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Ca: Santa Lucía, n coast (t.l.) Collections. Zayas (type) Oedopeza Serville 1835: 88 S. scurra (Chevrolat) 1862: 249 (Alcidion); O. pogonocheiroides Serville 1835: 88; Zayas 1975: Gundlach 1891: 209; Fisher 1926: 21 273 (Leptostylus); Zayas 1975: 252; Chemsak 1969: Distribution. Cuba, México to Uruguay 190 Cuba localities. Ha: Havana. PR: Sierra S. s. maculifer (Fisher) 1926: 2; (Ha: Santiago de Rangel; Guanahacabibes. Hosts; see Mészáros la Vega, t.l.; Washington, type) 1977 Distribution. Cuba. Endemic Collections. Zayas Cuba localities. SC: Pico Cuba Collections. Brno Pygmaleptostylus Gilmour 1963: 74. Endemic S. variabilis (Fisher) 1925: 11 (Urgleptes); Zayas genus 1975: 262 P. pygmaeus (Fisher) 1926: 24 (Leptostylus); Distribution. Cuba. Endemic Gilmour 1963: 75; Zayas 1975: 250 Cuba localities. Widespread. Ci: Cayamas Distribution. Cuba. Endemic (t.l.). A borer of cassava plants Cuba localities. Ci: Cayamas (t.l.) Collections. New York (paratypes). Washing- Collections. Washington (type). Zayas. ton (type). Zayas S. zorillai Zayas 1975: 259 (Urgleptes) Styloleptus Dillon 1956: 158 Distribution. Cuba. Endemic S. atrovittatus (Fisher) 1925: 9 (Urgleptes); Zayas Cuba localities. PR; Ancón; Cajálbana (t.l.); 1975: 258 Piedra de Molino Distribution. Cuba, Hispaniola Collections. Zayas (type) Cuba localities. Gr: Yara (t.l.). Gu: Florida Blanca. M: Peninsula de Zapata, San Blas. SC: Trypanidius Blanchard 1843: 209 Alto Songo T. insularis Fisher 1925: 8; Zayas 1975: 266 Collections. Havana (paratype). New York Distribution. Cuba. Endemic (paratype). Washington (type) Cuba localities. Gr: Nauas (t.l.). Gu: S. biustus (LeConte) 1852: 169 (Leptostylus); Guantánamo Chevrolat 1862: 249; Gundlach 1891: 209 Collections. New York (type) (Exocentrus); Zayas 1975: 235; Chemsak 1969: T. nocturnus Fisher 1942: 39; Zayas 1975: 267 189 Distribution. Cuba, Puerto Rico Leptostyulus minuens Hamilton 1896: 121 Cuba localities. SC: Loma del Gato; Sierra L. scurra dorsalis Fisher 1926: 21. Cuba Maestra, Loma de Calá. SS: Buenos Aires;Topes L. fisheri Gilmour 1965: 577. Cuba de Collantes Leptostylus biustus cubanus Fisher 1926: 22 (Ci: Collections. Zayas Cayamas, t.l.; Washington, type) Lethes israeli Zayas 1975: 25; Ho: Sierra Cristal Urgleptes Dillon 1956: 332; key in Gilmour 1963: (t.l.); Zayas (type); Micheli & Hovore 2003: 2 78 (synonomy) U. chamaeropsis ( Fisher) 1926: 27 (Lepturges); Distribution. Cuba Bahamas, se USA (FL) Zayas 1975: 269 Cuba localities. Ca: Playa Santa Lucía. Gu: Distribution. Cuba. Endemic Imias. Ha: Cojimar. Ho: Moa; Piloto; Sierra Cuba localities. Ci: Cayamas (t.l.). M: Pen- Cristal insula de Zapata Collections. Brno. Zayas Collections. Washington (type). Zayas S. cubanus (Fisher) 1926: 26 (Leiopus); Zayas 1975: U. guadeloupensis (Fleutiaux and Sallé) 1889: 472 260 (Urgleptes) (Lepturges); Zayas 1975: 270; Chemsak 1969: Distribution. Cuba. Endemic 90 Distribution. Cuba to Guadeloupe 180 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Brno. Zayas Cuba localities. Gr: Bayamo. M: Cárdenas. PR: Cordillera de los Organos; Rancho Mundito; TRIBE CYRTININI Aspiro. SC: Palma Mocha; Pico Turquino Collections. Brno. Gundlach no. 109b. Zayas Cyrtinus LeConte 1852: 166 C. venustus (Chevrolat) 1838: 283 (Amphionycha); C. acunai (Fisher) 1935: 206; Zayas 1975: 283 Zayas 1975: 295 (C. venustos) Distribution. Cuba. Endemic C. nuptus Chevrolat 1862: 251; Gundlach 1891: Cuba localities. SS: Trinidad Mts.; Buenos 211; Zayas 1975: 293 Aires (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. M: Cardenas (on Solanum C. humilis Zayas 1975: 284 toroum) Distribution. Cuba. Endemic Collections. Cambridge. Gundlach no. 1726 Cuba localities. SC: Loma del Gato (t.l.) (C. nuptus); Gundlach no. 106 (C. venustos) Collections. Zayas (type) Notes. This species seems to be part of a Batesian C. schwarzi Fisher 1926: 36 mimicry complex centered around species of Distribution. Cuba. Endemic Thonalmus (Lycidae) (Darlington 1938: 690) Cuba localities. Ci: Cayamas (t.l.). PR: Si- erra Rangel Oedudes Thomson 1868:199 Collections. Washington (type). Zayas O. alayoi (Zayas) 1956: 111 (Essostrutha); Zayas 1975: 300 (Adesmus); Chemsak 1978: 125; TRIBE Galileo and Martins 1997: 143 Distribution. Cuba. Endemic Calocosmus Chevrolat 1862: 250 Cuba localities. SS: Buenos Aires , Lomas C. chevrolati Fisher 1925: 14; Zayas 1975: 289 de Trinidad (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Gu: Guantánamo (t.l.). Ho: O. ramsdeni (Fisher) 1926: 39 (Essostrutha); Zayas Moa, Piloto; Nicaro. SC: Loma del Gato; Siboney 1956: 106; Zayas 1975: 302 (Adesmus); Collections. Harvard (paratype). New York Chemsak 1978: 125; Galileo and Martins 1997: (type). Zayas 143 C. dimidiatus (Chevrolat) 1838: 284 Distribution. Cuba. Endemic (Amphionycha); Jacquelin du Val 1857: 275; Cuba localities. Gu: Guantánamo (t.l.). Ho: Gundlach 1891: 212; Zayas 1975: 289 Moa Distribution. Cuba. Endemic Collections. New York (type) Cuba localities. Widespread O. roberto (Fisher) 1935: 209 (Essostrutha); Zayas Collections. Cambridge. Gundlach no. 364. 1956: 107; 1975: 303.(Adesmus); Chemsak Zayas 1978: 125; Galileo and Martins 1997: 143 Notes. This species seems to be part of a Batesian Distribution. Cuba. Endemic mimicry complex centered around species of Cuba localities. PR: Los Animas; Sierra de Thonalmus (Lycidae) (Darlington 1938: 690) Rangel (t.l.); Pan de Guajaibón C. nigripennis Chevrolat 1862: 252; Gundlach 1891: Collections. Washington (type). Zayas 213; Zayas 1975: 292 O. scaramuzzai (Fisher) (Essostrutha) 1936: 271; C. semimarginatus Bates 1881: 151; Zayas 1975: Zayas 1956: 109 (translation) (Adesmus); 294 Chemsak 1978: 125; Galileo and Martins 1997: C. janus Bates 1881: 151; Zayas 1975: 292 143 C. fulvicollis Fisher 1925: 13; Zayas 1975: 290 Adesmus montivagans (Fisher) 1942: 42 Distribution. Cuba. Endemic (Essostrutha); Zayas 1956: 110; 1975: 300; Cuba localities. Gu: Guantánamo. M: Chemsak 1978: 125 Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 797. New York Cuba localities. SC: Loma del Gato (t.l.); .Pico C. speciosus Chevrolat 1862: 252; Gundlach 1891: Turquino (t.l.) 212; Zayas 1975: 294; Chemsak 1969: 190 Collections. Harvard (type ). Washington. Hemilopus venustus Jacquelin du Val 1857: 275, Zayas not Chevrolat A Checklist of the Beetles of Cuba 181

REFERENCES islands no. 76. Natuurwet. Stud. , Breuning, S. von. 1974. Neue Lamiinae (Co- the Hague, 17: no. 33: 57-96. leoptera, Cerambycidae). Mitt. Zool. Mus. Ber- Mészáros Z. 1977. Datos para el conocimiento de lin 50: 237-241. las naturales plantas nutritivas de los Browne, J. and S.B. Peck. 1996. The long-horned Longicornis en Cuba (Coleoptera: beetles of south Florida (Cerambycidae: Co- Cerambycidae). Folia Ent. Hung. 30 (2): 75-76. leoptera); biogeography and relationships with Micheli, J. and F. T. Hovore. 2003. New syn- the Bahama Islands and Cuba. Canadian J. onymies and distributional records for Carib- Zool. 74: 2154-2169. bean Cerambycidae (Coleoptera). Coleop. Bull. Chemsak, J.A. 1969. Records and descriptions of 57: 1-4. Cuban Cerambycidae. Acta Musei Moraviae 54: Monné, M.A. and E.F. Giesbert. 1995. Check- 185-190. list of the Cerambycidae and (Co- Chemsak, J.A. 1978. Review of the genus leoptera) of the Western Hemisphere. Essostrutha Thomson (Coleoptera, Wolfsgarden Books, Burbank, CA. Cerambycidae). Pan-Pacific Ent. 54: 125-128. Philips, T.K. and Ivie, M.A. 1998. The Methiini Chemsak, J.A., E.G. Linsley, and F.A. of the West Indies (Coleoptera: Cerambycidae) Noguera. 1992. Listados faunisticos de México with notes on the circum-Caribbean species. II. Los Cerambyiidae y Disteniidae de Entomol Scand. 29: 57-87. Norteamérica, Centroamérica y las Indies Vàzquez, L.L., E. Valdés, and J.C. Amor. 1992. Occidentales (Coleoptera). Univ. Nac. Autó New manifestations of pests on economically México. Inst. Biol. 204 pp. important plants during the period from 1970 Darlington, P.J., Jr. 1938. Experiments on mim- to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo icry in Cuba, with suggestions for future study. Silvestri 49: 41-52. Trans. R. Ent. Soc. London 87: 681-695. Zayas, F. de. 1957. Revision de los Longicornios Dillon L. 1956. The Nearctic components of the Prionidos de Cuba (Coleopteros, Cerambycidae). tribe Acanthocinini (Coleoptera, Mem. Soc. Cubana Hist. Nat., Havana 23: 149- Cerambycidae). Ann. Ent. Soc. America 49: 181. 134-167; 207-235; 332-355. Zayas, F. de. 1975. Revision de la Familia Dillon, L. 1957. Revision of the neotropical Cerambycidae (Coleoptera, Phytophagoidea). Acanthocinini (Coleoptera, Cerambycidae). The Acad. Cien. Cuba, Instit. Zool. 443 pp. genus Lagocheirus. Bull. British Mus. Nat. Hist., London (Ent.) 6: 137-166. 156. FAMILY BRUCHIDAE Fernandez García, I. and N. Fiol Ferrer. 1999. The pea and bean weevils. Plantas hospedantes de las especies de All members of the family, as larvae, are seed cerambícidos (Coleoptera: Cerambycidae) de feeders, and most are associated with legumes. Cuba. Cocuyo 8: 14-16. Some Pachymerinae feed on palm nuts. Johnson Galileo, M.H.M. and U.R. Martins. 1997. Sobre and Kingsolver (1981) give a checklist of West In- o gênero Oedudes Thomson, 1868 (Coleoptera, dian bruchids; and Alvarez and Kingsolver (1997) Cerambycidae, Lamiinae, Hemilophni). Papéis give a checklist and additional literature for Cuba. Avulsos de Zoologia (São Paulo) 40: 137-146. Lawrence and Newton (1995) consider this group Galileo, M.H.M. and U.R. Martins. 1993. to be a subfamily of the Chrysomelidae. It is here Revisão da tribo Solenopterini (Coleoptera, used as a family, following Arnett and Thomas Cerambycidae, Prioninae). Parte III. Gênero (2002). Solenoptera A.-Serville, 1832. Revista Bras. Ent. 37: 425-458. SUBFAMILY PACHYMERINAE Gilmour, E.F. 1957. On the Neotropical Acanthoninini (Coleoptera, Cerambycidae, Caryobruchus Bridwell 1929: 148 Laminae). Some new genera and species. Arq. C. gleditsiae (Johansson and Linnaeus) 1767: 605; Biol. Technol. (Curitiba) 10(1): 3-36. (Dermestes). The palm seed weevil Gilmour, E. F. 1963. On the Neotropical Bruchus arthriticus Fabricius 1801: 398; Acanthocinini (Col. Cerambycidae, Lamiinae). Jacquelin du Val 1857: 166; Gundlach 1891: 265 Some Caribbean genera and species. Studies Distribution. Cuba, Bahamas, Bermuda, on the fauna of Curaçao and other Caribbean Hispaniola, Jamaica, México to Panamá, USA 182 Arthropods of Florida and Neighboring Land Areas, Vol. 18

(FL-NC-TX); a pest in seeds of many genera of St. Croix, Curaçao, USA (FL) palms Cuba localities. Ca: Jobabo. CA: Baragua. Cuba localities. IJ: Nueva Gerona. M: Ci: Cayamas (t.l.); Soledad. Gu: Baracoa; Moa. Cárdenas Ha: Havana. SC: coast below Pico Turquino. Collections. Gundlach no. 210. Pittsburgh Hosts: Hippomane mancinella, Tectona C. marieae Nilsson and Johnson 1990: 55 grandis, Ricinis communis, Randia aculeata, Distribution. Cuba. Endemic Terminalia catappa (Genaro and Kingsolver Cuba localities. Ci: Cayamas. Gu: Imias. Ha: 1997) Havana, Camacho (t.l.). In seeds of Sabal pal- Collections. Cambridge. Havana. Gainesville. metto and Chamaeodora elegans Washington (type) Collections. Washington (type) A. simulator (Jacquelin du Val) 1857: 167 (Spermophagus); Gundlach 1891: 266; SUBFAMILY AMBLYCERINAE Kingsolver 1970: 476 TRIBE AMBLYCERINI Distribution. Cuba, Costa Rica Cuba localities. CA: Baragua. Ci: Cayamas; Amblycerus Thunberg 1815: 109 Soledad. Ha: Havana. M: Cárdenas. Host: A. baracoensis Kingsolver 1970: 484 Luehea speciosa Distribution. Cuba, México, Costa Rica, Collections. Cambridge. Gundlach no. 211. Paraguay Havana. Washington Cuba localities. CA: Baragua. Ci: Cayamas. A. taeniatus (Suffrian) 1870: 167 (Spermophagus); Gu: Baracoa (t.l.). Host: Cordia gerascanthus Gundlach 1891: 266; Kingsolver 1970: 475 Collections. Havana. New York. Washing- Distribution. Cuba. Endemic ton (type) Cuba localities. Gr: Cabo Cruz. Host: A. cistelinus (Gyllenhal) 1833: 103 (Sperm- Cesalpinia bijug ophagus); Jacquelin du Val 1857: 167; Collections. Gundlach no. 1366 Gundlach 1891: 265; Kingsolver 1970: 473 Distribution. Cuba, México to Brazil TRIBE SPERMOPHAGINI Cuba localities. Ci: Cayamas; Soledad. Gu: Yateritas. M: Cárdenas. SC: Alto Songo, Florida Zabrotes Horn 1885: 156 Blanca. Host: Guazuma ulmifolia Z. subfasciatus (Boheman) 1833: 111 (Sperm- Collections. Cambridge. Champaign. ophagus); Gundlach 1891: 267. The Mexican Gundlach no. 362. Washington bean weevil A. eustrophoides (Schaeffer) 1904: 228 Distribution. Cuba (introduced), South (Spermophagus); Kingsolver 1970: 474 America, Panamá to USA (TX); nearly Distribution. Cuba, México, Costa Rica, USA circumtropical (FL) Cuba localities. Hosts: many leguminous Cuba localities. SC: Alto Songo, Florida genera Blanca. Host: Drypetes laterifolia Collections. Gundlach no. 731 Collections. Champaign A. pygidialis (Suffrian) 1870: 169 (Spermophagus); SUBFAMILY BRUCHINAE Gundlach 1891: 267; Kingsolver 1970: 481 TRIBE BRUCHIDIINI A. chapini Kingsolver 1970: 481 Distribution. Cuba, Jamaica Callosobruchus Pic 1902: 6 Cuba localities. Ci: Cayamas. Gu: Baracoa. C. chinensis (Linnaeus) 1758: 386 (Bruchus) Host: Cordia gerascanthus Bruchus pectinicornis Linnaeus 1767: 605; Collections. Gundlach no. 994. Washington Gundlach 1891: 260 A. sallei (Jekel) 1855: 30 (Spermophagus) Distribution. Cuba, Puerto Rico, Panamá, Distribution. Cuba, Hispaniola, Jamaica, South America, Old World, USA (DC, VA); cos- Puerto Rico, Venezuela mopolitan; a major pest of stored products Cuba localities. Host: Prosopis juliflora Cuba localities. Widespread. Hosts: many Collections. leguminous genera A. schwarzi Kingsolver 1970: 477 Collections. Gundlach no. 604 Distribution. Cuba, Hispaniola, Cayman Is- C. maculatus (Fabricius) 1775: 66 (Bruchus). The lands, Jamaica, Puerto Rico, Virgin Islands, cow pea weevil A Checklist of the Beetles of Cuba 183

Bruchus quadrimaculatus Fabricius 1792: 371: Cuba localities. Ca: Jaronú. Gr: Bayamo. Ha: Gundlach 1891: 260 Havana; Santiago de las Vegas. M: Cárdenas. Bruchus sinuatus Fahraeus 1839: 8; Gundlach SC: Santiago de Cuba. Host: Ipomoea alba 1891: 26l Collections. Gundlach no. 415. Washington Distribution. Cuba, Puerto Rico, México, South America, USA (TX-CT-FL); cosmopoli- TRIBE ACANTHOSCELIDINI tan pest of seeds of various legumes Cuba localities. Ci: Cienfuegos Acanthoscelides Schilsky 1905: 95 Collections. Gundlach no. 812 A. flavescens (Fahraeus) 1839: 32 (Bruchus) Distribution. Cuba, Hispaniola, Puerto Rico, TRIBE BRUCHINI St. Vincent, México to Panamá, USA Cuba localities. Hosts: Eriosema, Galactia, Bruchus Linnaeus 1767: 604 Rhynchosia, Tephrosia B. pisorum (Linnaeus) 1758: 356. The pea weevil Collections. B. pisi Linnaeus 1767: 604; Jacquelin du Val A. livens (Suffrian) 1870: 154 (Bruchus); Gundlach 1857: 164; Gundlach 1891: 263 1891: 262 Distribution. Cuba (introduced), Puerto Rico, Distribution. Cuba. Endemic South America, USA (widespread), Old World Cuba localities. SS: Trinidad Mts. Cuba localities. Hosts: usually Pisum, but Collections. Gundlach no. 1371 (type) also other legumes such as Cassia, Cytisus, A. obtectus Say 1831: 1 (Bruchus). The bean wee- Lathyrus, Phaseolus and Vicia vil Collections. Gundlach no. 1632 Bruchus tetricus Gyllenhal 1839: 22; Gundlach B. rufimanus (Boheman) 1833: 58; Jacquelin du 1891: 262 Val 1857: 165; Gundlach 1891: 264. The Bruchus pallidipes Fahraeus 1839: 91; Jacquelin broadbean weevil du Val 1857: 165 Distribution. Cuba (introduced), Puerto Rico, Distribution. Cuba (introduced); USA (NJ, LA, CA), Old World; cosmopolitan cosompolitan; a widespread pest of stored beans Cuba localities. M: Cárdenas. Hosts: usually and peas Vicia, but also Lupinus, Phaseolus, Pisum, Cuba localities. M: Cárdenas Vigna, etc. Collections. Gundlach no. 317 Collections. Gundlach no. 373 A. pantherinus Suffrian 1870: 163 (Bruchus); Gundlach 1891: 265 TRIBE MEGACERINI Distribution. Cuba. Endemic Cuba localities. SS: Trinidad Megacerus Fahraeus 1839: 34 Collections. Gundlach no. 1372 (type?) M. ferruginosus Teran and Kingsolver 1977: 79 A. quadratus (Suffrian) 1870: 161 (Bruchus); Distribution. Cuba. Endemic Gundlach 1891: 264 Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba. Endemic Collections. Cambridge (type) Cuba localities. Ci: Cienfuegos. SS: M. flabelliger (Fahraeus) 1839: 36 (Bruchus) Buenavista Distribution. Cuba, México to Brazil Collections. Gundlach no. 752 (type?) Cuba localities. Hosts: species of Ipomoea A. quadridentatus (Schaeffer) 1907: 304 (Bruchus) and Merramia Distribution. Cuba, México to Brazil, USA Collections. Cuba localities. Hosts: species of Mimosa M. porosus (Sharp) 1885: 490 (Bruchus) Collections. Distribution. Cuba, Puerto Rico, México, Colombia, Brazil Ctenocolum Kingsolver and Whitehead 1974: Cuba localities. Host: Merramia aegyptia 284 Collections. C. podagricus (Fabricius) 1801: 399 (Bruchus) M. tricolor (Suffrian )1870: 157 (Bruchus, Distribution. Cuba, México, El Salvador, Acanthoscelides); Gundlach 1891: 263; Teran Costa Rica,Venezuela and Kingsolver 1977: 137 Cuba localities. Hosts: Lonchocarpus, Distribution. Cuba, Hispaniola, Nicaragua Piscidia Collections. 184 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Meibomeus Bridwell 1946: 54 Collections. M. relictus (Suffrian) 1870: 156 (Bruchus); S. rufomaculatus (Motschulsky) 1874: 222 Gundlach 1891: 263; Kingsolver and Whitehead (Bruchus) 1976:5 Distribution. Cuba, Hispaniola, Jamaica, Distribution. Cuba. Endemic Puerto Rico, Grenada, México to Colombia Cuba localities. IJ: Nueva Gerona. ? Cuba localities. Host: Cassia Guamacaro sabana. Hosts: unknown for Cuba; Collections. other species attack seeds of , Desmodium, Poiretia and Zornia Stator Bridwell 1946: 55 Collections. Pittsburgh. Gundlach no. 563 S. bottimeri Kingsolver 1972: 225 Distribution. Cuba, Bahamas, USA (FL; Merobruchus Bridwell 1946: 54 Keys only) M. lysilomae Kingsolver 1988: 25 Cuba localities. Ca: Camagüey. Ci: Cayamas; Distribution. Cuba, Bahamas, Hispaniola, Soledad. SC: Santiago de Cuba (t.l.). Hosts: USA (FL) Acacia farnesiana, Acacia pinetorum Cuba localities. Hosts: Acacia, Albizia, Collections. Cambridge. New York. Ottawa. Lysiloma Washington (type) Collections. S. rugulosus Kingsolver 1972: 221 Distribution. Cuba. Endemic Mimosestes Bridwell 1946: 54 Cuba localities. CA: Baraguá (t.l.). Host: M. insularis Kingsolver and Johnson 1978: 35 Pithecellobium discolor Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Washington (type) Curaçao, Colombia, USA (Hi) Cuba localities. SC. Hosts: Acacia farnesiana, REFERENCES Prosopis Alvarez M.D. and J.M. Kingsolver. 1997. A Collections. Gainesville. Gatineau. preliminary list of the Bruchidae (Coleoptera) M. mimosae (Fabricius) 1781: 76 (Bruchus); of Cuba. Ent. News 108: 215-221. Kingsolver and Johnson 1978: 42 Genaro, J. and J.M. Kingsolver. 1997. Distribution. Cuba, Hispaniola, Puerto Rico, Amblycerus schwarzi (Coleoptera: Bruchidae) Curaçao, Trinidad, México to Brazil, USA (FL) attacking seeds of the tropical almond Cuba localities. SC. Hosts: Acacia, Terminalia (Combretaceae) in Cuba. Entomol Caesalpinia, Ceratonia, Parkinsonia News 108: 229-230. Collections. Gatineau. Gainesville Johnson, C.D. and J.M. Kingsolver. 1981. M. nubigens (Motschulsky) 1874: 237 (Bruchus); Checklist of the Bruchidae (Coleoptera) of Kingsolver and Johnson 1978: 49 Canada, United States, México, Central Distribution. Cuba, Puerto Rico, México to America, and the West Indies. Coleop. Bull. 35: Panamá, Brazil, USA (FL, TX, AZ, CA, HI), 409-422. Philippine Islands, New Caledonia Kingsolver, J.M. 1970. A synopsis of the subfam- Cuba localities. SC. Hosts: Acacia, Prosopis ily Amblycerinae Bridwell in the West Indies, Collections. Gatineau. Gainesville with descriptions of new species (Coleoptera: Bruchidae). Trans. Amer. Ent. Soc. 96: 469- Sennius Bridwell 1946: 55 497. S. fallax (Boheman) 1839: 59 (Bruchus) Kingsolver, J.M. 1972. Synopsis of the genus Sta- Bruchus xanthopus Suffrian 1870: 156; Gundlach tor Bridwell in the West Indies, with descrip- 1891: 262 tions of new species. Proc. Ent. Soc. Wash. 74: Distribution. Cuba, Puerto Rico, México, 219-229. Costa Rica, Panamá, USA (FL, GA) Kingsolver, J.M. 1988. Biosystematics of the ge- Cuba localities. M: Cienaga de Zapata. Hosts: nus Merobruchus of continental North America Cassia and the West Indies (Coleoptera: Bruchidae). Collections. Gundlach no. 841 U.S. Dept. Agric. Tech. Bull. 1744: 1-63. S. morosus (Sharp) 1885: 467 (Bruchus) Kingsolver, J.M. and C.D. Johnson. 1978. Sys- Distribution. Cuba, México to Panamá, USA tematics of the genus Mimosestes (Coleoptera: (TX) Bruchidae). US. Dept. Agric. Tech. Bull no. Cuba localities. Hosts: Cassia 1590. 106 pp. A Checklist of the Beetles of Cuba 185

Kingsolver, J.M. and D.R. Whitehead. 1976. Lema Fabricius 1798: 90 (see White 1993) The North and Central American species of L. alpinista Zayas 1988: 117 Meiobomeus (Coleoptera: Bruchidae: Distribution. Cuba. Endemic Bruchinae). US Dept. Agric. Tech. Bull. 1523. Cuba localities. SC: Pico Turquino (t.l.) 54 pp. Collections. Zayas (type) Nilsson, J.A. and C.D. Johnson. 1990. A new L. bifida (Olivier) 1808: 737 (Crioceris); Jacquelin species of palm bruchid from Cuba (West Indies) du Val 1857: 284 and a redescription of Caryobruchus gleditsiae Crioceris punctata Olivier 1791: 199 (L.) (Coleoptera: Bruchidae: Pachymerinae). Distribution. Cuba, Guadeloupe Coleop. Bull. 44: 50-59. Cuba localities. Teran, A.L. and J.M. Kingsolver. 1977. Revi- Collections. sion del genero Megacerus (Coleoptera: L. caeruleipennis Lacordaire 1845: 386; Jacquelin Bruchidae). Opera Lilloana 25: 1-287. du Val 1857: 280; Gundlach 1891: 338; Medvedev 1993: 4 157. FAMILY Distribution. Cuba. Endemic This family is represented in the Neotropics by Cuba localities. Gu: Yateras. M: Matanzas the subfamily Megalopodinae, and the larvae are Collections. Gundlach no. 916. Medvedev stem borers. Many species occur in the Neotropics, L. confusa Chevrolat 1835: 116; Jacquelin du Val but seemingly none occur in the West Indies. 1857: 282; Gundlach 1891: 340; White 1993: 27; Medvedev 1993: 4 158. FAMILY . Distribution. Cuba, Jamaica, Puerto Rico, The family is Holarctic and Neotropical in dis- México to Panamá, Colombia, USA (FL) tribution. The Aulacoscelininae range from the Cuba localities. Ha: Havana. M: Cárdenas southwestern USA to South America. None are Collections. Gundlach no. 297. Medvedev known from the West Indies. L. cubana deBorre 1881: 77 Distribution. Cuba. Endemic 159. FAMILY CHRYSOMELIDAE Cuba localities. The leaf beetles Collections. Reviewed by Shawn M. Clark, March, 1997, and L. internejuncta Pic 1947: 9 Charles Staines, March, 1998. Distribution. Cuba. Endemic Adults and many larvae feed on leaves of an- Cuba localities. giosperms. Some larvae occur in soil and feed on Collections. roots, some are case-bearers in litter, and Hispinae L. lorenzoi Zayas 1988: 116 are leaf miners. The family includes many agricul- Distribution. Cuba. Endemic tural pests. Wilcox (1983) includes the West In- Cuba localities. PP: Pan de Guajaiban (t.l.) dian fauna as a part of a summary of the North Collections. Zayas (type) American fauna. L. lunigera Suffrian 1866: 284; Gundlach 1891: 340; Medvedev 1993: 5 SUBFAMILY DONACIINAE Distribution. Cuba. Endemic TRIBE DONACIINI Cuba localities. Gu: Yateras. SC: Gran Pie- dra Donacia (s.str.) Fabricius 1775: 126 Collections. Gundlach no. 1483. Medvedev D. cinctincornis Newman 1838: 391 L. mannerheimi Lacordaire 1845: 384; Jacquelin D. antillarum Suffrian 1864: 282; Gundlach 1891: du Val 1857: 279 338 Crioceris tuberculata Guérin-Méneville 1844: 264 Distribution. Cuba, México, Honduras, USA Distribution. Cuba, Hispaniola (TX-ND-ME -FL), Canada (MB-NS) Cuba localities. Cuba localities. M: Cienaga de Zapata; lar- Collections. vae are aquatic and feed on submerged vegeta- L. occidentalis Zayas 1988: 118 tion Distribution. Cuba. Endemic Collections. Gundlach no. 1104 Cuba localities. PR: Consolacion del Norte (t.l.) SUBFAMILY CRIOCERINAE Collections. Zayas (type) TRIBE LEMIINI 186 Arthropods of Florida and Neighboring Land Areas, Vol. 18

