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The Isopods of Abyssal Depths in the Atlantic Ocean

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The Isopods of Abyssal Depths in the Atlantic Ocean The Isopods of Abyssal Depths in the Atlantic Ocean by ROBERT J. MENZIES Department of Biology, University of Southern California SUMMARY 83 23. Haploniscus polaris, n.sp. 105 24. Haploniscus telus, n.sp. 105 INTRODUCTION 84 25. Haploniscus tricornoides, n.sp. 106 Materials and Methods 84 26. Haploniscus tropicalis, n.sp. 106 Definition of the Area Sampled 85 27. Haploniscus rugosus, n.sp. 106 History of Previous Work 85 28. Haploniscus ovalis, n.sp. 106 3. Genus: Antennuloniscus, n.gen. 107 LIST OF ISOPODS FROM DEPTHS GREATER THAN 2000 METERS A Key to the Species of Antennuloniscus 107 BY REGION 87 29. Antennuloniscus dimeroceras (Barnard) 107 30. Antennuloniscus armatus, n.sp. 108 SYSTEMATICS 89 31. Antennuloniscus rostratus, n.sp. 109 Suborder: Gnathiidea Monod 89 32. Antennuloniscus ornatus, n.sp. 109 A Key to the Fourteen Described Species of Gnathiidea 4. Genus: Hydroniscus Hansen 109 Known from Depths of 500 Meters and Greater 89 A Key to the Species of Hydroniscus 109 1. Genus: Gnathia Leach 90 33. Hydroniscus ornatus, n.sp. 109 1. Gnathia vemae, n.sp. 90 34. Hydroniscus quadrifrons, n.sp. 109 2. Gnathia caeca Richardson 90 35. Hydroniscus abyssi Hansen 110 3. Gnathia stygia (G. O. Sars) 90 Family: Ischnomesidae 111 4. Gnathia albescenoides, n.sp. 92 A Key to the Genera of the Ischnomesidae 111 Suborder: Quatuordecapoda (Isopoda, sensu strictu) 92 5. Genus: Ischnomesus Richardson 111 Tribe: Asellota 92 A Key to the Species of Ischnomesus 111 A Key to the Subtribes of the Tribe Asellota 92 36. Ischnomesus profundus Hansen 112 A Key to the Families of the Tribe Asellota, Subtribe 37. Ischnomesus bispinosus (G. O. Sars) 112 Paraselloidea 94 38. Ischnomesus armatus Hansen 113 Family: Haploniscidae 94 39. Ischnomesus caribbicus, n.sp. 113 2. Genus: Haploniscus Richardson 94 40. Ischnomesus simplissimus, n.sp. 113 A Key to the Species of Haploniscus 95 41. Ischnomesus multispinis, n.sp. 113 5. Haploniscus bicuspis (G. O. Sars) 96 42. Ischnomesus wolffi, n.sp. 113 6. Haploniscus unicornis Menzies 96 43. Ischnomesus magnificus, n.sp. 114 7. Haploniscus excisus Richardson 96 44. Ischnomesus bidens, n.sp. 115 8. Haploniscus spinifer Hansen 98 45. Ischnomesus paucispinis, n.sp. 116 9. Haploniscus minutus, n.sp. 99 46. Ischnomesus decemspinosus, n.sp. 117 10. Haploniscus spatulifrons, n.sp. 99 47. Ischnomesus elegans, n.sp. 117 11. Haploniscus tuberculatus, n.sp. 99 6. Genus: Haplomesus Richardson 117 12. Haploniscus tridens, n.sp. 99 A Key to the Species of Haplomesus 117 13. Haploniscus elevatus, n.sp. 99 48. Haplomesus angustus Hansen 119 14. Haploniscus quadrifrons, n.sp. 100 49. Haplomesus modestus Hansen 119 15. Haploniscus princeps, n.sp. 101 50. Haplomesus quadrispinosus (G. O. Sars) 119 16. Haploniscus percavix, n.sp. 101 51. Haplomesus tenuispinis Hansen 119 17. Haploniscus tricornis, n.sp. 101 52. Haplomesus tropicalis, n.sp. 120 18. Haploniscus nondescriptus, n.sp. 102 53. Haplomesus insignis Hansen 121 19. Haploniscus acutus, n.sp. 103 54. Haplomesus ornatus, n.sp. 121 20. Haploniscus parallelus, n.sp. 103 55. Haplomesus bifurcatus, n.sp. 121 21. Haploniscus capensis, n.sp. 103 7. Genus: Heteromesus Richardson 121 22. Haploniscus trituberculatus, n.sp. 103 A Key to the Species of Heteromesus 