L. orientalis Zayas 1988: 119. This name may be a Neolema Monros 1951: 472 junior homonym and may need replacement N. dorsalis (Olivier) 1791: 201 (Crioceris); (White 1981: 37) Jacquelin du Val 1857: 286 (Lema); Gundlach Distribution. Cuba. Endemic 1891: 342; Medvedev 1993: 4 (Lema) Cuba localities. SC: Caney. SC: Santa María. Distribution. Cuba, Hispaniola, Puerto Rico, Type locality not given St. Vincent, Grenada, México, Panamá, South Collections. Zayas (type) America L. perizonata Lacordaire 1845: 397 (a replacement Cuba localities. ? Ramón. Gu: Guantanamo. name for the junior secondary homonym below SS: Trinidad (White 1981: 37); Jacquelin du Val 1857: 281; Collections. Gundlach no. 1484. Medvedev Gundlach 1891: 339 N. gundlachiana (Suffrian) 1874: 152 (Lema); L. marginata Guérin-Méneville in Chevrolat White 1993: 66 1838: 285 Lema intermedia Suffrian 1866: 287; Gundlach Distribution. Cuba. Endemic 1891: 342 Cuba localities. M: Cárdenas Distribution. Cuba, Hispaniola, USA (FL) Collections. Gundlach no. 296 Cuba localities. M: Colón L. placida Lacordaire 1845: 397 (404); Jacquelin Collections. Gundlach no. 1534 du Val 1857: 282; Gundlach 1891: 339 Distribution. Cuba. Endemic SUBFAMILY HISPINAE Cuba localities. M: Cárdenas. SS: Trinidad TRIBE CEPHALOLEIINI Collections. Gundlach no. 152 L. poeyi Lacordaire 1845: 510; Jacquelin du Val Cephaloleia Chevrolat 1837: 390 1857: 287; Gundlach 1891: 343; Medvedev C. barroi Uhmann 1959: 617 1993:5 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Lomas de Trinidad (t.l.) Cuba localities. Ha: Rio Cristal. M: Collections. Eberswalde (type?) Cárdenas. SC: Santiago de Cuba Collections. Gundlach no. 532. Medvedev Melanispa Baly 1856: 30 L. postica Guérin-Méneville in Chevrolat 1838: 285; M. bicolor Zayas 1960b: 132 Jacqueline du Val 1857: 285; Gundlach 1891: Distribution. Cuba. Endemic 431 Cuba localities. Ci: Pico Potrerillos (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Gr: Bayamo. M: Cárdenas Collections. Gundlach no. 435 TRIBE L. punctatofasciata Lacordaire 1845: 494; Gundlach 1891: 341 Anisostena Weise 1910: 120 Distribution. Cuba. Endemic A. cyanoptera (Suffrian) 1868: 227 (Odontota); Cuba localities. Gundlach 1891: 389; Sanderson 1967: 138; Collections. Gundlach no. 1147 Staines 1994: 130 L. quadricolor Lacordaire 1845: 402; Jacquelin du Distribution. Cuba, Jamaica, Hispaniola Val 1857: 281;l Medvedev 1993: 4 Cuba localities. Widespread Distribution. Cuba, México Collections. Amsterdam. Cambridge. Cuba localities. Widespread Champaign. Gainesville. Gundlach no. 423. Collections. Medvedev Washington L. trilinea White 1981: 50 (replacement name for C. trilineata (Olivier) (Crioceris); White 1993: Chalepus Thunberg 1805: 282 7 C. bicolor (Olivier) 1792: 96 (Odontota) Crioceris trilineata Olivier 1808: 739; Jacquelin Odontota rufogaster Thunberg 1805: 282 du Val 1857: 283 Odontota rufiventris Suffrian 1868: 229; Distribution. Cuba, México, Costa Rica, USA Gundlach 1891: 390 (widespread) Distribution. Cuba, widespread USA Cuba localities. Cuba localities. C. Staines has seen no Cu- Collections. ban specimens Collections. A Checklist of the Beetles of Cuba 187

C. sanguinicollis (Linnaeus) 1771: 530; Blake Cuba localities. CA: Baraguá (t.l.) 1970a: 126; Sanderson 1967: 137; Medvedev Collections. Washington (type) 1993:9 H. fallax (Suffrian) 1868: 237 (Porphyraspis); C. s. axillaris (Jacquelin du Val) 1856: 313 Gundlach 1891: 392 (Odontota); Gundlach 1891: 389 (O. auxillaris) Distribution. Cuba. Endemic C. s. australis Uhmann 1935: 239 (Cuba, Cuba localities. VC: Las Pozas Hispaniola, Colombia) Collections. Gundlach no. 1270 Distribution. Cuba, Hispaniola, Jamaica, H. flavipes Zayas 1988: 145 Puerto Rico, Virgin Islands, South America, Distribution. Cuba. Endemic USA (FL). Hosts: Sorghastrum setosum and Cuba localities. M: pinelands of Cienaga de Trichachne insularis (Gramineae) Zapata (t.l.) Cuba localities. Ha: Santiago de las Vegas. Collections. Zayas (type) M: Arroyo Bermejo. PR: Sierra Anafe; Rangel H. gundlachi (Boheman) 1862: 43 (Porphyraspis); Collections. Brno. Gundlach no. 490; no. 298 Gundlach 1891: 392; Medvedev 1993: 9 (auxillaris). Medvedev. Washington Distribution. Cuba. Endemic Cuba localities. IJ: in leaves of miraguano Octhispa Chapuis 1877: 23 Collections. Gundlach no. 1232. Medvedev O. testaceipes Pic 1929: 13 H. materna Zayas 1952: 71 Uroplata tricolor Suffrian 1868: 232 (Cuba); Distribution. Cuba. Endemic Gundlach 1891: 390 Cuba localities. M: Bahia de Cochinos, on Distribution. Cuba, Panamá palmettos (t.l.) Cuba localities. M: Cárdenas Collections. Zayas (type) Collections. Gundlach no. 907 H. quadrimaculata Blake 1971: 280 Distribution. Cuba. Endemic Octotoma Dejean 1837: 390 Cuba localities. Ha: Santiago de las Vegas O. gundlachii Suffrian 1868: 234; Gundlach 1891: (t.l.) 391; Staines 1989: 46; Medvedev 1993: 9 Collections. Washington (type) Distribution. Cuba. Endemic H. xanthocera (Boheman) 1850: 98 (Porphyraspis); Cuba localities. Ci: Soledad. Gu: Yateras. Ha: Jacquelin du Val 1857: 314; Gundlach 1891: Havana; Loma de Camoa. Ho: Loma de Moa. 391 (P. scanthocera); Medvedev 1993: 9 M: Cárdenas. In flowers of Lantana camara Distribution. Cuba. Endemic Collections. Cambridge. Gainesville. Cuba localities. IJ: La Fé. on miraguano Gundlach no. 192. Medvedev. Staines. Wash- Collections. Gundlach no. 219. Medvedev ington TRIBE OMOCERINI Pentispa Chapuis 1875: 322 (P. cristata Chapuis 1877: 21) Omocerus Chevrolat 1835: 119 Distribution. Cuba?, Antilles O. taurus (Fabricius) 1787: 380 (Tauroma); Cuba localities. Jacquelin du Val 1857: 315; Gundlach 1891: Collections. 399; Chaboo 2002: 719 Distribution. A South American species (Co- Spaethispa Uhmann 1940: 333 (Uroplata) lombia to Trinidad to Brazil); possibly an erro- S. pulchella (Suffrian) 1868: 230 (Uroplata); neous record or misidentification for Cuba. Ac- Gundlach 1891: 390 cepted as a Cuba record as O. (O.). taurus Distribution. Cuba. Endemic (Boheman) by Chaboo 2002: 719, without addi- Cuba localities. Ca: Manzanillo. Ci: tional data, and with a new record for Jamaica Yaguaramas Collections. Gundlach no. 1482 TRIBE MESOMPHALIINI

TRIBE HEMISPHAEROTINI Chelymorpha Boheman 1854: 1 C. comata Boheman 1854: 4; Medvedev 1993: 10 Hemisphaerota Chevrolat 1837: 397 Distribution. Cuba, Mexico to Honduras H. bimaculata Blake 1971: 279 Cuba localities. Ha: Miranao, coast Distribution. Cuba. Endemic Collections. Medvedev 188 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. fimbrialis Suffrian 1868: 241; Gundlach 1891: Cuba localities. SS: Trinidad, SS: Buenos 393 Aires (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. S. flavicollis (Klug) 1829: 14 (Cassida); Jacquelin Collections. Gundlach no. 1905 du Val 1857: 316 (Chelymorpha); Gundlach C. geniculata Boheman 1854: 39 1891: 392; Medvedev 1993: 9 C. argus Herbst; Jacquelin du Val 1857: 316; Distribution. Cuba. Endemic Gundlach 1891: 393 Cuba localities. Widespread in highlands Distribution. Cuba, Puerto Rico, USA (FL) Collections. Gundlach no. 897. Medvedev Cuba localities. M: Cárdenas S. indivisa Blake 1930: 218 Collections. Gundlach no. 122 (C. argus) Distribution. Cuba. Endemic C. lurida Suffrian 1868: 240; Gundlach 1891: 393 Cuba localities. Gu: Guantánamo (t.l.) (C. burida) Collections. New York. Washington (type) Distribution. Cuba. Endemic S. marginata Blake 1934: 53 Cuba localities. Distribution. Cuba. Endemic Collections. Gundlach no. 1904 Cuba localities. SS: Trinidad, SS: Buenos Aires (t.l.) Elytrogona Chevrolat 1837: 394 Collections. Washington (type) E. bulla Boheman 1862, 193; Chaboo 2000: 387 S. nigricans Zayas 1939: 255 E. ampulla Olivier 1808: 958; Gundlach 1891: Distribution. Cuba. Endemic 400 (misidentifications) Cuba localities. SC: Loma del Gato (t.l.) Distribution. Cuba, Endemic Collections. Washington (type) Cuba localities. SC: Pico Turquino and vi- S. oteroi Zayas 1952: 72 cinity Distribution. Cuba. Endemic Collections. Havana. New York. Washing- Cuba localities. SS: Lomas de Trinidad (t.l.) ton Collections. Zayas (type)

Stoiba Spaeth 1909: 720 TRIBE ISCHYROSONYCHINI S. angusticollis (Suffrian) 1868: 239 (Chelymorpha); Gundlach 1891: 393 Physonota Boheman 1854: 90 Distribution. Cuba. Endemic P. vitrea (Boheman) 1854: 239 (Eurypepla); Cuba localities. PR: Rangel Jacquelin du Val 1857: 318; Gundlach 1891: Collections. Gundlach no. 1266 395,400 S. barroi Zayas 1952: 72 Distribution. Cuba, Bahamas? Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: between Pico Mocha and Collections. Gundlach no. 1144 Pico Joaquim (t.l.) Collections. Zayas (type) TRIBE DORYNOTINI S. bruneri Blake 1930: 219 Distribution. Cuba. Endemic Paratrikona Apaeth 1923: 65 Cuba localities. SC: Sierra Maestra (t.l.) P. lerouxii (Boheman) 1854: 183 (Batonota); Collections. Washington (type) Jacquelin du Val 1857: 318; Gundlach 1891: S. clarildae Zayas 1939: 253 394 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Yunque de Baracoa (t.l.) Cuba localities. M: Cárdenas. SS: Trinidad Collections. Havana. Washington (type). (Baría) Zayas Collections. Gundlach no. 50 S. decemmaculata Blake 1930: 221 P. turritella Blake 1937: 76 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Sierra Maestra (t.l.) Cuba localities. SC: Sierra Maestra (t.l.) Collections. New York. Washington (type) Collections. Washington (type) S. fascicollis Blake 1934: 54 Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 189

TRIBE Coptocycla Chevrolat 1837: 396 C. conflagrata Boheman 1855: 294; Gundlach 1891: Agroiconota Spaeth 1913: 142 398 A. camagueyana Zayas 1988: 146 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. Ca: Sabana. SC: Siboney. Collections. Gundlach no. 417 Type locality not given Note. Possibly belonging to the genus Agroiconota Collections. Zayas (type) (Shawn Clark, in litt.) A. cubana Blake 1970a: 125 C. dorsopunctata (Klug) 1829: 14 (Psalidonota); Distribution. Cuba. Endemic Jacquelin du Val 1857: 320; Gundlach 1891: Cuba localities. Ca: Julioa Agos. Ha: Alamar 395; Medvedev 1993: 9 (Psalidonota) (t.l.); Cojimar Distribution. Cuba, México Collections. Brno (type) Cuba localities. Widespread A. propinqua (Boheman) 1855: 289 (Metriona); Collections. Gundlach no. 216. Medvedev Chaboo 2002: 716 Distribution. Cuba, Hispaniola, Jamaica, Deloyala Chevrolat 1837: 371 Puerto Rico, Nicaragua to Colombia and Ven- D. guttata (Olivier) 1790: 383 (Coptocycla); ezuela Gundlach 1891: 398 (C. gattuta); Medvedev Cuba localities. 1993: 10 Collections. Distribution. Cuba, Hispaniola, Puerto Rico, México, South America, e USA Charidotella Weise 1896: 13 Cuba localities. Widespread C. bisbinotata (Boheman) 1855: 144 (Coptocycla); Collections. Gundlach no. 217. Medvedev Gundlach 1891: 396; Blake 1966b: 221 (Metriona); Borowiec 1989: 214 (transferred Floridocassis Spaeth 1952: 348 from Coptocycla); Medvedev 1993: 10 F. repudiata (Suffrian) 1868: 249 (Coptocycla); (Metriona) Gundlach 1891: 398; Riley 1986: 111 Distribution. Cuba, Puerto Rico Distribution. Cuba, USA (SC-FL-MS). Host: Cuba localities. Gr: Bayamo. Ha: Miranao, Ipomoea sagittata Poir. coast. M: Cárdenas Cuba localities. Collections. Gundlach no. 1148. Medvedev Collections. Gundlach no. 1917 C. oblita (Suffrian) 1868: 247 (Coptocycla); Gundlach 1891: 398 Plagiometriona Spaeth 1899: 219 Distribution. Cuba. Endemic P. tortuguilla Zayas 1988: 148 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Gu: Tortuguilla (t. l.); on C. quadrisignata (Boheman) 1855: 150 Rochfortia steliata Bon., a spiny plant (Solan- (Coptocycla); Gundlach 1891: 397; Blake 1966b: aceae) 220 (Metriona); Medvedev 1993: 10 (Metriona) Collections. Zayas (type) Distribution. Cuba, Hispaniola, Puerto Rico Cuba localities. Widespread SUBFAMILY CHRYSOMELINAE Collections. Gundlach no. 218. Medvedev TRIBE CHRYSOMELINI C. sexpunctata (Fabricius) 1781: 109 (Cassida); SUBTRIBE DORYPHORINA Jacquelin du Val 1857: 319 Distribution. Cuba, widespread in West Leptinotarsa Stål 1838: 475 Indies, Canada to Argentina L. undecimlineata Stål 1858: 316; Medvedev 1993: Cuba localities. 7 Collections. Distribution. Cuba, Central and South C. subsignata (Boheman) 1862: 402 (Coptocycla) America. Solanum torvum is the usual host Distribution. Cuba, Hispaniola, Puerto Rico Cuba localities. Widespread Cuba localities. Collections. Medvedev Collections. Leucocera Stål 1858: 477 L. garciai Zayas 1988: 134 190 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba. Endemic L. semilutea Stål 1860: 459 Cuba localities. Gu: Pinares de Baracoa. Ha: L. semicaerulea Achard 1923: Daccordi 1995: 38 Cayo Piedra, Cocodrilo. Ho:. Moa, El Johnson. Distribution. Cuba. Endemic IJ. Type locality not given Cuba localities. Collections. Zayas (type) Collections. L. guanahacabibensi Zayas 1988: 135 L. sexguttata Chevrolat 1858: 211; Gundlach 1891: Distribution. Cuba. Endemic 366 (Crysomela); Daccordi 1995: 36; Medvedev Cuba localities. PR: Guanahacabibes (t.l.) 19934: 7 Collections. Zayas (type) Distribution. Cuba. Endemic L. hexaspila (Suffrian) 1867: 296 (Chrysomela); Cuba localities. Gr: Cabo Cruz. Gu: Gundlach 1891: 369; Daccordi 1995: 38 Caimanera; Guantánamo; Tortuguilla. SC: Si- L. amarella (Suffrian) 1867: 298 (Chrysomela); erra Mestra; Varsalles Gundlach 1891: 369; Daccordi 1995; 38 Collections. Gundlach no. 1264. Havana. Distribution. Cuba. Endemic Medvedev Cuba localities. L. testaceipes Stål 1858: 477 (Chrysomela); Collections. Gundlach no. 1876 Gundlach 1891: 366 L. ileanae Daccordi 1995: 40 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Gu: Tortuguilla (t.l.) Collections. Collections. Havana (type) L. insulana (Jacquelin du Val) 1856: 302 SUBTRIBE CHRYSOMELINA (Chrysomela); Gundlach 1891: 365; Daccordi 1995: 38 Chrysomela Linnaeus 1758: 368 L. insularis Stål 1858: 478; Daccordi 1995: 38 C. (Microdera) cruentipennis Jacquelin du Val 1856: Distribution. Cuba, Hispaniola 125; Gundlach 1891: 369 Cuba localities. Western Cuba Distribution. Cuba. Endemic Collections. Gundlach no. 220 Cuba localities. M: Cardenas; Cienaga de L. nydia (Stål) 1865: 203 (Chrysomela); Gundlach Zapata 1891: 366. Probably a synonym of L. Collections. Gundlach no. 323 testaceipes; Daccordi 1995: 42 Distribution. Cuba. Endemic Phaedon Latreille 1829: 151 Cuba localities. Widespread. P. cubensis Blake 1960: 98 Collections. Gundlach no. 1469 Distribution. Cuba. Endemic L. optica Suffrian 1867: 293; Gundlach 1891: 368 Cuba localities. SC: Ciudamar (t.l.) Distribution. Cuba. Endemic Collections. Washington (type). Zayas Cuba localities. M: Cárdenas P. splendicans Stål 1860: 470; Gundlach 1891: 370 Collections. Gundlach no. 433 (Chrysomela splendida) L. poeyi (Chevrolat) 1838: 285 (Chrysomela); Chrysomela splendidum Suffrian 1867: 300 Jacquelin du Val 1857: 301; Gundlach 1891: Distribution. Cuba. Endemic 368; Medvedev 1993: 7 Cuba localities. Gr: Bayamo L. apicicornis (Chevrolat) 1838: 286 Collections. Gundlach no. 1146 (Chrysomela); Gundlach 1891: 367 (Gundlach P. zayasi Blake 1960: 100 no. 221); Daccordi 1995: 36; Medvedev 1993: 7 Distribution. Cuba. Endemic L. cubana Stål 1858: 477; Daccordi 1995: 36 Cuba localities. SC: Gran Piedra (t.l.) L. rubropustalata (Suffrian) 1867: 295 Collections. Zayas (type female) (Chrysomela); Gundlach 1891: 368 (Gundlach no. 1875); Daccordi 1995; 36 Plagiodera Chevrolat 1837: 428 Distribution. Cuba. Endemic P. bicolor Zayas 1988: 135 Cuba localities. Ci: Cienfuegos. Gu: Distribution. Cuba. Endemic Tortuguilla. Ha: Las Canas; Litoral de Cuba localities. M: San Miguel de los Baños Marianao; Santiago de las Vegas. M: Cardenas; (t.l.) Cuabales de San Francisco. SC: Juraguá Collections. Zayas (type) Collections. Gundlach 1891: 131. Havana. P. cubensis (Blake) 1960: 98 (Plagiodera); Medvedev Balsbaugh and Daccordi 1987: 36 A Checklist of the Beetles of Cuba 191

Distribution. Cuba. Endemic Cuba localities. Ca: Camaguey. Ha: Santiago Cuba localities. SC: Ciudad del Mar (t.l.) de la Vegas (t.l.). LT: Omaja Collections. Washington (type) Collections. Washington (type) P. zayasi (Blake) 1960: 100; Balsbaugh and Daccordi N. insularis (Harold) 1875: 93 (Chthoneis); Blake 1987: 38 1963: 14 Distribution. Cuba. Endemic N. loricata Suffrian 1857: 311 (Diabrotica); Cuba localities. SC: Gran Piedra (t.l.) Gundlach 1891: 373 Collections. Zayas (type) Distribution. Cuba. Endemic Cuba localities. Widespread SUBFAMILY Collections. Gundlach no. 32. Washington TRIBE GALERUCINI N. vittata (Blake) 1959a: 180 (Chthoneis) Distribution. Cuba. Endemic Miraces Jacoby 1888: 611 Cuba localities. Ho Moa; Piloto (t.l.) M. barberi (Blake) 1951b: 141 (replacement name Collections. Washington (type male); Zayas for Dicoelotrachelus violaceus Blake) M. violaceus (Blake) 1948a: 121 Yingaresca Bechyne 1956: 298 (Dicoelotrachelus) Y. maculata (Blake) 1970a: 121 (Galerucella) Distribution. Cuba, Puerto Rico Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Pico Turquino (t.l.) Collections. Collections. Brno (type) M. dichroa (Suffrian) 1868: 203 (Haltica); Gundlach Y. maculipes (Blake) 1930: 217 (Galerucella) 1891: 384; Blake 1951b: 140 (Halticidea) Distribution. Cuba. Endemic Asbecesta violaea Allard 1890: 87 Cuba localities. SC: Pico Turquino summit Distribution. Cuba. Endemic (t.l.) Cuba localities. SC: Santiago de Cuba Collections. Washington (type) Collections. Gundlach no. 1500 Y. melanocephala (Blake) 1959a: 178 (Galerucella) Distribution. Cuba. Endemic Monocesta Clark 1965: 264 Cuba localities. Ho: Moa, La Breña (t.l.); M. cubensis Blake 1959a: 178 (properly belongs in Moa, Piloto. Gu: Yunque Trirhabda Leconte 1865: 219; S. Clarke, in litt.) Collections. Washington (type male) Distribution. Cuba. Endemic Y. oteroi (Blake) 1934: 49 (Galerucella) Cuba localities. Ho: Moa, La Breña (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type female) Cuba localities. SS: Trinidad, SS: Buenos M. opacipennis (Jacquelin du Val) 1856: 304 Aires (t.l.) (Coelomera); Gundlach 1891: 371 (Galleruca) Collections. Washington (type) Distribution. Cuba. Endemic Y. venustala (Suffrian) 1867: 303 (Galerucella); Cuba localities. M: Cárdenas Gundlach 1891: 371 (G. veniestula) Collections. Gundlach no. 149 Distribution. Cuba. Endemic Cuba localities. Gu: Baracoa. M: Cárdenas. Neolochmaea Laboissiere 1939: 153 On ateje (Cordia) (N. dilatipennis (Jacoby) 1886: 487) Collections. Gundlach no. 322 Distribution. Cuba (?), widespread from Florida to South America, and on many West TRIBE LUPERINI Indian islands Cuba localities. Acalymma Barber 1947: 154 Collections. A. annulatum (Suffrian) 1867: 307 (Diabrotica); N. immaculata (Blake) 1938: 48 (Galerucella) Gundlach 1891: 372 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico Cuba localities. Gu: Mts. N Imias (t.l.) Cuba localities. Widespread Collections. Cambridge (type) Collections. Gundlach no. 1575 A. bivittatum (Fabricius) 1801: 455 (Crioceris); Nyctiplanctus Blake 1963: 15 Gundlach 1891: 372 N. insulana (Blake) 1946b: 113 (Chthoneis) Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba. Endemic Grenada, St. Vincent, South America 192 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Widespread. In calabaza Cuba localities. Widespread (squash) flowers Collections. Gundlach no. 430 Collections. Gundlach no. 153 D. relicta Suffrian 1867: 310; Gundlach 1891: 373 A. innubum (Fabricius) 1775: 117 (Crioceris); Distribution. Cuba. Endemic Jacquelin du Val 1857: 305 (Diabrotica); Cuba localities. M: Cárdenas Gundlach 1891: 371 Collections. Gundlach no. 393 Distribution. Cuba, Puerto Rico, Guadeloupe, México to S. America Dicoelotrachelus Blake 1941: 171 Cuba localities. Widespread. In calabaza D. cubensis Blake 1946b: 111 (squash) flowers Distribution. Cuba. Endemic Collections. Gundlach no. 31 Cuba localities. SC: Loma del Gato (t.l.) Collections. Cambridge (type) Cerotoma Chevrolat 1837: 403 D. sulcatus Blake 1946b: 112 C. ruficornis (Olivier) 1791: 200; Medvedev 1993: 8 Distribution. Cuba. Endemic (Cerocoma); Fernandez Garcia 1998b: 8 Cuba localities. Gu: Mts. N of Imias (t.l.) C. denticornis (Fab.) 1801: 457; Jacquelin du Val Collections. Cambridge (type) 1857: 304; Gundlach 1891: 374 Distribution. Cuba, Jamaica, Hispaniola, Ectmesopus Blake 1940: 96 Puerto Rico, USA to Costa Rica E. cubensis Zayas 1988: 137 Cuba localities. Ha: Loma de Camoa. IJ: Distribution. Cuba. Endemic Nueva Gerona. M: Cárdenas. SS: Trinidad. An Cuba localities. SC: Juraguá (t.l.) important pest of beans (Heyer et al. 1993, listed Collections. Zayas (type) as Andrector ruficornis (Oliv.)) E. malachioides (Suffrian) 1867: 319 (Luperus); Collections. Gundlach no. 375. Medvedev. Gundlach 1891: 375 Pittsburgh Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Coelomera Chevrolat 1837: 399 Collections. Gundlach no. 138 C. liturata Suffrian 1876: 300; Gundlach 1891: 370; E. nigrolimbatus Blake 1959a: 181 Blake 1959a: 178 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ho: Moa; Piloto (t.l.) Cuba localities. Gu: Caimanera; Collections. Zayas (type female) Guantánamo E. occipitalis Blake 1940: 100 Collections. Gundlach no. 1487 Distribution. Cuba. Endemic Cuba localities. Ci: Soledad. Ha: Cojimar (t.l.) Diabrotica Chevrolat 1837: 404 Collections. Cambridge. Washington (type) D. balteata LeConte 1865: 213; Medvedev 1993: 7 E. placidus (Suffrian) 1867: 320 (Luperus); Distribution. Cuba; widespread in USA, Gundlach 1891: 375 south to Colombia Distribution. Cuba. Endemic Cuba localities. Ha: Santiago de las Vegas. Cuba localities. PR: San Cristobal. An important pest of beans Collections. and corn in Cuba (Heyer et al. 1989; 1993) E. Blake 1966a: 243 Collections. Medvedev Distribution. Cuba. Endemic D. graminea Baly 1886: 443 Cuba localities. Ha: San Jose de las Lajas; Distribution. Cuba, Puerto Rico Somorrostro. PR: Bahia Honda (t.l.) Cuba localities. Collections. Washington (type) Collections. E. zayasi Blake 1959a: 181 D. impressa Suffrian 1867: 309; Gundlach 1891: Distribution. Cuba. Endemic 373 Cuba localities. Ha: San Jose de las Lajas Distribution. Cuba. Puerto Rico (t.l.); Somorrostro Cuba localities. Gu: Baracoa; Monte Toro Collections. Zayas (type male) Collections. Gundlach no. 1486 D. pulchella Jacquein du Val 1856: 307; Gundlach Erynephala Blake 1936: 425 1891: 372; Blake 1956; 143 (illustration) (E. maritima (LeConte) 1865: 218) Distribution. Cuba, Central America A Checklist of the Beetles of Cuba 193

Distribution. Jamaica, Canada (NS), USA Altica Müller 1764: 14 (TX-FL-MA) A. purpurascens (Suffrian) 1878: 199 (Haltica); Cuba localities. Gundlach 1891: 383 (H. purpurasiens) Collections. Distribution. Cuba, Puerto Rico Cuba localities. Gr: Guisa. SC: Santiago de Isotes Weise 1922: 64 Cuba (I. albicornis (Fabricius) 1798: 96) Collections. Gundlach 1491: 1491 Diabrotica thoracica Fabricius 1801: 457; Note. Jacquelin du Val reported A. oleracea from Gundlach 1891: 374 Cuba, but this is a European species Distribution. Hispaniola, Puerto Rico (old A. subaurea (Suffrian) 1868: 199 (Haltica); Cuba records may be based on very similar Gundlach 1891: 383 Synbrotica darlingtoni (Blake) 1937: 69 of Distribution. Cuba. Endemic Hispaniola, and possibly of Cuba Cuba localities. Gr: Bayamo. M: Cárdenas Cuba localities. Collections. Gundlach no. 425 Collections. Gundlach no. 283 Aphthona Chevrolat 1842: 5. Note. True Leptonesiotes Blake 1958: 78 Aphthona are confined to the Old World. These L. cyanospila (Suffrian) 1867: 313 (Diabrotica); species will need to be reclassified (Shawn Gundlach 1891: 373 Clark, in litt.) Distribution. Cuba. Endemic A. auripennis (Suffrian) 1868: 212 (Haltica); Cuba localities. PR: Rangel Gundlach 1891: 386 Collections. Gundlach no. 1268 Distribution. Cuba, Puerto Pico L. quadrimaculata Blake 1959a: 182 Cuba localities. Ca: Manzanillo Distribution. Cuba. Endemic Collections. Gundlach no. 1499 Cuba localities. Ci: Rancho Luna (t.l.) A. compressa Suffrian 1868: 211; Gundlach 1891: Collections. Zayas (type female) 385 L. semicyaneus (Suffrian) 1867: 314 (Diabrotica); Distribution. Cuba, West Indies (where?) Gundlach 1891: 374 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 371 Cuba localities. Gu: Baracoa (A. maculipennis Jacoby 1885: 365; see Blake 1964: Collections. Gundlach no. 1485 10) Distribution. Puerto Rico, México, Guate- TRIBE ALTICINI mala, Panamá The flea beetles Cuba localities. Collections. Aedmon Clark 1860: 129 A. nana Blake 1949: 368 A. ferruginea (Suffrian) 1868: 174 (Hadropoda); Distribution. Cuba. Endemic Gundlach 1891: 378.;Blake 1943: 413 Cuba localities. CA: Baraguá. Ha: Havana; Distribution. Cuba. Endemic Ho: Sierra de Anafe, Caimito. (t.l.) Cuba localities. M: Cardenas; Cienaga de Collections. Washington (type) Zapata. ?Guamacaro A. ruficollis Blake 1970b: 123 Collections. Gundlach no. 979 Distribution. Cuba. Endemic A. turquinesis (Blake) 1943: 429; (Hadropoda) Cuba localities. M: Arroyo Bermejo (t.l.) Distribution. Cuba. Endemic Collections. Brno (type) Cuba localities. SC: Pico Turquino (t.l.) A. schwarzi Blake 1949: 368 Collections. Harvard (type) Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Alagoasa Bechyné 1955: 8 Collections. Washington (type) [(A. bicolor (Linnaeus) 1767: 593; Jacquelin du Val 1857: 308 (Oedionychis))] Apraea Baly 1877: 293 Distribution. Puerto Rico, Virgin Islands; A. bruneri (Blake) 1948a: 123; (Glyptobregma) probably not Cuba (see Blake 1940: 173) Distribution. Cuba. Endemic Cuba localities. Cuba localities. SS: Trinidad; Buenos Aires Collections. (t.l) 194 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Washington (type) Blepharida Chevrolat 1836: 394 A. clathrata (Suffrian) 1868: 189 (Glyptobregma); B. irrorata Chevrolat 1864: 182; Suffrian 1867: 323; Gundlach 1891: 380 (Haltica); Blake 1951: 140 Gundlach 1891: 376; Blake 1938: 49; 1949: 367; Distribution. Cuba. Endemic Furth 1998: 29; Medvedev 1993: 9 Cuba localities. Haltica adspersula Suffrian 1868: 185; Gundlach Collections. Gundlach no. 766 1891: 380 A. interstitiale (Suffrian) 1868: 187 (Glyptobregma); Distribution. Cuba, Puerto Rico Gundlach 1891: 380 (Haltica); Blake 1951b: 140 Cuba localities. Ca: Jobabo. CA: Baraguá; Distribution. Cuba. Endemic(Puerto Rico Central Jaronú. Ci: Soledad. Gr: Bayamo. M: records are misidentifications; Blake l947a: 92) Cardenas; Fermina. PR: Taco Taco. SC: Cuba localities. M: Cienaga de Zapata Santiago; Sierra Maestra. SS: Trinidad Collections. Gundlach no. 402 Collections. Basel. Cambridge. Gundlach no. A. minima Blake 1970a: 124 468; no. 1489 (B. adspersula). London (lecto- Distribution. Cuba. Endemic type). Medvedev. Washington Cuba localities. SS: Topes de Collantes (t.l.) Collections. Brno (type) Chaetocnema Stephens 1831: 325 A. pyritosa (Suffrian) 1868: 189 (Glyptobregma); C. apricaria (Suffrian) 1878: 219 (Plectroscelis); Gundlach 1891: 381(Haltica); Blake 1951b: 140 Gundlach 1891: 388; Blake 1941: 178; Medvedev Distribution. Cuba. Endemic 1993: 9 Cuba localities. M: Cardenas; Cienaga de Distribution. Cuba (Endemic); records from Zapata Jamaica, Hispaniola, and Puerto Rico may be Collections. Gundlach no. 871 errors (Blake 1941: 178) A. robusta (Suffrian) 1868: 191 (Glyptobregma); Cuba localities. Ci: Cayamas. M: Cárdenas. Gundlach 1891: 381 (Haltica); Blake 1951b: SC: 80 km W Santiago. SS: Trinidad Mts.; Mina 140 Carlota Distribution. Cuba. Endemic Collections. Cambridge. Gundlach no. 636. Cuba localities. PR: Rangel Medvedev. Washington Collections. Gundlach no. 1297 (C. brunnescens 1880: 259; Blake 1941: 180; White A. turquinense (Blake) 1947a: 94 (Glyptobregma); 1996: 46) Blake 1951b: 140 Distribution. Bahamas, West Indies (Blake Distribution. Cuba. Endemic 1941: 180), USA (FL) Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. Hosts: Conocarpus erectus; Collections. Harvard (type) Laguncularis recemosa, Flaveria linearis A. zapatensi Zayas 1988: 139 Collections. Distribution. Cuba. Endemic C. cyanoptera Blake 1964: 23 Cuba localities. M: Zapata Peninsula; San Distribution. Cuba. Endemic Blas (t.l.) Cuba localities. None given. Taken on avo- Collections. Zayas (type) cado Collections. Washington (type) Argopistes Motschulsky 1860: 236 C. obesula Leconte 1878: 414; Blake 1941: 180 A. coccinelloides (Suffrian) 1868: 223 (Argopus); Distribution. Cuba, Hispaniola, USA (FL) Gundlach 1891: 388 (A. couinelloides) Cuba localities. Widespread. Feeds on rice Distribution. Cuba. Endemic and sugarcane Cuba localities. M: Cárdenas Collections. Washington Collections. Gundlach no. 908 C. perplexa Blake 1941: 177 Distribution. Cuba, Puerto Rico; probably Asphaera Chevrolat 1842: 227 widespread in West Indies A. crucipennis (Suffrian) 1868: 168 (Oedionychis); Cuba localities. Ca: Camaguëy. Ci: Cayamas Gundlach 1891: 377 (t.l.); Soledad. Gu: Baracoa. Ha: Havana; S. de Distribution. Cuba. Endemic las Vegas. Ho: Sierra de Anafe, Caimito. LT: Cuba localities. M: Cienaga de Zapata Central Manatí. PR: Herradura. SS: Trinidad Collections. Gundlach no. 1496 Mts.; Mina Carlota Collections. Cambridge. Washington (type) A Checklist of the Beetles of Cuba 195

C. tuberculata (Suffrian) 1868: 218 (Plectroscelis); Cuba localities. Widespread Gundlach 1891: 389 Collections. Gundlach no. 822 Distribution. Cuba. Endemic D. matanzano Zayas 1988: 141 Cuba localities. Gu: Baracoa Distribution. Cuba. Endemic Collections. Gundlach no. 1502 Cuba localities. M: Cuabales de Corral Falso (C. vesca White 1996: 117) (t.l.) Distribution. USA (FL; only Dade and Mon- Collections. Zayas (type) roe Counties) D. sandersoni Zayas 1960a: 129 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ca: Camaguey, in field near airport (t.l.) Cyrsylus Jacoby 1891: 306 Collections. Champaign. Zayas (type male) C. cubensis Blake 1959b: 247 D. spilotrachela Blake 1928: 96; 1955: 39 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola, Puerto Rico Cuba localities. Ho: Sierra Cristal (t.l.) Cuba localities. Ca: Camagüey Collections. Washington (type). Zayas Collections. Washington

Disonycha Chevrolat 1844: 80 Epitrix Foudras 1860: 147 D. chlorotica (Olivier) 1808: 690 (Haltica); E. fasciata Blatchley 1918: 56 Gundlach 1891: 379 Distribution. Cuba, Bahamas, Puerto Rico, Distribution. Cuba, Puerto Rico, Guiana St. Vincent, Grenada, México to Panamá, USA Cuba localities. Widespread (Gulf Coast to FL); on Solanum; along saltwa- Collections. Gundlach no. 147 ter lagoons D. collata (Fabricius) 1801: 463 (Crioceris); Cuba localities. Gundlach 1891: 379; Vogt et al. 1979: 19 Collections. Distribution. Cuba, México to Panamá, USA E. fuscata (Jacquelin du Val) 1856: 312 Cuba localities. Gr: Cauto. M: Cienaga de Distribution. Cuba, Puerto Rico, Grenada, Zapata; Fermin. Feeds on alligator weed Guatemala? (Amaranthaceae, Polygonum sp.) Cuba localities. Collections. Gundlach no. 1488 Collections. D. conjugata Fabricius 1801: 495; Blake 1930: 210 E. parvula (Fabricius) 180l: 468; Furth and Savini Haltica costipennis Jacquelin du Val 1857: 310; 1996: 58 Gundlach 1891: 381 Distribution. Cuba, Puerto Rico, Grenada, Distribution. Cuba, USA (LA, FL, NC); on Guadeloupe, St. Vincent, Venezuela, México to Polygonum Panamá, USA Cuba localities. M: Cardenas; Cienaga de Cuba localities. Zapata Collections. Collections. Gundlach no. 419 (costipennis) D. fimbriolata (Suffrian) 1868: 177 (Haltica); Gioia Bechyné 1955: 77 Gundlach 1891: 379 G. polutima (Blake) 1964: 27 (Sidfaya); Savini and Distribution. Cuba. Endemic Furth 1999: 30 Cuba localities. Gu: Baracoa Distribution. Cuba. Endemic Collections. Gundlach no. 533 Cuba localities. SS: Trinidad, SS: Buenos D. guabiniquinaje Zayas 1988: 139 Aires (t.l.) Distribution. Cuba. Endemic Collections. Cambridge (type). Washington Cuba localities. M: Peninsula de Zapata (t.l.) Collections. Zayas (type) Heikertingerella Csiki 1940: 350 D. juragua Zayas 1988: 143 H. fusca Blake 1960: 102 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Juraguá (t.l.) Cuba localities. Ha: Havana,; at light on Collections. Zayas (type) shipboard (t.l.) D. marginipennis (Suffrian) 1868: 179 (Haltica); Collections. Washington (type) Gundlach 1891: 379 H. minima (Suffrian) 1868: 184 (Haltica); Gundlach Distribution. Cuba. Endemic 1891: 380; Blake 1960: 101 196 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Hispaniola H. latum Blake 1950: 20; Blanco and Duckett 2001: Cuba localities. Ci: Playa Ingles. PR: Sierra 11 Rangel. SC: Santiago de Cuba. SS: Trinidad; Distribution. Cuba. Endemic Buenos Aires Cuba localities. Ci: Clayamas. Gr: Upper Collections. Gundlach no. 150l. Washington. Yara Valley (t.l.). Gu: Guantanamo Zayas Collections. Washington (type) H. manni Blake 1950: 24; Medvedev 1993: 8; Blanco Hemilactica Blake 1937: 72 and Duckett 2001: 13 H. clara Blake 1959b: 244 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas. Ho: Felton; Cuba localities. Ho: Moa, Piloto; La Breña, Antilla Bay (t.l.); Sierra de Nipe, Pinares de (t.l.) Mayari. PR: Bahia Honda. SC: 80 km W Collections. Washington (type). Zayas Santiago H. crucifera Blake 1959b: 245 Collections. Cambridge. Ivie. Medvedev. Distribution. Cuba. Endemic Washington (type) Cuba localities. Ho: Sierra Cristal (t.l.) H. orientense Blake 1950: 23; Blanco and Ducket Collections. Zayas (type) 2001: 14 H. fasciata Blake 1938: 50 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Guantánamo (t.l.). SC: Cuba localities. Gu: Upper Ovando River (t.l.) Daiquirí; Gran Piedra; Juragua Collections. Cambridge (type) Collections. Cambridge. Havana. Washing- H. pulchella Blake 1937: 73 ton (type) Distribution. Cuba. Endemic H. ornatum Blake 1950: 16; Blanco and Duckett Cuba localities. SC: Jarahueca (t.l.) 2001: 15 Collections. Washington (type) Distribution. Cuba. Endemic H. quatuordecimpunctata (Suffrian) 1868: 206 Cuba localities. Ci: Cayamas (t.l.). Gu: (Haltica); Gundlach 1891: 384 Baracoa, Cupeyal. PR: Candelaria; Cerro Distribution. Cuba. Endemic Cabras; Pinares de Guanito Cuba localities. Gr: Bayamo. M: Cárdenas Collections. Havana. New York. Washington Collections. Gundlach no. 534 (type) H. rugosa Blake 1937: 74 H. sasha Blanco and Duckett 2001: 16 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Palma Mocha (t.l.) Cuba localities. Ci: Cumanayuagua, Collections. Washington (type) Guajímico (t.l.), on Stigmaphylon H. stomachosa (Suffrian) 1868: 204 (Haltica); Collections. Washington (type) Gundlach 1891: 384; Blake 1959b: 246 Distribution. Cuba. Endemic Leptophysa Baly 1877: 165 Cuba localities. PR: Pan de Guajaibon; L. darlington (Blake) 1945: 92 (Pseudoepitrix) Guajaibon; Rangel Distribution. Cuba. Endemic Collections. Gundlach no. 1275. Washington. Cuba localities. SC: Pico Turquino (t.l.) Zayas Collections. Cambridge (type). Washington L. longicornis (Blake) 1945: 90 (Pseudoepitrix) Homoschema Blake 1950: 10 (key in Blake 1950; Distribution. Cuba. Endemic Blanco and Duckett 2001) Cuba localities. Gu: Mt. N Imias (t.l.) H. cubensis Medvedev 1993: 8, 14 Collections. Cambridge (type). Washington Distribution. Cuba. Endemic L. tetraspilota (Blake) 1945: 90 (Pseudoepitrix) Cuba localities. Ha: Santa Lucia (t.l.) Distribution. Cuba. Endemic Collections. Medvedev (type, female) Cuba localities. SC: Pico Turquino (t.l.) H. darlingtoni Blake 1950: 24 Collections. Cambridge (type). Washington Distribution. Cuba. Endemic Cuba localities. SC: coast below Pico Longitarsus Latreille 1829: 155 Turquino (t.l.) L. impennatus Blake 1970a: 122 Collections. Cambridge (type). Washington Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 197

Cuba localities. Ha: Alamar. .M: Arroyo M. adulta (Suffrian) 1868: 222 (Dibolia); Gundlach Bermejo (t.l.); Varadero 1891: 388; Blake 1952: 2 Collections. Brno (type). Washington Distribution. Cuba. Endemic L. prokopi Blake 1970a: 123 Cuba localities. ? Guamacoro Distribution. Cuba. Endemic Collections. Gundlach no. 1214 Cuba localities. Ha: Alamar (t.l.); Cojimar M. coeruleipennis Blake 1959b: 247 Collections. Brno (type). Washington Distribution. Cuba. Endemic L. pterotus Blake 1970a: 122 Cuba localities. Ci: Cumanayagua (t.l.). PR: Distribution. Cuba. Endemic Guao; Corralillo Cuba localities. Ha: Alamar (t.l.) Collections. Washington (type). Zayas Collections. Brno (type). Washington M. rubropustulata (Suffrian) 1868: 221 (Dibolia); L. seminulus Suffrian 1868: 216; Gundlach 1891: Gundlach 1891: 388; Blake 1952: 2 389 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. M: Cienaga de Zapata Collections. Gundlach no. 57 Collections. Gundlach no. 1106 M. tababuiae Blake 1937: 75 L. subaeneus Suffrian 1868: 216; Gundlach 1891: Distribution. Cuba. Endemic; (not Puerto 389 (L. subaencus) Rico) Distribution. Cuba. Endemic Cuba localities. Ha: Santiago de las Vegas Cuba localities. M: Cárdenas (t.l). Feeding on Tabebuia pentaphylla Collections. Gundlach no. 838 Collections. Washington (type) L. varicornis Suffrian 1868: 215; Gundlach 1891: 389; Furth and Savini 1996: 60 Monomacra Chevrolat 1837: 413 Distribution. Cuba, Puerto Rico, Antilles, M. religata Jacquelin du Val 1856: 311; Gundlach México to Venezuela, USA 1891: 381; Blake 1946c: 267 (Lactica) Cuba localities. Ha: Madruga. M: Cárdenas [M. tibialis Olivier 1808: 697]. in USA (TX-FL- Collections. Gundlach no. 71 MD), probably not in Cuba; Blake 1946c, 267; Medvedev 1993: 8 (Cuba records as M. tibialis) Lysathia Bechyné 1959: 303 Distribution. Cuba. Endemic L. occidentalis (Suffrian) 1868: 197 (Haltica); Cuba localities. Gu: Monte Toro. Ha: Banes. Gundlach 1891: 382 M: Cárdenas. SC: Sierra Maestra, San Diego Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Gundlach no. 146. Medvedev Virgin Islands, Jamaica, Dominica, St. Lucia, México, Guadeloupe Nesaecrepida Blake 1964: 21 Cuba localities. N. asphaltina (Suffrian) 1868: 201 (Haltica); Collections. Gundlach no. 237 Gundlach 1891: 383 Distribution. Cuba, Puerto Rico, Jamaica Macrohaltica Bechyné 1959: 305 Cuba localities. IJ: Nueva Gerona. M: (M. amethystina (Olivier) 1808: 687) Cárdenas Distribution. West Indies (where?), Central Collections. Gundlach no. 427 and South America N. rufomarginata Blake 1964: 22 Cuba localities. Distribution. Cuba, Jamaica Collections. Cuba localities.“Jababa” (=Ha: Jabaco?) (t.l.) M. jamaicensis (Fabricius) 1792: 16; Furth and Collections. Washington (type) Savini 1996: 61; Medvedev 1993: 8 Distribution. Cuba, Jamaica, Hispaniola, Oedionychus Berthold 1827: 401 (an Old World Puerto Rico, Nicaragua to Colombia genus and the species below probably belong in Cuba localities. Ha: Lomas de Canoa. VE: Kuschelina, Capraita, or Alagoasa) Santa Clara O. amplilimbatus Blake 1959b: 244 Collections. Medvedev Distribution. Cuba. Endemic Cuba localities. PR: Guanahacabibes (t.l.) Megistops Boheman 1859: 186 (see key in Blake Collections. Zayas (type) 1952) O. complanatus Suffrian 1868: 169; Gundlach 1891: 377 198 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba. Endemic O. zayasi Blake 1959b: 242 Cuba localities. SC: Santiago de Cuba Distribution. Cuba. Endemic Collections. Gundlach no. 1579 Cuba localities. PR: Guanahacabibes (t.l.) O. cristalensis Blake 1959b: 242 Collections. Washington (type). Zayas Distribution. Cuba. Endemic Cuba localities. Ho: Sierra Cristal (t.l.) Omophoita Chevrolat 1837: 410 Collections. Zayas (type) (O. albicollis (Fabricius) 1787: 76; Blake 1931: 77) O. cubanus Harold 1875: 90 Distribution. Hispaniola, Puerto Rico, Vir- Distribution. Cuba. Endemic gin Islands, Lesser Antilles, Trinidad, Guyana, Cuba localities. None given Brazil Collections. Cuba localities. O. decempunctulatus Csiki 1940: 441. Collections. O. decempunctatus Suffrian 1868: 170; Gundlach O. cyanipennis( Fabricius) 1798: 97 (Galleruca); 1891: 378 Jacquelin du Val 1857: 308; Gundlach 1891: Distribution. Cuba. Endemic 377 (Oedionychis) Cuba localities. Gr: Sur de Bayamo Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Gundlach no. 1490 Virgin Islands, Jamaica, USA (FL, TX) [O. fasciatus (Fabricius) 1798: 96 (Galleruca); Cuba localities. Widespread Gundlach 1891: 377] Collections. Gundlach no. 222 O. interruptus Illiger 1807: 94 (see Blake 1940: 174) Phyllotreta Chevrolat in Dejean 1837: 391 (see Distribution. Hispaniola; reported from but White 1970) probably not in Cuba (see Blake 1940: 174; P. fallaciae Csiki 1939: 64 records probably of O. pseudofasciata, see be- Haltica fallax Suffrian 1868: 214; Gundlach 1891: low) 386 O. orientensis Blake 1959b: 241 Distribution. Cuba, Puerto Rico Distribution. Cuba. Endemic Cuba localities. Gr: Bayamo. M: Cienaga de Cuba localities. Ho: Moa; El Johnson (t.l.) Zapata Collections. Washington (type). Zayas Collections. Gundlach no. 978 O. pictus Suffrian 1868: 173; Gundlach 1891: 378 (O. prieta) Pseudodisonycha Blake 1954: 248 Distribution. Cuba. Endemic P. darlingtoni (Blake) 1938: 50 (Disonycha); Cuba localities. M: Cárdenas Medvedev 1993: 8 Collections. Gundlach no. 33 Distribution. Cuba. Endemic O. pseudofasciatus Blake 1940: 173; Gundlach Cuba localities. SC: Pico Turquino (t.l.); Si- 1891: 377 (as O. fasciataus Fabricius) erra Maestra Distribution. Cuba. Endemic Collections. Cambridge (type). Medvedev Cuba localities. Ca: Jaronú. Ci: Soledad. Gu: P. turquinesis Blake 1956: 144 Imias. Ha: Havana. M: Cárdenas. SS: Buenos Distribution. Cuba. Endemic Aires Cuba localities. SC: Pico Turquino (t.l.) Collections. Gundlach no. 295 Collections. Washington (type) O. pseudothoracica Blake 1945: 90 Distribution. Cuba. Endemic Strabala Chevrolat 1837: 413 Cuba localities. Ca: Camaguey (t.l.) S. ambulans (Suffrian) 1868: 182 (Altica); Collections. Washington (type) Gundlach 1891: 380; Blake 1953: 131 O. tortuosus Suffrian 1868: 171; Gundlach 1891: Distribution. Cuba, USA (FL) 378 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Cambridge. Gundlach no. 145. Cuba localities. M: Cárdenas Washington Collections. Gundlach no. 856 S. intermedia Jacquelin du Val 1857: 311; Gundlach O. turquino Zayas 1988: 144 1891: 380 (as synonym of S. amblans); Blake Distribution. Cuba. Endemic 1953: 133 Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba, (subspecies S. i. Collections. Zayas (type) costaricensis Blake (1953) in Costa Rica) A Checklist of the Beetles of Cuba 199

Cuba localities. Ca: Aguada. CA: Baraguá; Collections. Cambridge (type) Central Jaronú. Ci: Cayamas; Soledad. Gr: M. adustum Suffrian 1866: 339; Gundlach 1891: Upper Yara Valley. Gu: Mt. N. of Imias 363; Blake 1930: 214; 1970b: 51 Collections. Cambridge M. terminale Horn 1892: 215 S. rufa Illiger 1807: 152; Furth and Savini 1996: Distribution. Cuba, Grand Cayman, Baha- 66 mas (Andros), USA (FL) Distribution. Cuba, Jamaica, Puerto Rico, Cuba localities. CA: Baraguá. Ci: Cayamas. México to Brazil, USA, Canada Ha: Hershey (now Camilo Cienfuegos). M: Cuba localities. Cárdenas. PR: San Vicente. SC: Santiago Collections. Collections. Gundlach no. 256. Washington M. castaneum Blake 1965: 5; 1970b: 59 Syphrea Baly 1876: 447 Distribution. Cuba. Endemic S. nigrorubra (Blake) 1951b: 144 (Hermaeophaga) Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. SC: Palma Moche (t.l.) M. cornutum Blake 1958: 93; 1970b: 54. Collections. Washington (type) Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Systena Chevrolat 1837: 414 Collections. Washington S. basalis (Jacquelin du Val) 1857: 312 (Haltica); M. cubaecola Clavareau 1914: 88 Gundlach 1891: 386; Blake 1931: 80, 1935: 105, M. puncticollis Suffrian 1866: 341; Gundlach 1951b: 140, 1964: 18; Furth and Savini 1998: 1891: 363; species unknown to Blake 1970b: 60 136; Medvedev 1993: 8 Distribution. Cuba. Endemic S. ornata Baly 1877: 288 Cuba localities. M: Cienaga de Zapata Distribution. Cuba, Jamaica, Hispaniola, Collections. Gundlach no. 977 Puerto Rico (México, Guatemala, Nicaragua?) M. ensiforma Blake 1947b: 310; 1970b: 59 Cuba localities. Widespread. A pest on cot- Distribution. Cuba. Endemic ton, tobacco, and many vegetables such as beans Cuba localities. SC: Pico Turquino (t.l.) and maize (Blake 1951: 140; Heyer et al. 1989, Collections. Cambridge (type). Washington 1993) M. gracile Blake 1937: 67; 1970b: 54 Collections. Gundlach no. 279. London. M. cavicollis Blake 1960: 97. PR: Sierra Medvedev. Washington Cajalbana (t.l.). Washington (type). Zayas S. coeruleipennis (Suffrian) 1868: 213 (Haltica); Distribution. Cuba. Endemic Gundlach 1891: 386 (H. coerulcipennis) Cuba localities. Ha: Santiago de las Vegas Distribution. Cuba. Endemic (t.l.). PR: Sierra Cajalbana. SS: Buenos Aires Cuba localities. Ci: Cumanayagua; Corralillo. Collections. Washington (type) Ha: Madruga. M. grande Blake 1958: 98; 1970b: 53 Collections. Gundlach no. 60 Distribution. Cuba. Endemic S. plicata Blake 1959b: 246 Cuba localities. Gu: Yunque de Baracoa (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Ho: Sierra Cristal (t.l.). M: M. gundlachi Fernandez García 1998: 1 Rio San Juan, Morrillo Distribution. Cuba. Endemic Collections. Washington (type). Zayas Cuba localities. CA: Cayo Coco (t.l.); Cayo Note. This is a homonym of S. plicata Blatchley Guillermo 1921: 26, and a replacement name is needed Collections. Havana (type) (S. Clark, in litt.) M. imitans Blake 1947b: 310; 1970b: 58 Distribution. Cuba. Endemic SUBFAMILY EUMOLPINAE Cuba localities. SC: Aguadores (t.l.) TRIBE TYPOPHORINI Collections. Cambridge (type M. laeviusculum Suffrian 1866: 343; Gundlach Metachroma Chevrolat 1837: 436; Blake 1970b: 1891: 364; species unknown to Blake 1970b: 60 50 (key to Cuban species) M. sordidum Suffrian 1866: 345; Gundlach 1891: M. acunai Blake 1946a: 22; 1970b: 55 364 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ho: Moa (t.l.) Cuba localities. M: Cienaga de Zapata 200 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Collections. Gundlach no. 975 (no 976, Cuba localities. Ha: Santiago de las Vegas. sordidum) PR:Bahia Honda (t.l.); Baños de San Vicente; M. lituratum Suffrian 1866: 343; Gundlach 1891: Pinar del Rio 363; species unknown to Blake 1970b: 60 Collections. Cambridge. Washington (type) Distribution. Cuba. Endemic M. sordidum Suffrian 1857: 345; species unknown Cuba localities. PR: Rangel to Blake 1970b: 60 Collections. Gundlach no. 1279 Distribution. Cuba. Endemic M. longicornis Blake 1966b: 216; 1970b: 58 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Soledad (t.l.) M. suffriani Clavareau 1914: 89; replacement name Collections. Cambridge (type) M. suturalis Suffrian 1866: 342; Gundlach 1891: M. longitarsus Blake 1946a: 24; 1970b: 53 363; species unknown to Blake 1970b: 60 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. Western Cuba Collections. Cambridge (type) Collections. Gundlach no. 1873 M. maniocae Blake 1966b: 216; 1970b: 56 M. testaceum Blatchley 1920: 70 Distribution. Cuba. Endemic M. elachistum Blake 1960: 97. Ha: Havana, Cuba localities. Ci: Cayamas; Soledad (t.l.). shoreline (t.l.). Washington (type). Zayas M: Jovellanos. A pest of manioc M. testaceum elachistum Blake 1970 Collections. Cambridge. Washington (type) Distribution. Cuba, USA (FL) M. moaense Blake 1962: 176; 1970b: 57 Cuba localities. Ci: Cayamas. Ha: Havana, Distribution. Cuba. Endemic Marianao Cuba localities. Ho: Moa (t.l.) Collections. Washington. Zayas Collections. Washington (t.l.) M. xanthacrum Blake 1947b: 311; 1970b: 52 M. multipunctatum Blake 1958: 96; 1970b: 57 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Loma del Gato (t.l.) Cuba localities. Ha: Somorrostro. Ho: Sierra Collections. Cambridge (type) Cristal (t.l.) M. zayasi Blake 1958: 92; 1970b: 53 Collections. Washington (type). Zayas. Distribution. Cuba, USA (FL); on M. obscuricolle Blake 1970b: 58 Conocarpus erecta Distribution. Cuba. Endemic Cuba localities. Ho: Sierra Cristal (t.l.) Cuba localities. Ci: Cayamas (t.l.) Collections. Washington (type) Collections. Washington (type) M. odontotum Blake 1970b: 60 Typophorus Erichson 1847: 163 Distribution. Cuba. Endemic T. habanae Blake 1970a: 119 Cuba localities. Unspecified; taken on pine- Distribution. Cuba. Endemic apple Cuba localities. Ha: Santa Fe (t.l); Alamar. Collections. Washington (type) PR: Soroa M. oteroi Blake 1944: 252; 1970b: 55 Collections. Brno (type). Washington Distribution. Cuba. Endemic [T. nigritus Jacquelin du Val 1857: 299; Gundlach Cuba localities. Ha: Santiago de las Vegas 1891: 365; Medvedev 1993: 6] (t.l.) Distribution. Reported from Cuba as a pest Collections. Washington (type) of sweet potatos (as Tipophorus nigritus; M. rubellum Blake 1970b: 120 Vàzquez et al. 1992: 45), but the record was Distribution. Cuba. Endemic probably based on specimens of T. habanae Cuba localities. M: Arroyo Bermejo (t.l.) (Shawn Clark, in litt.) Collections. Brno (type) M. schwarzi Blake 1948b: 276; 1970b: 52 TRIBE EUMOLPINI Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Alethaxius Lefevre 1885: 42 Collections. Cambridge. Washington (type) A. acunai Blake 1947b: 314 M. simile Blake 1970b: 56 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ho: Moa (t.l.) Collections. Cambridge. Washington (type) A Checklist of the Beetles of Cuba 201