121 80 ROBERT J. MENZIES 56. Heteromesus longiremis Hansen 122 102. Storthyngura magnispinis (Richardson) 147 57. Heteromesus bifurcatus, n.sp. 122 103. Storthyngura triplispinosa, n.sp. 149 58. Heteromesus granulatus Richardson 122 104. Storthyngura symmetrica, n.sp. 149 59. Heteromesus spinescens Richardson 123 105. Storthyngura vemae, n.sp. 149 60. Heteromesus similis Richardson 123 106. Storthyngura birsteini, n.sp. 149 8. Genus: Stylomesus Wolff 123 107. Storthyngura snanoi, n.sp. 149 A Key to the Species of Stylomesus 123 13. Genus: Syneurycope Hansen 150 61. Stylomesus inermis (Vanhoffen) 123 A Key to the Species of Syneurycope 151 62. Stylomesus granulosus, n.sp. 124 108. Syneurycope heezeni, n.sp. 151 63. Stylomesus spinulosus, n.sp. 125 109. Syneurycope parallela Hansen 151 64. Stylomesus elegans, n.sp. 126 110. Syneurycope hanseni Menzies 151 65. Stylomesus regularis, n.sp. 126 111. Syneurycope multispina, n.sp. 152 66. Stylomesus simplex, n.sp. 126 14. Genus: Acanthocope Beddard 152 67. Stylomesus productus, n.sp. 127 A Key to the Species of Acanthocope 153 68. Stylomesus simulans, n.sp. 127 112. Acanthocope'argentinae, n.sp. 154 Family: Macrostylidae 127 113. Acanthocope unicornis, n.sp. 155 9. Genus: Macrostylis G. O. Sars 128 114. Acanthocope annulalus, n.sp. 155 A Key to the Species of Macrostylis 128 Family: Ilyarachnidae 155 69. Macrostylis truncatex, n.sp. 128 15. Genus: Ilyarachna G. O. Sars 156 70. Macrostylis abyssicola Hansen 129 115. Ilyarachna abyssorum Richardson 156 71. Macrostylis setifer, n.sp. 130 116. Ilyarachna bicornis Hansen 156 72. Macrostylis hirsuticaudis, n.sp. 131 117. Ilyarachna spinosissima Hansen 157 73. Macrostylis caribbicus, n.sp. 131 118. Ilyarachna longicornis G. O. Sars 157 74. Macrostylis vemae, n.sp. 131 119. Ilyarachna derjugini Gurjanova 158 75. Macrostylis bifurcatus, n.sp. 131 120. Ilyarachna nodifronoides, n.sp. 158 76. A4acrostylis minutus, n.sp. 131 121. Ilyarachna qfricana, n.sp. 159 77. Macrostylis bipunctatus, n.sp. 132 122. Ilyarachna spinoajricana, n.sp. 160 Family: Nannoniscidae 133 123. Ilyarachna simplex, n.sp. 160 A Key to the Genera of Nannonisridae 1 33 124. Ilyarachna multispinosa, n.sp. 160 10. Genus: Nannoniscnides Hansen 133 125. Ilyarachna argentinae, n.sp. 160 78. Nannoniscoides hinutus. n.sp. 133 126. Ilyarachna gurjanovae, n.sp. 160 1 1. Genus: Nannoniscus G. O. Sars 133 127. Ilyaiachna identijrons, n.sp. 161 A Key to the Species of Nannoniscus 1 34 128. Ilyaiachna tiiangulaia, n.sp. 161 79. Nannoniscus inermis Hansen 13 > 129. Ilyarachna scotia, n.sp. 162 80. Nannoniscus nrmatus Hansen 135 family: Desmosomidae 162 81. Nannoniscus analis Hansen 135 16. Genus: Desmosoma G. O. Sars 162 82. Nannoniscus oblongus G. O. Sars 136 A Key to the Species of Desmosoma 163 83. Nannoniscus camayae. n.sp. 137 130. Desmosoma giaciltpcs Hansen 163 84. Nannoniscus laevis, n.sp. 138 131. Desmosoma insigne Hansen 163 85. Nannoniscus primiticus, n.sp. 138 132. Desmosoma simile Hansen 164 86. Nannoniscus spinicornis Hansen 138 133. Desmosoma intermedium Hult 165 Family: Kurycopidae 138 134. Desmosoma coarcticum (G. O. Sars) 165 A Key to the Genera of the Kurycopidae 138 135. Desmosoma longispinum Hansen 165 12. Genus: Eurycope G. O. Sars 139 136. Desmosoma striata, n.sp. 165 A Kev to the Abyssal Species of Eurycope 13'' 137. Desmosoma magnispinum, n.sp. 165 87. Eurycope antarctica Vanhoffen 139 138. Desmosoma birsteini, n.sp. 166 88. Eurycope furcata