A. bruneri Blake 1949: 367 Distribution. Hispaniola; reported from Distribution. Cuba. Endemic Cuba, but probably in error, see Blake 1967: Cuba localities. SC: Sierra Maestra, Palma 232 Mocha to Pico Joaquin (t.l.) Cuba localities. IJ (Holland and Schwarz Collections. Washington (type) 1917) A. turquinensis Blake 1945: 326 Collections. Pittsburgh. Gundlach no. 232 Distribution. Cuba. Endemic C. suffriani Weise 1914: 122 Cuba localities. SC: Pico Turquino (t.l.) C. nigricornis Suffrian 1867: 325; Gundlach 1891: Collections. Cambridge (type). Washington 360 Distribution. Cuba. Endemic Colaspis Fabricius 1801: 411 Cuba localities. M: Matanzas C. alcyonea Suffrian 1866: 326; Gundlach 1891: Collections. Gundlach no. 1896 36l (C. aleyonea); Blake 1967: 227 C. unicolor Olivier 1808: 885 Distribution. Cuba. EndemicPuerto Rico Distribution. Cuba. Endemic records are probably misidentifications, see Cuba localities. Blake 1967 Collections. Cuba localities. Widespread C. viridula Suffrian 1866: 331; (Chalcophana); Collections. Gundlach no. 301. Washington Gundlach 1891: 362; Blake 1967: 231 C. amplicosta Blake 1970a: 118 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gr: Bayamo. SC: Loma del Cuba localities. Ha: Cojimar; Alamar (t.l.); Gato Cuatro Caminos; Santa Fe; Santiago de las Collections. Gundlach no. 1872. Washington Vegas. PR: Bahia Honda; Sierra de Gazabana Collections. Brno (type). Washington Exochognathus Blake 1946b: 114 C. barberi Blake 1967: 232; Medvedev 1993: 6 E. limbatus Blake 1946b: 116, 1948a: 121 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. CA: Baraguá (on “musail Cuba localities. PR: Sierra Rangel, Las cayes”) (t.l.) Animas (t.l.); Sierra de los Organos; Rancho Collections. Medvedev. Washington (type) Mundito C. brunnea Fabricius 1798: 94; Medvedev 1993: 6 Collections. Washington (type) Distribution. Cuba, Mexico (Baja California), USA Phanaeta Lefevre 1878: 132 Cuba localities. Ha: Lomas de Canoa P. abdominalis (Suffrian) 1866: 328 (Chalcophana); Distribution. Medvedev Gundlach 1891: 361 C. cubensis Blake 1967: 226; Medvedev 1993: 6 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Rangel Cuba localities. Ci: Cayamas (t.l.). Wide- Collections. Gundlach no. 1284 spread. On eggplant and other Solanum P. elongata (Suffrian) 1866: 327 (Chalcophana); Collections. Medvedev. Washington (type) Gundlach 1891: 361 C. fervida (Suffrian) 1866: 330 (Chalcophona); Distribution. Cuba. Endemic Gundlach 1891: 361; Blake 1967: 230; Medvedev Cuba localities. PR: Rangel 1993: 6 Collections. Gundlach no. 1285 Distribution. Cuba. Endemic P. striata (Suffrian) 1866: 336 (Chalcophana); Cuba localities. Ci: Cayamas. Ha: Havana. Gundlach 1891: 362 M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 157. Medvedev Cuba localities. M: Cardenas; Colon C. orientalis Blake 1967: 227 Collections. Gundlach no. 531 Distribution. Cuba. Endemic P. varicornis (Suffrian) 1866: 334 (Chalcophana); Cuba localities. Ci: Soledad (t.l.). Gu: Baracoa Gundlach 1891: 362 Collections. Cambridge (type) Distribution. Cuba. Endemic [C. smaragdula Olivier 1808: 883; Jacquelin du Val Cuba localities. Widespread 1857: 299; Gundlach 1891: 360] Collections. Gundlach no. 539 P. viridula (Suffrian) 1866: 331 (Chalcophana) Distribution. Cuba. Endemic 202 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Myochrous Erichson 1874: 164 Collections. M. cubensis Blake 1947c: 23; 1950: 60 Distribution. Cuba. Endemic TRIBE ADOXINI Cuba localities. Widespread. Ci: Cayamas (t.l.); hosts: a pest of sugarcane, etc. Chalcosicya Blake 1930: 215 Collections. Washington (type). Havana? C. alayoi Blake 1958: 91 Note. M. dubius (Fabricius) 1801: 53 of the Virgin Distribution. Cuba. Endemic Islands is reported by Jacquelin du Val 1857: Cuba localities. SC: Ciudamar (t.l.); Daiquiri 300 and Gundlach 1891: 364 (Gundlach no. 171) Collections. Washington (type). Zayas from Cuba, but it is a doubtful record and prob- C. ditrichota Blake 1938: 46; 1951a: 302 ably applies to the above species (Blake 1947c: Distribution. Cuba. Endemic 24; 1950: 61) Cuba localities SC: Pico Turquino. SS: Trinidad; Buenos Aires (t.l.); San Blas TRIBE CUBISPINI Collections. Cambridge (type) C. insularis (Blatchley) 1922: 32 (Colaspidea); Blake Cubispa Barber 1946: 19 1951a: 297 C. turquino Barber 1946: 22; Staines 2000: 59 C. costata Blake 1938: 49. IJ; Nueva Gerona (t.l.); Distribution. Cuba. Endemic Washington (type) Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba. Endemic Collections. Washington Cuba localities. IJ (t.l.). On citrus Collections. Lafayette (Blatchley colln.; type). SUBFAMILY LAMPROSOMATINAE Cambridge C. maestrensis Blake 1930: 216, 1951a: 290 Oomorphus Curtis 1831: 347 Distribution. Cuba. Endemic O. auricollis Suffrian 1866: 288 (Lamprosoma); Cuba localities. SC: Pico Turquino summit Gundlach 1891: 342 (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. PR: Rangel C. metallica Medvedev 1993: 7, 14 Collections. Gundlach no. 1148 Distribution. Cuba. Endemic Cuba localities. IJ: La Fé (t.l.) SUBFAMILY CRYPTOCEPHALINAE Collections. Medvedev (type) TRIBE CRYPTOCEPHALINI C. nana Suffrian 1866: 337 (Heteraspis); Gundlach 1891: 362; Blake 1951a: 296 Cryptocephalus Muller 1765: XIII (see Zayas Distribution. Cuba. Endemic 1960a for additional localities) Cuba localities. CA: Baraguá. Ci: Cayamas; C. adelphicus Blake 1970a: 117 Soledad. Gr: Bayamo. Ha: San Rafael, San Jose. Distribution. Cuba. Endemic M: Cárdenas. ? Alemon Cuba localities. Gr: Belic (t.l.) Collections. Gundlach no. 536. Cambridge. Collections. Brno (type) Washington C. alcaldei Zayas 1960a: 187 C. parsonsi Blake 1951a: 301 C. alvarezcondei Zayas 1960a: 189 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Mina Carlota (t.l.) Cuba localities. SC: Sierra Maestra; Loma Collections. Cambridge (type) de Calá (t.l.) C. rotunda Blake 1938: 44, 1951a: 296 Collections. Zayas (type female) Distribution. Cuba. Endemic [C. anceps Jacquelin du Val 1857: 293] Cuba localities. SC: Pico Turquino (t.l.) Distribution. South American species; prob- Collections. Cambridge. Washington (type) ably not in Cuba, a misidentification (Shawn C. semicostata Blake 1951a: 300 Clark, in litt.) Distribution. Cuba. Endemic C. anaqui Zayas 1960a: 175 Cuba localities. SC: Loma del Gato (t.l.) Distribution. Cuba. Endemic Collections. Cambridge (type) Cuba localities. Ho: Sierra Cristal (t.l.) Collections. Zayas (type female) A Checklist of the Beetles of Cuba 203

C. azureipennis Suffrian 1852: 83; 1858: 105; 1866: Zayas 1960a: 145; Medvedev 1993: 5 308; Jacquelin du Val 1857: 294; Gundlach Cribouri chlorotique Olivier 1891: 353; Zayas 1960a: 163 Distribution. Cuba, Hispaniola Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Gr: Bayamo Collections. Gundlach no. 1530. Medvedev. Collections. Gundlach no. 1529 Zayas C. behique Zayas 1960a: 166 C. coai Zayas 1960a: 164 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Joaquin. SS: Cuba localities. PR: Sierra de Rangel; Rancho Trinidad; Buenos Aires (t.l.) Mundito (t.l.) Collections. Zayas (type female) Collections. Zayas (type female) C. bicinctus Suffrian 1851: 275; 1858: 354; 1866: C. commutatus Suffrian 1866: 297; Gundlach 1891: 303; Jacqueline du Val 1857: 291; Gundlach 347; Zayas 1960a: 151 1891: 349 (C. bicirnetus); Zayas 1960a: 170 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gr: Bayamo, Buenavista. Ha: Cuba localities. M: Zarabanda. PR: Sierra littoral de La Havana; Pedrera de Mendoza. de Rangel; Viñales PR: Rangel, Viñales Collections. Gundlach no. 974. Zayas Collections. Gundlach no. 1865. Zayas C. blakeae Zayas 1960a: 206 C. complanatus Suffrian 1852: 93; 1858: 385; 1866: Distribution. Cuba. Endemic 312; Jacquelin du Val 1857: 296; Gundlach Cuba localities. SC: Daiquirí (t.l.) 1891: 356; Zayas 1960a: 196 Collections. Zayas (type female) Distribution. Cuba. Endemic C. bruneri Zayas 1952: 73; 1960a: 213 Cuba localities. Gr: Bayamo. LV: Distribution. Cuba. Endemic Cumanayagua. M: Canimar. SC: Gran Piedra Cuba localities. SC: Pico Joaquín (= Collections. Gundlach no. 1209. Zayas Turquino) (t.l.); Cueva de Aura (on Turquino) C. coyugi Zayas 1960a: 186 Collections. Zayas (type) Distribution. Cuba. Endemic C. carib Zayas 1960a: 201 Cuba localities. Ci: Rancho Luna (t.l.); wide- Distribution. Cuba. Endemic spread in Cuba Cuba localities. Ho: Moa, La Breña (t.l.) Collections. Zayas (type female) Collections. Zayas (type male) C. crenulatus Suffrian 1858: 345; 1866: 295; C. catey Zayas 1960a: 173 Gundlach 1891: 345; Zayas 1960a: 154 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Trinidad, Pico de Cuba localities. M: Cienaga de Zapata Potrerillo (t.l.) Collections. Gundlach no. 434 Collections. Zayas (type male) C. cubanensis Jacoby 1907: 842; Zayas 1960a: 215 C. cayofragosae Zayas 1988: 125 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Ciénaga de Zapata, Cuba localities. VC: Cayo Fragoso (t.l.) Zarabanda Collections. Zayas (type) Collections. C. cemi Zayas 1960a: 146 C. cucuba Zayas 1960a: 198 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ho: Moa (t.l.), in pinelands Cuba localities. SC: Loma del Gato (t.l.) Collections. Zayas (type female) Collections. Zayas (type female) C. censorius Suffrian 1858: 383; 1866: 310; C. curtus Suffrian 1851: 268; 1858: 353; 1866: 299; Gundlach 1891: 355; Zayas 1960a: 194; Jacquelin du Val 1857: 290; Gundlach 1891: Medvedev 1993: 5 347; Zayas 1960a: 160 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ho: Sierra Cristal. M: Cuba localities. Widespread Cárdenas. SC: Sierra Maestra; Loma de Calá Collections. Gundlach no. 764. Zayas Collections. Gundlach no. 763. Medvedev. C. cylindricus Suffrian 1858: 385; 1866: 313; Zayas Gundlach 1891: 356; Zayas 1960a: 181 C. chloroticus (Olivier) 1808: 787; Suffrian 1851: Distribution. Cuba. Endemic 247; 1858: 350; 1866: 296; Gundlach 1891: 346; 204 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Cuba localities. Ca: Sierra de Cubitas, Distribution. Cuba. Endemic “Paredones”. Ho: Sierra Cristal. PR: Rangel Cuba localities. Ha: Marianao. M: Fermina. Collections. Gundlach no. 1269. Zayas PR: Sierra de Rangel, Rancho Mundito; Guane, C. distensus Chevrolat 1864a: 181 Paredon de Los Acostas C. quinquepunctatus Suffrian 1852: 91; 1858: Collections. Gundlach no. 1869. Zayas 285; Jacquelin du Val 1856: 123; Gundlach C. jaumei Zayas 1960a: 155 1891: 355; Zayas 1960a: 194 Distribution. Cuba. Endemic C. concolor Zayas 1960a: 195; SC: Juraguá (t.l.); Cuba localities. PR: Herradura (t.l.) Zayas, type Collections. Zayas (type female) C. decoratus Zayas 1960a: 196; PR: Las Animas, C. mabuya Zayas 1960a: 161 Sierra Rangel; Zayas, type Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Mena; Pan de Matanzas Cuba localities. Ho: Sierra Cristal. PR: Si- (t.l.) erra Rangel. SC: Juraguá Collections. Zayas (type male) Collections. Zayas C. maestrensis Blake 1970a: 115 C. dives Suffrian 1852: 82; 1858: 105; 1866: 308; Distribution. Cuba. Endemic Jacquelin du Val 1857: 294; Gundlach 1891: Cuba localities. SC: Cabezal de Palma Mo- 353; Zayas 1960a: 169 cha (t.l.) Distribution. Cuba. Endemic Collections. Brno (type) Cuba localities. PR: Sierra de Rangel, Rancho C. maquey Zayas 1960a: 211 Mundito. Ha: Mariano, littoral de Havana; Distribution. Cuba. Endemic Mesa de Mariel Cuba localities. Ho: Sierra Cristal (t.l) Collections. Gundlach no. 1272. Zayas Collections. Zayas (type female) C. grossulus Suffrian 1851: 248; 1858: 350; 1866: C. maisi Zayas 1988: 126 296; Jacquelin du Val 1857: 289; Gundlach Distribution. Cuba. Endemic 1891: 346; Zayas 1960a: 177 Medvedev 1993: 5 Cuba localities. Gu: Maisi, Tortuguilla (t.l.) C. gundlachi Jacoby 1907: 841 Collections. Zayas (type) Distribution. Cuba. Endemic C. marginicollis Suffrian 1851: 258; Jacquelin du Cuba localities. Ha: Almendares; Havana; Val 1857: 289; Suffrian 1858: 351; Gundlach Marianao. M: Cienaga de Zapata 1891: 346; Zayas 1960a: 346; Blake 1970a: 117; Collections. Gundlach no. 765. Medvedev. Medvedev 1993: 5 Zayas Distribution. Cuba. Endemic C. guanahacabibes Zayas 1960a: 178 Cuba localities. Widespread. Distribution. Cuba. Endemic Collections. Gundlach no. 282. Medvedev. Cuba localities. PR: Peninsula de Washington. Zayas Guanahacabibes (t.l.) C. moa Zayas 1960a: 189 Collections. Zayas (type male) Distribution. Cuba. Endemic C. guanahatabey Zayas 1960a: 200 Cuba localities. Ho: Moa, El Johnson (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type female) Cuba localities. M: Loma El Palenque (t.l.) C. morenoi Zayas 1960a: 184 Collections. Zayas (type female) Distribution. Cuba. Endemic C. guatiaho Zayas 1960a: 203 Cuba localities. Ho: Sierra Cristal (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type female) Cuba localities. Ca Nuevitas, Playa Santa C. nabori Zayas 1960a: 181 Lucía (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type female) Cuba localities. SS: Buenos Aires (t.l.) C. hatuey Zayas 1960a: 148 Collections. Zayas (type female) Distribution. Cuba. Endemic C. pavidus Suffrian 1866: 305; Gundlach 1891: 351; Cuba localities. Gu: Baracoa, La Farola (t.l.), Zayas 1960a: 157 in pinelands Distribution. Cuba. Endemic Collections. Zayas (type female) Cuba localities. Gr: Cabo Cruz C. hypocritus Suffrian 1851: 277; 1858: 354; 1866: Collections. Gundlach no. 1531. Zayas 303; Jacquelin du Val 1857: 291; Gundlach C. pictus Suffrian 1858: 355; 1866: 305; Gundlach 1891: 349; Zayas 1960a: 171 1891: 351; Zayas 1960a: 153 A Checklist of the Beetles of Cuba 205

Distribution. Cuba. Endemic Collections. Gundlach no. 291 (C. saucius); Cuba localities. PR: Rangel no. 1868 (C. ephippium). Zayas Collections. Gundlach no. 1270 C. sericatus Suffrian 1858: 351; 1866: 298; C. platus Blake 1970a: 116 Gundlach 1891: 341; Zayas 1960a: 216 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ha: Guajaybon (t.l.) Cuba localities. M: Cienaga de Zapata Collections. Brno (type) Collections. Gundlach no. 1083 C. poeyi Suffrian 1866: 302; Gundlach 1891: 349; C. signatellus Suffrian 1866: 307; Gundlach 1891: Zayas 1960a: 149 351; Zayas 1960a: 214 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cienfuegos. SC: Alto Cuba localities. Ci: Cayamas. Gr: Bayamo Songo, Florida Blanca (t.l.?) Collections. Gundlach no. 1867. Zayas Collections. Gundlach no. 1527 C. rubetra Suffrian 1858: 381; 1866: 310; Gundlach C. subelatus Tappes 1885: XLI; replacement name 1891: 354; Zayas 1960a: 213 C. elatus Suffrian 1866: 298; Gundlach 1891: 347 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. Ci: Cienfuegos. SC: Daiquirí Collections. Gundlach no. 292. Zayas Collections. Gundlach no. 1866. Zayas C. rubrofasciatus Suffrian 1851: 273; 1858: 353; C. teresae Zayas 1960a: 192 1866: 301; Jacquelin du Val 1857: 290; Distribution. Cuba. Endemic Gundlach 1891: 348; Zayas 1960a: 172 Cuba localities. SS: Bueños Aires; Pico de Distribution. Cuba. Endemic Potrerillo (t.l.) Cuba localities. Ci: Cumanayagua. M: Collections. Zayas (type female) Cárdenas; El Morrillo; Maicas. Gu: Baracoa. C. thalassinus Suffrian 1852: 80; Jacquelin du Val Ho: Moa; La Breña. SC: Cuabitas 1857: 293; Gundlach 1891: 352; Zayas 1960a: Collections. Gundlach no. 1168. Zayas 209 C. rufitarsis Suffrian 1851: 278; 1858: 354; 1866: Distribution. Cuba. Endemic 304; Jacquelin du Val 1857: 292; Gundlach Cuba localities. 1891: 350; Zayas 1960a: 168 Collections. Zayas Distribution. Cuba. Endemic C. tibiellus Suffrian 1858: 354; 1866: 354; Gundlach Cuba localities. Ha: Havana; littoral. Ho: 1891: 350; Zayas 1960a: 164 Moa. M: Cienaga de Zapata; M: Fermina Distribution. Cuba. Endemic Collections. Gundlach no. 1623. Zayas Cuba localities. PR: Sierra de Rangel C. rusticus Suffrian 1851: 270; Jacquelin du Val Collections. Gundlach no. 1263. Zayas 1857: 290; Suffrian 1858: 353; 1866: 300; C. tortuosus Suffrian 1852: 89; 1858: 383; 1866: Gundlach 1891: 348; Zayas 1960a: 348 310; Jacquelin du Val 1857: 296; Gundlach Distribution. Cuba. Endemic 1891: 355; Zayas 1960a: 191; Medvedev 1993: Cuba localities. Gu: Yateras (t.l.?). SC: Loma 5 del Gato C. bardus Chevrolat 1864: 181 Collections. Gundlach no. 1533. Zayas Distribution. Cuba, Puerto Rico C. sao Zayas 1988: 125 Cuba localities. Widespread in Cuba Distribution. Cuba. Endemic Collections. Gundlach no. 148. Medvedev. Cuba localities. PR: Viñales, Sao (t.l.) Zayas Collections. Zayas (type) C. turey Zayas 1960a: 197 C. saucius Suffrian 1852: 84; 1858: 380; 1866: 309; Distribution. Cuba. Endemic Jacquelin du Val 1857: 295; Gundlach 1891: Cuba localities. Ca: Nuevitas, Playa Santa 353 (C. sancius); Zayas 1960a: 183 Lucía (t.l.) C. ephippium Suffrian 1858: 380; Gundlach 1891: Collections. Zayas (type female) 354; Zayas 1960a: 184 C. vinctus Suffrian 1866: 300; Gundlach 1891: 348; Distribution. Cuba. Endemic Zayas 1960a: 158 Cuba localities. Ci: coastal Cienfuegos. M: Distribution. Cuba. Endemic Cárdenas; Maicas. Ho: Sierra de Cristal; Moa, Cuba localities. Gr: Cabo Cruz (t.l.). SC: Piloto; Moa, El Johnson Juraguá Collections. Gundlach no. 1528; Zayas 206 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. vinulus Suffrian 1852: 87; 1858: 381; 1866: 309; Cuba localities. M: Peninsula de Zapata, San Jacquelin du Val 1857: 295; Gundlach 1891: Blas (t.l.) 354; Zayas 1960a: 180 Collections. Zayas (type) Distribution. Cuba. Endemic P. amorae Zayas 1988: 121 Cuba localities. Ha: Havana, coastal, on Distribution. Cuba. Endemic Coccoloba uvifera, etc. Cuba localities. M: Peninsula de Zapata, San Collections. Gundlach no. 320. Zayas Blas (t.l.) C. viridipennis Suffrian 1851: 280; Jacquelin du Collections. Zayas (type) Val 1857: 292; Suffrian 1858: 355; 1866: 304; P. armae Zayas 1988: 122 Gundlach 1891: 350; Zayas 1960a: 159; Distribution. Cuba. Endemic Medvedev 1993: 5 Cuba localities. Ho: Cupeyal (t.l.) Distribution. Cuba, Jamaica, Puerto Rico Collections. Zayas (t.l.) Cuba localities. Widespread, littoral and rud- P. brunneola Suffrian 1866: 322; Gundlach 1891: eral 359 (Pachybrachys) Collections. Gundlach no. 319. Medvedev. Distribution. Cuba. Endemic Zayas Cuba localities. Ci: Cienfuegos C. xerampelinus Suffrian 1852: 45; 1858: 373; 1866: Collections. Gundlach no. 1321 308; Jacquelin du Val 1857: 293; Gundlach P. conglomerata Suffrian 1858: 395; Gundlach 1891: 351; Zayas 1960a: 209 1891: 358 (Pachybrachys). Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Widespread; ruderal, at edges Cuba localities. Widespread of fallow fields in herbs and in flowers of Collections. Gundlach no. 15 Parthenium hysterophorus (L.) P. decipiens Suffrian 1866: 317; Gundlach 1891: Collections. Gundlach no. 689. Zayas 358 (Pachybrachys) Distribution. Cuba. Endemic Diachus LeConte 1880: 196 Cuba localities. Widespread. D. pusio (Suffrian) 1858: 378; 1866: 308; Gundlach Collections. Gundlach no. 917 1891: 352; Zayas 1960a: 216 P. devota Suffrian 1866: 321; Gundlach 1891: 359 Distribution. Cuba. Endemic (Pachybrachys duvotus) Cuba localities. Ha: Somorrostro. M: Pan de Distribution. Cuba. Endemic Matanzas. ? Guamacaro Cuba localities. Gr: Cabo Cruz Collections. Gundlach no. 1213. Zayas Collections. Gundlach no. 1871 D. squalens (Suffrian) 1852: 73; 1858: 377; 1866: P. flavocallens Suffrian 1866: 322; Gundlach 1891: 308; Gundlach 1891: 217; Zayas 1960a: 217 360 (Pachybrachys) Distribution. Cuba, Bahamas, USA Distribution. Cuba. Endemic Cuba localities. Ha: Mariel, littoral. M: Cuba localities. Gu: Baracoa Cárdenas. SS: Trinidad Collections. Gundlach no. 1524 Collections. Gundlach no. 843. Zayas P. gundlachi Suffrian 1858: 398; Gundlach 1891: 359 (Pachybrachys) Mastacanthus Suffrian 1852: 135 Distribution. Cuba. Endemic M. arcustriatus Chevrolat 1864a: 181 Cuba localities. PR: Rangel Distribution. Cuba. Endemic Collections. Gundlach no. 1271 Cuba localities. P. insolitus Zayas 1988: 122 Collections. Distribution. Cuba. Endemic M. insularis Suffrian 1852: 136; Jacquelin du Val Cuba localities. Ha: Marianao (t.l.). Ho: 1857: 297; Gundlach 1891: 356 Cupeyal. M: Punta Hicacos. VC: Sagua la Distribution. Cuba. Endemic Grande Cuba localities. Ci: Cienfuegos Collections. Zayas (type) Collections. Gundlach no. 1143 P. militans Suffrian 1866: 320; Gundlach 1891: 359 Distribution. Cuba. Endemic Pachybrachis Chevrolat 1837: 444 Cuba localities. Gr: sur de Bayamo P. alayoni Zayas 1988: 120 Collections. Gundlach no. 1525 Distribution. Cuba. Endemic P. parallelopipeda Suffrian 1866: 318; Gundlach 1891: 358 (Pachybrachys) A Checklist of the Beetles of Cuba 207

Distribution. Cuba. Endemic Distribution. Cuba. Endemic; not Brazil Cuba localities. Gr: Bayamo Cuba localities. Gu: Yateras. Ha: Arroyo Collections. Gundlach no. 1870 Naranjo. M: Fermina; Pan de Matanzas P. pumicatus Suffrian 1852: 147; Jacquelin du Val Collections. Gundlach no. 868. Washington 1857: 298 Gundlach 1891: 358 (Pachybrachys) C. flavicollis (Lacordaire) 1848: 823 (Chlamys); Distribution. Cuba. Endemic Jacquelin du Val 1857: 288; Gundlach 1891: Cuba localities. Ha: 344 Collections. Gundlach no. 1906 Distribution. Cuba. Endemic P. tortuguilla Zayas 1988: 123 Cuba localities. M: Fermina Distribution. Cuba. Endemic Collections. Gundlach no. 1242 Cuba localities. Type locality not given C. melanospila (Suffrian) 1866: 290 (Chlamys); Collections. Zayas (type) Gundlach 1891: 344 P. tostus Suffrian 1852: 143; Gundlach 1891: 357 Distribution. Cuba. Endemic (Pachybrachys) Medvedev 1993: 6 Cuba localities. M: Cárdenas P. costipennis Suffrian 1858: 393 Collections. Gundlach no. 525 Distribution. Cuba. Endemic C. minimus Zayas 1988: 130 Cuba localities. Gr: Bayamo. M: Cárdenas. Distribution. Cuba. Endemic PR: Rangel. SC: Sierra Maestra Cuba localities. SC: Gran Piedra; Pico Collections. Gundlach no. 151.Medvedev Turquino. Type locality not given P. turquinensis Zayas 1988: 124 Collections. Zayas (type) Distribution. Cuba. Endemic C. nigritella (Suffrian) 1866: 292 (Chlamys); Cuba localities. SC: Pico de Cuba; Pico Gundlach 1891: 345 Turquino (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. M: Cienaga de Zapata. ? Guamacaro Triachus LeConte 1880: 197 Collections. Gundlach no. 973 T. cerinus LeConte 1880: 197 C. pauperrimus Zayas 1988: 131 Distribution. Cuba, USA Distribution. Cuba. Endemic Cuba localities. Gu: Sierra de Nipe, Loma Cuba localities. Ho: Cupeyal (t.l.) de Mensura, 800-950 m, 1967, leg. R. Bielawski Collections. Zayas (type) and A. Riedel (new record from Shawn Clark, C. pictus Zayas 1988: 131 in litt.) Distribution. Cuba. Endemic Collections. Cuba localities. PR: Pan de Guajaibón (t.l.) Collections. Zayas (type) TRIBE CHLAMISINI C. pseudoflavicollis Zayas 1988: 133 Distribution. Cuba. Endemic Chlamisus Rafinesque 1815: 116 Cuba localities. Ho: Moa, Cupeyal. Type lo- C. aeneus Zayas 1988: 128 cality not given Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Ca: Camaguey (t.l.) C. straminea Suffrian 1866: 293 (Chlamys); Collections. Zayas (type) Gundlach 1891: 345 C. barroi Zayas 1988: 128 Distribution. Cuba. Endemic; also reported Distribution. Cuba. Endemic from Puerto Rico, but this is probably a Cuba localities. M: Bahia de Cochinos. SC: misidentification (Shawn Clark, in litt.) Loma del Gato; Pico Turquino (t.l.) Cuba localities. Gr: Cabo Cruz Collections. Zayas (type) Collections. Gundlalch no. 1523 C. cuprosus Zayas 1988: 129 Distribution. Cuba. Endemic REFERENCES Cuba localities. Ca: Camaguëy SC Gran Pie- Balsbaugh, E.U., Jr. and M. Daccordi. 1987. dra; Loma del Gato; Pico Turquino. Type local- Reclassification of some Chrysomelini and a ity not given new species of Plagiodera (Coleoptera: Collections. Zayas (type) Chrysomelidae). J. Kansas Ent. Soc. 60: 30-40. C. conifera (Lacordaire) 1848: 733 (Chlamys); Barber, H.S. 1946. A problematic apterous His- Gundlach 1891: 344, Blake 1944: 249 pid beetle from Cuba (Coleoptra: 208 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Chrysomelidae). Mem. Soc. Cubana Hist. Blake, D.H. 1956. Three new Neotropical flea Natur. 18: 19-22. beetles. J. Washington Acad. Sci. 46: 142-144. Blake, D.H. 1946a. Nine new species of Blake, D.H. 1958. Some new West Indian eumolpid Metachroma (Coleoptera) from the West beetles. Psyche 65: 91-98. Indies.J. Washington Acad. Sci 36: 22-27. Blake, D.H. 1959a. Seven new galerucid beetles Blake, D.H. 1946b. Seven new species of West In- from the West Indies. J. Wash. Acad. Sci. 49: dian Chrysomelidae (Coleoptera). Proc. Ent. Soc. 178-182. Washington 48: 111-119. Blake, D.H. 1959b. Ten new flea-beetles from Cuba. Blake, D.H. 1946c. Species of beetles of the genus Proc. Ent. Soc. Washington 61: 241-248. Lactica from the West Indies closely related to Blake, D.H. 1960. Seven new species of West In- L. tibialis (Olivier). J. Washington Acad. Sci dian Chrysomelidae. Proc. Ent. Soc. Washing- 36: 267-269. ton 62: 97-103. Blake, D.H. 1947a. A new genus of flea beetles Blake, D.H. 1962: Eight new species of from the West Indies. J. Washington Acad. Sci. Metachroma from the West Indies. Proc. Ent. 37: 92-95. Soc. Washington 64: 175-180. Blake, D.H. 1947b. New eumolpid beetles from the Blake, D.H. 1963: Eight new chrysomelid beetles West Indies. J. Washington Acad. Sci. 37: 310- from the West Indies. Proc. Ent. Soc. Wash- 315. ington 65: 14-20. Blake, D.H. 1947c. The species of Myochrous from Blake, D.H. 1964. Notes on new and old species of the West Indies (Coleoptera). Proc. Ent. Soc. Alticinae (Coleoptera) from the West Indies. Washington 49: 22-28. Proc. U.S. Natl. Mus. 115: no. 3477, 9-29. Blake, D.H. 1948a. Six new species of West In- Blake, D.H. 1965. Twelve new species of chry- dian Chrysomelidae (Coleoptera). Proc. Ent. Soc. somelid beetles from the West Indies (Co- Washington 50: 121-127. leoptera, Chrysomelidae). Am. Mus. Novitates Blake, D.H. 1948b. New species of Metachroma no. 2217, 13 pp. and other chrysomelid beetles from the West Blake, D.H. 1966a. More new galerucine beetles Indies. J. Washington Acad. Sci. 38: 274-279. with excised tibiae in the male. Proc. U. S. Blake, D.H. 1949. Notes on some West Indian Natl. Mus. 118: no. 3528, 233-255. Chrysomelidae. J. Washington Acad. Sci. 39: Blake, D.H. 1966b. Ten new chrysomelid beetles 367-371. from the West Indies and Key West. Proc. Ent. Blake, D.H. 1950a. A new genus of flea-beetles Soc. Washington 68: 213-222. from the West Indies. Psyche 57: 10-25. Blake, D.H. 1967. Some new and old species of Blake, D.H. 1950b. A revision of the beetles of the Colaspis in the West Indies. Proc. Ent. Soc. genus Myochrous. Proc. US Nat. Mus. no. Washington 69: 225-237. 3271, 101: 1 64. Blake, D.H. 1970a. Some new chrysomelid beetles Blake, D.H. 1951a. A revision of the beetles of the from Cuba. Casopis Moravskeho Musea (Brno) genus Chalcosicya Blake (Chrysomelidae) from 60: 115-126. the West Indies. Bull. Mus. Comp. Zool. Blake, D.H. 1970b. A review of the beetles of the (Harvard) 106: 287-312. genus Metachroma Chevrolat (Coleoptera: Blake, D.H. 1951b. Synonymies and new species Chrysomelidae). Smiths. Cont. Zool. no. 57. 109 of flea-beetles (Coleoptera, Chrysomelidae). pp. Proc. Ent. Soc. Wash. 53: 138-147. Blake, D.H. 1971. Fifteen new West Indian chry- Blake, D.H. 1952. Six new species of Megistops somelid beetles. Proc. Ent. Soc. Washington 73: with keys to the known species (Coleoptera). 269-282. Psyche 59: 1-12. Blanco, J.G. and C.N. Duckett. 2001. Taxo- Blake, D.H. 1953. The chrysomelid beetles of the nomic revision and cladistic analysis of genus Strabala Chevrolat. Proc. U.S. Natl. Homoschema Blake (Coleoptera: Mus. 103: no. 3319, 121-134. Chrysomelidae: Alticini). Caribb. J. Sci. 37: 1- Blake, D.H. 1954. Five new species of chrysomelid 29. beetles. J. Washington Acad. Sci. 44: 246-250. Borowiec, L. 1989. Three new species of Blake, D.H. 1955. Revision of the vittate species Charidotella Weise (Coleoptera, Chrysomelidae, of the Chrysomelid genus Disonycha from the ), with checklist of the genus. Americas south of the United States. Proc. U.S. Polskie Pismo Ent. 59: 203-222. Natl. Mus. 104 (3338): 1-86. A Checklist of the Beetles of Cuba 209

Chaboo, C.S. 2000. Revision and phylogeny of the Chrysomelidae). Carib. J. Sci. 7: 135-139. Caribbean genus Elytrogona Chevrolat (Co- Savini, V. and Furth, D. 1999. Revisión del leoptera: Chrysomelidae; Cassidinae: Stolaini). Género Gioia Bechyné, 1955 (Coleoptera: Coleop. Bull. 54: 379-394. Chrysomelidae: Alticinae) de las Antillas. Chaboo, C.S. 2002. Range extensions of New Elytron 13: 19-34. World tortoise beetles (Coleoptera: Staines, C.L. 1989. A revision of the genus Chrysomelidae: Cassidinae). Proc. Ent. Soc. Octotoma (Coleoptera, Clhrysomelidae, Washington 104: 716-720. Hispinae). Insecta Mundi 3: 41-56. Daccordi, M. 1995. Su alcume specie di Leucocera Staines, C.L. 1994. A revision of the genus Chevrolat dell’Isola di Cuba (Coleoptera Anisostena Weise (Coleoptera: Chrysomelidae: Chrysomelidae, Chrysomelinae). Boll. Mus. Hispinae). Part II. The subgenus Anisostena: Reg. Sci. Nat. Torino 13: 35-43. key to the species groups and the ariadne spe- Fernández García, I. 1998a. Nueva especie de cies group. Insecta Mundi 8: 125-135. Metachroma Chevrolat (Coleopera: Chrysom- Staines, C.L. 2000. A new species of Cubispa Bar- elidae) de Cuba. Poeyana 455: 1-3. ber, 1946 (Coleoptera, Chrysomelidae) from Fernandez Garcia, I. 1998b. Algunos aspectos Guatemala. Dtsch. Ent. Zeit. 47: 57-60. ecologicos de la comunidad de coleopteros de un Uhmann, E. 1959. Amerikanische Hispinae dus agroecosistema cañero en la provincia de La der Sammlungen Amerikanischer Habana. Poeyana no. 460, 1 14. Entomologen. Pesquisas 3: 611-630. Furth, D.G. 1998. New World Blepharida Vàzquez, L.L., E. Valdés, and J.C. Amor. 1992. Chevrolat 1836 (Coleoptera: Chrysomelidae: New manifestations of pests on economically Alticinae). Mem. Ent. Soc. Washington 21,109 important plants during the period from 1970 pp. to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo Furth, D.G. and V. Savini. 1996. Checklist of Silvestri 49: 41-52. the Alticinae of Central America, including Vogt, G.B., J.U. McGuire, Jr., and A.D. México (Coleoptera; Chrysomelidae). Insecta Cushman. 1979. Probable evolution and mor- Mundi. 10: 45-68. phological variation in South American Furth, D.G. and V. Savini. 1998. Corrections, Dysonychine flea beetles (Coleoptera: clarifications, and additions to the 1996 check- Chrysomelidae) and their amaranthaceous list of the Alticinae of Central America, includ- hosts. US Dept. Agric. Tech. Bull. 1593. 148 ing México (Coleoptera: Chrysomelidae). Insecta pp. Mundi 12: 133-138. White, R.E. 1970. Authorship of the genus Heyer, W., M.-L. Chiang Lok, and B. Cruz. Phyllotreta (Chrysomeloidae: Coleoptera). Proc. 1989. Some aspects of biology and population Ent. Soc. Wash. 72: 388-389. dynamics of Diabrotica balteata LeC. and White, R.E. 19891. Homonymy in world species- Systena basalis du Val (Coleoptera, group names of Criocerinae (Coleoptera: Chrysomelidae), maize and bean pests in Cuba. Chrysomelidae). US Dept. Agric. Tech Bull Acta Phytopathol. Ent. Hung. 24: 93-97. 1629. 69 pp. Heyer, W., M.-L. Chiang Lok, and B. Cruz. White, R.E. 1993. A revision of the subfamily 1993. Zur Fertilität und Mortilität der Criocerinae (Chrysomelidae) of North America Chrysomeliden Diabrotica balteata Lec., north of Mexico. U. S. Dept. Agric. Tech. Bull. Andrector ruficornis (Oliv.) und Systena basa- 1805. 158 pp. lis Duval. Beitr. Entomol (Berlin) 43: 379-385. White, R.E. 1996. A revision of the genus Medvedev, L.N. 1993. On the fauna and zoogeog- Chaetocnema of America north of México (Co- raphy of leaf-beeetles of the West Indies. 4-16. leoptera; Chrysomelidae). Cont. Amer. Ent. In: Sokolov, V.E. (ed.). Investigations of the Soc. 29:1-125. Cuban fauna [Issledovaniya fauny Kuby]. Wilcox, J.A. 1983. Chrysomelidae. In: R.H. Arnett Nauka, Moscow. [in Russian]. Jr., (ed.). Checklist of the beetles of North and Riley, E.G. 1986: Review of the tortoise beetle gen- Central America and the West Indies. Flora era of the tribe Cassidini occurring in America and Fauna Pubs., Gainesville, FL. Vol. 8. 166 north of México (Coleoptera: Chrysomelidae: pages. Cassidinae). J. New York Ent. Soc. 94: 98-114. Zayas, F. de. 1939. Dos nuevas especies de Sanderson, M.W. 1967. New West Indian Chrysomelidae de Cuba (Cassidinae). Mem. Soc. Hispinae, with notes and keys (Coleoptera: Cubana Hist. Nat. 13: 253-256. 210 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Zayas, F. de. 1952: Nuevos crisomelidos de las Valentine 1999: 255. subfamilias Cassidinae y Clytrinae. Mem. Soc. N. pallipes (Suffrian) 1870: 170 (Notioxenus); Cubana Hist. Nat. 21: 71-75. Gundlach 1891: 268; Valentine 1999: 255; 2004: Zayas, F. de. 1960a. Contribucion al estudio de 53 los Cryptocephalus Cubanos (Coleoptera: Distribution. Cuba. Endemic Chrysomelidae) y adicion de 25 nuevas especies Cuba localities. M: Cienaga de Zapata y formas a la fauna. Mem. Soc. Cub. Hist. Natu. Collections. Gundlach no. 690 24: 135-217 and 14 plates. Zayas, F. de. 1960b. Dos nuevos especies de TRIBE CISANTHRIBINI Chrysomelidae Col. de Cuba. Mem. Soc. Cubana Hist. Nat. 24: 129-133. Sicanthus Valentine 1999: 258 (S. rhizophorae Valentine 1999: 259) SUPERFAMILY CURCULIONOIDEA Distribution. USA: Florida Keys only Reviewed by R. S. Anderson, February 2002 Cuba localities. 160. FAMILY NEMONYCHIDAE Collections. This family occurs in temperate parts of North and South America but not the West Indies. The SUBFAMILY larvae usually feed in the male cones of conifers. TRIBE DECATAPHANINI

161. FAMILY ANTHRIBIDAE Ptychoderes Schönherr 1823: 1135 The fungus weevils P. cristalensis Zayas 1988: 150; Valentine 2004: 55 Larvae and adults of this family usually occur Distribution. Cuba. Endemic on or in rotten wood and fungal fruiting bodies. Cuba localities. Gu: Yateras. Ho: Sierra Members of the are often seed feeders. Cristal (type locality designasted in Valentine Valentine (1999) is a summary of the Nearctic fauna 2004: 55) and includes data on some Cuban species. Valen- Collections. Zayas (type) tine (2004) indicates that there are an additional 22 undescribed species and an additional three gen- TRIBE ISCHNOCERINI era for Cuba, which are not included below. Ischnocerus Schonherr 1839: 191 SUBFAMILY CHORAGINAE (I. infuscatus Fahraeus 1939: 192; Valentine 2004: TRIBE ARAECERINI 56) Distribution. Bahamas (Andros, Abaco, Acaromimus Jordan 1907: 381 Eleuthera), USA (MD to TX), Mexico to Guate- (A. americanus (Motschulsky) 1873: 251 mala (Xenorchestes)); Valentine 2004: 54 Cuba localities. Distribution. USA: coastal TX, AL, FL Collections. (Florida Keys including Dry Tortugas), Baha- mas (Andros, New Providence) TRIBE GYMNOGNATHINI Cuba localities. Collections. Gymnognathus Schönherr 1826: 37 G. caniqui Zayas 1988: 151; Valentine 2004: 56 (in- Araecerus Schönherr 1823: 1135 dicating that the species is in an undescribed A. fasciculatus DeGeer 1775: 276 (Curculio); genus) Jacquelin du Val 1857: 17; Valentine 2004: 52. Distribution. Cuba. Endemic The coffee bean weevil Cuba localities. M: Cienaga de Zapata (t.l.) A. coffeae Fabricius 1801: 411; Valentine 1999: Collections. Zayas (type) 252 Distribution. Cuba (introduced), Hispaniola, TRIBE RHINOTROPIDINI Jamaica, Genada, Bermuda, México to South America; cosmopolitan. A generalist feeder on Homocloeus Jordan 1904: 264 many dried and living plant tissues H. acerbus (Boheman) 1833: 149 (Tropideres); Cuba localities. Widespread Jacquelin du Val 1857: 170; Gundlach 1891: Collections. 269; Valentine 2004: 56 A Checklist of the Beetles of Cuba 211

Distribution. Cuba. Endemic H. sexverrucatus Suffrian 1870: 190 (Tropideres); Cuba localities. Widespread Gundlach 1891: 272; Valentine 2004: 57 Collections. Gundlach no. 176. Stockholm Distribution. Cuba, USA (FL) (type) Cuba localities. Gr: Bayamo. M: Matanzas H. balteatus (Gyllenhal) 1833: 154 (Tropideres); Collections. Gundlach no. 1158 Jacquelin du Val 1857: 170; Gundlach 1891: H. sordidulus (Suffrian) 1870: 193 (Tropideres); 273; Valentine 2004: 56 Gundlach 1891: 272; Valentine 2004: 57 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas Collections. Collections. Gundlach no. 754 H. caliginosus (Boheman) 1833: 149 (Tropideres); Jacquelin du Val 1857: 169; Gundlach 1891: TRIBE PIESOCORYNINI 27; Valentine 2004: 56 Tropideres modesta Suffrian 1870: 182; Gundlach Brachycorynus Valentine 1999: 271 1891: 270; Valentine 2004: 56 (synonomy) B. distentus (Frieser) 1983: 53 (Homocloeus); Val- Distribution. Cuba. Endemic entine 1999: 272; 2004: 58 Cuba localities. Distribution. Cuba, USA (s FL and Keys) Collections. Gundlach no. 1862 Cuba localities. Ha: Playa Marianao (t.l.) H. confusus (Suffrian) 1870: 198 (Tropideres); Collections. Frieser colln, (type to be depos- Gundlach 1891: 274; Valentine 2004: 56 ited in ZSMC: Zoologische Staatssamlung, Distribution. Cuba. Endemic Munich, Germany) Cuba localities. M: Cardenas; Fermina Collections. Gundlach no. 1159. Halle Phaenotheriopsis Wolfrum 1931: 70 (cotypes) P. bierigi Wolfrum 1931: 70; Valentine 2004: 58 H. dermestoides (Suffrian) 1870: 176 (Eugonus); Distribution. Cuba endemic. Gundlach 1891: 269; Valentine 2004: 56 Cuba localities. Ha: Tapaste (t.l.) Distribution. Cuba. Endemic Collections. Basel (type) Cuba localities. SC Collections. Gundlach no. 1861 Piesocorynus Dejean 1834: 235 H. feralis (Boheman) 1833: 148 (Tropideres); P. suffriani Valentine 2004: 58 (replacement name Jacquelin du Val 1857: 168; Gundlach 1891: for P. gracilicornis) 269; Valentine 2004: 56 P. gracilicornis Suffrian 1870: 186 (Tropideres); Distribution. Cuba. Endemic Gundlach 1891: 271 Cuba localities. Widespread Distribution. Cuba, Bahamas (Andros) Collections. Gundlach no. 674. Halle (type) Cuba localities. M: Cárdenas H. laetus (Suffrian) 1870: 195 (Tropideres); Collections. Gundlach no. 483 Gundlach 1891: 273; Valentine 2004: 56 Distribution. Cuba. Endemic TRIBE PLATYRHINI Cuba localities. Gr: Bayamo Collections. Gundlach no. 1370 Eucloeus Wolfram 1930: 26 H. luscus (Fahraeus) 1839: 209 (Tropideres); E. angulatus (Suffrian) 1870: 199 (Ptychoderes); Jacquelin du Val 1857: 169; Gundlach 1891: Gundlach 1891: 274; Valentine 2004: 59 270 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Yateras. SS: Trinidad Cuba localities. M: Cárdenas. Stockhlom Collections. Gundlach no. 1174. Halle (type) (type) Collections. Gundlach no. 13 TRIBE PLATYSTOMINI H. obsoletus (Suffrian) 1870: 184 (Tropideres); Gundlach 1891: 271; Valentine 2004: 57 Phoenicobiella Cockerell 1906: 243 Distribution. Cuba. Endemic P. trituberculatus (Suffrian) 1870: 172 (); Cuba localities. M: Cárdenas Gundlach 1891: 268; Valentine 2004: 60 Collections. Gundlach no. 172. Stockholm Distribution. Cuba. Endemic (type) Cuba localities. Ci: Cienfuegos. Notes. P. chamaeropis LeConte (1876: 401) is distributed 212 Arthropods of Florida and Neighboring Land Areas, Vol. 18

in USA (SC-LA) and Bahamas (South Bimini, Collections. Eleuthera, Crooked) (Valentine 2004: 60) O. fuscipennis (Suffrian) 1870: 192 Tropideres); Collections. Gundlach no. 811 Gundlach 1891: 272; Valentine 2004: 61 Distribution. Cuba. Endemic Toxonotus Lacordaire 1866: 575 Cuba localities. M: Cienaga de Zapata T. fascicularis (Schoenherr 1833: 132 (); Collections. Gundlach no. 997 Jacquelin du Val 1857: 171; Gundlach 1891: O. griseus (Suffrian) 1870: 195 (Tropideres); 268; Valentine 1999: 277; 2004: 60 Gundlach 1891: 273; Valentine 2004: 61 Distribution. Cuba, Bahamas (widespread), Distribution. Cuba. Endemic USA (south Florida) Cuba localities. M: Cienaga de Zapata Cuba localities. Widespread Collections. Gundlach no. 996 Collections. Gundlach no. 96. Stockholm O. parvulus (Suffrian) 1870: 193 (Tropideres); (type) Gundlach 1891: 272; Valentine 2004: 62 T. mendax (Wolfrum) 1930: 31 (Neanthribus); Val- Tropideres blanchardi Blackwelder 1947: 768 (re- entine 2004: 60 placement name) Distribution. Cuba, Bahamas (South Bimini, Distribution. Cuba. Endemic Cat, Abaco, Acklins, Crooked, Great Inagua) Cuba localities. M: Cárdenas; Cienaga de Cuba localities. Zapata Collections. Collections. Gundlach no. 658

TRIBE TRIGONORHININI REFERENCES Wollaston 1861: 102 Valentine, B. 1999 [1998]: A review of Nearctic T. rotundatus (LeConte) 1876: 407; Valentine 2004: and some related Anthribidae (Coleoptera). In- 61 secta Mundi 12: 251-296. Distribution. Cuba?, Mexico (northern), USA Valentine, B. 2004. A catalogue of West Indies (ME-IL-FL-TX) Anthribidae (Coleoptera). Insecta Mundi 17: 49- Cuba localities. PR: Sierra del Rosario, San 67. Vicente Collections. Champaign 162. FAMILY BELIDAE The cycad weevils TRIBE Members of this family breed in cones of cycads and Aurucaria, in palm fruits, and in the flowers LeConte 1876: 399 of Hydnoraceae root parasites. Rhopalotria slossoni E. schwarzi Pierce 1930: 28; Valentine 1999: 285; of Florida breeds in the cones of Zamia cycads. 2004: 61 Distribution. Cuba, USA (s FL and Keys) SUBFAMILY ALLOCORYNINAE Cuba localities. Collections. Rhopalotria Chevrolat 1878: XCVII R. dimidiata Chevrolat 1878: XCVII; Muñiz and Waterhouse 1845: 37 Barrera 1969: 205 O. angulatus (Suffrian) 1870: 187 (Tropideres); Distribution. Cuba. Endemic Gundlach 1891: 271; Valentine 2004: 61 Cuba localities. M: Varadero. A pollinator of T. variolosa Suffrian 1970: 188; Gundlach 1891: Zamia cycads and probably breeding in Zamia 271; Valentine 2004: 61 (synonomy); Gr: cones Bayamo. M: Cienaga de Zapata. Gundlach no. Collections. 995 Distribution. Cuba, Hispaniola, Jamaica REFERENCES Cuba localities. Gr: Bayamo. Ma: Cárdenas Muñiz, R. and A. Barrera. 1969. Rhopalotria Collections. Gundlach no. 173. Halle (type) dimidiata Chevrolat, 1878: estudio morfológico O. floridanus (Leng) 1916: 28 (Toxotropis); Valen- del adulto y descripción de la larva (Ins. Col. tine 2004: 61 Curcul.: Oxycoryninae). Rev. Soc. Mexicana Distribution. Cuba, USA (FL and Keys) Hist. Natur. 30: 205-222. Cuba localities. A Checklist of the Beetles of Cuba 213

163. FAMILY ATTELABIDAE Collections. Gundlach no. 1156 The leaf rolling weevils, tooth-nose snout beetles, E. sexmaculatus (Chevrolat) 1876: CCXXVII and thief weevils. Distribution. Cuba, Puerto Rico O’Brien and Wibmer (1982) is the primary Cuba localities. source for this list. Collections.

SUBFAMILY ATTELABINAE SUBFAMILY RHYNCHITINAE TRIBE RHYNCHITINI Emphyleuscelus Voss 1925: 69 E. cubensis Janczyk 1960: 36 Rhynchites Schneider 1791: 83 Distribution. Cuba. Endemic R. grandis Zayas 1988: 191 Cuba localities. Distribution. Cuba. Endemic Collections. Vienna (type) Cuba localities. SC: Gran Piedra (t.l.) Collections. Zayas (type) Euscelus Schönherr 1833: 205 R. trifasciatus Suffrian 1870: 229; Gundlach 1891: E. angulosus (Gyllenhal) 1833: 208 (Attelabus); 282 Jacquelin du Val 1857: 172; Gundlach 1891: Distribution. Cuba. Endemic 280 Cuba localities. M: Cienaga de Zapata Distribution. Cuba, Hispaniola Collections. Gundlach no. 1014 Cuba localities. Widespread Collections. Gundlach no. 116 TRIBE AULETINI E. armatus (Gyllenhal) 1833: 208 (Attelabus); Jacquelin du Val 1857: 172; Gundlach 1891: Auletobius Desbrochers 1869: 396 281 A. cubanus Voss 1922: 63 Distribution. Cuba, Hispaniola Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Collections. Pittsburgh Collections. Berlin (type) E. aureolus (Gyllenhal) 1833: 209 (Attelabus); Jacquelin du Val 1857: 173; Gundlach 1891: REFERENCES 281 O’Brien, C.W. and G.J. Wibmer. 1982. Anno- Distribution. Cuba. Endemic tated checklist of the weevils (Curculinidae Cuba localities. Widespread sensu lato) of North America, Central America, Collections. Gundlach no. 227 and the West Indies (Coleoptera: E. callosipennis Voss 1925: 44 Curculionoidea). Mem. Amer. Ent. Inst. 34. Distribution. Cuba, Hispaniola Cuba localities. 164. FAMILY BRENTIDAE Collections. Berlin The straight-snouted weeevils and the pear E. cancellatus Voss 1925: 43 shaped weevils Distribution. Cuba. Endemic Kissinger (1974) and O’Brien and Wibmer Cuba localities. (1982) are the primary sources for this list. Collections. Berlin (type) E. cribrarius (Olivier) 1807: 8 SUBFAMILY BRENTINAE Distribution. Cuba, Hispaniola TRIBE BRENTINI Cuba localities. Collections. Brentus Fabricius 1787: 95 E. pulchellus (Suffrian) 1870: 224; Gundlach 1891: B. anchorago (L.) 1758: 383; Jacquelin du Val 1857: 281 175 Distribution. Cuba. Endemic Belopherus militaris Olivier, Jacquelin du Val Cuba localities. Widespread 1857: 174; Gundlach 1891: 276 Collections. Gundlach no. 158 Belopherus strigicollis Lacordaire, Gundlach E. scutellatus (Klug) 1829: 12 (Attelabus); Jacquelin 1891: 276 (Belophorus) du Val 1857: 171; Gundlach 1891: 280 Brenthus turbatus Olivier, Jacquelin du Val Distribution. Cuba, Hispaniola 1857: 176; Gundlach 1891: 275 Cuba localities. Widespread 214 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Brenthus simplicicollis Chevrolat, Gundlach P. uncimanus (Boheman) 1840: 496 (Trachelizus); 1891: 277 Jacquelin du Val 1857: 174; Gundlach 1891: Brenthus vulneratus Schoenherr, Jacquelin du 278 Val 1857: 177 Distribution. Cuba, USA (FL); on Solanum Brenthus monilis Fabricius, Jacquelin du Val fruits, in hammock forests 1857: 178 Cuba localities. Gr: Bayamo Distribution. Cuba, widespread West Indies, Collections. Gundlach no. 1351 México, Central America, South America, USA (FL); under bark SUBFAMILY CYLADINAE Cuba localities. Widespread Collections. Gundlach no. 1350 (B. Cylas Latreille 1802: 196 simplicicollis); no. 121 (B. strigicollis) C. formicarius (Fabricius) 1798: 174. The sweetpotato weevil TRIBE ARRHENODINI Distribution. Cuba (introduced), widespread West Indies, Central and South America, USA Estenorhinus Lacordaire 1866: 431 (NM-FL); coastal in Ipomoea pes-caprae. This E. forcipitigerus Gyllenhal (L. & M.) 1814: 477 is a serious agricultural pest species Arrhenodes forcipitigerus Schoenherr, Jacquelin Cuba localities. du Val 1857: 173?; Gundlach 1891: 277 Collections. Gundlach no. 1863 (Estenorhinus); 283 Distribution. Cuba. Endemic SUBFAMILY ULOCERINAE Cuba localities. Widespread Collections. Gundlach no. 122 Ulocerus Schoenherr 1823: c.1137 U. bicaudatus Suffrian 1870: 230 SUBFAMILY CYPHAGOGINAE Distribution. Cuba. Endemic TRIBE STEREODERMINI Cuba localities. M: Cienaga de Zapata Collections. Gundlach no. 999 Stereodermus Lacordiare 1866: 419 S. exilis Suffrian 1870: 220; Gundlach 1891: 279 SUBFAMILY APIONINAE (Stercodermus) The Apioninae mostly feed on leaves as adults Distribution. Cuba, USA (FL); hammock for- and the larvae bore into seeds, stems, and other ests plant parts. Cuba localities. M: Cárdenas Collections. Gundlach no. 327 TRIBE APIONINI

SUBFAMILY TRACHELIZINAE Apion Herbst 1797: 100. The generic placement TRIBE TRACHELIZINI of these species may need to be reviewed A. agraticum Kissinger 1974: 28 Paritrachelizus Kleine 1921: 32 Distribution. Cuba. Endemic P. linearis (Suffrian) 1870: 218 (Trachelizus); Cuba localities. Ho: Moa (t.l.) Gundlach 1891: 279 Collections. Chanmpaign (type) Distribution. Cuba, Puerto Rico A. analomum Kissinger 1974: 6 Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 523 Cuba localities. Ci: Cayamas. PR: Bahia P. simplex (Suffrian) 1870: 218 (Trachelizus); Honda. SC: Gran Piedra (t.l.); Loma del Gato Gundlach 1891: 279 Collections. Champaign (type). Washington Distribution. Cuba. Endemic A. dilutum Kissinger 1974: 17 Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 294 Cuba localities. Ci: Cayamas (t.l.) P. tenuis (Suffrian) 1870: 216 (Trachelizus) Collections. Washington (type) Distribution. Cuba. Endemic (A. gulare Fall 1898: 140; Kissinger 1974: 8) Cuba localities. Widespread Distribution. Jamaica, México, USA (FL); Collections. Gundlach no. 123 larvae develop in seeds of Jamaican dogwood Cuba localities. A Checklist of the Beetles of Cuba 215

Collections. sensu lato) of North America, Central America, A. latrum Kissinger 1974: 8 and the West Indies (Coleoptera: Distribution. Cuba. Endemic Curculionoidea). Mem. Amer. Ent. Inst. 34. Cuba localities. Ci: Cayamas. PR: Aspiro- Rangel (t.l.); Bahia Honda. ?Corralillo 165 FAMILY Collections. Champaign (type). Washington The four genera of this family occur in south- A. maculaalba Suffrian 1870: 232; Gundlach 1891: ern South America. 283; Kissisnger 1974: 14 Distribution. Cuba. Endemic 166. FAMILY ITHYCERIDAE Cuba localities. Ci: Cayamas; Soledad. Gu: This family contains only one species in north- Imias. M: Cárdenas. SC: Gran Piedra eastern North America, and it is associated with Collections. Gundlach no. 1219. Washington oaks. A. naulum Kissinger 1974: 10 Distribution. Cuba. Endemic 167. FAMILY CURCULIONIDAE Cuba localities. Ho: Moa (t.l.) The snout beetles or weevils Collections. Champaign (type). Washington Reviewed by R.S. Anderson July 1994 A. nigrosparsum Suffrian 1870: 230; Gundlach This is the largest of all beetle families. Habits 1891: 282 are variable, but most species feed as larvae in rot- Distribution. Cuba. Endemic ting wood and cambium, or the roots, stems, flow- Cuba localities. Ci: Cayamas. Gr: Bayamo ers, and seeds of living plants. Many larvae occur Collections. Gundlach no. 1390. Washington in the soil and are root feeders. Some species are A. satagium Kissinger 1974: 16 economic pests of grain, rice, cotton, etc. O’Brien Distribution. Cuba. Endemic and Wibmer (1982) is the primary source for this Cuba localities. Ci: Cayamas (t.l.) list. Records for species presence in Cuba which Collections. Washington (type) are listed without supporting data are from O’Brien A. vetitum Kissinger 1974: 13 and Wibmer (1982). The bark and ambrosia beetles Distribution. Cuba. Endemic (Scolytinae and Platypodinae) have traditionally Cuba localities. PR: Bahia Honda (t.l.) been ranked as families. An alternative higher clas- Collections. Washington (type) sification of this group is that of of Alonso-Zarazaga and Lyal (1999) which uses additional family units, TRIBE PIEZOTRACHELINI and in which Scolytidae and Platypodidae are used as families. Chrysapion Kissinger 1968: 23 C. tantillum (Sharp) 1890: 50; Kissinger 1974: 4 SUBFAMILY DRYOPHTHORINAE Distribution. Cuba, México, Guatemala, TRIBE DRYOPHTHORINI USA (TX) Cuba localities. CA: Baraguá Stenommatus Wollaston 1873: 442 Collections. Washington S. parasulcifrons Osella 1981: 181 Distribution. Cuba. Endemic SUBFAMILY NANOPHYINAE Cuba localities. Ho: Quemado de Pineda (t.l.) TRIBE NANOPHYINI Collections. Verona (type) S. musae Marshall 1920: 271; Osella 1981: 183 Nanophyes Schönherr 1838: 780 Distribution. Cuba (introduced); native to N. dispersenotatus (Pic) 1916: 14 (Nanodes) Pacific Islands Distribution. Cuba. Endemic Cuba localities. PR: Guanahacabibes Cuba localities. Collections. Zayas or Verona Collections. TRIBE ORTHOGNATHINI REFERENCES Kissinger, D. A. 1974. Apioninae occuring in the Mesocordylus Lacordaire 1866: 314 West Indies (Curculionidae, Coleoptera). Taxo- M. cubensis Vaurie 1970: 26 nomic Publication, Riverside, CA. 57 pp. Distribution. Cuba. Endemic O’Brien, C.W. and G.J. Wibmer. 1982. Anno- Cuba localities. PR: Viñales (t.l.) tated checklist of the weevils (Curculinidae Collections. New York (type) 216 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Rhinostomus Rafinesque 1815: 115 SUBTRIBE SPHENOPHORINA R. oblitus (Jacquelin du Val) 1857: 229 (Rhina); Gundlach 1891: 333 Cosmopolites Chevrolat 1885: 289 Distribution. Cuba, Hispaniola C. sordidus (Germar) 1824: 299 (Calandra) Cuba localities. IJ Distribution. Cuba, widespread West Indies, Collections. Gundlach no. 478 México to South America, USA (FL) Cuba localities. TRIBE RHYNCHOPHORINI Collections. SUBTRIBE RHYNCOPHORINA Metamasius Horn 1873: 410 Rhyncophorus Herbst 1795: Pls LX-LXI M. bruneri Buchanan 1941: 169 (R. cruentatus (Fabricius) 1775: 128 (Curculio)) Distribution. Cuba, Jamaica? Distribution. West Indies, South America, Cuba localities. SC: Loma del Gato (t.l.) (hid- USA (TX-SC-FL) ing in “Manaca” palm) Cuba localities. Collections. Washington (type) Collections. M. hemipterus (Linnaeus) 1758: 377 (Curculio) R. palmarum (Linnaeus) 1758: 377 (Curculio) M. h. sericeus (Olivier) 1807: 84; Jacquelin du Distribution. Cuba, West Indies, México to Val 1857: 231; Gundlach 1891: 334 South America, USA (CA, TX) (Sphenophorus sericeus). Endemic subspecies of Cuba localities. Cuba Collections. Distribution. Cuba, Jamaica, Hispaniola, Nicaragua to Panamá, USA (FL) SUBTRIBE LITOSOMINA Cuba localities. Widespread. A pest of plan- tain and bananas; boring into their trunks Sitophilus Schönherr 1838: 967 Collections. S. granarius (Linnaeus) 1758: 378 (Curculio). The M. mosieri Barber 1920: 151 granary weevil Distribution. Cuba, Hispaniola, USA (FL) Distribution. Cuba (introduced); cosmopoli- Cuba localities. Ci: Cayamas (t.l.) tan; a stored products pest Collections. Washington (type) Cuba localities. M. ritchiei Marshall 1916: 197 Collections. Distribution. Cuba, Jamaica S. linearis (Herbst) 1797: 5 (Rhyncophorus); Cuba localities. Attacks pineapples Gundlach 1891: 335 Collections. Distribution. Cuba, Dominica,Guadeloupe, Jamaica, Puerto Rico, St. Barthélemy, Costa Scyphophorus Schönherr 1838: 855 Rica, South America, Old World, USA (FL, LA). S. acupunctatus Gyllenhal 1838: 857 A stored products pest, distributed by commerce S. anthracinus Gyllenhal 1838: 858; Jacquelin Cuba localities. Gr: Bayamo (in tamarind du Val 1857: 231; Gundlach 1891: 334 pods). VC: Las Pozas Distribution. Cuba, Jamaica, Hispaniola, Collections. Gundlach no. 1063 México to South America, USA (CA-KS-FL) S. oryzae (Linnaeus) 1763: 395 (Curculio); Cuba localities. Gu: Guantánamo; M: Jacquelin du Val 1857: 234; Gundlach 1891: Guamutas 336 (S. orizae). The rice weevil Collections. Gundlach no. 552 Distribution. Cuba (introduced); cosmopoli- tan; a stored products pest Sphenophorus Schönherr 1838: 874 Cuba localities. Widespread S. australis Chittenden 1905: 53 Collections. Gundlach no. 231 Distribution. Cuba, Puerto Rico, México, S. zeamais Motschulsky 1855: 77. The maize wee- widespread USA vil Cuba localities. Distribution. Cuba (introduced); cosmopoli- Collections. tan; a stored products pest S. cubensis (Buchanan) 1936: 150 (Calendra) Cuba localities. Distribution. Cuba, Jamaica, USA (FL) Collections. Cuba localities. Ci: Soledad. Ha: Central Amistad; Guanajay; Havana (t.l.); San Anto- A Checklist of the Beetles of Cuba 217