G. O. Sars 140 Family: Dendrotionidae 167 89. Eurycope producta G. O. Sars 141 A Key to the Genera of the Dendrotionidae 167 90. Eurycope vacarius Vanhoffen 141 17. Genus: Dendromunna, n.gen. 167 91. Eurycope complanata Bonnier 141 139. Dendromunna spinipes, n.sp. 167 92. Eurycope murrayi Walker 141 Family: Ianirellidae 167 93. Eurycope parva Bonnier 142 18. Genus: lanirella Bonnier 168 94. Eurycope nodifrons Hansen 143 A Key to the Species of lanirella 168 95. Eurycope hanseni Ohlin 143 140. lanirella lobata Richardson 168 96. Eurycope incisa Gurjanova 143 141. lanirella bifida, n.sp. 168 97. Eurycope acutitelson, n.sp. 143 142. lanirella laevis Hansen 168 98. Eurycope nodosa, n.sp. 145 143. lanirella vemae Menzies 169 99. Eurycope ovaloides, n.sp. 145 144. lanirella magnifrons, n.sp. 169 Genus: Storthyngura Vanhoffen 145 19. Genus: Spinianirella, n.gen. 170 A Key to the Species of Storthyngura 145 145. Spinianirella walfishensis, n.sp. 171 100. Storthyngura digitata, n.sp. 146 20. Genus: Rhacura Richardson 171 101. Storthyngura truncata (Richardson) 146 146. Rhacura pulchra Richardson 172 ABYSSAL ATLANTIC ISOPODS 81 Family: Munnidae 172 Tribe: Flabellifera 186 A Key to the Genera of the Munnidae 173 A Key to the Flabelliferan Subtribes 186 21. Genus: Munna Kroyer 173 Family: Serolidae 186 147. Munna [Munna) acanthi/era Hansen 173 33. Genus: Serolis Leach 186 148. Munna (Munna) argentinae, n.sp. 173 164. Serolis (Serolis) neaera Beddard 186 22. Genus: Acanthomunna Beddard 174 165. Serolis (Serolis) margaretae, n.sp. 187 A Key to the Species of Acanthomunna 175 166. Serolis (Serolis) macdonnellae, n.sp. 188 149. Acanthomunna beddardi, n.sp. 176 167. Serolis (Serolis) vemae, n.sp. 189 23. Genus: Notoxenoides, n.gen. 176 168. Serolis (Serolis) maryannae, n.sp. 189 150. Notoxenoides abyssi, n.sp. 177 34. Genus: Glabroserolis, n.gen. 189 151. Notoxenoides vemae, n.sp. 177 169. Glabroserolis specialis, n.sp. 189 Family: Acanthaspidae, new family 177 Subtribe: Anthuroidea 190 24. Genus: Acanthaspida Stebbing 177 A Key to the Genera of the Anthuroidea Having Abyssal A Key to the Species of Acanthaspidia 177 Species (after Barnard) 190 152. Acanthaspidia bifurcata, n.sp. 177 35. Genus: Pseudanthura Richardson 191 153. Acanthaspidia decorata Hansen 178 170. Pseudanthura lateralis Richardson 191 Family: Abyssianiridae Menzies 179 36. Genus: Calathura Norman and Stebbing 191 25. Genus: Abyssianira Menzies 179 171. Calathura brachiata (Stimpson) 191 154. Abyssianira dentifrons, Menzies 179 37. Genus: Anthelura Norman and Stebbing 192 155. Abyssianira argentenensis, n.sp. 179 172. Anthelura truncata (Hansen) 192 26. Genus: Xostylus, n.gen. 179 38. Genus: Ananthura Barnard 193 156. Xostylus parallelus, n.sp. 181 173. Ananthura abyssorum (Norman and Stebbing) Family: Ianiridae 181 193 27. Genus: Ianira Leach 181 39. Genus: Hyssura Norman and Stebbing 193 157. Ianira hanseni, n.sp. 181 174. Hyssura producta Norman and Stebbing 193 28. Genus: Abyssijaera, n.gen. 181 40. Genus: Leptanthura G. O. Sars 193 158. Abyssijaera ocarina, n.sp. 181 Subtribe: Cirolanoidea 194 Family: Echinothambemidae Menzies 183 Tribe: Valvifera 194 29. Genus: Echinothambema Menzies 183 Family: Arcturidae 194 159. Echinothambema ophiuroides Menzies 183 41. Genus: Antarcturus zur Strassen 194 30. Genus: Vemathambema, n.gen. 183 42. Genus: Mesidothea Richardson 195 160. Vemathambema elongata, n.sp.
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