nio; Santiago de Las Vegas. PR: San Juan. ? Anthonomus Germar 1817: 340 Martinez A. alboannulatus Boheman 1843: 218; Gundlach Collections. Washington (type) 1891: 286 S. venatus (Say) 1831: 22 (Rhynchophorus) Distribution. Cuba, Guadeloupe S. placidus (Say) 1831: 23 (Rhyncophorus); Cuba localities. Gr: Bayamo Gundlach 1891: 335 Collections. Gundlach no. 1407 Distribution. Cuba, widespread USA A. albocapitis Clark 1991a: 267 Cuba localities. M: Cárdenas Distribution. Cuba, Hispaniola, Puerto Rico, Collections. Gundlach no. 399 S. vestitus Chittenden 1904: 134 Cuba localities. Ca: near California. Ci: Distribution. Cuba, Bahamas, Hispaniola, Cayamas Puerto Rico, México, Martinique, se USA Collections. Cuba localities. A. costulatus Suffrian 1871: 137; Gundlach 1891: Collections. 287 Distribution. Cuba, Puerto Rico, USA (FL, SUBFAMILY ERIRRHININAE TX) TRIBE ERIRHININI Cuba localities. M: Cárdenas; Cienaga de SUBTRIBE STENOPELMINA Zapata Collections. Gundlach no. 1017 Helodytes Kuschel 1952: 56 A. fastosus Clark 1991b: 51 H. minox Kuschel 1952: 61 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Guantánamo Naval Base Cuba localities. (t.l.) Collections. Collections. O’Brien (type) A. grandis Boheman 1843: 232; Gundlach 1891: Lissorhoptrus LeConte 1876: 183 285 L. brevirostris (Suffrian) 1871: 157 (Hydronomus); Distribution. Cuba, Hispaniola, México to Gundlach 1891: 293; Kuschel 1952: 50 South America, USA (TX-TN-FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas; Real Campiña. Collections. Gundlach no. 215 A pest of rice plants (bionomics: Carbonnel 1982; A. irroratus Dietz 1891: 198 Meneses 1981) Distribution. Cuba, Puerto Rico, México, Collections. Gundlach no. 444. Washington USA (FL) L. lacustris Kuschel 1952: 38 Cuba localities. Distribution. Cuba, USA (FL, GA) Collections. Cuba localities. A. julichi Dietz 1891: 198 Collections. Distribution. Cuba, USA (FL) L. oryzophilus Kuschel 1952: 44 Cuba localities. Distribution. Cuba, México, USA (FL-TX- Collections. CT) A. leucocephale Clark 1991a: 270 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ci: Cayamas (t.l.) Collections. Washington (type) Neohydronomus Hustache 1926: 229 A. luteus Suffrian 1871: 131; Gundlach 1891: 286 N. elegans O’Brien and Wibmer 1989: 301 Distribution. Cuba. Endemic Distribution. Cuba, México to Panamá; feeds Cuba localities. on water lettuce (Pistia ) Collections. Gundlach no. 1644 Cuba localities. Ci: Cumanayagua A. modicellus Gyllenhal 1836: 355; Gundlach 1891: Collections. 289 Distribution. Cuba, Guadeloupe, Martinique, SUBFAMILY CURCULIONINAE St. Barthélemy TRIBE ANTHONOMINI Cuba localities. PR: Rangel Collections. Gundlach no. 1385 218 Arthropods of Florida and Neighboring Land Areas, Vol. 18

A. morbillosus Suffrian 1871: 146; Gundlach 1891: Botanebius Schönherr [1835]: 358 285 B. tuberculatus Gyllenhal 1836: 359; Jacquelin du Distribution. Cuba. Endemic Val 1857: 201; Gundlach 1891: 284 Cuba localities. Gr: Bayamo Distribution. Cuba. Endemic Collections. Gundlach no. 1406 Cuba localities. Widespread A. pulchellus Suffrian 1871: 138; Gundlach 1891: Collections. Gundlach no. 5260 287 Distribution. Cuba. Endemic Huaca Clark 1993a: 2 Cuba localities. H. collana Clark 1993a: 21 Collections. Gundlach no. 1312 Distribution. Cuba; Jamaica A. punctipennis Gyllenhal 1836: 334; Jacquelin du Cuba localities. Ha: Havana area? Making Val 1857: 199; Gundlach 1891: 286 flower galls on Triopteris jamaicensis A. aristus Suffrian 1871: 134 Collections. Washington A. divisus Suffrian 1871: 134 Distribution. Cuba. Endemic Loncophorus Chevrolat 1832: 215 Cuba localities. M: Cárdenas; Cienaga de L. daviesii (Swederus) 1787: 194 (Curculio) Zapata Distribution. Cuba. Endemic Collections. Gundlach no. 383 Cuba localities. A. rhamphoides Suffrian 1871: 142; Gundlach 1891: Collections. 288 L. humeralis Chevrolat 1832: 442 Distribution. Cuba. Endemic L. petiminosus Germar 1836: 395; Jacquelin du Cuba localities. M: Cienaga de Zapata Val 1857: 202; Gundlach 1891: 290 Collections. Gundlach no. 1019 Distribution. Cuba. Endemic A. suturellus Gyllenhal 1836: 357; Gundlach 1891: Cuba localities. In seeds of Ceiba 289 Collections. Gundlach no. 48 Distribution. Cuba, Martinique Cuba localities. Ci: Cienfuegos. Gr: Bayamo Neomastix Dietz 1891: 254 Collections. Gundlach no. 1400 N. veritas Clark 1993b: 18 A. testaceus Boheman 1843: 225; Gundlach 1891: Distribution. Cuba. Endemic 289 Cuba localities. “Cuba” Distribution. Cuba, St. Vincent, Guadeloupe Collections. São Paulo (type) Cuba localities. M: Cienaga de Zapata. PR: Rangel Pseudanthonomus Dietz 1891: 248 Collections. Gundlach no. 1016 P. photon Clark 1990: 678 A. tigrinus Suffrian 1871: 135; Gundlach 1891: 287 Distribution. Cuba, Hispaniola Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. Gr: Bayamo. M: Cárdenas Collections. Washington (type) Collections. Gundlach no. 772 A. variegatus Suffrian 1871: 139; Gundlach 189l: TRIBE CAMAROTINI 288 SUBTRIBE PRIONOMERA Distribution. Cuba. Endemic Cuba localities. Widespread Odontopus Say 1831: 15 Collections. Gundlach no. 189 O. cubanus (Zayas) 1988: 192 (Prionomerus) A. varipes Jacquelin du Val 1857: 200; Gundlach Distribution. Cuba. Endemic 1891: 288 Cuba localities. Ca: Imías. SC: Pico Distribution. Cuba, Bahamas, USA (FL) Turquino. Type locality not given. In leaf litter Cuba localities. Widespread Collections. Zayas (type) Collections. Gundlach no. 162 A. verrucosus Suffrian 1871: 126; Gundlach 1891: TRIBE DERELOMINI 285 Distribution. Cuba. Endemic Celetes Schönherr 1836: 634 Cuba localities. Widespread C. dimidiatus (Suffrian) 1872: 160 (Eurges); Collections. Gundlach no. 16 Gundlach 1891: 300 (Euerges) Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 219

Cuba localities. Ci: Cienfuegos TRIBE PIAZORHININI Collections. Gundlach no. 229 Piazorhinus Schoenerr 1835: 471 Derelomus Schönherr 1825: column 583 P. species; Vàzquez et al. 1992: 45 D. albidus Suffrian 1872: 159; Gundlach 1891: 300 Distribution. Cuba, probably introduced Distribution. Cuba, Puerto Rico Cuba localities. Las Tunas and Holguin Prov- Cuba localities. M: Cárdenas inces. A destructive defoliator of Coccoloba Collections. Gundlach no. 467 uvifera Jacq. and other species of Coccoloba in the coastal zone (Vàzquez et al. 1992: 45) Phyllotrox Schönherr 1843: 189 Collections. P. liturellus Suffrian 1871: 152; Gundlach 1891: 292 TRIBE PYROPINI Distribution. Cuba. Endemic Cuba localities. Pyropus Schönherr 1836: 641 Collections. Gundlach no. 1643 P. sapphirinus Gyllenhal 1836: 642 P. variegatus Suffrian 1871: 154; Gundlach 1891: Distribution. Cuba, Jamaica, Hispaniola 292 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. M: Cienaga de Zapata Collections. Gundlach no. 1013 TRIBE SMICRONYCHINI

TRIBE ERODISCINI Smicronyx Schönherr 1843: 313 S. albosignatus Suffrian 1871: 170; Gundlach 1891: Erodiscus Schönherr 1825: column 583 297 E. morio (Suffrian) 1871: 151 (Toxeutes); Gundlach Distribution. Cuba. Endemic 1891: 292 Cuba localities. SS: Trinidad Distribution. Cuba. Endemic Collections. Gundlach no. 1387 Cuba localities. M: Cienaga de Zapata. In dry leaves of royal palm TRIBE TYCHIINI Collections. Gundlach no. 1000 SUBTRIBE LIGNYODINA

TRIBE OTIDOCEPHALINI Chionanthobius Pierce 1912: 168 C. darlingtoni Clark 1981: 690 Micromyrmex Sleeper 1953: 118 Distribution. Cuba. Endemic M. poeyi (Chevrolat) 1832: 107 (Otidocephalus); Cuba localities. Ha: Havana (t.l.) Jacquelin du Val 1857: 198; Gundlach 1891: Collections. Cambridge (type) 291 Distribution. Cuba, USA (FL) SUBTRIBE TYCHIINA Cuba localities. Widespread. On Asclepias curasavica Tychius Germar 1817: 340 Collections. Gundlach no. 228 T. auricapillus Suffrian 1871: 124; Gundlach 1891: M. pulicarius (Boheman) 1843: 207 (Otidocephalus); 284 Jacquelin du Val 1857: 198 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico Cuba localities. SC: Santiago de Cuba Cuba localities. Collections. Gundlach no. 1399 Collections. T. discoloma Suffrian 1871: 122; Gundlach 1891: 284 Myrmex Sturm 1826: 32 Distribution. Cuba. Endemic M. simplex Suffrian 1871: 150 (Otidocephalus) Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 947 Cuba localities. Gr: Bayamo Collections. Gundlach no. 1273 SUBFAMILY BAGOINAE 220 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Bagous Germar 1817: 340 SUBTRIBE DIORYMERINA B. argillaceus (Suffrian) 1871: 159 (Hydronomus); Gundlach 1891: 293 Diorymerus Schönherr 1825: column 586 Distribution. Cuba. Endemic D. substriatus Boheman 1844: 286 Cuba localities. M: Cienaga de Zapata D. pelos Jacquelin du Val 1857: 208; Gundlach Collections. Gundlach no. 1411 1891: 317 B. peregrinus (Suffrian) 1871: 156 (Hydronomus); Distribution. Cuba, South America Gundlach 1891: 293 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 189 Cuba localities. Collections. Gundlach no. 1916 SUBTRIBE EURHININA B. tessulatus (Suffrian) 1871: 160 (Hydronomus); Gundlach 1891: 293 Eurhinus Illiger 1807: 309 Distribution. Cuba. Endemic E. festivus (Fabricius) 1792: 404 (Ehynchaenus); Cuba localities. Gundlach 1891: 318 Cuba localities. Gundlach no. 1409 Distribution. Cuba, Hispaniola, México to South America SUBFAMILY BARIDINAE Cuba localities. Widespread TRIBE BARIDINI Collections. Gundlach no. 118 SUBTRIBE BARIDINA TRIBE MADOPTERINI Baris Germar 1817: 340 SUBTRIBE ZYGOBARIDINA B. azurea (Boheman) 1836: 668 (Baridius); Jacquelin du Val 1857: 205; Gundlach 1891: Apinocis Lea 1927: 371 316 A. insularis (Buchanan) 1947: 167 (Anacentrinus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. IJ: Nueva Gerona. M: Cuba localities. Cárdenas Collections. Collections. Gundlach no. 54. Pittsburgh B. chalybea (Boheman) 1844: 126 (Baridius); Geraeus Pascoe 1889: 323 Jacquelin du Val 1857: 205; Gundlach 1891: G. cubensis (Casey) 1920: 402 (Centrinaspis) 316 (B. chalybacus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Guantánamo (t.l.) Cuba localities. Widespread Collections. Washington (type) Collections. Gundlach no. 1179 G. penicilla (Herbst) 1797: 29 (Curculio) B. penicillata (Boheman) 1836: 665 (Baridius) Centrinus holosericeus (Gyllenhal) 1836: 760; Distribution. Cuba, Hispaniola Jacquelin du Val 1857: 207; Gundlach 1891: 317 Cuba localities. C. oblongus Schoenherr, Jacquelin du Val 1857: Collections. 206; Gundlach 1891: 317 (B. puncticollis (Schönherr) 1826: 277 (Baridius)) Distribution. Cuba, México, USA (TX-IN-FL) Distribution. West Indies (where?) Cuba localities. Cuba localities. Collections. Gundlach no. 169 Collections. G. punctatissimus (Boheman) 1836: 750 B. quadrimaculata (Boheman) 1836: 661 (Baridius); (Centrinus); Jacquelin du Val 1857: 206; Jacquelin du Val 1857: 205; Gundlach 1891: Gundlach 1891: 316 315 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Widespread Cuba localities. Widespread Collections. Gundlach no. 225 Collections. Gundlach no. 209 G. tomentosa (Casey) 1920: 403 (Centrinaspis) B. tabaci (Sallé) 1855: 269 Distribution. Cuba. Endemic Distribution. Cuba, Hispaniola Cuba localities. Cuba localities. Collections. Collections. A Checklist of the Beetles of Cuba 221

TRIBE PERIDINETINI TRIBE ZYGOPINI

Peridinetus Schönherr 1837: 467 Mnemynurus Heller 1895: 54 P. roeselii (Boheman) 1836: 206 (Heilipus) M. maestralis Zayas 1988: 198 P. maculatus Rosenschoeld, Jacquelin du Val Distribution. Cuba. Endemic 1857: 225; Gundlach 1891: 321 Cuba localities. Type locality not given P. poeyi Jacquelin du Val 1857: 225; Gundlach Collections. Zayas (type) 1891: 322 M. rangelianus Zayas 1988: 197 Distribution. Cuba, Puerto Rico Distribution. Cuba. Endemic Cuba localities. Widespread. On caisimón Cuba localities. PR: Soroa; Rancho Mundito. (Piperaceae) Type locality not given Collections. Gundlach no. 160 (P. Collections. Zayas (type) maculatus); no. 160.5 (P. poeyi) P. signatus Rosenschoeld 1837: 472; Jacquelin du Poecilogaster Heller 1895: 16 Val 1857: 227; Gundlach 1891: 322 P. rubi Zayas 1988: 200 Distribution. Cuba, Puerto Rico Distribution. Cuba. Endemic Cuba localities. Cuba localities. PR: Soroa. Type locality not Collections. given Collections. Zayas (type) SUBFAMILY CEUTORHYNCHINAE TRIBE CNEMOGONINI SUBFAMILY COSSONINAE TRIBE COSSONINI Panophthalmus Buchanan 1936: 145 P. cubanus Buchanan 1936: 146 Cossonus Clairville 1798: 58 Distribution. Cuba. Endemic C. cubae Boheman 1845: 275; Jacquelin du Val Cuba localities. Ha: Santiago de Las Vegas 1857: 236; Gundlach 1891: 337 (t.l.). In aerial roots of Cissus Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. Widespread Collections. Gundlach no. 166 SUBFAMILY CONODERINAE C. hamiltoni Slosson 1899: 193 TRIBE LECHRIOPINI Distribution. Cuba, St. Thomas, USA (FL) Cuba localities. Copturus Schönherr 1825: column 586 Collections. C. floridanus (Fall) 1906: 61 (Piazurus) C. impressus Boheman 1838: 1019; Jacquelin du Distribution. Cuba, Bahamas, USA (FL) Val 1857: 235; Gundlach 1891: 336 Cuba localities. Distribution. Cuba, Jamaica, Mona, Puerto Collections. Rico, St. Thomas, USA (FL) C. peninsularis Zayas 1988: 195 Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 168 Cuba localities. M: Cienaga de Zapata, San C. spathula Boheman 1838: 1035; Jacquelin du Val Blas (t.l.) 1857: 235; Gundlach 1891: 337 (C. spatula) Collections. Zayas (type) Distribution. Cuba, Hispaniola, Jamaica, C. sordidus Zayas 1988: 193 Puerto Rico, Grenada, México to South America, Distribution. Cuba. Endemic USA (FL) Cuba localities. SC: Pico Cuba (t.l.) Cuba localities. Widespread Collections. Zayas (type) Collections. Gundlach no. 238 C. turquinensis Zayas 1988: 194 C. subcostatus Boheman 1845: 270; Jacquelin du Distribution. Cuba. Endemic Val 1857: 234; Gundlach 1891: 336 Cuba localities. SC: Loma del Gato; Turquino Distribution. Cuba. Endemic (t.l.) Cuba localities. M: Cárdenas Collections. Zayas (type) Collections. Gundlach no. 167

Stenotrupis Wollaston 1873: 447 S. acicula Wollaston 1873: 603 222 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Hispaniola, Puerto Rico SUBTRIBE CRYPTORHYNCHINA Cuba localities. Found in dead fronds of Thrinax palms Apteromechus Faust 1896: 82 Collections. A. longus (LeConte) 1876: 248 (Tyloderma) Distribution. Cuba, USA (TX-AR-FL) TRIBE ACAMPTINI Cuba localities. Collections. Acamptopsis Champion 1909: 2 A. cubana Champion 1909: 2 Cryptorhynchus Illiger 1807: 330 Distribution. Cuba. Endemic C. albonotatus Klug 1829: 12 Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. Collections. TRIBE DRYOTRIBINI C. apicalis Klug 1829: 12 Distribution. Cuba. Endemic Caulophilus Wollaston 1854: 315 Cuba localities. C. oryzae (Gyllenhal) 1838: 1075 (Rhyncolus). The Collections. broadnosed grain weevil C. bisignatus Suffrian 1876: 149 Distribution. Cuba (introduced), Jamaica, Distribution. Cuba. Endemic Puerto Rico, México to Panamá, USA (CA-FL- Cuba localities. SC); stored products pest Collections. Cuba localities. C. bufonius Jacquelin du Val 1857: 211; Gundlach Collections. 1891: 319 Distribution. Cuba. Endemic TRIBE ONYCHOLIPINI Cuba localities. VC: Las Pozas Collections. Gundlach no. 1395 Pseudopentarthrum Wollaston 1873: 445 C. calandroides Suffrian 1876: 126 P. sp. Champion 1909: 14 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. Washington C. cordiger Suffrian 1876: 151 Distribution. Cuba. Endemic TRIBE PROECINI Cuba localities. Collections. Proeces Schoenherr 1838: 1080 C. curticollis Suffrian 1876: 158 P. depressus Wollaston 1873: 616; Buchanan 1947: Distribution. Cuba. Endemic 50 (Eucoptus Cuba localities. Distribution. Cuba, Montserrat, Nicaragua, Collections. Panamá, USA (FL) C. decipiens Suffrian 1876: 129 Cuba localities. Ci: Cayamas. Gu: Baracoa, Distribution. Cuba. Endemic Los Llanos. In leaf sheaths of royal palm Cuba localities. Collections. Washington Collections. C. dimidiatus Boheman 1837: 156; Jacquelin du TRIBE RHYNCOLINI Val 1857: 214; Gundlach 191: 320 Distribution. Cuba. Endemic Stenancylus Casey 1892: 693 Cuba localities. S. colomboi Casey 1892: 693 Collections. Distribution. Cuba, St. Vincent, Jamaica, C. discophorus Suffrian 1876: 156 USA (FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. frontalis Klug 1829: 12; Jacquelin du Val 1857: SUBFAMILY 212; Gundlach 1891: 319 TRIBE CRYPTORHYNCHINI Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 223

Cuba localities. Gu: Yateras. PR: Rangel Collections. Collections. Gundlach no. 1393 C. pilosus Suffrian 1876: 130 C. fuscoaeneus Suffrian 1876: 127 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. plagiellus Suffrian 1876: 155 C. gracilirostris Suffrian 1876: 135 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. posticatus Suffrian 1876: 144 C. griseolus Suffrian 1876: 133 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. pulchellus Suffrian 1876: 157 C. histrionicus Suffrian 1876: 141 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. ravus Boheman 1844: 347; Jacquelin du Val C. inconspicuus Suffrian 1876: 136 1857: 213; Gundlach 1891: 320 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. lepidus Suffrian 1876: 131 C. salebrosus Suffrian 1876: 161 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. levidipus Boheman 1837: 75; Jacquelin du Val C. scrofula Suffrian 1876: 166 1857: 210; Gundlach 1891: 318 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas Collections. Collections. Gundlach no. 120 C. squamiger Suffrian 1876: 153 C. multituberculatus Suffrian 1876: 148 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. stercorarius Suffrian 1876: 145 C. muticus Suffrian 1876: 162 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. strigirostris Suffrian 1876: 152 C. nigritellus Suffrian 1876: 160 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. (C. stultus (Fabricius) 1787: 107 (Curculio)) C. palmicola Suffrian 1876: 164 Distribution. West Indies (where?) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. suberosus Suffrian 1876: 142 C. pectinatus Suffrian 1876: 159 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. C. trilineatus Suffrian 1876: 132 C. peregrinus Suffrian 1876: 167 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. 224 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. variegatus Suffrian 1876: 147 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. Staseas Champion 1905: 530 Discophorellus Wibmer & O’Brien 1986: 247 S. difficilis (Klug) 1829: 12 (Cryptorhynchus); (D. circulus (Boheman) 1837: 64) Jacquelin du Val 1857: 213; Gundlach 1891: Distribution. Cuba?, South America 320 Cuba localities. Distribution. Cuba, México to South America Collections. Cuba localities. M: Cárdenas Collections. Gundlach no. 49 Macromerus Schönherr 1825: column 585 M. clavipes Boheman 1837: 185; Jacquelin du Val Tyloderma Say 1831: 19 1857: 214; Gundlach 1891: 321 T. aeneotinctum Champion 1905: 528 Distribution. Cuba. Endemic Distribution. Cuba, México to Panamá, USA Cuba localities. Gu: Monte Libana (FL) Collections. Gundlach no. 1323 Cuba localities. Collections. Neoulosomus O’Brien and Wibmer 1982: 143 T. circumcaribbeum Wibmer 1981: 43 N. furo (Suffrian) 1872: 193 (Ulosomus); Gundlach Distribution. Cuba, México to Panamá, USA 1891: 310 (FL) Distribution. Cuba. Endemic Cuba localities. Ci: Soledad Cuba localities. PR: Rangel Collections. Cambridge Collections. Gundlach no. 1382 T. fulvicornis Suffrian 1872: 192 (Analcis); N. immundus (Boheman) 1837: 317 (Ulosomus); Gundlach 1891: 309 Jacquelin du Val 1857: 217; Gundlach 1891: Distribution. Cuba. Endemic 310 Cuba localities. Distribution. Cuba. Endemic Collections.Gundlach no. 3651 Cuba localities. M: Cárdenas Collections. Gundlach no. 790 SUBTRIBE N. laticaudis (Suffrian) 1872: 196 (Ulosomus); Gundlach 1891: 310 Acalles Schönherr 1825: column 586 Distribution. Cuba, Hispaniola, USA (FL) A. apicalis Boheman 1837: 326; Jacquelin du Val Cuba localities. Ci: Cienfuegos 1857: 221; Gundlach 1891: 305 Collections. Gundlach no. 1923 Distribution. Cuba, Bahamas, Jamaica N. pallidicornis (Chevrolat) 1880: 236 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 114 Cuba localities. A. brunneus Suffrian 1872: 186; Gundlach 1891: Collections. 308 Distribution. Cuba. Endemic Phyrdenus LeConte 1876: 249 Cuba localities. P. divergens (Germar) 1824: 282 (Cryptorhynchus) Collections. Distribution. Cuba, México to South A. frontalis Suffrian 1872: 188; Gundlach 1891: 309 America, USA (FL-TX-DC) Distribution. Cuba. Endemic Cuba localities. Cuba localities. Gu: Yateras Collections. Collections. Gundlach no. 1397 P. mariceus (Germar) 1824: 281 (Cryptorhynchus) A. interruptus Suffrian 1872: 185; Gundlach 1891: Distribution. Cuba, México to South 308 America, USA (FL, AZ) Distribution. Cuba. Endemic Cuba localities. Cuba localities. M: Cárdenas Collections. Collections. Gundlach no. 617 A. lateritius Suffrian 1872: 177; Gundlach 1891: Riehlia Hustache 1936: 213 306 R. grisescens Hustache 1936: 213 Distribution. Cuba Endemic A Checklist of the Beetles of Cuba 225

Cuba localities. Faustinus Berg 1898: 18 Collections. F. cubae (Boheman) 1844: 342; (Cryptorhynchus); A. miserabilis Suffrian 1872: 178; Gundlach 1891: Jacquelin du Val 1857: 212 306 Distribution. Cuba, South America, USA Distribution. Cuba. Endemic (FL) Cuba localities. Cuba localities. Collections. Gundlach no. 1924 Collections. A. plebejus Suffrian 1872: 179; Gundlach 1891: 306 Distribution. Cuba. Endemic Gerstaeckeria Champion 1905: 471 Cuba localities. G. cubaecola Fisher 1928: 7 Collections. Gundlach no. 1925 Distribution. Cuba. Endemic A. ptochoides Suffrian 1872: 181; Gundlach 1891: Cuba localities. SC: Santiago (t.l.). Feeds on 306 Opuntia cactus Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. M: Cárdenas Collections. Gundlach no. 1926 Lembodes Schönherr 1844: 436 A. squamiger Suffrian 1872: 190; Gundlach 1891: L. solitarius Boheman 1844: 437 309 Distribution. Cuba, Jamaica, Hispaniola, St. Distribution. Cuba. Endemic Thomas, Guadeloupe, USA (FL) Cuba localities. Cuba localities. Collections. Gundlach no. 1928 Collections. A. stipulosus Suffrian 1872: 176; Gundlach 1891: 305 Pseudomus Schönherr 1837: 263 Distribution. Cuba. Endemic P. albosparsus Chevrolat 1880: 279 Cuba localities. M: Cárdenas. SS: Trinidad Distribution. Cuba. Endemic Collections. Gundlach no. 299 Cuba localities. A. sulcifrons Suffrian 1872: 182; Gundlach 1891: Collections. 307 (A. suleifrons) P. apiatus Boheman 1844: 389; Jacquelin du Val Distribution. Cuba. Endemic 1857: 216; Gundlach 1891: 312 Cuba localities. M: Cárdenas Distribution. Cuba. Endemic Collections. Gundlach no. 1927 Cuba localities. M: Cardenas; Fermina A. terrosus Suffrian 1872: 175: Gundlach 1891: 305 Collections. Gundlach no. 226 Distribution. Cuba. Endemic P. bimaculatus Suffrian 1872: 206; Gundlach 1891: Cuba localities. M: Cárdenas 314 Collections. Gundlach no. 730 Distribution. Cuba. Endemic Cuba localities. Ci: Cienfuegos Euscepes Schönherr 1844: 429 Collections. Gundlach no. 1646 E. erinacellus Chevrolat 1880: 151 P. bohemani Kuschel 1955: 287; replacement name Distribution. Cuba. Endemic P. notatus Boheman 1837: 266, not Klug 1829 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. E. porcellus Boheman 1844: 430; Jacquelin du Val Collections. 1857: 221; Gundlach 1891: 311 (E. porullus) P. cacuminatus Boheman 1844: 390; Gundlach Distribution. Cuba, Jamaica to Martinique, 1891: 313 (P. caurminatus) México to Panamá, USA (FL) P. acuminatus Jacquelin du Val 1857: 216 Cuba localities. Widespread Distribution. Cuba. Endemic Collections. Gundlach no. 744 Cuba localities. Widespread E. postfasciatus (Fairmaire) 1849: 513 Collections. Gundlach no. 662 (Cryptorhynchus). The West Indian sweet po- P. deltoides Buchanan 1947: 49 tato weevil Distribution. Cuba. Endemic Distribution. Cuba (introduced), throughout Cuba localities. SS: Mina Carlota; Topes de the West Indies, USA (CA); native to Old World) Collantes (t.l.) Cuba localities. Collections. Washington (type) Collections. 226 Arthropods of Florida and Neighboring Land Areas, Vol. 18

P. fistulosus (Klug) 1829: 12 (Cleogonus); Jacquelin TRIBE GASTEROCERCINI du Val 1857: 217; Gundlach 1891: 313 Distribution. Cuba. Endemic Cophes Champion 1905: 515 Cuba localities. Widespread C. oblongus (LeConte) 1876: 256 (Cryptorhynchus) Collections. Gundlach no. 170 Distribution. Cuba, USA (TX-SC-FL) P. maximus Suffrian 1872: 204; Gundlach 1891: Cuba localities. 314 Collections. Distribution. Cuba. Endemic Cuba localities. Gu Baracoa.; Mata. SC: SUBFAMILY CYCLOMINAE Santiago de Cuba TRIBE RHYTIRRININI Collections. Gundlach no. 1388 SUBTRIBE LISTRODERINA P. notatus (Klug) 1829: 13 (Cleogonus); Jacquelin du Val 1857: 215; Gundlach 1891: 312 Listronotus Jekel 1865: 566 Distribution. Cuba. Endemic L. nigropunctatus (Suffrian) 1870: 228 Cuba localities. M: Cárdenas (Listroderes); Gundlach 1891: 299 Collections. Gundlach no. 87 Distribution. Cuba. Endemic P. rugifer Suffrian 1872: 201; Gundlach 1891: 313 Cuba localities. M: Guamutas Distribution. Cuba. Endemic Collections. Gundlach no. 929 Cuba localities. Gr: Bayamo Collections. Gundlach no. 1403 SUBFAMILY ENTIMINAE P. trisignatus (Klug) 1829: 12 (Cleogonus) TRIBE ANYPOTACTINI Cleogonus viduus Boheman 1837: 265; Jacquelin du Val 1857: 215; Gundlach 1891: 311 (P. Polydacrys Schönherr 1834: 130 viduos) P. scansorius (Klug) 1829: 13 (Sitona) Distribution. Cuba. Endemic P. modestus Gyllenhal 1834: 131; Jacquelin du Cuba localities. M: Fermina; Guamutas Val 1857: 179;Gundlach 1891: 323 Collections. Gundlach no. 14 Distribution. Cuba, Dominica, Guadeloupe Cuba localities. Widespread Tylodes Sahlberg 1823: 49 Collections. Gundlach no. 1178 T. bullatus (Boheman) 1837: 340 (Acalles); P. sulcifrons Zayas 1988: 170 Jacquelin du Val 1857: 218; Gundlach 1891: Distribution. Cuba. Endemic 307 Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. M: Cárdenas Collections. Gundlach no. 163 TRIBE EMBRITHINI T. laticollis (Boheman) 1837: 341 (Acalles); Jacquelin du Val 1857: 218; Gundlach 1891: Peribrotus Gerstaecker 1871: 72 307 P. circularis Zayas 1988: 176 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. M: Cárdenas Cuba localities. Gr: Cabo Cruz (t.l.) Collections. Gundlach no. 1240 Collections. Zayas (type) T. magnicollis (Boheman) 1837: 342 (Acalles); Jacquelin du Val 1857: 219; Gundlach 1891: TRIBE EUSTYLINI 308 Distribution. Cuba. Endemic Diaprepes Schönherr 1823: column 1140 Cuba localities. M: Cárdenas D. famelicus (Olivier) 1790: 544 (Curculio); not Collections. Gundlach no. 1636 Leng and Mutchler 1914: 469 T. quadriplicatus Jacquelin du Val 1857: 219 D. f. elegantulus Gyllenhal 1834: 13; subspecies Distribution. Cuba. Endemic of Cuba and Martinique Cuba localities. Distribution. Cuba, Lesser Antilles Collections. Cuba localities. Collections. D. revestitus Chevrolat 1880: 165 Distribution. Cuba. Endemic A Checklist of the Beetles of Cuba 227

Cuba localities. E. pictus (Guérin-Méneville) 1847: 9 (Prepodes); Collections. Jacquelin du Val 1857: 181; Gundlach 1891: D. sommeri (Rosenschoeld) 1840: 339 (Exophthal- 324 mus); Jacquelin du Val 1857: 180; Gundlach E. p. fulvovirgatus (Marshall) 1934: 625; Cuba, 1891: 324 endemic subspecies; Sierra Maestra (t.l.). Lon- Distribution. Cuba. Endemic don (type) Cuba localities. Ha: Cojimar Distribution. Cuba. Endemic Collections. Gundlach no. 1245 Cuba localities. Collections. Exophthalmus Schönherr 1823: c.1140 E. pseudohybridus Zayas 1988: 183 E. azureus Zayas 1988: 188 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Gran Piedra (t.l.) Cuba localities. SS: Buenos Aires (t.l.) Collections. Zayas (type) Collections. Zayas (type) E. quadriguttatus Zayas 1988: 188 E. bayamensis Zayas 1988: 177 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino (t.l.) Cuba localities. Gr: Bayamesa (t.l.) Collections. Zayas (type) Collections. Zayas (type) E. scalaris (Boheman) 1840: 349 (Prepodes); E. cristalensis Zayas 1988: 181 Jacquelin du Val 1857: 181; Gundlach 1891: Distribution. Cuba. Endemic 325 Cuba localities. Ho: Sierra Cristal (t.l.) Distribution. Cuba, Costa Rica Collections. Zayas (type) Cuba localities. M: Cárdenas. A pest of cit- E. griseus Zayas 1988: 187 rus crops (Vàzquez et al. 1992: 45) Distribution. Cuba. Endemic Collections. Gundlach no. 202 Cuba localities. SC: Gran Piedra (t.l.) E. siberianus Zayas 1988: 178 Collections. Zayas (type) Distribution. Cuba. Endemic E. gundlachi Chevrolat 1880: 165 Cuba localities. Gr: La Siberia (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. E. sparsimviridis Zayas 1988: 185 Collections. Distribution. Cuba. Endemic E. humeridens (Marshall) 1934: 623 (Prepodes) Cuba localities. Ho: Sierra Cristal (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. SC: Pico Turquino (t.l.) (E. sphacelatus (Olivier) 1807: 319 (Curculio)) Collections. London (type) Distribution. Jamaica to Puerto Rico E. hybridus (Chevrolat) 1858: 210 ( Praepodes) Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Cuba E. sulphuratus Chevrolat 1879: XCVII Collections. Distribution. Cuba. Endemic E. lepidus (Chevrolat) 1858: 210 (Praepodes) Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. E. viridis Zayas 1988: 187 Collections. Distribution. Cuba. Endemic E. maculosus Chevrolat 1879: XCVIII Cuba localities. Gu: Cuchillas de Toa (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. E. zayasi Zayas 1988: 179 Collections. Distribution. Cuba. Endemic E. mixtus Chevrolat 1880: 165 Cuba localities. Gr: La Siberia (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Collections. Pseudeustylus Champion 1911: 296 E. multipunctatus Zayas 1988: 184 P. planicollis Marshall 1934: 626 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gr: La Siberia (t.l.) Cuba localities. SC: Sierra Maestra (t.l.) Collections. Zayas (type) Collections. 228 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE GEONEMINI Cuba localities. SC: Sierra Maestra (t.l.) Collections. London (type) Compsonomus Jekel 1875: 138 L. argus (Reiche) 1840: 275 (Ptilopus); Jacquelin C. alternans (Klug) 1829: 13 (Geonemus); Jacquelin du Val 1857: 184; Gundlach 1891: 327 du Val 1857: 196; Gundlach 1891: 333; Distribution. Cuba, USA (FL) Buchanan 1947: 44 (Eugeonemus) Cuba localities. SC: Santiago de Cuba Geonemus alternans (Boheman) 1834: 293, not Collections. Gundlach no. 1337 Klug 1829 L. aurifer (Drury) 1773: 68 (Curculio) Distribution. Cuba. Endemic Distribution. Cuba, Jamaica Cuba localities. “Cuba” as t.l. Cuba localities. Collections. Gundlach no. 1263 Collections. C. aureosquamosus (Jacquein du Val) 1857: 195 L. bivirgatus Marshall 1934: 621 (Geonemus); Gundlach 1891: 332; Buchanan Distribution. Cuba. Endemic 1947: 44. (Eugeonemus) Cuba localities. SC: Pico Turquino (t.l.) Distribution. Cuba. Endemic Collections. London (type) Cuba localities. “Cuba” as t.l. L. bruneri Marshall 1933: 59 Collections. Gundlach no. 354 Distribution. Cuba. Endemic C. irregularis (Perroud) 1853: 518 (Geonemus); Cuba localities. SS: Buenos Aires (t.l.) Buchanan 1947: 44 (Eugeonemus) Collections. London (type). Washington Distribution. Cuba. Endemic L. buchanani Marshall 1933: 59 Cuba localities. “Cuba” as t.l. Distribution. Cuba. Endemic Collections. Cuba localities. SS: Trinidad Mts., Buenos C. robustus (Perroud) 1853: 515 (Geonemus); Aires (t.l.); San Blas Buchanan 1947: 44 (Eugeonemus) Collections. London. Washington Distribution. Cuba. Endemic L. cabocruz Zayas 1988: 162 Cuba localities. “Cuba”as t.l. Distribution. Cuba. Endemic Collections. Cuba localities. Gr: Cabo Cruz (t.l.). Ha: Ciudamar. SC: Daiquiri; Pico Turquino Doleropus Buchanan 1947: 46. Endemic genus Collections. Zayas (type) D. apterus Buchanan 1947: 47 L. cristalensis Zayas 1988: 161 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. PR: Guane (t.l.). Found on Cuba localities. Ho: Sierra Cristal (t.l.) tobacco and squash Collections. Zayas (type) Collections. Washington (type) (L. distortus Gyllenhal 1840: 393 Distribution. West Indies (where?) Ischionoplus Chevrolat 1878: IX Cuba localities. I. niveoguttatus Chevrolat 1878: X Collections. Distribution. Cuba. Endemic L. festivus Zayas 1988: 159 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. SC: Pico Turquino (t.l.) Collections. Zayas (type) Lachnopus Schoenherr 1840: 380 L. floridanus Horn 1876: 101 L. acunae Zayas 1988: 160 Distribution. Cuba, USA (FL) Distribution. Cuba. Endemic Cuba localities. Cuba localities. SC: Sierra Maestra (t.l.) Collections. Collections. Zayas (type) L. guerinii Jacquelin du Val 1857: 185; Gundlach L. acuticollis (Gyllenhal) 1834: 37 (Ptilopus); 1891: 327 Jacquelin du Val 1857: 189; Gundlach 1891: Distribution. Cuba. Endemic 329 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Gr: Bayamo. M: Cárdenas L. guttatupunctatus Zayas 1988: 155 Collections. Gundlach no. 112 Distribution. Cuba. Endemic L. alboguttatus Marshall 1934: 622 Cuba localities. Gu: Hacienda de San Carlos Distribution. Cuba. Endemic (t.l.) A Checklist of the Beetles of Cuba 229

Collections. Zayas (type) L. porcus Zayas 1988: 158 L. hispidus (Gyllenhal) 1834: 34 (Ptilopus); Distribution. Cuba. Endemic Jacquelin du Val 1857: 188; Gundlach 1891: Cuba localities. SC: Daiquirí (t.l.) 329 Collections. Zayas (type) Distribution. Cuba, USA (FL) L. siboney Zayas 1988: 166 Cuba localities. Gr: Bayamo. IJ: Nueva Distribution. Cuba. Endemic Gerona. M: Cárdenas. A pest of citrus crops Cuba localities. SC: Juraguá; Siboney (t.l.) (Vàzquez et al. 1992: 45) Collections. Zayas (type) Collections. Gundlach no. 213. Pittsburgh L. sparsimguttatus Perroud 1853: 481 L. leonorae Zayas 1988: 165 (Lachnopus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Cuba localities. Gu: Baracoa, Farola (t.l.); Collections. in pinelands L. splendidus Boheman 1840: 382; Jacquelin du Collections. Zayas (type) Val 1857: 188; Gundlach 1891: 328 L. lineatoguttatus Perroud 1853: 468 Distribution. Cuba. Endemic Lachnopus seriepunctatus (Jacquelin du Val) Cuba localities. Gr: Bayamo 1857: 187; Gundlach 1891: 328 Collections. Gundlach no. 206 Distribution. Cuba. Endemic L. sublineatus Perroud 1853: 478; Jacquelin du Val Cuba localities. 1857: 192; Gundlach 1891: 330 Collections. Gundlach no. 205 1/2 Distribution. Cuba. Endemic L. luctuosus (Klug) 1829: 13 (Sitona); Jacquelin Cuba localities. du Val 1857: 183 (Prepodes); Gundlach 1891: Collections. 327 L. vittatus (Klug) 1829: 13 (Ptilopus); Jacquelin Distribution. Cuba. Endemic du Val 1857: 186; Gundlach 1891: 328 Cuba localities. ? Guamacaro Distribution. Cuba. Endemic Collections. Gundlach no. 117 Cuba localities. Widespread L. magdae Zayas 1988: 164 Collections. Gundlach no. 205 Distribution. Cuba. Endemic Cuba localities. SC: Pico Turquino (t.l.) Tetrabothynus Labram and Imhoff [1852]: nr. Collections. Zayas (type) 68 L. mayari Zayas 1988: 156 T. spectabilis (Klug) 1829: 13 (Prepodes); Jacquelin Distribution. Cuba. Endemic du Val 1857: 183; Gundlach 1891: 326 Cuba localities. Ho: Mayarí (t.l.); in pines (Prepodes sputabilis) Collections. Zayas (type) Distribution. Cuba. Endemic L. multipunctatus Jacquelin du Val 1857: 190; Cuba localities. M: Cárdenas Gundlach 1891: 330 Collections. Gundlach no. 204 Distribution. Cuba. Endemic T. tripunctata Zayas 1988: 154 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. Ho: Sierra Cristal (t.l.) L. niveoirroratus Jacquelin du Val 1857: 189; Collections. Zayas (type) Gundlach 1891: 330 Distribution. Cuba. Endemic Tropirhinus Schönherr 1823: c.1140 Cuba localities. M: Fundador Canímar T. elegans (Guérin-Méneville) 1847: 10 (Prepodes); Collections. Gundlach no. 207 Jacquelin du Val 1857: 182; Gundlach 1891: L. oteroi Marshall 1933: 60 326 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SS: Buenos Aires (t.l.) Cuba localities. Collections. London (type). Washington Collections. L. pollinarius Gyllenhal 1840: 387; Jacquelin du T. tredecimmaculatus (Guérin-Méneville) 1847: 10 Val 1857: 191: Gundlach 1891: 330 (Prepodes); Jacquelin du Val 1857: 182; Distribution. Cuba. Endemic Gundlach 1891: 325 Cuba localities. Ci: Cienfuegos Distribution. Cuba. Endemic Collections. Gundlach no. 1330 Cuba localities. Collections. 230 Arthropods of Florida and Neighboring Land Areas, Vol. 18

TRIBE LORDOPINI P. litus (Germar) 1824: 431 (Cyphus); Jacquelin du Val 1857: 193; Gundlach 1891: 331 Nesolordops Marshall 1934: 627. Endemic ge- (Pachnacus). The citrus root weevil nus Distribution. Cuba (widespread), Jamaica, N. myersi Marshall 1934: 627 México, USA (FL) Distribution. Cuba. Endemic Cuba localities. IJ: Nueva Gerona. A wide- Cuba localities. Ci: Soledad (t.l.). On Pani- spread pest of citrus crops and of cassava in cum maximum Matanzas and elsewhere (Vàzquez et al. 1992: Collections. London (type) 45) Collections. Gundlach no. 1699. Pittsburgh TRIBE NAUPACTINI P. pater Zayas 1988: 167 Distribution. Cuba. Endemic Artipus Sahlberg 1823: 22 Cuba localities. Ho: Mayarí (t.l.). SC: A. minima Zayas 1988: 153 Juraguá Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Gu: Baracoa; Imías (t.l.) P. psittacus (Olivier) 1807: 339 (Curculio) Collections. Zayas (type) Distribution. Cuba, Puerto Rico A. unguiculatus Chevrolat 1880: 254 Cuba localities. IJ Distribution. Cuba. Endemic Collections. Cuba localities. P. rosadonetoi Lopez Castilla 1992: 4 Collections. Distribution. Cuba. Endemic Cuba localities. Gu: Baracoa (t.l.) Pantomorus Schoenherr 1840: 942 Collections. Havana (type) (Pantomorus cervinus (Boheman) 1847: 17) P. godmanni (Crotch) 1867: 389 (Asynonychus) Pandeleteius Schönherr 1834: 129 Distribution. West Indies (where?), intro- P. maculatus Zayas 1988: 172 duced from South America, North to South Distribution. Cuba. Endemic America, Old World Cuba localities. M: Cienaga de Zapata (t.l.) Cuba localities. Collections. Zayas (type) P. mayari Zayas 1988: 172 TRIBE TANYMECINI Distribution. Cuba. Endemic SUBTRIBE TANYMECINA Cuba localities. Ho: Altiplano de Mayarí (t.l.) Collections. Zayas (type) Pachnaeus Schönherr 1826: 121 P. pinarensis Zayas 1988: 171 P. alayoi Lopez Castilla 1992: 2 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Gu: Pinares del Yunque de Cuba localities. PR: Guane (t.l.) Baracoa (t.l.) Collections. Havana Collections. Zayas (type) P. azurescens Gyllenhal 1834: 58; Jacquelin du Val P. sulphurosus Zayas 1988: 175 1857: 193; Gundlach 1891: 331 (Pachnacus) Distribution. Cuba. Endemic P. a. griseus Gyllenhal 1834: 59 Cuba localities. Gu: Imías (t.l.) Distribution. Cuba. Endemic Collections. Zayas (type) Cuba localities. Widespread. A pest of citrus crops (Vàzquez et al. 1992: 45) Scalaventer Howden 1970: 3 Collections. Gundlach no. 1187 S. cubensis Howden 1970: 31 P. costatus Perroud 1853: 495 Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. SC: coast below Pico Cuba localities. Turquino (t.l.) Collections. Collections. Cambridge (type) P. juvenalis Zayas 1988: 169 S. subtropicus (Fall) 1907: 263 (Pandeletejus); Distribution. Cuba. Endemic Howden 1971: 21 Cuba localities. IJ (t.l.) Distribution. Cuba, USA (FL) Collections. Zayas (type) Cuba localities. PR: Viñales Collections. Cambridge A Checklist of the Beetles of Cuba 231

SUBFAMILY HYPERINAE Cuba localities. Ci: Cienfuegos. Ha: Santiago TRIBE CEPURINI de las Vegas (in banana corms) Collections. Gundlach no. 1177. Washington Cephalalges Schönherr 1840: 467 C. cubae (Chevrolat) 1838: 281 (Phytonomus); Jacquelin du Val 1857: 194; Gundlach 1891: TRIBE CLEOGONINI 332 (Hypera) Distribution. Cuba. Endemic Cleogonus Schönherr 1825: column 86 Cuba localities. M: Cárdenas C. grossulus Suffrian 1872: 170; Gundlach 1891: Collections. Gundlach no. 224 303 (Cleogenus) Distribution. Cuba. Endemic SUBFAMILY LIXINAE Cuba localities. TRIBE LIXINI Collections. Gundlach no. 1922

Lixus Fabricius 1801: 498 Rhyssomatus Schönherr 1837: 364 L. merula Suffrian 1871: 168; Gundlach 1891: 296 R. crispicollis Boheman 1837: 367; Jacquelin du Distribution. Cuba, USA (MA-FL-TX) Val 1857: 222; Gundlach 1891: 304 Cuba localities. Distribution. Cuba. Endemic Collections. Cuba localities. M: Cárdenas L. tardus Suffrian 1871: 166; Gundlach 189l: 296 Collections. Gundlach no. 869 Distribution. Cuba. Endemic R. ebeninus (Boheman) 1837: 391 (Chalcodermus); Cuba localities. M: Cardenas; Cienaga de Jacquelin du Val 1857: 222; Gundlach 1891: Zapata 304 Collections. Gundlach no. 159 Distribution. Cuba, Puerto Rico Cuba localities. Gr: Bayamo. M: Cienaga de SUBFAMILY Zapata TRIBE ANCHONINI Collections. Gundlach no. 728

Anchonus Schönherr 1825: column 584 TRIBE CONOTRACHELINI A. aspericollis Suffrian 1871: 176; Gundlach 1891: 298 Conotrachelus Dejean 1835: 296 Distribution. Cuba. Endemic C. albicans Suffian 1872: 168; Gundlach 1891: 302 Cuba localities. M: Cárdenas Distribution. Cuba, Puerto Rico Collections. Gundlach no. 1199 Cuba localities. SC: Santiago de Cuba A. bicornis Suffrian 1871: 179; Gundlach 1891: 298 Collections. Gundlach no. 810 Distribution. Cuba. Endemic C. cariniscutum Marshall 1934: 630 Cuba localities. M: Cárdenas Distribution. Cuba, Hispaniola Collections. Cuba localities. Ci: Soledad (t.l.) A. incrassatus Suffrian 1871: 175; Gundlach 1891: Collections. London (type) 298 (A. incrusatus) C. coronatus LeConte 1878: 430 Distribution. Cuba. Endemic Distribution. Cuba, USA (FL, AR, SC) Cuba localities. IJ Cuba localities. Collections. Gundlach no. 1236 Collections. A. rusticus Suffrian 1871: 177; Gundlach 1891: 299 C. diaconitus (Klug) 1829: 12 (Balanius); Jacquelin Distribution. Cuba. Endemic du Val 1857: 223; Gundlach 1891: 303 Cuba localities. M: Cárdenas Distribution. Cuba, México to South America Collections. Gundlach no. 188 Cuba localities. A. suillus (Fabricius) 1792: 402 (Curculio); Collections. Jacquelin du Val 1857: 203; Gundlach 1891: C. lassulus Boheman 1837: 409; Jacquelin du Val 298; Buchanan 1947: 48 1857: 224; Gundlach 1891: 303 Distribution. Cuba, Bahamas, Hispaniola, Distribution. Cuba. Endemic Puerto Rico, Guadeloupe, St. Barthélemy, USA Cuba localities. (FL) Collections. 232 Arthropods of Florida and Neighboring Land Areas, Vol. 18

C. lineola Suffrian 1872: 167; Gundlach 1891: 302 Distribution. Cuba, Hispaniola, México to Distribution. Cuba. Endemic Panamá, South America Cuba localities. Gr: Bayamo. M: Cienaga de Cuba localities. M: Cárdenas Zapata Collections. Gundlach no. 9 Collections. Gundlach no. 1008 M. rusticus (Klug) 1829: 13; Gundlach 1891: 295 C. presbyta Suffrian 1872: 165; Gundlach 1891: 302 (Heilipus) Distribution. Cuba. Endemic Distribution. Cuba, México, South America Cuba localities. M: Cienaga de Zapata. SS: Cuba localities. Widespread; under bark of Trinidad dead trees Collections. Gundlach no. 1003 Collections. Gundlach no. 8 C. sapotae Barber 1924: 184 Distribution. Cuba. Endemic TRIBE LYMANTINI Cuba localities. Collections. Caecossonus Gilbert 1955: 193 C. serpentinus (Klug) 1829: 12 (Balanius); C. decuanus Osella 1973: 373 (Caecocosonnus, Jacquelin du Val 1857: 224; Gundlach 1891: lapsus), 1976: 674; 1977b: 399; 1980: 366; 301 Howden 1992: 18 Distribution. Cuba, Jamaica, Hispaniola, Distribution. Cuba. Endemic Bahamas, México to Panamá, USA (FL, GA) Cuba localities. PR: Rio Cuyaguateje, Dolina Cuba localities. M: Cardenas; Cienaga de de Cueva de la Pluma (t.l.); an eyeless soil in- Zapata; Zarabanda habitant Collections. Gundlach no. 165 Collections. Bucharest (type?). Verona C. verticalis (Klug) 1829: 12 (Balanius); Jacquelin du Val 1857: 224; Gundlach 1891: 301 Decuanellus Osella 1977b: 399 Distribution. Cuba, México to South America D. vinai Osella 1977b: 399; Howden 1992: 32 Cuba localities. Distribution. Cuba. Endemic Collections. Gundlach no. 1921 Cuba localities. Gu: Rio Jojo (t.l.); an eyeless soil inhabitant TRIBE CYCLOTERINI Collections. Bucharest (type)? Verona

Gononotus LeConte 1876: 336 Dioptrophorus Faust 1892: 35 G. angulicollis (Suffrian) 1871: 181 (Anchonus); D. sp. Champion 1902: 92 Gundlach 1891: 299 Distribution. Cuba. Endemic Distribution. Cuba, Puerto Rico, México, Cuba localities. USA (FL) Collections. Cuba localities. M: Cárdenas Collections. Gundlach no. 1198 Pseudoalaocybites Osella 1980: 369 P. negreai (Osella) 1977b: 397 (Caecocosonnus); TRIBE HYLOBIINI Osella 1980: 372 (Alaocybites); Howden 1992: SUBTRIBE HYLOBIINA 29 Distribution. Cuba. Endemic; eyeless soil in- Heilipus Germar 1824: 399 habitant H. draco (Fabricius) 1801: 461 (Rhynchaenus); Cuba localities. SC: Cabesada de Rio Baconao Gundlach 1891: 294 (t.l.) Distribution. Cuba, México to Panamá, Collections. Bucharest (type)? Verona South America Cuba localities. Widespread Pseudocaecocossonus Osella 1977a: 393 Collections. Gundlach no. 119 P. zayasi Osella 1977a: 393, 1976: 674, 1977b: 399, 1980: 416; Howden 1992: 22 Marshallius Kuschel 1955: 291,k Distribution. Cuba. Endemic M. guttatus (Boheman) 1843: 52 (Heilipus); Cuba localities. Ho: Sierra Cristal (t.l.); an Jacquelin du Val 1857: 197; Gundlach 1891: eyeless soil inhabitant 294 Collections. Verona (type). Zayas A Checklist of the Beetles of Cuba 233

TRIBE STERNECHINI Collections. Washington (type)

Chalcodermus Dejean 1835: 297 SUBTRIBE BOTHROSTERNINA C. pupillatus (Suffrian) 1872: 173 (Rhyssonatus); Gundlach 1891: 304 (Rhyssomatus) Cnesinus LeConte 1868: 171 Distribution. Cuba, Puerto Rico C. cubensis Blackman 1943b: 371 Cuba localities. Ci: Cienfuegos Distribution. Cuba. Endemic Collections. Gundlach no. 1180 Cuba localities. Ci: Cayamas (t.l.) Collections. Washington (type) Sternechus Schönherr 1826: 251 S. insularis Boheman 1843: 356; Jacquelin du Val Pagiocerus Eichhoff 1868: 148 1857: 202; Gundlach 1891: 295 (Sternuchus); P. frontalis (Fabricius) 1801: 389 (Bostrichus) 315 P. rimosus Eichhoff 1868b: 148 (Cuba t.l., type Distribution. Cuba. Endemic lost) Cuba localities. Ca: Manzanillo. M: Cárdenas Distribution. Cuba (introduced), Guadeloupe, Collections. Gundlach no. 528; no. 1182 México to Panamá to Brazil, USA (NC, SC, GA, S. pectoralis Suffrian 1872: 156; Gundlach 1891: FL, TX); economic pest of avocado and stored 296 (Sternuchus) grain Distribution. Cuba. Endemic Cuba localities. Cuba localities. Collections. Collections. Gundlach no. 1637 SUBTRIBE PHLOEOTRIBINA TRIBE TRYPETIDINI Phloeotribus Latreille 1804: 108 Nanus Schönherr 1844: 89 P. setulosus Eichhoff 1868: 14 N. uniformis Boheman 1844: 90; Jacquelin du Val P. atlanticus Schedl 1951: 81 (Cuba t.l., type in 1857: 204; Gundlach 1891: 314 Vienna) Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba, Jamaica, México to Guadeloupe, México to Panamá, USA (FL) Panamá to Brazil Cuba localities. Widespread Cuba localities. Collections. Gundlach no. 230 Collections.

SUBFAMILY SCOLYTINAE SUBTRIBE PHLOEOSININA The bark and ambrosia beetles Reviewed by D. Bright, January, 2002. Chramesus LeConte 1868: 168 Members of this subfamily usually bore galler- C. opacicollis Eggers 1940: 124 ies under the bark of trees and shrubs in the cam- Distribution. Cuba, Jamaica, Dominica, bium layer. Synonyms which have not been used Grenada, Guadeloupe in a Cuban context are not given. See Wood and Cuba localities. Bright (1992) for these and other details and for Collections. host plants. Few Cuban host records are available. C. robustus Schedl 1948: 264 Bright (1985) is a summary of West Indian species Distribution. Cuba. Endemic and literature. Atkinson and Peck (1994) present Cuba localities. details for these beetles in the southern four sub- Collections. Vienna (type) tropical counties of Florida. Wood (1979) summa- rizes the fauna of the USA. Cladoctonus Strohmeyer 1911: 17 C. cubensis (Wood) 1961: 105 (Hopliophthorus) TRIBE HYLESININI Distribution. Cuba, México SUBTRIBE PHRIXOSOMINA Cuba localities. Ci: Cayamas (t.l.) Collections. Washington (type) Phrixosoma Blandford 1897: 148 P. parva Blackman 1943b: 393 Dendrosinus Chapuis 1869: 236 Distribution. Cuba. Endemic (D. bourreriae Schwarz 1920: 225) Cuba localities. Ci: Cayamas (t.l.) 234 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Bahamas, Jamaica, Puerto Distribution. Cuba, Jamaica, México to Ven- Rico, USA (FL) ezuela Cuba localities. Cuba localities. Ci: Soledad. SC: Loma del Collections. Gato Collections. Ottawa SUBTRIBE HYPOBORINA SUBTRIBE CTENOPHORINA Chaetophloeus LeConte 1876: 382 C. cubensis Bright 1981: 159 Pycnarthrum Eichhoff 1878: 104 Distribution. Cuba. Endemic P. hispidium (Farrari) 1867: 19 Cuba localities. SS: Mapos (t.l.) P. gracile Eichhoff 1878: 104; Cuba (t.l.), Wash- Collections. Ottawa (type) ington (type) C. insularis (Blackman) 1940: 400 (Renocis) Distribution. Cuba (t.l.), Jamaica, Puerto Distribution. Cuba, Bahamas, Virgin Is- Rico, Virgin Islands, Guyana, Venezuela, lands, USA (FL) México to Panamá, USA (TX, FL) Cuba localities. Cuba localities. Collections. Collections. Type in Vienna, lost

Liparthrum Wollaston 1854: 294 Scolytodes Ferrari 1867: 77 L. necopinus (Bright) 1982: 166 (Trypanophellos) S. cubensis (Schedl) 1972: 56 (Hexacolus) Distribution. Cuba. Endemic Distribution. Cuba. Endemic Cuba localities. Ci: Cayamas (t.l.) Cuba localities. PR: Malas (t.l.) Collections. Washington (type) Collections. Vienna (type) S. glaber (Eichhoff) 1868: 400 (Hexacolus) TRIBE SCOLYTINI Distribution. Cuba. Endemic SUBTRIBE SCOLYTINA Cuba localities. Collections. Brussels (type) Cnemonyx Eichhoff 1868: 150 S. nitidissimus (Eggers) 1940: 135 (Hexacolus) C. ficus (Schwarz) 1896: 44 (Loganicus) Distribution. Cuba, Guadeloupe Distribution. Cuba, Bahamas, Virgin Is- Cuba localities. SC: Gran Piedra. ?Terrazas, lands, USA (FL); host tree, Hippomane in Cecropia leaves mancinella; see Atkinson, 1993, Coleop. Bull. Collections. Ottawa. 47: 287-288 S. notatus (Eggers) 1940: 133 Cuba localities. Distribution. Cuba, Puerto Rico, Guadeloupe, Collections. Dominica C. vagabundus (Wood) 1961: 89 (Loganius) Cuba localities. Distribution. Hispaniola, Puerto Rico, Vir- Collections. gin Islands, Panamá, USA (FL) Cuba localities. SUBTRIBE MICRACINA Collections. Hylocurus Eichhoff 1872: 133 Scolytopsis Blandford 1896: 123 H. alienus Eichhoff 1878: 301 S. puncticollis Blandford 1896: 123 Distribution. Cuba. Endemic S. cubensis Wood 1967: 123 (Ci: Cayamas, t.l.; Cuba localities. Washington, type) Collections. Vienna (type) Distribution. Cuba, México to Argentina, H. cuspidatus Eggers 1951: 153 (USA; FL?); in Conocarpus erecta (buttonwood) Distribution. Cuba. Endemic and Laguncularia racemosa (white mangrove) Cuba localities. Cuba localities. CA: Baraguá Collections. Vienna (type) Collections. Ottawa H. quadrispinosus Blackman 1928: 191 Distribution. Cuba. Endemic Scolytus Geoffroy 1762: 309 Cuba localities. Ci: Cayamas (t.l.) S. dimidiatus Chapuis 1869: 57 Collections. Washington (type) A Checklist of the Beetles of Cuba 235

Micracis LeConte 1868: 164 Distribution. Cuba (introduced), USA (FL); M. cubensis Blackman 1928: 193 native to se Asia Distribution. Cuba. Endemic Cuba localities. SC: Santiago, Roig Botani- Cuba localities. Ci: Cayamas (t.l.) cal Garden Collections. Washington (type) Collections. Gatineau. Ottawa M. swainei Blackman 1920: 32 C. carpophagus (Hornung) 1842: 116 (Bostrichus) Distribution. Cuba, Honduras, México, C. thrinacis Hopkins 1915: 46 (Isle of Pines (t.l.), Canada (ON), USA (TX-NY-FL) Washington (type)) Cuba localities. C. bakeri Hopkins 1915: 46 (Havana (t.l.), Wash- Collections. ington (type)) C. anonae Hopkins 1915: 46 (Cuba (t.l.), Wash- Pseudothysanoes Blackman 1920: 46 ington (type)) P. insularis (Blackman) 1943: 359 (Cryptocleptes) Distribution. Cuba (introduced), Hispaniola, Distribution. Cuba. Endemic Jamaica, Puerto Rico, Virgin Islands, Bermuda, Cuba localities. Ci: Cayamas (t.l.) Grenada, México to Honduras, USA (DC-FL); Collections. Washington (type) native to se Asia; in palm seeds P. minor (Blackman) 1928: 207 Cuba localities. SC Distribution. Cuba. Endemic Collections. Gatineau. Ottawa Cuba localities. Ci: Cayamas (t.l.) C. cyperi (Beeson) 1929: 230 Collections. Washington (type) Distribution. Jamaica, Puerto Rico, (P. securigerus (Blackman) 1943: 364) Guadeloupe, México to Honduras, Brazil, USA Distribution. Hispaniola, Puerto Rico, USA (LA-FL) (FL) Cuba localities. SC Cuba localities. Collections. Gatineau. Ottawa Collections. C. dactyliperda (Fabricius) 1801: 387 (Bostrichus) Distribution. Cuba (introduced), Bahamas, SUBTRIBE IPINA Jamaica, Puerto Rico, México to Panamá,USA (CA-FL); circumtropical in seeds of Phoenix Ips DeGeer 1775: 190 palms; native to Africa I. calligraphus (Germar) 1824: 461 (Tomicus) Cuba localities. Ips interstitialis Eichoff 1869: 273; Vàzquez et Collections. al. 1992: 45 C. rhizophorae (Hopkins) 1915: 48 (Spermatoplex) Distribution. Cuba, Bahamas, Dominican Distribution. México, Panamá, Galapagos, Republic, Jamaica, México, Honduras, wide- USA (FL), Asia; in seeds of red mangrove (Woo- spread USA (sw, TX-NY-FL), Canada (PQ) druff 1970), and other seeds Cuba localities. Gu: M: and PR. This spe- Cuba localities. cies is indicated as a pest of weakened pine Collections. trees, and an associate of the pathenogenic fun- C. robustus Eichhoff 1878: 313 gus Ceratocystis (Vàzquez et al. 1992: 45) C. cylindricus Schedl 1949: 116 (Caicos (t.l.), Collections. Vienna (type)) I. grandicollis (Eichhoff) 1868a: 402 (Tomicus) Distribution. Cuba (introduced), Puerto Rico; Distribution. Cuba, Bahamas, Hispaniola, native to Old World; in seeds of Euterpe globosa Jamaica, México to Nicaragua, widespread palm USA; an important pest of Pinus tropicalis Cuba localities. Cuba localities. Collections. Collections. Dendrocranulus Schedl 1938: 165 SUBTRIBE DRYOCOETINA D. carbonarius (Ferrari) 1867: 41 (Xylocleptes) Distribution. Cuba, Jamaica, USA (FL) Cocotrypes Eichhoff 1878: 308 Cuba localities. Ci: Cayamas C. advena Blandford 1849: 100 Collections. Washington Thamnurgides cubanus Eggers 1934: 79 (Sierra Rosario, Cuyajabas, (t.l.); Washington, (type)) 236 Arthropods of Florida and Neighboring Land Areas, Vol. 18

SUBTRIBE CRYPTURGINA Collections. Gatineau. Ottawa. Washington (type) Crypturgus Erichson 1836: 60 X. ferrugineus (Fabricius) 1801: 388 (Bostrichus) C. alutaceus Schwarz 1894a: 17 Distribution. Cuba, Bahamas, Hispaniola, Distribution. Cuba (t.l.), USA (TX-NJ-FL) Jamaica, Puerto Rico, widespread México, Cen- Cuba localities. “Cuba”; no other data tral America, South America, Africa, USA (TX- Collections. Vienna (type) MI-MA-FL); in many tree species Cuba localities. SC SUBTRIBE XYLEBORINA Collections. Gatineau. Ottawa (X. pubescens Zimmerman 1868: 145) Ambrosiodmus Hopkins 1915: 55 Distribution. Bahamas (Andros), widespread (A. devexulus (Wood) 1978: 398 (Xyleborus)) e USA Distribution. Hispaniola, Puerto Rico, USA Cuba localities. Hosts: in Pinus species (FL) Collections. Cuba localities. X. spinulosus Blandford 1898: 201 Collections. Distribution. Cuba, Hispaniola, Jamaica, A. lecontei Hopkins 1915: 56 Grenada, Guadeloupe A. gundlachi Eggers 1931: 20 (Cuba (t.l.), Berlin Cuba localities. SC (type)) Collections. Gatineau. Ottawa Distribution. Cuba, Bahamas, Hispaniola, X. volvulus (Fabricius) 1775: 454 (Bostrichus) Jamaica, Puerto Rico, Guadeloupe, USA (LA- Distribution. Cuba, Jamaica, Puerto Rico, FL) widespread México, Central and South America, Cuba localities. USA (FL) Collections. Cuba localities. Collections. Premnobius Eichhoff 1878: 404 X. xylographus (Say) 1826: 256 (Bostrichus) P. cavipennis Eichhoff 1878: 404 X. inermis Eichhoff 1868: 401 (Cuba, t.l.; Ham- Distribution. Cuba (introduced), Jamaica, burg, type lost) Puerto Rico, México, Brazil, USA (FL); native Distribution. Cuba, Guadeloupe, Puerto Rico, to Africa; in wood of hardwoods USA (TX-MN-FL); mostly in oaks Cuba localities. Cuba localities. M: Cienaga Zapata, Playa Collections. Larga Collections. Washington Theoborus Hopkins 1915: 57 (T. ricini (Eggers) 1932: 298 (Xyleborus)) Xylosandrus Reitter 1913: 83 Distribution. Hispaniola, México to Brazil, X. compactus (Eichhoff) 1875: 201 (Xyleborus). The USA (FL) black twig borer Cuba localities. Distribution. Cuba (introduced), Virgin Is- Collections. lands, Brazil, USA (FL to TX); native to Af- rica; in wood of many hardwoods Xyleborinus Reitter 1913: 79 Cuba localities. A pest of increasing impor- X. andrewesi (Blandford) 1896: 227 (Xyleborus) tance in the robusta variety of coffee trees Distribution. Cuba (introduced), Jamaica, (Vàzquez et al. 1992: 46), and it may become a widespread Asia, Africa, Indomalaya problem of major importance Cuba localities. SC: Gran Piedra Collections. Collections. Ottawa (X. zimmermanni (Hopkins) 1915: 67 (Anisandrus)) Distribution. Bahamas, México to Costa Xyleborus Eichhoff 1864: 37 Rica, South America, USA (FL) X. affinis Eichhoff 1868: 401 Cuba localities. Distribution. Cuba (t.l.), Hispaniola, Ja- Collections. maica, Barbados, widespread México, Central and South America, USA (TX-MA-FL); in many SUBTRIBE CRYPHALINA tree species Cuba localities. SC A Checklist of the Beetles of Cuba 237

Cryptocarenus Eggers 1933: 10 Distribution. Cuba (introduced), Bahamas, C. heveae (Hagedorn) 1912: 338 (Stephanoderes) México to Panamá, South America, USA (TX- Trachyderes parvus Blackman 1943: 36 (Ci: FL); native to Old World Cayamas (t.l.), Washington (type) Cuba localities. Distribution. Cuba, Guadeloupe, Virgin Is- Collections. lands, México to Panama to Brazil, USA (FL) H. crudiae (Panzer) 1791: 35 (Bostrichus) Cuba localities. Distribution. Cuba (introduced), Grenada, Collections. Puerto Rico, México to Costa Rica, South C. seriatus Eggers 1933: 10 America, USA (LA-KS-DC-FL); native to Asia; Distribution. Cuba, Hispaniola, Jamaica, in bark and seeds of hardwoods Virgin Islands, México to Costa Rica to Brazil, Cuba localities. USA (TX, FL) Collections. Cuba localities. H. erectus LeConte 1876: 356 Collections. H. cubensis Hopkins 1915: 32 (Ci: Cayamas (t.l.), Washington (type)) Hypocryphalus Hopkins 1915: 8 Distribution. Cuba, St. Thomas, México, H. mangiferae (Stebbing) 1914: 542 (Cryphalus). Venezuela, USA (TX) The mango bark beetle Cuba localities. Distribution. Cuba (introduced), Barbados, Collections. Guadeloupe, México to Panamá, Venezuela, H. eruditus (Westwood) 1836: 34 Brazil, USA (FL); native to Asia; circumtropical Cryphalus obscurus Ferrari 1867: 17 (Cuba (t.l.), in mango bark Vienna (type)) Cuba localities. Ho: Holguin. A pest of plan- H. lineatifrons Hopkins 1915: 17 (Ci: Cayamas tations of young mango trees in Havana Prov- (t.l.), Washington (type)) ince: boring into trunks and branches (Vàzquez H. parvus Hopkins 1915: 17 (Ci: Cayamas. (t.l), et al. 1992: 45) Washington (type)) Collections. Ottawa H. flavipes Hopkins 1915: 18 (Ci: Cayamas (t.l.), Washington (type)) Hypothenemus Westwood 1836: 34 Stephanoderes flavicollis Hopkins 1915: 24 (Ci: (H. arrecae (Hornung) 1842: 117 (Bostrichus)) Cayamas (t.l.), Washington (type)) Distribution. Bahamas, Puerto Rico, Virgin S. elongatus Hopkins 1915: 25 (Ci: Cayamas (t.l.), Islands, Brazil, USA (FL); native to Africa or Washington (type)) Asia; in bark and seeds of hardwoods S. subconcentralis Hopkins 1915: 25 (Ci: Cuba localities. Cayamas (t.l.), Washington (type)) Collections. S. unicolor Hopkins 1915: 25 (Ci: Cayamas (t.l.), H. birmanus (Eichhoff) 1878: 486 (Triarmocerus). Washington (type)) The kaiwe scolytid S. gracilis Eggers 1929: 51 (Cuba (t.l.), Vienna Distribution. Cuba (introduced), Jamaica, (type)) Costa Rica to Panamá, USA (FL); native to SE Distribution. Cuba, Jamaica, Puerto Rico, Asia México to Panamá, South America, Africa, Cuba localities. Asia, USA (CA-TX, MI-NJ-FL) Collections. Cuba localities. H. ceibae Hopkins 1915: 20 Collections. Distribution. Cuba, Grenada, Puerto Rico H. gossyppi (Hopkins) 1915: 25 (Stephanoderes) Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba, México, USA (FL) Collections. Washington (type) Cuba localities. Ci: Cayamas (t.l.) H. columbi Hopkins 1915: 18 Collections. Washington (type) H. abdominalis Hopkins 1915: 18 (Ci: Cayamas H. hirsutus (Wood) 1954; 1020 (Stephanoderes) (t.l.), Washington (type)) Distribution. Cuba, USA (FL) H. brunneipennis Hopkins 1915: 18 (Ci: Cayamas Cuba localities. (t.l.), Washington (type)) Collections. H. amplipennis Hopkins 1915: 19 (Ci: Cayamas H. interstitialis (Hopkins) 1915: 28 (Stephanoderes) (t.l.), Washington (type)) Stephanoderes obliquus Hopkins 1915: 30 (Ci: Cayamas (t.l.), Washington (type)) 238 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Distribution. Cuba, Jamaica, México to Scolytogenes Eichhoff 1878: 475, 497 Panamá, Colombia, Brazil, USA (AL-KS-CT- S. knabi (Hopkins) 1915: 34 (Ernoporides) FL) Distribution. Cuba (introduced), Hispaniola, Cuba localities. Jamaica, Tortola (BWI), Guadeloupe, México, Collections. Central America, Venezuela, USA (FL); prob- H. javanus (Eggers) 1908: 215 (Stephanoderes) ably native to Old World Stephanoderes obesus Hopkins 1915: 30 (Ci: Cuba localities. Cayamas (t.l.), Washington (type)) Collections. Stephanoderes pistor Schedl 1951: 102 (Havana (t.l.), Vienna (type)) Trischidias Hopkins 1915: 7 Distribution. Cuba (introduced), Hispaniola, T. atoma (Hopkins) 1915: 15 (Hypothenemus). The Guadeloupe, México, Venezuela, Brazil, USA Bermuda beetle (FL); native to Africa; in bark and seeds of hard- Distribution. Cuba (introduced), Brazil, wide- woods spread USA Cuba localities. Cuba localities. An important and wide- Collections. spread pest species, especially in Matanzas H. obr Atkinson unpublished Province. A control program reduced its im- Distribution. Cuba, USA (FL) portance to a harmless level (Vàzquez et al. Cuba localities. 1992: 46). Hosts: many diverse tree species Collections. Collections. Havana H. obscurus (Fabricius) 1801: 395 (Bostrichus) Distribution. Cuba, Hispaniola, Jamaica, SUBTRIBE PITYOPHTHORINA Puerto Rico, Virgin Islands, Guadeloupe, México to Panamá, South America, USA (FL) Araptus Eichhoff 1878: 305 Cuba localities. A. cubensis (Blackman) 1942: 191 (Neodryocoetes) Collections. Distribution. Cuba. Endemic H. pilosus Hopkins 1915: 20 Cuba localities. Ci: Cayamas (t.l.) Distribution. Cuba. Endemic Collections. Washington (type) Cuba localities. Ci: Cayamas (t.l.) A. hymenaeae (Eggers) 1933: 9 (Neodryocoetes) Collections. Washington (type) Distribution. Cuba, Jamaica, Hispaniola, H. seriatus (Eichhoff) 1872: 133 (Stephanoderes) Lesser Antilles, Panama to Brazil Stephanoderes minutus Hopkins 1915: 26 (Ci: Cuba localities. Ha: Marianao, from Ficus Cayamas (t.l.), Washington (type)) sp. Hosts: in seeds of Hymenaea, Erythrina, Stephanoderes nitidipennis Hopkins 1915: 29 (Ci: etc. Cayamas (t.l.), Washington (type)) Collections. Ottawa. S. nitidulus Hopkins 1915: 29 (Ci: Cayamas (t.l.), A. pallidus (Blackman) 1942: 193 (Neodryocoetes) Washington (type)) A. devius Schedl 1972: 61 (PR: Valle Ancon (t.l.), S. subopacicollis Hopkins 1915: 30 (Ci: Cayamas Vienna (type)) (t.l.), Washington (type)) Distribution. Cuba, Hispaniola, Puerto Rico, Distribution. Cuba, Hispaniola, Puerto Rico, Brazil Virgin Islands, Barbados, México to Costa Rica, Cuba localities. Ci: Cayamas (t.l.) South America, USA (TX-KY-WV-FL) Collections. Washington (type) Cuba localities. A. politus (Blandford) 1904: 244 (Pityophthorus) Collections. Distribution. Cuba, Hispaniola, Jamaica, H. setosus (Eichhoff) 1868: 391 (Hypoborus) México, Costa Rica, USA (FL) Distribution. Cuba (introduced), Hispaniola, Cuba localities. Jamaica, Puerto Rico, México to Panamá, South Collections. America, USA (FL); native to Africa Cuba localities. Pityophthorus Eichhoff 1864: 39 Collections. P. concentralis Eichhoff 1878: 188 H. squamosus (Hopkins) 1915: 26 (Stephanoderes) Distribution. Cuba, Bahamas, USA (FL) Distribution. Cuba, México, USA (FL, TX) Cuba localities. Cuba localities. Ci: Cayamas (t.l.) Collections. Hamburg (type lost) Collections. Washington (type) P. pudens (Blackman) 1942: 199 (Pityophthoroides) A Checklist of the Beetles of Cuba 239

Distribution. Cuba, Hispaniola, Virgin Is- Distribution. Cuba, Hispaniola, Jamaica, lands Puerto Rico, México to Argentina, USA (CA- Cuba localities. Ci: Cayamas (t.l.) TX-FL); cosmotropical Collections. Washington (type) Cuba localities. P. pulicarius Zimmermann 1868: 144 Collections. P. cubensis Schedl 1972: 65; PR: Viñales (t.l.), Vienna (type) REFERENCES Distribution. Cuba, widespread USA (TX- Alonso-Zarazaga, M.A. and C.H.C. Lyal. 1999. MN-MA-FL), Canada (NS) A world catalogue of families and genera of Cuba localities. Curculionoidea (Insecta: Coleoptea) (excepting Collections. Scolytidae and Platypodidae). Entomopraxis, P. regularis Blackman 1942: 206 S.C. P. Barcelona, Spain. Distribution. Cuba. Endemic Atkinson, T.H. and Peck, S.B. 1994. Annotated Cuba localities. Ci: Cayamas (t.l.) checklist of the bark and ambrosia beetles (Co- Collections. Washington (type) leoptera: Platypodidae and Scolytidae) of tropi- P. n. sp. cal southern Florida. Florida Ent. 77: 313-329. Distribution. Cuba. Endemic Bright, D.E. 1985. Studies in West Indian Cuba localities. Gu: Moa; Mts. N of Baracoa Scolytidae (Coleoptera) 3. Checklist of Scolytidae Collections. Ottawa of the West Indies, with description of new spe- cies and taxonomic notes. Ent. Arb. Mus. Frey SUBTRIBE CORTHYLINA 33/34: 169-187. Buchanan, L. L. 1947. Notes on Cuban Corthylus Erichson 1836: 64 Curculionidae with descriptions of new genera C. papulans Eichoff 1869: 280 and species (Coleoptera). Mem. Soc. Cubana C. spinifer Schwarz 1891: 114 Hist. Natur. 19: 43-51. Distribution. Cuba, Bahamas, Grenada, Carbonnel, R.M. 1982. Bionomics of the rice wa- México to Panamá, South America, USA (FL) ter weevil Lissorhoptrus brevirostris (SFFR) Cuba localities. PR: Sierra Rangel in Cuba. Int. Rice Res. Newsl. 7: 20-21. Collections. Ottawa Clark, W. E. 1981. The genus Chionanthus Pierce (Coleoptera: Curculionidae): a new species from Microcorthylus Ferrari 1867: 58 Cuba and a key to species. Proc. Ent. Soc. M. minimus Schedl 1950: 160 Washington 83: 690-693. Distribution. Cuba, Jamaica, México to Costa Clark, W.E. 1990. The Neotropical species of Rica, Brazil Pseudanthonomus Dietz (Coleoptera: Cuba localities. Ci: Cayamas Curculionidae). Trans. Amer. Ent. Soc. 116: Collections. Washington 655-696. Clark, W.E. 1991a. The Anthonomus guttatus Monarthrum Kirsch 1866: 213 species group (Coleoptera: Curculionidae). Proc. M. mali (Fitch) 1855: 326 (Tomicus) Ent. Soc. Wash. 93: 262-271. Distribution. Cuba, USA (widespread) Clark, W.E. 1991b. The Anthonomus curvirostris Cuba localities. species group (Coleoptera: Curcurlionidae). Collections. Trans. Amer. Ent. Soc. 117: 39-66. Clark, W.E. 1993a. Huaca, new genus Clark, of SUBFAMILY PLATYPODINAE Neotropical Anthonomini (Coleoptera: The ambrosia beetles Curculionidae). Trans. Amer. Ent. Soc. 119: 1- Members of this subfamily bore galleries un- 45. der bark and feed on ambrosia fungi. Clark, W.E. 1993b. The weevil genus Neomastix Platypus Herbst 1793: 128 Dietz (Coleoptera: Curculionidae: Anthonomini). P. parallelus (Fabricius) 1801: 384 Coleop. Bull. 47: 1-19. P. poeyi Guérin-Méneville 1838: 183; Jacquelin Howden, A. T. 1970. The Tanymecini of the West du Val 1857: 237; Gundlach 1891: 98 (syntypes Indies (Coleoptea: Curculionidae). Cont. Amer. in London and Brussels) Ent. Soc. 5: 1-73. P. subcostatus Jacquelin du Val 1857: 238 (Cuba Howden, A. T. 1992. Review of the New World (t.l.)) eyeless weevils with uncinate tibiae (Coleoptera, 240 Arthropods of Florida and Neighboring Land Areas, Vol. 18

Curculionidae: Molytine, Cryptorhynchinae, Orthognathus. Amer. Mus. Novitates. No. Cossoninae). Mem. Ent. Soc. Canada 162, 76 2441, 1-78. pp. Vàzquez, L.L., E. Valdés, and J.C. Amor. 1992. Kuschel, G. 1952. Revision de Lissorhoptrus New manifestations of pests on economically LeConte y generos vecinos de America. Rev. important plants during the period from 1970 Chilena Ent. 1: 23-74. to 1991 in Cuba. Boll. Lab. Ent. Agr. Filippo Kuschel, G. 1955: Nuevas sinonomias y Silvestri 49: 41-52. anotaciones sobre Curculionidea (Coleoptera). Wibmer, G.J. 1981. Revision of the New World Rev. Chilena Ent. 4: 261-312. weevil genus Tyloderma in America north of Lopez Castila, R. 1992. Dos nuevas especies de Mexico (Coleoptera: Curculionidae: Pachnaeus (Coleoptera: Curculionidae) de Cuba. Cryptorhynchinae). Southwest. Entomol., Suppl Poeyana 421: 8 pp. 3: 1-95. Meneses, R. 1981. Ecological studies of Wood, S.E. 1979. A catalog of the Coleoptera of Lissorhoptrus brevirostris (Coleoptera: America north of México, Family Platypodidae. Curculionidae) in rice region of southern Sancti USA Dept. Agric. Handbook 529-141. Spiritus, Cuba. Agrotechnica de Cuba 13: 23- Wood, S.E. and D.E. Bright, Jr., 1992. A cata- 35. log of Scolytidae and Platypodidae, part 2: Taxo- O’Brien, C.W. and G.J. Wibmer. 1982. Anno- nomic Index. Naturalist Mem. 13, tated checklist of the weevils (Curculionidae vol. A and vol. B. 1483 pp. sensu lato) of North America, Central America, and the West Indies (Coleoptera: Curculionidae). Mem. Amer. Ent. Inst. 34. 382 ORDER STREPSIPTERA pp. These insects are in leaf-hoppers O’Brien, C.W. and G.J. Wibmer. 1989. Revision and various plant-feeding insects as well as other of the Neotropical genus Neohydronomus groups such as wasps and ants (Alayo 1977, Hustache (Coleoptera: Curculionidae). Coleop. Kathirithamby 1992, Zayas 1981). Because larvae Bull. 43: 291-304. and females are endoparasitic, the strepsipterans Osella, G. 1973. Una nuova specie di cossonino have been thought to have potential in Cuba as cieco dell’isola di Cuba Coleoptera- biological control organisms, especially against pests Curculionidae. Résultats des expédition of sugarcane. Additional species occur elsewhere in biospéologiquess Cubano-Roumaines à Cuba. 1: the Caribbean and will eventually be found in Cuba 373-377. (for instance, 10 species occur in Florida Osella, G. 1976. Decuanellus viti n.sp. di (Kathirithamby and Peck 1994) and many of these curculionide endogeo dell’isola di Guadelupa would be expected to occur in Cuba). This order is (Coleoptera). Rev. Suisse Zool. 83: 673-677. included here because, in the past, it has been con- Osella, G. 1977a. Psuedocaecocossonus zayasi n. sidered by some authors to be a family within the gen., n.sp. di Curculionidae endogeo di Cuba beetles. Kukalova-Peck and Lawrence (2004: 111) (Coleoptera). Résultats des expédition review and add to evidence that Strepsiptera are biospéologiquess Cubano-Roumaines à Cuba. 2: the sister group to Coleoptera. 393-395. Osella, G. 1977b. Nuove specie di Curculionidae endogei dell’isola di Cuba (Coleoptera). Résultats SUBORDER MENGENILLIDA des expédition biospéologiquess Cubano- 1. FAMILY MENGENILLIDAE Roumaines à Cuba. 2: 397-402. This family does not occur in the New World. Osella, G. 1980. Nuovi curculionidi ciechi della regione neotropicale (Coleoptera). Frag. Ent. 15: SUBORDER STYLOPIDIA 365-422. 2. FAMILY CORIOXENIDAE Osella, G. 1981; Il genere Stenommatus Wollaston Triozocera Pierce 1911: 490 a Cuba (Coleoptera: Curculionidae). Résultats T. mexicana (Pierce) 1909: 86 (Trioxcera); Peck des expédition biospéologiquess Cubano- 1997: 9 Roumaines à Cuba. 3: 181-183. T. texana Pierce 1911: 491, Luna de Carvalho Vaurie, P. 1970: Weevils of the tribe Sipalini (Co- 1967: 22 leoptera, Curculionidae, Rhynchophorinae). Distribution. Cuba, Puerto Rico, Bermuda, Part 2. The genera Mesocordylus and Trinidad, México, Guatemala, Brazil. Parasi- A Checklist of the Beetles of Cuba 241

toid of Pangaeus bilineatus (Cydnidae, Hemi- 8. FAMILY STYLOPIDAE ptera) This family is cosmopolitan in Distribution. Cuba localities. Ci: Soledad. SC: Santiago de They all parasitize Hymenoptera. They are not yet Cuba, Roig Botanical Garden. SS: Montes de known from Cuba. Trinidad, Mina de Carlota Collections. Gatineau REFERENCES (T. tecpanensis Brailovsky and Márquez 1974; Alayo D., P. 1977. Introduccion al estudio del Kifune and Brailovsky 1987: 65) Orden Strepsiptera en Cuba. Acad. Cien. Cuba, Distribution. México Inst. Zool., Informe Cien.-Tech. 8. Cuba localities. Possible misidentification of Genaro, J.A. and S.B. Peck. 1995. Caenocholax above record from Mina de Carlota fenyesi en Cuba (Strepsiptera: Myrmecolacidae). Collections. Cocuyo 2: 10. Kathirithamby, J. 1992. Strepsiptera of Panamá 3. FAMILY BOHARTILLIDAE and . pp. 421-431. In: Quintero, This family is known only from Honduras, and D. and A. Aiello (eds.). The Insects of Panamá in amber from the Dominican Republic. It may yet and Mesoamerica. Oxford Univ. Press, New be found in Cuba. York. Kathirithamby, J. and S.B. Peck. 1994. 4. FAMILY MYRMECOLACIDAE Strepsiptera of south Florida and the Bahamas Caenocholax Pierce 1909: 88 with the description of a new genus and new C. fenyesi Pierce 1909: 89; Genaro and Peck 1995:10 species of Corioxenidae. Can. Ent. 126: 125-134. Distribution. Cuba, Bahamas through Kifune, T., and H. Brailovsky. 1987. Two new Florida to Arizona, to Argentina and Chile. A species of Mexican Strepsiptera in the collec- parasitoid of Pheidole ants tion of the Instituto de Biologia, Universidad Cuba localities. Ci: Soledad Nacional Autonoma de México (Notulae Collections. Strepsipterologicae – IX). Esakia 20: 61-71. Kukalova-Peck, J. and Lawrence, J.F. 2004. 5. FAMILY HALICTOPHAGIDAE Relationships among coleopteran suborders and Stenocranophilus Pierce 1914: 126 major endoneopteran lineages: evidence from S. quadratus Pierce 1914: 127; Bruner, hind wing characters. European Journal Ento- Scaramuzza and Otero 1945: 160 mology 101: 95144. Distribution. Cuba, Puerto Rico, Trinidad, Luna de Carvalho, Ed. 1967. Terceira Venezuela. An endoparasite of the plant hop- contribuição para o estudo dos Estrepsípteros per delphacid Saccharosydre saccharivora angolenses (Insecta Strepsiptera). Publ. Cyult. (Westwood), a pest of sugarcane Cia. Diamante Angola 77: 13-56. Cuba localities. Peck, S.B. 1997. Triozocera mexicana in Cuba Collections. (Strepsiptera: Corioxenidae). Cocuyo 6: 9. Zayas, F. de. 1981. Entomofauna Cubana. Vol. 8. 6. FAMILY CALLIPHARIXENIDAE Parte tercera. Orden Strepsiptera. Ed. Cien.- This family is known only from southeast Asia. Tech., La Havana.

7. FAMILY ELENCHIDAE This family is cosmopolitan in Distribution. They parasitize Homoptera, and are not yet known from Cuba.